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A Checklist of Diatom Species Reported (and Presumed Native) from Canadian Coastal Waters
L. Mather, K. MacIntosh, I. Kaczmarska, G. Klein and J.L. Martin
Fisheries and Oceans Canada Science Branch Maritimes Region St. Andrews Biological Station 531 Brandy Cove Road St. Andrews, NB Canada E5B 2L9
2010
Canadian Technical Report of Fisheries and Aquatic Sciences 2881
Canadian Technical Report of Fisheries and Aquatic Sciences
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Canadian Technical Report of Fisheries and Aquatic Sciences 2881
March 2010
A Checklist of Diatom Species Reported (and Presumed Native)
from Canadian Coastal Waters
by
L. Mather1, K. MacIntosh1, I. Kaczmarska1, G. Klein1 and J.L. Martin
Fisheries and Oceans Canada, Maritimes Region, St. Andrews Biological Station,
531 Brandy Cove Road, St. Andrews, NB Canada E5B 2L9
1Dept. Biology, Mount Allison University, 63B York St., Sackville, NB Canada E4L 1G7
This is the two hundred and ninetieth Technical Report of the Biological Station, St. Andrews, NB
ii
© Her Majesty the Queen in Right of Canada, 2010
Cat. No. Fs 97-6/2881E…………………ISSN 0706-6457 (print version)
Cat. No. F597-6/2881E-PDF…………ISNN 1488-5379 (on-line version) Correct citation for this publication: Mather, L., MacIntosh, K., Kaczmarska, I., Klein, G. and Martin, J.L. 2010. A checklist of diatom species reported (and presumed native) from Canadian coastal waters. Can. Tech. Rep. Fish. Aquat. Sci. 2881: iii + 78p.
iii
ABSTRACT
Mather, L., MacIntosh, K., Kaczmarska, I., Klein, G. and Martin, J.L. 2010. A checklist of diatom species reported (and presumed native) from Canadian coastal waters. Can. Man. Rep. Fish. Aquat. Sci. 2881: iii+ 78 p.
The transportation of phytoplankton via ballast water is one of the main vectors allowing organisms to establish themselves in new environments. Since the 1970s, both the frequency of ocean crossings and the volume transported has increased. Therefore, it is to be expected that the potential for invasion would have also increased. Although reviews on the species richness of native biota have been done for some areas of the world, a clear and concise list of the presumed native biota for Canadian waters has yet to be documented. The aim of this report is to provide a list of diatom species found as their names appear in the cited literature, as well as their current valid name, from the Canadian and the neighbouring north, east, and west coasts of North America. This will allow for a better understanding of the presumed native diatoms, and thus the potentially non-native diatom species, documented in Canadian waters. As well, it provides a time-line for when these organisms were documented as occurring in our waters.
RÉSUMÉ
Mather, L., MacIntosh, K., Kaczmarska, I., Klein, G. and Martin, J.L. 2010. A checklist of diatom species reported (and presumed native) from Canadian coastal waters. Can. Tech. Rep. Fish. Aquat. Sci. 2881: iii+ 78 p.
Le transport des eaux de ballast est un des vecteurs principal dans le transfert du phytoplancton qui permet à ces organismes de s’établir dans un nouvel environnement. Depuis les années 1970 la capacité et la fréquence des navires long courrier ont augmenté, ainsi que le potentiel d’invasion. Ailleurs au monde, des recensements de la richesse des espèces natives ont été achevés mais jusqu’à date il n’y a aucune liste documentée pour les diatomées présumées indigènes dans les eaux canadiennes. Le but de ce rapport est de dresser une liste des diatomées trouvées dans les eaux canadiennes et des côtes adjacentes du nord, est et ouest de l’Amérique du Nord. Cette liste comprend le nom de l’espèce comme il apparaît dans la littérature citée ainsi que le nom courant valide. Elle offre une meilleure compréhension des diatomées dites natives et subséquemment des espèces non indigènes dans les eaux canadiennes. Également, cette liste permet de dresser une ligne temporelle depuis quand ces organismes ont étés observés dans nos eaux canadiennes.
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INTRODUCTION
The confirmed introduction of non-indigenous and toxic or harmful phytoplankton species via ballast water in Australia (Hallegraeff et al. 1988) and other waters around the world has established a growing concern regarding our ability to recognize newcomers for the North American coasts (Edlund et al. 2000, Mills et al. 1993, Ricciardi 2001). The possibility for invasion of a species is not only determined by whether or not a transfer vector is available to the organism, but whether or not the environmental conditions, such as nutrient availability and temperature, of the new location facilitate survival. These introduced organisms pose obvious ecological risks to native habitats but the risk is further magnified by the potential that such organisms may become toxigenic or otherwise harmful once introduced. However, before identifying a species as alien, a good understanding of species richness in native biota is needed. Examples of documentation of native species include: diatoms of the United States (Patrick and Reimer 1966, 1975), diatoms of Europe (Hustedt 1930, 1939, Krammer 2000, 2002, Lange-Bertalot 2001) and diatoms of British coast (Hendey 1964). Such systematic work from the microbial community is yet to be undertaken for marine diatoms in Canadian waters. This work was initiated through a project funded by the Canadian Aquatic Invasive Species Network (CAISN) to study the potential for diatoms to invade Canadian marine and freshwater environments through the transport of ballast water. CAISN is an organization that is conducting a comprehensive study examining and identifying existing invasions with the goal of preventing future invasions from foreign locations into Canadian ports. The overall mission of CAISN is to reveal to the public the results of an expansive network of research and to raise awareness and initiate policy changes when it comes to aquatic invasive species. As part of this project, ballast water samples from ships coming to the ports of Halifax and Montreal (East Coast), Vancouver (West Coast), and the Great Lakes were examined for the potential of diatoms species to survive and thrive in these tanks. As this research was conducted, a list of species’ names and references was collected to aid in mapping the potential for change in species’ composition in our waters. This report documents diatoms (Bacillariophyta) that have been observed in Canada and the neighbouring north, east and west coasts of North America, and thus would be expected to naturally occur in Canadian waters. Reported species are documented, including: 1) the diatom species name listed as it appears in its original publication; 2) the species current and valid name; and, 3) the common synonyms, verified through the two taxonomic databases: algae base and catalogue of life (algaebase.org, catalogueoflife.com). Use of invalid names or synonyms can cause erroneous or confusing reports of species of micro-biota. Diatom species reported here were collected from a total of thirty published sources that represent the years spanning 1910-2008 (see Appendix 1), and five lists obtained through personal communications (B. Ginn, Dept. of Biology, Queen's University, 116 Barrie St., Kingston, ON Canada K7L 3N6; L.A. Hobson, University of Victoria, PO Box 1700 STN CSC, Victoria, BC Canada V8W 2Y2; R.A. Horner, University of Washington, Box 357940, Seattle, WA 98195-7940; J.L. Martin, Biological Station, 531 Brandy Cove Road, St. Andrews, NB Canada E5B 2L9; and C. Rafuse, National Research Council Institute for Marine Biosciences, 1411 Oxford Street, Halifax, NS Canada, B3H 3Z1). Ballast waters from transoceanic vessels can be a major means by which diatoms are transported to ports throughout
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the world. The volume and frequency of these ocean crossing vessels has significantly increased since the 1970s, with increasing introductions at the close of the 20th century (Carlton 1996), a time span sufficient for microorganisms such as diatoms to establish themselves in Canadian waters. Examples from the macroscopic level include species such as the zebra mussel (Dreissena polymorpha) and the Emerald ash boring beetle (Agrilus planipennis) - non-indigenous invaders that are now well established in Canada. There is growing evidence that the potential for invasions to occur at a microscopic level exists as well - as has been documented for the species Didymosphenia geminata (Lyngbye) M. Schmidt (Blanco and Ector 2009) and Thalassiosira baltica, (Edlund et. al. 2000) - all of which necessitates more research focusing on microbes, both native and those arriving from exotic locations. In this list, some diatoms presumed by many to be non-indigenous diatoms, have been found to be reported in some locations in Canada as early as 1910. One such example is Achnanthes subsessilis K., (Bailey 1910). Another example is Pseudo-nitzschia fraudulenta (Cleve) Hasle, a species thought to be cosmopolitan, but was first observed in the Bay of Fundy as recently as the year 2008 (Martin and LeGresley 2008). Some species such as Tabellaria flocculosa (Roth) Kützing, (Hay et. al. 2003; Bailey 1913) are most likely native to our waters as they have been reported both before and after 1970, when the volume and frequency of ocean crossing vessels began to increase substantially (Carlton 1996). As more organisms become established, it is increasingly difficult to determine whether or not species are indigenous or introduced. Thus, uncovering the time-lines to which these species occur can indicate the possibility for establishment of new species in our coastal waters as well as the potential for species (native/non-native) to sustain themselves within the environment. More species, established through ballast water transport, are indicated in a manuscript written by Klein et al. (2009). We appreciate that some taxa included in this report might have been introduced centuries ago given that seafaring communities have existed on the Canadian East coast since the 1600s. Until presented with evidence to the contrary, we assume these species to be indigenous. The goal of this compilation is to provide baseline data of our current understanding of marine diatom species composition in Canada. This will aid in recognizing non-native arrivals found through various vectors including incoming ships’ ballast water and sediments.
MATERIALS AND METHODS
Books and papers collected from scientific journals were checked for species’ lists. As well, various colleagues were contacted and contributed their personal species’ lists. Details on the collection of samples are described in the books and papers cited in this manuscript and the names of the colleagues who contributed their lists are listed in the Acknowledgements.
RESULTS
Approximately 893 valid names are listed, representing 160 diatom genera that include 825 species and 69 subspecies level taxa. Of these, some are marine and brackish, likely collected in
3
situ, but others are known to be from freshwater and most likely introduced into the marine environment from inland sources. Appendix I presents the species’ names as they are found originally as well as current accepted names and synonyms. Each species’ name has been cited indicating the source of the information and the known geographic location in Canada (East, West or Arctic) of the species is documented.
DISCUSSION
This checklist of species represents the start of documenting species currently observed in North American marine waters, specifically those found on the Canadian marine coasts. The intent of this publication is to make the present data available for examining the composition of diatom species in Canadian waters, both present and from the past. This will provide valuable baseline data on native diatom species and give a greater understanding of the potential for introduction of new organisms from foreign locations that may exist in our coastal regions and establish themselves. This list may also aid in identifying species that are cryptogenic, that is species that have not been confidently identified as being native or introduced (Carlton 1996). Cryptogenic species are ones that have an unknown origin, or an origin that is not explicitly clear. It seems inevitable that marine organisms that are moved transoceanically into habitats wherever vectors implicated in their transport have gone would be classified as being cryptogenic (Carlton 1996). The understanding that cryptogenic species may be more common than originally thought makes it possible for one to significantly underestimate the impacts of biological invasions on natural and previously invaded communities (Carlton 1996). Neglecting this information may lead to a misleading interpretation of invasion corridors, the types of communities susceptible or resistant to invasions, and invasion success rates (Carlton 1996). Although we have included the vast majority of papers and books that list species from the North American marine coast, specifically from Canadian waters, we acknowledge that this may not be representative of all the data available and as more research is being conducted more species may be detected.
ACKNOWLEDGEMENTS
Partial funding for this work was through Canadian Aquatic Invasive Network (CAISN) to I. Kaczmarska. The CAISN organization is partners with a number of organizations including Natural Sciences and Engineering Research Council of Canada (NSERC), the Department of Fisheries and Oceans, Transport Canada, the Ontario Ministry of the Environment, and Ontario Ministry of Natural Resources. We gratefully acknowledge the following colleagues who kindly shared their species lists, in alphabetic order of surname: Brian Ginn (PhD research at Queen's University pending his acceptance), Louis Hobson (University of Victoria), Rita Horner (University of Washington), and Cheryl Rafuse (Dalhousie University). We thank Rita Horner and Murielle LeGresley for reviewing; Charlotte McAdam for editing and Joanne Power for formatting the manuscript.
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357
940,
Sea
ttle,
WA
981
95-7
940
4 Raf
use
(per
s. co
mm
.) - C
. Raf
use,
Nat
iona
l Res
earc
h C
ounc
il - I
nstit
ute
for M
arin
e B
iosc
ienc
es, 1
411
Oxf
ord
Stre
et, H
alifa
x, N
S C
anad
a B
3H 3
Z15 H
obso
n (p
ers.
com
m.)-
L.A
. Hob
son,
Uni
vers
ity o
f Vic
toria
, PO
Box
170
0 ST
N C
SC, V
icto
ria, B
C C
anad
a V
8W 2
Y2
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Ach
nant
hes
bahu
sien
sis
(Gru
now
) La
nge-
Ber
talo
t A
chna
nthe
s ba
husi
ensi
s (G
runo
w)
Lang
e-B
erta
lot (
ab)
Nav
icul
a ko
enig
ii B
rock
man
n,
Nav
icul
a ba
husi
ensi
s (G
runo
w)
Cle
ve, A
chna
nthe
s ps
eudo
sole
a S
imon
sen
Eas
t (B
oF)
Gin
n (p
ers.
com
m.)
1
Ach
nant
hes
bior
etii
Ger
mai
nA
chna
nthe
s bi
oret
ii G
erm
ain
(ab)
Nav
icul
a ro
taea
na v
ar. e
xcen
trica
G
runo
w, N
avic
ula
vanh
eurc
kii
R.M
. Pat
rick
Arc
ticA
nton
iade
s et
al.
2005
Ach
nant
hes
brev
ipes
Ach
nant
hes
brev
ipes
C. A
gard
h (a
b)C
ymbo
sira
aga
rdhi
i K
ützi
ng,
Ach
nant
ella
bre
vipe
s (A
gard
h)
Gai
llon,
Ach
nant
hidi
um b
revi
pes
(Aga
rdh)
Hei
berg
, Ach
nant
hidi
um
brev
ipes
(A
gard
h) C
leve
Eas
tB
rune
l 196
2 (B
aile
y 19
10, C
arve
r &
Mal
let 2
002)
Ach
nant
hes
brev
ipes
var
. in
term
edia
Ach
nant
hes
brev
ipes
var
. in
term
edia
(K
ützi
ng) C
leve
(ab)
Ach
nant
hes
inte
rmed
ia K
ützi
ng,
Ach
nant
hes
subs
essi
lis K
ützi
ng
Eas
tB
rune
l 196
2
Ach
nant
hes
broe
nlun
dens
is F
oged
Ach
nant
hes
broe
nlun
dens
is F
oged
(a
b)A
rctic
Ant
onia
des
et a
l. 20
05
Ach
nant
hes
chlid
anos
Hoh
n &
H
elle
rman
Psa
mm
othi
dium
chl
idan
os (
M.H
. H
ohn
& H
elle
rman
) Lan
ge-B
erta
lot
(ab)
Ach
nant
hes
chlid
anos
M.H
. Hoh
n &
Hel
lerm
anA
rctic
Ant
onia
des
et a
l. 20
05
Ach
nant
hes
flexe
lla K
ützi
ngE
ucoc
cone
is fl
exel
la (K
ützi
ng)
Mei
ster
(ab)
Ach
nant
hes
flexe
lla (
Küt
zing
) B
run
Arc
ticA
nton
iade
s et
al.
2005
Tabl
e I :
Lis
t of s
peci
es d
ocum
ente
d fr
om e
ast,
wes
t and
arc
tic N
orth
Am
eric
an c
oast
al w
ater
s, w
ith e
mph
asis
on
Can
adia
n w
ater
s. (B
oF=
Bay
of F
undy
, US=
U
nite
d St
ates
, per
s. co
mm
.= p
erso
nal c
omm
unic
atio
n, (c
ol)=
cat
alog
ue o
f life
, (ab
)= a
lgae
bas
e). R
efer
ence
s out
side
par
enth
eses
are
the
orig
inal
pap
ers f
or
refe
renc
e of
spec
ies'
nam
e w
hile
refe
renc
es w
ithin
par
enth
eses
are
from
add
ition
al p
aper
s tha
t sup
port
the
pres
ence
of a
spec
ies.
Pag
e 9
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Ach
nant
hes
hauc
kian
a va
r. ro
stra
ta S
hulz
Ach
nant
hes
hauc
kian
a v
ar. r
ostra
ta
Sch
ulz
(col
)E
ast
Zwic
ker 1
998
Ach
nant
hes
helv
etic
a H
uste
dtA
chna
nthe
s he
lvet
ica
(Hus
tedt
) La
nge-
Ber
talo
t (ab
)A
chna
nthe
s au
stria
ca v
ar.
vent
ricos
a K
rass
ke, A
chna
nthe
s au
stria
ca v
ar. h
elve
tica
Hus
tedt
Arc
ticA
nton
iade
s et
al.
2005
Ach
nant
hes
krie
geri
Kra
sske
A
chna
nthe
s kr
iege
ri K
rass
ke (a
b)W
est
(Effi
ngha
m
Inle
t)
Hay
et a
l. 20
03
Ach
nant
hes
kryo
phila
Pet
erse
nA
chna
nthe
s kr
yoph
ila J
.B. P
eter
sen
(ab)
Nav
icul
a se
min
ulum
var
. in
term
edia
Hus
tedt
, Ach
nant
hes
plitv
icen
sis
Hus
tedt
Arc
ticA
nton
iade
s et
al.
2005
Ach
nant
hes
laev
is Ø
stru
pA
chna
nthe
s la
evis
Øst
rup
(ab)
Arc
ticA
nton
iade
s et
al.
2005
Ach
nant
hes
long
ipes
Aga
rdh
Ach
nant
hes
long
ipes
Aga
rdh
(ab)
Eas
t/Wes
tLi
nkle
tter e
t al.
1977
(Bai
ley
1910
, 19
13)
Ach
nant
hes
mar
ginu
lata
Gru
now
in
Cle
ve &
Gru
now
Ach
nant
hes
mar
ginu
lata
Gru
now
in
Cle
ve &
Gru
now
(ab)
Arc
ticA
nton
iade
s et
al.
2005
Ach
nant
hes
min
utis
sim
a K
ützi
ngA
chna
nthi
dium
min
utis
sim
um
(Küt
zing
) Cza
rnec
ki (a
b)A
chna
nthe
s m
inut
issi
ma
Küt
zing
Wes
t (E
ffing
ham
In
let)/
Arc
tic/E
ast
Hay
et a
l. 20
03 (A
nton
iade
s et
al.
2005
, Cat
tane
o et
al.
2004
)
Ach
nant
hes
pete
rsen
ii H
uste
dtR
ossi
thid
ium
pet
erse
nii
(Hus
tedt
) R
ound
& L
. Buk
htiy
arov
a ex
Abo
al
(ab)
Ach
nant
hes
pete
rsen
ii H
uste
dt,
Ach
nant
hes
hust
edtii
Bily
&
Mar
van,
Ach
nant
hes
jako
vlej
evic
ii H
uste
dt, A
chna
nthe
s kr
yoph
ila
var.
dens
estri
ata
Hus
tedt
, A
chna
nthe
s pa
ralle
la J
.R. C
arte
r, A
chna
nthe
s pr
ocer
a H
uste
dt,
Ach
nant
hes
pusi
lla v
ar.
pete
rsen
ii (H
uste
dt) L
ange
-B
erta
lot,
Ach
nant
hes
son
yda
B.
Har
tley,
Nav
icul
a de
nses
triat
a H
uste
dt, A
chna
nthe
s gr
ubei
S
imon
sen
Arc
ticA
nton
iade
s et
al.
2005
Pag
e 10
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Ach
nant
hes
ross
ii H
uste
dtA
chna
nthe
s ro
ssii
Hus
tedt
(ab)
Ach
nant
hes
cass
ida
Car
ter
Arc
ticA
nton
iade
s et
al.
2005
Ach
nant
hes
spp.
Ach
nant
hes
spp.
Eas
t/Arc
ticB
ates
& S
train
200
6 (B
ooth
&
Hor
ner 1
997)
Ach
nant
hes
subs
essi
lis K
.A
chna
nthe
s br
evip
es v
ar.
inte
rmed
ia (
Küt
zing
) Cle
veA
chna
nthe
s su
bses
silis
Küt
zing
Eas
tB
aile
y 19
10 (B
aile
y 19
13)
Ach
nant
hes
taen
iata
A
chna
nthe
s ta
enia
ta G
runo
w (a
b)A
chna
nthe
s ta
enia
ta v
ar.
hype
rbor
ea G
runo
w, A
chna
nthe
s hy
perb
orea
(Gru
now
) Gru
now
Eas
t (B
oF)/A
rctic
Car
ver &
Mal
let 2
002
(Bur
sa 1
963,
G
ran
& B
raar
ud 1
935)
Ach
nant
hes
vent
ralis
Kra
sske
Ach
nant
hes
vent
ralis
(K
rass
ke)
Lang
e-B
erta
lot
(ab)
Ach
nant
hes
kryo
phila
var
. pr
otra
cta
Hus
tedt
, Ach
nant
hes
subl
aevi
s H
uste
dt, N
avic
ula
vent
ralis
Kra
sske
, Ach
nant
hes
subl
aevi
s va
r. cr
assa
C.W
. R
eim
er
Arc
ticA
nton
iade
s et
al.
2005
Ach
nant
hidi
um b
iaso
letti
anum
(G
runo
w in
Cle
ve a
nd G
runo
w)
Rou
nd &
Buk
htiy
arov
a
Ach
nant
hidi
um b
iaso
letti
anum
(G
runo
w) F
.E. R
ound
& L
. B
ukht
iyar
ova
(ab)
Ach
nant
hidi
um b
iaso
letti
anum
(G
runo
w) L
ange
-Ber
talo
t, A
chna
nthi
dium
bia
sole
ttian
um
(Küt
zing
) Buk
htiy
arov
a, M
icro
neis
bi
asol
ettia
na (K
ützi
ng) M
eist
er,
Mic
rone
is b
iaso
letti
ana
(Küt
zing
) C
leve
, Ach
nant
hes
bias
olet
tiana
G
runo
w, F
alca
tella
bia
sole
ttian
a (K
ützi
ng) R
aben
hors
t, S
yned
ra
bias
olet
tiana
Küt
zing
Wes
t (E
ffing
ham
In
let)
Hay
et a
l. 20
03
Act
inoc
yclu
s cu
rvat
ulus
Jan
isch
Act
inoc
yclu
s cu
rvat
ulus
Jan
isch
Act
inoc
yclu
s su
boce
llatu
s (G
runo
w) R
attra
y, C
osci
nodi
scus
cu
rvat
ulus
var
. sub
ocel
latu
s G
runo
w
Wes
t/Arc
ticS
ance
tta &
Cal
vert
1988
(Lov
ejoy
et
al.
2002
, McQ
uoid
& H
obso
n 20
01)
Act
inoc
yclu
s eh
renb
ergi
Ral
fsA
ctin
ocyc
lus
octo
nariu
s E
hren
berg
(a
b)A
ctin
ocyc
lus
ehre
nber
gii
Ral
fsE
ast
Gra
n &
Bra
arud
193
5
Act
inoc
yclu
s sp
.A
ctin
ocyc
lus
sp.
Arc
ticR
iede
l et a
l. 20
03A
ctin
ocyc
lus
tenu
issi
mus
Act
inoc
yclu
s te
nuis
sim
us C
leve
Eas
t/Arc
ticB
érar
d-Th
erria
ult e
t al.
1999
(L
ovej
oy e
t al.
2002
)
Pag
e 11
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Act
inoc
yclu
s oc
tona
rius
Ehr
enbe
rgA
ctin
ocyc
lus
octo
nariu
s E
hren
berg
(a
b)A
ctin
ocyc
lus
ehre
nber
gii
Ral
fsW
est
Hob
son
2009
Act
inop
tych
us s
enar
ius
Ehr
enbe
rgA
ctin
opty
chus
sen
ariu
s (E
hren
berg
) E
hren
berg
(ab)
Act
inop
tych
us u
ndul
atus
Bai
ley
ex H
uste
dt, A
ctin
ocyc
lus
sena
rius
Ehr
enbe
rg, A
ctin
ocyc
lus
undu
latu
s J.
W. B
aile
y,
Act
inoc
yclu
s un
dula
tus
Küt
zing
, A
ctin
opty
chus
und
ulat
us (
J.W
. B
aile
y) R
alfs
Eas
t/Wes
t H
orne
r et a
l. 20
05 (L
inkl
ette
r et a
l. 19
77, S
ance
tta &
Cal
vert
1988
, M
cQuo
id &
Hob
son
2001
, Hay
et
al. 2
003)
Act
inop
tych
us s
p.A
ctin
opty
chus
sp.
Arc
ticB
ursa
196
3A
ctin
opty
chus
spl
ende
ns
(Sha
dbol
t) R
alfs
A
ctin
opty
chus
spl
ende
ns (S
hadb
olt)
Ral
fs e
x P
ritch
ard
(ab)
Act
inop
tych
us h
alio
nyx
Gru
now
, A
ctin
opty
chus
qua
tuor
dena
rius
Ehr
enbe
rg, A
ctin
osph
aeni
a sp
lend
ens
Sha
dbol
t
Wes
t (U
S)/E
ast
Cup
p 19
43 (M
cQuo
id &
Hob
son
2001
, Car
ver &
Mal
let 2
002)
Act
inop
tych
us u
ndul
atus
(B
aile
y)
Ral
fsA
ctin
opty
chus
sen
ariu
s (E
hren
berg
) E
hren
berg
(ab)
Act
inop
tych
us u
ndul
atus
(J.
W.
Bai
ley)
Ral
fs
Eas
t/Wes
t (U
S)/A
rctic
Cup
p 19
43 (B
aile
y 19
10, 1
913,
B
ursa
196
3, G
ran
& B
raar
ud
1935
)A
ctin
opty
chus
vul
garis
Sch
uman
nA
ctin
opty
chus
vul
garis
Sch
uman
n (a
b)W
est
(Effi
ngha
m
Inle
t)
Hay
et a
l. 20
03
Am
phip
leur
a pe
lluci
da K
utz.
Am
phip
leur
a pe
lluci
da (K
ützi
ng)
Küt
zing
(ab)
Ber
kele
ya p
ellu
cida
(Küt
zing
) G
iffen
, Car
rodo
ria p
ellu
cida
(K
ützi
ng) K
untz
e, A
ulac
ocys
tis
pellu
cida
(Küt
zing
) Has
sall,
N
avic
ula
pellu
cida
(Küt
zing
) E
hren
berg
, Fru
stul
ia p
ellu
cida
K
ützi
ng
Eas
tB
aile
y 19
13
Am
phip
leur
a si
gmoi
dea
W. S
m.
Am
phip
leur
a si
gmoi
dea
W. S
mith
Eas
tB
aile
y 19
10 (B
aile
y 19
13)
Am
phip
rora
ala
ta K
utz.
Ent
omon
eis
alat
a (E
hren
berg
) E
hren
berg
Am
phip
rora
ala
ta (E
hren
berg
) K
ützi
ngE
ast
Bai
ley
1910
(Bai
ley
1913
)
Am
phip
rora
gig
ante
a va
r. su
lcat
a (O
'Mea
ra) C
leve
Am
phip
rora
gig
ante
a v
ar. s
ulca
ta
(O'M
eara
) Cle
ve (a
b)W
est (
US
)C
upp
1943
Am
phip
rora
hyp
erbo
rea
Gra
nA
mph
ipro
ra h
yper
bore
a (G
runo
w)
Gra
n (c
ol)
Am
phip
rora
pal
udos
a va
r. hy
perb
orea
Gru
now
A
rctic
Bur
sa 1
963
(Boo
th &
Hor
ner 1
997)
Am
phip
rora
pal
udos
aA
mph
ipro
ra p
alud
osa
W. S
mith
Eas
tB
rune
l 196
2
Pag
e 12
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Am
phite
tras
sp.
Am
phite
tras
sp.
Wes
tH
obso
n 20
09 (M
cQuo
id &
Hob
son
2001
)A
mph
ora
sp.
Am
phor
a sp
.E
ast
Bru
nel 1
962
Am
phor
a sp
p.A
mph
ora
spp.
Arc
tic/W
est
Love
joy
et a
l. 20
02 (M
cQuo
id &
H
obso
n 20
01)
Am
orph
a af
finis
Kut
z.A
mph
ora
libyc
a E
hren
berg
(ab)
Am
phor
a af
finis
Küt
zing
E
ast
Bai
ley
1913
Am
phor
a co
ffeae
form
is (
C.A
. A
gard
h) K
ützi
ng
Am
phor
a ap
onin
a K
ützi
ng (a
b)Fr
ustu
lia c
offe
aefo
rmis
C. A
gard
h W
est
(Effi
ngha
m
Inle
t)
Hay
et a
l. 20
03
Am
phor
a co
stat
a W
. S.
Am
phor
a co
stat
a W
. Sm
ith (a
b)E
ast
Bai
ley
1913
Am
phor
a he
lene
nsis
Giff
enA
mph
ora
hele
nens
is M
.H. G
iffen
(a
b)W
est
(Effi
ngha
m
Inle
t)
Hay
et a
l. 20
03
Am
phor
a in
arie
nsis
Kra
mm
erA
mph
ora
inar
iens
is K
ram
mer
(ab)
Arc
ticA
nton
iade
s et
al.
2005
Am
phor
a la
evis
var
. lae
viss
ima
Am
phor
a la
evis
var
. lae
viss
ima
(Gre
gory
) Cle
veA
rctic
Rie
del e
t al.
2003
(Lov
ejoy
et a
l. 20
02)
Am
phor
a lib
yca
Ehr
enbe
rgA
mph
ora
libyc
a E
hren
berg
(ab)
Am
phor
a co
pula
ta (
Küt
zing
) S
choe
man
& A
rchi
bald
, Am
phor
a ov
alis
var
. lib
yca
(Ehr
enbe
rg)
Cle
ve, A
mph
ora
oval
is v
ar. a
ffini
s (K
ützi
ng) v
an H
eurc
k, A
mph
ora
affin
is K
ützi
ng, F
rust
ulia
cop
ulat
a K
ützi
ng
Arc
ticA
nton
iade
s et
al.
2005
Am
phor
a ob
tect
a H
.L.S
.A
mph
ora
obte
cta
J.W
. Bai
ley
(ab)
Eas
tB
aile
y 19
13
Am
phor
a ov
alis
Kut
z.A
mph
ora
oval
is (K
ützi
ng) K
ützi
ng
(ab)
Am
phor
a am
phor
a (C
.G.
Ehr
enbe
rg) J
. Pan
tocs
ek,
Am
phor
a oc
ella
ta E
hren
berg
, N
avic
ula
amph
ora
Ehr
enbe
rg,
Frus
tulia
ova
lis K
ützi
ng
Eas
t (B
oF)/W
est
Bai
ley
1913
(McQ
uoid
& H
obso
n 20
01, B
aile
y 19
10)
Pag
e 13
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Am
phor
a pe
dicu
lus
(Küt
zing
) G
runo
w e
x S
chm
idt
Am
phor
a pe
dicu
lus
(Küt
zing
) G
runo
w e
x A
. Sch
mid
t (ab
)A
mph
ora
oval
is v
ar. p
edic
ulus
(K
ützi
ng) P
ero,
Enc
yone
ma
pedi
culu
s (K
ützi
ng) H
. Per
agal
lo,
Am
phor
a ov
alis
var
. ped
icul
us
(Küt
zing
) van
Heu
rck,
Cym
bella
ca
espi
tosu
m v
ar. p
edic
ulus
(K
ützi
ng) B
run,
Cym
bella
pe
dicu
lus
Küt
zing
, Am
phor
a ov
alis
var
. ped
icul
us K
ützi
ng
Arc
ticA
nton
iade
s et
al.
2005
Am
phor
a ro
bust
aA
mph
ora
robu
sta
Gre
gory
(ab)
Eas
tC
arve
r & M
alle
t 200
2A
mph
ora
spitz
berg
ensi
s V
anLa
ndin
gham
Am
phor
a sp
itzbe
rgen
sis
Van
Land
ingh
am (a
b)A
rctic
Ant
onia
des
et a
l. 20
05
Am
phor
a sp
.A
mph
ora
sp.
Eas
tB
rune
l 196
2A
mph
ora
spp.
Am
phor
a sp
p.E
ast
Mar
tin (p
ers.
com
m.)2
Ara
chno
idis
cus
ehre
nber
gii
Bai
ley
ex E
hren
berg
Ara
chno
idis
cus
ehre
nber
gii
Bai
ley
ex E
hren
berg
(ab)
Wes
t (U
S)
Cup
p 19
43
Arc
ocel
lulu
s co
rnuc
ervi
sA
rcoc
ellu
lus
corn
ucer
vis
G.R
. Has
le,
H.A
. von
Sto
sch
& E
.E. S
yver
tsen
(a
b)
Eas
t/Arc
ticB
érar
d-Th
erria
ult e
t al.
1999
(L
ovej
oy e
t al.
2002
)
Ast
erio
nella
form
osa
Ast
erio
nella
form
osa
Has
sall
(ab)
Ast
erio
nella
gra
cilli
ma
var
. fo
rmos
a (H
assa
ll) W
islo
uch
Eas
tC
atta
neo
et a
l. 20
04
Ast
erio
nella
gra
cilli
ma
Hei
b.A
ster
ione
lla g
raci
llim
a (H
antz
sch)
H
eibe
rg (a
b)
Arc
tic/E
ast
Bur
sa 1
963
(Gra
n &
Bra
arud
19
35)
Ast
erio
nella
japo
nica
Ast
erio
nello
psis
gla
cial
is (
F.
Cas
traca
ne) F
.E. R
ound
Eas
tB
rune
l 196
2 (G
ran
& B
raar
ud
1935
)A
ster
ione
lla k
aria
na G
runo
wA
ster
opla
nus
karia
nus
(Gru
now
) C.
Gar
dner
& R
.M. C
raw
ford
(ab)
Ast
erio
nella
kar
iana
Gru
now
, A
ster
ione
llops
is k
aria
na (G
runo
w)
F.E
. Rou
nd
Arc
tic/E
ast
Bur
sa 1
963
(Gra
n &
Bra
arud
19
35)
Ast
erio
nello
psis
gla
cial
isA
ster
ione
llops
is g
laci
alis
(F.
C
astra
cane
) F.E
. Rou
ndA
ster
ione
lla g
laci
alis
F.
Cas
traca
ne, A
ster
ione
lla ja
poni
ca
Cle
ve
Eas
t/Wes
tB
érar
d-Th
erria
ult e
t al.
1999
(H
orne
r 200
2)
Ast
erio
nello
psis
kar
iana
Ast
erop
lanu
s ka
rianu
s (G
runo
w) C
. G
ardn
er &
R.M
. Cra
wfo
rd (a
b)E
ast
Car
ver &
Mal
let 2
002
Ast
erio
nello
psis
soc
ialis
(Le
win
&
Nor
ris) R
ound
Ast
erio
nella
soc
ialis
Lew
in &
Nor
ris
(ab)
Wes
tH
orne
r (pe
rs. c
omm
.)3
Ast
erol
ampr
a sp
.A
ster
olam
pra
sp.
Arc
ticB
ursa
196
3
Pag
e 14
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Ast
erol
ampr
a m
aryl
andi
ca
Ehr
enbe
rgA
ster
olam
pra
mar
ylan
dica
E
hren
berg
(ab)
Ast
erol
ampr
a va
nheu
rcki
i B
run
Wes
t (U
S)/E
ast
Cup
p 19
43
Ast
erom
phal
us fl
abel
latu
sA
ster
omph
alus
flab
ella
tus
(Bré
biss
on) G
revi
lle (a
b)S
pata
ngid
ium
flab
ella
tum
B
rébi
sson
E
ast
Car
ver &
Mal
let 2
002
Ast
erom
phal
us h
epta
ctis
(B
rébi
sson
) Ral
fsA
ster
omph
alus
hep
tact
is
(Bré
biss
on) R
alfs
(a
b)A
ster
olam
pra
areo
lata
A. M
ann,
A
ster
olam
pra
hept
actis
Gre
ville
, A
ster
omph
alus
retic
ulat
us C
leve
, S
pata
ngid
ium
hep
tact
is
Bré
biss
on, S
pata
ngid
ium
ra
lfsia
num
G. N
orm
an,
Ast
erol
ampr
a ra
lfsia
na (
G.
Nor
man
) Gru
now
Wes
tC
upp
1943
(Hob
son
2009
, S
ance
tta &
Cal
vert
1988
, McQ
uoid
&
Hob
son
2001
)
Ast
erom
phal
us h
yalin
usA
ster
omph
alus
hya
linus
Kar
sten
(a
b)E
ast
Bér
ard-
Ther
riaul
t et a
l. 19
99
Ast
erom
phal
us s
arco
phag
us
Wal
lich
Ast
erom
phal
us s
arco
phag
us W
allic
h (a
b)W
est
Hor
ner (
pers
. com
m.)
3
Atth
eya
arm
atus
(W
est)
Cra
wfo
rdA
tthey
a ar
mat
a (T
. Wes
t) R
. M.
Cra
wfo
rd (a
b)W
est
Hor
ner (
pers
. com
m.)
3
Atth
eya
deco
raA
tthey
a de
cora
T. W
est (
ab)
Eas
tB
érar
d-Th
erria
ult e
t al.
1999
Atth
eya
long
icor
nis
Atth
eya
long
icor
nis
R. M
. Cra
wfo
rd
& C
. Gar
dner
(ab)
Eas
tM
artin
& L
eGre
sley
200
8
Atth
eya
sept
entri
onal
isA
tthey
a se
pten
trion
alis
(Øst
rup)
C
raw
ford
(ab)
Cha
etoc
eros
sep
tent
riona
le
Øst
rup,
Gon
ioce
ros
sept
entri
onal
is (Ø
stru
p) R
ound
, C
raw
ford
& M
ann
Eas
t/Arc
ticB
érar
d-Th
erria
ult e
t al.
1999
(R
iede
l et a
l. 20
03, L
ovej
oy e
t al.
2002
, Boo
th &
Hor
ner 1
997,
R
ozan
ska
et a
l. 20
08)
Atth
eya
sp.
Atth
eya
sp.
Arc
ticLo
vejo
y et
al.
2002
Aul
acod
iscu
s ki
ttoni
i A
rnot
tA
ulac
odis
cus
kitto
nii
Arn
ott e
x R
alfs
(a
b)W
est
Hor
ner (
pers
. com
m.)
3
Aul
acos
eira
am
bigu
a (G
runo
w)
Sim
onse
nA
ulac
osei
ra a
mbi
gua
(Gru
now
) S
imon
sen
(ab)
Mel
osira
am
bigu
a (G
runo
w) O
.F.
Mul
ler
Eas
t (B
oF)
Gin
n (p
ers.
com
m.)
1
Pag
e 15
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Aul
acos
eria
dis
tans
(E
hren
b.)
Sim
onse
nA
ulac
osei
ra d
ista
ns (E
hren
berg
) S
imon
sen
(ab)
Mel
osira
pol
ymor
pha
subs
p.
dist
ans
(C.G
. Ehr
enbe
rg) H
. B
ethg
e, G
allio
nella
dis
tans
E
hren
berg
, Mel
osira
dis
tans
(E
hren
berg
) Küt
zing
, Lys
igon
ium
di
stan
s (E
hren
berg
) Tre
visa
n,
Lysi
goni
um d
ista
ns (E
hren
berg
) K
untz
e, M
elos
ira p
olym
orph
a su
bsp.
dis
tans
(C.G
. Ehr
enbe
rg)
H. B
ethg
e
Eas
tG
inn
(per
s. c
omm
.)1
Aul
acos
eira
gra
nula
taA
ulac
osei
ra g
ranu
lata
(Ehr
enbe
rg)
Sim
onse
n (a
b)M
elos
ira p
olym
orph
a su
bsp.
gr
anul
ata
(C.G
. Ehr
enbe
rg) H
. B
ethg
e, G
ailli
onel
la g
ranu
lata
E
hren
berg
, Mel
osira
gra
nula
ta
(Ehr
enbe
rg) R
alfs
, Mel
osira
pu
ncta
ta v
ar. g
ranu
lata
(E
hren
berg
) Cle
ve &
Möl
ler,
Lysi
goni
um g
ranu
latu
m
(Ehr
enbe
rg) K
untz
e, O
rthos
ira
gran
ulat
a (E
hren
berg
) Sch
onfe
ldt
Eas
t (B
oF)
Car
ver &
Mal
let 2
002
Aul
acos
eira
isla
ndic
aA
ulac
osei
ra is
land
ica
(O.M
ülle
r)
Sim
onse
nE
ast
Bér
ard-
Ther
riaul
t et a
l. 19
99
Aul
acos
eira
spp
.A
ulac
osei
ra s
pp.
Wes
tM
cQuo
id &
Hob
son
2001
Aul
acos
eira
sub
arct
ica
(O. M
ülle
r)
Haw
orth
Mel
osira
ital
ica
subs
p. s
ubar
ctic
a O
.F. M
ülle
r (ab
)A
ulac
osei
ra s
ubar
ctic
a (O
. Mül
ler)
H
awor
thW
est
(Effi
ngha
m
Inle
t)
Hay
et a
l. 20
03
Aul
iscu
s sc
ulpt
us R
alfs
A
ulis
cus
scul
ptus
(W. S
mith
) Ral
fs
ex P
ritch
ard
(ab)
Aul
iscu
s ca
elat
us J
.W. B
aile
y,
Eup
odis
cus
scul
ptus
W. S
mith
E
ast (
BoF
)B
aile
y 19
13 (L
aksh
min
aray
ana
1983
)A
ulis
cus
spin
osus
Chr
is.
Aul
iscu
s sp
inos
us T
.Chr
istia
n (a
b)E
ast
Bai
ley
1913
Aul
iscu
s sp
.A
ulis
cus
sp.
Wes
tS
ance
tta &
Cal
vert
1988
Pag
e 16
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Bac
illar
ia p
arad
oxa
Gm
el.
Bac
illar
ia p
arad
oxa
J.F.
Gm
elin
(ab)
Vib
rio p
axill
ifer
O.F
. Mül
ler,
Osc
illat
oria
pax
illife
r (O
.F. M
ülle
r)
Sch
rank
, Dia
tom
a pa
xilli
feru
m
(O.F
. Mül
ler)
Bré
biss
on, N
itzsc
hia
paxi
llife
r (O
.F. M
ülle
r) H
eibe
rg,
Nitz
schi
a pa
rado
xa (J
.F. G
mel
in)
Gru
now
, Osc
illat
oria
pax
illife
ra
(O.F
. Mül
ler)
Sch
rank
ex
Gom
ont,
Bac
illar
ia p
axill
ifer
(O.F
. Mül
ler)
T.
Mar
sson
, Hom
oeoc
ladi
a pa
xilli
fer
(O.F
. Mül
ler)
Elm
ore,
Bac
illar
ia
paxi
llife
r (O
.F. M
ülle
r) H
ende
y
Eas
tB
aile
y 19
10 (B
aile
y 19
13)
Bac
illar
ia p
axill
ifer
(O.F
. Mül
ler)
H
ende
yB
acill
aria
par
adox
a J
.F. G
mel
in (a
b)O
scill
aria
pax
illife
ra (
O.F
. Mül
ler)
S
chra
nkE
ast/W
est
Hor
ner e
t al.
2005
(Mar
tin [p
ers.
co
mm
.]2 , Car
ver &
Mal
let 2
002,
M
cQuo
id &
Hob
son
2001
)B
acte
riast
rum
com
osum
Pav
illar
dB
acte
riast
rum
com
osum
Pav
. (a
b)W
est
(US
)/Eas
tC
upp
1943
Bac
teria
stru
m d
elic
atul
umB
acte
riast
rum
del
icat
ulum
Cle
ve
(ab)
Eas
t/Wes
tC
arve
r & M
alle
t 200
2 (H
orne
r pe
rs. c
omm
.)3
Bac
teria
stru
m e
long
atum
Cle
veB
acte
riast
rum
elo
ngat
um C
l. (a
b)W
est
(US
)/Eas
tC
upp
1943
(Car
ver &
Mal
let 2
002)
Bac
teria
stru
m fu
rcat
umB
acte
riast
rum
furc
atum
Sha
dbol
t (a
b)B
acte
riast
rum
var
ians
Lau
der
Eas
tC
arve
r & M
alle
t 200
2
Bac
teria
stru
m h
yalin
um L
aude
rB
acte
riast
rum
hya
linum
Lau
d. (a
b)W
est
(US
)/Eas
tC
upp
1943
(Car
ver &
Mal
let 2
002)
Bac
teria
stru
m h
yalin
um v
ar.
prin
ceps
(C
astra
cane
) Ika
riB
acte
riast
rum
hya
linum
var
. pr
ince
ps (
Cas
tr.) I
kari
(ab)
Wes
t (U
S)
Cup
p 19
43
Bac
teria
stru
m s
pp.
Bac
teria
stru
m s
pp.
Eas
tM
artin
(per
s. c
omm
.)2
Bac
tero
sira
bat
hyom
phal
aB
acte
rosi
ra b
athy
omph
ala
(Cle
ve)
E.E
. Syv
erts
en &
G.R
. Has
le (a
b)B
acte
rosi
ra fr
agili
s (G
ran)
Gra
n,
Laud
eria
frag
ilis
Gra
n,
Cos
cino
disc
us b
athy
omph
alus
C
leve
Eas
t/Arc
ticB
érar
d-Th
erria
ult e
t al.
1999
(L
ovej
oy e
t al.
2002
)
Pag
e 17
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Bac
terio
sira
frag
ilis
Gra
nB
acte
rosi
ra b
athy
omph
ala
(Cle
ve)
E.E
. Syv
erts
en &
G.R
. Has
leB
acte
rosi
ra fr
agili
s (G
ran)
Gra
n E
ast
(BoF
)/Arc
ticLi
nkle
tter e
t al.
1977
(Bru
nel 1
962,
B
ursa
196
3, G
ran
& B
raar
ud 1
935,
B
ooth
& H
orne
r 199
7)
Bel
lero
chea
hor
olog
ical
isB
elle
roch
ea h
orol
ogic
alis
von
S
tosc
h (a
b)B
elle
roch
ea m
alle
us (B
right
wel
l) V
an H
eurc
k E
ast
Car
ver &
Mal
let 2
002
Bel
lero
chea
mal
leus
Bel
lero
chea
mal
leus
(B
right
wel
l) V
an H
eurc
kTr
icer
atiu
m m
alle
us B
right
wel
lE
ast
Bér
ard-
Ther
riaul
t et a
l. 19
99
(Bur
sa 1
963)
Ber
kele
ya ru
tilan
s (T
rent
epoh
l) G
runo
wB
erke
leya
rutil
ans
(Tre
ntep
ohl e
x R
oth)
Gru
now
(ab)
Car
rodo
ria ru
tilan
s (R
oth)
Kun
tze,
A
mph
iple
ura
rutil
ans
(Tre
ntep
ohl
ex R
oth)
Cle
ve, C
onfe
rva
rutil
ans
Tren
tepo
hl, B
erke
leya
dill
wyn
ii va
r. ru
tilan
s (T
rent
epoh
l ex
Rot
h)
Eib
en, M
onem
a ru
tilan
s (T
rent
epoh
l ex
Rot
h) M
eneg
hini
, G
irode
lla ru
tilan
s (R
oth)
Gai
llon,
S
chiz
onem
a ru
tilan
s (T
rent
epoh
l ex
Rot
h) A
gard
h, H
ydro
linum
ru
tilan
s(T
rent
epoh
l ex
Rot
h) L
ink,
B
angi
a ru
tilan
s (R
oth)
Lyn
gbye
, C
onfe
rva
rutil
ans
Tre
ntep
ohl e
x R
oth
Eas
t (B
oF)
Car
dina
l et a
l. 19
84
Bid
dulp
hia
alte
rnan
s (B
aile
y) V
an
Heu
rck
Tric
erat
ium
alte
rnan
s J.
W. B
aile
y (a
b)B
iddu
lphi
a al
tern
ans
(J.W
. Bai
ley)
V
an H
eurc
k E
ast/W
est
(US
)C
upp
1943
(Bér
ard-
Ther
riaul
t et
al. 1
999,
Gra
n &
Bra
arud
193
5)B
iddu
lphi
a au
rita
(Lyn
g.) B
réb.
&
God
.O
dont
ella
aur
ita (L
yngb
ye) C
.A.
Aga
rdh
(ab)
Bid
dulp
hia
aurit
a (L
yngb
ye)
Bré
biss
on
Eas
t/Arc
ticR
awle
nce
1987
(Bru
nel 1
962,
Frit
z 19
21, B
ursa
196
3, G
ran
& B
raar
ud
1935
, Bai
ley
1910
, 191
3)
Bid
dulp
hia
aurit
a va
r. ob
tusa
Bid
dulp
hia
obtu
sa (
Küt
zing
) Ral
fs
(ab)
Bid
dulp
hia
aurit
a va
r. ob
tusa
(K
ützi
ng) H
uste
dtE
ast
Bru
nel 1
962
Bid
dulp
hia
baile
yiB
iddu
lphi
a m
obili
ensi
s (J
.W. B
aile
y)
Gru
now
Bid
dulp
hia
baile
yi W
. Sm
ithE
ast
Bai
ley
1910
Bid
dulp
hia
dubi
a (B
right
wel
l) C
leve
Bid
dulp
hia
dubi
a (T
. Brig
htw
ell)
P.T
. C
leve
(ab)
Wes
t (U
S)
Cup
p 19
43
Bid
dulp
hia
laev
isP
leur
osira
laev
is (E
hren
berg
) C
ompè
re (a
b)B
iddu
lphi
a la
evis
Ehr
enbe
rgE
ast
Bai
ley
1913
Pag
e 18
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Bid
dulp
hia
long
icru
ris v
ar. h
yalin
a (S
chrö
der)
Cup
p O
dont
ella
long
icru
ris (
Gre
ville
) H
oban
(ab)
Bid
dulp
hia
long
icru
ris v
ar. h
yalin
a (S
chrö
d.) C
upp
W
est (
US
)C
upp
1943
Bid
dulp
hia
mob
ilien
sis
Odo
ntel
la m
obili
ensi
s (J
.W. B
aile
y)
Gru
now
Bid
dulp
hia
mob
ilien
sis
Gru
now
, Zy
goce
ros
mob
ilien
sis
J.W
. B
aile
y, B
iddu
lphi
a ba
ileyi
W.
Sm
ith
Eas
tFr
itz 1
921
(Bai
ley
1913
)
Bid
dulp
hia
obtu
sa
Bid
dulp
hia
obtu
sa (K
ützi
ng) R
alfs
(a
b)B
iddu
lphi
a ro
peria
na G
revi
lle,
Odo
ntel
la o
btus
a K
ützi
ng,
Bid
dulp
hia
aur
ita v
ar. o
btus
a (K
ützi
ng) H
uste
dt
Wes
t (U
S)
Car
ver &
Mal
let 2
002
Bid
dulp
hia
pulc
hella
Gra
yB
iddu
lphi
a pu
lche
lla S
.F. G
ray
Eas
tB
aile
y 19
10 (C
arve
r & M
alle
t 20
02)
Bid
dulp
hia
regi
a (S
chul
ze)
Ost
enfe
ldO
dont
ella
regi
a (S
chul
tze)
S
imon
sen
Den
ticel
la re
gia
Sch
ultz
e (a
b)E
ast
Gra
n &
Bra
arud
193
5
Bid
dulp
hia
retic
ulum
Bid
dulp
hia
retic
ulum
(Ehr
enbe
rg)
Boy
er (a
b)Tr
icer
atiu
m s
culp
tum
Sha
dbol
t, Tr
icer
atiu
m p
unct
atum
Brig
htw
ell
Eas
tC
arve
r & M
alle
t 200
2
Bid
dulp
hia
rhom
bus
(Ehr
enbe
rg)
W. S
mith
Zygo
cero
s rh
ombu
s E
hren
berg
(ab)
Wes
t (U
S)/E
ast
Cup
p 19
43 (B
aile
y 19
13, C
arve
r &
Mal
let 2
002)
Bra
chys
ira v
itrea
B
rach
ysira
vitr
ea (G
runo
w) R
. Ros
s (a
b)A
nom
oeon
eis
vitre
a (G
runo
w)
Ros
s, A
nom
oeon
eis
varia
bilis
(R
oss)
Rei
mer
, Nav
icul
a va
riabi
lis
R. R
oss,
Gom
phon
ema
vitre
um
Gru
now
Eas
t C
atta
neo
et a
l. 20
04
Bré
biss
onia
boe
ckii
(Ehr
enbe
rg)
Gru
now
Bré
biss
onia
boe
ckii
(Ehr
enbe
rg) E
. O
'Mea
ra (a
b)C
occo
nem
a bo
ecki
i E
hren
berg
, D
oryp
hora
boe
ckii
(Ehr
enbe
rg)
W.S
mith
Eas
tLi
nkle
tter e
t al.
1977
(Bai
ley
1913
, 19
10, C
ardi
nal e
t al.
1984
)
Cal
onei
s ae
roph
ila B
ock
Cal
onei
s ae
roph
ila B
ock
(ab)
Arc
ticA
nton
iade
s et
al.
2005
Cal
onei
s he
ndey
i La
nge-
Ber
talo
tC
alon
eis
hend
eyi
Lang
e-B
erta
lot
(ab)
Arc
ticA
nton
iade
s et
al.
2005
Cal
onei
s sc
hum
ania
na G
run.
Cal
onei
s sc
hum
ania
na (
Gru
now
) C
leve
(ab)
Eas
tB
aile
y 19
13
Cal
onei
s si
licul
a (E
hren
berg
) Cle
veN
avic
ula
silic
ula
Ehr
enbe
rg (a
b)A
rctic
Ant
onia
des
et a
l. 20
05
Cal
onei
s su
bcle
vei
Kra
mm
erC
alon
eis
subc
leve
i K
ram
mer
(ab)
Arc
ticA
nton
iade
s et
al.
2005
Cam
pylo
disc
us c
ribro
sum
C
ampy
lodi
scus
ech
inei
s E
hren
berg
(a
b)C
ampy
lodi
scus
crib
rosu
s W
. S
mith
Eas
tB
aile
y 19
10
Pag
e 19
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Cam
pylo
disc
us fa
stuo
sus
Her
.C
ampy
lodi
scus
fast
uosu
s E
hren
berg
(ab)
Cam
pylo
disc
us th
uret
ii B
rébi
sson
, Cam
pylo
disc
us
parv
ulus
W.S
mith
Eas
tB
aile
y 19
13
Cam
pylo
disc
us ra
lfsii
W. S
m.
Cam
pylo
disc
us ra
lfsii
W.S
mith
(ab)
Eas
tB
aile
y 19
13
Cam
pylo
disc
us s
pira
lis W
. Sm
.C
ampy
lodi
scus
spi
ralis
(Küt
zing
) W.
Sm
ith (a
b)E
ast
Bai
ley
1913
Cam
pylo
disc
us th
uret
iiC
ampy
lodi
scus
fast
uosu
s E
hren
berg
(ab)
Cam
pylo
disc
us th
uret
ii B
rébi
sson
Eas
tB
rune
l 196
2
Cam
pylo
neis
spp
.C
ampy
lone
is s
pp.
Wes
tM
cQuo
id &
Hob
son
1997
(M
cQuo
id &
Hob
son
2001
)C
ampy
losi
ra c
ymbe
llifo
rmis
(A
. S
chm
idt)
Gru
now
Cam
pylo
sira
cym
belli
form
is (
A.
Sch
mid
t) G
runo
w e
x V
an H
eurc
k (a
b)
Syn
edra
cym
belli
form
is
A.S
chm
idt (
col)
Wes
t (U
S)
Cup
p 19
43 (C
arve
r & M
alle
t 200
2)
Cer
atau
lina
berg
oni
H. P
erag
allo
Cer
atau
lina
pela
gica
(Cle
ve)
Hen
dey
(ab)
Cer
atau
lus
berg
onii
H. P
erag
allo
E
ast
Gra
n &
Bra
arud
193
5
Cer
atau
lina
pela
gica
(C
leve
) H
ende
yC
erat
aulin
a pe
lagi
ca (C
leve
) H
ende
y (a
b)C
erat
aulu
s be
rgon
ii H
. Per
agal
lo,
Cer
atau
lina
berg
onii
(H.
Per
agal
lo) S
chüt
t
Eas
t/Wes
tH
orne
r et a
l. 20
05 (L
inkl
ette
r et a
l. 19
77, H
argr
aves
& M
aran
da 2
002,
B
érar
d-Th
erria
ult e
t al.
1999
, B
ates
& S
train
200
6)C
erat
aulin
a sp
p.C
erat
aulin
a sp
p.E
ast
Mar
tin (p
ers.
com
m.)2
Cha
etoc
eros
abn
orm
isC
haet
ocer
os a
bnor
mis
A.I.
P
rosh
kina
-Lav
renk
o (a
b)E
ast
Bru
nel 1
962
Cha
etoc
eros
affi
nis
Lau
der
Cha
etoc
eros
affi
nis
Lau
der (
ab)
Cha
etoc
eros
ang
ulat
us S
chüt
t, C
haet
ocer
os ra
lfsii
Cle
ve,
Cha
etoc
eros
java
nicu
s C
leve
, C
haet
ocer
os s
chut
tii C
leve
Eas
t/Wes
tS
ance
tta &
Cal
vert
1988
(Bér
ard-
Ther
riaul
t et a
l. 19
99, B
rune
l 196
2,
Gra
n &
Bra
arud
193
5)
Cha
etoc
eros
affi
nis
var.
circ
inal
is
(Meu
nier
) Hus
tedt
Cha
etoc
eros
affi
nis
var.
circ
inal
is
(Meu
n.) H
ust.
(ab)
Wes
t (U
S)
Cup
p 19
43
Cha
etoc
eros
affi
nis
var.
will
eiC
haet
ocer
os w
illei
Gru
now
(ab)
Eas
tB
érar
d-Th
erria
ult e
t al.
1999
(B
rune
l 196
2)C
haet
ocer
os a
nast
omos
ans
Cha
etoc
eros
ana
stom
osan
s G
runo
w
(ab)
Eas
t/Wes
tB
rune
l 196
2 (H
orne
r 200
2)
Cha
etco
eros
ang
ulat
usC
haet
ocer
os a
ffini
s La
uder
(ab)
Eas
tB
rune
l 196
2
Pag
e 20
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Cha
etoc
eros
atla
ntic
us C
leve
Cha
etoc
eros
atla
ntic
us C
leve
(ab)
Cha
etoc
eros
dis
par
Cas
traca
ne,
Cha
etoc
eros
pol
ygon
us S
chüt
t, C
haet
ocer
os a
udax
Sch
ütt
Eas
t/Wes
t/Ar
ctic
Cup
p 19
43 (B
érar
d-Th
erria
ult e
t al
. 199
9, B
rune
l 196
2, B
ursa
196
3,
Love
joy
et a
l. 20
02, G
ran
&
Bra
arud
193
5, B
ooth
& H
orne
r 19
97)
Cha
etoc
eros
atla
ntic
us f
orm
aud
ax
(Sch
ütt)
Gra
nC
haet
ocer
os a
tlant
icus
f. a
udax
(S
chüt
t) G
ran.
(ab)
Wes
t (U
S)
Cup
p 19
43
Cha
etoc
eros
atla
ntic
us v
ar.
skel
eton
(S
chüt
t) H
uste
dtC
haet
ocer
os a
tlant
icus
var
. sk
elet
on (
Sch
ütt)
Hus
t. (a
b)W
est (
US
)C
upp
1943
Cha
etoc
eros
bor
eale
Bai
l.C
haet
ocer
os b
orea
lis J
.W. B
aile
y (a
b)E
ast
Bai
ley
1913
(Bai
ley
1910
)
Cha
etoc
eros
bor
ealis
Cha
etoc
eros
bor
ealis
J.W
. Bai
ley
(ab)
Eas
t/Arc
ticB
érar
d-Th
erria
ult e
t al.
1999
(B
rune
l 196
2, B
ursa
196
3, G
ran
&
Bra
arud
193
5)C
haet
ocer
os b
revi
s S
chüt
tC
haet
ocer
os b
revi
s S
chüt
t (ab
)E
ast/W
est(U
S)/
Arc
ticC
upp
1943
(Bér
ard-
Ther
riaul
t et
al. 1
999,
Bru
nel 1
962,
Lov
ejoy
et
al. 2
002,
Gra
n &
Bra
arud
193
5)
Cha
etoc
eros
cap
reol
us
Cha
etoc
eros
cap
reol
us (
Ehr
enbe
rg)
Bai
ley
(ab)
Eas
tB
aile
y 19
10
Cha
etoc
eros
cer
atos
poru
s O
sten
feld
Cha
etoc
eros
cer
atos
poru
s O
sten
feld
(ab)
Cha
etoc
eros
gra
cile
Aps
tein
E
ast
Gra
n &
Bra
arud
193
5
Cha
etoc
eros
cer
atos
poru
s va
r. br
achy
setu
sC
haet
ocer
os c
erat
ospo
rus
var.
brac
hyse
tus
J.E
.B. R
ines
& P
.E.
Har
grav
es (a
b)
Eas
tB
érar
d-Th
erria
ult e
t al.
1999
Cha
etoc
eros
chr
ioph
yllu
m C
ast.
Cha
etoc
eros
chr
ioph
yllu
m C
ast.
(ab)
Eas
tB
aile
y 19
10
Cha
etoc
eros
cin
ctus
Gra
nC
haet
ocer
os c
inct
us G
ran.
(ab
)W
est
(US
)/Eas
tC
upp
1943
(Bru
nel 1
962,
Gra
n &
B
raar
ud 1
935)
Cha
etoc
eros
cle
vei
Cha
etoc
eros
cle
vei
F. S
chüt
t (ab
)E
ast
Bru
nel 1
962
Cha
etoc
eros
coa
rcta
tus
Laud
erC
haet
ocer
os c
oarc
tatu
s L
aude
r (ab
)C
haet
ocer
os ru
de C
leve
Wes
t (U
S)/E
ast
Cup
p 19
43 (B
rune
l 196
2)
Cha
etoc
eros
com
pres
sus
Cha
etoc
eros
com
pres
sus
Lau
der
(ab)
Cha
etoc
eros
con
tortu
s S
chüt
tE
ast/W
est/A
rct
icM
artin
(per
s. c
omm
.)2 (Bru
nel
1962
, Rie
del e
t al.
2003
, Gra
n &
B
raar
ud 1
935,
Bat
es &
Stra
in
2006
, Joh
nson
198
8)
Pag
e 21
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Cha
etoc
eros
con
cavi
corn
is M
angi
nC
haet
ocer
os c
onca
vico
rne
Man
gin
(ab)
Cha
etoc
eros
crio
philu
s
Eas
t/Wes
t/Ar
ctic
Har
grav
es &
Mar
anda
200
2 (S
ance
tta &
Cal
vert
1988
, Bér
ard-
Ther
riaul
t et a
l. 19
99, B
rune
l 196
2,
Bur
sa 1
963,
Lov
ejoy
et a
l. 20
02,
Gra
n &
Bra
arud
193
5, B
ooth
&
Hor
ner 1
997,
Roz
ansk
a et
al.
2008
)C
haet
ocer
os c
onca
vico
rnis
for
m
vola
ns (
Sch
ütt)
Hus
tedt
Cha
etoc
eros
con
cavi
corn
is f
orm
vo
lans
(S
chüt
t) H
ust.
(ab)
Wes
t (U
S)
Cup
p 19
43
Cha
etoc
eros
con
stric
tus
Cha
etoc
eros
con
stric
tus
Gra
n (a
b)E
ast
Bru
nel 1
962
(Gra
n &
Bra
arud
19
35)
Cha
etoc
eros
con
tortu
s S
hütt
Cha
etoc
eros
com
pres
sus
Laud
er
(ab)
Cha
etoc
eros
con
tortu
s S
hütt
Eas
t/Wes
tH
orne
r et a
l. 20
05 (B
érar
d-Th
erria
ult e
t al.
1999
, Bru
nel 1
962)
Cha
etoc
eros
con
volu
tus
Cha
etoc
eros
con
volu
tus
Cas
traca
ne
(ab)
Eas
t/Wes
t/Ar
ctic
Hor
ner e
t al.
2005
(Cha
ng e
t al.
2007
, Bér
ard-
Ther
riaul
t et a
l. 19
99, B
rune
l 196
2, B
ursa
196
3,
Love
joy
et a
l. 20
02, G
ran
&
Bra
arud
193
5, R
ozan
ska
et a
l. 20
08)
Cha
etoc
eros
con
volu
tus
f. cl
avip
esC
haet
ocer
os c
onvo
lutu
s f.
cla
vipe
s
E. P
aasc
he (a
b)E
ast
Bru
nel 1
962
Cha
etoc
eros
con
volu
tus
var
. tri
seto
saC
haet
ocer
os c
onvo
lutu
s f.
trise
tosa
J.
F. B
rune
l C
haet
ocer
os c
onvo
lutu
s v
ar.
trise
tosa
Eas
t/Arc
ticM
artin
(per
s. c
omm
.)2 (Roz
ansk
a et
al.
2008
)C
haet
ocer
os c
oron
atus
Cha
etoc
eros
cor
onat
us G
ran
(ab)
Eas
tB
rune
l 196
2
Cha
etoc
eros
cos
tatu
s P
avill
ard
Cha
etoc
eros
cos
tatu
s P
avill
ard
(ab)
Cha
etoc
eros
adh
aere
ns M
angi
n E
ast/W
est
(US
)C
upp
1943
(Bru
nel 1
962,
McQ
uoid
&
Hob
son
1997
)C
haet
ocer
os c
rinitu
s S
chüt
t in
Ost
enfe
ldC
haet
ocer
os c
rinitu
s S
chüt
t (ab
)E
ast
Bru
nel 1
962
Cha
etoc
eros
crio
philu
sC
haet
ocer
os c
rioph
ilus
Cas
traca
ne
(ab)
Eas
tB
rune
l 196
2
Cha
etoc
eros
cur
vise
tus
Cle
veC
haet
ocer
os c
urvi
setu
s C
leve
(ab)
Eas
t (B
oF)/W
est/
Arc
tic
Hor
ner (
pers
. com
m.)
3 (Bru
nel
1962
, Bur
sa 1
963)
Cha
etoc
eros
dad
ayi
Pav
illar
dC
haet
ocer
os d
aday
i P
avill
ard
(ab)
Eas
t/Wes
tH
orne
r (pe
rs. c
omm
.)3 (B
rune
l 19
62)
Pag
e 22
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Cha
etoc
eros
dan
icus
C
haet
ocer
os d
anic
us C
leve
(ab)
Eas
t/Wes
tB
érar
d-Th
erria
ult e
t al.
1999
(B
rune
l 196
2, G
ran
& B
raar
ud
1935
, Bat
es &
Stra
in 2
006,
Hor
ner
2002
)C
haet
ocer
os d
ebili
s C
leve
Cha
etoc
eros
deb
ilis
Cle
ve (a
b)C
haet
ocer
os v
erm
icul
atum
Sch
ütt
Eas
t/Wes
t/Ar
ctic
Har
grav
es &
Mar
anda
200
2 (S
ance
tta &
Cal
vert
1988
, Bér
ard-
Ther
riaul
t et a
l. 19
99, B
rune
l 196
2,
Bur
sa 1
963,
Lov
ejoy
et a
l. 20
02,
Gra
n &
Bra
arud
193
5, M
cQuo
id &
H
obso
n 19
97)
Cha
etoc
eros
dec
ipie
ns
Cha
etoc
eros
dec
ipie
ns C
leve
(ab)
Cha
etoc
eros
gru
now
ii S
chüt
t E
ast/W
est/A
rct
icB
aile
y 19
13 (B
aile
y 19
10,
Link
lette
r et a
l. 19
77, B
érar
d-Th
erria
ult e
t al.
1999
, Bru
nel 1
962,
Fr
itz 1
921,
Bur
sa 1
963,
Lov
ejoy
et
al. 2
002,
Gra
n &
Bra
arud
193
5,
Boo
th &
Hor
ner 1
997)
Cha
etoc
eros
dec
ipie
ns f
orm
si
ngul
aris
Gra
nC
haet
ocer
os d
ecip
iens
for
m
sing
ular
is G
ran
(ab)
Wes
t (U
S)
Cup
p 19
43
Cha
etoc
eros
del
icat
ulus
Cha
etoc
eros
del
icat
ulus
C.E
.H.
Ost
enfe
ld (a
b)E
ast
Bru
nel 1
962
Cha
etoc
eros
den
sus
Cha
etoc
eros
den
sus
Cle
ve
Cha
etoc
eros
bor
ealis
var
. de
nsum
Cle
ve
Eas
t/Wes
t/Ar
ctic
San
cetta
& C
alve
rt 19
88 (B
rune
l 19
62, B
ursa
196
3, G
ran
& B
raar
ud
1935
)C
haet
ocer
os d
iade
ma
(Ehr
enbe
rg)
Gra
nC
haet
ocer
os d
iade
ma
(Ehr
enbe
rg)
Gra
n C
haet
ocer
os p
arad
oxus
P
erag
allo
, Cha
etoc
eros
gr
oenl
andi
cus
Cle
ve,
Cha
etoc
eros
dis
tans
var
. su
bsec
undu
s G
runo
w e
x V
an
Heu
rck,
Syn
dend
rium
dia
dem
a E
hren
berg
Eas
t/Wes
t/Ar
ctic
San
cetta
& C
alve
rt 19
88 (B
érar
d-Th
erria
ult e
t al.
1999
, Bru
nel 1
962,
Fr
itz 1
921,
Lov
ejoy
et a
l. 20
02,
Gra
n &
Bra
arud
193
5, M
cQuo
id &
H
obso
n 19
97)
Cha
etoc
eros
dic
haet
a E
hren
berg
Cha
etoc
eros
dic
haet
us E
hren
berg
(a
b)C
haet
ocer
os ja
nisc
hian
us
Cas
traca
ne, C
haet
ocer
os
rem
otus
Cle
ve &
Gru
now
, C
haet
ocer
os d
ista
ns C
leve
Wes
t (U
S)
Cup
p 19
43 (B
rune
l 196
2)
Pag
e 23
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Cha
etoc
eros
dic
ladi
a C
haet
ocer
os d
icla
dia
F.S
. C
astra
cane
deg
li A
ngel
min
elli
(ab)
Eas
tB
aile
y 19
10
Cha
etoc
eros
did
ymus
Ehr
enbe
rgC
haet
ocer
os d
idym
us C
.G.
Ehr
enbe
rg
Cha
etoc
eros
did
ymus
f.
sing
ular
is H
. Tak
ano,
C
haet
ocer
os d
idym
us v
ar. a
nglic
a (G
runo
w) G
ran
Eas
t/Wes
t/Ar
ctic
San
cetta
& C
alve
rt 19
88 (B
érar
d-Th
erria
ult e
t al.
1999
, Bru
nel 1
962,
B
ursa
196
3,G
ran
& B
raar
ud 1
935,
B
ates
& S
train
200
6, M
cQuo
id &
H
obso
n 19
97)
Cha
etoc
eros
did
ymus
var
. ang
lica
(Gru
now
) Gra
nC
haet
ocer
os d
idym
us E
hren
berg
(a
b)C
haet
ocer
os d
idym
us v
ar.
angl
ica
(Gru
now
) Gra
nW
est (
US
)C
upp
1943
Cha
etoc
eros
did
ymus
var
. pr
otub
eran
s (L
aude
r) G
ran
and
Yen
do
Cha
etoc
eros
pr
otub
eran
s L
aude
r (a
b)C
haet
ocer
os d
idym
us v
ar.
prot
uber
ans
(Lau
der)
Gra
n an
d Y
endo
Wes
t (U
S)
Cup
p 19
43
Cha
etoc
eros
diff
icili
s C
leve
Cha
etoc
eros
diff
icili
s C
l. (a
b)W
est
(US
)/Eas
tC
upp
1943
(Bru
nel 1
962)
Cha
etoc
eros
dip
yren
ops
Cha
etoc
eros
dip
yren
ops
A.F
. M
euni
er (a
b)E
ast
Bru
nel 1
962
Cha
etoc
eros
div
ersu
s C
leve
Cha
etoc
eros
div
ersu
s C
leve
(ab)
Cha
etoc
eros
laev
e G
. Leu
duge
r-Fo
rtmor
elE
ast/W
est
(US
)C
upp
1943
(Car
ver &
Mal
let 2
002)
Ch.
eib
eni
Gru
now
Cha
etoc
eros
eib
enii
(Gru
now
) M
euni
er (a
b)C
haet
ocer
os p
arad
oxum
var
. ei
beni
i (G
runo
w) V
an H
eurc
k A
rctic
Bur
sa 1
963
Cha
etoc
eros
furc
ella
tus
Bai
ley
Cha
etoc
eros
furc
ella
tus
Bai
ley
(ab)
Eas
t/Arc
ticM
artin
(per
s. c
omm
.)2 (Bér
ard-
Ther
riaul
t et a
l. 19
99, R
iede
l et a
l. 20
03, B
ursa
196
3, G
ran
& B
raar
ud
1935
, Boo
th &
Hor
ner 1
997)
Cha
etoc
eros
gra
cilis
Sch
ütt
Cha
etoc
eros
neo
grac
ile
Van
Land
ingh
am (
ab)
Cha
etoc
eros
gra
cilis
Sch
ütt
Wes
t (U
S)
Cup
p 19
43 (B
rune
l 196
2, B
ursa
19
63)
Cha
etoc
eros
hol
satic
us S
chüt
tC
haet
ocer
os h
olsa
ticus
Sch
ütt (
ab)
Cha
etoc
eros
bal
ticus
Cle
veW
est
(US
)/Eas
tC
upp
1943
(Bér
ard-
Ther
riaul
t et
al. 1
999,
Bru
nel 1
962)
Cha
etoc
eros
ingo
lfian
usC
haet
ocer
os in
golfi
anus
Ost
enfe
ldE
ast/A
rctic
Bat
es &
Stra
in 2
006
(Roz
ansk
a et
al
. 200
8)C
haet
ocer
os k
aria
nus
Gru
now
Cha
etoc
eros
kar
ianu
s G
runo
w (a
b)E
ast/A
rctic
Mar
tin (p
ers.
com
m.)2 (B
érar
d-Th
erria
ult e
t al.
1999
, Bru
nel 1
962,
Lo
vejo
y et
al.
2002
)
Pag
e 24
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Cha
etoc
eros
laci
nios
us G
runo
wC
haet
ocer
os la
cini
osus
Sch
ütt (
ab)
Eas
t/Wes
t/Ar
ctic
Hor
ner e
t al.
2005
(Bér
ard-
Ther
riaul
t et a
l. 19
99, B
rune
l 196
2,
Bur
sa 1
963,
Lov
ejoy
et a
l. 20
02,
Gra
n &
Bra
arud
193
5)C
haet
ocer
os la
evis
Leu
duge
r-Fo
rtmor
elC
haet
ocer
os la
evis
Leu
d.-F
ort (
ab)
Eas
t/Wes
t (U
S)
Cup
p 19
43 (C
arve
r & M
alle
t 200
2)
Cha
etoc
eros
laud
eri
Ral
fsC
haet
ocer
os la
uder
i R
alfs
(ab)
Cha
etoc
eros
wei
ssflo
gii
Sch
utt
Wes
t (U
S)
Cup
p 19
43 (B
rune
l 196
2)
Cha
etoc
eros
lore
nzia
nus
Gru
now
Cha
etoc
eros
lore
nzia
nus
Gru
now
(a
b)C
haet
ocer
os c
ellu
losu
s La
unde
rE
ast/W
est/A
rct
icS
ance
tta &
Cal
vert
1988
(Bér
ard-
Ther
riaul
t et a
l. 19
99, B
ursa
196
3,
Bat
es &
Stra
in 2
006)
Cha
etoc
eros
lore
nzia
nus
f. fo
rcep
sC
haet
ocer
os lo
renz
ianu
s va
r. fo
rcep
s A
.F. M
euni
er (a
b)E
ast
Bru
nel 1
962
Cha
etoc
eros
mes
sane
nsis
C
astra
cane
C
haet
ocer
os m
essa
nens
e C
astra
cane
(ab
)C
haet
ocer
os fu
rca
Cle
veW
est
(US
)/Eas
tC
upp
1943
(Bru
nel 1
962,
Car
ver &
M
alle
t 200
2)C
haet
ocer
os m
itra
Cha
etoc
eros
mitr
a (J
.W. B
aile
y)
Cle
ve (a
b)D
icla
dia
mitr
a J.
W. B
aile
y,
Dic
ladi
a gr
oenl
andi
ca C
leve
E
ast/A
rctic
Bru
nel 1
962
(Bur
sa 1
963,
Lov
ejoy
et
al.
2002
)C
haet
ocer
os m
uelle
riC
haet
ocer
os m
uelle
ri E
.J.
Eas
tB
rune
l 196
2 (J
ohns
on 1
988)
Cha
etoc
eros
pau
lsen
iiC
haet
ocer
os p
auls
enii
Ost
enfe
ld
(ab)
Eas
tB
rune
l 196
2
Cha
etoc
eros
pav
illar
dii
Ikar
iC
haet
ocer
os p
avill
ardi
i J.
Ikar
i (ab
)W
est
San
cetta
& C
alve
rt 19
88
Cha
etoc
eros
pel
agic
us C
leve
Cha
etoc
eros
pel
agic
us C
l. (a
b)W
est (
US
)C
upp
1943
(Bru
nel 1
962)
Cha
etoc
eros
per
pusi
llus
Cle
veC
haet
ocer
os p
erpu
sillu
s C
l. (a
b)W
est(U
S)/E
ast
Cup
p 19
43 (B
rune
l 196
2)
Cha
etoc
eros
per
uvia
nus
Brig
htw
ell
Cha
etoc
eros
per
uvia
nus
Brig
htw
ell
(ab)
Cha
etoc
eros
chi
lens
is K
rass
ke,
Cha
etoc
eros
con
vexi
corn
is
Man
gin,
Cha
etoc
eros
pe
ruvi
anum
var
. cur
rens
P
erag
allo
, Cha
etoc
eros
per
uvio
-at
lant
icum
Kar
sten
Wes
t (U
S)/E
ast
Cup
p 19
43 (B
rune
l 196
2, C
arve
r &
Mal
let 2
002)
Cha
etoc
eros
per
uvia
nus
form
gr
acili
s (S
chrö
der)
Hus
tedt
Cha
etoc
eros
per
uvia
nus
var
. gr
acili
s J
.L.B
. Sch
röde
r (a
b)W
est (
US
)C
upp
1943
Cha
etoc
eros
pro
ceru
sC
haet
ocer
os p
roce
rus
F. S
chüt
t (ab
)E
ast
Bru
nel 1
962
Cha
etoc
eros
pse
udoc
rinitu
sC
haet
ocer
os p
seud
ocrin
itus
Ost
enfe
ld (a
b)E
ast
Bru
nel 1
962
(Gra
n &
Bra
arud
19
35)
Pag
e 25
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Cha
etoc
eros
pse
udoc
urvi
setu
s M
angi
nC
haet
ocer
os p
seud
ocur
vise
tus
Man
g. (a
b)E
ast/W
est
(US
)C
upp
1943
Cha
etoc
eros
radi
cans
Sch
ütt
Cha
etoc
eros
radi
cans
Sch
ütt (
ab)
Cha
etoc
eros
sco
lope
ndra
Cle
veE
ast/W
est/A
rct
icH
orne
r et a
l. 20
05 (S
ance
tta &
C
alve
rt 19
88, B
érar
d-Th
erria
ult e
t al
. 199
9, B
rune
l 196
2, L
ovej
oy e
t al
. 200
2, G
ran
& B
raar
ud 1
935,
M
cQuo
id &
Hob
son
1997
)C
haet
ocer
os ra
lfsii
Cha
etoc
eros
affi
nis
Laud
er (a
b)C
haet
ocer
os ra
lfsii
Cle
veE
ast
Bru
nel 1
962
Cha
etoc
eros
rigi
dus
Cha
etoc
eros
rigi
dus
Ost
enfe
ld (a
b)E
ast
Bru
nel 1
962
Cha
etoc
eros
rost
ratu
sC
haet
ocer
os ro
stra
tus
Laud
er (a
b)E
ast
Bru
nel 1
962
(Car
ver &
Mal
let
2002
)C
haet
ocer
os s
alta
nsC
haet
ocer
os s
alta
ns P
.T. C
leve
(ab)
Eas
tB
rune
l 196
2
Cha
etoc
eros
sch
üttii
Cha
etoc
eros
affi
nis
Lau
der (
ab)
Cha
etoc
eros
sch
uttii
Cle
veE
ast
Bru
nel 1
962
Cha
etoc
eros
sco
lope
ndra
Cha
etoc
eros
radi
cans
Sch
ütt (
ab)
Cha
etoc
eros
sco
lope
ndra
Cle
veE
ast
Bru
nel 1
962
Cha
etoc
eros
sei
raca
nthu
sC
haet
ocer
os s
eria
cant
hus
Gra
n (a
b)E
ast
Bru
nel 1
962
Ch.
sep
tent
riona
lis
Atth
eya
sept
entri
onal
is (
Øst
rup)
C
raw
ford
(ab)
Arc
ticB
ursa
196
3
Cha
etoc
eros
sim
ilis
Cle
veC
haet
ocer
os s
imili
s C
leve
(ab)
Eas
t/Wes
t/Ar
ctic
Hor
ner e
t al.
2005
(Bér
ard-
Ther
riaul
t et a
l. 19
99, G
ran
&
Bra
arud
193
5, R
ozan
ska
et a
l. 20
08)
Cha
etoc
eros
sim
plex
C
haet
ocer
os s
impl
ex O
sten
feld
(ab)
Eas
t/Arc
ticM
artin
(per
s. c
omm
.)2 (Raf
use
[per
s.co
mm
.]4 , Bér
ard-
Ther
riaul
t et
al. 1
999,
Bru
nel 1
962,
Gra
n &
B
raar
ud 1
935,
Roz
ansk
a et
al.
2008
)C
haet
ocer
os s
ocia
leC
haet
ocer
os s
ocia
le L
aude
r (ab
)E
ast
Fritz
192
1
Pag
e 26
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Cha
etoc
eros
soc
ialis
Lau
der
Cha
etoc
eros
soc
ialis
Lau
der (
ab)
Eas
t/Wes
t/Ar
ctic
Hor
ner e
t al.
2005
(Cha
ng e
t al.
2007
, San
cetta
& C
alve
rt 19
88,
Bér
ard-
Ther
riaul
t et a
l. 19
99,
Bru
nel 1
962,
Rie
del e
t al.
2003
, B
ursa
196
3, L
ovej
oy e
t al.
2002
, G
ran
& B
raar
ud 1
935,
Bat
es &
S
train
200
6, B
ooth
& H
orne
r 199
7)
Cha
etoc
eros
spp
.C
haet
ocer
os s
pp.
Eas
tB
ates
& S
train
200
6C
haet
ocer
os s
ubse
cund
usC
haet
ocer
os d
iade
ma
(Ehr
enbe
rg)
Gra
n (T
omas
199
7)E
ast
Bru
nel 1
962
(Bur
sa 1
963)
Cha
etoc
eros
sub
tilis
Cha
etoc
eros
sub
tilis
Cle
ve (a
b)E
ast/A
rctic
/B
érar
d-Th
erria
ult e
t al.
1999
C
haet
ocer
os te
nuis
sim
usC
haet
ocer
os te
nuis
sim
us A
.F.
Meu
nier
(ab)
Eas
t/Arc
ticB
érar
d-Th
erria
ult e
t al.
1999
(R
iede
l et a
l. 20
03)
Cha
etoc
eros
tere
sC
haet
ocer
os te
res
Cle
ve (a
b)E
ast/W
est/A
rct
icB
érar
d-Th
erria
ult e
t al.
1999
(B
rune
l 196
2, B
ursa
196
3, L
ovej
oy
et a
l. 20
02, G
ran
& B
raar
ud 1
935,
B
ates
& S
train
200
6, H
orne
r et a
l. 20
05)
Cha
etoc
eros
tetra
stic
hon
Cha
etoc
eros
tetra
stic
hon
Cle
veE
ast/W
est
(US
)B
rune
l 196
2 (C
upp
1943
)
Cha
etoc
eros
torti
liset
usC
haet
ocer
os to
rtilis
etus
L.A
. Man
gin
Eas
tB
rune
l 196
2
Cha
etoc
eros
torti
liset
usC
haet
ocer
os to
rtiss
imus
Gra
n (a
b)E
ast
Bru
nel 1
962
Cha
etoc
eros
torti
ssim
usC
haet
ocer
os to
rtiss
imus
Gra
n (a
b)E
ast/W
est
Car
ver &
Mal
let 2
002
(Cup
p 19
43)
Cha
etoc
eros
uru
guay
ensi
sC
haet
ocer
os u
rugu
ayen
sis
F.C
. M
ülle
r Mel
cher
s (a
b)E
ast
Bru
nel 1
962
Cha
etoc
eros
vis
tula
eC
haet
ocer
os v
istu
lae
C. A
pste
in
(ab)
Eas
tB
rune
l 196
2
Cha
etoc
eros
wig
ham
iiC
haet
ocer
os w
igha
mii
Brig
htw
ell
(ab)
Cha
etoc
eros
bot
tnic
us C
leve
, C
haet
ocer
os b
icon
cavu
s G
ran
Eas
t/Arc
ticB
érar
d-Th
erria
ult e
t al.
1999
(B
rune
l 196
2, L
ovej
oy 2
002,
R
ozan
ska
et a
l. 20
08)
Cha
etoc
eros
will
eiC
haet
ocer
os w
illei
Gru
now
(ab)
Cha
etoc
eros
affi
nis
var
. will
ei
(Gra
n) H
uste
dtE
ast
Har
grav
es &
Mar
anda
200
2 (M
artin
[per
s. c
omm
.]2 , Bér
ard-
Ther
riaul
t et a
l. 19
99, B
rune
l 196
2,
Link
lette
r et a
l. 19
77)
Pag
e 27
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Cha
etoc
eros
sp.
C
haet
ocer
os s
p.
Eas
t/Arc
ticM
artin
(per
s. c
omm
.)2 (Frit
z 19
21
& B
ursa
196
3)C
haet
ocer
os s
pp.
(pha
eoce
ros)
Cha
etoc
eros
spp
. (p
haeo
cero
s)E
ast
Mar
tin (p
ers.
com
m.)2
Cha
etoc
eros
sub
secu
ndus
(G
runo
w) H
uste
dtC
haet
ocer
os s
ubse
cund
us (G
runo
w
ex V
an H
eurc
k) H
uste
dt (
ab)
Cha
etoc
eros
dis
tans
var
. su
bsec
undu
s G
runo
w e
x V
an
Heu
rck
Wes
t (U
S)
Cup
p 19
43
Cha
etoc
eros
van
heur
ckii
Gra
nC
haet
ocer
os v
anhe
urck
ii G
ran
(ab)
Eas
t/Wes
t (U
S)
Hor
ner e
t al.
2005
(Cup
p 19
43,
San
cetta
& C
alve
rt 19
88, M
cQuo
id
& H
obso
n 19
97)
Cha
etoc
eros
vis
tula
e A
pste
inC
haet
ocer
os v
istu
lae
Aps
tein
(ab)
Wes
t (U
S)
Cup
p 19
43
Cha
etoc
eros
wig
ham
ii B
right
wel
lC
haet
ocer
os w
igha
mii
Brig
htw
ell
(ab)
Cha
etoc
eros
bot
tnic
us C
leve
, C
haet
ocer
os b
icon
cavu
s G
ran
Eas
t/Wes
t (U
S)
Cup
p 19
43 (B
aile
y 19
13)
Clim
acod
ium
frau
enfe
ldia
num
G
runo
wC
haet
ocer
os fr
auen
feld
ianu
m
Gru
now
(ab)
Eas
t/Wes
t (U
S)
Cup
p 19
43 (C
arve
r & M
alle
t 200
2)
Clim
acos
phen
ia e
long
ata
Clim
acos
phen
ia e
long
ata
Mer
esch
kow
sky
(ab)
Eas
tB
aile
y 19
13
Clim
acos
phen
ia m
onili
gera
E
hren
berg
Clim
acos
phen
ia m
onili
gera
Ehr
. (ab
)W
est (
US
)C
upp
1943
Coc
cone
is c
osta
ta G
rego
ryC
occo
neis
cos
tata
Gre
gory
(ab)
Wes
t (E
ffing
ham
In
let)
Hay
et a
l. 20
03
Coc
cone
is d
ecip
iens
Cle
veC
occo
neis
dec
ipie
ns P
.T. C
leve
(ab)
Wes
tM
cQuo
id &
Hob
son
2001
Coc
cone
is d
imin
uta
Pen
tosc
ek,
Kie
sela
lg, B
alat
Coc
cone
is n
eodi
min
uta
Kra
mm
er
(ab)
Coc
cone
is d
imin
uta
Pan
tocs
ekE
ast
John
son
1988
Coc
cone
is d
ista
ns G
rego
ryC
occo
neis
dis
tans
Gre
gory
(ab)
Coc
cone
is g
ranu
lifer
a G
revi
lle,
Coc
cone
is v
illos
a H
. Per
agal
loE
ast (
BoF
)R
awle
nce
1987
Coc
cone
is n
eodi
min
uta
var.
mar
ina
(Kra
mm
er) d
e S
tefa
no,
Mar
ino
& M
azze
lla
Coc
cone
is n
eodi
min
uta
var.
mar
ina
(Kra
mm
er)
de S
tefa
no, M
arin
o &
Maz
zella
(ab)
Wes
t (E
ffing
ham
In
let)
Hay
et a
l. 20
03
Coc
cone
is p
edic
ulus
C
occo
neis
ped
icul
us E
hren
berg
(ab)
Enc
yone
ma
caes
pito
sum
var
. pe
dicu
lus
(Ehr
enbe
rg) D
e To
ni,
Coc
cone
is c
omm
unis
f. p
edic
ulus
(E
hren
berg
) Chm
iele
vski
Eas
tB
aile
y 19
10
Pag
e 28
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Coc
cone
is p
lace
ntul
aC
occo
neis
pla
cent
ula
Ehr
enbe
rg
(ab)
Coc
cone
is p
edic
ulus
var
. pl
acen
tula
(C
.G. E
hren
berg
) A.
Gru
now
, Coc
cone
is c
omm
unis
f.
plac
entu
la (
Ehr
enbe
rg)
Chm
iele
vski
Eas
t/Wes
t (E
ffing
ham
In
let)
Hay
et a
l. 20
03 (B
aile
y 19
13,
Bai
ley
1910
, Zw
icke
r 199
8)
Coc
cone
is p
lace
ntul
a va
r. eu
glyp
pta
(Ehr
.) C
l.C
occo
neis
pla
cent
ula
var.
eugl
ypta
(E
hren
berg
) Cle
veC
occo
neis
eug
lypt
a E
hren
berg
Eas
tZw
icke
r 199
8
Coc
cone
is p
lace
ntul
a va
r. lin
eata
(E
hr.)
V.H
.C
occo
neis
pla
cent
ula
var.
linea
ta
(Ehr
enbe
rg) P
. Cle
ve (c
ol)
Coc
cone
is li
neat
a E
hren
berg
Eas
tZw
icke
r 199
8
Coc
cone
is p
lace
ntul
a E
hr. v
. pl
acen
tula
Coc
cone
is p
lace
ntul
a E
hr. v
. pl
acen
tula
(ab)
Coc
cone
is e
long
ata
Ehr
enbe
rg,
Coc
cone
is p
unct
ata
Ehr
enbe
rg,
Coc
cone
is p
umila
Küt
zing
Eas
tR
awle
nce
1987
Coc
cone
is s
cute
llum
Ehr
enbe
rgC
occo
neis
scu
tellu
m E
hren
berg
(ab)
Eas
t/Wes
t (E
ffing
ham
In
let)/
Arc
tic
Hay
et a
l. 20
03 (L
inkl
ette
r et a
l. 19
77, Z
wic
ker 1
998,
Joh
nson
19
88, B
aile
y 19
10, B
aile
y 19
13)
Coc
cone
is s
cute
llum
var
. par
va
(Gru
now
in V
an H
eurc
k) C
leve
Coc
cone
is s
cute
llum
var
. par
va
(Gru
now
in V
an H
eurc
k) C
leve
(ab)
Wes
t (E
ffing
ham
In
let)
Hay
et a
l. 20
03
Coc
cone
is s
pp.
Coc
cone
is s
pp.
Wes
tH
obso
n 20
09 (M
cQuo
id &
Hob
son
1997
, McQ
uoid
& H
obso
n 20
01)
Coc
cone
is s
taur
onei
form
is
(Rab
enho
rst)
Oku
noC
occo
neis
sta
uron
eifo
rmis
(R
aben
hors
t) O
kuno
(ab)
Coc
cone
is s
cute
llum
var
. st
auro
neifo
rmis
Van
Heu
rck
Wes
t (E
ffing
ham
In
let)
Hay
et a
l. 20
03
Coc
cone
ma
cist
ula
Coc
cone
ma
cist
ula
(Ehr
enbe
rg)
Ehr
enbe
rg (a
b)E
ast
Bai
ley
1913
Coc
cone
ma
lanc
eola
tum
Cym
bella
lanc
eola
ta (E
hren
berg
) K
irchn
er (a
b)C
occo
nem
a la
nceo
latu
m
Ehr
enbe
rgE
ast
Bai
ley
1910
Col
leto
nem
a su
bcoh
aere
ns
Col
leto
nem
a su
bcoh
aere
ns
Thw
aite
s (a
b)E
ast
Bai
ley
1913
Cor
ethr
on c
rioph
ilum
C
oret
hron
crio
philu
m C
astra
cane
(a
b)C
oret
hron
hys
trix
Hen
sen
Eas
t/Wes
tC
hang
et a
l. 20
07
Cor
ethr
on h
ystri
x H
ense
nC
oret
hron
crio
philu
m C
astra
cane
(a
b)E
ast
Gra
n &
Bra
arud
193
5
Pag
e 29
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Cos
cino
disc
us a
ster
omph
alus
C
osci
nodi
scus
ast
erom
phal
us
Ehr
enbe
rg (
ab)
Cos
cino
disc
us a
ster
omph
alus
va
r. co
nspi
cua
Gru
now
ex
Van
H
eurc
k, C
osci
nodi
scus
radi
atus
va
r. as
tero
mph
alus
(E
hren
berg
) E
hren
berg
, C
osci
nodi
scus
as
tero
mph
alus
var
. gen
uina
G
runo
w
Eas
tB
aile
y 19
13 (B
rune
l 196
2, L
ovej
oy
2002
)
Cos
cino
disc
us a
ster
omph
alus
var
. ce
ntra
lisC
osci
nodi
scus
cen
tralis
Ehr
enbe
rg
(ab)
Cos
cino
disc
us a
ster
omph
alus
va
r. ce
ntra
lis G
runo
wE
ast
Bru
nel 1
962
Cos
cino
disc
us c
entra
lis
(Ehr
enbe
rg)
Cos
cino
disc
us o
culu
s-iri
dis
var.
tenu
istri
ata
Gru
now
(ab)
Cos
cino
disc
us a
ster
omph
alus
va
r. ce
ntra
lis G
runo
wE
ast/W
est/A
rct
icH
orne
r et a
l. 20
05 (H
argr
aves
&
Mar
anda
200
2, B
érar
d-Th
erria
ult
et a
l. 19
99, B
rune
l 196
2, L
ovej
oy
et a
l. 20
02)
Cos
cino
disc
us c
entra
lis v
ar.
paci
fica
Gra
n &
Ang
stC
osci
nodi
scus
cen
tralis
var
. pa
cific
a G
ran
& A
ngst
(ab)
Eas
t/Wes
t (U
S)
Cup
p 19
43 (B
rune
l 196
2)
Cos
cino
disc
us c
onci
nnus
Cos
cino
disc
us c
onci
nnus
W. S
mith
(a
b)E
ast/W
est/A
rct
icH
argr
aves
& M
aran
da 2
002
(Bai
ley
1913
, Bér
ard-
Ther
riaul
t et
al. 1
999,
Bur
sa 1
963)
Cos
cino
disc
us c
urva
tulu
s G
runo
w
Cos
cino
disc
us c
urva
tulu
s G
runo
w
ex A
. Sch
mid
t (ab
)W
est (
US
)C
upp
1943
Cos
cino
disc
us e
ccen
tricu
s Th
alas
sios
ira e
ccen
trica
(E
hren
berg
) Cle
ve (a
b)C
osci
nodi
scus
hel
iozo
ides
S
idda
ll, C
osci
nodi
scus
ec
cent
ricus
Ehr
enbe
rg
Eas
tB
aile
y 19
13
Cos
cino
disc
us e
xcen
tricu
s E
hren
berg
Cos
cino
disc
us e
xcen
tricu
s E
hr. (
ab)
Eas
t/Wes
t (U
S)/A
rctic
Cup
p 19
43 (B
aile
y 19
13, B
rune
l 19
62, B
ursa
196
3, G
ran
& B
raar
ud
1935
)C
osci
nodi
scus
fasc
icul
atus
O'M
eC
osci
nodi
scus
nor
man
ii G
rego
ry e
x G
revi
lle (a
b)C
osci
nodi
scus
fasc
icul
atus
S
chm
idt
Eas
t (B
oF)
Link
lette
r et a
l. 19
77
Cos
cino
disc
us g
igas
Ehr
enbe
rgC
osci
nodi
scus
gig
as E
hren
berg
(ab)
Wes
tS
ance
tta &
Cal
vert
1988
(McQ
uoid
&
Hob
son
2001
)C
osci
nodi
scus
gra
nii
Cos
cino
disc
us g
rani
i G
ough
(ab)
Eas
t/Wes
t/Ar
ctic
Mar
tin (p
ers.
com
m.)2
(Rie
del e
t al.
2003
, Bur
sa 1
963,
McQ
uoid
&
Hob
son
2001
)C
osci
nodi
scus
gris
eus
Cos
cino
disc
us g
riseu
s R
.K. G
revi
lle
(ab)
Eas
tB
aile
y 19
10
C. l
inea
tus
Ehr
enbe
rgTh
alas
sios
ira le
ptop
us (
Gru
now
ex
Van
Heu
rck)
Has
le &
G. F
ryxe
ll (a
b)C
osci
nodi
scus
line
atus
E
hren
berg
, Cos
cino
disc
us
ehre
nber
gii
O'M
eara
Arc
ticB
ursa
196
3
Pag
e 30
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Cos
cino
disc
us m
argi
natu
s K
.B.
Cos
cino
disc
us m
argi
natu
s E
hren
berg
(ab
)C
osci
nodi
scus
lim
batu
s E
hren
berg
, Cos
cino
disc
us
fimbr
iatu
s-lim
batu
s E
hren
berg
Eas
t/Wes
t/Ar
ctic
Bai
ley
1913
(Bai
ley
1910
, Bru
nel
1962
, Bur
sa 1
963)
Cos
cino
disc
us m
inut
usC
osci
nodi
scus
min
utus
Küt
zing
(ab)
Eas
tB
aile
y 19
13
Cos
cino
disc
us n
eora
diat
usC
osci
nodi
scus
neo
radi
atus
A. C
leve
-E
uler
(ab)
Eas
tB
rune
l 196
2
Cos
cino
disc
us n
itidu
s G
rego
ryC
osci
nodi
scus
niti
dus
Gre
g. (a
b)E
ast/W
est
(US
)C
upp
1943
(Bai
ley
1913
)
Cos
cino
disc
us o
culu
s-iri
dis
Cos
cino
disc
us o
culu
s-iri
dis
(Ehr
enbe
rg) E
hren
berg
(ab
)C
osci
nodi
scus
radi
atus
var
. oc
ulus
-irid
is
Ehr
enbe
rg,
Cos
cino
disc
us o
culu
s-iri
dis
var.
genu
ina
Gru
now
, Cos
cino
disc
us
ocul
us-ir
idis
var
. typ
icus
Cle
ve-
Eul
er
Eas
t/Wes
tB
aile
y 19
10
Cos
cino
disc
us o
mph
alan
thus
Ehr
.C
osci
nodi
scus
om
phal
anth
us
Ehr
enbe
rg (a
b)E
ast
Bai
ley
1913
Cos
cino
disc
us p
ellu
cida
C
osci
nodi
scus
pel
luci
dus
A. G
runo
w
(ab)
Eas
tB
aile
y 19
10
Cos
cino
disc
us p
erfo
ratu
s E
hren
berg
Cos
cino
disc
us p
erfo
ratu
s E
hren
berg
(a
b)E
ast/W
est
Cup
p 19
43
Cos
cino
disc
us p
erfo
ratu
s va
r. ce
llulo
sa G
runo
wC
osci
nodi
scus
per
fora
tus
var.
cellu
losa
Gru
n. (
ab)
Wes
t (U
S)
Cup
p 19
43
Cos
cino
disc
us p
erfo
ratu
s va
r. pa
villa
rdi
(For
ti) H
uste
dtC
osci
nodi
scus
per
fora
tus
var.
pavi
llard
i (F
orti)
Hus
t. (a
b)W
est (
US
)C
upp
1943
Cos
cino
disc
us p
olyc
hord
usTh
alas
sios
ira a
ngus
te-li
neat
a (A
. S
chm
idt)
G. F
ryxe
ll &
Has
le (a
b)C
osci
nodi
scus
pol
ycho
rdus
Gra
nE
ast
Bru
nel 1
962
Cos
cino
disc
us ra
diat
us E
hren
berg
Azp
eitia
nod
ulife
ra(A
.W.F
. Sch
mid
t) G
.A. F
ryxe
ll &
P.A
. Sim
s (a
b)C
osci
nodi
scus
radi
atus
E
hren
berg
Eas
t/Wes
t/Ar
ctic
San
cetta
& C
alve
rt 19
88 (B
aile
y 19
10, B
aile
y 19
13, B
érar
d-Th
erria
ult e
t al.
1999
, Bru
nel 1
962,
Lo
vejo
y et
al.
2002
, McQ
uoid
&
Hob
son
2001
, Hay
et a
l. 20
03)
Cos
cino
disc
us s
p.C
osci
nodi
scus
sp.
Eas
tB
rune
l 196
2C
osci
nodi
scus
spp
.C
osci
nodi
scus
spp
.W
est
McQ
uoid
& H
obso
n 19
97
(McQ
uoid
& H
obso
n 20
01)
Cos
cino
disc
us s
ubbu
llien
sC
osci
nodi
scus
sub
bulli
ens
E.G
. Jø
rgen
sen
(ab)
Eas
tB
rune
l 196
2
Pag
e 31
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Cos
cino
disc
us s
ubgl
obos
usC
osci
nodi
scus
sub
glob
osus
P.T
. C
leve
& A
. Gru
now
(ab)
Eas
tB
rune
l 196
2
Cos
cino
disc
us s
ubtil
is E
hr.
Cos
cino
disc
us s
ubtil
is E
hren
berg
(a
b)E
ast
Bai
ley
1913
Cos
cino
disc
us w
aile
sii
Gra
n &
A
ngst
Cos
cino
disc
us w
aile
sii
Gra
n &
Ang
st
(ab)
Eas
t/Wes
tH
orne
r et a
l. 20
05 (H
argr
aves
&
Mar
anda
200
2)C
osci
nosi
ra a
tlant
ica
Cos
cino
sira
atla
ntic
a P
.J.L
. D
ange
ard
(ab)
Eas
tB
rune
l 196
2
Cos
cino
sira
med
iterr
anea
Cos
cino
sira
med
iterr
anea
J.L
.B.
Sch
röde
r (ab
)E
ast
Bru
nel 1
962
Cos
cino
sira
oes
trupi
Ost
enfe
ldTh
alas
sios
ira o
estru
pii
(Ost
enfe
ld)
Has
le (a
b)C
osci
nosi
ra o
estru
pii
Ost
enfe
ldE
ast
Gra
n &
Bra
arud
193
5
Cos
cino
sira
pol
ycho
rda
Thal
assi
osira
ang
uste
-line
ata
(A.
Sch
mid
t) G
. Fry
xell
& H
asle
(ab)
Cos
cino
sira
pol
ycho
rda
(Gra
n)
Gra
n, C
osci
nodi
scus
pol
ycho
rda
Gra
n
Eas
t/Wes
tB
rune
l 196
2 (B
ursa
196
3, G
ran
&
Bra
arud
193
5, H
orne
r 200
2)
Cte
noph
ora
pulc
hella
(R
alfs
ex
Kut
zing
) Will
iam
s &
Rou
ndC
teno
phor
a pu
lche
lla (
Küt
zing
) D.M
. W
illia
ms
& R
ound
Syn
edra
pul
chel
la R
alfs
ex
Küt
zing
Eas
tZw
icke
r 199
8
Cyc
lote
lla a
ntiq
ua W
.Sm
ithC
yclo
tella
ant
iqua
W.S
mith
(ab)
Arc
ticA
nton
iade
s et
al.
2005
Cyc
lote
lla c
hoct
awha
tche
eana
P
rasa
dC
yclo
tella
cho
ctaw
hatc
heea
na
Pra
sad
(ab)
Cyc
lote
lla c
aspi
a G
runo
w,
Cyc
lote
lla h
akan
sson
iae
Wen
dker
Eas
t/Wes
t (E
ffing
ham
In
let)
Hay
et a
l. 20
03 (B
érar
d-Th
erria
ult
et a
l. 19
99)
Cyc
lote
lla c
ompt
a G
run.
Cyc
lote
lla c
omta
(E
hren
berg
) K
ützi
ng (a
b)D
isco
plea
com
ta E
hren
berg
, P
unct
icul
ata
com
ta (
Ehr
enbe
rg)
H. H
akan
sson
Eas
tB
aile
y 19
10 (B
aile
y 19
13)
Cyc
lote
lla k
ützi
ngia
na T
hw.
Cyc
lote
lla k
ützi
ngia
na T
hw. (
ab)
Eas
tB
aile
y 19
10
Cyc
lote
lla li
tora
lis L
ange
&
Syv
erts
enC
yclo
tella
lito
ralis
Lan
ge &
S
yver
tsen
(ab)
Wes
tM
cQuo
id &
Hob
son
2001
Cyc
lote
lla m
eneg
hini
ana
Kg.
Cyc
lote
lla m
eneg
hini
ana
Küt
zing
(a
b)E
ast
Bai
ley
1913
(Joh
nson
198
8, B
aile
y 19
10)
Cyc
lote
lla o
perc
ulat
a (A
g.) K
tz.
Cyc
lote
lla o
perc
ulat
a (C
. Aga
rdh)
B
rébi
sson
(ab)
Eas
t (B
oF)
Raw
lenc
e 19
87
Cyc
lote
lla s
p.C
yclo
tella
sp.
Wes
t/Arc
ticS
ance
tta &
Cal
vert
1988
(Bur
sa
1963
)C
yclo
tella
spp
.C
yclo
tella
spp
.E
ast/W
est/A
rct
icM
artin
(per
s. c
omm
.)2(R
iede
l et a
l. 20
03, M
cQuo
id &
Hob
son
2001
, H
ay e
t al.
2003
)
Pag
e 32
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Cyc
lote
lla s
triat
a K
ützi
ng (G
runo
w)
Cyc
lote
lla s
triat
a K
ützi
ng (G
runo
w)
(ab)
Cos
cino
disc
us s
triat
us K
ützi
ng,
Cyc
lote
lla d
alla
sian
a W
.Sm
ith
Eas
t/Wes
t (E
ffing
ham
In
let)
Hay
et a
l. 20
03 (B
aile
y 19
13,
Bér
ard-
Ther
riaul
t et a
l. 19
99)
Cyl
indr
othe
ca c
lost
eriu
m
(Ehr
enbe
rg) L
ewin
& R
eim
ann
Cyl
indr
othe
ca c
lost
eriu
m
(Ehr
enbe
rg) R
eim
ann
& L
ewin
(ab
)N
itzsc
hiel
la te
nuiro
stris
M
eres
chko
wsk
y, C
erat
onei
s cl
oste
rium
Ehr
enbe
rg, N
itzsc
hia
clos
teru
im W
. Sm
ith
Eas
t/Wes
t/Ar
ctic
Hor
ner e
t al.
2005
(Bér
ard-
Ther
riaul
t et a
l. 19
99, T
hale
r 200
7,
Rie
del e
t al.
2003
, Lov
ejoy
et a
l. 20
02, B
ates
& S
train
200
6, B
ooth
&
Hor
ner 1
997,
Roz
ansk
a et
al.
2008
)C
ylin
drot
heca
gra
cilis
(B
réb.
) Gru
n.C
ylin
drot
heca
gra
cilis
(Bré
biss
on e
x K
ützi
ng) G
runo
w (
ab)
Cyl
indr
othe
ca g
erst
enbe
rger
i R
aben
hors
t, C
erat
onei
s gr
acili
s B
rébi
sson
ex
Kut
zing
, Nitz
schi
a ta
enia
W.S
mith
, Nitz
schi
ella
gr
acili
s (B
rébi
sson
ex
Küt
zing
) R
aben
hors
t
Eas
tR
awle
nce
1987
(Bér
ard-
Ther
riaul
t et
al.
1999
, Tha
ler 2
007,
Trit
es
2005
, Bat
es &
Stra
in 2
006)
Cym
atop
leur
a el
liptic
a S
.B.D
.C
ymat
ople
ura
ellip
tica
(Bré
biss
on)
W.S
mith
(ab)
Eas
tB
aile
y 19
13
Cym
atos
ira lo
renz
iana
C
ymat
osira
lore
nzia
na G
runo
w (a
b)E
ast
Car
ver &
Mal
let 2
002
Cym
atos
ira s
p.C
ymat
osira
sp.
Wes
tS
ance
tta &
Cal
vert
1988
Cym
bella
am
phic
epha
la N
aege
liC
ymbe
lla a
mph
icep
hala
Nae
geli
(ab)
Arc
ticA
nton
iade
s et
al.
2005
Cym
bella
ang
usta
ta (
W.S
mith
) C
leve
Cym
bella
ang
usta
ta (
W.S
mith
) C
leve
(ab)
Arc
ticA
nton
iade
s et
al.
2005
Cym
bella
arc
tica
(Lan
gers
tedt
) S
chm
idt
Cym
bella
arc
tica
(Lan
gers
tedt
) S
chm
idt (
ab)
Arc
ticA
nton
iade
s et
al.
2005
Cym
bella
bot
ellu
s (L
ange
rste
dt)
Sch
mid
tC
ymbe
lla b
otel
lus
(Lan
gers
tedt
) C
leve
(ab)
Arc
ticA
nton
iade
s et
al.
2005
Cym
bella
ces
atii
(Rab
enho
rst)
Gru
now
ex
Sch
mid
tE
ncyo
nops
is c
esat
ii (R
aben
hors
t) K
ram
mer
(ab)
Cym
bella
ces
atii
(Rab
enho
rst)
Gru
now
ex
Sch
mid
tA
rctic
Ant
onia
des
et a
l. 20
05
Cym
bella
cis
tula
(E
hren
berg
) K
irchn
erC
occo
nem
a ci
stul
a (E
hren
berg
) E
hren
berg
(ab)
Arc
tic/E
ast
Ant
onia
des
et a
l. 20
05 (B
aile
y 19
13)
Cym
bella
cus
pida
ta K
ützi
ngC
ymbe
lla c
uspi
data
Küt
zing
(ab)
Arc
tic/E
ast
Ant
onia
des
et a
l. 20
05 (B
aile
y 19
13)
Cym
bella
des
crip
ta (
Hus
tedt
) K
ram
mer
& L
ange
-Ber
talo
tC
ymbe
lla b
roen
lund
ensi
s F
oged
(a
b)C
ymbe
lla d
escr
ipta
(H
uste
dt)
Kra
mm
er &
Lan
ge-B
erta
lot
Arc
ticA
nton
iade
s et
al.
2005
Pag
e 33
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Cym
bella
des
igna
ta K
ram
mer
C
ymbe
lla d
esig
nata
Kra
mm
er (a
b)A
rctic
Ant
onia
des
et a
l. 20
05
Cym
bella
ehr
enbe
rgii
Küt
zing
Cym
bella
ehr
enbe
rgii
Küt
zing
(ab)
Eas
tB
aile
y 19
10
Cym
bella
gas
troid
esC
ymbe
lla a
sper
a (E
hren
berg
) Cle
veC
ymbe
lla g
astro
ides
(Küt
zing
) B
rébi
sson
& G
odey
Eas
tB
aile
y 19
10
Cym
bella
ince
rta (G
runo
w) C
leve
Cym
bella
ince
rta G
runo
w (a
b)A
rctic
Ant
onia
des
et a
l. 20
05
Cym
bella
lanc
eola
ta
Cym
bella
lanc
eola
ta (E
hren
berg
) K
irchn
er (
ab)
Coc
cone
ma
lanc
eola
tum
E
hren
berg
, Gom
phon
ema
lanc
eola
tum
Ehr
enbe
rg
Eas
tB
aile
y 19
10
Cym
bella
lapp
onic
a G
runo
w
Cym
bella
lapp
onic
a G
runo
w e
x C
leve
(ab)
Arc
ticA
nton
iade
s et
al.
2005
Cym
bella
late
ns K
rass
keE
ncyo
nem
a la
tens
(Kra
sske
) D.G
. M
ann
(ab)
Cym
bella
late
ns K
rass
keA
rctic
Ant
onia
des
et a
l. 20
05
Cym
bella
mic
roce
phal
a G
runo
w in
V
an H
eurk
Enc
yono
psis
mic
roce
phal
a (G
runo
w) K
ram
mer
(ab)
Cym
bella
ruttn
eri
Hus
tedt
, N
avic
ula
inco
mpt
a K
rass
keA
rctic
Ant
onia
des
et a
l. 20
05
Cym
bella
obs
cura
Kra
sske
Enc
yone
ma
obsc
urum
(K
rass
ke)
D.G
. Man
n (a
b)C
ymbe
lla o
bscu
ra K
rass
keA
rctic
Ant
onia
des
et a
l. 20
05
Cym
bella
sile
siac
a B
leis
ch in
R
aben
hors
tE
ncyo
nem
a si
lesi
acum
(B
leis
ch)
D.G
. Man
n (a
b)C
ymbe
lla s
ilesi
aca
Ble
isch
Arc
ticA
nton
iade
s et
al.
2005
Cym
bella
sta
uron
eifo
rmis
La
gers
tedt
Cym
bella
sta
uron
eifo
rmis
Lag
erst
. (a
b)A
rctic
Ant
onia
des
et a
l. 20
05
Cym
bella
spp
. Aga
rdh
Cym
bella
spp
. Aga
rdh
Wes
t (E
ffing
ham
In
let)
Hay
et a
l. 20
03 (M
cQuo
id &
H
obso
n 20
01)
Cym
bella
tum
ida
Bre
b.C
ymbe
lla tu
mid
a (B
rébi
sson
in
Küt
zing
) van
Heu
rck
(ab)
Coc
cone
ma
tum
idum
Bré
biss
onE
ast
Bai
ley
& M
acK
ay 1
921
Cys
topl
eura
turg
ida
(Ehr
.) K
untz
eE
pith
emia
turg
ida
(Ehr
enbe
rg)
Küt
zing
(ab)
Nav
icul
a tu
rgid
a E
hren
berg
Eas
tP
atric
k &
Rei
mer
197
5
Dac
tylio
sole
n an
tarc
ticus
Dac
tylio
sole
n an
tarc
ticus
C
astra
cane
(ab)
Dac
tylio
sole
n an
tarc
ticus
var
. ty
pica
, Dac
tylio
sole
n an
tarc
ticus
f.
typi
ca H
eide
n &
Kol
be
Wes
tH
obso
n 20
09
Dac
tylio
sole
n bl
avya
nus
(H.
Per
agal
lo) H
asle
Dac
tylio
sole
n bl
avya
nus
(H.
Per
agal
lo) H
asle
(ab)
Gui
nard
ia b
lavy
ana
H. P
erag
allo
Eas
t/Wes
tH
orne
r et a
l. 20
05
Pag
e 34
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Dac
tylio
sole
n fra
gilis
sim
us
(Ber
gon)
Has
leD
acty
lioso
len
fragi
lissi
mus
(Ber
gon)
H
asle
(ab)
Rhi
zoso
leni
a fra
gilis
sim
a B
ergo
nE
ast/W
est/A
rct
icH
orne
r et a
l. 20
05 (B
érar
d-Th
erria
ult e
t al.
1999
, Lov
ejoy
20
02, B
ates
& S
train
200
6)D
acty
lioso
len
med
iterr
anea
us
Per
agal
loLe
ptoc
ylin
drus
med
iterr
aneu
s (H
. P
erag
allo
) Has
le (a
b)D
acty
lioso
len
med
iterr
aneu
s (P
erag
allo
) Per
agal
loE
ast
Gra
n &
Bra
arud
193
5
Dac
tylio
sole
n ph
uket
ensi
sD
acty
lioso
len
phuk
eten
sis
(B.G
. S
unds
tröm
) G.R
. Has
le (a
b)R
hizo
sole
nia
phuk
eten
sis
B.G
. S
unds
tröm
Eas
t/Wes
tH
obso
n 20
09 (C
arve
r & M
alle
t 20
02)
Dac
tylio
sole
n sp
p.D
acty
lioso
len
spp.
Eas
tM
artin
(per
s. c
omm
.)2
Den
ticul
a ku
etzi
ngii
Gru
now
Den
ticul
a ku
etzi
ngii
Gru
now
(ab)
Nitz
schi
a de
ntic
ula
Gru
now
, D
entic
ula
dent
icul
a (G
runo
w)
Sch
onfe
ldt
Arc
ticA
nton
iade
s et
al.
2005
Dia
desm
is c
onte
nta
(Gru
now
) M
ann
Dia
desm
is c
onte
nta
(Gru
now
ex
Van
H
eurc
k) D
.G. M
ann
(ab)
Nav
icul
a co
nten
ta G
runo
w e
x V
an
Heu
rck,
Nav
icul
a tri
nodi
s f.
m
inut
a G
runo
w, S
chiz
onem
a co
nten
tum
(Gru
now
) Kun
tze
Arc
ticA
nton
iade
s et
al.
2005
Dia
desm
is p
arac
onte
nta
Lang
e-B
erta
lot &
Wer
umD
iade
smis
par
acon
tent
a La
nge-
Ber
talo
t & W
erum
(ab)
Arc
ticA
nton
iade
s et
al.
2005
Dia
tom
a el
onga
tum
Ag.
Dia
tom
a te
nuis
C. A
gard
h (a
b)D
iato
ma
elon
gatu
m A
g.E
ast
Raw
lenc
e 19
87D
iato
ma
hyem
ale
Lgng
.D
iato
ma
hyem
ale
(Rot
h) H
eibe
rg
(ab)
Eas
tB
aile
y 19
10
Dia
tom
a m
esod
on (
Ehr
enbe
rg)
Kut
zing
Dia
tom
a m
esod
on (E
hren
berg
) K
ützi
ngD
iato
ma
hyem
alis
f. m
esod
on
(Ehr
enbe
rg) G
runo
w, D
iato
ma
vulg
are
var
. mes
odon
(E
hren
berg
) Gru
now
, Odo
ntid
ium
m
esod
on (
Küt
zing
) Küt
zing
, D
iato
ma
hyem
ale
var.
mes
odon
(E
hren
berg
) Kirc
hner
, Dia
tom
a hy
emal
is v
ar. m
esod
on
(Ehr
enbe
rg) G
runo
w
Wes
tH
ay e
t al.
2003
Dia
tom
a m
onili
form
is K
ützi
ngD
iato
ma
mon
ilifo
rmis
Küt
zing
(ab)
Dia
tom
a te
nue
var.
mon
ilifo
rme
Küt
zing
Arc
ticA
nton
iade
s et
al.
2005
Dia
tom
a s
pp.
Dia
tom
a s
pp.
Wes
tM
cQuo
id &
Hob
son
2001
Dia
tom
a te
nuis
Aga
rdh
Dia
tom
a te
nuis
Aga
rdh
(ab)
Dia
tom
a te
nuis
var
. elo
ngat
a Ly
ngby
e, D
iato
ma
elon
gatu
m
(Lyn
gbye
) C. A
gard
h A
gard
h
Arc
ticA
nton
iade
s et
al.
2005
Dia
tom
a te
nue
Ag.
Dia
tom
a te
nue
Aga
rdh
(ab)
Eas
tB
aile
y 19
10D
iato
ma
tenu
e va
r. el
onga
tum
Ly
ngby
eD
iato
ma
tenu
e va
r. el
onga
tum
(L
yngb
.) (a
b)E
ast (
BoF
)P
oulin
et a
l. 19
84a
Pag
e 35
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Dia
tom
ella
min
uta
Hus
tedt
in A
. S
chm
idt,
Atla
sD
iato
mel
la m
inut
a H
uste
dt in
A.
Sch
mid
t, A
tlas
(ab)
Wes
t (E
ffing
ham
In
let)
Hay
et a
l. 20
03
Dic
ladi
a ca
preo
lus
Ehr
.D
icla
dia
capr
eolu
s E
hren
berg
(ab)
Eas
tB
aile
y 19
13
Dip
lone
is b
ombu
sD
iplo
neis
bom
bus
(Ehr
enbe
rg)
Cle
veP
innu
laria
bom
bus
Ehr
enbe
rg,
Nav
icul
a bo
mbu
s (E
hren
berg
) K
ützi
ng, N
avic
ula
abno
rmis
C
astra
cane
Eas
tB
érar
d-Th
erria
ult e
t al.
1999
Dip
lone
is d
idym
a (E
hren
berg
) C
leve
Dip
lone
is d
idym
a(E
hren
berg
) Cle
ve
(ab)
Pin
nula
ria d
idym
us E
hren
berg
, N
avic
ula
didy
mus
Ehr
enbe
rgE
ast
Link
lette
r et a
l. 19
77 (B
aile
y 19
13)
Dip
lone
is in
terr
upta
(K
ützi
ng) C
leve
Dip
lone
is in
terr
upta
(Küt
zing
) Cle
ve
(ab)
Nav
icul
a in
terr
upta
Küt
zing
, S
chiz
onem
a in
terr
uptu
m (C
leve
) K
untz
e
Eas
tTh
aler
200
7
Dip
lone
is li
tora
lis v
ar. a
rctic
aD
iplo
neis
lito
ralis
var
. arc
tica
P.T
. C
leve
(ab)
Arc
ticR
iede
l et a
l. 20
03
Dip
lone
is li
tora
lis v
ar. c
lath
rata
Dip
lone
is li
tora
lis v
ar. c
lath
rata
(Ø
stru
p) C
leve
(ab)
Arc
ticR
iede
l et a
l. 20
03
Dip
lone
is o
cula
ta (
Bré
biss
on)
Cle
veD
iplo
neis
ocu
lata
(Bré
biss
on) C
leve
(a
b)A
rctic
Ant
onia
des
et a
l. 20
05
Dip
lone
is p
arm
a C
leve
Dip
lone
is p
arm
a C
leve
Arc
ticA
nton
iade
s et
al.
2005
Dip
lone
is s
pp.
Dip
lone
is s
pp.
Wes
tM
cQuo
id &
Hob
son
2001
Dip
lone
is s
mith
ii (B
rébi
sson
) Cle
veD
iplo
neis
sm
ithii
(Bré
biss
on in
W.
Sm
ith) C
leve
(ab)
Nav
icul
a el
liptic
a W
.Sm
ith,
Nav
icul
a sm
ithii
Bré
biss
on in
W.
Sm
ith
Eas
t/Wes
t (E
ffing
ham
In
let)
Hay
et a
l. 20
03 (B
aile
y 19
13)
Dity
lum
brig
htw
ellii
(W
est)
Gru
now
Dity
lum
brig
htw
ellii
(T. W
est)
Gru
now
(ab
)D
itylu
m tr
igon
um L
.W. B
aile
y ex
L.
W. B
aile
y, T
ricer
atiu
m
brig
htw
ellii
T. W
est,
Dity
lum
in
aequ
ale
J.W
. Bai
ley
ex L
.W.
Bai
ley
Eas
t/Wes
t/Ar
ctic
Hor
ner e
t al.
2005
(Lin
klet
ter e
t al.
1977
, Cha
ng e
t al.
2007
, San
cetta
&
Cal
vert
1988
, Lov
ejoy
et a
l. 20
02, G
ran
& B
raar
ud 1
935,
M
cQuo
id &
Hob
son
1997
, 200
1,
Hay
et a
l. 20
03)
Dity
lum
sol
Dity
lum
sol
(G
runo
w) D
e To
ni (a
b)E
ast
Car
ver &
Mal
let 2
002
Del
phin
eis
surir
ella
(E
hr.)
G.W
. A
ndre
ws
Del
phin
eis
surir
ella
(E
hren
berg
) G
.W. A
ndre
ws
Rha
phon
eis
surir
ella
(Ehr
enbe
rg)
Gru
now
Eas
tTr
ites
2005
Pag
e 36
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Den
ticul
a ku
etzi
ngii
Gru
now
Den
ticul
a ku
etzi
ngii
Gru
now
Nitz
schi
a de
ntic
ula
Gru
now
, D
entic
ula
dent
icul
a (G
runo
w)
Sch
onfe
ldt
Arc
ticA
nton
iade
s et
al.
2005
Det
onul
a co
nfer
vace
aD
eton
ula
conf
erva
cea
(Cle
ve) G
ran
(ab)
Laud
eria
con
ferv
acea
Cle
ve,
Det
onul
a cy
stife
ra G
ran
Eas
t/Arc
ticB
érar
d-Th
erria
ult e
t al.
1999
(B
rune
l 196
2, L
ovej
oy e
t al.
2002
, G
ran
& B
raar
ud 1
935,
Bat
es &
S
train
200
6)D
eton
ula
cyst
ifera
Det
onul
a co
nfer
vace
a (C
leve
) Gra
n (a
b)D
eton
ula
cyst
ifera
Gra
nE
ast
Bru
nel 1
962
Det
onul
a pu
mila
(C
astra
cane
) S
chüt
tD
eton
ula
pum
ila (
Cas
traca
ne)
Sch
ütt
Sch
roed
erel
la d
elic
atul
a P
avill
ard,
Th
alas
sios
ira c
onde
nsat
a C
leve
, La
uder
ia p
umila
Cas
traca
ne
Wes
tS
ance
tta &
Cal
vert
1988
Dor
ypho
ra a
mph
icer
osR
haph
onei
s am
phic
eros
(E
hren
berg
) Ehr
enbe
rg (a
b)D
oryp
hora
am
phic
eros
(E
hren
berg
) Küt
zing
Eas
tB
aile
y 19
13
Dor
ypho
ra b
oeck
iiB
rébi
sson
ia b
oeck
ii (E
hren
berg
) E.
O'M
eara
(ab)
Dor
ypho
ra b
oeck
ii (E
hren
berg
) W
. Sm
ithE
ast
Bai
ley
1913
Enc
yone
ma
caes
pito
sum
Kut
z.E
ncyo
nem
a ca
espi
tosu
m K
utz.
(ab)
Coc
cone
ma
caes
pito
sum
(K
ützi
ng) G
.S. W
est,
Cym
bella
ca
espi
tosa
(Küt
zing
) Sch
utt,
Cym
bella
cae
spito
sum
(Küt
zing
) B
run,
Cym
bella
cae
spito
sa
(Küt
zing
) Bru
n, C
ymbe
lla
vent
ricos
a v
ar. c
aesp
itosa
(K
ützi
ng) D
ippe
l
Eas
tB
aile
y 19
10 (B
aile
y 19
13)
Enc
yone
ma
elgi
nens
e (K
ram
mer
) M
ann
Cym
bella
elg
inen
sis
Kra
mm
er (a
b)E
ncyo
nem
a el
gine
nse
(Kra
mm
er)
Man
nA
rctic
Ant
onia
des
et a
l. 20
05
Enc
yone
ma
foge
dii
Kra
mm
erE
ncyo
nem
a fo
gedi
i K
ram
mer
(ab
)A
rctic
Ant
onia
des
et a
l. 20
05
Enc
yone
ma
gaue
man
nii
(Mei
ster
) K
ram
mer
Enc
yone
ma
gaue
man
nii
(Mei
ster
) K
ram
mer
(ab
)A
rctic
Ant
onia
des
et a
l. 20
05
Enc
yone
ma
lang
e-be
rtalo
tii
Kra
mm
erE
ncyo
nem
a la
nge-
berta
lotii
K
ram
mer
(ab
)A
rctic
Ant
onia
des
et a
l. 20
05
Pag
e 37
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Enc
yone
ma
min
utum
(H
ilse)
Man
nE
ncyo
nem
a m
inut
um (H
ilse
in
Rab
enho
rst)
D.G
. Man
n (
ab)
Enc
yone
ma
vent
ricos
um v
ar.
min
utum
(H
ilse)
May
er, C
ymbe
lla
vent
ricos
a f.
min
uta
(Hils
e in
R
aben
hors
t) C
leve
-Eul
er,
Cym
bella
ven
trico
sa f.
min
uta
(Hils
e in
Rab
enho
rst)
May
er,
Cym
bella
ven
trico
sa f.
min
uta
(Hils
e in
van
Heu
rck)
M
eres
chko
wsk
y, E
ncyo
nem
a ve
ntric
osum
var
. min
uta
(Hils
e in
R
aben
hors
t) S
chm
idt,
Enc
yone
ma
vent
ricos
um f.
m
inut
a (H
ilse
in R
aben
hors
t) G
runo
w, C
ymbe
lla g
raci
lis v
ar.
min
uta
(Hils
e in
Rab
enho
rst)
Rab
enho
rst,
Cym
bella
min
uta
Hils
e in
Rab
enho
rst,
Cym
bella
va
r. m
inut
a (H
ilse
in R
aben
hors
t) C
leve
-Eul
er
Arc
ticA
nton
iade
s et
al.
2005
Enc
yone
ma
vent
ricos
um (
Aga
rdh)
G
runo
wE
ncyo
nem
a ve
ntric
osum
(Aga
rdh)
G
runo
w (a
b)C
occo
nem
a ve
ntric
osum
(K
ützi
ng) G
.S. W
est
Eas
t/Arc
ticA
nton
iade
s et
al.
2005
(Bai
ley
1910
)E
ndic
tya
ocea
nica
Ehr
enbe
rgE
ndic
tya
ocea
nica
Ehr
enbe
rg (a
b)E
ast
Gra
n &
Bra
arud
193
5
Ent
omon
eis
kjel
lman
iiE
ntom
onei
s kj
ellm
anii
(Cle
ve) P
oulin
&
Car
dina
l (ab
)A
mph
ipro
ra k
jellm
anii
Cle
ve,
Ent
omon
eis
kjel
lman
ii va
r. ty
pica
C
leve
-Eul
er
Eas
t/Arc
ticB
érar
d-Th
erria
ult e
t al.
1999
(R
iede
l et a
l. 20
03)
Ent
omon
eis
palu
dosa
var
. dup
lex
Ent
omon
eis
palu
dosa
var
. dup
lex
(Don
kin)
D.B
. Cza
rnec
ki &
D.C
. R
eink
e (a
b)
Eas
tB
érar
d-Th
erria
ult e
t al.
1999
Ent
omon
eis
palu
dosa
var
. hy
perb
orea
Ent
omon
eis
palu
dosa
var
. hy
perb
orea
(G
runo
w) P
oulin
&
Car
dina
l (ab
)
Am
phip
rora
hyp
erbo
rea
Gru
now
, A
mph
ipro
ra p
alud
osa
f. m
inut
a G
runo
w, A
mph
ipro
ra h
yper
bore
a va
r. ge
nuin
a C
leve
-Eul
er
Arc
ticR
iede
l et a
l. 20
03
Ent
omon
eis
sp.
Ent
omon
eis
sp.
Arc
ticR
iede
l et a
l. 20
03 (B
ooth
& H
orne
r 19
97)
Pag
e 38
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Eph
emer
a pl
anam
embr
anac
eaE
phem
era
plan
amem
bran
acea
(H
ende
y) P
addo
ck (a
b)N
avic
ula
plan
amem
bran
acea
H
ende
y E
ast/A
rctic
Bér
ard-
Ther
riaul
t et a
l. 19
99
(Rie
del e
t al.
2003
)
Epi
them
ia a
rgus
W. S
m.
Epi
them
ia a
rgus
(Ehr
enbe
rg)
Küt
zing
(ab)
Cys
topl
eura
arg
us (E
hren
berg
) K
untz
e, E
pith
emia
oce
llata
(E
hren
berg
) Küt
zing
, Eun
otia
ar
gus
Ehr
enbe
rg, E
unot
ia
ocel
lata
Ehr
enbe
rg, E
pith
emia
ar
gus
var
. oce
llata
(E
hren
berg
) F.
Fric
ke
Eas
tB
aile
y 19
13
Epi
them
ia g
ibba
Kut
z.E
pith
emia
gib
ba K
ützi
ng (a
b)E
ast
Bai
ley
1910
Epi
them
ia h
yndm
ani
Epi
them
ia h
yndm
anii
W.S
mith
(ab)
Eas
tB
aile
y 19
13
Epi
them
ia m
uscu
lus
Küt
zing
Rho
palo
dia
mus
culu
s (K
ützi
ng) O
.F.
Mül
ler (
ab)
Epi
them
ia m
uscu
lus
Küt
zing
Eas
tB
aile
y 19
10 (B
aile
y 19
13)
Epi
them
ia tu
rgid
a E
hr.
Epi
them
ia tu
rgid
a (E
hren
berg
) K
ützi
ng (a
b)N
avic
ula
turg
ida
Ehr
enbe
rg,
Cym
bella
turg
ida
(Ehr
enbe
rg)
Has
sall,
Cys
topl
eura
turg
ida
(Ehr
enbe
rg) K
untz
e
Eas
tB
aile
y 19
10 (B
aile
y 19
13)
Epi
them
ia z
ebra
Ehr
.E
pith
emia
adn
ata
(Küt
zing
) B
rébi
sson
(ab)
Epi
them
ia z
ebra
(E
hren
berg
) K
ützi
ngE
ast
Bai
ley
1910
(Bai
ley
1913
)
Euc
ampi
a co
rnut
aE
ucam
pia
corn
uta
(ab)
Eas
tC
arve
r & M
alle
t 200
2E
ucam
pia
groe
nlan
dica
Euc
ampi
a gr
oenl
andi
ca C
leve
(ab)
Arc
ticB
érar
d-Th
erria
ult e
t al.
1999
(B
rune
l 196
2, L
ovej
oy e
t al.
2002
)
Euc
ampi
a re
cta
Euc
ampi
a re
cta
H.H
. Gra
n &
T.
Bra
arud
Eas
tG
ran
& B
raar
ud 1
935
Euc
ampi
a zo
diac
us
Euc
ampi
a zo
diac
us E
hren
berg
(ab)
Euc
ampi
a br
itann
ica
W.S
mith
, E
ucam
pia
nodo
sa A
. Sch
mid
tE
ast/W
est/A
rct
icC
hang
et a
l. 20
07 (B
rune
l 196
2,
Bur
sa 1
963,
Gra
n &
Bra
arud
19
35)
Euc
ampi
a zo
odia
ca E
hren
berg
Euc
ampi
a zo
odia
cus
Ehr
enbe
rg (a
b)E
ucam
pia
zood
iaca
Ehr
enbe
rgW
est
San
cetta
& C
alve
rt 19
88
Eun
otia
arc
us
Eun
otia
arc
us E
hren
berg
(ab)
Him
antid
ium
arc
us (
Ehr
enbe
rg)
Ehr
enbe
rgE
ast
Bai
ley
1913
Eun
otia
cur
vata
(K
tz.)
Lage
rst
v.
curv
ata
Eun
otia
cur
vata
var
. cur
vata
(K
ützi
ng) L
ager
st (a
b)E
ast
Raw
lenc
e 19
87
Eun
otia
dio
don
Ehr
.E
unot
ia d
iodo
n E
hren
berg
(ab
)E
ast
Bai
ley
1913
(Bai
ley
1910
)E
unot
ia lu
naris
Eun
otia
bilu
naris
(E
hren
berg
) Mill
s (a
b)E
unot
ia lu
naris
(E
hren
berg
) G
runo
wE
ast
Bai
ley
1910
Pag
e 39
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Eun
otia
maj
or
Eun
otia
maj
or (
W.S
mith
) R
aben
hors
t (a
b)E
ast
Bai
ley
1910
(Bai
ley
1913
)
Eun
otia
mon
odon
Eun
otia
mon
odon
Ehr
enbe
rg (a
b)E
unot
ia m
ajor
(W. S
mith
) R
aben
hors
t E
ast
Bai
ley
1910
(Bai
ley
1913
)
Eun
otia
pec
tinal
is
Eun
otia
pec
tinal
is (K
ützi
ng)
Rab
enho
rst
(ab)
Eun
otia
pec
tinal
is v
ar. m
inor
(K
ützi
ng) R
aben
hors
t, H
iman
tidiu
m p
ectin
ale
Küt
zing
Eas
tB
aile
y 19
10
Eun
otia
pec
tinal
is v
ar. u
ndul
ata
Ral
fE
unot
ia p
ectin
alis
var
. und
ulat
a (R
alfs
) Rab
enho
rst (
ab)
Eas
tB
aile
y &
Mac
Kay
192
1
Eun
otia
pra
erup
ta E
hren
berg
Eun
otia
pra
erup
ta E
hren
berg
(ab
)A
rctic
Ant
onia
des
et a
l. 20
05
Eun
otia
spp
.E
unot
ia s
pp.
Wes
tH
obso
n 20
09 (H
ay e
t al.
2003
, M
cQuo
id &
Hob
son
2001
)E
unot
ia te
traod
on E
hr.
Eun
otia
tetra
odon
(Ehr
enbe
rg) R
alfs
(a
b)E
ast
Bai
ley
1913
Eup
odis
cus
radi
atus
Eup
odis
cus
radi
atus
J.W
. Bai
ley
(ab)
Aul
acod
iscu
s ra
diat
us (J
.W.
Bai
ley)
Brig
htw
ell
Eas
tC
arve
r & M
alle
t 200
2
Falla
cia
sp.
Falla
cia
sp.
Arc
ticLo
vejo
y et
al.
2002
Foss
ula
arct
ica
Has
le, S
yver
tsen
&
Qui
llfel
dtFo
ssul
a ar
ctic
a G
.R. H
asle
, E.E
. S
yver
tsen
& C
.H. v
on Q
uillf
eldt
(ab)
Arc
ticLo
vejo
y et
al.
2002
Frag
ilaria
cap
ucin
a D
esm
azie
res
Frag
ilaria
cap
ucin
a D
esm
aziè
res
(ab)
Frag
ilaria
cap
ucin
a f.
lanc
eola
ta
(Gru
now
in v
an H
eurc
k) H
uste
dt,
Frag
ilaria
cap
ucin
a v
ar.
lanc
eola
ta G
runo
w, F
ragi
laria
ca
puci
na f.
lanc
eola
ta (
Gru
now
) S
kabi
chev
skii
Eas
t/Arc
ticLi
nkle
tter e
t al.
1977
(Bai
ley
1913
, A
nton
iade
s et
al.
2005
)
Frag
ilaria
cap
ucin
a va
r. gr
acili
s (Ø
stru
p) H
uste
dtFr
agila
ria c
apuc
ina
var.
grac
ilis
(Øst
rup)
Hus
tedt
(ab)
Arc
ticA
nton
iade
s et
al.
2005
Frag
ilaria
con
stru
ens
E.
Sta
uros
ira c
onst
ruen
s E
hren
berg
(a
b)Fr
agila
ria c
onst
ruen
s E
hren
berg
Eas
tB
aile
y &
Mac
Kay
192
1
Frag
ilaria
oce
anic
aFr
agila
riops
is o
cean
ica
(Cle
ve)
Has
le (a
b)Fr
agila
ria o
cean
ica
Cle
veE
ast
Bru
nel 1
962
(Gra
n &
Bra
arud
19
35)
Frag
illar
ia h
arris
onii
Gru
n.Fr
agill
aria
har
rison
ii (W
.Sm
ith)
Gru
now
(ab)
Eas
tB
aile
y 19
13
Pag
e 40
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Frag
ilaria
cf.
sopo
tens
is W
itkow
ski
& L
ange
- Ber
talo
tFr
agila
ria c
f. so
pote
nsis
Witk
owsk
i &
Lan
ge- B
erta
lot (
ab)
Wes
t (E
ffing
ham
In
let)
Hay
et a
l. 20
03
Frag
ilaria
spp
.Fr
agila
ria s
pp.
Wes
tM
cQuo
id &
Hob
son
2001
Frag
ilaria
stri
atul
a Ly
ngby
eFr
agila
ria s
triat
ula
Lyng
bye
(ab)
Eas
t/Arc
ticP
oulin
et a
l. 19
84b
(Bér
ard-
Ther
riaul
t et a
l. 19
99, R
iede
l et a
l. 20
03)
Frag
ilaria
vau
cher
iae
(Küt
zing
) P
eter
sen
Frag
ilaria
vau
cher
iae
(Küt
zing
) P
eter
sen
(ab)
Frag
ilaria
cap
ucin
a v
ar.
vauc
heria
e (K
ützi
ng) L
ange
-B
erta
lot
Arc
ticA
nton
iade
s et
al.
2005
Frag
ilario
psis
atla
ntic
aFr
agila
riops
is a
tlant
ica
Paa
sche
(ab)
Nitz
schi
a pa
asch
ei H
asle
Eas
tB
érar
d-Th
erria
ult e
t al.
1999
Frag
ilario
psis
cyl
indr
usFr
agila
riops
is c
ylin
drus
(G
runo
w)
Krie
ger (
ab)
Frag
ilaria
cyl
indr
us G
runo
w,
Frag
ilaria
nan
a S
teem
ann
Nie
lsen
, Fra
gila
riops
is c
ylin
drus
va
r. pl
anct
onic
a W
illi K
riege
r,
Frag
ilario
psis
cyl
indr
us f
. min
or
Man
guin
, Fra
gila
riops
is li
near
is
var.
inte
rmed
ia M
angu
in,
Nitz
schi
a cy
lindr
us (
Gru
now
) H
asle
Eas
t/Arc
ticB
érar
d-Th
erria
ult e
t al.
1999
(R
iede
l et a
l. 20
03, L
ovej
oy e
t al.
2002
, Roz
ansk
a et
al.
2008
)
Frag
ilario
psis
dol
iolu
s (W
allic
h)
Med
lin &
Sim
sFr
agila
riops
is d
olio
lus
(Wal
lich)
M
edlin
& S
ims
(ab)
Syn
edra
dol
iolu
s W
allic
h, P
seud
o-eu
notia
dol
iolu
s (W
allic
h) G
runo
w E
ast/W
est
McQ
uoid
& H
obso
n 20
01 (C
arve
r &
Mal
let 2
002)
Frag
ilario
psis
oce
anic
aFr
agila
riops
is o
cean
ica
(Cle
ve)
Has
le (a
b)Fr
agila
ria o
cean
ica
Cle
ve,
Frag
ilaria
arc
tica
Gru
now
, N
itzsc
hia
grun
owii
Has
le,
Eas
t/Arc
ticB
érar
d-Th
erria
ult e
t al.
1999
(R
iede
l et a
l. 20
03, L
ovej
oy e
t al.
2002
)Fr
agila
riops
is p
seud
onan
aFr
agila
riops
is p
seud
onan
a (H
asle
) H
asle
(ab)
Frag
ilario
psis
nan
a (S
teem
ann
Nie
lsen
) Paa
sche
, Nitz
schi
a ps
eudo
nana
Has
le
Eas
tB
érar
d-Th
erria
ult e
t al.
1999
Frag
ilario
psis
spp
. Fr
agila
riops
is s
pp.
Wes
t/Arc
ticH
obso
n 20
09 (B
ooth
& H
orne
r 19
97, H
ay e
t al.
2003
)Fr
ustu
lia v
ulga
ris (
Thw
ait)
De
T. v
. vu
lgar
isFr
ustu
lia v
ulga
ris v
ar. v
ulga
ris
(Thw
aite
s) D
e To
ni (
ab)
E
ast (
BoF
)R
awle
nce
1987
Gom
phon
ema
acum
inat
um E
hr.
Gom
phon
ema
acum
inat
um E
hren
b.
(ab)
Eas
t Li
nkle
tter e
t al.
1977
(Bai
ley
1913
, B
aile
y 19
10)
Pag
e 41
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Gom
phon
ema
angu
stat
um
(Küt
zing
) Rab
enho
rst
Gom
phon
ema
angu
stat
um (K
ützi
ng)
Rab
enho
rst (
ab)
Sph
enel
la a
ngus
tata
Küt
zing
, G
omph
onem
a b
ohem
icum
H
uste
dt, S
phen
onei
s an
gust
ata
(Küt
zing
) Tre
visa
n, G
omph
onem
a in
stab
ilis
Hoh
n &
Hel
lerm
ann
Eas
t/Arc
ticA
nton
iade
s et
al.
2005
(Bai
ley
1910
)
Gom
phon
ema
capi
tatu
m
Gom
phon
ema
capi
tatu
m E
hren
berg
(a
b)G
omph
onem
a co
nstri
ctum
f.
clav
ata
(Ehr
enbe
rg) C
leve
-Eul
er,
Gom
phon
ema
trunc
atum
var
. ca
pita
ta (E
hren
berg
) Woo
dhea
d &
Tw
eed,
Gom
phon
ema
cons
trict
um v
ar. c
apita
tum
(E
hren
berg
) Gru
now
, G
omph
onem
a co
nstri
ctum
f.
capi
tata
(E
hren
berg
) Hus
tedt
Eas
tB
aile
y 19
13
Gom
phon
ema
cons
trict
um E
hr.
Gom
phon
ema
trunc
atum
Ehr
enbe
rg
(ab)
Gom
phon
ema
cons
trict
um E
hr.
Eas
tB
aile
y 19
13
Gom
phon
ema
cruc
iger
umG
omph
onem
a cr
ucig
erum
Eas
t (B
oF)
Link
lette
r et a
l. 19
77G
omph
onem
a ex
iguu
m K
.G
omph
onem
opsi
s ex
igua
(K
ützi
ng)
Med
linG
omph
onem
a ex
iguu
m K
ützi
ngE
ast
Bai
ley
1910
Gom
phon
ema
gem
inat
um A
.D
idym
osph
enia
gem
inat
a (L
yngb
ye)
M. S
chm
idt (
ab)
Gom
phon
ema
gem
inat
um
(Lyn
gbye
) C. A
gard
hE
ast
Bai
ley
1910
(Bai
ley
1913
)
Gom
phon
ema
mar
inum
W. S
m.
Gom
phon
ema
mar
inum
(K
ützi
ng)
Sm
ith (a
b)E
ast
Bai
ley
1913
(Bai
ley
1910
)
Gom
phon
ema
spp.
Aga
rdh
Gom
phon
ema
spp.
Aga
rdh
Wes
t (E
ffing
ham
In
let)
Hay
et a
l. 20
03 (M
cQuo
id &
H
obso
n 20
01)
Gom
phon
emop
sis
pseu
doex
igua
(S
imon
sen)
Med
linG
omph
onem
opsi
s ps
eudo
exig
ua
(Sim
onse
n) M
edlin
(ab)
Wes
t (E
ffing
ham
In
let)
Hay
et a
l. 20
03
Gra
mm
atop
hora
ang
ulos
a E
hren
berg
Gra
mm
atop
hora
ang
ulos
a E
hren
berg
(ab)
Eas
t/Wes
t(US
)C
upp
1943
Gra
mm
atop
hora
mar
ina
(Lyn
gbye
) K
ützi
ngG
ram
mat
opho
ra m
arin
a (L
yngb
ye)
Küt
zing
(ab)
Dia
tom
a m
arin
um L
yngb
yeE
ast/W
est(U
S)/
Arc
ticLi
nkle
tter e
t al.
1977
(Bai
ley
1913
, B
aile
y 19
10, C
upp
1943
, Bur
sa
1963
, Bat
es &
Stra
in 2
006,
M
cQuo
id &
Hob
son
2001
)
Gra
mm
atop
hora
mar
ina
var.
adria
tica
Gru
now
Gra
mm
atop
hora
mar
ina
var.
adria
tica
Gru
n. (a
b)W
est (
US
)C
upp
1943
Pag
e 42
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Gra
mm
atop
hora
oce
anic
a (E
hren
berg
, 185
4, in
par
t) G
runo
w
Gra
mm
atop
hora
oce
anic
a E
hren
berg
(ab
)G
ram
mat
opho
ra m
arin
a v
ar.
com
mun
is G
runo
w e
x V
an H
eurc
kW
est (
US
)C
upp
1943
Gra
mm
atop
hora
ser
pent
ina
Ehr
enbe
rgG
ram
mat
opho
ra s
erpe
ntin
a E
hren
berg
(ab)
Eas
tLi
nkle
tter e
t al.
1977
(Bai
ley
1913
, B
aile
y 19
10)
Gra
mm
atop
hora
sp.
Gra
mm
atop
hora
sp.
Arc
ticB
ursa
196
3G
ram
mat
opho
ra s
pp.
Gra
mm
atop
hora
spp
.E
ast/W
est
Hob
son
2009
(McQ
uoid
& H
obso
n 20
01)
Gui
nard
ia d
elic
atul
a (C
leve
) Has
leR
hizo
sole
nia
delic
atul
a C
leve
(ab)
Eas
t/Wes
tH
orne
r et a
l. 20
05 (B
érar
d-Th
erria
ult e
t al.
1999
, Bat
es &
S
train
200
6, H
orne
r 200
2)G
uina
rdia
flac
cida
(C
astra
cane
) H.
Per
agel
loG
uina
rdia
flac
cida
(Cas
traca
ne) H
. P
erag
ello
(ab)
Hen
seni
ella
bal
tica
Sch
utt,
Rhi
zoso
leni
a ca
stra
cane
i C
leve
, R
hizo
sole
nia
flacc
ida
Cas
traca
ne,
Gui
nard
ia b
altic
a S
chut
t
Eas
t/Wes
t(US
)C
upp
1943
(Lin
klet
ter 1
977,
Gra
n &
Bra
arud
193
5)
Gui
nard
ia s
triat
a G
uina
rdia
stri
ata
(Sto
lterfo
th) H
asle
Euc
ampi
a st
riata
Sto
lterfo
th,
Rhi
zoso
leni
a st
olte
rfoth
ii H
. P
erag
allo
Eas
t/Wes
tC
arve
r & M
alle
t 200
2 (H
orne
r 20
02)
Gyr
osig
ma
balti
cum
Gyr
osig
ma
balti
cum
(E
hren
berg
) R
aben
hors
tN
avic
ula
balti
ca E
hren
berg
, P
leur
osig
ma
balti
cum
(E
hren
berg
) W. S
mith
Eas
tB
rune
l 196
2 (T
rites
200
5, B
ates
&
Stra
in 2
006)
Gyr
osig
ma
diap
hanu
mG
yros
igm
a di
apha
num
Cle
ve (a
b)A
rctic
Rie
del e
t al.
2003
Gyr
osig
ma
dist
ortu
m (W
. Sm
.) W
. G
riffit
h &
Hen
frey
Gyr
osig
ma
dist
ortu
m (
W. S
mith
) G
riffit
h &
Hen
frey
(col
)E
ast
Thal
er 2
007
(Trit
es 2
005)
Gyr
osig
ma
fasc
iola
(E
hren
berg
) J.
W. G
riffit
h &
Hen
frey
Gyr
osig
ma
fasc
iola
(E
hren
berg
) J.
W. G
riffit
h &
Hen
frey
(col
)C
erat
onei
s fa
scio
la E
hren
berg
, P
leur
osig
ma
fasc
iola
(E
hren
berg
) W
. Sm
ith
Eas
tTh
aler
200
7
Gyr
osig
ma
hipp
ocam
pus
(Ehr
enbe
rg) H
assa
llG
yros
igm
a hi
ppoc
ampu
s (E
hren
berg
) Has
sall
(ab)
Nav
icul
a hi
ppoc
ampu
s E
hren
berg
, Ple
uros
igm
a hi
ppoc
ampu
s (E
hren
berg
) W
.Sm
ith
Eas
t P
atric
k &
Rei
mer
196
6
Gyr
osig
ma
litto
rale
G
yros
igm
a lit
tora
le (
W. S
mith
) G
riffit
h &
Hen
frey
(ab)
Ple
uros
igm
a lit
tora
le W
. Sm
ithE
ast
Raw
lenc
e 19
87
Gyr
osig
ma
obliq
uum
Gru
now
Gyr
osig
ma
obliq
uum
(Gru
now
) B
oyer
(col
)E
ast
Thal
er 2
007
Pag
e 43
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Gyr
osig
ma
obtu
satu
m (
Sul
livan
&
Wor
mle
y) B
oyer
G. o
btus
atum
(S
ulliv
an &
Wor
mle
y)
Boy
er (a
b)A
rctic
Ant
onia
des
et a
l. 20
05
Gyr
osig
ma
prol
onga
tum
var
. cl
oste
rioid
esG
yros
igm
a pr
olon
gatu
m v
ar.
clos
terio
ides
(G
runo
w) C
leve
Ple
uros
igm
a pr
olon
gatu
m v
ar.
clos
terio
ides
Gru
now
E
ast
Bru
nel 1
962
Gyr
osig
ma
spen
cerii
(Q
ueke
tt)
Cle
veG
yros
igm
a sp
ence
rii (
J.W
. Bai
ley
ex
Que
kett)
Grif
fith
& H
enfre
y (a
b)S
calp
trum
spe
ncer
i (Q
ueke
tt)
Kun
tze,
Ple
uros
igm
a sp
ence
ri va
r. ku
etzi
ngii
(Gru
now
) Gru
now
, P
leur
osig
ma
kuet
zing
ii G
runo
w,
Gyr
osig
ma
spen
ceri
(W. S
mith
) G
riffit
h &
Hen
frey,
Ple
uros
igm
a sp
ence
rii (B
aile
y ex
Que
kett)
W.
Sm
ith, N
avic
ula
spen
cerii
J.W
. B
aile
y ex
Que
kett,
Ple
uors
igm
a sp
ence
rii W
. Sm
ith, G
yros
igm
a ku
etzi
ngii
(Gru
nov)
Cle
ve
Eas
t/Wes
t (U
S)/A
rctic
Cup
p 19
43 (B
ursa
196
3)
Gyr
osig
ma
tenu
issi
mum
W. S
m.
Gyr
osig
ma
tenu
issi
mum
(W. S
mith
) G
riffit
h &
Hen
frey
(ab)
Ple
uros
igm
a te
nuis
sim
um
W.S
mith
Eas
tR
awle
nce
1987
(Bat
es &
Stra
in
2006
)G
yros
igm
a w
ansb
ecki
iG
yros
igm
a w
ansb
ecki
i (D
onki
n)
Cle
veP
leur
osig
ma
wan
sbec
kii
Don
kin,
P
leur
osig
ma
balti
cum
var
. w
ansb
ecki
i (D
onki
n) V
an H
eurc
k
Eas
tB
rune
l 196
2 (T
rites
200
5)
Han
naea
arc
us (
Ehr
enbe
rg)
Pat
rick
in P
atric
k &
Rei
mer
Han
naea
arc
us (
Ehr
enbe
rg) R
.M.
Pat
rick
(ab)
Wes
t (E
ffing
ham
In
let)/
Arc
tic
Hay
et a
l. 20
03 (A
nton
iade
s et
al.
2005
)
Han
tzsc
hia
wey
prec
htii
Han
tzsc
hia
wey
prec
htii
Gru
now
(ab)
Arc
ticR
iede
l et a
l. 20
03
Has
lea
cruc
iger
oide
sH
asle
a cr
ucig
eroi
des
(Hus
tedt
) S
imon
sen
(ab)
Arc
ticR
iede
l et a
l. 20
03
Has
lea
kjel
lman
iiH
asle
a kj
ellm
anii
(Cle
ve) S
imon
sen
(ab)
Arc
ticR
iede
l et a
l. 20
03
Has
lea
sp.
Has
lea
sp.
Arc
ticB
ooth
& H
orne
r 199
7H
asle
a vi
trea
Has
lea
vitre
a (C
leve
) Sim
onse
n (a
b)P
leur
osig
ma
vitre
um C
leve
A
rctic
Rie
del e
t al.
2003
Hem
iaul
us m
embr
anac
eus
Cle
veH
emia
ulus
mem
bran
aceu
s C
l. (a
b)E
ast/W
est
(US
)C
upp
1943
(Car
ver &
Mal
let 2
002)
Hem
iaul
us s
inen
sis
Gre
ville
Hem
iaul
us s
inen
sis
Gre
v. (
ab)
Eas
t/Wes
t (U
S)
Cup
p 19
43 (C
arve
r & M
alle
t 200
2)
Pag
e 44
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Hem
idis
cus
cune
iform
is v
ar.
vent
ricos
a (C
astra
cane
) Hus
tedt
Hem
idis
cus
cune
iform
is v
ar.
vent
ricos
a (C
astr.
) Hus
t. (a
b)W
est (
US
)C
upp
1943
Him
antid
ium
arc
us W
. Sm
.E
unot
ia a
rcus
Ehr
enbe
rg (a
b)H
iman
tidiu
m a
rcus
(Ehr
enbe
rg)
Ehr
enbe
rg
Eas
tB
aile
y 19
13
Him
antid
ium
gra
cile
Ehr
.H
iman
tidiu
m g
raci
le E
hren
berg
(ab
)E
ast
Bai
ley
1913
Him
antid
ium
maj
us W
. Sm
.H
iman
tidiu
m m
ajus
W. S
mith
(ab)
Eas
tB
aile
y 19
13
Hya
lodi
scus
sp.
Hya
lodi
scus
sp.
(ab)
Wes
tS
ance
tta &
Cal
vert
1988
Hya
lodi
scus
sub
tilis
Bai
l.H
yalo
disc
us s
ubtil
is J
.W.B
aile
y (a
b)E
ast
Bai
ley
1913
Isth
mia
ene
rvis
Ehr
.Is
thm
ia e
nerv
is E
hren
berg
(ab)
Isth
mia
obl
iqua
ta (J
.E.S
mith
) C
.Aga
rdh,
Isth
mie
lla e
nerv
is
Cle
ve
Eas
tB
aile
y 19
10
Isth
mia
ner
vosa
Küt
zing
Isth
mia
ner
vosa
Küt
z. (
ab)
Eas
t/Wes
t (U
S)/A
rctic
Bai
ley
1910
(Cup
p 19
43, B
ursa
19
63)
Laud
eria
con
ferv
acea
Det
onul
a co
nfer
vace
a (C
leve
) Gra
n (a
b)La
uder
ia c
onfe
rvac
ea C
leve
Eas
tB
rune
l 196
2
Laud
eria
ann
ulat
aLa
uder
ia a
nnul
ata
Cle
ve (a
b)La
uder
ia b
orea
lis G
ran
Eas
tR
awle
nce
1987
Laud
eria
gla
cial
isP
oros
ira g
laci
alis
(G
runo
w)
Jorg
ense
n (a
b)La
uder
ia g
laci
alis
(Gru
now
) Gra
nE
ast
Bru
nel 1
962
Lenn
oxia
fave
olat
aLe
nnox
ia fa
veol
ata
H.A
. Tho
mse
n &
K
.R. B
uck
(ab)
Eas
tB
érar
d-Th
erria
ult e
t al.
1999
Lept
ocyl
indr
us d
anic
us C
leve
Lept
ocyl
indr
us d
anic
us C
leve
(ab
)E
ast/W
est/A
rct
icH
orne
r et a
l. 20
05 (B
rune
l 196
2,
Bur
sa 1
963,
Lov
ejoy
200
2, G
ran
& B
raar
ud 1
935,
Bat
es &
Stra
in
2006
)Le
ptoc
ylin
drus
med
iterr
aneu
sLe
ptoc
ylin
drus
med
iterr
aneu
s (H
. P
erag
allo
) Has
le (a
b)La
uder
ia m
edite
rran
ea H
. P
erag
allo
, Dac
tylio
sole
n m
edite
rran
eus
(Per
agal
lo)
Per
agal
lo, D
acty
lioso
len
med
iterr
aneu
s va
r. te
nuis
Cle
ve,
Dac
tylio
sole
n te
nuis
(Cle
ve)
Gra
n, D
acty
lioso
len
mel
eagr
is
Kar
sten
Eas
tB
érar
d-Th
erria
ult e
t al.
1999
(B
ates
& S
train
200
6)
Pag
e 45
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Lept
ocyl
indr
us m
inim
us G
ran
Lept
ocyl
indr
us m
inim
us G
ran
(ab)
Lept
ocyl
indr
us b
elgi
cus
Meu
nier
Eas
t/Wes
tH
orne
r et a
l. 20
05 (L
inkl
ette
r et a
l. 19
77, C
hang
et a
l. 20
07, B
érar
d-Th
erria
ult e
t al.
1999
, Gra
n &
B
raar
ud 1
935,
Bat
es &
Stra
in
2006
)Li
cmop
hora
abb
revi
ata
Aga
rdh
Licm
opho
ra a
bbre
viat
a C
. Aga
rdh
(ab)
Eas
t/Arc
ticB
ursa
196
3 (B
ates
& S
train
200
6)
Licm
opho
ra fl
abel
lata
W.S
.Li
cmop
hora
flab
ella
ta C
. Aga
rdh
(ab)
Exi
laria
flab
ella
ta G
revi
lleE
ast
Bai
ley
1913
Licm
opho
ra h
yalin
a (K
ützi
ng)
Gru
now
Lico
mor
pha
hyal
ina
(Küt
zing
) G
runo
w (a
b)E
ast
Pou
lin e
t al.
1984
a
Licm
opho
ra ly
ngby
ei (
Küt
zing
) G
runo
w e
x V
an H
eurc
kLi
cmop
hora
lyng
byei
(Küt
zing
) G
runo
w e
x V
an H
eurc
k (a
b)Li
com
orph
a ab
brev
iata
C.A
gard
h,
Pod
osph
enia
lyng
byei
Kut
zing
Eas
t/Wes
t (U
S)
Cup
p 19
43 (B
aile
y 19
10)
Licm
opho
ra s
pp.
Licm
opho
ra s
pp.
Wes
tH
obso
n 20
09 (M
cQuo
id &
Hob
son
2001
)Li
cmop
hora
tinc
ta
Licm
opho
ra ti
ncta
E
ast
Bai
ley
1913
Lyre
lla c
lava
taLy
rella
cla
vata
(Gre
gory
) Man
n (a
b)N
avic
ula
clav
ata
Gre
gory
E
ast
Bér
ard-
Ther
riaul
t et a
l. 19
99
Lyre
lla s
p.Ly
rella
sp.
Wes
tH
obso
n 20
09 (M
cQuo
id &
Hob
son
2001
)M
angu
inea
sp.
Pad
dock
M
angu
inea
sp.
Pad
dock
A
rctic
Love
joy
et a
l. 20
02M
asto
gloi
a el
liptic
a M
asto
gloi
a el
liptic
a (C
. Aga
rdh)
C
leve
(ab
)Fr
ustu
lia e
llipt
ica
C.A
gard
h,
Am
phor
a el
liptic
a (A
gard
h)
Küt
zing
, Mas
togl
oia
dans
eii
var.
ellip
tica
Van
Heu
rck
Eas
tB
aile
y 19
10
Mas
togl
oia
exig
ua L
ewis
Mas
togl
oia
exig
ua F
.W. L
ewis
(ab)
Eas
tB
aile
y 19
13
Mas
togl
oia
lanc
eola
ta S
.B.D
.M
asto
gloi
a la
nceo
lata
Thw
aite
s (a
b)E
ast
Bai
ley
1913
Mas
togl
oia
smith
ii Th
w.
Mas
togl
oia
smith
ii Th
wai
tes
ex W
. S
mith
(ab)
Eas
tB
aile
y 19
10
Med
iopy
xis
hely
sia
M
edio
pyxi
s he
lysi
a (a
b)E
ast
Mar
tin &
LeG
resl
ey 2
008
Pag
e 46
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Mel
osira
arc
tica
Mel
osira
arc
tica
Dic
kie
(ab)
Mel
osira
hyp
erbo
rea
Sch
ütt,
Gai
llone
lla a
rctic
a (D
icki
e)
Ehr
enbe
rg
Eas
t/Arc
ticB
érar
d-Th
erria
ult e
t al.
1999
(B
rune
l 196
2, Z
wic
ker 1
998,
R
iede
l et a
l. 20
03, B
ursa
196
3,
Love
joy
et a
l. 20
02, B
ooth
&
Hor
ner 1
997,
Roz
ansk
a et
al.
2008
)M
elos
ira b
orre
riM
elos
ira b
orre
ri G
revi
lleE
ast
Fritz
192
1 (B
rune
l 196
2, B
aile
y 19
10, B
aile
y 19
13)
Mel
osira
cre
nula
ta
Mel
osira
cre
nula
ta (E
hren
berg
) K
ützi
ng (
ab)
Eas
tLi
nkle
tter e
t al.
1977
(Bai
ley
1910
, Fr
itz 1
921)
Mel
osira
dis
tans
(E
hr.)
Ktz
.A
ulac
oser
ia d
ista
ns (E
hren
b.)
Sim
onse
n (a
b)M
elos
ira d
ista
ns (E
hren
berg
) K
ützi
ng
Eas
tR
awle
nce
1987
Mel
osira
gla
cial
isM
elos
ira g
laci
alis
(A
. Gru
now
) A.
Cle
ve-E
uler
(ab)
Eas
tB
rune
l 196
2
Mel
osira
gra
nula
ta (
Ehr
.) R
alfs
Aul
acos
eira
gra
nula
ta (E
hren
berg
) S
imon
sen
(ab)
Mel
osira
gra
nula
ta (E
hren
berg
) R
alfs
E
ast
Raw
lenc
e 19
87 (B
aile
y 19
10)
Mel
osira
gra
nula
ta v
. ang
ustis
sim
a O
. Mül
ler
Aul
acos
eira
gra
nula
ta v
ar.
angu
stis
sim
a (
O.F
. Mül
ler)
S
imon
sen
(ab)
Mel
osira
gra
nula
ta v
ar.
angu
stis
sim
a O
.F. M
ülle
rE
ast
Raw
lenc
e 19
87
Mel
osira
hyp
erbo
rea
Gru
now
Mel
osira
hyp
erbo
rea
Gru
now
(ab)
Eas
t (B
oF)
Link
lette
r et a
l. 19
77 (F
ritz
1921
)
Mel
osira
ital
ica
(Ehr
.) K
tz.
Aul
acos
eira
ital
ica
(Ehr
enbe
rg) S
imon
sen
(ab)
Mel
osira
ital
ica
(Ehr
.) K
tz.
Eas
t (B
oF)
Raw
lenc
e 19
87
Mel
osira
juer
gens
ii A
g.M
elos
ira li
neat
a (D
illw
yn) A
gard
h (a
b)M
elos
ira ju
erge
nsii
Ag.
Eas
tB
aile
y 19
10 (Z
wic
ker 1
998)
Mel
osira
line
ata
(Dill
w.)
C.A
. Ag.
Mel
osira
line
ata
(Dill
wyn
) Aga
rdh
(ab)
Mel
osira
juer
gens
ii C
. Aga
rdh,
G
allio
nella
line
ata
(Dill
wyn
) Bor
y in
Ehr
enbe
rg, C
onfe
rva
linea
ta
Dill
wyn
, Lys
igon
ium
line
atum
(D
illw
yn) T
revi
san
Eas
t (B
oF)
Raw
lenc
e 19
87 (B
rune
l 196
2)
Mel
osira
mar
ina
Mel
osira
mar
ina
(W.S
mith
) Jan
isch
(a
b)E
ast (
BoF
)Li
nkle
tter e
t al.
1977
Pag
e 47
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Mel
osira
mon
ilifo
rmis
(M
ülle
r)
Aga
rdh
Mel
osira
mon
ilifo
rmis
(O.F
. Mül
ler)
C
. Aga
rdh
(ab)
Lysi
goni
um m
onili
form
e (O
.F.
Mül
ler)
Tre
visa
n, L
ysig
oniu
m
mon
ilifo
rme
(O.F
. Mül
ler)
Lin
k,
Con
ferv
a m
onili
form
is O
.F. M
ülle
r, M
elos
ira b
orre
ri va
r. m
onili
form
is
(O.F
. Mül
ler)
A. G
runo
w
Eas
t/Wes
tLi
nkle
tter e
t al.
1977
(McQ
uoid
&
Hob
son
2001
)
Mel
osira
mon
ilifo
rmis
var
. su
bglo
bosa
Mel
osira
mon
ilifo
rmis
var
. su
bglo
bosa
(G
runo
w) H
uste
dtM
elos
ira b
orre
ri va
r. su
bglo
bosa
A
. Gru
now
E
ast
Bru
nel 1
962
Mel
osira
num
mul
oide
s A
g.M
elos
ira n
umm
uloi
des
C.A
. Aga
rdh
(ab)
Eas
t/Wes
t (E
ffing
ham
in
let)/
Arc
tic
Hay
et a
l. 20
03 (B
aile
y 19
13,
Bai
ley
1910
, Bér
ard-
Ther
riaul
t et
al. 1
999,
Bru
nel 1
962,
Bur
sa
1963
)M
elos
ira o
ctag
ona
Mel
osira
oct
ogon
a S
chm
idt (
ab)
Eas
tB
aile
y 19
10M
elos
ira o
richa
lcea
Ken
.M
elos
ira o
richa
lcea
(Mer
tens
ex
Jürg
ens)
Küt
zing
(ab
)C
onfe
rva
oric
halc
ea M
erte
ns e
x Jü
rgen
sE
ast
Bai
ley
1910
Mel
osira
sp.
Mel
osira
sp.
Eas
t/Wes
t/Ar
ctic
San
cetta
& C
alve
rt 19
88 (B
ursa
19
63, B
ates
& S
train
200
6)M
elos
ira s
pp.
Mel
osira
spp
.W
est
McQ
uoid
& H
obso
n 20
01M
elos
ira s
ubfle
xilis
Kut
z.M
elos
ira s
ubfle
xilis
Kut
zing
(ab)
Eas
tB
aile
y 19
13 (B
aile
y 19
10)
Mel
osira
sub
sals
aM
elos
ira s
ubsa
lsa
Cle
ve-E
uler
(ab)
Eas
tB
rune
l 196
2
Mel
osira
sul
cata
Par
alia
sul
cata
(E
hren
berg
) Cle
ve
(ab)
Mel
osira
sul
cata
(Ehr
enbe
rg)
Küt
zing
Eas
tFr
itz 1
921
(Bru
nel 1
962,
Gra
n &
B
raar
ud 1
935,
Bai
ley
1910
)M
elos
ira v
aria
ns A
g.M
elos
ira v
aria
ns C
. Aga
rdh
(ab)
Lysi
goni
um v
aria
ns (
C.A
gard
h)
DeT
oni
Eas
tB
aile
y 19
13 (B
aile
y 19
10)
Mem
bran
eis
chal
leng
eri
Pftz
erM
embr
anei
s ch
alle
nger
i (G
runo
w)
Pad
dock
(ab)
Nav
icul
a ch
alle
nger
i G
runo
w,
Trop
idon
eis
anta
rctic
a (G
runo
w in
C
leve
& M
ölle
r) C
leve
Eas
tB
érar
d-Th
erria
ult e
t al.
1999
Mer
idio
n ci
rcul
are
Ag.
Mer
idio
n ci
rcul
are
(Gre
ville
) C.
Aga
rdh
(ab)
Exi
laria
circ
ular
ia (
Gre
ville
) G
revi
lle, E
chin
ella
circ
ular
is
Gre
ville
, Exi
laria
circ
ular
is
(Gre
ville
) C. A
gard
h
Eas
tB
aile
y 19
13 (B
rune
l 196
2)
Mer
idio
n in
term
ediu
m H
.L.S
.M
erid
ion
inte
rmed
ium
H.L
. Sm
ith
(ab)
Eas
t B
aile
y 19
13 (B
aile
y 19
10)
Mer
idio
n sp
p.M
erid
ion
spp.
Wes
tM
cQuo
id &
Hob
son
2001
Pag
e 48
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Meu
nier
a m
embr
anac
eaM
euni
era
mem
bran
acea
(Cle
ve)
P.C
. Silv
a (a
b)S
taur
opsi
s m
embr
anac
ea (
Cle
ve)
Meu
nier
, Nav
icul
a m
embr
anac
ea
Cle
ve, S
taur
onei
s m
embr
anac
ea
(Cle
ve) H
uste
dt
Arc
ticR
iede
l et a
l. 20
03
Min
idis
cus
chile
nsis
Min
idis
cus
chile
nsis
Riv
era
(ab)
Eas
tB
érar
d-Th
erria
ult e
t al.
1999
Min
idis
cus
com
icus
Min
idis
cus
com
icus
Tak
ano
(ab)
Eas
tB
érar
d-Th
erria
ult e
t al.
1999
Min
idis
cus
spp.
Has
leM
inid
iscu
s sp
p. H
asle
Wes
t (E
ffing
ham
In
let)
Hay
et a
l. 20
03 (S
ance
tta &
C
alve
rt 19
88, M
cQuo
id &
Hob
son
2001
)M
inid
iscu
s tri
ocul
atus
Min
idis
cus
trioc
ulat
us (
F.J.
R. T
aylo
r)
Has
le (a
b)C
osci
nodi
scus
trio
cula
tus
F.J
.R.
Tayl
orE
ast
Bér
ard-
Ther
riaul
t et a
l. 19
99
Min
utoc
ellu
lus
Has
le, v
on S
tosc
h an
d S
yver
tsen
sp.
Min
utoc
ellu
lus
Has
le, v
on S
tosc
h an
d S
yver
tsen
sp.
(ab)
Arc
ticLo
vejo
y et
al.
2002
Nav
icul
a ac
uta
S.B
.D.
Nav
icul
a ac
uta
Skv
ortz
ov (a
b)E
ast
Bai
ley
1913
Nav
icul
a al
gida
Nav
icul
a al
gida
Gru
now
(ab)
Arc
ticR
iede
l et a
l. 20
03N
avic
ula
amer
ican
a E
hr.
Sel
laph
ora
amer
ican
a (E
hren
berg
) D
.G. M
ann
(ab)
Nav
icul
a am
eric
ana
Ehr
enbe
rg
Eas
tB
aile
y 19
13
Nav
icul
a ba
cillu
m E
hren
berg
Sel
laph
ora
baci
llum
(Ehr
enbe
rg)
D.G
. Man
n (a
b)N
avic
ula
baci
llum
Ehr
enbe
rgA
rctic
Ant
onia
des
et a
l. 20
05
Nav
icul
a ba
ileya
na G
run.
Nav
icul
a ba
ileya
na G
runo
w e
x A
. S
chm
idt (
ab)
Eas
tB
aile
y 19
13
Nav
icul
a bi
nodi
s E
hren
berg
Nei
dium
bin
odis
(Ehr
enbe
rg)
Hus
tedt
(ab)
Eas
tR
awle
nce
1987
Nav
icul
a br
yoph
ila (
Pet
erse
n)
Lang
e-B
erta
lot
Nav
icul
a br
yoph
ila P
eter
sen
(ab)
Arc
ticA
nton
iade
s et
al.
2005
Nav
icul
a ch
iara
e La
nge-
Ber
talo
t &
Gen
kal
Nav
icul
a ch
iara
e La
nge-
Ber
talo
t &
Gen
kal (
ab)
Arc
ticA
nton
iade
s et
al.
2005
Nav
icul
a ci
ncta
(Ehr
enbe
rg) R
alfs
Nav
icul
a ci
ncta
(Ehr
enbe
rg) R
alfs
(a
b)N
avic
ula
umid
a B
ock,
Nav
icul
a in
utili
s K
rass
ke, N
avic
ula
cinc
ta
var.
heuf
lerii
Gru
now
, Pin
nula
ria
cinc
ta E
hren
berg
, Sch
izon
ema
cinc
tum
(E
hren
berg
) Kun
tze,
N
avic
ula
cari
var
. cin
cta
(Ehr
enbe
rg) L
ange
-Ber
talo
t
Arc
ticA
nton
iade
s et
al.
2005
Pag
e 49
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Nav
icul
a cl
emen
tis G
runo
wN
avic
ula
clem
entis
Gru
now
(ab
)P
laco
neis
cle
men
tis (
Gru
now
) E
.J.C
ox, N
avic
ula
incl
emen
tis
Hen
dey,
Nav
icul
a cl
emen
tis v
ar.
rhom
bica
Bro
ckm
ann,
S
chiz
onem
a cl
emen
tis (G
runo
w)
Kun
tze,
Pla
cone
is rh
ombi
ca
(Bro
ckm
ann)
Lan
ge-B
erta
lot
Arc
ticA
nton
iade
s et
al.
2005
Nav
icul
a cl
unth
ensi
s f.
rost
rata
Nav
icul
a cl
uthe
nsis
f. r
ostra
ta
(Sim
onse
n) H
uste
dt (a
b)N
avic
ula
pseu
dogl
acia
lis f
. ro
stra
ta S
imon
sen
Eas
tB
érar
d-Th
erria
ult e
t al.
1999
Nav
icul
a co
ccon
eifo
rmis
Gre
gory
ex
Gre
ville
Cav
inul
a co
ccon
eifo
rmis
(Gre
gory
ex
Gre
ville
) D.G
. Man
n &
A.J
. Stic
kle
(ab)
Nav
icul
a co
ccon
eifo
rmis
Gre
gory
ex
Gre
ville
Arc
ticA
nton
iade
s et
al.
2005
Nav
icul
a cr
ucig
era
Cle
veH
asle
a cr
ucig
era
(W. S
mith
) S
imon
sen
(ab)
Sta
uron
eis
cruc
iger
a (W
. Sm
ith)
Hei
berg
, Nav
icul
a cr
ucig
era
(W.
Sm
ith) C
leve
Eas
t (B
oF)
Laks
hmin
arya
na 1
983
Nav
icul
a cr
ypto
ceph
ala
Küt
zing
Nav
icul
a cr
ypto
ceph
ala
Küt
zing
(ab)
Nav
icul
a cr
ypto
ceph
ala
form
a te
rres
tris
Lund
, Sch
izon
ema
cryp
toce
phal
um(K
ützi
ng) K
untz
e,
Nav
icul
a cr
ypto
cefa
lsa
Lan
ge-
Ber
talo
t
Arc
ticA
nton
iade
s et
al.
2005
Nav
icul
a cy
prin
us
Nav
icul
a cy
prin
us (
Ehr
enbe
rg)
Küt
zing
(ab)
Eas
tB
aile
y 19
13
Nav
icul
a di
ceph
ala
(Ehr
enbe
rg)
W.S
mith
Nav
icul
a di
ceph
ala
(Ehr
enbe
rg)
W.S
mith
(ab)
Eas
tB
aile
y 19
13
Nav
icul
a di
gito
-rad
iata
(G
reg.
) R
alfs
.N
avic
ula
digi
to-r
adia
ta (
Gre
g.) R
alfs
(a
b)E
ast (
BoF
)R
awle
nce
1987
Nav
icul
a di
rect
a P
innu
laria
dire
cta
W.S
mith
(ab)
Eas
t/Arc
ticB
aile
y 19
13 (B
érar
d-Th
erria
ult e
t al
. 199
9, R
iede
l et a
l. 20
03,
Love
joy
et a
l. 20
02, R
ozan
ska
et
al. 2
008)
Nav
icul
a di
stan
s (W
. Sm
ith) R
alfs
Nav
icul
a di
stan
s(W
. Sm
ith) S
chm
idt
(ab)
Pin
nula
ria d
ista
ns W
.Sm
ithE
ast/W
est
(US
)/Arc
ticC
upp
1943
(Bai
ley
1913
, Bai
ley
1910
, Bur
sa 1
963,
Gra
n &
Bra
arud
19
35, H
obso
n &
McQ
uoid
200
1)
Nav
icul
a eg
regi
a H
uste
dtFa
llaci
a eg
regi
a (H
uste
dt) D
.G.
Man
n (a
b)N
avic
ula
egre
gia
Hus
tedt
Arc
ticA
nton
iade
s et
al.
2005
Pag
e 50
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Nav
icul
a el
liptic
aD
iplo
neis
elli
ptic
a (K
ützi
ng) C
leve
(a
b)N
avic
ula
ellip
tica
Küt
zing
Eas
tB
aile
y 19
10 (B
aile
y 19
13)
Nav
icul
a ga
ndru
pii
Kra
sske
Nav
icul
a ga
ndru
pii
(Pet
erse
n)
Kra
sske
(ab)
Arc
ticA
nton
iade
s et
al.
2005
Nav
icul
a ge
lida
Nav
icul
a ge
lida
Gru
now
Arc
ticR
iede
l et a
l. 20
03N
avic
ula
gem
mat
a G
rev.
Nav
icul
a ge
mm
ata
Gre
v. (a
b)E
ast (
BoF
)R
awle
nce
1987
Nav
icul
a ge
rloffi
i S
chim
ansk
iN
avic
ula
gerlo
ffii
Sch
iman
ski (
ab)
Arc
ticA
nton
iade
s et
al.
2005
Nav
icul
a gl
obul
ifera
Hus
tedt
Nav
icul
a gl
obul
ifera
Hus
tedt
(ab)
Arc
ticA
nton
iade
s et
al.
2005
Nav
icul
a gr
aeffi
i G
runo
w e
x A
. S
chm
idt
Nav
icul
a gr
aeffi
i G
runo
w e
x A
. S
chm
idt
(ab)
Eas
t (B
oF)
Raw
lenc
e 19
87
Nav
icul
a gr
anii
Nav
icul
a gr
anii
(Jør
gens
en) G
ran
(ab)
Sta
uron
eis
gran
ii E
. Jor
gens
en
Eas
t/Arc
ticB
rune
l 196
2 (R
iede
l et a
l. 20
03,
Love
joy
et a
l. 20
02)
Nav
icul
a gr
egar
ia D
onki
nN
avic
ula
greg
aria
Don
kin
(ab)
Sch
izon
ema
greg
ariu
m (D
onki
n)
Kun
tze
Wes
t (E
ffing
ham
In
let)
Hay
et a
l. 20
03 (B
érar
d-Th
erria
ult
et a
l. 19
99)
Nav
icul
a ha
loph
ilioi
des
Hus
tedt
Nav
icul
a ha
loph
ilioi
des
Hus
tedt
(ab)
Arc
ticA
nton
iade
s et
al.
2005
Nav
icul
a he
nned
yi S
.B.D
.Ly
rella
hen
nedy
i (W
. Sm
ith) S
tickl
e &
Man
n (a
b)N
avic
ula
henn
edyi
S.B
.D.
Eas
tB
aile
y 19
13
Nav
icul
a im
pexa
Nav
icul
a im
pexa
Hus
tedt
(ab)
Arc
ticR
iede
l et a
l. 20
03N
avic
ula
inso
ciab
ilis
Falla
cia
inso
ciab
ilis
(Kra
sske
) D.G
. M
ann
Nav
icul
a fri
tsch
ii J.
W.G
. Lun
d,
Nav
icul
a in
soci
abili
s K
rass
keA
rctic
Ant
onia
des
et a
l. 20
05
Nav
icul
a ka
riana
var
. frig
ida
Nav
icul
a ka
riana
var
. frig
ida
(Gru
now
) Cle
ve (a
b)E
ast
Bér
ard-
Ther
riaul
t et a
l. 19
99
Nav
icul
a kr
assk
ei H
uste
dtN
avic
ula
kras
skei
Hus
tedt
(ab)
Arc
ticA
nton
iade
s et
al.
2005
Nav
icul
a kr
yoph
ilaN
avic
ula
crio
phila
(Cas
traca
ne) D
e To
niN
avic
ula
kryo
phila
Cle
veA
rctic
Boo
th &
Hor
ner 1
997
Nav
icul
a le
nzii
Hus
t. in
Sch
mid
tN
avic
ula
lenz
ii H
ust (
ab)
Arc
ticA
nton
iade
s et
al.
2005
Nav
icul
a lib
er S
.B.D
.N
avic
ula
liber
W.S
mith
(ab)
Eas
tB
aile
y 19
13N
avic
ula
long
a R
alfs
.N
avic
ula
long
a (G
rego
ry) R
alfs
ex
Prit
char
d (a
b)P
innu
laria
long
a G
rego
ryE
ast
Bai
ley
1913
Nav
icul
a m
acul
ata
Bai
l.N
avic
ula
mac
ulat
a H
uste
dt (a
b)E
ast
Bai
ley
1913
(Bai
ley
1910
)N
avic
ula
maj
or K
tz.
Nav
icul
a m
ajor
Küt
zing
Eas
tB
aile
y 19
10 (B
aile
y 19
13)
Nav
icul
a m
arin
a R
alfs
Pet
rone
is m
arin
a (R
alfs
ex
Prit
char
d) D
.G. M
ann
(ab)
Nav
icul
a pu
nctu
lata
W. S
mith
, N
avic
ula
mar
ina
Ral
fs e
x P
ritch
ard
Eas
tB
aile
y 19
10
Pag
e 51
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Nav
icul
a m
esol
epta
Kut
z.P
innu
laria
mes
olep
ta (
Ehr
enbe
rg)
W. S
mith
(ab)
Nav
icul
a m
esol
epta
Kut
z.E
ast
Bai
ley
1913
Nav
icul
a m
inus
cula
A
dlaf
ia m
inus
cula
(G
runo
w) H
. La
nge-
Ber
talo
t (ab
)N
avic
ula
pseu
doex
iliss
ima
Hus
tedt
, Nav
icul
a im
portu
nata
H
uste
dt, N
avic
ula
mur
alis
G
runo
w, N
avic
ula
min
uscu
la v
ar.
mur
alis
(G
runo
w) L
ange
-Ber
talo
t, N
avic
ula
cloa
cina
Lan
ge-B
erta
lot
& B
onik
, Nav
icul
a m
inus
cula
G
runo
w
Eas
tB
aile
y 19
10
Nav
icul
a m
utic
a K
utzi
ngLu
ticol
a m
utic
a (K
ützi
ng) D
.G. M
ann
Nav
icul
a im
bric
ata
Boc
k,
Nav
icul
a m
utic
a K
ützi
ngA
rctic
Ant
onia
des
et a
l. 20
05
Nav
icul
a no
bilis
Ehr
.N
avic
ula
nobi
lis K
ützi
ng (a
b)E
ast
Bai
ley
1910
(Bai
ley
1913
)N
avic
ula
patu
la S
.B.D
.N
avic
ula
patu
la W
.Sm
ith (a
b)E
ast
Bai
ley
1913
Nav
icul
a pe
lagi
caN
avic
ula
pela
gica
Cle
ve (a
b)S
taur
opsi
s pe
lagi
ca (
Cle
ve)
Meu
nier
E
ast/A
rctic
Bér
ard-
Ther
riaul
t et a
l. 19
99
(Rie
del e
t al.
2003
, Lov
ejoy
et a
l. 20
02, B
ooth
& H
orne
r 199
7)N
avic
ula
pellu
cidu
laN
avic
ula
pellu
cidu
la H
uste
dt (a
b)A
rctic
Rie
del e
t al.
2003
Nav
icul
a pe
regr
ina
Ktz
.N
avic
ula
pere
grin
a (E
hren
berg
) K
ützi
ng (a
b)P
innu
laria
per
egrin
a E
hren
berg
Eas
t Li
nkle
tter e
t al.
1977
(Bai
ley
1913
&
Bai
ley
1910
)N
avic
ula
perm
agna
Bai
l.C
alon
eis
perm
agna
(J.
W. B
aile
y)
Cle
ve (a
b)N
avic
ula
perm
agna
(B
aile
y) A
.M.
Edw
ards
Eas
tB
aile
y 19
10 (B
aile
y 19
13)
Nav
icul
a pe
rmin
uta
Gru
now
in V
an
Heu
rck
Nav
icul
a pe
rmin
uta
Gru
now
(ab
)N
avic
ula
dist
erta
Hus
tedt
, N
avic
ula
cryp
toce
phal
a v
ar.
perm
inut
a (G
runo
w) C
leve
Wes
t (E
ffing
ham
In
let)
Hay
et a
l. 20
03 (B
érar
d-Th
erria
ult
et a
l. 19
99)
Nav
icul
a ph
ylle
pta
Kut
zing
Nav
icul
a ph
ylle
pta
Küt
zing
(ab)
Nav
icul
a ph
ylle
pta
Küt
zing
, N
avic
ula
lanc
eola
ta v
ar. p
hylle
pta
(Küt
zing
) van
Heu
rck
Arc
ticA
nton
iade
s et
al.
2005
Nav
icul
a ps
eudo
tene
lloid
es
Kra
sske
Nav
icul
a ps
eudo
tene
lloid
es
Cho
lnok
y (a
b)A
rctic
Ant
onia
des
et a
l. 20
05
Nav
icul
a pu
pula
var
. mut
ata
(Kra
sske
) Hus
tedt
Nav
icul
a pu
pula
var
. mut
ata
(Kra
sske
) Hus
tedt
(ab)
Arc
ticA
nton
iade
s et
al.
2005
Nav
icul
a qu
adrip
edis
Nav
icul
a se
pten
trion
alis
(G
runo
w)
Gra
n (a
b)N
avic
ula
quad
riped
is C
leve
-Eul
erE
ast
Bru
nel 1
962
Pag
e 52
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Nav
icul
a ra
dios
a S
.B.D
.N
avic
ula
radi
osa
Küt
zing
(ab)
Nav
icul
a ra
dios
a va
r. te
nella
(B
rébi
sson
) Van
Heu
rck,
S
chiz
onem
a ra
dios
um (K
ützi
ng)
Kun
tze
Eas
tB
aile
y 19
13
Nav
icul
a re
curv
ata
Nav
icul
a re
curv
ata
Gra
n (a
b)A
rctic
Rie
del e
t al.
2003
Nav
icul
a se
men
Ehr
.N
avic
ula
sem
en E
hren
berg
(ab)
Eas
tB
aile
y 19
13N
avic
ula
sept
entri
onal
isN
avic
ula
sept
entri
onal
is (
Gru
now
) G
ran
(ab)
Sta
uron
eis
sept
entri
onal
is
Gru
now
, Nav
icul
a qu
adrip
edis
C
leve
-Eul
er
Eas
t/Arc
ticB
érar
d-Th
erria
ult e
t al.
1999
(B
rune
l 196
2, R
iede
l et a
l. 20
03,
Love
joy
et a
l. 20
02, B
ooth
&
Hor
ner 1
997)
Nav
icul
a sp
p.N
avic
ula
spp.
Eas
t/Wes
t (U
S)/A
rctic
Hor
ner e
t al.
2005
(Lov
ejoy
et a
l. 20
02, B
ates
& S
train
200
6, B
ooth
&
Hor
ner 1
997,
McQ
uoid
&
Hob
son
2001
)N
avic
ula
soeh
rens
is K
rass
keN
avic
ula
soeh
rens
is K
rass
ke (a
b)A
rctic
Ant
onia
des
et a
l. 20
05
Nav
icul
a su
perb
a va
r. su
bacu
taN
avic
ula
supe
rba
var
. sub
acut
a G
ran
Arc
ticR
iede
l et a
l. 20
03
Nav
icul
a tra
nsita
ns C
l.N
avic
ula
trans
itans
Cl.
(ab)
Eas
t (B
oF)/A
rctic
Raw
lenc
e 19
87 (B
ooth
& H
orne
r 19
97)
Nav
icul
a tra
nsita
ns v
ar. d
eras
aN
avic
ula
trans
itans
var
. der
asa
(Gru
now
) Cle
ve (a
b)N
avic
ula
dera
sa G
runo
w
Eas
tB
érar
d-Th
erria
ult e
t al.
1999
Nav
icul
a tra
nsita
ns v
ar. d
eras
a f.
delic
atul
aN
avic
ula
trans
itans
var
. der
asa
f. de
licat
ula
Hei
mda
l (ab
)E
ast/A
rctic
Bér
ard-
Ther
riaul
t et a
l. 19
99
(Lov
ejoy
et a
l. 20
02)
Nav
icul
a tri
gono
ceph
ala
Nav
icul
a tri
gono
ceph
ala
Cle
veE
ast/A
rctic
Rie
del e
t al.
2003
(Boo
th &
Hor
ner
1997
)N
avic
ula
trigo
noce
phal
a va
r. de
pres
sa
Nav
icul
a tri
gono
ceph
ala
var
. de
pres
sa Ø
stru
pA
rctic
Rie
del e
t al.
2003
Nav
icul
a tri
punc
tata
(O
.F. M
ull.)
B
ory
v. tr
ipun
ctat
aN
avic
ula
tripu
ncta
ta v
ar. t
ripun
ctat
a (O
.F. M
ülle
r) B
ory
(ab)
Eas
tR
awle
nce
1987
Nav
icul
a va
nhoe
ffeni
iN
avic
ula
vanh
oeffe
nii
Gra
n (a
b)S
taur
opsi
s va
nhoe
ffeni
i (G
ran)
M
euni
er
Eas
t/Arc
ticB
érar
d-Th
erria
ult e
t al.
1999
(R
iede
l et a
l. 20
03, L
ovej
oy e
t al.
2002
, Gra
n &
Bra
arud
193
5, B
ooth
&
Hor
ner 1
997)
Nav
icul
a vi
ridis
Ktz
.N
avic
ula
virid
is (
Nitz
sch)
Küt
zing
Eas
tB
aile
y 19
10 (B
aile
y 19
13)
Pag
e 53
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Nav
icul
a vu
lpin
a K
ützi
ngN
avic
ula
vulp
ina
Küt
zing
(ab)
Sch
izon
ema
vulp
inum
(Küt
zing
) K
untz
e, N
avic
ula
grac
ilis
var
. vu
lpin
a (K
ützi
ng) R
aben
hors
t, N
avic
ula
virid
ula
var
. vul
pina
(K
ützi
ng) L
ange
-Ber
talo
t
Arc
ticA
nton
iade
s et
al.
2005
Nei
dium
am
plia
tum
(E
hren
berg
) K
ram
mer
Nei
dium
am
plia
tum
(Ehr
enbe
rg)
Kra
mm
er (
ab)
Nav
icul
a am
plia
ta E
hren
berg
, N
eidi
um ir
idis
var
. am
plia
tum
(E
hren
berg
) Cle
ve
Arc
ticA
nton
iade
s et
al.
2005
Nei
dium
ber
gii
(Cle
ve) K
ram
mer
Nei
dium
ber
gii
(Cle
ve) K
ram
mer
(a
b)A
rctic
Ant
onia
des
et a
l. 20
05
Nei
dium
bin
ode
(Ehr
.) H
ust.
v.
bino
deN
eidi
um b
inod
e (E
hr.)
Hus
t. v.
bi
node
E
ast
Raw
lenc
e 19
87
Neo
dent
icul
a se
min
ae
Neo
dent
icul
a se
min
ae(R
. Sim
onse
n &
T. K
anay
a) F
. Aki
ba &
Y.
Yan
agis
awa
(ab)
Den
ticul
a se
min
ae R
. Sim
onse
n &
T. K
anay
a, D
entic
ula
mar
ina
Sem
ina,
Den
ticul
opsi
s se
min
ae
(Sim
onse
n &
Kan
aya)
Sim
onse
n
Eas
t/Wes
tM
artin
& L
eGre
sley
200
8
Nitz
schi
a am
phib
ia G
runo
wN
itzsc
hia
amph
ibia
Gru
now
(ab)
Arc
ticA
nton
iade
s et
al.
2005
(Zw
icke
r 19
98)
Nitz
schi
a an
gula
ris S
.B.D
.N
itzsc
hia
angu
laris
W.S
mith
(ab)
Eas
tB
aile
y 19
13 (B
aile
y 19
10)
Nitz
schi
a bi
loba
ta
Nitz
schi
a bi
loba
ta W
. Sm
ith (a
b)E
ast
Lin
klet
ter e
t al.
1977
(Bai
ley
1913
&
Bai
ley
1910
)N
itzsc
hia
bilo
bata
var
. min
or G
run.
Nitz
schi
a bi
loba
ta v
ar. m
inor
Gru
n.
(ab)
Wes
t (U
S)
Cup
p 19
43
Nitz
schi
a br
evis
sim
a G
runo
w in
V
an H
eurc
kN
itzsc
hia
brev
issi
ma
Gru
now
Nitz
schi
a pa
rvul
a Le
wis
, N
itzsc
hia
parv
ula
var.
terr
icol
a Lu
nd
Eas
tTh
aler
200
7
Nitz
schi
a ci
rcum
suta
Bai
l.N
itzsc
hia
circ
umsu
ta (B
aile
y)
Gru
now
(ab
)S
urire
lla c
ircum
suta
J.W
.Bai
ley
Eas
tB
aile
y 19
13 (B
aile
y 19
10)
Nitz
schi
a cl
ausi
i H
antz
sch
Nitz
schi
a cl
ausi
i H
antz
sch
(ab)
Arc
ticA
nton
iade
s et
al.
2005
Pag
e 54
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Nitz
schi
a cl
oste
rium
Nitz
schi
a cl
oste
rium
(E
hren
berg
) W.
Sm
ith (a
b)C
erat
onei
s cl
oste
rium
Ehr
enbe
rg,
Nitz
schi
ella
clo
ster
ium
(E
hren
berg
) Rab
enho
rst,
Nitz
schi
a lo
ngis
sim
a v
ar.
clos
teriu
m (E
hren
berg
) Van
H
eurc
k, N
itzsc
hia
curv
irost
ris v
ar.
clos
teriu
m (E
hren
berg
) De
Toni
, N
itzsc
hiel
la lo
ngis
sim
a v
ar.
clos
teriu
m (E
hren
berg
) Per
agal
lo
& P
erag
allo
, Hom
oeoc
ladi
a cl
oste
rium
(Ehr
enbe
rg) K
untz
e,
Cyl
indr
othe
ca c
lost
eriu
m
(Ehr
enbe
rg) R
eim
ann
& L
ewin
Eas
t/Arc
ticB
rune
l 196
2 (F
ritz
1921
, Bur
sa
1963
, Bai
ley
1910
, Bai
ley
1913
)
Nitz
schi
a co
mm
utat
a G
runo
wN
itzsc
hia
com
mut
ata
Gru
now
(ab
)N
itzsc
hia
pseu
doam
phio
xys
Hus
tedt
Arc
ticA
nton
iade
s et
al.
2005
Nitz
schi
a co
nstri
cta
Gru
n.N
itzsc
hia
cons
trict
a (G
rego
ry)
Gru
now
E
ast
Bai
ley
1910
Nitz
schi
a bi
loba
ta
Nitz
schi
a bi
loba
ta W
. Sm
ith (a
b)E
ast
Fritz
192
1N
itzsc
hia
debi
lis (
Arn
ott)
Gru
now
Tryb
lione
lla d
ebili
s A
rnot
t ex
O'M
eara
(ab)
Nitz
schi
a de
bilis
(A
rnot
t) G
runo
w
Arc
ticA
nton
iade
s et
al.
2005
Nitz
schi
a de
licat
issi
ma
Pse
udo-
nitz
schi
a de
licat
issi
ma
(Cle
ve) H
eide
n (a
b)N
itzsc
hia
delic
atis
sim
a C
leve
Eas
t/Wes
t/Ar
ctic
San
cetta
& C
alve
rt 19
88 (B
rune
l 19
62, B
ursa
196
3, G
ran
& B
raar
ud
1935
)N
itzsc
hia
diss
ipat
a va
r. m
edia
(H
antz
sch)
Gru
now
in V
an H
eurc
kN
itzsc
hia
diss
ipat
a va
r. m
edia
(H
antz
sch)
Gru
now
(ab
)N
itzsc
hia
tenu
is v
ar. m
edia
(H
antz
sch)
Rab
enho
rst,
Nitz
schi
a m
edia
Han
tzsc
h
Arc
ticA
nton
iade
s et
al.
2005
Nitz
schi
a ep
ithem
ioid
es G
runo
wN
itzsc
hia
epith
emoi
des
Gru
now
(ab)
Nitz
schi
a ep
ithem
iform
is L
ange
-B
erta
lot
Eas
tTh
aler
200
7
Nitz
schi
a fil
iform
is (
W.S
mith
) Van
H
eurc
kN
itzsc
hia
filifo
rmis
(W
.Sm
ith)
Hus
tedt
(ab)
Eas
tG
inn
(per
s. c
omm
.)1
Nitz
schi
a fo
ntifu
gaN
itzsc
hia
font
ifuga
Cho
lnok
y (a
b)E
ast
Bér
ard-
Ther
riaul
t et a
l. 19
99N
itzsc
hia
frigi
daN
itzsc
hia
frigi
da G
runo
w (a
b)E
ast/A
rctic
Bér
ard-
Ther
riaul
t et a
l. 19
99
(Rie
del e
t al.
2003
, Bur
sa 1
963,
Lo
vejo
y et
al.
2002
, Boo
th &
H
orne
r 199
7, R
ozan
ska
et a
l. 20
08)
Pag
e 55
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Nitz
schi
a fru
stul
um (
Küt
zing
) G
runo
w in
Cle
ve a
nd G
runo
wN
itzsc
hia
aust
riaca
Hus
tedt
(ab)
Nitz
schi
a fru
stul
um (
Küt
zing
) G
runo
w
Wes
t (E
ffing
ham
In
let)
Hay
et a
l. 20
03 (Z
wic
ker 1
998)
Nitz
schi
a gr
acili
s N
itzsc
hia
grac
ilis
Han
tzsc
h (a
b)E
ast
Bai
ley
1913
Nitz
schi
a gr
anul
ata
Tryb
lione
lla g
ranu
lata
(G
runo
w)
D.G
. Man
n (a
b)N
itzsc
hia
gran
ulat
a G
runo
wE
ast
Bai
ley
1910
Nitz
schi
a hu
dson
iiN
itzsc
hia
huds
onii
M. P
oulin
& A
. C
ardi
nal (
ab)
Arc
ticR
iede
l et a
l. 20
03
Nitz
schi
a in
cons
picu
a G
runo
wN
itzsc
hia
inco
nspi
cua
Gru
now
(ab)
Wes
t (E
ffing
ham
In
let)/
Arc
tic
Hay
et a
l. 20
03 (A
nton
iade
s et
al.
2005
)
Nitz
schi
a la
evis
sim
aN
itzsc
hia
laev
issi
ma
Gru
now
(ab)
Arc
ticR
iede
l et a
l. 20
03 (B
ooth
& H
orne
r 19
97)
Nitz
schi
a le
vide
nsis
Ant
.Tr
yblio
nella
levi
dens
is W
. Sm
ith (a
b)N
itzsc
hia
levi
dens
is (W
. Sm
ith)
Gru
now
Eas
tB
aile
y 19
10
Nitz
schi
a lig
owsk
ii W
itkow
ski,
Lang
e-B
erta
lot,
Koc
iole
k &
B
rzez
insk
a
Nitz
schi
a lig
owsk
ii W
itkow
ski,
Lang
e-B
erta
lot,
Koc
iole
k &
Brz
ezin
ska
(col
)E
ast
Thal
er 2
007
Nitz
schi
a lin
earis
W. S
m.
Nitz
schi
a lin
earis
(A
gard
h) W
. Sm
ith
(ab)
Eas
tB
aile
y 19
13
Nitz
schi
a li
neol
a G
ran
Pse
udo-
nitz
schi
a lin
eola
(C
leve
) H
asle
(ab)
Nitz
schi
a lin
eola
Cle
veA
rctic
Bur
sa 1
963
Nitz
schi
a lit
tora
lis G
run.
Tryb
lione
lla li
ttora
lis (
Gru
now
) D.G
. M
ann
(ab)
Nitz
schi
a lit
tora
lis G
runo
wE
ast
Bai
ley
1910
Nitz
schi
a lo
ngis
sim
a R
alfs
Nitz
schi
a lo
ngis
sim
a (B
rébi
sson
in
Küt
zing
) Ral
fs (
ab)
Nitz
schi
a bi
rost
rata
W. S
mith
, C
erat
onei
s lo
ngis
sim
a B
rébi
sson
Eas
t/Wes
t/Ar
ctic
Bai
ley
1910
(Bér
ard-
Ther
riaul
t et
al. 1
999,
Bru
nel 1
921,
Lov
ejoy
et
al. 2
002,
Bai
ley
1913
, Roz
ansk
a et
al
. 200
8)N
itzsc
hia
long
issi
ma
var.
clos
teriu
mN
itzsc
hia
clos
teriu
m (
Ehr
enbe
rg) W
. S
mith
(ab)
Nitz
schi
a lo
ngis
sim
a v
ar.
clos
teriu
m (
Ehr
enbe
rg) V
an
Heu
rck
Eas
tB
rune
l 196
2
Nitz
schi
a lo
ngic
ruris
N
itzsc
hia
long
icru
ris
Eas
t (N
S)
Raf
use
(per
s. c
omm
.)4
Nitz
schi
a ne
ofrig
ida
Nitz
schi
a ne
ofrig
ida
Med
lin (a
b)A
rctic
Boo
th &
Hor
ner 1
997
Nitz
schi
a pa
cific
a C
upp
Nitz
schi
a pa
cific
a C
upp
(ab)
Wes
t (U
S)
Cup
p 19
43N
itzsc
hia
pale
a (K
ützi
ng) W
.Sm
ithN
itzsc
hia
pale
a (K
ützi
ng) W
.Sm
ith
(ab)
Arc
ticA
nton
iade
s et
al.
2005
Nitz
schi
a pa
leac
ea G
runo
wN
itzsc
hia
pale
acea
Gru
nov
(ab)
Arc
ticA
nton
iade
s et
al.
2005
Pag
e 56
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Nitz
schi
a pa
lust
ris H
uste
dtN
itzsc
hia
palu
stris
Hus
tedt
(ab)
Arc
ticA
nton
iade
s et
al.
2005
Nitz
schi
a pa
rado
xaTr
yblio
nella
gra
nula
ta (G
runo
w)
D.G
. Man
nE
ast
Bai
ley
1910
Nitz
schi
a pa
rado
xa v
ar. t
umid
ula
Bac
illar
ia p
axill
ifer
var.
tum
idul
a (G
runo
w) W
itkow
ski,
Lang
e-B
erta
lot
& M
etze
ltin
(ab)
Arc
ticR
iede
l et a
l. 20
03
Nitz
schi
a pe
lluci
daN
itzsc
hia
pellu
cida
Gru
now
(ab)
Eas
tB
érar
d-Th
erria
ult e
t al.
1999
(T
rites
200
5)N
itzsc
hia
perm
inut
a (G
runo
w) M
. P
erag
allo
Nitz
schi
a pe
rmin
utum
(G
runo
w) M
. P
erag
allo
(ab)
Nitz
schi
a hi
emal
is H
uste
dt,
Nitz
schi
a fru
stul
um v
ar.
perm
inut
um G
runo
w
Wes
t (E
ffing
ham
In
let)/
Arc
tic
Hay
et a
l. 20
03 (A
nton
iade
s et
al.
2005
)
Nitz
schi
a pl
ana
W. S
m.
Nitz
schi
a pl
ana
W.S
mith
(ab)
Eas
tB
aile
y 19
13N
itzsc
hia
pola
risN
itzsc
hia
pola
ris G
runo
w e
x C
leve
(a
b)A
rctic
Boo
th &
Hor
ner 1
997
Nitz
schi
a pu
ncta
ta G
runo
wN
itzsc
hia
punc
tata
Gru
now
(ab)
Eas
tLi
nkle
tter e
t al.
1977
(Bai
ley
1910
)
Nitz
schi
a pu
ngen
sP
seud
o-ni
tzsc
hia
pung
ens
(Gru
now
ex
Cle
ve) H
asle
(ab
)N
itzsc
hia
pung
ens
Gru
now
ex
Cle
ve
Eas
tS
ance
tta &
Cal
vert
1988
(Joh
nson
19
88)
Nitz
schi
a pu
ngen
s va
r. at
lant
ica
Cle
veN
itzsc
hia
pung
ens
var.
atla
ntic
a C
l. (a
b)W
est (
US
)C
upp
1943
Nitz
schi
a pu
ra H
uste
dtN
itzsc
hia
pura
Hus
tedt
(ab)
Arc
ticA
nton
iade
s et
al.
2005
Nitz
schi
a pu
silla
Gru
now
Nitz
schi
a pu
silla
Gru
now
(ab)
Nitz
schi
a kü
tzin
gian
a H
ilse,
N
itzsc
hia
obtu
sang
ula
Hus
tedt
, N
itzsc
hia
indi
stin
cta
Mic
hailo
v,
Nitz
schi
a am
phio
xys
var
. pus
illa
(Gru
now
) May
er, H
antz
schi
a am
phio
xys
var
. pus
illa
(Gru
now
) D
ippe
l, N
itzsc
hia
indi
stin
cta
Mic
hailo
v
Arc
ticA
nton
iade
s et
al.
2005
Nitz
schi
a re
ctilo
nga
Nitz
schi
a re
ctilo
nga
Taka
no (a
b)E
ast
Bér
ard-
Ther
riaul
t et a
l. 19
99N
itzsc
hia
scab
raN
itzsc
hia
scab
ra C
leve
(ab)
Arc
ticR
iede
l et a
l. 20
03N
itzsc
hia
scal
aris
N
itzsc
hia
scal
aris
(E
hren
b.) W
.Sm
. (a
b)E
ast
Link
lette
r et a
l. 19
77 (B
aile
y 19
10)
Nitz
schi
a sc
utel
lum
S.B
.D.
Nitz
schi
a sc
utel
lum
(W
.Sm
ith) W
olle
(a
b)E
ast
Bai
ley
1913
Nitz
schi
a se
riata
Pse
udo-
nitz
schi
a se
riata
(C
leve
) H.
Per
agal
lo (a
b)N
itzsc
hia
seria
ta C
leve
E
ast/A
rctic
Fritz
192
1 (B
rune
l 196
2, B
ursa
19
63, G
ran
& B
raar
ud 1
935)
Pag
e 57
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Nitz
schi
a si
gma
W. S
mith
Nitz
schi
a si
gma
(Küt
zing
) W. S
mith
(a
b)S
igm
atel
la s
igm
a (K
ützi
ng)
Fren
guel
li, H
omoe
ocla
dia
sigm
a (K
ützi
ng) K
untz
e, S
yned
ra s
igm
a K
ützi
ng
Eas
tLi
nkle
tter e
t al.
1977
(Bai
ley
1913
, B
aile
y 19
10, F
ritz
1921
, Tha
ler
2007
, Trit
es 2
005)
Nitz
schi
a si
gmoi
dea
S.B
.D.
Nitz
schi
a si
gmoi
dea
(Nitz
sch)
W.
Sm
ith (a
b)B
acill
aria
sig
moi
dea
Nitz
sch,
N
itzsc
hia
elon
gata
Has
sal,
Hom
oeoc
ladi
a si
gmoi
dea
(Nitz
sch)
Elm
ore,
Fru
stul
ia
sigm
oide
a (N
itzsc
h) J
enne
r, S
yned
ra s
igm
oide
a (N
itzsc
h)
Küt
zing
, Cym
bella
sig
moi
dea
(Nitz
sch)
C. A
gard
h
Eas
tB
aile
y 19
13 (B
aile
y 19
10)
Nitz
schi
a sp
.N
itzsc
hia
sp.
Arc
ticB
ursa
196
3 (B
ooth
& H
orne
r 199
7)
Nitz
schi
a su
btili
s (K
tz.)
Gru
n.N
itzsc
hia
subt
ilis
Gru
now
(ab)
Eas
t (B
oF)
Raw
lenc
e 19
87N
itzsc
hia
ther
mal
is K
tz.
Nitz
schi
a th
erm
alis
Küt
zing
(ab)
Eas
tLi
nkle
tter e
t al.
1977
(Bai
ley
1910
)
Nitz
schi
a try
blio
nella
H.
Nitz
schi
a try
blio
nella
Han
tzsc
h (a
b) T
rybl
ione
lla h
antz
schi
ana
var.
calid
a (G
runo
v) V
an H
eurc
k,
Tryb
lione
lla h
antz
schi
ana
var.
min
or P
layf
air,
Try
blio
nella
ha
ntzs
chia
na v
ar. o
vata
(L
ager
sted
t) P
layf
air,
Tryb
lione
lla
hant
zsch
iana
var
. vic
toria
e (G
runo
v) P
layf
air,
Tryb
lione
lla
grac
ilis
W. S
mith
, Try
blio
nella
ha
ntzs
chia
na v
ar. a
ustra
lica
Pla
yfai
r, Tr
yblio
nella
ha
ntzs
chia
na G
runo
w
Eas
tB
aile
y 19
13 (B
aile
y 19
10)
Nitz
schi
a tu
bico
laN
itzsc
hia
tubi
cola
Gru
now
(ab)
Eas
tZw
icke
r 199
8N
itzsc
hia
verm
icul
aris
Han
.N
itzsc
hia
verm
icul
aris
(K
ützi
ng)
Han
tzsc
h (a
b)E
ast
Bai
ley
1913
Nitz
schi
a vi
trea
Non
n.N
itzsc
hia
vitre
a G
. Nor
man
(ab)
Eas
tB
aile
y 19
13
Odo
ntel
la a
urita
(Lyn
gbye
) C.A
. A
gard
hO
dont
ella
aur
ita (L
yngb
ye) C
.A.
Aga
rdh
(ab)
Dia
tom
a au
ritum
Lyn
gbye
, B
iddu
lphi
a au
rita
(Lyn
gbye
) B
rébi
sson
Eas
t/Wes
tH
orne
r et a
l. 20
05 (S
ance
tta &
C
alve
rt 19
88, B
érar
d-Th
erria
ult e
t al
. 199
9, B
ates
& S
train
200
6,
McQ
uoid
& H
obso
n 20
01)
Pag
e 58
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Odo
ntel
la lo
ngic
ruris
(G
revi
lle)
Hob
anO
dont
ella
long
icru
ris (G
revi
lle)
Hob
an (
ab)
Bid
dulp
hia
long
icru
ris v
ar. h
yalin
a (J
.L.B
. Sch
röde
r) E
.E. C
upp,
B
iddu
lphi
a hy
alin
a J
.L.B
. S
chrö
der,
Bid
dulp
hia
long
icru
ris
Gre
ville
Eas
t/Wes
tH
orne
r et a
l. 20
05 (S
ance
tta &
C
alve
rt 19
88, M
cQuo
id &
Hob
son
2001
, Car
ver &
Mal
let 2
002)
Odo
ntel
la m
obili
ensi
s O
dont
ella
mob
ilien
sis
(J.W
. Bai
ley)
G
runo
wZy
goce
ros
mob
ilien
sis
J.W
. B
aile
y, B
iddu
lphi
a m
obili
ensi
s (J
.W. B
aile
y) G
runo
w
Wes
tH
orne
r 200
2
Odo
ntel
la o
btus
a K
utz.
Bid
dulp
hia
obtu
sa (K
ützi
ng) R
alfs
(a
b)O
dont
ella
obt
usa
Kut
z.E
ast
Mar
tin e
t al.
1999
Odo
ntel
la re
gia
Odo
ntel
la re
gia
(Sch
ultz
e)
Sim
onse
n (a
b)B
iddu
lphi
a re
gia
(Sch
ultz
e)
Ost
enfe
ld
Eas
tB
érar
d-Th
erria
ult e
t al.
1999
Odo
ntel
la s
inen
sis
(Gre
ville
) G
runo
wO
dont
ella
sin
ensi
s (G
revi
lle) G
runo
w
(ab)
Bid
dulp
hia
sine
nsis
Gre
ville
Eas
tB
érar
d-Th
erria
ult e
t al.
1999
Odo
ntid
ium
mes
odon
S.B
.D.
Dia
tom
a m
esod
on (
Ehr
enbe
rg)
Küt
zing
(ab)
Odo
ntid
ium
mes
odon
S.B
.D.
Eas
tB
aile
y 19
13
Odo
ntid
ium
mut
abile
S.
Odo
ntid
ium
mut
abile
W. S
mith
(ab)
Eas
tB
aile
y 19
10 (B
aile
y 19
13)
Ope
phor
a cf
. hor
stia
na W
itkow
ski
O. m
utab
ilis
(Gru
now
) S
abbe
& W
yver
man
(ab)
Ope
phor
a c
f. ho
rstia
na W
itkow
ski
Wes
t (E
ffing
ham
In
let)
Hay
et a
l. 20
03
Ope
phor
a m
arin
a (G
rego
ry) P
etit
Ope
phor
a m
arin
a (G
rego
ry) P
etit
(a
b)M
erid
ion
mar
inum
Gre
gory
Wes
t (E
ffing
ham
In
let)
Hay
et a
l. 20
03
Ope
phor
a ol
seni
i M
ølle
rO
peph
ora
mut
abili
s (G
runo
w) S
abbe
&
Wyv
erm
anO
peph
ora
olse
nii
Møl
ler
Wes
t (E
ffing
ham
In
let)
Hay
et a
l. 20
03
Par
alia
mar
ina
Par
alia
sul
cata
(E
hren
berg
) Cle
veP
aral
ia m
arin
a (W
. Sm
ith)
Hei
berg
Eas
t/Wes
tB
ates
& S
train
200
6 (M
cQuo
id &
H
obso
n 19
97)
Par
alia
sul
cata
(E
hren
berg
) Cle
veP
aral
ia s
ulca
ta (E
hren
berg
) Cle
ve
(ab)
Mel
osira
sul
cata
(E
hren
berg
) K
ützi
ng, G
allio
nella
sul
cata
E
hren
berg
, Par
alia
mar
ina
(W.S
mith
) Hei
berg
, Orth
osei
ra
mar
ina
W.S
mith
Eas
t/Wes
tH
orne
r et a
l. 20
05 (L
inkl
ette
r et a
l. 19
77, S
ance
tta &
Cal
vert
1988
, B
érar
d-Th
erria
ult e
t al.
1999
, B
rune
l 192
1, T
rites
200
5, M
cQuo
id
& H
obso
n 20
01, H
ay e
t al.
2003
)
Pau
liella
taen
iata
Pau
liella
taen
iata
(G
runo
w) F
. E.
Rou
nd &
P. W
. Bas
son
(ab)
Eas
t/Arc
ticB
érar
d-Th
erria
ult e
t al.
1999
(R
iede
l et a
l. 20
03, L
ovej
oy e
t al.
2002
)
Pag
e 59
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Pet
rodi
ctyo
n ge
mm
a (E
hren
berg
) D
.G. M
ann
in R
ound
et a
l.P
etro
dict
yon
gem
ma
(Ehr
enbe
rg)
D.G
. Man
n (a
b)S
urire
lla g
emm
a (E
hren
berg
)E
ast
Thal
er 2
007
Pin
nula
ria a
cuta
W. S
m.
Pin
nula
ria a
cuta
W.S
mith
(ab)
Eas
tB
aile
y 19
13P
innu
laria
bor
ealis
Ehr
enbe
rgP
innu
laria
bor
ealis
Ehr
enbe
rg (
ab)
Pin
nula
ria c
hile
nsis
Ble
isch
, N
avic
ula
bore
alis
(Ehr
enbe
rg)
Küt
zing
, Sch
izon
ema
bore
ale
(Ehr
enbe
rg) K
untz
e
Arc
ticA
nton
iade
s et
al.
2005
Pin
nula
ria c
yprin
us
Pin
nula
ria c
yprin
us C
.G. E
hren
berg
(a
b)E
ast
Bai
ley
1913
Pin
nula
ria d
iver
gens
W.S
mith
Pin
nula
ria d
iver
gens
W.S
mith
(ab)
Arc
ticA
nton
iade
s et
al.
2005
Pin
nula
ria in
term
edia
(La
gers
tedt
) C
leve
Pin
nula
ria in
term
edia
(La
gers
tedt
) C
leve
(ab)
Arc
ticA
nton
iade
s et
al.
2005
Pin
nula
ria in
terr
upta
W.S
mith
Pin
nula
ria in
terr
upta
W.S
mith
(ab)
Arc
ticA
nton
iade
s et
al.
2005
Pin
nula
ria m
ajor
(K
utz)
Rab
enho
rst
Pin
nula
ria m
ajor
(K
utz)
Rab
enho
rst
(ab)
Eas
tG
inn
(per
s. c
omm
.)1
Pin
nula
ria n
obili
s E
hr.
Pin
nula
ria n
obili
s E
hren
berg
(ab
)N
avic
ula
nobi
lis E
hren
berg
Eas
tB
aile
y 19
13
Pin
nula
ria o
bscu
ra K
rass
keP
innu
laria
obs
cura
Kra
sske
(ab)
Arc
ticA
nton
iade
s et
al.
2005
Pin
nula
ria o
valis
Hils
eD
iplo
neis
ova
lis (
Hils
e) C
leve
(ab)
Pin
nula
ria o
valis
Hils
eE
ast
Pat
rick
& R
eim
er 1
966
Pin
nula
ria q
uadr
atar
ea v
ar.
bico
ntra
cta
Pin
nula
ria q
uadr
atar
ea v
ar.
bico
ntra
cta
(E.V
. Øst
rup)
H.H
. Gra
nA
rctic
Rie
del e
t al.
2003
Pin
nula
ria q
uadr
atar
ea v
ar.
cons
trict
aP
innu
laria
qua
drat
area
var
. co
nstri
cta
(Øst
rup)
Hei
den
(ab)
Nav
icul
a pi
nnul
aria
var
. co
nstri
cta
Øst
rup
Arc
ticR
iede
l et a
l. 20
03
Pin
nula
ria s
emiin
flata
Pin
nula
ria s
emiin
flata
(E
.V. Ø
stru
p)
H.H
. Gra
n (a
b)A
rctic
Rie
del e
t. al
. 200
3 (B
ooth
&
Hor
ner 1
997)
Pin
nula
ria s
p.P
innu
laria
sp.
Eas
tB
rune
l 196
2P
innu
laria
spp
.P
innu
laria
spp
.A
rctic
Rie
del e
t al.
2003
Pin
nula
ria s
ubro
stra
ta (
A.C
leve
) C
leve
-Eul
erP
innu
laria
sub
rost
rata
(A.C
leve
) C
leve
-Eul
er (
ab)
Pin
nula
ria d
iver
gent
issi
ma
var.
subr
ostra
ta A
. Cle
ve, P
innu
laria
di
verg
entis
sim
a v
ar. w
ulffi
i J.
B.
Pet
erse
n
Arc
ticA
nton
iade
s et
al.
2005
Pag
e 60
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Pin
nula
ria v
iridi
s K
tz.
Pin
nula
ria v
iridi
s (N
itzsc
h)
Ehr
enbe
rg
(ab)
Nav
icul
a vi
ridis
(N
itzsc
h)
Ehr
enbe
rg, F
rust
ulia
viri
dis
(Nitz
sch)
Küt
zing
, Bac
illar
ia v
iridi
s N
itzsc
h, N
avic
ula
virid
is (N
itzsc
h)
Küt
zing
, Sch
izon
ema
virid
e (N
itzsc
h) K
untz
e
Eas
tLi
nkle
tter 1
977
(Bai
ley
1913
)
Pla
giog
ram
ma
sp.
Pla
giog
ram
ma
sp.
Arc
ticB
ursa
196
3P
lagi
ogra
mm
opsi
s va
nheu
rcki
i (G
runo
w) H
asle
, von
Sto
sch,
&
Syv
erts
en
Pla
giog
ram
ma
vanh
eurc
kii
Gru
now
(a
b)E
ast/W
est
Hor
ner (
pers
. com
m.)
3 (Trit
es
2005
, McQ
uoid
& H
obso
n 20
01)
Pla
giot
ropi
s lo
nga
Pla
giot
ropi
s lo
nga
(Cle
ve) K
untz
e (a
b)E
ast
Bér
ard-
Ther
riaul
t et a
l. 19
99
Pla
giot
ropi
s va
nheu
rkii
(Gru
now
) C
leve
Trop
idon
eis
vanh
eurc
kii
(Gru
now
) C
leve
(ab)
Pla
giot
ropi
s va
nheu
rcki
i G
runo
wE
ast
Trite
s 20
05
Pla
nkto
niel
la s
ol (
Wal
lich)
Sch
ütt
Pla
nkto
niel
la s
ol (W
allic
h) S
chüt
t (a
b)C
osci
nodi
scus
sol
Wal
lich,
P
lank
toni
ella
wol
tere
cki
Sch
impe
rE
ast/W
est
(US
)C
upp
1943
(Car
ver &
Mal
let 2
002)
Pla
noth
idiu
m d
elic
atul
um (
Küt
zing
) R
ound
& B
ukht
iyar
ova
Pla
noth
idiu
m d
elic
atul
um (
Küt
zing
) R
ound
& L
. Buk
htiy
arov
a (a
b)M
icro
neis
del
icat
ula
(Küt
zing
) C
leve
, Ach
nant
hes
delic
atul
a (K
ützi
ng) G
runo
w, A
chna
nthe
s de
licat
ula
(Küt
zing
) Bru
n,
Ach
nant
hes
delic
atul
a (K
ützi
ng)
Gru
now
, Fal
cate
lla d
elic
atul
a (K
ützi
ng) R
aben
hors
t, A
chna
nthi
dium
del
icat
ulum
K
ützi
ng, A
chna
nthe
iops
is
delic
atul
a (K
ützi
ng) L
ange
-B
erta
lot
Wes
t (E
ffing
ham
In
let)
Hay
et a
l. 20
03
Pla
noth
idiu
m c
f. en
gelb
rech
tii
(Cho
lnok
y) R
ound
& B
ukht
iyar
ova
Pla
noth
idiu
m e
ngel
brec
htii
(Cho
lnok
y) R
ound
& B
ukht
iyar
ova
(ab)
Wes
t (E
ffing
ham
In
let)
Hay
et a
l. 20
03
Pag
e 61
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Pla
noth
idiu
m la
nceo
latu
m
(Bré
biss
on e
x K
ützi
ng) L
ange
-B
erta
lot
Pla
noth
idiu
m la
nceo
latu
m
(Bré
biss
on e
x K
ützi
ng) L
ange
-B
erta
lot
(ab)
Pla
noth
idiu
m la
nceo
latu
m
(Bré
biss
on e
x K
ützi
ng) F
.E.
Rou
nd &
L. B
ukht
iyar
ova,
M
icro
neis
lanc
eola
ta (B
rébi
sson
in
Küt
zing
) Fre
ngue
lli,
Ach
nant
hes
lanc
eola
ta
(Bré
biss
on e
x K
ützi
ng) G
runo
w,
Ach
nant
hes
lanc
eola
ta
(Bré
biss
on e
x K
ützi
ng) G
runo
w,
Ach
nant
hidi
um la
nceo
latu
m
Bré
biss
on e
x K
ützi
ng,
Ach
nant
heio
psis
lanc
eola
ta
(Bré
biss
on e
x K
ützi
ng) L
ange
-B
erta
lot
Wes
t (E
ffing
ham
In
let)
Hay
et a
l. 20
03
Ple
uros
igm
a ac
umin
atum
G
yros
igm
a ac
umin
atum
(K
ützi
ng)
Rab
enho
rst (
ab)
Ple
uros
igm
a ac
umin
atum
(K
ützi
ng) G
runo
w
Eas
tLi
nkle
tter e
t al.
1977
(Bai
ley
1913
)
Ple
uros
igm
a ae
stua
rii W
. Sm
ithP
leur
osig
ma
aest
uarii
(B
rébi
sson
ex
Küt
zing
) W. S
mith
(ab)
Nav
icul
a ae
stua
rii B
rébi
sson
ex
Küt
zing
Eas
tLi
nkle
tter e
t al.
1977
(Bai
ley
1910
)
Ple
uros
igm
a af
finis
Gru
n.P
leur
osig
ma
affin
is G
run.
(ab)
Eas
tB
aile
y 19
13P
leur
osig
ma
alle
nuat
umP
leur
osig
ma
alle
nuat
umE
ast
Link
lette
r et a
l. 19
77P
leur
osig
ma
angu
latu
m (Q
ueke
tt)
W. S
mith
Ple
uros
igm
a an
gula
tum
W. S
mith
(a
b)E
ast
Link
lette
r et a
l. 19
77 (B
aile
y 19
10,
Fritz
192
1, T
rites
200
5, B
ates
&
Stra
in 2
006)
Ple
uros
igm
a an
gula
tum
var
. st
rigos
a (W
. Sm
ith) V
an H
eurc
kP
leur
osig
ma
strig
osum
W. S
mith
(a
b)P
leur
osig
ma
angu
latu
m v
ar.
strig
osum
(W. S
mith
) Van
Heu
rck
Eas
tM
artin
et a
l. 20
01
Ple
uros
igm
a ar
cuat
um D
onki
nP
leur
osig
ma
arcu
atum
Don
kin
(ab)
Eas
tB
aile
y 19
13
Ple
uros
igm
a ba
lticu
mG
yros
igm
a ba
lticu
m (
Ehr
enbe
rg)
Rab
enho
rst (
ab)
Ple
uros
igm
a ba
lticu
m
(Ehr
enbe
rg) W
. Sm
ith
Eas
tFr
itz 1
921
(Bai
ley
1910
, Bai
ley
1913
)P
leur
osig
ma
deco
rum
Wm
. Sm
ithP
leur
osig
ma
deco
rum
Wm
. Sm
ith
(ab)
Eas
tLi
nkle
tter e
t al.
1977
Ple
uros
igm
a de
licat
ulum
W.S
.P
leur
osig
ma
delic
atul
um W
.Sm
ith
(ab)
Eas
tB
aile
y 19
13
Pag
e 62
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Ple
uros
igm
a el
onga
tum
W. S
mith
Ple
uros
igm
a el
onga
tum
W. S
mith
(a
b)P
leur
osig
ma
angu
latu
m v
ar.
elon
gatu
m (W
. Sm
ith) v
an
Heu
rck,
Gyr
osig
ma
elon
gatu
m
(W. S
mith
) Grif
fith
& H
enfre
y,
Gyr
osig
ma
elon
gatu
m (W
. Sm
ith)
G.S
. Wes
t
Eas
t (B
oF)/W
est
(US
)
Link
lette
r et a
l. 19
77 (C
upp
1943
, C
arve
r & M
alle
t 200
2)
Ple
uros
igm
a ex
imiu
m (
Thw
aite
s)
Cle
ve &
Gru
now
P
leur
osig
ma
exim
ium
(Th
wai
tes)
C
leve
& G
runo
w (
ab)
Eas
tP
atric
k &
Rei
mer
196
6
Ple
uros
igm
a fa
scio
laG
yros
igm
a fa
scio
la (
Ehr
enbe
rg)
J.W
. Grif
fith
& H
enfre
y (a
b)P
leur
osig
ma
fasc
iola
(Ehr
enbe
rg)
W. S
mith
Eas
tFr
itz 1
921
(Bai
ley
1910
, Bai
ley
1913
)P
leur
osig
ma
form
osum
W. S
mith
Ple
uros
igm
a fo
rmos
um W
. Sm
ith
(ab)
Ple
uros
igm
a au
stra
licum
Witt
, P
leur
osig
ma
tahi
tens
e W
itt
Eas
tLi
nkle
tter e
t al.
1977
(Bai
ley
1910
, B
aile
y 19
13, B
aile
y 19
10, F
ritz
1921
, Bru
nel 1
962)
Ple
uros
igm
a ha
mul
iferu
m B
run
and
Tem
père
Ple
uros
igm
a ha
mul
iferu
m B
run
and
Tem
p. (a
b)W
est (
US
)C
upp
1943
Ple
uros
igm
a in
term
ediu
m W
. Sm
ithP
leur
osig
ma
inte
rmed
ium
W. S
mith
(a
b)P
leur
osig
ma
nube
cula
var
. in
term
ediu
m (
W. S
mith
) Cle
ve,
Ple
uros
igm
a nu
becu
la W
. Sm
ith,
Ple
uros
igm
a in
term
ediu
m v
ar.
nube
cula
(W
. Sm
ith) G
runo
w e
x V
an H
eurc
k
Eas
t (B
oF)
Car
dina
l et a
l. 19
89
Ple
uros
igm
a m
acru
m W
.S.
Ple
uros
igm
a m
acru
m W
.Sm
ith (a
b)E
ast
Bai
ley
1913
Ple
uros
igm
a ni
coba
ricum
Gru
now
Ple
uros
igm
a ni
coba
ricum
Gru
n. (a
b)W
est (
US
)C
upp
1943
Ple
uros
igm
a no
rman
ii R
alfs
Ple
uros
igm
a no
rman
ii R
alfs
(ab)
Ple
uros
igm
a af
fine
Gru
now
E
ast/W
est
(US
)C
upp
1943
(Car
ver &
Mal
let 2
002)
Ple
uros
igm
a sp
.P
leur
osig
ma
sp.
Eas
t/Arc
ticB
ursa
196
3 (G
ran
& B
raar
ud
1935
)P
leur
osig
ma
W. S
mith
spp
.P
leur
osig
ma
W. S
mith
spp
.A
rctic
Love
joy
et a
l. 20
02P
leur
osig
ma
spen
ceri
Gru
n.P
leur
osig
ma
spen
ceri
(Que
k.) W
. S
mith
Eas
tB
aile
y 19
10
Ple
uros
igm
a st
rigili
s W
. Sm
ithP
leur
osig
ma
strig
ilis
Wm
. Sm
ith (a
b)E
ast (
BoF
)Li
nkle
tter e
t al.
1977
(Bai
ley
1910
, B
aile
y 19
13)
Ple
uros
igm
a st
rigos
um W
. Sm
ithP
leur
osig
ma
strig
osum
W. S
mith
(a
b)P
leur
osig
ma
angu
latu
m v
ar.
strig
osum
(W
.Sm
ith) V
an H
eurk
E
ast
Link
lette
r et a
l. 19
77 (B
aile
y 19
10,
Fritz
192
1, B
aile
y 19
13)
Ple
uros
igm
a st
uxbe
rgii
Ple
uros
igm
a st
uxbe
rgii
Cle
ve &
G
runo
w (a
b)A
rctic
Rie
del e
t al.
2003
(Boo
th &
Hor
ner
1997
)
Pag
e 63
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Por
osira
gla
cial
is J
orge
nsen
Por
osira
gla
cial
is (G
runo
w)
Jorg
ense
n (a
b)P
odos
ira h
orm
oide
s va
r. gl
acia
lis
Gru
now
, Pod
osira
gla
cial
is
(Gru
now
) Cle
ve, L
aude
ria
glac
ialis
(G
runo
w) G
ran,
Por
osira
an
tarc
tica
O.G
. Koz
lova
Eas
t/Arc
ticB
ursa
196
3 (L
ovej
oy e
t al.
2002
, G
ran
& B
raar
ud 1
935,
Boo
th &
H
orne
r 199
7)
Pro
bosc
ia a
lata
Pro
bosc
ia a
lata
(B
right
wel
l) S
ünds
trom
(ab)
Rhi
zoso
leni
a al
ata
Brig
htw
ell,
Rhi
zoso
leni
a al
ata
f. gr
acill
ima
(Cle
ve) G
ran,
Rhi
zoso
leni
a al
ata
f. G
enui
na G
ran
Eas
t/Wes
tB
érar
d-Th
erria
ult e
t al.
1999
(H
orne
r 200
2)
Pro
bosc
ia e
umor
pha
Pro
bosc
ia e
umor
pha
K. T
akah
ashi
, R
. Jor
dan
& J
. Prid
dle
(ab)
Rhi
zoso
leni
a ob
tusa
Hen
sen
Eas
tM
artin
& L
eGre
sley
200
8 (C
arve
r &
Mal
let 2
002)
Pse
udog
omph
onem
a ar
ctic
um
Pse
udog
omph
onem
a ar
ctic
um
(Gru
now
) L.K
. Med
lin (a
b)A
rctic
Rie
del e
t al.
2003
(Lov
ejoy
et a
l. 20
02)
Pse
udo-
nitz
schi
a am
eric
ana
(G.R
.Has
le) G
.A. F
ryxe
llP
seud
o-ni
tzsc
hia
amer
ican
a (G
.R.
Has
le) G
.A. F
ryxe
ll (a
b)N
itzsc
hia
amer
ican
a G
.R.H
asle
E
ast/W
est
Mar
tin (p
ers.
com
m.)2 (H
orne
r pe
rs. c
omm
.)3
Pse
udo-
nitz
schi
a au
stra
lis
Fren
guel
liP
seud
o-ni
tzsc
hia
aust
ralis
Fr
engu
elli
(ab)
Nitz
schi
a ps
eudo
seria
ta H
asle
W
est
Hor
ner (
pers
. com
m.)
3
Pse
udo-
nitz
schi
a de
licat
issi
ma
(Cle
ve) H
eide
nP
seud
o-ni
tzsc
hia
delic
atis
sim
a (C
leve
) Hei
den
(ab)
Nitz
schi
a de
licat
issi
ma
Cle
ve,
Nitz
schi
a ac
tydr
ophi
la H
asle
Eas
t/Wes
t/Ar
ctic
Har
grav
es &
Mar
anda
200
2 (C
hang
et a
l. 20
07, B
érar
d-Th
erria
ult e
t al.
1999
, Lov
ejoy
et
al. 2
002)
Pse
udo-
nitz
schi
a fra
udul
enta
(C
leve
) Has
leP
seud
o-ni
tzsc
hia
fraud
ulen
ta
(Cle
ve) G
.R. H
asle
(ab)
Nitz
schi
a fra
udul
enta
Cle
ve,
Pse
udo-
nitz
schi
a se
riata
var
. fra
udul
enta
(Cle
ve) H
. Per
agal
lo
Eas
t/Wes
tH
argr
aves
& M
aran
da 2
002
(Mar
tin &
LeG
resl
ey 2
008)
Pse
udo-
nitz
schi
a gr
anii
(Has
le)
Has
le &
Syv
erts
enP
seud
o-ni
tzsc
hia
gran
ii (G
.R. H
asle
) G
.R. H
asle
(ab)
Nitz
schi
a gr
anii
G.R
. Has
le
Arc
ticLo
vejo
y et
al.
2002
Pse
udo-
nitz
schi
a m
ultis
erie
s (H
asle
) Has
leP
seud
o-ni
tzsc
hia
mul
tiser
ies
(Has
le)
Has
le (
ab)
Nitz
schi
a pu
ngen
s f.
mul
tiser
ies
G.R
. Has
le, P
seud
o-ni
tzsc
hia
pung
ens
f. m
ultis
erie
s H
asle
Eas
t/Wes
tH
argr
aves
& M
aran
da 2
002
(Mar
tin [p
ers.
com
m.]2 , B
érar
d-Th
erria
ult e
t al.
1999
)
Pse
udo-
nitz
schi
a ps
eudo
delic
atis
sim
a (H
asle
) Has
leP
seud
o-ni
tzsc
hia
pseu
dode
licat
issi
ma
(Has
le) H
asle
(a
b)
Nitz
schi
a de
licat
ula
Has
le,
Nitz
schi
a ps
eudo
delic
atis
sim
a G
.R.
Has
le
Eas
t/Wes
t/Ar
ctic
Hor
ner e
t al.
2005
(Mar
tin [p
ers.
co
mm
.]2 , Har
grav
es &
Mar
anda
20
02, B
érar
d-Th
erria
ult e
t al.
1999
, Boo
th &
Hor
ner 1
997)
Pag
e 64
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Pse
udo-
nitz
schi
a pu
ngen
s (G
runo
w) H
asle
Pse
udo-
nitz
schi
a pu
ngen
s (G
runo
w
ex C
leve
) Has
le (
ab)
Nitz
schi
a pu
ngen
s G
runo
w e
x C
leve
Eas
t/Wes
tH
argr
aves
& M
aran
da 2
002
(Mar
tin [p
ers.
com
m.]2 , B
érar
d-Th
erria
ult e
t al.
1999
)P
seud
o-ni
tzsc
hia
seria
ta (
Cle
ve)
H.P
erag
allo
Pse
udo-
nitz
schi
a se
riata
(Cle
ve)
H.P
erag
allo
(ab)
Nitz
schi
a se
riata
Cle
ve
Eas
t/Wes
tH
obso
n (p
ers.
com
m.)
5 (Cha
ng e
t al
. 200
7, K
aczm
arsk
a et
al.
2007
, B
érar
d-Th
erria
ult e
t al.
1999
)
Pse
udo-
nitz
schi
a se
riata
f. o
btus
aP
seud
o-ni
tzsc
hia
obtu
sa (
Has
le)
Has
le &
Lun
dhol
m (a
b)P
seud
o-ni
tzsc
hia
seria
ta f
. ob
tusa
(H
asle
) Has
leE
ast
Bér
ard-
Ther
riaul
t et a
l. 19
99
Pse
udo-
nitz
schi
a sp
p.P
seud
o-ni
tzsc
hia
spp.
Arc
ticLo
vejo
y et
al.
2002
Pse
udo-
nitz
schi
a su
bpac
ifica
P
seud
o-ni
tzsc
hia
subp
acifi
ca (G
.R.
Has
le) G
.R. H
asle
(ab)
Nitz
schi
a su
bpac
ifica
G.R
. Has
leE
ast/W
est
Mar
tin &
LeG
resl
ey 2
008
(Hor
ner
[per
s. c
omm
.]3 )P
unct
astri
ata
sp.
Pun
ctas
triat
a sp
.E
ast
Bér
ard-
Ther
riaul
t et a
l. 19
99R
apho
neis
sp.
Rha
phon
eis
sp.
Wes
tM
cQuo
id &
Hob
son
2001
Rha
phon
eis
amph
icer
os E
hren
berg
Rha
phon
eis
amph
icer
os
(Ehr
enbe
rg) E
hren
berg
(ab
)R
haph
onei
s la
nceo
lata
E
hren
berg
, Dor
ypho
ra
amph
icer
os (E
hren
berg
) Küt
zing
, C
occo
neis
am
phic
eros
E
hren
berg
, Rha
phon
eis
rhom
bus
Ehr
enbe
rg, R
haph
onei
s am
phic
eros
var
. rho
mbi
ca
Gru
now
ex
Van
Heu
rck
Eas
t/Wes
tLi
nkle
tter e
t al.
1977
(Bai
ley
1913
, B
aile
y 19
10)
Rha
phon
eis
amph
icer
os v
. el
onga
ta P
erag
. & P
erag
.R
haph
onei
s am
phic
eros
var
. el
onga
ta P
erag
. & P
erag
(ab)
Eas
tR
awle
nce
1987
Rha
phon
eis
mar
garit
alim
bata
M
ertz
Rha
phon
eis
mar
garit
alim
bata
Mer
tz
(ab)
Wes
t (E
ffing
ham
In
let)
Hay
et a
l. 20
03
Rha
bdon
ema
adria
ticum
Küz
ting
Rha
bdon
ema
adria
ticum
Küz
ting
(ab)
Eas
tB
aile
y 19
13
Rha
bdon
ema
arcu
atum
(Ly
ngby
e)
Küt
zing
Rha
bdon
ema
arcu
atum
(Lyn
gbye
) K
ützi
ng (a
b)D
iato
ma
arcu
atum
Lyn
gbye
Eas
t/Arc
ticLi
nkle
tter e
t al.
1977
(Bai
ley
1913
, B
aile
y 19
10, B
rune
l 196
2, B
ursa
19
63)
Rha
bdon
ema
spp.
Rha
bdon
ema
spp.
Eas
t/Wes
tM
artin
(per
s. c
omm
.)2 (McQ
uoid
&
Hob
son
2001
)R
hizo
sole
nia
acum
inat
a (H
. P
érag
ello
) Gra
nR
hizo
sole
nia
acum
inat
a (H
. P
érag
ello
) Gra
n (a
b)R
hizo
sole
nia
tem
pere
i va
r. ac
umin
ata
H. P
erag
allo
W
est (
US
)C
upp
1943
Pag
e 65
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Rhi
zoso
leni
a al
ata
Brig
htw
ell
Pro
bosc
ia a
lata
(Brig
htw
ell)
Sün
dstro
m (a
b)R
hizo
sole
nia
alat
a B
right
wel
lE
ast/A
rctic
Bur
sa 1
963
(Gra
n &
Bra
arud
19
35)
Rhi
zoso
leni
a al
ata
form
cur
viro
stris
G
ran
Rhi
zoso
leni
a al
ata
form
cur
viro
stris
G
ran
(ab)
Wes
t (U
S)
Cup
p 19
43
Rhi
zoso
leni
a al
ata
form
gra
cilli
ma
(Cle
ve) G
run
Pro
bosc
ia a
lata
(Brig
htw
ell)
Sün
dstro
m (a
b)R
hizo
sole
nia
alat
a va
r. gr
acill
ima
(Cle
ve) G
runo
w e
x V
an H
eurc
k,
Rhi
zoso
leni
a gr
acill
ima
Cle
ve
Eas
t (B
oF)/W
est
(US
)
Cup
p 19
43
Rhi
zoso
leni
a al
ata
form
indi
ca (
H.
Pér
agal
lo) G
ran
Rhi
zoso
leni
a al
ata
form
indi
ca (
H.
Pér
agal
lo) G
ran
(ab)
Rhi
zoso
leni
a in
dica
H. P
erag
allo
Wes
t (U
S)
Cup
p 19
43
Rhi
zoso
leni
a al
ata
form
iner
mis
(C
astra
cane
) Hus
tedt
Rhi
zoso
leni
a al
ata
form
iner
mis
(C
astr.
) Hus
t. (a
b)W
est (
US
)C
upp
1943
Rhi
zoso
leni
a be
rgon
ii H
. Pér
agal
lo
Rhi
zoso
leni
a be
rgon
ii H
. Pér
agal
lo
(ab)
Rhi
zoso
leni
a am
puta
ta O
sten
feld
Wes
t (U
S)
Cup
p 19
43
Rhi
zoso
leni
a ca
lcar
avi
s M
. S
chul
tze
Pse
udos
olen
ia c
alca
r-av
is
(Sch
ultz
e) S
unds
tröm
(ab)
Rhi
zoso
leni
a ca
lcar
avi
s M
. S
chul
tze
Wes
t (U
S)
Cup
p 19
43
Rhi
zoso
leni
a ca
stra
cane
i H
. P
érag
allo
Gui
nard
ia fl
acci
da (
Cas
traca
ne) H
. P
erag
allo
Rhi
zoso
leni
a ca
stra
cane
i C
leve
Wes
t (U
S)
Cup
p 19
43
Rhi
zoso
leni
a de
licat
ula
Gui
nard
ia d
elic
atul
a (C
leve
) Has
leR
hizo
sole
nia
delic
atul
a C
leve
Arc
ticB
ooth
& H
orne
r 199
7
Rhi
zoso
leni
a fa
eroe
nsis
Ost
enfe
ldR
hizo
sole
nia
faer
oens
is O
sten
feld
(a
b)E
ast (
BoF
)Li
nkle
tter e
t al.
1977
(Frit
z 19
21,
Bru
nel 1
962)
Rhi
zoso
leni
a fra
gilis
sim
aD
acty
lioso
len
fragi
lissi
mus
(B
ergo
n)
G.R
. Has
leR
hizo
sole
nia
fragi
lissi
ma
Ber
gon
Eas
t/Wes
tB
rune
l 196
2 (G
ran
& B
raar
ud
1935
, Hor
ner 2
002)
Rhi
zoso
leni
a fra
gilis
sim
a va
r. be
rgon
iiR
hizo
sole
nia
fragi
lissi
ma
var.
berg
onii
A. C
leve
-Eul
erE
ast
Bru
nel 1
962
Rhi
zoso
leni
a fra
gilis
sim
a va
r. fa
eroe
nsis
Rhi
zoso
leni
a fra
gilis
sim
a va
r. fa
eroe
nsis
(C
.E.H
. Ost
enfe
ld) A
. C
leve
-Eul
er
Eas
tB
rune
l 196
2
Rhi
zoso
leni
a he
beta
ta B
aile
yR
hizo
sole
nia
hebe
tata
J.W
. Bai
ley
(ab)
Rhi
zoso
leni
a he
bata
ta f
. hie
mal
is
Gra
nE
ast/W
est/A
rct
icLi
nkle
tter e
t al.
1977
(Frit
z 19
21,
Bru
nel 1
962,
Bur
sa 1
963,
Gra
n &
B
raar
ud 1
935,
Bat
es &
Stra
in
2006
)R
hizo
sole
nia
hebe
tata
f. h
iem
alis
Rhi
zoso
leni
a he
beta
ta J
.W. B
aile
yR
hizo
sole
nia
hebe
tata
f. h
iem
alis
G
ran
Eas
tB
rune
l 196
2
Pag
e 66
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Rhi
zoso
leni
a he
beta
ta f
. sem
ispi
naR
hizo
sole
nia
hebe
tata
f. s
emis
pina
(H
ense
n) G
ran
(ab)
Rhi
zoso
leni
a se
mis
pina
Hen
sen,
R
hizo
sole
nia
styl
iform
is v
ar.
sem
ispi
na (H
ense
n) G
. Kar
sten
Eas
t/Wes
tH
obso
n (p
ers.
com
m.)
5 (Bér
ard-
Ther
riaul
t et a
l. 19
99, B
rune
l 196
2,
Bat
es &
Stra
in 2
006)
Rhi
zoso
leni
a im
bric
ata
Rhi
zoso
leni
a im
bric
ata
Brig
htw
ell
(ab)
Rhi
zoso
leni
a sh
rubs
olei
Cle
ve,
Rhi
zoso
leni
a im
bric
ata
var.
stria
ta (
Gre
ville
) Gru
now
, R
hizo
sole
nia
imbr
icat
a va
r. sh
rubs
olei
(C
leve
) Sch
röde
r
Eas
tB
érar
d-Th
erria
ult e
t al.
1999
(Gra
n &
Bra
arud
193
5, B
ates
& S
train
20
06)
Rhi
zoso
leni
a ob
tusa
Pro
bosc
ia e
umor
pha
K. T
akah
ashi
, R
. Jor
dan
& J
. Prid
dle
(ab)
Rhi
zoso
leni
a ob
tusa
Hen
sen
Eas
tFr
itz 1
921
Rhi
zoso
leni
a pu
ngen
sR
hizo
sole
nia
setig
era
f. pu
ngen
s (C
leve
-Eul
er) B
rune
l (ab
)R
hizo
sole
nia
pung
ens
Cle
ve-
Eul
erE
ast
Bér
ard-
Ther
riaul
t et a
l. 19
99
(Bru
nel 1
962,
Bat
es &
Stra
in
2006
)R
hizo
sole
nia
robu
sta
Nor
man
N
eoca
lypt
rella
robu
sta
(G. N
orm
an e
x R
alfs
) D.U
. H
erná
ndez
-Bec
erril
& M
. E. M
eave
del
Cas
tillo
(ab)
Rhi
zoso
leni
a ro
bust
a G
. Nor
man
ex
Ral
fs
Eas
t/Wes
t (U
S)
Cup
p 19
43
Rhi
zoso
leni
a se
tiger
a B
right
wel
lR
hizo
sole
nia
setig
era
Brig
htw
ell
(ab)
Rhi
zoso
leni
a ja
poni
ca
Cas
traca
ne, R
hizo
sole
nia
hens
enii
Sch
ütt
Eas
t/Wes
tH
orne
r et a
l. 20
05 (B
aile
y 19
13,
Bai
ley
1910
, Lin
klet
ter e
t al.
1977
, S
ance
tta &
Cal
vert
1988
, Bru
nel
1962
, Gra
n &
Bra
arud
193
5, B
ates
&
Stra
in 2
006,
Hay
et a
l. 20
03)
Rhi
zoso
leni
a se
tiger
a f.
pung
ens
Rhi
zoso
leni
a se
tiger
a f.
pung
ens
(C
leve
-Eul
er) B
rune
l (ab
)R
hizo
sole
nia
pung
ens
Cle
ve-
Eul
er
Eas
tB
rune
l 196
2
Rhi
zoso
leni
a sh
rubs
olei
Rhi
zoso
leni
a im
bric
ata
Brig
htw
ell
(ab)
Rhi
zoso
leni
a sh
rubs
olei
Cle
veE
ast
Fritz
192
1
Rhi
zoso
leni
a st
ylifo
rmis
Rhi
zoso
leni
a st
ylifo
rmis
Brig
htw
ell
(ab)
Rhi
zoso
leni
a st
ylifo
rmis
var
. po
lyda
ctyl
a va
n H
eurc
k,
Rhi
zoso
leni
a st
ylifo
rmis
var
. lo
ngis
pina
Hus
tedt
Eas
t/Arc
ticB
ursa
196
3 (G
ran
& B
raar
ud
1935
, Car
ver &
Mal
let 2
002)
Rhi
zoso
leni
a sp
.R
hizo
sole
nia
sp.
Wes
tM
cQuo
id &
Hob
son
1997
Rhi
zoso
leni
a sp
p.R
hizo
sole
nia
spp.
Wes
tS
ance
tta &
Cal
vert
1988
(McQ
uoid
&
Hob
son
2001
)R
hoic
osph
enia
cur
vata
(K
utzi
ng)
Gru
now
ex
Rab
h.R
hoic
osph
enia
abb
revi
ata
(C.
Aga
rdh)
Lan
ge-B
erta
lot
Rho
icos
phen
ia c
urva
ta (
Küt
zing
) G
runo
wW
est
McQ
uoid
& H
obso
n 20
01 (B
aile
y 19
13)
Pag
e 67
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Rho
palo
dia
gibb
a (E
hren
berg
) M
ülle
rR
hopa
lodi
a gi
bba
(Ehr
enbe
rg) O
.F.
Mül
ler
(ab)
Eun
otia
gib
ba (E
hren
berg
) E
hren
berg
, Nav
icul
a gi
bba
Ehr
enbe
rg, C
ysto
pleu
ra g
ibba
(E
hren
berg
) Kun
tze,
Epi
them
ia
vent
ricos
a v
ar. g
ibba
(Ehr
enbe
rg;
Ehr
enbe
rg) J
. Sch
uman
n,
Frus
tulia
gib
ba (E
hren
berg
) Je
nner
, Epi
them
ia g
ibba
(E
hren
berg
) Küt
zing
, Pin
nula
ria
gibb
a (E
hren
berg
) Ehr
enbe
rg,
Sch
izon
ema
gibb
um (E
hren
berg
) K
untz
e
Eas
tLi
nkle
tter e
t al.
1977
Rho
palo
dia
mus
culu
s R
hopa
lodi
a m
uscu
lus
(Küt
zing
) O.F
. M
ülle
r (ab
)R
hopa
lodi
a gi
bber
ula
var.
mus
culu
s (K
ützi
ng) M
usch
ler,
Epi
them
ia m
uscu
lus
Kut
zing
, C
ysto
pleu
ra m
uscu
lus
(Küt
zing
) K
untz
e, E
unot
ia w
este
rman
ni v
ar.
mus
culu
s (K
ützi
ng) G
.L.
Rab
enho
rst,
Rho
palo
dia
gibb
erul
a v
ar. m
uscu
lus
(Küt
zing
) C
leve
-Eul
er
Eas
tLi
nkle
tter e
t al.
1977
Rho
palo
dia
spp.
Rho
palo
dia
spp.
Wes
tM
cQuo
id &
Hob
son
2001
Rho
palo
dia
turg
ida
Rho
palo
dia
turg
ida
Eas
t (B
oF)
Link
lette
r et a
l. 19
77R
hopa
lodi
a ze
bra
Rho
palo
dia
zebr
aE
ast
Link
lette
r et a
l. 19
77S
chiz
onem
a cr
ucig
erum
W. S
m.
Has
lea
cruc
iger
a (W
. Sm
ith)
Sim
onse
n (a
b)
Sch
izon
ema
cruc
iger
a W
. Sm
.E
ast
Bai
ley
1910
Sch
röde
rella
del
icat
ula
Pav
illar
dD
eton
ula
pum
ila (C
astra
cane
) Gra
n (a
b)D
eton
ula
schr
oede
ri G
ran,
La
uder
ia s
chro
eder
i B
ergo
n,
Det
onul
a de
licat
ula
(H. P
erag
allo
) G
ran,
Lau
deria
del
icat
ula
H.
Per
agal
lo, S
chrö
dere
lla
schr
oede
ri (B
ergo
n) P
avill
ard
Wes
t C
upp
1943
(Hor
ner 2
002)
Sco
liopl
eura
tum
ida
(deB
rébi
sson
) R
aben
hors
tS
colio
pleu
ra tu
mid
a (B
rébi
sson
ex
Kut
zing
) Rab
enho
rst (
ab)
Eas
t (B
oF)
Link
lette
r et a
l. 19
77
Pag
e 68
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Sel
laph
ora
pupu
la (
Küt
z.)
Mer
esch
kovs
kyS
ella
phor
a pu
pula
(Küt
zing
) M
eres
chko
vsky
(ab
)N
avic
ula
pupu
la K
ützi
ng,
Sch
izon
ema
pupu
la (K
ützi
ng)
Kun
tze
Eas
t (B
oF)
Gin
n (p
ers.
com
m.)
1
Ske
leto
nem
a co
stat
um
Ske
leto
nem
a co
stat
um (G
revi
lle)
Cle
ve (
ab)
Mel
osira
cos
tata
Gre
ville
Eas
t/Wes
t/Ar
ctic
Bai
ley
1913
(San
cetta
& C
alve
rt 19
88, B
érar
d-Th
erria
ult e
t al.
1999
, Bru
nel 1
962,
Lov
ejoy
et a
l. 20
02, B
ates
& S
train
200
6,
McQ
uoid
& H
obso
n 19
97,
McQ
uoid
& H
obso
n 20
01)
Ske
leto
nem
a su
bsal
sum
Ske
leto
nem
a su
bsal
sum
(A
.Cle
ve)
Bet
hge
(ab)
Eas
tB
rune
l 196
2
Sta
uron
eis
acut
a W
.S.
Sta
uron
eis
koch
ii P
anto
csek
(col
)S
taur
onei
s ac
uta
W. S
mith
E
ast
Bai
ley
1913
Sta
uron
eis
ance
ps E
hr.
Sta
uron
eis
ance
ps E
hren
berg
(ab
)S
chiz
onem
a an
ceps
(E
hren
berg
) K
untz
e, S
taur
onei
s an
ceps
var
. am
phic
epha
la (K
ützi
ng) C
leve
, S
taur
onei
s an
ceps
var
. am
phic
epha
la (K
ützi
ng) v
an
Heu
rck,
Sta
uron
eis
amph
icep
hala
K
ützi
ng, S
taur
onei
s an
ceps
f.
grac
ilis
(Ehr
enbe
rg) C
leve
, N
avic
ula
ance
ps (E
hren
berg
) M
ann
Eas
tB
aile
y 19
13 (B
aile
y 19
10)
Sta
uron
eis
obliq
uaS
taur
onei
s ob
liqua
Gre
gory
(ab)
Eas
tLi
nkle
tter e
t al.
1977
(Bai
ley
1910
)
Sta
uron
eis
phoe
nice
nter
on
(Nitz
sch)
Ehr
enbe
rgS
taur
onei
s ph
oeni
cent
eron
(Nitz
sch)
E
hren
berg
(ab
)S
taur
onei
s ph
oeni
cent
eron
var
. ge
nuin
a P
.T. C
leve
, Nav
icul
a ph
oeni
cent
eron
(Nitz
sch)
E
hren
berg
, Cym
bella
ph
oeni
cent
eron
(N
itsch
) C.
Aga
rdh,
Bac
illar
ia p
hoen
icen
tero
n N
itzsc
h, S
taur
onei
s no
bilis
f.
alab
amae
(Hei
den)
A. C
leve
, S
chiz
onem
a ph
oeni
cent
erum
(E
hren
berg
) Kun
tze
Eas
t/Arc
ticA
nton
iade
s et
al.
2005
(Bai
ley
1913
, Bai
ley
1910
)
Pag
e 69
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Sta
uron
eis
smith
ii G
runo
wS
taur
onei
s sm
ithii
Gru
now
(ab
)S
taur
onei
s lin
earis
Ehr
enbe
rg
sens
u W
. Sm
ith, P
leur
osta
urum
sm
ithii
(Gru
now
) Gru
now
Eas
t (B
oF)
Raw
lenc
e 19
87
Sta
uron
eis
spic
ula
Has
lea
spic
ula
(Hic
kie)
L. B
ukht
iyar
ova
(ab)
Sta
uron
eis
spic
ula
Hic
kie
Eas
tB
aile
y 19
10
Sta
uros
ira c
onst
ruen
s E
hren
berg
Sta
uros
ira c
onst
ruen
s E
hren
berg
(a
b)Fr
agila
ria c
onst
ruen
s (E
hren
berg
) G
runo
w, S
taur
osira
ven
ter
var.
cons
truen
s (E
hren
berg
) Cle
ve &
M
ölle
r, N
emat
opla
ta c
onst
ruen
s (E
hren
berg
) Kun
tze
Eas
tG
inn
(per
s. c
omm
.)1
Sta
urop
sis
sept
entri
onal
isN
avic
ula
sept
entri
onal
is (
Øst
rup)
C
leve
(ab)
Sta
urop
sis
sept
entri
onal
is
Meu
nier
Eas
tB
rune
l 196
2
Sta
uros
irella
sp.
Sta
uros
irella
sp.
Wes
tM
cQuo
id &
Hob
son
2001
Ste
llarim
a st
ella
ris (
Rop
er) H
asle
S
tella
rima
stel
laris
(Rop
er) G
.R.
Has
le &
P.A
. Sim
s (a
b)C
osci
nodi
scus
ste
llaris
Rop
erE
ast
(BoF
)/Wes
tH
orne
r (pe
rs. c
omm
.)3
(San
cetta
&
Cal
vert
1988
)S
teph
anop
yxis
app
endi
cula
ta
Ehr
enbe
rgS
teph
anop
yxis
app
endi
cula
ta (C
.G.
Ehr
enbe
rg) C
.G. E
hren
berg
(ab
)E
ast (
BoF
)Li
nkle
tter e
t al.
1977
Ste
phan
opyx
is n
ippo
nica
Gra
n &
Y
endo
Ste
phan
opyx
is n
ippo
nica
Gra
n &
Y
endo
(ab)
Eas
t/Wes
tH
orne
r et a
l. 20
05 (C
arve
r & M
alle
t 20
02)
Ste
phan
opyx
is tu
rris
(G
revi
lle e
t A
rnot
t) R
alfs
Ste
phan
opyx
is tu
rris
(G
revi
lle) R
alfs
ex
Prit
char
d (a
b)C
resw
ellia
turr
is G
revi
lle &
Arn
ott
Eas
t/Wes
tS
ance
tta &
Cal
vert
1988
(McQ
uoid
&
Hob
son
2001
, Car
ver &
Mal
let
2002
)S
trept
othe
ca th
amen
sis
Shr
ubso
leH
elic
othe
ca ta
mes
is (
Shr
ubso
le)
Ric
ard
(ab)
Wes
t (U
S)
Cup
p 19
43
Stri
atel
la d
elic
atul
a (K
ützi
ng)
Gru
now
S
triat
ella
del
icat
ula
(Küt
zing
) G
runo
w e
x V
an H
eurc
k (a
b)W
est (
US
)C
upp
1943
Stri
atel
la u
nipu
ncta
ta (
Lyng
bye)
A
gard
hS
triat
ella
uni
punc
tata
(Lyn
gbye
) C.
Aga
rdh
(ab)
Frag
ilaria
uni
punc
tata
Lyn
gbye
Eas
t/Wes
tLi
nkle
tter e
t al.
1977
(Bai
ley
1913
, B
aile
y 19
10, B
rune
l 196
2, B
ates
&
Stra
in 2
006)
Sur
irella
bis
eria
ta W
.Sm
.S
urire
lla b
iser
iata
Bré
biss
on (a
b)S
urira
ya b
iser
iata
(B
rébi
sson
) P
fitze
rE
ast
Bai
ley
1910
Sur
irella
con
stric
ta
Sur
irella
con
stric
ta W
. Sm
ith (
ab)
Eas
tB
aile
y 19
10
Sur
irella
cra
ticul
a E
hr.
Sur
irella
cra
ticul
a (C
.G. E
hren
berg
) C
.G. E
hren
berg
Eas
tB
aile
y 19
13
Pag
e 70
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Sur
irella
cru
ciat
a A
.S.
Sur
irella
cru
ciat
a S
chm
idt (
ab)
Eas
tB
aile
y 19
13S
urire
lla c
rum
ena
Bre
b.S
urire
lla c
rum
ena
de
Bré
biss
on e
x K
ützi
ng (a
b)E
ast
Bai
ley
1913
Sur
irella
ele
gans
Ehr
.S
urire
lla e
lega
ns E
hren
berg
(ab)
Eas
tB
aile
y 19
13S
urire
lla fa
stuo
sa v
ar. r
eced
ens
(A. S
chm
idt)
Cle
veS
urire
lla fa
stuo
sa v
ar. r
eced
ens
(A.
Sch
m.)
Cl.
(ab)
Wes
t (U
S)
Cup
p 19
43
Sur
irella
febi
gerii
Lew
isS
urire
lla fe
bige
rii L
ewis
(ab)
Eas
tB
aile
y 19
13S
urire
lla g
emm
a (E
hren
berg
) K
ützi
ngS
urire
lla g
emm
a (E
hren
berg
) K
ützi
ng (
col)
Nav
icul
a ge
mm
a E
hren
berg
E
ast
Link
lette
r et a
l. 19
77 (B
aile
y 19
13)
Sur
irella
min
uta
Bre
b.S
urire
lla m
inut
a B
rébi
sson
(ab)
Eas
tB
aile
y 19
13 (B
aile
y 19
10)
Sur
irella
mol
eria
na G
runo
w
Sur
irella
mol
leria
na G
runo
w (
ab)
Eas
tLi
nkle
tter e
t al.
1977
(Bai
ley
1913
, B
aile
y 19
10)
Sur
irella
ova
lis d
eBré
biss
onS
urire
lla o
valis
Bré
biss
on (a
b)E
ast
Link
lette
r et a
l. 19
77 (B
aile
y 19
13,
Bai
ley
1910
)S
urire
lla p
ulch
ella
Sur
irella
pul
chel
la L
.A. B
rébi
sson
(a
b)E
ast (
BoF
)Li
nkle
tter e
t al.
1977
Sur
irella
sm
ithii
Ral
fsS
urire
lla s
mith
ii R
alfs
(ab)
Eas
t (B
oF)
Pou
lin e
t al.
1987
Sur
irella
spl
endi
da K
ütz.
Sur
irella
spl
endi
da K
ützi
ng (a
b)E
ast
Bai
ley
1913
(Bai
ley
1910
)S
urire
lla s
pp.
Sur
irella
spp
.E
ast
Thal
er 2
007
Sur
irella
stri
atul
a Tu
rp.
Sur
irella
stri
atul
a P
.J. T
urpi
n (a
b)N
avic
ula
stria
tula
(Tu
rpin
) E
hren
berg
, N
ovill
a st
riatu
la
(Tur
pin)
Hei
berg
, Sur
iraya
st
riatu
la (T
urpi
n) P
fitze
r
Eas
tB
aile
y 19
13 (B
aile
y 19
10)
Syn
dend
rium
dia
dem
aC
haet
ocer
os d
iade
ma
(Ehr
enbe
rg)
Gra
n (a
b)S
ynde
ndriu
m d
iade
ma
Ehr
enbe
rgE
ast
Bru
nel 1
962
Syn
edra
acu
s K
tz.
Uln
aria
acu
s (K
ützi
ng) M
. Abo
al (a
b)S
yned
ra a
cus
Küt
zing
E
ast
Bai
ley
1913
(Bai
ley
1910
)
Syn
edra
acu
s va
r. ra
dian
s (K
ütz)
H
uste
dtU
lnar
ia d
elic
atis
sim
a va
r. an
gust
issi
ma
(Gru
now
) M. A
boal
& P
.C. S
ilva
(ab)
Syn
edra
acu
s va
r. ra
dian
s (K
ützi
ng) H
uste
dtE
ast
Bai
ley
1910
Syn
edra
fasc
icul
ata
(Aga
rdh)
K
ützi
ngS
yned
ra fa
scic
ulat
a (A
gard
h)
Küt
zing
Eas
tR
awle
nce
1987
Syn
edra
gai
llion
ii (B
ory)
Ehr
enbe
rgC
atac
ombu
s ga
illon
ii (B
ory)
D.M
.Will
iam
s &
Rou
nd (a
b)S
yned
ra g
aillo
nii
(Bor
y de
Sai
nt-
Vin
cent
) Ehr
enbe
rg
Eas
t (B
oF)
Link
lette
r et a
l. 19
77
Syn
edra
luna
ris E
hr.
Syn
edra
luna
ris E
hren
berg
(ab)
Eas
tLi
nkle
tter e
t al.
1977
(Bai
ley
1913
)
Pag
e 71
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Syn
edra
radi
ans
Küt
zing
Syn
edra
radi
ans
Küt
zing
(ab)
Eas
tLi
nkle
tter e
t al.
1977
(Bai
ley
1910
, B
aile
y 19
13)
Syn
edra
sal
ina
Syn
edra
uln
a va
r. sa
lina
(W. S
mith
) W
. Sm
ithS
yned
ra s
alin
a W
. Sm
ithE
ast
Bai
ley
1910
(Bai
ley
1913
)
Syn
edra
sp.
Syn
edra
sp.
Arc
ticB
ursa
196
3S
yned
ra u
lna
(Nitz
sch)
Ehr
enbe
rgU
lnar
ia u
lna
(Nitz
sch)
P. C
ompè
re
(ab)
Syn
edra
uln
a(N
itzsc
h) E
hren
berg
(a
b)E
ast
Link
lette
r et a
l. 19
77 (B
rune
l 196
2,
Bai
ley
1910
, Bai
ley
1913
)
Syn
edra
uln
a va
r. co
ntra
ta O
str.
Syn
edra
uln
a va
r. co
ntra
cta
Øst
rup
Eas
tZw
icke
r 199
8
Syn
edro
psis
hyp
erbo
rea
Syn
edro
psis
hyp
erbo
rea
(Gru
now
) H
asle
, Med
lin &
Syv
erts
en (a
b)S
yned
ra h
yper
bore
a G
runo
w,
Syn
edra
hyp
erbo
rea
var.
rost
ella
ta G
runo
w, S
yned
ra
hype
rbor
ea v
ar. f
lexu
osa
Gru
now
Arc
ticR
iede
l et a
l. 20
03 (L
ovej
oy e
t al.
2002
, Boo
th &
Hor
ner 1
997)
Tabe
llaria
fene
stra
ta K
.B.
Tabe
llaria
fene
stra
ta (L
yngb
ye)
Küt
zing
(ab
)E
ast
Bai
ley
1913
(Bai
ley
1910
)
Tabe
llaria
floc
culo
sa K
utz.
Tabe
llaria
floc
culo
sa (R
oth)
Küt
zing
(a
b)C
onfe
rva
flocc
ulos
a R
oth,
S
triat
ella
floc
culo
sa (
Rot
h)
Kun
tze,
Bac
illar
ia fl
occu
losa
(R
oth)
Lei
blei
n
Eas
t/Wes
t (E
ffing
ham
In
let)
Hay
et a
l. 20
03 (B
aile
y 19
13,
Cat
tane
o et
al.
2004
)
Tabe
llaria
spp
.Ta
bella
ria s
pp.
Eas
t/Wes
tM
artin
(per
s. c
omm
.)2 (McQ
uoid
&
Hob
son
2001
)
Pag
e 72
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Tabu
laria
fasc
icul
ata
(Aga
rdh)
W
illia
ms
& R
ound
Tabu
laria
fasc
icul
ata
(C. A
gard
h)
D.M
. Will
iam
s &
Rou
nd (
ab)
Frag
ilaria
tabu
lata
(C
.Aga
rdh)
La
nge-
Ber
talo
t, Fr
agila
ria
fasc
icul
ata
(C. A
gard
h) L
ange
-B
erta
lot,
Syn
edra
tabu
lata
var
. fa
scic
ulat
a (C
. Aga
rdh)
, Syn
edra
ta
bula
ta (C
. Aga
rdh)
Küt
zing
, D
iato
ma
tabu
lata
C. A
gard
h,
Dia
tom
a ta
bula
ta C
.A. A
gard
h,
Dia
tom
a ta
bula
tum
C. A
gard
h,
Exi
laria
fasc
icul
ata
(C. A
gard
h)
Gre
ville
, Lyn
gbye
a fa
scic
ulat
a (C
. A
gard
h) S
omm
erfe
lt, E
chin
ella
fa
scic
ulat
a (A
gard
h) J
urge
ns,
Dia
tom
a fa
scic
ulat
a C
.A. A
gard
h,
Syn
edra
fasc
icul
ata
(Aga
rdh)
K
ützi
ng, T
abul
aria
tabu
lata
(C
. A
gard
h) S
noei
js
Eas
t/Wes
t (E
ffing
ham
In
let)
Hay
et a
l. 20
03
Tabu
laria
inve
stie
ns (
W. S
mith
) W
illia
ms
& R
ound
Tabu
laria
inve
stie
ns (
W. S
mith
) D
.M. W
illia
ms
& R
ound
(ab)
Syn
edra
inve
stie
ns W
.Sm
ithW
est
(Effi
ngha
m
Inle
t)
Hay
et a
l. 20
03 (B
érar
d-Th
erria
ult
et a
l. 19
99)
Tabu
laria
tabu
lata
(K
utzi
ng) D
.M.
Will
iam
s &
F.E
. Rou
ndTa
bula
ria ta
bula
ta (
C. A
gard
h) D
.M.
Will
iam
s &
Rou
nd (c
ol)
Eas
tZw
icke
r 199
8
Thal
assi
onem
a ni
tzsc
hioi
des
Gru
now
Thal
assi
onem
a ni
tzsc
hioi
des
(Gru
now
) Mer
esch
kow
sky
(ab)
Thal
assi
othr
ix n
itzsc
hioi
des
var.
java
nica
Gru
now
, Syn
edra
ni
tzsc
hioi
des
Gru
now
, Th
alas
siot
hrix
nitz
schi
oide
s (G
runo
w) G
runo
w, T
hala
ssio
thrix
cu
rvat
a C
astra
cane
, Syn
edra
ni
tzsc
hioi
des
var.
min
or C
leve
, Th
alas
siot
hrix
frau
nfel
dii
var.
Nitz
schi
oide
s (G
runo
w)
Jörg
ense
n
Eas
t/Wes
t/Ar
ctic
Link
lette
r 197
7 (S
ance
tta &
C
alve
rt 19
88, B
érar
d-Th
erria
ult e
t al
. 199
9, B
rune
l 196
2, M
cQuo
id &
H
obso
n 19
97, M
cQuo
id &
Hob
son
2001
, Hay
et a
l. 20
03, R
ozan
ska
et a
l. 20
08, H
orne
r 200
2,
Raw
lenc
e 19
87)
Thal
assi
osira
aes
tival
is G
ran
&
Ang
stTh
alas
sios
ira a
estiv
alis
Gra
n (a
b)W
est (
US
)H
orne
r et a
l. 20
05 (S
ance
tta &
C
alve
rt 19
88, M
cQuo
id &
Hob
son
1997
)
Pag
e 73
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Thal
assi
osira
ang
ulat
a (G
rego
ry)
Has
leTh
alas
sios
ira a
ngul
ata
(Gre
gory
) H
asle
(ab)
Orth
osei
ra a
ngul
ata
Gre
gory
, Th
alas
sios
ira d
ecip
iens
(Gru
now
) E
.G. J
ørge
nsen
Eas
t/Wes
t (E
ffing
ham
in
let)/
Arc
tic
Hay
et a
l. 20
03 (L
ovej
oy e
t al.
2002
)
Thal
assi
osira
ang
uste
-line
ata
(A.
Sch
mid
t) Fr
yxel
l & H
asle
Thal
assi
osira
ang
uste
-line
ata
(A.
Sch
mid
t) G
. Fry
xell
& H
asle
(ab)
Thal
assi
osira
pol
ycho
rda
(G
ran)
Jø
rgen
sen,
Cos
cino
disc
us p
olyc
hord
us G
ran,
Cos
cino
disc
us a
ngus
te-li
neat
a A
. S
chm
idt,
Cos
cino
sira
pol
ycho
rda
(Gra
n) G
ran
Eas
t/Wes
t /A
rctic
Hor
ner e
t al.
2005
(San
cetta
&
Cal
vert
1988
, Bér
ard-
Ther
riaul
t et
al. 1
999,
Lov
ejoy
et a
l. 20
02,
Boo
th &
Hor
ner 1
997,
McQ
uoid
&
Hob
son
1997
, McQ
uoid
& H
obso
n 20
01)
Thal
assi
osira
ant
arct
ica
Thal
assi
osira
ant
arct
ica
Com
ber
(ab)
Thal
assi
osira
ant
arct
ica
Kar
sten
, Th
alas
sios
ira fa
llax
Meu
nier
, Th
alas
sios
ira a
ntar
ctic
a v
ar.
bore
alis
G. F
ryxe
ll, D
ouce
tte, &
H
ubba
rd
Eas
tB
érar
d-Th
erria
ult e
t al.
1999
(R
iede
l et a
l. 20
03)
Thal
assi
osira
ant
arct
ica
var.
bore
alis
Thal
assi
osira
ant
arct
ica
Com
ber
(ab)
Thal
assi
osira
ant
arct
ica
var
. bo
real
is G
. Fry
xell,
Dou
cette
, &
Hub
bard
Arc
ticR
iede
l et a
l. 20
03 (L
ovej
oy e
t al.
2002
)
Thal
assi
osira
bal
tica
(Gru
n.) O
sten
.Th
alas
sios
ira b
altic
a (G
runo
w)
Ost
enfe
ld (a
b)C
osci
nodi
scus
pol
yaca
nthu
s v
ar.
balti
cus
Gru
now
, Cos
cino
disc
us
balti
cus
(Gru
now
) Gru
now
ex
Cle
ve
Eas
tR
awle
nce
1987
(Bru
nel 1
962)
Thal
assi
osira
bio
cula
taTh
alas
sios
ira b
iocu
lata
(Gru
now
) O
sten
feld
(ab)
Cos
cino
disc
us b
iocu
latu
s G
runo
wE
ast/A
rctic
Bér
ard-
Ther
riaul
t et a
l. 19
99
(Bru
nel 1
962,
Bur
sa 1
963,
Lov
ejoy
et
al.
2002
, Gra
n &
Bra
arud
193
5)
Thal
assi
osira
bio
cula
ta v
ar. e
xigu
aTh
alas
sios
ira b
iocu
lata
var
. exi
gua
(Gru
now
) Hus
tedt
(ab)
Eas
tB
érar
d-Th
erria
ult e
t al.
1999
Thal
assi
osira
bul
bosa
Syv
erts
enTh
alas
sios
ira b
ulbo
sa E
.E.
Syv
erts
en (a
b)A
rctic
Love
joy
et a
l. 20
02
Thal
assi
osira
con
dens
ata
Det
onul
a pu
mila
(Cas
traca
ne) G
ran
Thal
assi
osira
con
dens
ata
Cle
veE
ast
Bru
nel 1
962
Thal
assi
osira
con
ferta
Has
le in
H
asle
and
Fry
xell
Thal
assi
osira
con
ferta
Has
le (a
b)W
est
(Effi
ngha
m
Inle
t)
Hay
et a
l. 20
03 (B
érar
d-Th
erria
ult
et a
l. 19
99)
Pag
e 74
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Thal
assi
osira
dec
ipie
nsTh
alas
sios
ira a
ngul
ata
(Gre
gory
) H
asle
(ab)
Thal
assi
osira
dec
ipie
ns (G
runo
w)
E.G
. Jør
gens
en, C
osci
nodi
scus
de
cipi
ens
Gru
now
ex
Van
Heu
rck,
Th
alas
sios
ira g
elat
inos
a H
ense
n
Eas
t/Wes
tB
rune
l 196
2 (G
ran
& B
raar
ud
1935
, Hay
et a
l. 20
03)
Thal
assi
osira
ecc
entri
ca
(Ehr
enbe
rg) C
leve
Thal
assi
osira
ecc
entri
ca
(Ehr
enbe
rg) C
leve
(ab)
Cos
cino
disc
us e
ccen
tricu
s E
hren
berg
, Cos
cino
disc
us
laby
rinth
us R
oper
Eas
t/Wes
tH
orne
r et a
l. 20
05 (S
ance
tta &
C
alve
rt 19
88, M
cQuo
id &
Hob
son
1997
, McQ
uoid
& H
obso
n 20
01,
Hay
et a
l. 20
03)
Thal
assi
osira
falla
xTh
alas
sios
ira a
ntar
ctic
a C
ombe
r (a
b)Th
alas
sios
ira fa
llax
Meu
nier
Eas
tB
rune
l 196
2
Thal
assi
osira
fluv
iatil
isTh
alas
sios
ira w
eiss
flogi
i (G
runo
w)
G. F
ryxe
ll &
Has
le (a
b)Th
alas
sios
ira fl
uvia
tilis
Hus
tedt
Eas
tB
rune
l 196
2
Thal
assi
osira
gra
vida
Cle
veTh
alas
sios
ira g
ravi
da C
leve
(ab)
Thal
assi
osira
tche
rnia
i M
angu
inE
ast/W
est(U
S)/
Arc
ticR
awle
nce
1987
(Cup
p 19
43,
San
cetta
& C
alve
rt 19
88, B
érar
d-Th
erria
ult e
t al.
1999
, Bru
nel 1
962,
B
ursa
196
3, G
ran
& B
raar
ud 1
935,
M
cQuo
id &
Hob
son
1997
, M
cQuo
id &
Hob
son
2001
)
Thal
assi
osira
hya
lina
Thal
assi
osira
hya
lina
(Gru
now
) G
ran
(ab)
Cos
cino
disc
us h
yalin
us G
runo
w
Eas
t/Arc
ticB
érar
d-Th
erria
ult e
t al.
1999
(B
rune
l 196
2, B
ursa
196
3, L
ovej
oy
et a
l. 20
02, G
ran
& B
raar
ud 1
935)
Thal
assi
osira
kus
hire
nsis
Thal
assi
osira
kus
chire
nsis
Tak
ano
(ab)
Thal
assi
osira
sol
itaria
Gay
oso
Eas
tB
érar
d-Th
erria
ult e
t al.
1999
Thal
assi
osira
leva
nder
iTh
alas
sios
ira le
vand
eri
Van
Goo
r (a
b)E
ast
Bru
nel 1
962
Thal
assi
osira
cf.
lept
opus
(G
runo
w) H
asle
& F
ryxe
llTh
alas
sios
ira le
ptop
us (G
runo
w e
x V
an H
eurc
k) H
asle
& G
. Fry
xell
(ab)
Cos
cino
disc
us le
ptop
us v
ar.
disc
repa
ns R
attra
y,
Cos
cino
disc
us p
seud
olin
eatu
s P
anto
csek
, Cos
cino
disc
us
lept
opus
Gru
now
, Cos
cino
disc
us
linea
tus
Ehr
enbe
rg,
Cos
cino
disc
us p
rael
inea
tus
Jous
é
Eas
t/Wes
t (E
ffing
ham
In
let)
Hay
et a
l. 20
03
Pag
e 75
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Thal
assi
osira
nan
aTh
alas
sios
ira n
ana
Lohm
ann
(ab)
Eas
tB
rune
l 196
2
Thal
assi
osira
nor
dens
kioe
ldii
Cle
veTh
alas
sios
ira n
orde
nski
oeld
ii C
leve
(a
b)E
ast/W
est/A
rct
icH
orne
r et a
l. 20
05 (S
ance
tta &
C
alve
rt 19
88, B
érar
d-Th
erria
ult e
t al
. 199
9, B
rune
l 196
2, R
iede
l et a
l. 20
03, L
ovej
oy e
t al.
2002
, Gra
n &
B
raar
ud 1
935,
Bat
es &
Stra
in
2006
, McQ
uoid
& H
obso
n 19
97,
McQ
uoid
& H
obso
n 20
01, H
ay e
t al
. 200
3)Th
alas
sios
ira o
cean
ica
Thal
assi
osira
oce
anic
a H
asle
(ab)
Cyc
lote
lla n
ana
Hus
tedt
E
ast
Bér
ard-
Ther
riaul
t et a
l. 19
99
Thal
assi
osira
oes
trupi
i (O
sten
feld
) H
asle
Thal
assi
osira
oes
trupi
i (O
sten
feld
) H
asle
(ab
)C
osci
nosi
ra o
estru
pii
Ost
enfe
ld,
Thal
assi
osira
ant
iqua
var
. se
ptat
a P
rosc
hkin
a-La
vren
ko
Eas
t/Wes
t (E
ffing
ham
in
let)
Hay
et a
l. 20
03
Thal
assi
osira
pac
ifica
Gra
n &
A
ngst
Thal
assi
osira
pac
ifica
Gra
n &
Ang
st
(ab)
Eas
t/Wes
tH
orne
r et a
l. 20
05 (S
ance
tta &
C
alve
rt 19
88, B
érar
d-Th
erria
ult e
t al
. 199
9, M
cQuo
id &
Hob
son
2001
, Hay
et a
l. 20
03)
Thal
assi
osira
plic
ata
Thal
assi
osira
plic
ata
H. J
. Sch
rade
r (a
b)W
est
McQ
uoid
& H
obso
n 19
97
Thal
assi
osira
pro
schk
inae
Thal
assi
osira
pro
schk
inae
Mak
arov
a (a
b)Th
alas
sios
ira s
pinu
lata
Tak
ano,
Th
alas
sios
ira p
rosc
hkin
ae v
ar.
spin
ulat
a (T
akan
o) M
akar
ova
Eas
tB
érar
d-Th
erria
ult e
t al.
1999
Thal
assi
osira
pse
udon
ana
Thal
assi
osira
pse
udon
ana
Has
le &
H
eim
dal (
ab)
Eas
tB
érar
d-Th
erria
ult e
t al.
1999
Thal
assi
osira
pun
ctig
era
(Cas
traca
ne) H
asle
Thal
assi
osira
pun
ctig
era
(Cas
traca
ne) H
asle
(ab
)Th
alas
sios
ira ja
poni
ca K
isse
lev,
E
thm
odis
cus
japo
nicu
s C
astra
cane
, Eth
mod
iscu
s pu
nctig
er C
astra
cane
, Th
alas
sios
ira a
ngst
ii (G
ran)
M
akar
ova
Eas
t (B
oF)/W
est
Hor
ner e
t al.
2005
(San
cetta
&
Cal
vert
1988
, Car
ver &
Mal
let
2002
)
Thal
assi
osira
rotu
la M
euni
erTh
alas
sios
ira ro
tula
Meu
nier
(ab)
Eas
t/Wes
t/Ar
ctic
Hor
ner e
t al.
2005
(San
cetta
&
Cal
vert
1988
, Bru
nel 1
962,
Bur
sa
1963
)Th
alas
sios
ira s
p.Th
alas
sios
ira s
p.E
ast/A
rctic
Bur
sa 1
963
(Gra
n &
Bra
arud
19
35)
Pag
e 76
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Thal
assi
osira
sub
tilis
(O
sten
.) G
ran.
Thal
assi
osira
sub
tilis
(Ost
enfe
ld)
Gra
n (a
b)P
odos
ira s
ubtil
is O
sten
feld
Eas
t/Wes
t(US
)R
awle
nce
1987
(Cup
p 19
43,
Bru
nel 1
962,
Bur
sa 1
963)
Thal
assi
osira
tene
ra P
rosc
hkin
a-La
vren
koTh
alas
sios
ira te
nera
Pro
schk
ina-
Lavr
enko
(ab)
Eas
t/Wes
t (E
ffing
ham
In
let)
Hay
et a
l. 20
03 (B
érar
d-Th
erria
ult
et a
l. 19
99)
Thal
assi
othr
ix lo
ngis
sim
a C
leve
an
d G
runo
wTh
alas
siot
hrix
long
issi
ma
Cle
ve &
G
runo
w (
ab)
Syn
edra
thal
assi
othr
ix C
leve
Eas
t/Wes
t(US
)/ A
rctic
Link
lette
r et a
l. 19
77 (C
upp
1943
, B
érar
d-Th
erria
ult e
t al.
1999
, Frit
z 19
21, B
ursa
196
3, L
ovej
oy e
t al.
2002
, Gra
n &
Bra
arud
193
5)
Thal
assi
othr
ix n
itzsc
hioi
des
Thal
assi
onem
a ni
tzsc
hioi
des
(Gru
now
) Mer
esch
kow
sky
(ab)
Thal
assi
othr
ix n
itzsc
hioi
des
(Gru
now
) Gru
now
Eas
tFr
itz 1
921
(Bru
nel 1
962)
Trac
hyne
is a
sper
a (E
hren
berg
) C
leve
Trac
hyne
is a
sper
a (E
hren
berg
) C
leve
(ab)
Nav
icul
a st
auro
neifo
rmis
Le
udug
er-F
ortm
orel
, Pin
nula
ria
aspe
ra (
Ehr
enbe
rg) E
hren
berg
, S
taur
onei
s pu
lche
lla W
.Sm
ith,
Sta
uron
eis
achn
anth
es
(Ehr
enbe
rg) K
ützi
ng, S
taur
onei
s as
pera
(E
hren
berg
) Küt
zing
, S
taur
opte
ra a
sper
a E
hren
berg
, N
avic
ula
aspe
ra E
hren
berg
, N
avic
ula
aspe
ra v
ar. g
enui
na
(Cle
ve) F
ricke
Eas
tB
aile
y 19
13 (T
hale
r 200
7)
Tric
erat
ium
alte
rnan
sB
iddu
lphi
a al
tern
ans
(J.W
. Bai
ley)
V
an H
eurc
kTr
icer
atiu
m a
ltern
ans
J.W
. Bai
ley
Eas
tB
érar
d-Th
erria
ult e
t al.
1999
Tric
erat
ium
favu
sTr
icer
atiu
m fa
vus
Ehr
enbe
rgTr
icer
atiu
m c
ompt
um E
hren
berg
, Tr
icer
atiu
m m
uric
atum
Brig
htw
ell,
Tr
icer
atiu
m fi
mbr
iatu
m W
allic
h,
Bid
dulp
hia
favu
s (E
hren
berg
) G
runo
w,
Tric
erat
ium
fero
x C
astra
cane
, Tric
erat
ium
sa
rcop
hagu
s C
astra
cane
Eas
tB
rune
l 196
2
Tric
erat
ium
sp.
Tric
erat
ium
sp.
Wes
tM
cQuo
id &
Hob
son
2001
Trig
oniu
m s
p.Tr
igon
ium
sp.
Wes
tH
obso
n 20
09Tr
opid
onei
s an
tarc
tica
var.
poly
plas
ta G
ran
& A
ngst
Trop
idon
eis
anta
rctic
a va
r. po
lypl
asta
Gra
n &
Ang
st (a
b)W
est
Hor
ner (
pers
. com
m.)
3
Pag
e 77
Nam
e us
ed in
ori
gina
l pap
erC
urre
nt u
sed
(val
id) n
ame
(=Sy
nony
ms)
Loc
atio
nR
efer
ence
(s)
Trop
idon
eis
lepi
dopt
era
(Gre
gory
) C
leve
Trop
idon
eis
lepi
dopt
era
(Gre
gory
) C
leve
(ab)
Am
phip
rora
lepi
dopt
era
Gre
gory
, Tr
opid
onei
s le
pido
pter
a v
ar.
med
iterr
anea
Per
agal
lo
Wes
t (U
S)
Cup
p 19
43
Trop
idon
eis
sp.
Trop
idon
eis
sp.
Arc
ticB
ooth
& H
orne
r 199
7Tr
yblio
nella
gra
cilis
N
itzsc
hia
trybl
ione
lla H
antz
sch
(ab)
Tryb
lione
lla g
raci
lis W
. Sm
ithE
ast
Bai
ley
1910
(Bai
ley
1913
)
Tryb
lione
lla le
vide
nsis
W.S
mith
Tryb
lione
lla le
vide
nsis
W. S
mith
(ab)
Nitz
schi
a le
vide
nsis
(W
.Sm
ith)
Gru
n, N
itzsc
hia
trybl
ione
lla v
ar.
levi
dens
is (
W. S
mith
) Gru
now
, D
entic
ula
levi
dens
is (W
. Sm
ith)
De
Toni
, Try
blio
nella
tryb
lione
lla
var.
levi
dens
is (
W. S
mith
) P
roch
azka
, Try
blio
nella
ha
ntzs
chia
na v
ar. l
evid
ensi
s (W
. S
mith
) Fre
ngue
lli
Eas
tTh
aler
200
7 (P
oulin
et a
l. 19
90)
Tryb
lione
lla s
cute
llum
Sm
.Tr
yblio
nella
scu
tellu
m W
. Sm
ith (a
b)E
ast
Bai
ley
1910
(Bai
ley
1913
)
Uro
sole
nia
erie
nsis
(H.L
. Sm
ith)
Rou
nd &
Cra
wfo
rdU
roso
leni
a er
iens
is(H
.L. S
mith
) F.E
. R
ound
& R
.M. C
raw
ford
(ab)
Rhi
zoso
leni
a er
iens
is H
.L. S
mith
, R
hizo
sole
nia
erie
nsis
var
. ge
nuin
a A
. Cle
ve-E
uler
Eas
t (B
oF)
Gin
n (p
ers.
com
m.)
1 (Bér
ard-
Ther
riaul
t et a
l. 19
99)
Zygo
cero
s m
obile
nsis
Bai
l.O
dont
ella
mob
ilien
sis
(J.W
. Bai
ley)
G
runo
wZy
goce
ros
mob
ilien
sis
J.W
. B
aile
yE
ast
Bai
ley
1910
Pag
e 78