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peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;
2
PRLS =maximum lifespan − age at last birth
maximum lifespan − age of female reproductive maturity
Offspring Ratio =offspring weight at birth
adult weightOR
offspring length3
adult length3
Litter Size Adjusted Offspring Ratio = (Offspring Ratio)* (Average Litter Size)
Reproductive Window =1 −
Age at Last Birth − Age of Reproductive Maturity
Maximum Age
peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;
3
PRLS =maximum lifespan − age at last birth
maximum lifespan − age of female reproductive maturity
Offspring Ratio =offspring weight at birth
adult weightOR
offspring length3
adult length3
peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;
4
ReproductiveMaturity
AdulthoodDevelopment
Reproductive Senescence
DEATH
IZATION
Gestation Weaning
Birth
Maximum Documented Lifespan
Pro
po
rtio
n R
epro
du
ctiv
e o
r A
live
Fertilization
Reproductive Period PRLS
A B
CCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCCC
peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;
5
peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;
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A B
C D
*
E F
frac
tion
repr
oduc
tive
frac
tion
aliv
efr
actio
n re
prod
uctiv
e
mat
ricid
e fr
actio
n
% re
prod
uctiv
e/al
ive
% re
prod
uctiv
e/al
ive
time (days of adulthood) time (days of adulthood)
time (days of adulthood) time (days of adulthood)
time (days of adulthood) time (days of adulthood)
peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;
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peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;
8
peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;
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peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;
10
peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;
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peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;
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peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;
peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;
peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;
peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;
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peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;
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peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;
15
peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;
peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;
peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;
peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;
peer-reviewed) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprint (which was not.http://dx.doi.org/10.1101/048835doi:bioRxiv preprint first posted online Apr. 15, 2016;