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POPULATION ECOLOGYCh 40 (Urry)
© 2014 Pearson Education, Inc.
Concept 40.4: Population density, dispersion, and demographics
▪ biotic and abiotic factors influence density, distribution, and size of populations
Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings
Population size• 4 factors in pop size:
Births and immigration add individuals to a population.
Natality Immigration
PopuIation size
Emigration
Deaths
Deaths and emigration remove individuals from a population.
Population• A population
• Is a group of individuals of a single species living in the same general area• Density:• Dispersion:
• clumped• uniform• random
Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings
Life Tables• Summary of the survival pattern of a population • Follows the fate of a cohort • Age specific
Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings
Life Table: Belding’s ground squirrels• Range: E. OR & CA, S. ID, cent. NV,
Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings
Survivorship Curves• for Belding’s ground squirrels
– death rate is relatively constant
Figure 52.4
1000
100
10
1
Num
ber o
f sur
vivo
rs (l
og s
cale
)
0 2 4 6 8 10Age (years)
Males
Females
© 2014 Pearson Education, Inc.
Figure 40.16
1,000
Percentage of maximum life span
III
Num
ber o
f sur
vivo
rs (l
og sc
ale)
100
10
01
10050
II
I
Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings
Pop Growth Rates• How fast does pop size change? • Simplest calculation:
– N = pop – t = time – B = birth – D = death∆N∆t
= B - D
∆N∆t
= bN - dN b = per capita birth rate d = per capita death rate
∆N∆t
= rN Instantaneous growth rate (r = b - d)
Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings
Population growth rate• J-curve
– under what conditions? – can it be sustained?
Figure 52.9
0 5 10 150
500
1,000
1,500
2,000
Number of generations
Pop
ulat
ion
size
(N)
dNdt
= 1.0N
dNdt
= 0.5N
© 2014 Pearson Education, Inc.
Figure 40.18
6,000
Year
Elep
hant
pop
ulat
ion
0
2,000
8,000
4,000
1900 1930 1940 1950 1960 19701910 1920
Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings
Logistic Growth• Carrying capacity (K)
– = maximum pop size the env. can support
• Use logistic growth model
Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings
Logistic Growth model• Sigmoid (S-shaped) curve
Figure 52.12
dNdt
= 1.0N Exponential growth
Logistic growth
dNdt
= 1.0N1,500 − N
1,500
K = 1,500
0 5 10 150
500
1,000
1,500
2,000
Number of generations
Pop
ulat
ion
size
(N)
Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings
The Logistic Model and Life Histories• K-selection, or density-dependent selection
– Selects for life history traits that are sensitive to population density
• r-selection, or density-independent selection – Selects for life history traits that maximize
reproduction
Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings
Regulation of Pop Size• complex interaction of biotic and abiotic
influences • Two general questions:
– What factors stop a population from growing? – Why do some populations show radical
fluctuations in size over time, while others remain stable?
Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings
Density-Dependent Population Regulation• Density-dependent birth and death rates
– Competition for resources – health (disease) – predation – toxic wastes
Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings
Population Cycles• Many populations
– Undergo regular boom-and-bust cycles
Figure 52.21 Year1850 1875 1900 1925
0
40
80
120
160
0
3
6
9
Lynx
pop
ulat
ion
size
(th
ousa
nds)
Har
e po
pula
tion
size
(th
ousa
nds)
Lynx
Snowshoe hare
Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings
Human Pop Growth• What is our growth curve? • What’s going on with our growth rate? • No population can grow indefinitely
– humans are no exception
Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings
The Global Human Population• The human population
– Increased relatively slowly until about 1650 and then began to grow exponentially
Figure 52.228000 B.C.
4000 B.C.
3000 B.C.
2000 B.C.
1000 B.C.
1000 A.D.
0
The Plague Hum
an p
opul
atio
n (b
illio
ns)
2000 A.D.
0
1
2
3
4
5
6
Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings
• Though the global population is still growing – The rate of growth began to slow approximately
40 years ago
Figure 52.231950 1975 2000 2025 2050
Year
2003
Per
cent
incr
ease
2.2
2
1.6
1.4
1.2
1
0.8
0.6
0.4
0.2
0
1.8
Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings
Regional Patterns of Population Change• To maintain population stability
– A regional human population can exist in one of two configurations • High birth rate, high death rate • Low birth rate, low death rate
Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings
Demographic Transition• = the move from the first toward the second state
Figure 52.24
50
40
20
0
30
10
1750 1800 1850 1900 1950 2000 2050
Birth rateDeath rate
Birth rateDeath rate
Year
Sweden Mexico
Birt
h or
dea
th ra
te p
er 1
,000
peo
ple
Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings
Age structure• the relative number of individuals at each age • Is commonly represented in pyramids
Figure 52.25
Rapid growth Afghanistan
Slow growth United States
Decrease Italy
Male Female Male Female Male FemaleAge Age
8 6 4 2 0 2 4 6 8 8 6 4 2 0 2 4 6 8 8 6 4 2 0 2 4 6 8Percent of population Percent of population Percent of population
80–8485+
75–7970–7465–6960–6455–5950–5445–4940–4435–3930–34
20–2425–29
10–145–90–4
15–19
80–8485+
75–7970–7465–6960–6455–5950–5445–4940–4435–3930–34
20–2425–29
10–145–90–4
15–19
Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings
Global Carrying Capacity• Just how many humans can the biosphere
support? • $64k question
Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings
Ecological Footprint• The ecological footprint concept
– Summarizes the aggregate land and water area needed to sustain the people of a nation
– Is one measure of how close we are to the carrying capacity of Earth
Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings
• Ecological footprints for 13 countries – Show that the countries vary greatly in their footprint
size and their available ecological capacity
Figure 52.27
16
14
12
10
8
6
4
2
00 2 4 6 8 10 12 14 16
New Zealand
AustraliaCanada
Sweden
WorldChina
India
Available ecological capacity (ha per person)
SpainUK
Japan
GermanyNetherlands
Norway
USA
Eco
logi
cal f
ootp
rint (
ha p
er p
erso
n)
Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings
• At more than 6 billion people – The world is already in ecological deficit
Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings
Stability and Fluctuation• Long-term population studies
– Isle Royale moose first migrated to island c. 1900