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 CELLULOSE BIOSYNTHESIS

IN OOMYCETES

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  Oomycetes can be biotrophs,hemibiotrophs or necrotrophs.

  Biotrophic species obtain their nutrients

from living tissues  Hemibiotrophs rst feed on living tissues

and !ill their host at a later stage. 

"ecrotrophs live e#lusively on dead hosttissues.

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   The Oomycete cellulose is microbrillar,of low crystallinity .

  Since little se-uence information from

Oomycetes has been available untilrecently, genes encoding the $esAcatalytic subunit in these microorganismsremain un!nown.

  "o $esA terminal comple# has beenobserved in Oomycetes

  $esA $ellulose synthases

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   The e#act mode of assembly of thecarbohydrate components in the cellwalls of Oomycetes is not !nown, even

though it is believed that the cell wallsconsist of an inner layer of cellulose .

  $ellulose in Oomycetes has most li!elythe same sca/olding function as chitin in

fungal cell walls.