1

21

..

2-3

2

2-3

1 2

3-4

1

6-7

3

2

elytra (

elytron)

?

4

2

5

6

7

: Chalcosoma caucasus

: Giant rhinoceros beetle

120 .

3

8

4

9

10

5

11

12

6

..

13

14

7

17

18

Dynastes hercules

Scarabaeidae

190

65-70

16

(Hercules beetle)

19

150

60-65

(Neptune beetle)

20

100-120

60-65

(Elephant beetle)

70-100

87-100 70-75

(goliath beetle)

21

130

(Causcasus three-horned beetle)

22

8

23

24

(compound

eyes) 1

1-2

11

(scape)

(pedicel)

3-11 (flagellum)

(mandible)

(stag)

stag beetle

(labrum)

(labium)

(head)

25

3

(prothorax)

(pronotum)

(meso-

thorax)

1

2

7 8

(tergite)

(sternite)

,

, , , 5

(thorax)

(abdomen)

1

2

3

26

(Complete metamorphosis)

(prolegs)

Eruciform

4

Sca rabae i f o rm (Wh i te grub- l i ke )

C

3

27

Vermiform (maggot-like)

(Cerambycidae)

(round-headed bor-

er) (Buprestidae)

(flat-headed borer)

Eruciform (Scarabaeidae)

(Lucanidae)

(Passalidae) Campodeiform (diplu-

ran-like)

(Diplura) (Thysanura)

3

(Meloidae)

(Coccinellidae)

Elateriform (wireworm-like)

(Elateridae)

(Tenebrionidae)

28

4

1. (egg)

2. (larva)

3. (pupa)

(Exatate pupa)

4. (adult)

29

(predators)

Coccinellidae

Monochilus sexmaculatus

3

30

9

31

32

(Flighting beetle)

33

12

3

1

3

3

300

34

""

35

100 -120

15 20

C

(Scarabaeiform type)

30

6

8

30 35

2 5

36

10

37

38

11

39

40

41

12

42

2

(Dynastinae,

Scarabaeidae)

(Lucanidae)

48-70

1

2

2-3

(Eupatorus gracilicornis)

43

2.5-4.0

10.5

7.5

1

38.5-79.5

42.2-54.0

()(Japanese rhinoceros beetle)

(Hexarthrius parryi)

(Giraffe stag beetle)

44

1

15-20

1.

2-3

2.

10-20

3.

4.

45

1.

1:1 1:2

4-5

2.

5

15-20

()

3.

45-60

5-6

4.

5.

46

47

48