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8/9/2019 Chapter 3 Plant Nomenclature
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Plant NomenclaturePlant Nomenclature
Systematic BotanySystematic Botany
Chapter 3Chapter 3
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NomenclatureNomenclature
assignment of names to plantsassignment of names to plants
involves principles governed by rulesinvolves principles governed by rules
developed and adopted by thedeveloped and adopted by theInternational Botanical CongressesInternational Botanical Congresses
rules formally listed in therules formally listed in the InternationalInternational
Code of Botanical NomenclatureCode of Botanical Nomenclature nomenclature and classification arenomenclature and classification are
different but inseparabledifferent but inseparable
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International Code of BotanicalInternational Code of Botanical
Nomenclature or CodeNomenclature or Code
ultimate goal is to provide ONE correctultimate goal is to provide ONE correct
name for each taxonname for each taxon
the rules are subdivided intothe rules are subdivided into articlesarticles,,
which must be adhered to, andwhich must be adhered to, and
recommendationsrecommendations, which are optional, which are optional
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Basis of Scientific NamesBasis of Scientific Names
the present system is the result of athe present system is the result of ahistoric series of changes that graduallyhistoric series of changes that graduallybecome formalizedbecome formalized
oldest plant namesoldest plant names common namescommon names
used in ancient Greece and Romeused in ancient Greece and Rome
present namespresent names LatinizedLatinized spellingspelling
before 1753before 1753 polynomialspolynomials
LinnaeusLinnaeus Species PlantarumSpecies Plantarumbinomial nomenclaturebinomial nomenclature
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Scientific Names versusScientific Names versus
Common NamesCommon Names5 reasons why botanists use Latinized scientific5 reasons why botanists use Latinized scientific
names (SN) instead of common names (CN)names (SN) instead of common names (CN)
1.1. SN is universal and are recognized throughoutSN is universal and are recognized throughoutthe worldthe world
2.2. CN usually do not provide information indicatingCN usually do not provide information indicatingthe generic and family relationshipsthe generic and family relationships
3.3. A wellA well--known plant may have several commonknown plant may have several commonnamesnames
4.4. Sometimes, two or more plants may have theSometimes, two or more plants may have thesame common namesame common name
5.5. Many species, particularly rare ones, do not haveMany species, particularly rare ones, do not havecommon namescommon names
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Composition of Scientific NamesComposition of Scientific Names
Consists of three elements:Consists of three elements:
1. Generic names1. Generic names
2. Specific epithets2. Specific epithets
3. Authors name3. Authors name
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a singular Latinized noun or a word treateda singular Latinized noun or a word treatedas a nounas a noun
always written with an initial capital letteralways written with an initial capital letter
may not consist of two words unless they aremay not consist of two words unless they arejoined with a hyphenjoined with a hyphen
may be taken from any source, and it maymay be taken from any source, and it maycommemorate some person of distinctioncommemorate some person of distinction
customarilycustomarily underlinedunderlined when written or typed;when written or typed;when printed, they are inwhen printed, they are in italicsitalics ororboldfaceboldface
Composition of Scientific NamesComposition of Scientific Names
Generic NamesGeneric Names
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may be derived from any source and maymay be derived from any source and mayhonor a person, or they may be derived from anhonor a person, or they may be derived from anold common name, a geographic location, orold common name, a geographic location, or
some characteristic of the plantsome characteristic of the plant often an adjective illustrating a distinguishingoften an adjective illustrating a distinguishingfeature of a speciesfeature of a species
may consists of two words but must bemay consists of two words but must be
hyphenatedhyphenated usually agrees with theusually agrees with the gendergenderof the genericof the genericname (if adjective)name (if adjective)
Composition of Scientific NamesComposition of Scientific Names
Specific epithetSpecific epithet
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Composition of Scientific NamesComposition of Scientific Names
Specific epithetSpecific epithet if named after a person, (1) ending in a vowelif named after a person, (1) ending in a voweloror erer, the letter, the letteriiis added, (2) ending in ais added, (2) ending in aconsonant, the letterconsonant, the letteriiiiare added, (3) for aare added, (3) for a
female, it ends infemale, it ends iniae oriae or aeae when derived from geographical names theywhen derived from geographical names theyare usually terminated byare usually terminated byensis,ensis, --(a)nus,(a)nus, --inus,inus,--ianus or icusianus or icus
must be written with a small initial lettermust be written with a small initial letter customarilycustomarily underlinedunderlined when written or typed;when written or typed;when printed, they are inwhen printed, they are in italicsitalics ororboldfaceboldface
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Composition of Scientific NamesComposition of Scientific Names
Authors NameAuthors Name a source of historical information regarding the namea source of historical information regarding the name
of the plantof the plant
may be abbreviated or full namemay be abbreviated or full name
may consists of two names, with the first inmay consists of two names, with the first inparentheses indicating the person who first describedparentheses indicating the person who first describedthe plant, and the second name indicate the name ofthe plant, and the second name indicate the name ofthe person who made the changesthe person who made the changes
name changes should be made only after carefulname changes should be made only after carefulconsideration of taxonomic relationships and mustconsideration of taxonomic relationships and mustfollow the requirements of the ICBNfollow the requirements of the ICBN
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Rules of NomenclatureRules of Nomenclature
Linnaeus in 1737 and 1751Linnaeus in 1737 and 1751
1818thth century,century, prioritypriority or the use of the oldest name wasor the use of the oldest name wasrecognized as the cornerstone of nomenclaturerecognized as the cornerstone of nomenclature
AP de CandolleAP de Candolle detailed set or rules on the processdetailed set or rules on the processof assigning names, thus evolving the ICBNof assigning names, thus evolving the ICBN
LinnaeusLinnaeus synonymy of namessynonymy of names
Nomenclator botanicusNomenclator botanicus Steudel (1821, 1840Steudel (1821, 1840--41) an41) an
index of plant names; useful for checking names andindex of plant names; useful for checking names andsynonyms; forerunner ofsynonyms; forerunner ofIndex kewensisIndex kewensis
18671867 11stst International Botanical Congress in ParisInternational Botanical Congress in Parisfollowed by 1892, 1905, 1907 and 1910followed by 1892, 1905, 1907 and 1910
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Rules of Nomenclature:Rules of Nomenclature:
PrinciplesPrinciples1.1. Botanical nomenclature is independent of zoologicalBotanical nomenclature is independent of zoological
nomenclaturenomenclature
2.2. The application of names of taxonomic groups isThe application of names of taxonomic groups is
determined by means of nomenclatural types.determined by means of nomenclatural types.3.3. The nomenclature of a taxonomic group is basedThe nomenclature of a taxonomic group is based
upon the priority of publication.upon the priority of publication.
4.4. Each taxonomic group with a particularEach taxonomic group with a particularcircumscription, position and rank can bear only onecircumscription, position and rank can bear only one
correct name, the earliest that is in accordance withcorrect name, the earliest that is in accordance withthe rulesthe rules
5.5. Scientific names of taxonomic groups are treated asScientific names of taxonomic groups are treated asLatin regardless of their derivation.Latin regardless of their derivation.
6.6. The rules of nomenclature are retroactive unlessThe rules of nomenclature are retroactive unlessexpressly limited.expressly limited.
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Rules of Nomenclature:Rules of Nomenclature:
ProcedureProcedureDetailed procedures based upon these principlesDetailed procedures based upon these principles
are divided intoare divided into Rules andRecommendationsRules andRecommendations. The. TheCode states, the objective of the Rules is to putCode states, the objective of the Rules is to putthe nomenclature of the past into order and tothe nomenclature of the past into order and to
provide for that of the future; names contrary to aprovide for that of the future; names contrary to arule cannot be maintained. the Recommendationsrule cannot be maintained. the Recommendations
dealing with minor points provide guidance anddealing with minor points provide guidance and
uniformity in plants. However, names that areuniformity in plants. However, names that arecontrary to the Recommendations cannot becontrary to the Recommendations cannot be
rejected for that reason.rejected for that reason.
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Rules of Nomenclature:Rules of Nomenclature:
Ranks of TaxaR
anks of Taxa
Division Class Order Family TribeDivision Class Order Family Tribe
Genus Section Series Species Genus Section Series Species
Variety Form Variety Form
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Rules of Nomenclature:Rules of Nomenclature:
The Type MethodThe Type Method Taxonomists use the type method as a legalTaxonomists use the type method as a legal
device to provide the correct name for a taxondevice to provide the correct name for a taxon
type specimentype specimen a single specimen or the plantsa single specimen or the plants
on a single herbarium sheeton a single herbarium sheet type of a genustype of a genus a species; type of a familya species; type of a family aa
genusgenus
nomenclatural typenomenclatural type not necessarily the mostnot necessarily the most
representative of a taxon; it is the specimen orrepresentative of a taxon; it is the specimen orspecimens with which the name of the taxon isspecimens with which the name of the taxon ispermanently associated; no way reflects thepermanently associated; no way reflects thetypological concept of an idealized specimentypological concept of an idealized specimen
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Rules of Nomenclature:Rules of Nomenclature:
The Type MethodThe Type MethodSeveral kinds of types:Several kinds of types:
1.1. holotypeholotype
2.2. isotypeisotype3.3. lectotypelectotype
4.4. syntypesyntype
5.5. paratypeparatype
6.6. neotypeneotype
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Rules of Nomenclature:Rules of Nomenclature:
Priority of NamesPriority of Names PriorityPriority concerned with the precedenceconcerned with the precedence
of the date of valid publication andof the date of valid publication anddetermines the acceptance of one of two ordetermines the acceptance of one of two or
more names that are otherwise acceptablemore names that are otherwise acceptable Legitimate versus illegitimateLegitimate versus illegitimate
nomina conservandanomina conservanda conserved namesconserved names
based on the International Botanicalbased on the International BotanicalCongressCongress
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Rules of Nomenclature:Rules of Nomenclature:
Effective and Valid PublicationEffective and Valid PublicationA.A. Effective PublicationEffective Publication
distribution of printed matter to the generaldistribution of printed matter to the general
public or at least to botanical institutions withpublic or at least to botanical institutions with
B.B. Valid PublicationValid Publication
a name must be effectively published in thea name must be effectively published in the
form specified by the Codeform specified by the Code
must be accompanied by a description or amust be accompanied by a description or areference to a previously publishedreference to a previously published
description for that taxondescription for that taxon
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Rules of Nomenclature:Rules of Nomenclature:
Citation of Authors NameCitation of Authors Name To be accurate and complete, the name of aTo be accurate and complete, the name of a
taxon should include a citation of the author ortaxon should include a citation of the author or
authors who originally described that taxonauthors who originally described that taxon
Purposes:Purposes:
expedites locating the original plant descriptionexpedites locating the original plant description
distinction between two namesdistinction between two names
can aid botanists in tracing the transference ofcan aid botanists in tracing the transference ofspecies from one genus to anotherspecies from one genus to another
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Rules of Nomenclature:Rules of Nomenclature:
Retention, Choice & Rejection ofRetention, Choice & Rejection of
NamesNames1.1. A change in the diagnostic limit separating the taxonA change in the diagnostic limit separating the taxon
from its relatives is not justifiable cause for a change infrom its relatives is not justifiable cause for a change inthe name of the taxon.the name of the taxon.
2.2. When a species is described in one genus and laterWhen a species is described in one genus and latertransferred to another genus, the specific epithet, iftransferred to another genus, the specific epithet, iflegitimate, must be retained.legitimate, must be retained.
3.3. When two or more taxa of the same rank are united,When two or more taxa of the same rank are united,the oldest legitimate name or epithet is selected.the oldest legitimate name or epithet is selected.
4.4. A legitimate name or epithet must not be rejectedA legitimate name or epithet must not be rejectedmerely because it is inappropriate or disagreeable, ormerely because it is inappropriate or disagreeable, orbecause another is preferable or better known, orbecause another is preferable or better known, orbecause it has lost its original meaning.because it has lost its original meaning.
5.5. Hybrids must follow the same rules of the Code.Hybrids must follow the same rules of the Code.
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Rules of Nomenclature:Rules of Nomenclature:
Cultivated PlantsCultivated Plants Retain the names applied to the same taxa inRetain the names applied to the same taxa in
their native habitattheir native habitat
cultivarcultivar denotes an assemblage of cultivateddenotes an assemblage of cultivatedplants that is clearly distinguished by anyplants that is clearly distinguished by any
characters, and that, following reproduction,characters, and that, following reproduction,
retain its distinguishing characteristicsretain its distinguishing characteristics
International Code of Nomenclature forInternational Code of Nomenclature forCultivated PlantsCultivated Plants (Brickell, 1980)(Brickell, 1980)