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@IJAPSA-2016, All rights Reserved Page 170
CHECK-LIST OF TENEBRIONIDAE(COLEOPTERA) OF SIKKIM WITH TWO
NEW RECORDS V. D. HEGDE
Western Regional centre, Zoological Survey of India, Vidya Nagar, Ravet Road, Pune- 411 044
(Maharashtra)
ABSTRACT
The paper contains the report of 106 species of 37 genera under 18 tribes of 05 sub families
from Sikkim state and interestingly, identification of earlier collections in National Zoological
collections (NZC) in Zoological Survey of India revealed 02 species are the new reports to
theSikkim state. The Gonocephalum is the dominant genus from this state having 22
speciesfollowed by the genus Laena(14) and Strongylium (07). Further, this compilation revealed
that, in India, 30 species were reported only from Sikkim and 31 species were reported from West
Bengal and Sikkim only showing the richness of Biodiversity of this North Eastern states.
Key words: Check-list, Tenebrionidae, Sikkim, New records.
I. INTRODUCTION Occurrence of many species of the order Coleoptera are with special ecological requirements
for their continued existence and can be used as indicators of ecological conditions. The family
Tenebrionidae is the fifth largest family under the order Coleoptera and representatives of the family
are found in almost all the habitats. The representatives of this family are hard to differentiate but can
be identified by these characters. Body hard, antennal insertion hidden under frons, elytra usually
completely covering abdomen, abdomen with five visible sternites and first three segments connate,
front coxal cavities closed behind, heteromerous tarsi, tarsal segments and claws simple.
Eventhough there are several studies on the different fauna of Sikkim state (2003)including
somefamilies of Coleoptera by different research workers, no comprehensive report has beendone on
the Darkling Beetles (Coleoptera:Tenebrionidae) of this state. Hence, this compilation was
undertaken to throw some light on the Darkling beetles ofSikkim state by preparing the check-list
with two new records (marked with asterik).
II. MATERIALS AND METHODS
To make the study comprehensive, all earlier scattered reportswere also taken into
consideration and few important references were noted (Iwan, 2002, Iwan et al 2010, Masumoto and
Schawaller 2010, Schawaller, 2002, 2006, 2007 a & b, 2009, 2012, Schawaller and Ando, 2006,
2009).The Tenebrionidae collections in theNational Zoological collections, Zoological Survey of
India, Kolkata were also sorted out and identified. The classification was followed as per Bouchard
et al (2005). Theregistration numbers were also given for the materials examined.
Check-list of the species
Subfamily: Lagriinae Latreille, 1825 (1820)
Tribe: Laenini Seidlitz, 1895
Laena affinis Schuster, 1935
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Laena brahmaeMasumoto, 1990
Laena espagnoli Kaszab, 1965
Laena kangchendzongensis Masumoto,1990
Laena kaszabi Masumoto,1990
Laena laevipennis Schuster, 1926
Laena longipilisSchuster, 1926
Laena merklottoi Masumoto, 1990
Laena nishikawai Masumoto, 1990
Laena opaca Kaszab, 1970
Laena phithangensis Masumoto, 1990
Laena sakaii Masumoto, 1990
Laena singalilensis Masumoto, 1990
Laena vishnua Masumoto, 1990
Tribe: Lagriini Latreille, 1825 (1820)
Arthromacra sikkimensis Borchmann, 1916
Bothynogria bicolor (Kollar and Redtenbacher 1848)
Cerogria montana Merkl, 1991
Cerogria nepalensis (Hope, 1831) .
Cerogria rufina (Fairmaire, 1894)
Lagria comosella Fairmaire, 1894
Lagria schawalleri Merkl, 1991
Tribe:LupropiniArdoin, 1958
Luprops tristis (Fabricius, 1801)
Sub Family:Pimelinae Latreille, 1802
Tribe: Tentyriini Eschscholtz, 1831
Ascelosodis everestina Blair, 1923
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Ascelosodis nitida Blair, 1923
Ascelosodis schmidi Kaszab, 1965
Tribe: Stenosini Schaum,1859 (1834)
Dichillus indicus Wasmann, 1896
Subfamily:Tenebrioninae Latreille,1802
Tribe: Toxicini Lacordaire, 1859
Toxicum assamense Pic, 1913
Tribe: Alphitobini Reitter, 1917
Alphitobius diaperinus Panzer,1797
Alphitobius laevigatus (Fabricius, 1781)
Alphitobius piceus Olivier, 1972
Tribe : Ulomini Blanchard, 1845
Latheticus oryzae Waterhouse,1880
Uloma polita (Wiedemann, 1821)
Uloma rubripes (Hope, 1831)
Uloma ruficornis Allard, 1896
Tribe: Palorini Mathews, 2003
Palorus ratzeburgii Wissman, 1848
Tribe: Tribolini Mulsant, 1854
Tribolium castaneum (Herbst, 1797)
Tribolium confusum Jacquelin du Val, 1868
Tribe:Blaptini Leach, 1815
Balptyscellis alpinus Kaszab, 1965
Balptyscellis costulata Fairmaire, 1901
Blaps apicecostata Blair, 1922
Gnaptorina(Hesperoptorina) brucei Blair, 1923
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Gnaptorina (Hesperoptorina) sikkimensis Kaszab, 1965
Tribe: Pedinini Eschscholtz, 1829
Adamus sikkimensis (Kaszab,1975)
Platolenes sikkimensis Saha, 1988
Tribe: Opatrini Brulle, 1832
Gonocephalum alaticolle (Fairmaire, 1893)
Gonocephalum bilineatum(Walker, 1858)
Gonocephalum catenulatum (Fairmaire, 1896)
Gonocephalum civicum Kaszab, 1952
Gonocephalum depressum (Fabricius, 1801)
Gonocephalum dorsogranosum (Fairmaire, 1896)
Gonocephalum gebienianum Kaszab, 1952
Gonocephalum gracile (Bates, 1879)
Gonocephalum granulatipenne Kaszab, 1978
Gonocephalum guerryi Chatanay, 1917
Gonocephalum hauschildi Kaszab, 1952
Gonocephalum helopioide (Fairmaire, 1894)
Gonocephalum himalayense Kaszab, 1952
Gonocephalum hingstoni Kaszab, 1952
Gonocephalum hoffmannseggi (Steven, 1829)
Gonocephalum nepalicum Kaszab, 1973
Gonocephalum oblongum (Fabricius, 1801)
Gonocephalum patricium Kaszab, 1952
Gonocephalum roseni Kaszab, 1952
Gonocephalum sikkimense Kaszab, 1952
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Gonocephalum tuberculatum (Hope, 1831)
Gonocephalum vagum (Steven, 1829)
Sub Family: Diaperinae Latreille, 1802
Tribe : Diaperini Latreille, 1802
Basanus flaviventris Blair, 1937
Ceropria laticollis Fairmaire , 1903
Tribe Gnathidiini Gebien, 1921
Menimus wittmeri Kaszab, 1982
Tribe Leiochrinini Lewis, 1894
Crypsis bimaculatus Kaszab, 1946
Crypsis rufomarginatus Kaszab, 1946
Crypsis violaceipennis Waterhouse, 1877
Leiochrinus sauteri Kaszab, 1946
Derispia assamica Kaszab,1946
Derispia blairi Kaszab,1946
Derispia diaperoides Kaszab, 1946
Derispia indica Kaszab, 1946
Derispia sikkimensis Kaszab,1946
Derispia tenuipunctata Kaszab,1946
Derispia truncata Kaszab, 1961*
Derispiola blairi Kaszab, 1946
Derispiola darjeelingiana (Kaszab, 1946)
Leiochrinus metallicus Schawaller, 2005
Leiochrinus sauteri Kaszab, 1946
Leiochrodes assimilis Kaszab, 1961
Leiochrodes sikkimensis Kaszab, 1961
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Leiochrodes subaeneus Pic, 1918
Pimplema hemisphaericum (Laporte 1831)
Subfamily: Stenochiinae Kirby, 1837
Tribe: Cnodalonini Gistel, 1855
Catapiestus indicus Fairmaire, 1896
Derosphaerus asperipennis Allard, 1896
Derosphaerus rugosus Gravely, 1915
Foochounus confusus Schwaller and Ando, 2009
Promethis glabra (Hope, 1831)
Promethis harmandi (Alland, 1896)
Promethis punctatostriata (Motschulsky, 1872)
Promethis puncticollis (Motschulsky, 1872)
Promethis transversicollis (Motschulsky, 1872)
Pseudonautes viridiaeneus (Allard, 1896)
Tribe: Stenochiini Kirby, 1837
Rhopalobates villardi Fairmaire, 1897
Strongylium angusticolle Maklin, 1864
Strongylium angustissimum Pic. 1922
Strongylium dubium (Gebien)*
Strongylium mediofoveatum Blair, 1931
Strongylium metallescens Dohrn, 1880
Strongylium rufipenne L. Redtenbacher, 1844
Strongylium westermanni Maklin, 1864
Details of the species
Laena affinis Schuster, 1935
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Distribution: INDIA: Sikkim and West Bengal.
Elsewhere: Not yet recorded.
Laena brahmae Masumoto, 1990 Laena gomcheyensis Masumoto 1990 n.syn.
Distribution: INDIA: Sikkim (Phithang, Gomchey-Phukpa))
Elsewhere: Not yet recorded.
Laena espagnoli Kaszab, 1965 Distribution: INDIA: Sikkim (Donkung).
Elsewhere: Not yet recorded.
Laena kangchendzongensis Masumoto,1990 Distribution: INDIA: Sikkim (Phithang).
Elsewhere: Nepal.
Laena kaszabi Masumoto,1990 Distribution:INDIA: Sikkim ( Phithang).
Elsewhere: Nepal.
Laena laevipennis Schuster, 1926 Laena sandakphuensis Masumoto 1990 syn.
Distribution: INDIA: Sikkim (Tonglu-Phalut) and West Bengal.
Elsewhere: Not yet recorded.
Laena longipilis Schuster, 1926 Laena nepalensis(Jedlicka 1965) n.syn.
Distribution: INDIA: Sikkim (Tonglu-Phalut) and West Bengal.
Elsewhere: Nepal.
Laena merklottoiMasumoto, 1990 Distribution: INDIA: Sikkim (Tadong, Dzongri).
Elsewhere: Nepal.
Laena nishikawai Masumoto, 1990 Distribution: INDIA: Sikkim (Choka).
Elsewhere: Nepal.
Laena opaca Kaszab, 1970 Distribution:INDIA: Sikkim (Gnatong, Padamtsin to Lingtou).
Elsewhere: Nepal.
Laena phithangensis Masumoto, 1990 Laena shunichiiMasumoto 1990 n.syn.
Distribution: INDIA: Sikkim(Phithang) and West Bengal.
Elsewhere: Nepal.
Laena sakaii Masumoto, 1990 Distribution: INDIA: Sikkim (Singalia Dara) and West Bengal.
Elsewhere: Nepal.
Laena singalilensis Masumoto, 1990 Distribution: INDIA: Sikkim (Singalia) and West Bengal.
Elsewhere: Nepal.
Laena vishnua Masumoto, 1990 Laena kephakensis Masumoto 1990 n.syn.
Distribution: INDIA: Sikkim (Kephak) and West Bengal.
Elsewhere: Not yet recorded.
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Arthromacra sikkimensis Borchmann, 1916 Arthromacra sikkimensis Borchmann, 1916 : 132.
Donaciolagria sikkimensisBorchmann, 1936: 179, 180.
Donaciolagria sikkimensisMerkl 2008: 117.
Distribution:INDIA: Sikkim.
Elsewhere: Not yet recorded.
Bothynogria bicolor (Kollar and Redtenbacher 1848)
Lagria bicolor Kollar and Redtenbacher, 1848: 534; Borchmann, 1910:7 1915; 87 (as synonym of
Lagria ruficollis) 1936: 137 (as synonym of Cerogria ruficollis).
Distribution:INDIA: Sikkim, West Bengal and Uttarakhand.
Elsewhere: Nepal.
Cerogria montana Merkl, 1991 Distribution:INDIA: Sikkim and West Bengal.
Elsewhere: Nepal.
Cerogria nepalensis (Hope 1831) Distribution:INDIA: Sikkim, West Bengal and Uttarakhand.
Elsewhere: Nepal.
Cerogria rufina ( Fairmaire 1894) Distribution:INDIA: Sikkim, West Bengal,Himachal Pradesh and Tamil Nadu.
Elsewhere: Nepal.
Lagria comosella Fairmaire 1894 Distribution: INDIA: Sikkimand West Bengal.
Elsewhere: Nepal.
Lagria schawalleri Merkl, 1991 Distribution: INDIA: Sikkim (Rhenock) and West Bengal.
Elsewhere: Nepal.
Luprops tristis (Fabricius, 1801) Lagria tristis Fabricius, 1801: 70
Lyprops piceus Fairmaire, 1894: 24.Synonimised by Gebien (1941: 826 [681]).
Distribution:INDIA: Sikkim, Tamil Nadu, Kerala, Karnatakaand Andhra Pradesh.
Elsewhere: Sri Lanka.
Ascelosodis everestina Blair, 1923
Ascelosodis everestina Blair, 1923: 278: Kaszab, 1959: 361: 1965: 108. Figs. 4-5: 1968: 50: Ren and
Shi, 2006: 12-13. Figs. 55-67, 144,150.
Distribution: INDIA: Sikkim.
Elsewhere: China.
Ascelosodis nitida Blair, 1923
Ascelosodis nitida Blair, 1923: 281: Kaszab, 1959: 358; 1968: 48. Ren and Shi 2006 14-15, Figs.
81-93, 152.
Distribution:INDIA: Sikkim.
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Elsewhere: China.
Ascelosodis schmidi Kaszab, 1965
Ascelosodis schmidi Kaszab, 1965. Figs 1-3: 1968: 51: Ren and Shi, 2006: 16-17, Figs. 102-114,
145,154.
Distribution:INDIA: Sikkim.
Elsewhere: China.
Dichillus indicus Wasmann, 1896
Dichillus indicus Wasmann, 1896, Deutsche Ent. Zeitschr. P.239
Distribution:INDIA: Sikkim and West Bengal.
Elsewhere: Not yet recorded
Toxicum assamense Pic, 1913
Toxicum assamensis Pic, 1913, Mel. Ent. 6
Distribution:INDIA: Assam, Sikkim and West Bengal.
Elsewhere: Nepal.
Alphitobius diaperinusPanzer,1797
Tenebrio diaperinus Panzer 1797, Fauna Germ37: 16
Alphitobius diaperinus Oertzen, 1903, Deutschen Ent. Zeitsch : 233.
Distribution :INDIA:Assam, West Bengal and Uttarakhand. Cosmopolitan-Throughout India.
Alphitobius laevigatus(Fabricius, 1781)
Alphitobius laevigatus Fabricius, 1781, Spec. Ins. 1: 90.
Alphitobius laevigatus (Fabricius), Kaszab, Tijdschr. V. Ent. C. VII (5): 295.
Distribution:INDIA: Assam, West Bengal, Uttar Pradesh, Rajasthan, and AndamanIsland.
Cosmopolitan- Throughout India.
Elsewhere: Not yet recorded.
Alphitobius piceusOlivier, 1972
Alphitobius piceus Olivier, 1972, Encycl. Meth7: 50.
Distribution: INDIA: Assam, West Bengal and Uttarakhand.Cosmopolitan-Throughout India.
Elsewhere: Cosmopolitan.
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Latheticus oryzae Waterhouse,1880
Latheticus oryzae Waterhouse,1880, Ann. Mag. Nat. Hist. 5(V) : 148
Kaszab, 1972, Alti. Soc. It. Sc. Nat.c. Museo. Civ. St. Nat. Milono. 113 (4) : 366-384.
Distribution:INDIA: West Bengal, Cosmopolitan- Throughout India.
Elsewhere: Cosmopolitan.
Uloma polita (Wiedemann, 1821) Uloma polita Wiedemann, Germ. Mag. IV, 1821, p. 149.
Distribution:INDIA:Sikkim (Tadung), Assam, West Bengal and Uttarakhand
Elsewhere: Not yet recorded.
Uloma rubripes (Hope, 1831) Uloma prehimalayana Kaszab, 1975, Syn. n.
Uloma rubripes Hope, 1831, Misc.1: 31.
Distribution:INDIA: Sikkim (Rongay), Uttarakhand,Assam, Meghalaya and West Bengal.
Uloma ruficornis Allard, 1896
Uloma kulzeri Ardoin, 1960, syn. n.
Distribution:INDIA: Sikkim and West Bengal.
Elsewhere: Nepal.
Palorus ratzeburgii Wissman, 1848
Hypophloeus ratzeburgii Wissmam, 1848, Steltein ent. Zeitschr.9: 77
Distribution:INDIA: West Bengal.Cosmopolitan-Throughout India.
Elsewhere: Cosmopolitan.
Tribolium castaneum (Herbst, 1797)
Colydium castaneum (Herbst) 1797 Nature kafer 7: 2
Tribolium castaneum Macleay, 1825 Annulora javanica: 50
Distribution:INDIA: West Bengal. Cosmopolitan- Throughout India.
Elsewhere: Cosmopolitan.
Tribolium confusum Jacquelin du Val, 1868
Tribolium confusum Jacquelin du Val, 1868, Gen. col d’. Eur. Cat. P. 181.
Distribution:INDIA: West Bengal, Cosmopolitan- Throughout India.
Elsewhere: Cosmopolitan.
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Balptyscellis alpinus Kaszab, 1965 Balptyscellis alpinus Kaszab, 1965, Misc. Zool. Barcelona. 2(1).
Distribution:INDIA:Sikkimand West Bengal.
Elsewhere: Not yet recorded.
Balptyscellis costulata Fairmaire, 1901 Distribution:INDIA: Sikkim (Gnatong)
Elsewhere: Not yet recorded.
Blaps apicecostata Blair, 1922
Distribution: INDIA: Sikkim (Tungu).
Elsewhere: Tibet, Nepal.
Gnaptorina(Hesperoptorina)brucei Blair, 1923
Distribution:INDIA: Sikkim.
Elsewhere: Nepal and China.
Gnaptorina (Hesperoptorina) sikkimensis Kaszab, 1965
Gnaptorina (Hesperoptorina) sikkimensis Kaszab, 1965 :110
Distribution: INDIA: Sikkim (Donkung).
Elsewhere: Not yet recorded.
Adamus sikkimensis (Kaszab,1975)
Platydendarus sikkimensis Kaszab, 1975b: 315.
Adamus sikkimensis: Iwan 1997a: 257, 2002: 50.
Distributio:INDIA: Sikkim.
Elsewhere: Not yet recorded.
Platolenes sikkimensisSaha, 1988
Platolenes sikkimensis Saha, 1988, Rec. Zool. Surv. India, 85(3): 433- 437.
Material examined: 1 ex. Sikkim, F.H. Gravely coll., 14.vi.1916 Reg. no. 10769/H4A, ZSI, Head
Quarters, Kolkata.
Distribution:INDIA: Sikkim.
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Remark: Described earlier by Saha (1988) from Sikkim and the present material was collected in the
year 1916.
Gonocephalum alaticolle (Fairmaire, 1893) Bradymerus alaticolle Fairmaire, 1893: 23. – Chatanay 1917: 245.
Gonocephalum alaticolle (Fairmaire, 1893). – Gebien 1939: 448; Kaszab 1952a: 475, 1964: 235;
Kaszab and Chûjô 1964: 235; Iwanand Löbl 2008: 262.
Distribution:INDIA: Sikkimand West Bengal.
Elsewhere: Cambodia.
Gonocephalum bilineatum(Walker, 1858) Opatrum bilineatum Walker, 1858: 284.
Hopatrum bilineatum Walker, 1858. – Gemminger 1870: 1931.
Gonocephalum bilineatum (Walker, 1858). – Gebien 1910b: 322,1921: 269, 1939: 446; Chatanay
1917: 238; Blair 1921: 269; Schuster1928: 989; Reichardt 1936: 109; Kaszab 1952a: 641, 1954: 75,
1965a:114, 1965d: 287, 1970a: 424, 1977: 257, 1979a: 260, 1979b: 65, 1980b:171; Boddy 1965:
171; Kwon and Choi 1986: 106; Schawaller 1997a:3; Hua Li 2002: 138; Iwan and Löbl 2008: 263.
Gonocephalum KamtschaticumMotschulsky, 1860: 139.
Hopatrum kamtschaticum Motschulsky, 1860. – Gemminger 1870:1932.
Gonocephalum kamtschaticum Motschulsky, 1860. – Reitter 1904:146; Gebien 1910b: 323; Schuster
1928: 989; Reichardt 1936: 109;Kaszab 1952a: 682, 1965d: 287.
Blapstinus latifrons LeConte, 1874: 70. – Casey 1890: 393;
Gonocephalum latifrons (LeConte, 1874). – Champion 1895: 115;Gebien 1910b: 323, 1939: 452;
Kaszab 1952a: 682; Ferrer 2000: 129.
Opatrum orarium Lewis, 1894: 380. – Champion 1895: 117.
Gonocephalum orarium (Lewis, 1894). – Gebien 1910b: 324;Reichardt 1936: 109; Schuster 1928:
989; Kaszab 1952a: 682.
Distribution:INDIA: Sikkim, West Bengal, Andaman Islands, Uttarakhand and Uttar Pradesh.
Elsewhere:Bhutan, Borneo, Indonesia, Malaysia, Philippines, Sri Lanka, Nepal, Japan and Korea.
Gonocephalum catenulatum (Fairmaire, 1896) Hopatrum catenulatum Fairmaire, 1896: 19.
Gonocephalum catenulatum (Fairmaire, 1896). – Gebien 1910b: 322,1939: 447; Kaszab 1952a: 463;
Iwan and Löbl 2008: 263.
Distribution:INDIA:Sikkim, West Bengal,Karnataka and Maharashtra
Elsewhere: Not yet recorded.
Gonocephalum civicum Kaszab, 1952 Gonocephalum civicum Kaszab, 1952a: 612. – Kaszab 1970a: 423,1977: 257; Schawaller 1997a: 4;
Iwan and Löbl 2008: 263.
Distribution:INDIA: Sikkim, West Bengal,Jharkand, Maharashtra Karnataka and Uttarakhand.
Elsewhere: Nepal, Myanmar, Nepal, Thailand.
Gonocephalum depressum (Fabricius, 1801) Opatrum depressum Fabricius, 1801: 117. – Steven 1829: 95.
Hopatrum depressum Fabricius, 1801- Gemminger 1870: 1931; Fairmaire 1893: 20, 1894: 17.
Gonocephalum depressum (Fabricius, 1801). – Gebien 1906: 213;1910b: 323, 1913: 376, 1939: 447;
Gravely 1913: 520; Blair 1922: 292; Kaszab 1952a: 635, 1974a: 193, 1979a: 260, 1979b: 65;
Kwieton1981: 17; Schawaller 1997a: 5; Hua Li 2002: 138; Iwan and Löbl2008: 263.
Opatrum contrahens Walker, 1858: 284. – Blair 1921: 269.
Hopatrum contrahens Walker, 1858. – Gemminger 1870: 1931.
Gonocephalum contrahens (Walker, 1858). – Gebien 1910b: 322,1939: 447; Kaszab 1952a: 681.
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Distribution: INDIA: Sikkim, Arunachal Pradesh, Assam,Manipur,Meghalaya,West Bengal,
Himachal Pradesh and Kashmir.
Elsewhere:Bhutan,Nepal, Pakistan, Indonesia, Laos, Philippines, Sri Lanka, Myanmar, Taiwan,
Afghanistan and China.
Gonocephalum dorsogranosum (Fairmaire, 1896) Hopatrum dorsogranosum Fairmaire, 1896: 14.
Gonocephalum dorsogranosum (Fairmaire, 1896). – Gebien 1910b:323, 1939: 448; Kaszab 1940:
149, 1952a: 627, 1960a: 149, 1968b: 20,1970c: 152, 1974a: 193; Kwieton 1981: 17; Iwan and Löbl
2008: 263.
Gonocephalum foveolatus Kaszab, 1963: 343.
Distribution:INDIA: Sikkim, West Bengal, Tamil Nadu, Maharashtra and Uttarakhand.
Elsewhere: Pakistan and Myanmar.
Gonocephalum gebienianum Kaszab, 1952 Gonocephalum Gebienianum Kaszab, 1952a: 470. – Kaszab 1979a:260.
Gonocephalum gebienianum Kaszab, 1952a. – Iwan and Löbl 2008:263.
Distribution:INDIA: Sikkimand West Bengal
Elsewhere: Indonesia.
Gonocephalum gracile (Bates, 1879) Penthicus gracilis Bates, 1879: 482. – Bates 1890: 19.
Loboderus gracile Bates, 1879: 482. – Bates 1890: 75.
Lobodera gracile (Bates, 1879). – Reichardt 1936: 169.
Gonocephalum gracile (Bates, 1879). – Gebien 1939: 447; Kaszab1952a: 546, 1961b: 351, 1977:
257, 1978b: 230; Schawaller andSkopin 1978: 230; Schawaller 1997a: 6; Hua Li 2002: 138; Iwan
and Löbl 2008: 263.
Gonocephalum ruficorne Gravely, 1915: 250. – Kaszab 1952a:676.
Gonocephalum curvicolle Gridelli, 1934: 66. – Kaszab 1952a: 676;hom. nov. nom. preoccup., nec
Gonocephalum curvicolle Reitter,1889: 705.
Distribution:INDIA: Sikkim, West Bengal, Himachal Pradesh and Kashmir.
Elsewhere: Nepal and China.
Gonocephalum granulatipenne Kaszab, 1978 Gonocephalum granulatipenne Kaszab, 1978: 173. – Iwan and Löbl2008: 263.
Distribution:INDIA: Sikkimand West Bengal.
Elsewhere: Bhutan.
Gonocephalum guerryi Chatanay, 1917 Gonocephalum Guerryi Chatanay, 1917: 244. – Schuster 1928: 989;Gebien 1939: 447; Gridelli
1954: 282; Kaszab 1952a: 548, 1960a: 146.
Gonocephalum guerryi Chatanay, 1917. – Kaszab 1968b: 20, 1970c:148; Hua Li 2002: 138; Iwan
and Löbl 2008: 264.
Gonocephalum Potanini Reichardt, 1936: 108, 209. – Gebien 1939: 446.
Gonocephalum potanini Reichardt, 1936. – Reichardt 1936: 108;Kaszab 1952a: 676.
Distribution:INDIA: Sikkimand West Bengal.
Elsewhere: Myanmar, China, Hongkong, Hubei, Sichuan, Yunnan, Pakistan and Afghanistan.
Gonocephalum hauschildi Kaszab, 1952 Gonocephalum Hauschildi Kaszab, 1952a: 498. – Kaszab 1965a: 113.
Gonocephalum hauschildi Kaszab, 1952a. – Schawaller 1997a: 6;Iwan and Löbl 2008: 264.
Distribution:INDIA: Sikkim, West Bengal and Uttarakhand.
Elsewhere: Thailand, China and Nepal.
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Gonocephalum helopioide (Fairmaire, 1894) Hopatrum helopioide Fairmaire, 1894: 19.
Opatrum helopioides (Fairmaire, 1894): 19. – Champion 1895: 116.
Gonocephalum helopioide (Fairmaire, 1894). – Gebien 1910b: 323,1939: 448; Kaszab 1952a: 574;
Schawaller 1997a: 7; Iwan and Löbl2008: 264.
Distribution:INDIA: Sikkim, West Bengaland Uttar Pradesh.
Elsewhere: Myanmar and Nepal.
Gonocephalum himalayense Kaszab, 1952 Gonocephalum himalayense Kaszab, 1952a: 528. – Schawaller1997a: 8; Iwan and Löbl 2008: 264.
Distribution: INDIA: Sikkim and West Bengal.
Elsewhere:Bhutan and Nepal.
Gonocephalum hingstoni Kaszab, 1952 Gonocephalum Hingstoni Kaszab, 1952a: 524.
Gonocephalum hingstoni Kaszab, 1952. – Iwan and Löbl 2008: 264.
Distribution:INDIA: Sikkim, West Bengal andUttar Pradesh.
Elsewhere: Not yet recorded.
Gonocephalum hoffmannseggi (Steven, 1829) Opatrum Hoffmannseggi Steven, 1829: 96.
Hopatrum HoffmannseggiSteven, 1829. – Gemminger 1870: 1932.
Gonocephalum Hoffmannseggi(Steven, 1829). – Gebien 1910b: 323,1939: 448; Kaszab 1952a: 666,
1970a: 424, 1979a: 260.
Gonocephalum hoffmannseggii(Steven, 1829). – Kaszab 1979b: 67;Schawaller 1997a: 8; Iwan and
Löbl 2008: 264.
Distribution: INDIA: Sikkim and West Bengal.
Elsewhere: Bhutan, Myanmar, Indonesia, Philipines, Sri Lanka, Nepal, Taiwan and New Guinea.
Gonocephalum nepalicum Kaszab, 1973 Gonocephalum nepalicum Kaszab, 1973: 245. – Kaszab 1977: 256;Schawaller 1997a: 9; Bremer and
Ferrer 2002: 223; Iwan and Löbl2008: 264.
Distribution: INDIA: Sikkimand West Bengal.
Elsewhere:Bhutan and Nepal.
Gonocephalum oblongum (Fabricius, 1801) Opatrum oblongum Fabricius, 1801: 117.
Hopatrum oblongum Fabricius, 1801. – Gemminger 1870: 1932.
Gonocephalum oblongum (Fabricius, 1801). – Gebien 1906: 215,1910b: 324, 1939: 447; Kaszab
1952a: 459, 1979a: 260; Bremer and Ferrer 2002: 223; Iwan and Löbl 2008: 264.
Distribution:INDIA: Sikkimand West Bengal.
Elsewhere: Nepaland Myanmar.
Gonocephalum patricium Kaszab, 1952 Gonocephalum patricium Kaszab, 1952a: 604. – Iwan and Löbl 2008: 264.
Distribution: INDIA: Sikkim, West Bengal and Uttarakhand.
Elsewhere: Pakistan and Myanmar.
Gonocephalum roseni Kaszab, 1952 Gonocephalum Roseni Kaszab, 1952a: 546. – Kaszab 1977: 257; Schawaller 1997a: 10.
Gonocephalum roseni Kaszab, 1952a. – Kaszab 1965a: 113, 1970a:423; Iwan and Löbl 2008: 265.
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Distribution:INDIA: Sikkim,Arunachal Pradesh,West Bengal and Uttarakhand.
Elsewhere: Bhutanand Nepal.
Gonocephalum sikkimense Kaszab, 1952 Gonocephalum sikkimense Kaszab, 1952a: 517. – Iwan and Löbl2008: 266.
Distribution:INDIA: Sikkim and West Bengal.
Elsewhere: Not yet recorded.
Gonocephalum tuberculatum (Hope, 1831) Opatrum tuberculatum Hope, 1831: 31.
Gonocephalum tuberculatum(Hope, 1831). – Chatanay 1917: 243;Miwa 1931: 151; Gridelli 1934:
67; Reichardt 1936: 108; Gebien1939: 446; Kaszab 1941a: 51, 1952a: 464, 1960a: 139, 1961d:
236,1968b : 19, 1970a: 423, 1970c: 145, 1973b: 55, 1974a: 193, 1980b: 171;Gridelli 1954: 283;
Nakane 1966: 243; Schawaller 1997a: 13; Hua Li 2002: 139; Iwan and Löbl 2008: 266.
Hopatrum elongatumGuérin-Méneville, 1834. – Fairmaire 1894: 17;Gemminger, 1870: 1931.
Opatrum elongatumGuérin-Méneville, 1834: 32.
Gonocephalum elongatum(Guérin-Méneville, 1834) – Gebien 1910b:323, 1913: 4; Schuster 1928:
989; Reichardt 1936: 108; Kaszab 1952a: 672.
Gonocephalum quadrinodosum Reitter, 1904: 146. – Gebien 1910b: 325, 1939: 446; Schuster 1928:
989 (= Opatrum elongatum Guérin-Méneville, 1834); Reichardt 1936: 108 (= Opatrum elongatum
Guérin-Méneville, 1834); Kaszab 1952a: 672.
Distribution: INDIA:Sikkim, West Bengal, Kashmir, Himachal Pradesh and Uttarakhand.
Elsewhere: Nepal, Pakistan, Taiwan, Bangladesh, Afghanistan,Philippines, Myanmar, Vietnam,
China,Hainan, Hongkong, Hubei, Sichuan and Yunnan.
Gonocephalum vagum (Steven, 1829) Opatrum vagum Steven, 1829: 96.
Hopatrum vagum Steven, 1829. – Gemminger 1870: 1934.
Gonocephalum vagum (Steven, 1829). – Gebien 1910b: 327, 1939: 448;Kaszab 1952a: 662, 1979b:
66; Ferrer 1995a: 59; Hua Li 2002: 139; Iwan and Löbl 2008: 266.
Hopatrum arenarium Fairmaire, 1894: 17. – Blair 1922: 269. hom. nov. nom. preoccup., nec
Opatrum arenarium Fabricius, 1775.
Gonocephalum arenarium (Fairmaire, 1894). – Kaszab 1952a: 684;Ferrer 1995a: 59 (syn.).
Opatrum planatum Walker, 1858: 284.
Hopatrum planatum Walker, 1858. – Gemminger 1870: 1933.
Gonocephalum planatum (Walker, 1858). – Gebien 1910b: 324, 1939:448 (= Opatrum moluccanum
Blanchard, 1853); Blair 1922: 292; Kaszab 1952a: 684; Ferrer 1995a: 59 (syn.).
Distribution:INDIA: Sikkimand West Bengal
Elsewhere: China, Yunnan, Sri Lanka and Nepal.
Basanus flaviventris Blair, 1937 Distribution: INDIA: Sikkim (Rangeli River), West Bengal and Meghalaya.
Elsewhere: Laos and Vietnam.
Ceropria laticollis Fairmaire, 1903 Ceropria laticollis Fairmaire, 1903, Ann. Soc. ent. Belg.,47p. 13.
Distribution: INDIA: Sikkim and West Bengal.
Elsewhere: Thailand, Myanmar, China, Japan, Tonkin and Aman.
Menimus wittmeri Kaszab, 1982
Distribution: INDIA: Sikkim (Dzongori).
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Elsewhere: Not yet recorded.
Crypsis bimaculatus Kaszab, 1946 Distribution:INDIA: Sikkim and Meghalaya.
Elsewhere: Nepal.
Crypsis rufomarginatus Kaszab, 1946 Crypsis migliaceioiFerrer, 1983 syn.
Distribution: INDIA: Sikkim and Himachal Pradesh.
Elsewhere: Bhutan and Nepal.
Crypsis violaceipennis Waterhouse, 1877 Crypsis violaceipennis Waterhouse,1.e. p.74.
Distribution: INDIA:Sikkim, Meghalaya,West Bengal and Uttarakhand.
Elsewhere: Laos and Nepal.
Leiochrinus sauteri Kaszab, 1946
Distribution:INDIA: Sikkimand Uttarakhand.
Elsewhere: Bangladesh,Nepal, Indochina and Taiwan.
Derispia assamica Kaszab,1946
Distribution: INDIA: Sikkim.
Elsewhere: Not yet recorded.
Derispia blairiKaszab,1946
Distribution: INDIA: Sikkim.
Elsewhere: Nepal, Indochina and Yunnan.
Diaperoides Kaszab, 1946 Distribution: INDIA: Sikkim and West Bengal.
Elsewhere: Nepal.
Derispia indica Kaszab, 1946 Distribution: INDIA: Sikkim.
Elsewhere: Myanmar.
Derispia sikkimensis Kaszab,1946
Distribution: INDIA: Sikkim.
Elsewhere: Not yet recorded.
Derispia tenuipunctata Kaszab,1946
Distribution: INDIA: Sikkim.
Elsewhere: Not yet recorded.
Derispia truncataKaszab, 1961* Material examined: 3 ex., Magnan, Sikkim, G. Ramakrishna Coll. 08.v.1962, Reg. no. 6841-
6843/H4A, ZSI, Head Quarters, Kolkata.
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Distribution: INDIA: Sikkim (Magnan) and Assam.
Elsewhere: Nepal
Remark: This species is recorded for the first time from Sikkim.
Derispiola blairi Kaszab, 1946 Material examined: 1 ex., Sikkim, A.K.K. Menon Coll. 07.v.1959, Reg. no. 10650/H4A, ZSI, Head
Quarters, Kolkata.
Distribution: INDIA: Sikkim.
Elsewhere: Nepal and Indochina.
Remark: Reported earlier by Schawaller (1992,2005) also.
Derispiola darjeelingiana (Kaszab, 1946) Material examined: 4 ex., Lingship, Sikkim, G.Ramakrishna Coll. 18.v.1962, Reg. no. 6846-
6849/H4A, ZSI, Head Quarters, Kolkata.
Distribution: INDIA: Sikkim (Lingship) and West Bengal.
Elsewhe: Nepal.
Remark: Reported earlier by Schawaller (1992) also.
Leiochrinus metallicus Schawaller, 2005 Distribution: INDIA: Sikkim (Pelling).
Elsewhere: Not yet recorded.
Leiochrinus sauteri Kaszab, 1946
Leiochrinus sauteri Kaszab, 1946, Mon. p.123, 132 ft. 204,216.
Distribution: INDIA: Assam, Sikkim and West Bengal.
Elsewhere: Nepal, Indochina and Taiwan.
Leiochrodes assimilis Kaszab, 1961 Distribution: INDIA: Sikkim.
Elsewhere: Nepal.
Leiochrodes sikkimensisKaszab, 1961 Distribution:INDIA: Sikkim and West Bengal.
Elsewhere: Nepal.
Leiochrodes subaeneusPic, 1918
Distribution: INDIA: Sikkim.
Elsewhere: Banguey, Borneo, Java and Thailand.
Pimplema hemisphaericum (Laporte 1831)
Distribution: INDIA: Sikkim and Assam.
Elsewhere: Sumatra, Borneo, Java, New Guinea, Luzon, Myanmar and Nepal.
Catapiestus indicus Fairmaire, 1896
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Catapiestus indicus Fairmaire,1896,Ann. Soc. Ent. Belg. 40, p.28
Distribution:INDIA: Sikkim and Karnataka.
Elsewhere: Nepal.
Derosphaerus asperipennis Allard, 1896 Derosphaerus asperipennis Allard, Bull. Soc. Ent. Fr. 1896, p.320
Distribution: INDIA: Sikkim, Assam, West Bengal andUttarakhand.
Elsewhere: Bhutan,Nepal and Tibet.
Derosphaerus rugosusGravely, 1915 Distribution: INDIA: Assam, Sikkim West Bengal and Meghalaya.
Elsewhere: Nepaland Aborland.
Foochounus confususSchwaller and Ando, 2009 Distribution: INDIA: Sikkim (Lagyap).
Elsewhere: Bhutan.
Promethis glabra (Hope, 1831) Comb.n. Tenebrio glaber Hope, 1831: Zool. Misc.: 31 (Locus typicus: Nepaul).
Nyctobates parvicollis Fairmaire, 1894: Annls. Soc. Ent. Fr. 38:37 (Locus typicus: Kurseing), Syn. n.
Distribution:INDIA: Sikkim and West Bengal.
Elsewhere: Nepal, Bhutan and Myanmar.
Promethis harmandi (Alland, 1896)
Promethis harmandi Alland, 1896: Bull. Soc. Ent. Fr. 1896: 378 (Locus typicus: Sikkim)
Seteris crenatostriata Kaszab, 1965, nec Motschulsky, 1872: Misc. Zool. 2(1) : 123
Seteris crenatostriata Kaszab, 1970, nec Motschulsky, 1872: Ent. Basiliania. 1: 317.
Distribution: INDIA: Sikkim, Punjab, Assam and West Bengal
Elsewhere: China, Bhutan, Nepal, Myanmar, Thailand & Pakistan.
Promethis punctatostriata (Motschulsky, 1872) Comb.n.
Seteris punctatostriata Motschulsky, 1872: Bull. Soc. Imp. Nat. Mosc. 45 (3): 29
Nyctobates indosinicus Fairmaire, 1893:Annls. Soc. Ent. Belg.37:296
Seteris indosinica Gravely, 1915: Rec. Indian Mus. 8: 526
Seteris foveiceps Pic. 1929: Melang. exot. ent. 53:17
Seteris pilipes Pic, 1929: Melang. exot. ent. 53:17
Seteris indosinica Ardoin, 1969: Bull. Soc. Ent. Fr. 1969 :74
Seteris indosinica Kaszab, 1970: Ent. Basiliensia 1:307
Seteris indosinica Kaszab, 1980: Annls Hist. nat. Mus. natn. Hung.72:177
Distribution:INDIA: Sikkim, Assam and West Bengal.
Elsewhere: Laos, China, Nepal, Korea, Myanmar, Kambodscha, Vietnam, Thailand and Malay
Penin.
Promethis puncticollis (Motschulsky, 1872) Comb.n.
Iphthinus calcaratus Dejean, 1837: Catalogue de coleopteres, Paris, ed. 3: 225
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Seteris puncticollis Motschulsky, 1872: Bull. Soc. Imp. Nat. Mosc. 45 (3) : 29
Seteris nanillarum Gebien,1913, nec. Fairmaire, 1886: Philipp. J. Sci. 8 (56) D: 402
Seteris penicilligora Gebien,1914, nec. Gebien,1911: Sarawak, Mus. J. 2(5): 38
Seteris manillarum Gebien, 1914, nec. Fairmaire, 1886 : Suppl. Ent. 15 : 36
Seteris manillarum Gebien, 1921, nec. Fairmaire, 1886 : Philipp. J. Sci.19 (4) : 456, Taf.1, Abb. 6.
Nyctobates brevicornis Deyrolle in litt.
Nyctobates anthracina Deyrolle in litt.
Nyctobates unisulcata Fairmaire in litt.
Seteris rugulicollis Fairmaire in litt.
Distribution: INDIA: East- India Sikkim.
Elsewhere: China, Malay Penin, Singapore, Sri Lanka, Nios, Sumatra, Java, Bali, Lombok,
Sumbawa, Flores, Karimon, Ceram, Borneo, Banguey, Philippines and Sulu.
Promethis transversicollis (Motschulsky, 1872) Comb.n.
Seteris transversicollis Motschulsky ,1872: Bull. Soc. Imp. Nat. Mosc.45 (3): 29
Nyctobates (Seteris?) laevis Fairmaire ,1896: Notes Leyden Mus. 18: 230
Seteris laevis Gravely, 1915: Rec. Indian Mus. 8 (8): 526
Seteris laevis Kaszab, 1970: Ent. Basiliensia 1:307
Distribution:INDIA: Sikkim, Assam and North India- Uttar Pradesh.
Elsewhere: China, Myanmar, Vietnam, Thailand, Laos, Nepal, Bhutan and Korea.
Pseudonautes viridiaeneus(Allard, 1896)
Distribution: INDIA: Sikkimand West Bengal.
Elsewhere: Nepal.
Rhopalobates villardi Fairmaire, 1897
Rhopalobates villardi Fairmaire, 1897, 1 c. p.231.
Encyalesthus klapperichiKaszab, 1954.
Distribution:INDIA: Sikkim (Villardi).
Elsewhere: Indochinaand China.
Strongylium angusticolle Maklin, 1864
Strongylium angusticolleMaklin, 1864 Monographie: 333 (Bangladesh, sylhet) t.2, f.20.
Distribution: INDIA: Sikkim, Meghalaya, Assam, Uttarakhand and West Bengal.
Elsewhere: Yunnan,Nepaland Bangladesh.
Strongylium angustissimum Pic. 1922
Strongylium angustissimum Pic, 1922, Mel.exo.ent., 37:27 (Yunnan).
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Distribution:INDIA: Sikkim, Assam, West Bengal, Meghalaya and Uttar Pradesh.
Elsewhere: Yunnan and Nepal.
Strongylium dubium (Gebien)*
Material examined: 1 ex.,Sikkim, (Coll. Name, Not available) 16.vi.1912, Reg. no. 10642 /H4A,
ZSI, Head Quarters, Kolkata.
Distribution:INDIA: Sikkimand West Bengal.
Remark: This species is recorded for the first time from Sikkim and the collection of this
material was in 1912. (more than 100 years ago).
Strongylium mediofoveatum Blair, 1931
Strongylium mediofoveatum Blair, 1931, The Ent. Mon. Mag.,67: 200 (Assam, Patkai Mts.)
Distribution:INDIA: Assam, Sikkim and West Bengal.
Elsewhere: Nepal.
Strongylium metallescens Dohrn, 1880
Strongylium metallescens Dohrn, 1880, Stettin. Ent. Zeit., 41p. 377 (Darjeeling).
Distribution:INDIA:Sikkim(Rongay) and West Bengal.
Elsewhere: Nepal.
Strongylium rufipenne L. Redtenbacher, 1844
Strongylium rufipenne L.Redtenbacher, 1844, Kashmir und Reich der. Siek. Vieter Band. Zweite
Abtheilung: 533. (Kashmir).
Distributrion: INDIA: Sikkim, Uttarakhand and West Bengal.
Elsewhere: Nepal.
Strongylium westermanni Maklin, 1864
Strongylium westermanni Maklin, 1864, Monographie: 341 (449). (INDIA orientali).
Strongylium cariosipenne Fairmaire, 1896, Not. Leyden Mus., 18: 112. (Pedong) {Syn. Nov.}
Distribution: INDIA: Sikkim and Uttarakhand.
Elsewhere: Nepal.
IV. ACKNOWLEDGEMENTS I am very much thankful to Director, Zoological Survey of India, Kolkata for the facilities.
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