Checklist of the Land Snails and Slugs

of California Barry Roth and Patricia S. Sadeghian

SANTA BARBARA M U S E U M OF N A T U R A L HISTORY

Contributions in Science Number 3

Checklist of the Land Snails and Slugs of California

Barry Roth and Patricia S. Sadeghian

SANTA BARBARA M U S E U M OF N A T U R A L HISTORY

2003

Checklist of the Land Snails and Slugs of California

Barry Roth and Patricia S. Sadeghian

Santa Barbara Museum of Natural History Contributions in Science - Number 3

© 2003 Santa Barbara Museum of Natural History

2559 Puesta del Sol Road

Santa Barbara, California 93105-2936 USA

http://www.sbnature.org

Original Date of Publication: 6 June 2003

Editor: Paul Valentich-Scott

Digital photography by Patricia S. Sadeghian

Digital image editing by Patricia S. Sadeghian and Paul Valentich-Scott

Layout by Marie Murphy and Paul Valentich-Scott

Spine photo: Xerarionta (Xerarionta) intercisa, photo by F. G. Hochberg

Front cover photo: Micraronta (Micrarionta) rufocincta, photo by F. G. Hochberg

Title page art: Haplotrema (Haplotrema) catalinense, illustration by Jamie Calhoun

This publication may be purchased at: www.sbnature.org/estore

Library of Congress Cataloging-in-Publication Data

Roth, Barry. Checklist of the land snails and slugs of California / Barry Roth and Patricia S. Sadeghian. p. cm — (Santa Barbara Museum of Natural History contributions in science ; no. 3) ISBN 0-936494-34-4 (pbk.) 1. Snai ls—Cali fornia—Classif icat ion. 2. Slugs (Moi lusks)—Cali fornia—Classif icat ion.

I. Roth, Barry. II Sadeghian, Patricia S., 1975- III. Title. IV. Series. QL430.4.R68 2003 594' .38'09794—dc21 2003008847

© 2003, All rights reserved.

This book may not be reproduced in whole or in part for any purpose whatever, without written permission from the publisher (Santa Barbara Museum of Natural History).

Printed and bound by Kinko's, Santa Barbara, Califorina

Xerarionta (Plesarionta) tryoni (Newcomb, 1864) Santa Barbara and San Nicolas Islands. Xerarionta (Plesarionta) tryoni tryoni (Newcomb, 1864)

Santa Barbara and San Nicolas Islands.

Type material: Helix tryoni Newcomb, 1864, syntypes CU 27858. Helix tryonii var. albida Hemphill, 1891, syntypes AMNH 52540 (4), CAS 032758 (4), CAS 037698 (2), SBMNH 33952 (7), UCM 20678 (13), USNM 174388-174394 (14). Helix tryonii var. californica Hemphill, 1891, syntypes AMNH 54521 (4), ANSP 10828 (1), ANSP 86799 (3), ANSP 86800 (2), ANSP 86801 (2), ANSP 86803 (2), CAS 037692 (5), SBMNH 33942 (9), UCM 20679, USNM 174375-174377 (6). Helix tryonii vav.fasciata Hemphill, 1891, syntypes AMNH 52042 (2), AMNH 52542 (2), CAS 032737 (6), SBMNH 33953 (9), UCM 20681 (13), USNM 174369-174574 (12). Helix tryonii war. nebulosa Hemphill, 1891, syntypes AMNH 51961 (2), AMNH 52543 (2), AMNH 52573 (2), CAS 032735 (6), CAS 037695 (6), SBMNH 33931 (10), USNM 174354-174359 (12). Helix tryonii var. varius Hemphill, 1891, syntypes ANMH 51802 (2), CAS 037694 (6), UCM 20686 (12), USNM 174360-174365 (12). Helix tryonii color var. maculata Hemphill, 1901, syntypes UCM 20685 (4), USNM 174378-174383 (12).

Xerarionta (Plesarionta) tryoni hemphilli (Edson & Hannibal, 1911) Extinct. Late Pleistocene and early Holocene: Santa Barbara and San Nicolas Islands.

Type material: Helix tryonii var. subcarinata Hemphill, 1891, non H. subcarinata Montagu, 1803 (renamed Epiphragmophora kelletti hemphilli Edson & Hannibal, 1911), syntypes CAS 037717 (4), CAS 058812 (5), SBMNH 33924 (33), USNM 174320-174324 (10) (Coan & Roth, 1987). Helix tryonii var. major Hemphill, 1901, syntypes CAS 058873 (2), SBMNH 33923 (16), USNM 174297-174299 (6), USNM 174308-174310 (6). Helix tryonii var. minor Hemphill, 1901, syntypes CAS 037701 (5), CAS 037710 (5), CAS 058819 (8), USNM 174302-174305 (8). Micrarionta (Xerarionta) tryoni carinata Pilsbry, 1939, ex Hemphill ms, holotype ANSP 86804a.

Subgenus Xerarionta, sensu stricto Xerarionta (Xerarionta) agnesae (Kanakoff, 1950)

Extinct. Late Pleistocene: San Clemente Island.

Type material: Micrarionta (Xerarionta) agnesae Kanakoff, 1950, holotype LACMIP 1088.

Xerarionta (Xerarionta) intercisa (W. G. Binney, 1857) San Clemente Island.

Type material: Helix intercisa W. G. Binney, 1857, holotype AMNH 57847. Helix crebri-striata*NQVicomb, 1864, syntypes CU 26377 (3). Helix intercisa var. albida Hemphill, 1891, syntypes CAS 058817 (7), possible syntypes AMNH 52117 (2), ANSP 10799 (3), CAS 037578 (2), CAS 037840 (2), USNM 17540 (2). Helix intercisa var. elegans Hemphill, 1891, syntypes CAS 058883 (4), possible syntypes AMNH 52539 (2), ANSP 10807 (3), CAS 037569 (2), CAS 037572 (7). Helix intercisa war. minor Hemphill, 1891, syntypes AMNH 51963 (2), ANSP 10805 (3), CAS 037341 (6), CAS 037626 (2), USNM 174524-174525 (4). Helix intercisa var. nepos Hemphill, 1891, syntypes AMNH 51962 (2), ANSP 86539 (2), CAS 037571 (2), CAS 037611 (4), CAS 058835

Land Snails and Slugs of California 19