Records of the Western Australian Museum Supplement No. 59: 83-95 (2000).

FISHES OF CHRISTMAS ISLAND, INDIAN OCEAN

Gerald R. AlIen

Western Australian Museum, Francis Street, Perth, Western Australia 6000, Australia

The first fish collections at Christmas Island werethose of CW. Andrews (British Museum) in 1897and 1908, CA. Gibson-Hill (Raffles Museum,Singapore) in 1938-40, and E.J. Car and G.F. Mees(Western Australian Museum) in 1961. As a resultof their work 127 fish species were recorded fromthe island. Three additional collections, includingthat of the present survey, between 1978 and 1987by G.R. AlIen have considerably increased theknown fish fauna (AlIen, 1988). The total presentlystands at 575 species belonging to 78 families.

The fish fauna of Christmas Island consistsprimarily of species associated with coral reefs.Most of the fishes found there have distributionsthat encompass wide areas in the Indo-West Pacificregion. At least 85-90% of the species fall in thiscategory (AlIen and Steene, 1979). Their widedistribution is facilitated by pelagic eggs and larvalstages that drift near the sea's surface.

Despite its Indian Ocean location, the overallfauna is more closely allied to that of the westernPacific, particularly Indonesia. However, severalIndian Ocean species are represented includingCephalopholis nigripinnis, Chaetodon madagaskariensis,C. mitratus, Chromis dimidiata, C. nigrura, C.opercularis, Halichoeres leucoxanthus, and Acanthurusleucosternon. Three species are apparently endemicto the island: Pseudochromis viridis(Pseudochromidae), Steeneichthys nativiatis

Table 6 List of Fishes

(Plesiopidae), and Praealticus natalis (Blenniidae).All three are tiny, secretive reef dwellers, whichexhibit parental care of demersal eggs, andpresumably have a minimal pelagic larval stage.However, future collecting, particularly along theIndian Ocean coast of Sumatra and Java, may revealwider distributions. Centropyge joculator is knownonly from Christmas Island and the Cocos (Keeling)Islands and the damselfish Stegastes insularis isapparently restricted to Christmas Island andMarcus Island in the north Pacific.

Although it is difficult to assess the impact of the1983 El Nifto-related coral die-back on the island'sfish population there appeared to be relatively littlechange between 1978 and 1986-87. However, algalfeeding surgeonfishes appeared to be moreabundant after the 1983 event. Large aggregationsof Acanthurus nigricans, A. gullatus, and A. triosteguswere prominent in this respect. In addition, thecoral eating leatherjacket (Oxymonacanthuslongirostris) although not abundant in 1978, was notsighted during the later surveys.

REFERENCESAllen, G.R (1988). Fishes of Christmas Island Indian Ocean.

Christmas Island Natural History Association.Allen, G.R and Steene, RC. (1979). The fishes of Christmas

Island, Indian Ocean. Australian National Parks andWildlife Service Special Publication 2: 1-81.

This table includes all fishes that have been collected or observed at Christmas Island. Numbers after each species andauthor entry refer to 1987 station numbers. Species not followed by station numbers were recorded during the 1978 and1986 visits by G. Allen.Key to SymbolsNumbers =sampling stations (see Table 4).

Station numbers

RHINCODONTIDAE ... WHALE SHARKSRhincodon typus (Smith, 1829)

SPHYRNIDAE ... HAMMERHEAD SHARKSSphyrna lewini (Griffith & Smith, 1834)

CARCHARHINIDAE ... WHALER SHARKSCarcharhinus albimarginatus (Rilppell, 1835)Carcharhinus amblyrhynchos Bleeker,Carcharhinus longimanus (Poey, 1861)Galeocerdo cuveri (Peron & Lesueur, 1822)Triaenodon obesus (Rilppell, 1837)

84 G.R. Allen

Table 6 (cont.)

Station numbers

MOBULIDAE ... MANTARAYSManta birostris (Donndorff, 1798)

MORINGUIDAE ... SAND-DWELLING WORM EELSMoringua floresiana Weber & de Beaufort, 1916Moringua javanica (Kaup, 1856)Moringua macrochir Bleeker, 1855

XENOCONGRIDAE ... REEF EELSKaupichthys atronasus Schultz, 1953Kaupichthys diodontus Schultz, 1943

MURAENIDAE ... MORAY EELSAnarchias seychellensis Smith, 1962Channomuraena vittata (Richardson, 1844)Echidna nebulosa (Ahl, 1789)Echidna polyzona (Richaradson, 1845)Echidna unicolor (Schultz, 1953Enchelycore bayeri (Schultz, 1953)Enchelycore pardalis Schlegel, 1842Enchelycore schismatorhynchus (Bleeker, 1:853)Enchelynassa canina (Quoy & Gaimard, 1824)Gymnomuraena zebra (Shaw, 1797)Gymnothorax breedeni McCosker & Randall, 1977Gymnothorax buroensis (Bleeker, 1857)Gymnothorax chilospilus Bleeker, 18/?4Gymnothorax enigmaticus McKosker & Randall, 1982Gymnothorax flavimarginatus (Ruppell, 1828)Gymnothorax fuscomaculatus (Schultz, 1953)Gymnothorax javanicus (Bleeker, 1865)Gymnothorax margaritophorus (Bleeker, 1865)Gymnothorax melatremus (Schultz, 1953)Gymnothorax meleagris (Shaw & Nodder, 1795)Gymnothorax molucci'nsis (Bleeker, 1865)Gymnothorax monostigmus (Regan, 1909)Gymnothorax rueppeliae (McClelland, 1845)Gymnothorax zonipectus Seale, 1906Rhinomuraena quaesita'Garman, 1888Siderea picta (Ahl, 1789)Siderea thrysoidea (Richardson, 1844)Uropterygius concolor Ruppell, 1835)Uropterygius fuscoguttatus Schultz, 1953Uropterygius macrocephalus (Bleeker, 1863)UroptenJgius marmoratus (Lacepede, 1803)Uropterygius micropterus (Bleeker, 1852)Uropterygius supraforatus (Regan, 1909)Uropterygius xanthopterus (Bleeker, 1859)

OPHICHTHIDAE ... BURROWING SNAKE EELSApterichtus klazingai (Weber, 1913)Brachysomophis crocodilinus (Bennett, 1833)Callechelys marmorata (Bleeker, 1853)Leiuranus semicinctus (Lay & Bennett, 1839)Myrichthys maculosus (Cuvier, 1817)Muraenichthys laticaudatus (Ogilby, 1897)Phyllophichthus xenodontus (Gosline, 1951)

CONGRIDAE ... CONGER EELSConger cinereus Ruppell, 1828Heteroconger haasi (Klausewitz & Eibl-Eibesfeldt, 1959)

CLUPEIDAE ... HERRINGSSpratelloides gracilis Temminck & Schlegel, 1846

b _

55

5

51,2,125

55

51

5

5

Fishes 85

Table 6 (cont.)

Station numbers

SYNODONTIDAE ... LIZARDFISHESSaurida gracilis (Quoy & Gaimard, 1824)Sallrida undosqllamis (Richardson, 1848)Synodlls englemani Schultz, 1953Synodlls variegatlls (Lacepede, 1803)

OPHIDIIDAE ... CUSKEELSBrotllla mllltibarbata Temminck & Schlegel, 1846

BYTHITIDAE ... CUSKEELSBrosmophyciops palltzkei Schultz, 1953Dinematichthys sp.Microbotllla sp.Ogilbia sp.

ANTENNARIIDAE ... ANGLERFISHESAntennarills analis (Gosline, 1957)Antennarills coccinells (Lesson, 1831)Antennarills nllmmifer (Cuvier, 1817)

GOBlESCOCIDAE ... CLINGFISHESDiscotrema crinophila Briggs, 1976

ATHERINIDAE ... SILVERSIDESAtherion elymlls Jordan & Starks, 1901Hypoatherina barnesi Schultz, 1953

EXOCOETIDAE ... FLYING FISHESCypselllrus oxycephallls (Bleeker, 1852)Cypselurlls poecilopterlls (Valenciennes, 1846)

BELONIDAE ... LONG TOMSPlatybelone argallls (Lesueur, 1821)Tylosllrlls crocodilllS (Lesueur, 1821)

BERYCIDAE ... ALFONSINOSBeryx decadactyllls Cuvier, 1829

HOLOCENTRIDAE ... SQUIRRELFISHESMyripristis berndti Jordan & Evermann, 1903Myripristis kllntee Valenciennes, 1831Myripristis murdjan (Forsskal, 1775)Myripristis pralinills Cuvier, 1829Myripristis vittatlls Valenciennes, 1831Plectrypops lima (Valenciennes, 1831)Sargocentron calldimaclllatllm (Riippell, 1835Sargocentron diadema (Lacepede, 1802)Sargocentron lepros (Allen & Cross, 1893)Sargocentron microstomlls (Giinther, 1859)Sargocentron praslin (Lacepede, 1802)Sargocentron punctatissimllm (Cuvier, 1829)Sargocentron tiere (Cuvier, 1829)Sargocentron tiereoides (Bleeker, 1853)

AULOSTOMIDAE ... PAINTED FLUTEMOUTHSAlllostomlls chinensis (Linnaeus, 1766)

FISTULARIIDAE ... FLUTEMOUTHSFistlllaria commersonii Riippell, 1835

SOLENOSTOMIDAE ... GHOST PIPEFISHESSolenostomlls cyanopterus (Bleeker, 1854)

SYNGNATHIDAE ... PIPEFISHESChoeroichthys brachysoma (Bleeker, 1855)

3

1,3,51,3,5

2

7,9,121,3,5,6,7,9

1

1,3,7,8,10

1,3,5,61,3,5,6,7,12,14

1,2,5,6,7,91,8

1,3

51,3,7,8,10,12

1,3

86 G.R. Allen

Table 6 (cant.)

Station numbers

Choeroiehthys seulptus (Giinther, 1870)Corythoichthys sehultzi Herald, 1953Cosmoeampus banneri (Herald & Randall, 1972)Doryrhamphus baldwini (Herald & Randall, 1972)Doryrhamphus excisus exeisus Kaup, 1856Mierognathus brevirostris pygmaeus Fritzsche, 1981

SCORPAENIDAE ... SCORPIONFlSHESDendroehirus bioeellatus (Fowler, 1938)Dendroehirus zebra (Cuvier, 1829)Ptreois antennata (Bloch, 1787)Pterois radiata (Cuvier, 1829)Pterois volitans (Linnaeus, 1758)Seorpaenodes albaiensis (Evermann & Seale, 1907)Seorpaenodes eorallinus Smith, 1957Seorpaenodes guamensis (Quoy & Gaimard, 1824)Seorpaenodes hirsutus Smith, 1957Seorpaenodes parvipinnis (Garrett, 1863)Seorpaenodes varipinnis (Smith, 1957)Seorpaenopsis oxyeephalus (Bleeker, 1849)Seorpaenopsis diabolus (Cuvier, 1829)Sebastapistes cyanostigma (Bleeker, 1856)Sebastapistes maruitiana (Cuvier, 1829)Sebastapistes strongia (Cuvier, 1829)Sebastapistes tinkhami (Fowler, 1946)Synaneeia verrucosa (Bloch and Schneider, 1801)Taenionotus triaeanthus Lacepede, 1802

APLOACTINIDAE ... VELVETFISHESCoeeotropus larvatus Poss & Allen, 1987

CARACANTHIDAE ,., CROUCHERSCaraeanthus maeulatus (Gray, 1831)Caraeanthus unipinna (Gray, 1831)

PLATYCEPHALIDAE ... FLATHEADSThysanophrys papillolabium Schultz, 1966Thysanophrys sp.

SERRANIDAE ... GROPERSAethaloperea rogaa (ForssHI, 1775)Anyperodon leueogrammieus (Valenciennes, 1828)Belonoperea ehabanaudi (Fowler & Bean, 1930)Cephalopholis argus (Bloch & Schneider, 1801)Cephalopholis analis (Valenciennes, 1828)Cephalopholis igarashiensis Katayama, 1957Cephalopholis leopardus (Lacepede, 1802)Cephalopholis miniatus (ForssHI, 1775)Cephalopholis nigripinnis (Valenciennes, 1828)Cephalopholis sexmaeulatus (Bloch & Schneider, 1801)Cephalopholis spiloparaea (Valenciennes, 1828)Cephalopholis urodeta (Schneider, 1801)Epinephelus eoralieola (Valenciennes, 1828)Epinephelus fasciatus (Forskkal, 1775)Epinephelus hexagonatus (Bloch & Schneider, 1801)Epinephelus laneeolatus (Bloch, 1790)Epinephelus merra Bloch, 1793Epinephelus morrhua (Valenciennes, 1833)Epinephelus retouti (Bleeker, 1868)Epinephelus spilotoeeps Schultz, 1953Epinephelus tauvina (ForssHI, 1775)Epinephelus tukula Morgans, 1959Graeila albomarginata (Fowler & Bean, 1930)

1

12

1

9,12

5

5

5551

3,10,123

1,3,5,6,7,8,10,123,12

1,31,3,7,9,10,121,3,5,63,5,12

1

1,3,5,6,7,8,9,10,12

Fishes 87

Table 6 (cont.)

Station numbers

Gracila polleni (Bleeker, 1868)Grammistes sexlineatus (Thunberg, 1792)Grammistops ocellatus Schultz, 1953Liopropoma mitratum Lubbock & Randall, 1978Liopropoma susumi (Jordan and Seale, 1905)Liopropoma tonstrinum Randall & Taylor, 1988Luzonichthys waitei (Fowler, 1931)Plectranthias fourmanoiri Randall, 1980Plectranthias inermis Randall, 1980Plectranthias nanus Randall, 1980Pogonoperca punctata (Valenciennes, 1830)Pseudanthias dispar (Herre, 1955)Pseudanthias evansi Smith,1954Pseudanthias lori Lubbock & Randall, 1976Pseudanthias smithvanizi Randall & Lubbock, 1981Pseudanthias squamipinnis (Peters, 1855)Pseudogramma polyacantha (Bleeker, 1856)Saloptia powelli Smith, 1964Suttonia lineata Gosline, 1960Variola louti (Forsskal, 1775)

CIRRHlTIDAE ... HAWKFISHESAmblycirrhitus bimacula (Jenkins, 1903)Cirrhitichthys oxycephalus (Bleeker, 1855)Cirrhitus pinnulatus (Schneider, 1801)Oxycirrhites typus Bleeker, 1857Paracirrhites arcatus (Cuvier, 1829)Paracirrhites forsteri (Schneider, 1801)Paracirrhites hemistictus (Glinther, 1874)

PSEUDOCHROMIDAE ... DOTTYBACKSPseudochromis tapeinosoma Bleeker, 1853P. viridis Gill & Allen, 1996Pseudoplesiops multisquamatus Allen, 1987Pseudoplesiops revellei Schultz, 1953

PLESIOPIDAE ... DEVILFISHESCalloplesiops altivelis (Steindachner, 1903)Plesiops coeruleolineatus Ruppell, 1835Plesiops corallicola Bleeker, 1853

TERAPONIDAE ... GRUNTERSTerapon theraps (Cuvier, 1828)

KUHLIIDAE ... FLAGFISHESKuhlia mugil (Forster, 1801)

PRIACANTHIDAE ... BIGEYESHeteropriacanthus cruentatus (Lacepede, 1800)

APOGONIDAE ... CARDINALFISHESApogon angustatus (Smith & Radcliffe, 1911)Apogon apogonides (Bleeker, 1856)Apogon aureus (Lacepede, 1803)Apogon crassiceps Garman, 1903Apogon doryssa (Jordan & Seale, 1906)Apogon evermanni Jordan & Snyder, 1904Apogon jraenatus Valenciennes, 1832Apogon kallopterus Bleeker, 1856Apogon nigrofasciatus Lachner, 1953Apogon novemfasciatus Cuvier, 1828Apogon savayensis Glinther, 1871Apogon semiornatus Peters, 1876Apogon taeniophonls Regan, 1908

1,3

. 1,3,5,6,7,8,12

1,121,3,5,6,7,8,9,10,12,14

1,3,5,6,9

51,121,3,10,12

1,3,5,6,7,8,9,121,3,5,6,121

5

1

10,1212121

1012

5

88 G.R. AlIen

Table 6 (cant.)

Station numbers

Apogon taeniopterus Bennett, 1835Apogon talboti Smith, 1961Apogonichthys ocellatus Weber, 1913Cheilodipterus heptazona Bleeker, 1849Cheilodipterus quinquelineatus Cuvier, 1828Cheilodiptenls sp.Fowleria aurita (Cuvier, 1831)Fowleria vaiulae Jordan and Seale, 1906Pseudamiops gracilicauda (Lachner, 1953)Rhabdamia eremia Allen, 1987

MALACANTHIDAE ... TILEFISHESMalacanthus brevirostris Guichenot, 1848Malacanthus latovittatus (Lacepede, 1798)

ECHENEIDAE ... REMORASEcheneis naucrates Linnaeus, 1758Remora remora Linnaeus, 1758

CORYPHAENIDAE ... DOLPHINFISHESConJphaena hippurus Linnaeus, 1758

CARANGIDAE ... TREVALLIESAlectis indicus (Riippell, 1830)Carangoides ferdau (Forsskal, 1775)Carangoides orthogrammus Jordan & Gilbert, 1882Caranx ignobilis (Forsskal, 1775)Caranx lugubris Poey, 1861Caranx melampygus Cuvier, 1833Caranx sexfasciatus Quay & Gaimard, 1825Caranx tille Cuvier, 1833Decapterus macarellus (Cuvier, 1833)Elegatis bipinnulata (Quay & Gaimard, 1825)Gnathanodon speciosus (ForsskiH, 1775)Scomberoides lysan (Forsskal, 1775)Trachinotus baillonii (Lacepede, 1802)

LUTJANIDAE ... SNAPPERSAphareus furca (Lacepede, 1802)Aprion virescens Valenciennes, 1830Etelis coruscans Valenciennes, 1862Etelis radiosus Anderson, 1981Lutjanus bohar (Forsski\.l, 1775)Lutjanus fulviflamma (Forsski\.l, 1775)Lutjanus fulvus (Schneider, 1801)Lutjanus gibbus (Forsski\.l, 1775)Lutjanus kasmira (Forsski\.l, 1775)Lutjanus rivulatus (Cuvier, 1828)Macolor macularis Fowler, 1931Macolor niger (Forsski\.l, 1775)Paracaesio sordidus Abe & Shinohara, 1962Pristipomoides auricilla Gordan, Evermann & Tanaka, 1927)Pristipomoides zonatus (Valenciennes, 1830)

CAESIONIDAE ... FUSILERSCaesio caerulaureus Lacepede, 1802Caesio lunaris Cuvier, 1830Caesio teres Seale, 1906Pterocaesio lativittata Carpenter, 1987Pterocaesio tile (Cuvier, 1830)

HAEMULIDAE ... SWEETLIPSPlectorhinchus gibbosus (Lacepede, 1802)Plectorhinchus orientalis (Bloch, 1793)

13

101,3,8,14

1

1,7,107,121,3,5,7,10,12

1,7,9,12

1,3,5,6,7,8,9,10,12,147

1,3,5,6,7,8,9,123

1,1010

3,5,6,7,123,5,6,7,10,121,3,12

9,121,3,9,1236,7

6,7,12

Fishes 89

Table 6 (cant.)

Station numbers

LETHRINIDAE ... EMPERORSGnathodentex aureolineatus (Lacepede, 1802)Monotaxis grandoculis (Forsskal, 1775)

NEMIPTERIDAE ... CORAL BREAMScolopsis bilineatus (Blach, 1793)

MULLIDAE ... GOATFISHESMulloides flavolineatus (Lacepede, 1802)Mulloides vanicolensis (Valenciennes, 1831)Parupeneus bifasciatus (Lacepede, 1801)Parupeneus cydostomus (Lacepede, 1801)Parupeneus macronema (Lacepede, 1802)Parupeneus multifasciatus Quay & Gaimard, 1824Parupeneus pleurostigma (Bennett, 1831)

KYPHOSIDAE ... DRUMMERSKyphosus cinerascens (FarsskaJ, 1775)Kyphosus vaigiensis (Quay & Gaimard, 1824)

PEMPHERIDAE ... SWEEPERSPempheris oualensis Cuvier, 1831Pempheris sp.

CHAETODONTIDAE ... BUTTERFLYFISHESChaetodon auriga Farssk1U, 1775Chaetodon baronessa Cuvier, 1831Chaetodon citrinellus Cuvier, 1831Chaetodon ephippium Cuvier, 1831Chaetodon guttatissimus Bennett, 1832Chaetodon kleinii Blach, 1790Chaetodon lineolatus Cuvier, 1831Chaetodon lunula (Lacepede, 1802)Chaetodon lunulatus (Quay & Gaimard, 1824)Chaetodon madagaskariensis Ah!, 1923Chaetodon melannotus Blach & Schneider, 1801Chaetodon meyeri Schneider, 1801Chaetodon mitratus Gunther, 1860Chaetodon ornatissimus Cuvier, 1831Chaetodon punctatofasciatus Cuvier, 1831Chaetodon semion Bleeker, 1855Chaetodon speculum Cuvier, 1831Chaetodon trifascialis Quay & Gaimard, 1824Chaetodon unimaculatus Blach, 1787Chaetodon vagabundus Linnaeus, 1758Forcipiger flavissimus Jardan & McGregar, 1898Forcipiger longirostris (Broussanet, 1782)Hemitaurichthys polylepis (Bleeker, 1857)Heniochus chrysostomus Cuvier, 1831Heniochus monoceros Cuvier, 1831Heniochus singularius Smith & Radcliffe, 1911Heniochus varius (Cuvier, 1829)

POMACANTHIDAE ... ANGELFISHESApolemichthys trimaculatus (Cuvier, 1831)Centropyge bicolor (Blach, 1787)Centropyge bispinosus (Gunther, 1860)Centropyge eibli Klausewitz, 1963Centropyge flavicauda Fraser-Brunner, 1933Centropyge flavissima (Cuvier, 1831)Centropyge joculator Smith-Vaniz & Randall, 1974Centropyge tibicen (Cuvier, 1831)Centropyge vrolikii (Bleeker, 1853)

1,6,7,9,10,121,3,5,6,7,8,9,10,12

1,3,5,6,8,10,141,3,5,6,8,10,12,141,3,5,6,7,8,9,10,12,141,3,5,7,8,10,12,1417,8,10,12,1414

3,5,7,8,10,12,141,3,6,14

1,3,7,8,10,12

3,8,9,10,12

1,3,5,8,12,148,91,7,12,141,5,9,12,141,3,5,81,3,5,6,8,9,121,12

1,3,5,8,10,121,3,5,6,8,10,12

1,3,5,7,9,10,141,3,5,7,8,9,10,111,3,7,9,10,12,141,3,5,9,12,141,3,5,7,8,9,10,121,3,5,6,7,8,9,121,141,3,5,6,7,8,9,10,12,141,3,5,7,9,141,3,5,7,8,12

1,3,5,6,8,10

10,13

1,3,5

1

1,3,5,6,7,8,9,12,14

1

90 G.R. Allen

Table 6 (cont.)

Station numbers

Pomacanthus imperator (Bloch, 1787)Pomacanthus semicirculatus (Cuvier, 1831)Pygoplites diacanthus (Boddaert, 1772)

MUGILlDAE ... MULLETSCrenimugil crenilabris (Forsskal, 1775)Mugil cephalus Linnaeus, 1758

POMACENTRIDAE ... DAMSELFISHESAbudefdufnotatus (Day, 1869)Abudefdufseptemfasciatus (Cuvier, 1830)Abudefdtifsexfascmtus (Lacepede, 1802)Abduefdtifsordidus (Forsskal, 1775)Abduefdtifvaigiensis (Quoy & Gaimard, 1824)Amblyglyphidodon aureus (Cuvier, 1830)Amphiprion clarkii (Bennett, 1830)Amphiprion perideraion Bleeker, 1855Amphiprion sandaracinos AlIen, 1972Chromis alpha Randall, 1988Chromis amboinensis (Bleeker, 1873)Chromis atripectoralis Welander & Schultz, 1951Chromis atripes Fowler & Bean, 1928Chromis caudalis Randall, 1988Chromis dimidiata (Klunzinger, 1871)Chromis delta Randall, 1988Chromis elerae Fowler & Bean, 1928Chromis lepidolepis Bleeker, 1877Chromis lineata Fowler & Bean, 1928Chromis margaritifer Fowler, 1946Chromis nigrura Smith, 1960Chromis opercularis (Giinther, 1867)Chromis ternatensis (Bleeker, 1856)Chromis weberi Fowler & Bean, 1928Chromis xanthochira (Bleeker, 1851)Chromis xanthura (Bleeker, 1854)Chrysiptera glauca (Cuvier, 1830)Chrysiptera leucopoma (Lesson, 1830)Chrysiptera unimaculata (Cuvier, 1830)Dascyllus reticulatus (Richardson, 1846)Dascyllus trimaculatus (Ruppell, 1828)Lepidozygus tapeinosoma (Bleeker, 1856)Plectroglyphidodon dickii (Lienard, 1839)Plectroglyphidodon imparipennis (Valliant & Sauvage, 1875)Plectroglyphidodon johnstonianus (Fowler & Ball, 1924)Plectroglyphidodon lacrymatus (Quoy & Gaimard, 1825)Plectroglyphidodon leucozonus (Bleeker, 1859)Plectroglyphidodon phoenixensis (Schultz, 1943)Pomacentrus bankanensis Bleeker, 1853Pomacentrus chrysurus Cuvier, 1830Pomacentrus coelestis Jordan & Starks, 1901Stegastes albifasciatus (Schlegel & Muller, 1839-44)Stegastes fasciolatus (Ogilby, 1889)Stegastes insularis Allen & Emery, 1985

LABRIDAE ... WRASSESAnampses caeruleopunctatus Ruppell, 1828Anampses meleagrides Valenciennes, 1839Anampses twistii Bleeker, 1856Bodianus anthioides (Bennett, 1830)Bodianus axillaris (Bennett, 1831)Bodianus diana (Lacepede, 1802)Bodianus mesothorax (Bloch & Schneider, 1801)Bodianus opercularis (Guichenot, 1847)

1,3,5,7,8,141,61,3,5,6,7,8,9,10,12,14

5,9,121,3,9,1211,3,9,10,12,141,3,121,3,5,6,7,8,9,10,12,1410,121,3,61

1,3,5,6,7,8,9,10,12

1,3,5,6,7,8,9,10,141

1,3,5

1,3,5,6,8,9,10,12

1,3,7,8,9,10,12,141,3,5,7,8,9,10,12,141,3,7,9,10,121,3,5,6,8,10,12,141,3,5,6,8,10,12,141,3,6,7,8,9,10,121,3,5,6,8,10,12,143,7,81,3,8,9,101

1,3,5,6,8,10,121,3,5,121,3,5,6,121,3,5,8,9,10,14101,3,5,6,8,123,5,9,10,123,5,9

91,3,7,8,101,31,3,81,3,5,6,7,8,9,10,12,14

1,3,5,9,10,12,143,533,5,8,9,121,91,3,5,91,10

Fishes 91

Table 6 (cont.)

Station numbers

Cheilinus oxycephalus Bleeker, 1853Cheilinus trilobatus Lacepede, 1802Cheilinus undulatus Riippell, 1835Cheilio inermis (Forsskal, 1775)Cirrhilabrus cyanopleura (Bleeker, 1851)Cirrhilabrus exquisitus Smith, 1957Coris aygula Lacepede, 1802Coris dorsomacula Fowler, 1908Coris gaimard (Quoy & Gaimard, 1824)Epibulus insidiator (pallas, 1770)Gomphosus varius Lacepede, 1802Halichoeres chrysus Randall, 1980Halichoeres hortulanus (Lacepede, 1801)Halichoeres hyrtlii (Bleeker, 1856)Halichoeres leucoxanthus Randall & Smith, 1982Halichoeres margaritaceus (Valenciennes, 1839)Halichoeres rnarginatus Riippell, 1835Halichoeres melasmapomus Randall, 1980Halichoeres ornatissimus (Garrett, 1863)Halichoeres scapularis (Bennett, 1831)Halichoeres trimaculatus (Quoy & Gaimard, 1834)Hemigymnus fasciatus (Bloch, 1792)Hemigymnus melapterus (Bloch, 1791)Hologymnosus annulatus (Lacepede, 1801)Hologymnosus doliatus (Lacepede, 1802Labrichthys unilineatus (Guichenot, 1847)Labroides bicolor Fowler & Bean, 1928Labroides dimidiatus (Valenciennes, 1839)Labroides pectoralis Randall & Springer, 1975Labropsis xanthonota Randall, 1981Macropharyngodon ornatus Randall, 1978Macropharyngodon negrosensis Herre, 1932Novaculichthys taeniourus (Lacepede, 1802)Oxycheilinus unifasciatus Streets, 1877Pseudocheilinus hexataenia (Bleeker, 1857)Pseudocheilinus octotaenia Jenkins, 1899Pseudocoris heteropterus (Bleeker, 1857)Pseudocoris yamashiroi Schmidt, 1930Pseudodax moluccanus (Valenciennes, 1840)Stethojulis bandanensis (Bleeker, 1851)Stethojulis strigiventer (Bennett, 1832)Thalassoma amblycephalus (Bleeker, 1856)Thalassoma hardwickei (Bennett, 1830)Thalassoma jansenii (Bleeker, 1856)Thalassoma lunare (Linnaeus, 1758)Thalassoma lutescens (Lay & Bennett, 1839)Thalassoma purpureum (Forsskal, 1775)Thalassoma quinquevittatum Fowler, 1928Thalassoma trilobatum (Lacepede, 1801)Wetmorella albofasciata Schultz & Marshall, 1954Xyrichthys aneitensis (Giinther, 1862)Xyrichthys pavo Valenciennes, 1839

SCARlDAE ... PARROTFISHESBolbometopon muricatum (Valenciennes, 1840)Chlorurus microrhinos (Bleeker, 1854)Chlorurus sordidus Forsskal, 1775Seams chameleon Choat & Randall, 1986Searus festivus Valenciennes, 1839Seams forsteni (Bleeker, 1861)Scarus frenatus Lacepede, 1802Seams ghobban Forsskal, 1775

3,14

3,6,961,3,5,6,8,9101,3,5,6,8,9,10,123,5,6,81,3,5,6,7,8,9,10,12,14101,3,10,12,14

33,101,3,5,6,121,3,8,9,1210,311,3,5,8,9,12

1033,52,591,3,5,6,8,91,3,5,6,8,999,10,121,3,81,3,5,6,7,8,10,12,141,3,5,61,3,5,6,8,9,12

1,3,5,6,7,8,9,10,12,141,891,3,5,6,8,10,141,3,5,6,8,101,3,5,6,8

1,3,5,6,7,8,9,10,12,141,3,7,8,141,3,149,10

8,1010

3,9,12,141,5381,3,5,7,8,9,10,12,141,3,5,7,8,9,12

92 G.R. Allen

Table 6 (cont.)

Station numbers

Seams niger Forsskal, 1775Seams ovieeps Valenciennes, 1839Seams prasiognathus Valenciennes, 1840Seams psittaeus Forsskal, 1775Seams mbroviolaeeus Bleeker, 1849Seams sehlegeli Bleeker, 1861Seams spinus (Kner, 1868)

MUGILOIDlDAE ... WEAVERFISHESParapercis clathrata Ogilby, 1910Parapereis tetraeanthus Jordan & Seale, 1906Parapereis sehauinslandi (Steindachner, 1900)

CREEDlDAE ... SANDBURROWERSChalixodytes tauensis Schultz, 1943

TRIPTERYGIIDAE ... TRIPLEFINSCeratobragma helenae Holleman, 1987Enneapterygius elegans (Peters, 1876)Enneapterygius tutuilae Jordan & Seale, 1906Heleogramma ehiea Rosenblatt, 1960Norfolkia braehylepis (Schultz, 1960)Ucla xenogrammus Holleman, 1993

BLENNIIDAE ." BLENNIESAltieus saliens (Forster, 1788)Andamia heteroptera (Bleeker, 1857)Andamia reyi (Sauvage, 1834)Aspidontus taeniatus Quoy & Gaimard, 1834Blenniella eyanostigma (Bleeker, 1849)Blenniella periophthalmus (Valenciennes, 1836)Cirrpeetes eastaneus (Valenciennes, 1836)Cirripeetes gilberti Williams, 1988Cirripectes polyzona (Bleeker, 1868)Cirripeetes stigmatieus Strasburg & Schultz, 1953Cirrisalarias bunares Springer, 1976Eesenius bieolor (Day, 1888)Eesenius midas (Starck, 1969)Eesenius oculus Springer, 1971Entomaerodus eaudofasciatus (Regan, 1909)Entomaerodus epalzeoeheilus (Bleeker, 1859)Entomaerodus vermieulatus (Valenciennes, 1836)Exallias brevis (Kner, 1868)Istiblennius bellus (Giinther, 1861)Istiblennius edentulus (Bloch & Schneider, 1801)Istiblennius lineatus (Valenciennes, 1836)Mimoblennius atrocinetus (Regan, 1909)Nannosalarias nativitatus (Regan, 1909)Parenehelyurus hepburni (Snyder, 1908)Plagiotremus rhinorhynehos (Bleeker, 1852)Plagiotremus tapeinosoma (Bleeker, 1857)Praealtieus natalis (Regan, 1909)Rhabdoblennius snowi (Fowler, 1928)

ELEOTRIDAE ... GUDGEONSCalumia godeffroyi (Giinther, 1877)Eleotris ftlsea (Bloch & Schneider, 1801)

GOBIIDAE ... GOBlESAmblyeleotris fasciata (Herre, 1953)Bathygobius eoeosensis (Bleeker, 1854)Bathygobius eyclopterus (Valenciennes, 1837)Bathygobius padangensis (Bleeker, 1851)

--------

1,3,5,8,9,141,3,5,61,3,81,31,3,5,6,7,8,9,10,12,1431,3,5,6,8,13,14

1,3,5,6,7,8,9,10,12,14

5

8,9,1212

555

55

55

5

55

Fishes 93

Table 6 (cont.)

Station numbers

Callogobius sclateri (Steindachner, 1880)Eviota albolineata Jewett & Lachner, 1983Eviota prasina (Klunzinger, 1871)Eviota sp. 1Eviota sp. 2Eviota sp. 3Fusigbbius duospilus Hoese & Reader, 1985Fusigobius sp. 1Fusigobius sp. 2Gnatholepis scapulostigma Herre, 1953Gobiodon citrinus (Riippell, 1838)Gobiodon rivulatus (Riippell, 1828)Gobiodon sp.Istigobius decoratus (Herre, 1927)Kellogella cardinalis Oordan & Seale, 1906)Pleurosicya sp.Priolepis cinctus (Regan, 1908)Priolepis semidoliatus (Valenciennes, 1837)Trimma emeryi Winterbottom, 1985Trimma macrophthalma Tomiyama, 1936Trimma sheppardi Winterbottom, 1984Trimma taylori Lobel, 1979Valenciennea helsdingenii (Bleeker, 1858)Valenciennea sexguttata (Valenciennes, 1937)Valenciennea strigata (Broussonet, 1837)

MICRODESMIDAE ... WORM GOBlESGunnelichthys monostigma Smith, 1958Nemateleotris decora Randall & Allen, 1973Nemateleotris magnifica Fowler, 1938Ptereleotris evides Oordan & Hubbs, 1925)Ptereleotris heteropterus Bleeker, 1855Ptereleotris microlepis (Bleeker, 1849)Ptereleotris zebra Fowler, 1938

EPHIPPIDAE ... BATFISHESPlatax orbicularis (Forsskill, 1775)Platax teira (Forsskill, 1775)

ZANCLIDAE ... MOORISH IDOLSZanclus cornutus Linnaeus, 1758

ACANTHURIDAE ... SURGEONFISHESAcanthurus blochii Valenciennes, 1835Acanthurus dussumieri Valenciennes, 1835Acanthurus guttatus Bloch & Schneider, 1801Acanthurus leucosternon Bennett, 1832Acanthurus lineatus (Linnaeus, 1758)Acanthurus maculiceps (Ahl, 1923)Acanthurus mata (Cuvier, 1829)Acanthurus nigricans (Linnaeus, 1758)Acanthurus nigricauda Duncker & Mohr, 1929Acanthurus nigrofuscus (Forsskal, 1775)Acanthurus olivaceus Bloch & Schneider, 1801Acanthurus pyroferus Kittlitz, 1834Acanthurus thompsoni (Fowler, 1923)Acanthurus triostegus (Linnaeus, 1758)Acanthurus xanthopterus Valenciennes, 1835Ctenochaetus striatus (Quoy & Gaimard, 1824)Ctenochaetus strigosus (Bennett, 1828)Nasa brevirostris (Valenciennes, 1835)Nasa hexacanthus (Bleeker, 1855)Nasa lituratus (Bloch & Schneider, 1801)

12

5,9,10,12

1,3,5,6,9,10,121,3,9

9,10,14

1,3,5,6,7,8,12,14

105,81,3,8,9,10,121,3,5,8,9,141,3,5,6,7,8,9,10,12,141,3,5,6,8,10,12,14

1,3,5,6,7,8,9,10,12,14

1,3,5,6,8,101,5,7,8,9,12,14

1,8,9,12,141,3,5,9,12,14

1,3,5,6,8,9,12,141,3,5,6,8,9,10,12,14

1,3,8,10,121,3,5,6,8,9,10,12

94 G.R. Allen

Table 6 (cont.)

Station numbers

Naso tuberosus Lacepede, 1802Naso unicornis (Forsskc'l.l, 1775)Naso vlaminghi (Valenciennes, 1835)Paracanthurus hepatus (Linnaeus, 1766)Zebrasoma scopas (Cuvier, 1829)Zebrasoma veliferum (Bloch, 1797)

SPHYRAENIDAE ... BARRACUDASSphyraena barracuda (Walbaum, 1792)Sphyraena flavicauda Riippell, 1838

GEMPYLIDAE ... SNAKE MACKERELSPromethichthys prometheus (Cuvier, 1831)

SCOMBRIDAE ... TUNASAcanthocybium solandri (Cuvier, 1831)Euthynnus affinis (Cantor, 1849)Gymnosarda unicolor (Riippell, 1835)Katsuwonus pelamis (Linnaeus, 1758)Thunnus albacares (Bonnaterre, 1788)

ISTIOPHORlDAE ... SAILFISHESIstiophorus platypterus (Shaw & Nodder, 1792)Makira mazara Gordan & Snyder, 1901)Tetrapturus audax (Phillipi, 1887)

BOTHIDAE ... LEFT HANDED FLATFISHESBothus mancus (Broussonet, 1782)Bothus pantherinus (Riippell, 1830)

SAMARIDAE... RIGHT HANDED FLATFISHESSamaricus triocellatus Woods, 1960

SOLEIDAE ... SOLESAseraggodes melanostictus (Peters, 1876)Aseraggodes sp.

BALISTIDAE ... TRlGGERFISHESBalastapus undulatus (Park, 1797)Balistoides conspicillum (Bloch & Schneider, 1801)Balistoides viridescens (Bloch & Schneider, 1801)Melichthys indicus Randall & Klausewitz, 1973Melichthys niger (Bloch, 1786)Melichthys vidua (Solander, 1844)Odonus niger (Riippell, 1836)Rhinecanthus rectangulus (Bloch & Schneider, 1801)Sufflamen bursa (Bloch & Schneider, 1801)Sufflamen chrysopterus (Bloch & Schneider, 1801)Xanthichthys auromarginatus (Bennett, 1831)Xanthichthys caeruleolineatus Randall, Matsuura &

Zama,1978

MONACANTHIDAE ... LEATHERJACKETSAlutera scripta (Osbeck, 1765)Amanses scopas (Cuvier, 1829)Cantherines dumerilii (Hollard, 1854)Cantherines pardalis (Riippell, 1835)Oxymonacanthus longirostirs (Bloch & Schneider, 1801)Paraluteres prionurus (Bleeker, 1851)Pervagor janthinosoma (Bleeker, 1854)

OSTRACIONTIDAE ... BOXFISHESOstracion cubicus Linnaeus, 1758Ostracion me1eagris Shaw, 1796Ostracion solorensis Bleeker, 1853

1,3,8,10,143

1,3,5,6,7,8,10,12,141,8

12

1,3,5,6,7,8,10,12,13,143,8,12,141,5,6,121,91,3,5,6,7,8,9,10,12,143,5,7,8,9,10,12,14121,3,5,7,8,10,12,131,3,5,8,10,12,13,141,3,5,7,9,10,12,141,3,6,12,14

8,12

31,3,5,91,3,5,141,3,5,6,10,14

121,3,5,13

Fishes 95

Table 6 (cant.)

Station numbers

TETRAODONTlDAE ,.. PUFFERFISHESArothron hispidus (Linnaeus, 1758)Arothron meleagris (Lacepede, 1798)Arothron nigropunctatus (Bloch & Schneider, 1801)Canthigaster amboinensis (Bleeker, 1865)Canthigaster bennetti (Bleeker, 1854)Canthigaster epilampra Genkins, 1903)Canthigaster janthinoptera (Bleeker, 1855)Canthigaster tyleri Allen & Randall, 1977Canthigaster valentini (Bleeker, 1853)

DIODONTlDAE ... PORCUPlNEFISHESDiodon hystrix Linnaeus, 1758Diodon liturosus Shaw, 1806

11,143,5,14

1,3