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Costello, Emblow and White (eds), 2001
HEMICHORDATA
Compiled by Jacob van der Land.
The European l ist was derived from URMO with names provided by J. Hallan. No expert has checked the accuracy of the ERMS list.
PHYLUM HEMICHORDATA
Class Enteropneusta
Family Harrimunlidae Harrimania
kupJferi (von Willemoes-Suhm, 1 87 1 ) SaccogJossus
horsl; BrambeH & Goodhart, 1941 kowalevskii (Agassiz, 1 873) mereschkawskii (Wagner, 1 885) pygmaeus Hinrichs & Jacobi, 1 938 ruber Tattersall, 1 905
Stereobalanus canadensis (Spengel, 1 893)
Prologlossus kaehleri (Caullery & Mesnil, 1900)
Family Ptychoderidae Balanoglossus
clavigerus delle Chiaje, 1 829 Glossobalanus
elongatus Spengel, 1904 marginalus Meek, 1 922 minutus Kowalevsky, 1 866 sarniensis (Koehler, 1 886)
Family SpengeJidae Glandiceps
ta/aboti (Marion, 1 876)
Family il/cer/ae sedls Planctosphaera
pelagica (Spengel, 1 932)
Class Pterobrancbia
Family Rhabdopleuridae Rhabdopleura
compacta Hincks, 1 880 normani Allmann, 1 869
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ECHINODERMATA
Compiled by Hans G. Hansson
The list of European Echinodermata was compiled by Hans G. Hansson. H used the Echinoderm part of the compilers NEAT l ists (available via Internet at address htttp:llwww.tmbl .gu .se/libdb/taxon/taxa.html) as a starting point. NEAT (North East Atlantic Taxa) are preliminary check-lists, covering Scandinavian marine animal taxa. The chapter by FJ. Fell on Echinodermata in Parker ( 1 982) was used as the systematic framework. The asteroids are mainly l isted from the recent monograph on Atlantic species by Clark & Downey ( 1 992). Mortensen ( 1 927) is used as a basal source for the other groups. Additions from several other sources have been made, and the electronic part of the Zoological Record (covering recent decades) has been utilised. However, examination of the reports from the large expeditions (except the Norwegian North Atlantic and the (ngolf) and other literature since WWII, may add to the species list.
The compiler is generally interested in marine fauna, but is not a specialist on any Echinoderm taxon, so there are certainly several omissions and lapses in this draft list. The assistance of experts in groups of Echinodermata was thus invaluable. Dr Sabine StOhr, Stockholm, assisted w ith ophiuroid names, Dr Claude Massin, Brussels, assisted with the holothurians and added 9 omitted species of this taxon to the list. Dr Andrey Gebruk, Moscow, together with his friends Alexander Mironov and Alexei Smirnov kindly helped with several corrections and additions to the holothurians, the crinoids, the echinoids and the genus Caulaster among the asteroids. Prof. Dusan Zavodnik, Rovinj, Croatia, provided severa l corrections, new distribution notes (particularly for the Adriatic Sea area), information about synonyms, and added a few omitted species. Dr Mateo Garrido, Gran Canada, provided additional records from the Canaries.
Authors of most genus and species names in the list below have been found, and most often also the years of description, but with several exceptions. Also, some of the years of authority in this list may be wrong, because different sources often have different opinions regarding which year a certain species is described (e.g. gen. Poliometra Clark; the original publication: 1 923, Neave: 1 924, McKenzie in Howson & Picton 1 997: 1 9 1 3; in this case the year from the original publ ication must be correct). Thus, further study of the primary l iterature may improve the accuracy of this list.
References Belyaev G.M. & A.N. Mironov, 1 996. Starfishes Porcellanaster
and Caulaster (Porcellanasteridae) from the Atlantic and the Antarctic. Zoologichesky zhurnal, 75 (6): 886 - 899 (In Russian, English summary).
Brattegard, T & T. Holthe (Eds.). 1 997. Distribution o/marine. benthic macro-organisms in Norway. A tabulated catalogue. Preliminary edition. Reseach Report for ON, Nr. 1 997- 1 . Trondheim. 409 pp. (Echinoderm chapters by T. HBis8!ter, T Brattegard & J.-A. Sneli, pp. 3 0 1 -3 1 5).
Clark, A.H. 1 923. Crinoidea. The Danish IngolfExpedition . . Vol. IV. Luno, Copenhagen. 58 pp.
Clark, A.M. 1 970. Echinodermata Crinoidea. Marine Invertebrates a/Scandinavia. 3. Universitetsforiaget, Oslo. 55 pp.
Clark, A.M. & M.E. Downey 1 992. Starfishes o/the Atlantic. Chapman & Hall, London. 794 pp.
- 3 36 -
D'yakonov, A.M. 1 967. Ophiuroids o/the USSR Seas. Israel Program for Scientific Translations, Jerusalem 123 pp.
D'yakonov, A.M. 1 968. Sea Stars (Asteroids) o/the USSR Seas. Israel Program for Scientific Translations, Jerusalem 1 83 pp.
D'yakonov, A.M. 1 969. Echinodermata, Vol. 1. Sea-Urchins (Echinoidea) No. I . Fauna 0/ Russia and adjacent Countries. Israel Program for Scientific Translations, Jerusalem 265 pp.
Fell, H.B. 1 960. Synoptic Keys to the Genera of Ophiuroid ea. Zoology Publicalionsfrom Victoria University of Wellington No. 26. 34 pp + literature index.
Fiege, D. & Y. Liao 1 996. Penilpidia IUdwigi (Marenzeller, 1 893) (Holothurioidea: Elpidiidae) rediscovered from the Eastern Mediterranean Sea. In: Uiblein, F., OIt,I., Stachowitsch, M. (Eds) Deep-sea and extreme shallowwater habitats: affinities and adaptations. Biosystematics and Ecology Series, 1 1 , 61 -66.
Gebruk, A.V. 1 989. Revision of the family Pelagothuriidae (Holothurioidea, Elasipodida) with a review of swimming holothurians. Part I . Revision of Pelagothuriidae. Zool. Zhurnal 68 ( 12): 57-66. (In Russian, English summary).
Gebruk, A.V., 1 990. Deep-sea holothurians ofthe/amily Elpidiidae. Moscow, Nauka, 1 60 p. [In Russian].
Gulliksen , B. Palerud, R., Brauegard, T. & I.-A. Sneli, 1 999. Distribution of marine benthic macro-organisms at Svalbard (including Bear Island) and Jan Mayen. Research report for DN 1 999 - 4. 148 pp. Trondheim, Norway.
Hansen, B, 1 975. Systematics and biology of the deep-sea holothurians. Galathea Report, 13, 1 -262.
Heding, S.G. 1 935. Holothurioidea I. The Danish Ingolf Expedition. Vol. IV. Hagerup, Copenbagen. 84 pp + 8 plates.
Heding, S.G. 1 935 Echinoderms. Meddele/ser om Gren/and. Bd. 1 04, Nr. 1 3. 68 pp + 1 plate.
Heding, S.G. 1 936 Echinoderms. Meddelelser om Gren/and. Bd. 1 08, Nr. I . 34 pp.
Heding, S.G. 1 942. Hololhurioidea II. The Danish Ingolf Expedition. Vol. IV. Hagerup, Copenbagen. 39 pp + 2 plates.
Heding, S.G. & A. Panning 1 954. Phyllophoridae. Spolia Zoologica Musei Hauniensis XIII. 209 pp.
Howson, C.M. & B.E. Picton 1 997 (Eds.). The Species Directory o/the Fauna and Flora o/the British Isles and Surrounding Seas. The Ulster Museum, Belfast & The Marine Conservation Society, Ross-on-Wye. 508 pp. (Echinoderm chapter by J.D. McKenzie, pp. 287-295).
Heisa:ter, T. 1 990. An annotated check-list of the echinoderms of the Norwegian coast and adjacent waters. Sarsia 75 : 83-1 06
Kier P.M. & M. H. Lawson, 1 978. Index of Living and Fossil Echinoids 1 924- 1 970. Smithson. Contrib. Paleobiol. 34 : vi+ 1 82 pp. (not yet seen)
Madsen, FJ. 1 947. The Echinoderms collected by the "Skagerak" expedition in the eastern Atlantic 1 946. I. Asteroidea, Ophiuroidea, Echinoidea, and Holothuroidea. Goteborgs Kungl. Vetenskaps- och Villerhets-Samhiilles Handlingar, Sj!ltte fOljden. Ser. B, Band 5. N:o 7. 1 6 pp.
Madsen, F.J. 1 987. The Henricia sanguino/enta complex (Echinodermata, Asteroidea) of the Norwegian Sea and adjacent waters. A re-evaluation, with notes on related species. Steenstrupia 1 3 (5), pp. 20 1-268.
Madsen, FJ. & B. Hansen 1 994. Echinodermata Holothurioidea. Marine Invertebrates of Scandinavia. 9. Scandinavian University Press. 1 43 pp.
Mil ler, J.E and Pawson, D.L. 1 990. Swimming sea cucumbers (Echinoderamata :Holothuroidea): a survey, with analysis of swimming behaviour in four bathyal species. Smithsonian Contributions to Marine Sciences, 35, 1 8p.
Mironov A.N., 1 974. Pourtalesiid sea urchins ofthe Antarctic and Subantarctic (Echinoidea: Pourtalesiidae). Transactions o/the P.P. Shirshov Institute 0/ oceanology, 98: 240 - 252. (In Russian, English summary).
Mironov A.N., 1 985. Circumcontinental zonation in distribution of Atlantic echinoids. Transactions o/the P.P. Shirshov
European Register of Marine Species
Institute 0/ oceanology, 1 20, 70-95. (In Russian, English summary).
Mironov A.N. 1 995. Holasteroid echinoids. 2. Pourtalesia. Zoologichesky zhurnal, 74 ( 1 2), 59-75. (In Russian, English summary).
Mironov A.N. 1 997. Holasteroid echinoids. 4. Echinosigra. Zoologichesky zhurna/, 76, (2): 1 73-1 86. (In Russian, English summary).
Mortensen, T. 1 903. Echinoidea I. The Danish lngolf Expedition. Vol. IVa. Luno, Copenhagen. 1 83 pp + 2 1 plates.
Mortensen, T. 1 907. Echinoidea II. The Danish Ingolf Expedition. Vol. IVa. Luno, Copenhagen. 200 pp + 1 9 plates.
Mortensen, T. 1 927, Handbook of the Echinoderms 0/ the British Isles. Clarendon Press, Oxford. 47 1 pp.
Mortensen, T. 1 933. Ophiuroidea. The Danish lngolf Expedition. Vol. IV. Luno, Copenhagen. 1 2 1. pp + 3 plates.
Neave, S.A. 1 939-50. Nomenclator Zoologicus. Vol . l ·5. The Zoological Society of London. + M.A. Edwards & A.T. Hopwood (Eds.) 1 966 Vol. 6 + M.A. Edwards & H.G. Vevers (Eds.) Vol. 7 .
Parker, S.P. (Ed.) 1 982. Synopsis and Classification 0/ Living Organisms. 2 Vols. McGraw-Hili, New York. 1 166 + 1 232 pp.
Paterson, G .L.J. 1 985. The deep-sea Ophiuroldea of the north Atlantic Ocean. Bulletin o/the British Museum (Natural History). Zoology. 49 ( I ). 1 62 pp. (not seen)
Pereira, Mariano Oliveiro Rego 1 997. Checklist of the l ittoral echinoderms of the Azores. Aforeana 8 (3): 33 1 -337.
Picton, B.E. 1 993. A Field Guide to the Shallow-water Echinoderms o/the British Isles. Immel Publishing. 96 pp.
Sherborn, C.D. 1 902. Index Animalium sive index nom inurn quae ab a.d MDCCL VIII generibus et speciebus anlmalium imposita sunt. Sectio prima a kalendis Ianuariis, MDCCLVIII usque adjinem Decembris. MDCCC. C. J. Clay & Sons, Cambridge Unlv. Press, London. 1 1 95 pp.
Sherborn, C.D. 1 922-32. Index Animalium sive index nominum quae ab a,d MDCCLVIIl generibus et speciebus animalium imposita sunt, Sectio secunda a kalendis Ianuariis, MDCCCI usque adjinem Decembris, MDCCCL. The Trustees of the British Museum, London. 7056 + 1098 pp.
Tortonese, E. 1 965. Echinodermata. Fauna d'italia. Vol. 6. Edizioni Calderini, Bologna. 422 pp.
PHYLUM ECIDNODERMATA
Class Crinoidea
Order Millericrinida
Family Hyocrinidae Gephyrocrinus
grima/dii Koehler & Bather, 1 902
Order Cyrtocrinida
Family Holopodidac Cyathidium
/oresti Cherbonnier & Guille, 1 972 C. meteorensis Fechter, 1 973
Order Bourgueticrinida
Family Bourgueticrinidae Conocrinus
cabioshi Roux, 1 976 cherbonnieri Roux, 1 976
Democrinus par/aUi Perrier, 1 883 = Rhizocrinus magnus Gislen, 1 947 = Rhizocrinus rawsoni Koehler, 1 909, non Pourtales,
1 874 conifer (A.H. C lark, 1909)
Bythocrinus intermedius A.H. Clark, 1 9 1 5 � Rhizocrinus conifer A.H. Clark, 1 909 = Rhizocrinus robustus A.H. Clark, 1 909 rawsonii (Pourtali!:s, 1 874) = Rhizocrinus rawsonii Pourtales, 1 874
Rhizocrinus
- 337 -
Costello, Emblow and White (eds), 2001
lofotensis M. Sars, 1 868 = R. verrilli A.H. Clark, 1 908
Family Bathycrinidae Bathycrinus
aldrichianus Wyville Thomson, 1 817 = B. serratus AH. Clark, 1 908 carpenlerii (Danielssen & Koren, L 811)
lIycrintls carpenferii Danielssen & Koren, 1 811 gracilis Wyville Thomson, 1 817
Monachocrinus perrieri (Koehler & Vaney, 1 9 1 0) = Balhycrintls perrieri Koehler & Vaney, 1 9 1 0
M coe/us H.L. Clark, 1 923 recuperattls (Perrier, L 885) = Ilycrinus rectlperalus Perrier, 1 885 sexradiatus AH. Clark, 1 9 1 9
Family Phrynocrinidae Zeuclocrinus
gis/eni AM. Clark, 1 913
Family Porphyrocrinidae Porphyrocrinus
incrassalus (Gislen, 1933) Ihalassae Roux, 1 911
Order Isocrinida
Family Isocrinidae Diplocrinus
(Annacrinus) wyvillelhomsoni (Jeffreys, 1 8??) Penlacrinus wyville-Ihomsoni Jeffreys, 1 8??
Order Comatulida
Family Pentamctrocrinidae Penlamelrocrinus
allanticus (Perrier, 1 883) Eudioerinus allanticus Perrier, 1 883
Thaumalacrinus jungerseni A.H. Clark, 1 923
Family Antedonidae Antedon
bifida (Pennant, 1 117) = ? Anledan duebeni Bohlsche, 1 866 = A. gorgonia de Freminville, 1 8 1 1
A. maroccana A.H. Clark, 1 9 1 0 = Aslerias bifida Pennant, 1 711 medilerranea (de Lamarck, 1 8 1 6) = A. adriatica A.H. Clark, 1 9 1 1 = Comalula annulala Risso, 1 826
Comalula medilerranea de Lamarck, 1 8 1 6 pelasus (Dahen & Koren, 1 846) = Aleelo petasus DUben & Koren, 1 846
Hathrometra sarsii (DUben & Koren, 1 846)
Antedon teneila Carpenter non (Retzius) = H. tenel/a (Carpenter, 1 888) non (Retzius, 1 783)
Heliometra glaeiaUs (Owen, 1 833 ex Leach MS) = Aleelo eschrichtii J. MUller, 1 84 1 = Alecto glacialis Owen, 1 833 ex Leach MS = Antedon quadrata P .H. Carpenter, 1 884
Leptometra cellica (M'Andrew & Barrett, 1 858) = Comalula cel1ica M'Andrew & B arrett, 1 85 8 = Comatula woodwardi Barrett, 1 857, non Forbes, 1 852 = L. phalangium (Carpenter, 1 88 1 ) non (J. MUller, 1 84 1) phalangium (J. MUlier, 1 84 1 ) = Aleclo phalal1gium J. MUlier, 1841
Meteororomelra montieola A.M. Clark, 1 980
Orthometra hibernica (A.H. Clark, 1 91 3) = Trichometra hibernica A.H. Clark, 1 9 1 3
Poliometra prolixa (Sladen, 1 88 l ) = Anledon prolixa Siaden, 1 88 1
Harhrometra prolixa (Sladen)
Trichometra cubensis (Pourtales, 1 869) = Antedon eubensis Pourtales, 1 869
T. aspera AH. Clark, 1 908 delieata A.H. Clark, 1 9 1 J
Thaumatomelra seplentrionalis A.H. Clark, 1 923
Family Atelecrinidae A telecrinus
balanoides P.H. Carpenter, 1 88 1 = A. helgae A.H. Clark, 1 9 1 3
Family Comasteridae Neocomalella
europaea A.H. Clark, 1 9??
Family Thalassometridae Koehlermelra
f1ava (Koehler) = Anledonflava Koehler porrecta (P.H. Carpenter, 1 8??) = Antedon porrecla P.H. Carpenter, 1 8??
Thalassometra lusilanica (P.H. Carpenter, 1 884) A.M. Clark, 1 980 = Antedon lusilanica P.H. Carpenter, 1 884
Sliremelra ltlsitaniea (P.H. Carpenter, 1 884) omissa (Koehler) = Antedon omissa Koehler
Class Asteroidea
Order Paxillosida
Family Aslropectinidae Asrropecten
- 3 3 8 -
arandacus (Linnaeus, 1 158) = Aslerias arandaca Linnaeus, 1158 = Aslerias auranliaca Tiedemann, 1 8 1 6
A. antarcticus Studer, 1 816 = A. crenaster Dujardin & Hupe, 1 862
A. gruveN Koehler, 1 9 1 2 = A . meridionalis Studer, 1 876 = A. perarmalus Perrier, 1 869 bispinosus (Otto, 1 823) = Aster/as bispinosa Otto, 1 823
A. myosurus Perrier, 1 869 hermalophilus Siaden, 1 883 iberieus Perrier, 1 894 irregularis subsp. irregularis (Pennant, 1 771) = Asterias irregularis Pennant, 1 171 = A. acleularis Nonnan, 1 865 = A. echinulatus J. MUller & Troschel, 1 844 = A. helgolandieus Greeff, 1 87 1 = A. muelleri }. MUlier & Troschel, 1 844 Irregu/�ris subsp. pentaeanlhus (Delle Chiaje, 1 827) = Aster/as pentacantha Delle Chiaje, 1 827 jonsloni (Delle Chiaje, 1 827)
= Asterias jonstoni Delle Chiaje, 1 827 = A. aster De Filippi, 1 859 = A. squamalus J. MOller & Troschel, 1 844 platyacanlhus (Philippi, 1 837) = Aslerias platyacantha Philippi, 1 837 spinulosus (Philippi, 1 831) = Asterias spinulosa Phil ippi, 1 837
Bathybiasler vexilfifer (W. Thomson, 1 873) = Archaster robuslus Verrill, 1 884
Archasler vexillifer W. Thomson, 1 873 = Aslropeelen pallidus Danielssen & Koren, 1 817 = Ifyasler mirabilis Danielssen & Koren, 1 884
Phoxaster pumilus Siaden, 1 889 Dytasler
grandis subsp. grandis (Verrill, 1 884) Archasler grandis Verrill, 1 884
= D. agassizi Perrier, 1 894 D. biserialis Siaden, 1 889
= D. exilis var. carinala S laden, 1 889 = D. madreporifer Sladen, 1 889
D. rigidus Perrier, 1 894 intermedius Perrier, 1 89 1 moliis (perrier, 1 885)
Crenaster mollis Perrier, 1 885 semispinosus (perrier, 1 894)
Crenaster semispinosus Perrier, 1 894 D. parvulus Koehler, 1 909
Leplychaster arctjcus (M. Sars, 1 85 1 ) � Astropecten arct/cus M . Sars, 1 85 1 � As/ropec/en lutken; Barrett, 1 857
Loncho/as/er /ar/areus Sladen, 1 889
Persephonaster /eptactis H.L. Clark, 1 94 1 patagia/us (Sladen, 1 889)
P. spinulosus H.L. Clark, 1 94 1 Psi/aster patagia/us Siaden, 1 889
sphenoplax (Bell, 1 892) Astropec/en sphenop/ax Bell, 1 892
= Psi/asteropsis humilis Koehler, 1 907 P/ulonaster
agassizi subsp. nota/us Siaden, 1 889 = ?Goniopecten edwardsi Perrier, in Milne-Edwards,
1 882 ?Goniopecten inermis Perrier, 1 885 (nomen nudum)
= P. abbreviatus Siaden, 1 889 P. granulosus Perrier, 1 89 1
= P. no/alus S laden, 1 889 bifrons (Wyvil\e Thomson, 1 873)
Archas/er bifrons Wyville Thomson, 1 873 = P. proteus H.L. Clark, 1 923
Psi/aster andromeda subsp. andromeda (J. MUlier &. Troschel,
1 842) � Astropecten andromeda J. MUller &. Trosche1, 1 842 � Astropecten crisUi DUben &. Koren, in DUben, 1 845 cassiope Sladen, 1 889
? Archaster pulcher Perrier, 1 88 1 = P. andromeda var. cassiope Grieg, 1 907 = P. squameus H.L. Clark, 1 94 1
Tethyaster subinermis (Philippi, 1 837)
Asterias subinermis Philippi, 1 837
Family Luidiidae Luidia
atlanlidea Madsen, 1 950 ciliaris (Philippi, 1 837)
Aslerias imperali Delle Chiaje, 1 84 1 = L. fragilissima Forbes, 1 839 sarsil subsp. sarsii D!lben &. Koren, in Dilben, 1 845
Astrella simp/ex Perrier, in Milne-Edwards, 1 882 = L. africana S iaden, 1 889
FamilY Ctenodiscidae Ctenodiscus
crispatus (Relzius, 1 805) Asterias aranda Dewhurst, 1 834
= Asterias crispala Retzius, 1 805 = Asterias palaris Sabine, 1 824
C. cornicu/atus Perrier, 1 875 = C. pygmaeus J. MUlier &. Troschel, 1 842
Family Radiasteridae Radiaster
lizardi (Sladen, 1 882) Mimasler lizardi Sladen, 1 882
Family Porcellanasteridae Cau/aster
eremicus (Sladen, 1 889) = A/balrotrossaster richardi Koehler, 1 909
Porcel/anaster eremicus Siaden, 1 889 pedlinculailis Perrier, 1 882
Hypha/asler inermis Siaden, 1 883 = H. anlonU Perrier, 1 885
European Register of Marine Species
= H. fortis Koehler, 1 907 = H. gracilis Koehler, 1 907 = H. parfaiti Perrier, 1 885
Parcel/an aster ceruleus Wyville Thomson, 1 877 = Albatrossia semimarginalis Ludwig, 1 905
Cau/aster sladeni Perrier, 1 885 = P. granulosus Perrier, 1 88 5 = P. inermis Perrier, 1 885
P. irreglilaris Lieberkind, 1 932 Slyracasler
armalus Siaden, 1 883 = Machairaster pictus Perrier, 1 884 = S. edwardsi Perrier, 1 885
S. spinosus Perrier, 1 885 chuni Ludwig, 1 907 elongallis Koehler, 1 907 horridus Siaden, 1 883 = Chunasler scapanephorus Ludwig, 1 907
Thoracas/er eylindrailis Sladen, 1 883 = Pseudaster cordi/er Perrier, 1 885
T. magnus Ludwig, 1 907
Order Notornyotida
Family Benthopectinidaf Benthopeelen
simplex subsp. simplex (Perrier, 1 88 1 ) Archaster simp/ex Perrier, 1 88 1
= B. spinasus Grieg, 1 9 1 8 (p.p.), non Verrill, 1 884 = Pararchas/er armatus Siaden, 1 889 spinosissimus (Sladen, 1 889) = Pararchas/er spinosissimus Siaden, 1 889
Cheiraster sepilus (Verrill, 1 885) = Archaster sepi/us Verrill, 1 885
Luidiaster dubius H.L. Clark, 1 94 1 = Peclinaster dispar Verrill, 1 9 1 5 = Pectinaster echinu/atus Ludwig, 1 9 1 0 (p.p.)
Pontaster perp/exlls Perrier, 1 894 = Pontaster venus/us Siaden, 1 889
Peetinas/er fllholi Perrier, 1 885
= Pan/aster forcipatus Siaden, 1 889 Pantaster prislinus Siaden, 1 889
Ponlaster lenuispinus (DUben &. Koren, 1 846) = Astropeelen lenuispinus DUben &. Koren, 1 846 = Crenaster marionis Perrier, 1 885 = P. hebilus Siaden, 1 889
P. limbatus Siaden, 1 889
Order Va)vatida
Family Chaetasteridae Chaetaster
/ongipes (Relzius, 1 805) = Asterias longipes Retzius, 1 805 = Asterias subu/ata de Lamarck, 1 8 1 6
Nepanlhia /esselala Gray, 1 840
Family Odontasteridae Hop/asler
spinosus Perrier, in Milne-Edwards, 1 882 = Penlagorwsler /epidus Siaden, 1 889
Odontaster mediterraneus (von Marenzeller, 1 893) = Gnathasler medilerraneus von Marenzeller, 1 893
Family Goniasleridae
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Ceramasler granularis subsp. granularis (Relzius, 1 783) = Aslerias granularis Retziu8, 1 783 = Aslerias tessellatus de Lamarck, 1 8 1 6 = Astrogonium borea/e B arrett, 1 857 = Pentagonaster eximius Verrill, 1 894
Pentagonaster simp/ex Verrill, 1 895 grenadensis subsp. grenadensis (Perrier, 1 88 1 )
Costello, Emblow and White (eds), 2001
= Astrogonium greeni Bell, 1 889 = Penlagonaster qffinis Perrier, 1 884 = Penlagonasler balteatus Siaden, 1891 = Pentagonaster crassus Perrier, 1 885 = Pentagonaster deplasi Perrier, 1 885 = Pentagonaster gosselini Perrier, 1 884 = Penlagonaster grenadensis Perrier, 1881
Pentagonaster haesitans Perrier, 1 885 = Penlagonaster hyslricis von MarenzeIler, 1 893 = Penlagonaster kergroheni Koehler, 1 895
Evoplosoma scorpio Downey, 198 1
Goniasler tessellatus (de Lamarck, 1 8 1 6) = Asterias granufaris Gmelin, 1 79 1 (p.p.)
Aslerias lesselfata de Lamarck, 1 8 16 = Astrogonium duhium Perrier, 1 869 = G. africanus Verrill, 1 87 1 = G. americanus Verrill, 1 87 1 = G. cuspidafus Gray, 1 840
G. sehae Gray, 1 840 Hippasferia
phrygiana subsp. phrygiana (Parelius, 1 768) Aslerias equestris de Lamarck, 1 8 1 6, non Retzius, 1 805
= Asterias johnstoni Gray, in Johnston, 1 836 = Asterias phrygianus Parelius, 1 768
Astrogonium acufeatum Barrett, 1 857 = Goniaster ahbensis Gray, 1 843
H. (Nehippasleria) insignis Dons, 1 938 = H. cornula Gray, 1 840 = H. europaea Gray, 1 840
H. plana Gray, 1 840 insignis Dons, 1 937
Mediasler bairdi subsp. bairdi (Verrill, 1 882) = Archasler bairdi Verrill, 1 882 = M slellatas Perrier, 1 89 1
Nymphaster arenalus (Perrier, 1 88 1 )
Dorigonajacqueli Perrier, 1 894 = Dorigona prehensilis Perrier, 1 885 = N. albidus S laden, 1 889
N. basilicus S laden, 1 889 = N. prolenlus Siaden, 1 889 = Penlagonasler arena/as Perrier, 1 88 1 = Penlagonasler subspinosus Perrier, 1 88 1 = Penlagonas/er lernalis Perrier, 188 1
Paragonaster subliUs (Perrier, 1 88 1 ) = Archasler /ormosus Verrill, 1 884
Goniopeclen sublilis Perrier, 1 88 1 = P. cylindralus Siaden, 1 889 = P. strictus Perrier, 1 894
Pentagonaster elongatus Perrier, 1 885
Pellasler placenla (J. MUller & Troschel, 1 842) � Gonias/er nidarosiensis Storm, 1 88 1 = Goniodiscus aculus Heller, 1 863
Goniodiscus placenla 1. MUller & Troschel, 1 842
= Goniodiscus placenlae/ormis Heller, 1 863 = P. hebes Verrill, 1 899
Penlaganasler minor Koehler, 1 895 = Penlagonasler mirabilill Perrier, 1 875 = Penlagonasler planus Verrill, 1 895 = Penlagonasler vincenti Perrier, 1 885
PUnlhasler denialUs (Perrier, 1 884) � Penlagonaster concinnus Siaden, 1 89 1 = Pentagonasler denlalus Perrier, 1 884
Pentagonasler grandis Perrier, 1 885 = Pentagonasler perrieri Siaden, 1 889
Tosia (PUn/haster) compla Verrill, 1 899
Tosia (Plinthaster) nilida Verrill, 1 899
Pseudarchaster gracilis subsp. gracilis (Sladen, 1 889)
= Aphroditasler gracilis Siaden, 1 889 = ASlrogonium aequabile Koehler, 1 907
Aslrogonium aphrodite Perrier, 1 894 = Astrogonium eminew Koehler, 1 907
ASlrogonium marginalum Koehler, 1 909 Astrogonium necator Perrier, 1 894
= P. cancinnus Verril l , 1 894 P. granuliferus Verrill, 1899
= P. ordina/us Verrill, 1 899 parelii (D!lben & Koren, 1 846)
Aslrogonium annectens Perrier, 1 894 = Astrogonium/allax Perrier, 1 885
As/rogonium hystrix Perrier, 1 894 Aslrogonium longobrachia/e Koehler, 1 907
= Astropecfen parelii Dilben & Koren, 1 846 P. intermedius var. insignis Verrill, 1 895
= P. intermedius Siaden, 1 889 = P. tessel/alus var. arcticus Sluiter, 1 895
Sphaeriodiscus bourgeli (Perrier, 1 885) = Slephanasier bourgeti Perrier, 1 885
Family Ophidiasteridlle Hazelia
a/fenuala Gray, 1840 = Aslerias coriacea Grube, 1 840 = Asterias laevigala subsp. varielas de Lamarck, 1 8 1 6 = ASlerias variolala Risso, 1 826, non Retzius, 1 805 = Ophidiasler (Hazelia) atfenualus Gray, 1 840
Ophidiaster lessonae Gasco, 1 876 Narcissia
canariensis (d'Orbigny, 1 839) = A slerias canariensis d'Orbigny, 1 839 = Fromia narcissae Perrier, 1 885
Fromia narcissiae Perrier, 1 894 N. lenerifJae Gray, 1 840
Irigonaria Sladen, 1 889 Ophidiaster
ophidianus (de Lamarck, 1 8 1 6) = Asterias ophidiana de Lamarck, 1 8 1 6
O. aurantius Gray, 1 840 = O. canariensis Greeff, 1 872 reyssi Sibuet, 1 977
Family POfaniidae Chand raster
grandis (Verrill, 1 878) = ?Marginasler fimbrialus Siaden, 1 889 = Porania grandis Verrill, 1 878
Marginasler capreensis (Gasco, 1 876) = Asterops/s capreensis Gasco, 1 876
Porania
- 340 -
(Porania) pu/villus subsp. pulvillus (O.F. Maller, 1 776)
= Asleropsis clenacanlila J. Maller & Troschel, 1 842 = Ganiaster templeloni Thomson, in Forbes, 1 839
P. gibbosa Gray, 1 840 (nomen nudum) (Pseudaporania) starmi(Dons, 1 935) = Pseudaparania stormi Dons
Poraniomorpha (Poraniomorpha) bidens Mortensen, 1 932 (Poraniomorpha) hispida subsp. hispida (M. Sars 1 872)
,
= ?Rhegasler murrayi Sladen, 1 883 = Asterina borealis Verrill, 1 878 = Goniaster hispidus M. Sars, 1 872 = Parania spinulosa Verrill, 1 880 (Poraniamorpha) hispida subsp. rosea Danielssen & Koren, 188 1
= Lasiaster villosus Siaden, 1 889 = P. rosea Danielssen & Koren, 1 88 1 (Poraniomorpha) tumida (Stux.berg, 1 878)
Rhegasler lumidus (Stuxberg) auctt. = So/asler tumida Stux.berg, 1 878 = Sphaeriasler bjoerklykkei (Dons, 1 938) (Culci/opsis) borealis (SOssbach & Breckner, 1 9 1 1 )
? Marginaster perllagonlls Perrier, in Milne-Edwards, 1882 CII/cila borealis SOssbach & Breckner, 1911 Sphaeraster berthae Dons, 1938
= Sphaer/aster berthae Dons, 1939 Ty/aster
wille; Danielssen & Koren, 1881
Family Sphaerasterldae Podosphaeraster
tha/assae Cherbonnier, 1970
Family Asterlnidae A lIopatiria
ocelllfera (Gray, 1847) = ?Palmipes lobiancoi Ludwig, 1897
Parasterina africana Engel & eroes, 1960 Paliria ocelllfera Gray, 1847
= Patiria pulla Koehler & Vaney, 1906 = Patiria rosea Koehler & Vaney, 1906
Anseropoda lobiancoi (Ludwig, 1897)
Palmipes lobiancoi Ludwig, 1897 placenta (Pennant, 1777) = Asterias carti/aginea Fleming, 1828 = Aster/as membranacea Retzius, 1783 = Aster/as placenta Pennant, 1777 = Aster/scus palmi pes J. MOller & Troschel, 1842
Asterina gibbosa (Pennant, 1777)
Asterias gibbosa Pennant, 1777 = Asterias verruculata Retzius, 1805 = A. minllta Nardo, 1834 = AsterisellS arrecijiensis Greeff, 1872 pancerli (Gasco, 1870) = AsterisellS paneerii Gasca, 1870 phylaeliea Emson & Crump, 1979 wega Perrier
Tremaster mirabilis Verrill, 1880
T. lawis H.L. Clark, 1941
Order Spinulollida
Family Echinasteridllc Echinaster
(Eehinaster) sepositus subsp. sepositus (Retzius, 1783)
= Asterias sagena Retzius, 1805 = Asler/as seposita Relzius, 1783 = Rhopia mediterranea Gray, 1840
Henricia caudani (Koehler, 1895) = Cribella caudani Koehler, 1895 cylindrel/a (Sladen, 1883) = Cribrella abyssa/is Perrier, 1894 = Cribrella oculala var. eylindrella Siaden, 1883
H. abyssicola Perrier, 1885 & Farran, 1913 & Mortensen, 1927
esehrichti (J. MOiler & Troschel, 1842) = Eehinaster esehriehti J. MUller & Trosehel, 1842 = Magda/enaster arct/cus Koehler, 1907 hedingi Madsen, 1987 lisa A.H. Clark, 1949 lisa subsp. ingolfl Madsen, 1987 ocu/ata (pennant, 1777) = Asterias oeulata Pennant, 1777
Ii. sanguinolenta auelt., non (O.F. MOiler) (p.p.) per/orata (O.F. MUller, 1776) = Asterias per/orata O.F. MOiler, 1776 = Cribrefla oeulata var. murina Perrier, 1875 = Cribrefla sanguino/enla f. seabrior Michailovskij, 1902 = Echinaster serobieulalus Danielssen & Koren, 1883
Ii. sanguino/enta auett., non (O.F. Muller) = Ii. cross/spika Heding, in Jltgerski61d, 1971 (nomen
nudum) = Ii. fisheri subsp. echinala Heding, in J!lgerski6Id, 1971
(nomen nudum)
European Register of Marine Species
= H. knipowitschi D'yakonov, 1950 Ii. sanguinolenta Rasmussen, 1965 non (O.F. Mllller)
pertusa (O.F. Mllller, 1776) ?Cribrefla biseayensis Koehler, 1895
= Asterias pertllsa O.F. MOiler, 1776 = Cribrella sanguinoienta var. abyss/cola Norman, 1869
Eehinaster oelllatus f. major Dilben & Koren, 1846 sanguinolenta (O.F. Mllller, 1776) = Asterias sanguinolenta O.F. MOiler, 1776
Cribrella pectinata Verrill, 1894 = Cribrella sanguino/enta var. curta Norman, 1869
Cribrella sanguinolenta f. lawior Michailovskij, 1902 = Echinaster oculatlls f. minor Dilben & Koren, 1846 = Echinaster sarsi J. MUlier & Troschel, 1844
Ii. pCI/orata Rasmussen, 1965 non (O.F. MUller) = Ii. sanguinoienla subsp. miliaris Verrill, 1914 skorikovi D'yakonov, 1950 solido D'yakonov, 1950 spongiosa (0. Fabricius, 1780)
Order Velatida
Family Soilisteridae Crossaster
papposus (Linnaeus, 1767) Asterias qffinis Brandt, 1835
= Asterias helianthemoides Pennant, 1777 = Asterias papposa Linnaeus, 1767 sqllamallls (Dllderlein, 1900) = So/asler affinis Danielssen & Koren, 1877 non (Brandt,
1835) = SO/OSler papposus var. squamata D1lderlein = So/asler squamatus D6derlein
Lophaster /urelfer (DUben & Koren, 1846) = Solaster /urclfer Dilben & Koren, 1846
Solaster ear/li Verrill, 1879 endeca (Linnaeus, 1771)
Aster/as alboverrueosa Brandt, 1835 = Asterias aspera O.F. MOiler, 1776 = Asterias endeea Linnaeus, 1771 = Asterias rumphii Parelius, 1768 = S. endeca subsp. deeemradiata Siaden, 1889
S. galaxides Verrill, 1909 = S. intermedius Sluiter, 1895 glaeialis Danielssen & Koren, 1881 = S. echinatus Storm, 1888 syrtensis Verrill, 1894
Family Korethrasteridae Korethrasler
h/spidlls Wyville Thomson, 1873 K. selosus Perrier, in Milne-Edwards, 1882
Family Plerasteridae
- 341 -
Diplopteraster multi pes (M. Sars, 1866) = Pterasler mllil/pes M. Sars, 1866
Hymenaster giboryi Perrier, 1894 pellucidus Thomson, 1873 = Ii. giganteus Siaden, 1891 = H. membranaceus Thomson, 1877 = H. perissonotus Fisher, 1910 regalis Verrill, 1895 = H. kieri Downey, 1979 reticulatus Sibuet, 1976 rex Perrier, 1885 = H. gennaeus H.L. Clark, 1923 rosells Koehler, 1907 tenllispinlls Sibuet, 1976
Pleraster ac/cllia (Downey, 1970) = Marsipasler acicula Downey, 1970 a/veolaills Perrier, 1894 hystrix Harvey, 1989 mifital'is (O.F. MUlier, 1776)
Costello, Emblow and White (eds), 200 I
= Aslerias militaris O.F. MUlier, 1776 = P. aporus Ludwig, 1 886 obscurus (Perrier, 1 89 1 ) = Hexasler obscurus Perrier, 1 89 1 = P. (Temnasler) hexaclis Verrill, 1 894 personalus Sladen, 1 89 1
?Plerasler sordidus Perrier, 1 894 = P. reduclus Koehler, 1907 pulvil/us (M. Sars, 1 86 1 )
Family Myxasteridae Myxaster
perrieri Koehler, 1 896 sol Perrier, 1 885
Pylhonasler al/antidis A.H. Clark, 1948
Order Brisingida
Family Brlsingidae Brisinga
endecacnemos Asbjemsen, 1 856 hirsula Perrier, 1 894
Brisingelia coronala (0.0. Sars, 1 87 1 ) = Brisinga coronala 0.0. Sars, 1 87 1 = Brisingia mediterranea Perrier, 1 885
Novodinia homonyma Downey, 1 986
Brisinga e/egans Perrier, 1 885, non Verrill, 1 884 = Odinia elegans Perrier, 1 894 pondina (Sladen, 1 889) Downey, 1986
Odinia pandina Siaden, 1 889 semicoronala (Perrier, 1 885) Downey, 1 986
Brisinga robusta Perrier, 1 885 = Brisinga semicoronata Perrier, 1 885
Family Freyellidae Co/paster
edwardsi (Perrier, 1 882) Downey, 1986 = Brisinga edwardsi Perrier, 1 882 scutigeru/a Siaden, 1 889
Freyaslera benlhophila (Sladen, 1 889) Downey, 1 986
Freyelia benthophila Siaden, 1 889 sexradiata (Perrier, 1 885) Downey, 1 986 = Freyel/a sexradiala Perrier, 1 885 luberculata (Sladen, 1 889) Downey, 1 986 = Freyel/a tuberculala Siaden, 1 889
Freyella elegans (Verrill, 1 884) = Brisinga elegans Verrill, 1 884
F. aspera Verrill, 1 895 = F. bracleala Siaden, 1 889 = F. laubieri Cherbonnier & Sibuet, 1973
F. spinosa var. abyssicola Perrier, 1894 = F. spinosa Perrier, 1 885 recta Koehler, 1 907
Order ForcipuJatida
Family Asleriidae Asteria.
rubens Linnaeus, 1 758 A. clalhrala Pennant, 1 777
= A. holsatica Retzius, 1 805 = /I. minula Linnaeus, 1 76 1 = A . rubens subsp. violacea O.F. MUlier, 1 776
= A. spinosa Say, 1 825 = A. vulgar/s Packard, 1 863
Cose/naslerias lenuispina (de Lamarck, 1 8 1 6) = /lslerias at/anlica Verrill, 1 868 = Asler/as savaresi Delle Chiaje, 1 827
Asler/as lenu/spina de Lamarck, 1 8 1 6 = Lylaster inaequalis Perrier, 1 894
/casterias panopla (Stuxberg, 1 878) = Asterias panopla Stuxberg
Leplaslerlas canuli Heding, 1 936 clavispina Heding, 1 936 (Leptasterias) danica (Levinsen, 1 887) degerboelli Heding, 1 935 floccosa (Levinsen, 1 8??) floccosa var. crassa Heding, 1 935 (Endogenasterias) groenlandica (Steenstrup, 1 857) = ASleracanthion groenlandica Steenstrup, 1 8057
Asterias cribraria Stimpson, 1 862 (Leplaslerias) hyperborea (Danielssen & Koren,
1 883) Aslerias hyperborea Danielssen & Koren, 1 883
= Asler/as normani Danielssen & Koren, 1 884 hyperborea var. /nlermedia Heding, 1935 (Leptaslerias ) islandlca (Levinsen, 1 887) = Aster/as mulleri var. /slandica Levinsen, 1 887 (Leplaster/as) muelleri subsp. muelleri (M. Sars,
1 846) = Asteracanth/on mulleri M. Sars, 1 846 (Leplaster/as) muelleri subsp. ce/tica Clark & Downey, 1992
= Aster/as hispida Pennant, 1 777 (Hexaslerias) polaris (J. Mllller & Troschel, 1 842) = ASleracanthion polaris J. MOller & Troschel, 1 842
Asler/as acervata Stimpson, 1 862 Aster/as borealis Perrier, 1 875
Marthaster/as giacialis (Linnaeus, 1758) = Asteracanth/on afticanus J. MUlier, & Troschel, 1 842 = Asler/as angulosa O.F. MUlier, 1 788
Aster/as echinophora Delle Chiaje, 1 827 = Aster/as glacialis Linnaeus, 1 758 = Aster/as madelrensls Stimpson, 1 862
Asler/as rarispina Perrier, 1 875 = /lslerias spinosa Pennant, 1777
M foliacea Jullien, 1878 = Sielionia webbiana d'Orbigny, 1 839
Scleraslerias guerne/ Perrier, 1 89 1 neglecta (Perrier, 1 89 1 ) = Aslerias edmundi Ludwig, 1 897
Sto/asterias neg/ecta Perrier, 1 89 1 richard! (Perrier, 1 882)
Aster/as richard/ Perrier, 1 882 Stephanasterlas
albula (Stimpson, 1 853) Asleracanlhion albulus Stimpson, 1 853
= Asteracanlhion problema Steenstrup, 1855 = Asterias gracilis Perrier, 1 88 1
Stichastrella ambigua (Farran, 1 9 1 3)
Stichaster roseus var. ambiguus Farran, 1 9 1 3 rosea (O.F. MUl ler, 1 776) = Asterias rosea O.F. MOiler, 1776
Chaelaster hermann; 1. Mllller & Troschel, 1 842 = Stichaster arcticus Danie1ssen & Koren, 1 883
Urasterias linckii (J. Mlllier & Troschel, 1 842) = Asteracanthion Jinckii J. MUller & Troschel, 1 842
ASleracanthion slellionura E. Perrier, 1869 = Aslerias enop/a Verrill, 1 895 = Aslerias gunner/ Danielssen & Koren, 1 884
Family Pedicellasteridae Hydraslerias
sexradiala (Perrier, in Milne-Edwards, 1 882) Pedicel/asler sexradiatus Perrier, in Milne-Edwards, 1 882
Pedicel/asler typicus M. Sars, 1 86 1 = Archasler biftons W . Thomson, 1 873 = Astel'acanlhion palaeocrystallus Duncan & Siaden,
1 877
Family Zoroasleridae Zoroaster
- 342 -
/u/gens Thomson, 1 873
Family Labldiasteridae Coronaster
volsel/alus (Sladen, 1 889) = Asterias (Slo/aslerias) valsella/us Sladen, 1 889 = C. antonii Perrier, 1 894
Family Neomorphasteridae Neomorphasler
margaritaceus (Perrier, in Milne-Edwards, 1 882) A.M. Clark & Downey, 1992
Calycaster monoecus Perrier, 1 89 1 = N. eustichus Siaden, 1 889 = N. par/aiti Koehler, 1 896 = Pedicellaster margarilaceus Perrier, in Mi lne-Edwards,
1 882 = Stichasler talisman; Perrier, 1 88S
Class Ophiuroidea
Order Phrynophiurida
Family Ophiomyxidae Aslrogeron
supinus (Lyman, 1 883) Ophiogeron supinus Lyman, 1 883
Ophiogeron edenlulus Lyman, 1 878
Ophfomyxa penlagona (de Lamarck, 1 8 1 6) = Ophfura penlagona de Lamarck, 1 8 1 6 serpentaria Lyman, 1 883
Ophiophrixus spinosus (Storm, 1 8 8 1 ) = Ophiobyrsa hyslricis Lyman, 1883
Ophfobyrsel/a hyslricis (Lyman, 1 883) Ophioscolex
g/acia/is J. MUller & Troschel, 1 842 purpureus DUben & Koren, 1 846
?Ophiosco/ex quadrispinus Verrill, 1 885
Order Euryalina
Family Asteronycbidae Asleronyx
loveni J. MUlier & Troschel, 1 842 = A. cooped Bell, 1 909 = A. dispar LU!ken & Mortensen, 1 899
A. /ocardi Koehler, 1 896 = Ophiuropsis Iymani Studer, J 884
Astrodia lenuispina (Verrill, 1 884) = A. bispinasa Koehler, 1 922
Family Gorgonocepbalidae Astrosparlus
medilerraneus (Risso, 1 826) = Euryale arborescens L. Agassiz, 1 839 = Euryale medilerraneus Risso, 1 826
Gorgonocepha/us verrucosus Grube, 1 840 Gorgonocephalus
arclicus Leach, 1 8 1 9 = Astrophylon agassizi Stimpson, 1 853 capu/medusae (Linnaeus, 1 758) = Aslerias caput-medusae Linnaeus, 1 758
Astrophyton linckii I. MUller & Troschel, 1 842 = Astrophyton sculalum Fleming, 1 828 eucnemis (I. MUlier & Troschel, 1 842) Lyman, 1 882 = Astrophyton eucnemis I. Milller & Troschel, 1 842 = Astrophylon malmgren; Danielssen & Koren, 1 877 = G. caryi Lyman, 1 860
G. japonicus Dliderlein, 1907 /amarckii (I. MUller & Troschel, 1 842) Lyman, 1 882 = Astrophyton lamarcki J. MUlier & Troschel, 1 842
Family Asteroschematidae Asteroschema
inornatum Koehler, 1 9?? Ophiocreas
oedipus Lyman, 1 879
European Register of Marine Species
Order Ophiurida
Family Opbiotricbidae Ophiolhrix
fragilis (Abildgaard, in O.F. MUller, J 789) = Asteria cuvieri Delle Chiaje, 1 828
Aslerias echinala Delle Chiaje, 1 822-29 = Asteriasfragifis Abildgaard, in O.F. MUlier, 1 789
Aslerias penlaphylla Pennant, 1 777 = Aslerias rllbra Delle Chiaje, 1 8?? = Ophiocoma minula Forbes, 1 839 = Ophiocoma rosula Forbes, 1 839 = O. alopecurus I. MUller & Troschel , 1 842 = o. /erllssacii J. MUller & Troschel, 1 842 = O. lusitanica Ljungman, J 872 = 0. rammefsbergii J. MUlier & Troschel, 1 842
0. rubra Ljungman, J 872 = Ophiura qllinquemaculala Delle Chiaje, 1 828 indigna Koehler, 1 906 luetken; Wyville Thomson, 1 873 macu/ala Ljungman, 1 872 = 0. indllcla Koehler, 1 906
Family Opbiactidae Hislampica
dllplicata (Lyman, 1 875) Ophiaclis
abyssicola (M. Sacs, 1 86 1 ) Ljungman, 1 866 = ?Ophiactis cuspidala Lyman, 1 882 = Amphiura abyssicola M. Sars, 1 86 1 = O. corallicola Koehler, 1 896
0. echinala Koehler, 1 898 = O. poa Lyman, 1 882 balli (W. Thompson, 1 840) = Ophiocoma balli W Thompson, 1 840 = Ophiocoma goadsiri Forbes, 1 840 (Nov.) canotia Lyman, 1 882 /ymani Ljungman, 1 872 nidarasiensis Mortensen, 1 920 pro fundi LU!ken & Mortensen, 1 899 savignyi (1. MUller & Troschel, 1 842)
O. brocki de Loriol, 1 893 = O. krebsii LUtken, 1 857
Ophiolepis savignyi J . MUlier & Troschel, 1 842 virens (M. Sars, 1 8S7) = Amphiura virens M. Sars, 1 857
Ophiopholis aculeala (Linnaeus, 1 767) = Ophiocama bellis Forbes, 1 839
Ophiolepis sc% pendrica 1. MUlier & Troschel, 1 842 � Polypholis echinala Duncan, 1 8??
Ophiopus arcticus Ljungman, 1 867 � Ophiaregma abyssorum G.O. Sars, 1 872
Family Amphiuridae
- 343 -
Amphichilus da/eus (Lyman, 1 882) Matsumoto, 1 9 1 7 � Amphiura dalea Lyman, 1 882
Amphioplus verrilli (Lyman, 1 879) � Silox pu/vinus Cherbonnicr & Sibuet. 1 972
Amphipholis instructa Koehler, 1906 murmanica D'yakonov, 1 929 squamata (Delle Chiaje, 1 829)
A. lineala Ljungman, 1 872 � A. tenera LUtken, 1 8??
Amphiura e/egans (Leach, 1 8 1 5) = Asterias squamala Delle Chiaje, 1 829 = Axiognalhus squamala (Delle Chiaje) torelli Ljungman, 1 872
Amphiura abyssorum Norman, in Jeffreys, 1 876 = ? A. digna Koehler, 1906 bellis Lyman, 1 879 bellis var. Irilonis Hoyle, 1 884
Emblow and White 200 I
borealis (G.O. Sars, 1 87 1 ) Ljungman, 1 872 = Ophiope/tis borealis G.O. Sars, 187 l (Acrocnida) brachiata (Montagu, 1 804) = Amphiocnida brachiata (Montagu) = A. neapolitana M. Sars, 1 857 = Asterias brachiala Montagu, 1 804 cherbonnieri Guille, 1 972 chiajei Forbes, 1 843 defamare; Cherbonnier, 1 958? denticulata Koehler, 1 89 filiformis (O.F. MUlier. 1 776) = Asleriasfiliformis O.F. Mill ler, 1 776 fragilis Verrill, 1 885
= A. denticulala Koehler, 1 898 grandisquama Lyman, 1 869 = A. iris Lyman, 1 879 = A. josephinae Ljungman, 1 872 = A. longispina Koehler griegi Mortensen, 1 920 incana Lyman, 1 879 mediterranea Lyman, 1 882 olleri Ljungman, 1 872 = A. grandis Koehler, 1 896 palmeri Lyman, 1 882 richardi Koehler, 1 906 sarsi Ljungman, 1 872 (Ophiope/tis) securigera (Dilben & Koren, 1 846)
Ophiopeltis securigera DUben & Koren, 1 846 (Acrocnida) sem/squamata (Koebler) senega/ensis Madsen, 1 970 stepanovi D'yakonov, 1 954 = A. stepanovi Chernyavskii, 1 86 1 (nomen nudum)
A. stepanovi Lyman, 1 869 (nomen nudum) sundevalli O. Millier & Troschel, 1 842) = Ophiofepis sundevalli J. Milller & Troschel, I S42
Monamphiura apicula (Cherbonnier, 1 957) = Amphiura apicula Cherbonnier, 1 957
Paramphlura punelala (Forbes, 1 84 I )
Ophiaeoma punetata Forbes, 1 84 1
Family Ampbilepididae Amphilepis
norveglca (Ljungman, IS65) = Amphiura norvegica Ljungman, 1 865 ingolfiana Mortensen, 1 93 3
Order Laemophiurina
Family Ophiacanlbidae Ophlacantha
abyssicola G.O. Sars, 1 87 1 O. mil/espina Verrill, 1 879
angolensis Koehler, 1 923 anomala G.O. Sars, 1 87 1
?Ophiacantha nodosa Lyman, I S?? arislata Koehler bidentata (Retzius, 1 805)
Asterias bidentala Retzius, 1 805 = 0. fratema Verrill, 1 885
0. groenlandica J. MUller & Troschel, 1 844 = O. hibernica Farran, 1 9 1 3 = 0. spinulosa J . MUlier & Troschel, 1 842 brevispina Koehler, 1 898? composifa Koehler, 1 907 cra.fsidens Verrill cuspidata Lyman, 1 882 (Ophiotreta) danae Mortensen, 1 933
0. valenciennesi Lieberkind, 1 929, non Lyman decipiens Koehler, 1 906 densa Farran, 1 9 1 3? enopia subsp. velerna (Koehler, 1 907) hirsllta Lyman, 1 875 lineata Koehler, 1 909 mesembria H.L. Clark, 19 15 nodosa Lyman, 1 882
- 344 -
nolata Koehler, 1 906 opercu/aris (Koehler, 1 909) = Ophiolimna opercularis Koehler, 1 909 parcita Koehler, 1906 setosa (Retzius, I S05)
Ophiura selosa Retzius, 1 805 smitti Ljungman, 1 872 slmulans Koehler, 1 896
O. composita Koehler, 1 906, non Koehler, 1904 = 0. composila Mortensen, 1 927 spec/abi/is G.O. Sars, IS71 spinosella Mortensen, 1933 valenciennesi Lyman, 1 879
Ophiambix devaneyi Paterson, 1 985 meteoris Bartsch, 1 983
Ophiochondrus armalus (Koehler, 1 909) = Ophioplus armalus Koehler, 1 909
Ophiocomina nigra (Abildgaard, in O.F. Milller, 1 789)
Asterias nigra Abildgaard, in O.F. Milller, 1 789 Ophiacanlha danae Mortensen, 1 933
= Ophiocoma granuiata Forbes, 1 839 Ophiocoma nigrll (Abildgaard)
= Ophiocoma nilssoni J. MUlier & Troschel, 1 842 = Ophiocoma raschi G.O. Sars, IS??
Ophiolebes bacata Koehler, 1909 retecta (Koehler, 1 896)
Ophioscolex retectus Koehler, 1 896 Ophiomitra
hamula Mortensen, 1 933 Ophiopsila
annulosa (M. Sars, 1 859) Ophianoplus annulosus M. Sars, 1 859
aranea Forbes, 1 845 = Ophianoplus marmoreus M. Sars, 1 8?? alT. guineensls (Koehler, 1 9 1 4)
Ophiocamax dominans Koehler, 1 9??
Ophiomifrella clavigem (Ljungman, 1 865)
?Ophiomitrella cordifera Koehler, 1909 = Ophiuclis clavigera Ljungman, I S65 = Ophio/ebes acanellae Verrill, I S85
Ophiolebes claviger (Ljungman, 1 865) globulifera (Koehler, 1 896)
Ophiop/inlhuca carduus (Lyman, 1 882) chelys (Lyman, 1 882)
Ophiothamnus ajJinis Ljungman, 1 872 = Ophioleda minima Koehler, 1 906
Ophioplinthaca occlusa Koehler, 1 907 = O. minima Matsumoto, 1 9 1 7 charUs H.L. Clark, 1 9 1 5
O. gracilis (Verrill, 1 885), non (Studer, 1 822) Ophlolimna
bairdi (Lyman, 1 883) Verril l , 1 899 = Ophiacantha bairdi Lyman, 1 883
Ophiomedea duplicata Koehler, 1906
Ophioprium rosea (Lyman, 1 879) Baker & Devaney, 1981
Ophiacanlha rosea Lyman, 1 879 = Ophiacantha t/'lmcata Koehler, 1 930
Ophiotoma bartlelli (Lyman, 1 8??) coriacea Lyman, 1 883
Ophiolrema alberti Koehler, 1 896
Ophiomyces frutectosus Lyman, 1 869 grandis Lyman, 1 878
Order Chilophiurina
Family Ophiuridae Amphiophiura
bullata (Wyville Thomson, 1 873) Matsumoto, 1 9 1 5 = Ophioglypha abdita Koehler, 1 909
Ophioglypha bul/ala Wyville Thomson, 1 873 bul/ala subsp. eonvexa (Lyman, J 878) = Ophioglypha convexa Lyman, 1 878 bullara subsp. viljazi Litvinova, J 971 = A. viljazi Litvinova, 1971 saurura (Verrill, J 894) = Ophioglypha aspera Koehler, 1 898 = Ophioglypha saurura Verrill, J 894
Anlhophiura ingolft Fasmer, 1 930
Aspidophiura minilia (Lyman, 1 878) = Ophiog/ypha minuta Lyman, 1 878
Homophiura abyssorurn (Lyman, 1 883) abyssorum var. africana (Madsen, 1 947)
Homalophiura abyssorum var. afrieana Madsen, 1 947 lessellata (Verrill, 1 894) = Homalophiura tessel/ata (Verrill, 1 894)
Ophioglypha tessellala Verrill, 1 894 Ophiocten
abyssicolum von Marenzeller, 1 8?? affinis (LUtken, 1 858) Sumida, Tyler, Gage & N0rrevang, 1 998
Ophiura affinis Liitken, 1 858 cenlobi Paterson, Tyler & Gage, 1 982 gracilis (G.O. Sars, 1 87 1 )
0. serieeum auctt., non (Forbes, 1 852) (p.p.) = Ophiog/ypha gracilis G.O. Sars, 1 8 7 1 = Ophiog/ypha signala Verrill, 1 882
Ophiura signata (Verrill, 1 882) haslatum Lyman, 1 878
o.phioclen longispinum Koehler, 1 898 = O. /atens Koehler, 1 906 = O. pacifieum Latken & Mortensen, 1 899 sculalum Koehler, 1 898 serieeum (Forbes, 1 852) = 0. kroeyeri Liltken, 1 855
Ophiura sericea Forbes, 1 852 Ophioclenella
ades Tyler, Paterson, Sibuet, Guille, Murton & Segonzac, 1995
Ophio/epis paucispina ?
Ophiomisidium pulchellum (Wyville Thomson, 1 8??)
Ophiomusium pulchellum Wyville Thomson, 1 8?? Ophiomusium
africanum Koehler, 1909 /ymani Wyville Thomson, 1 873
Ophiophycis mirabilis Koehler, 1 90 I
Ophiopleura aurantiaca (Verrill, 1 882) Verrill, 1 882
Ophioglypha aurantiaea Verrill, 1 882 borealis Danielssen & Koren, 1 877 = Luetkenia arc/iea Duncan, 1 878
Ophiosphalma armigerum (Lyman, 1 878) H.L. Clark, 1941 = Ophiomusium armigerum Lyman, 1 878 planum (Lyman, 1 878) H.L. Clark, 1 94 1 = Ophiomusium planum Lyman, 1 878
Ophiotjalfa vivipara Mortensen, 1 9 1 3
Ophiotypa simplex Koehler, 1 897
Ophiozonella molesfa (Koehler, 1 906)
Ophiozona molesfa Koehler, 1 906 sineera (Koehler, 1 906)
European Register of Marine Species
= Ophiozona sincera Koehler, 1 906 tjalftana (Mortensen, 1 9 13) Matsumoto, 1 9 1 5
Ophiozona tjalftana Mortensen, 1 9 1 3 Ophiura
albida Forbes, 1 839 africana (Koehler) = Ophiocten africana Koehler (Dictenaphiura) carnea LUtken, 1 858 ex M. Sars MS clemens (Koehler, 1 906) = Ophioglypha clemens Koehler, 1 906
jlagellala (Lyman, 1 878) = Gymnophiura caerulescens Lmken & Mortensen 1 899 = Ophiog/yphaj/agellata Lyman, 1 878
'
grubei Heller, 1 863 imprudens (Koehler, 1 906)
Ophioglypha imprlldens Koehler, 1 906 inermis (Lyman, 1 882) = Ophiog/ypha inermis Lyman, 1 882 (Ophiuroglypha) irrorata (Lyman, 1 878) = Ophioglypha grandis Verrill, 1 894 = Ophioglypha involuta Koehler, 1 899
Ophioglypha irrorala Lyman, 1 878 = Ophioglypha orbiculala Lyman, 1 882 = Ophioglypha lumu/osa Liitken & Mortensen, 1 899 (Ophiuroglypha) irrorata subsp. concreta (Koehler
1 90 1 ) ,
= Ophiog/ypha concreta Koehler, 1 90 1 (Ophiuroglypha) irrorata subsp. po/yacantha Mortensen, 1 933 Homalophiura mullispina Cherbonnier & Sibuet 1 972
(jungmani (Lyman, 1 878) ,
= Ophioglypha /epida Lyman, 1 882 Ophiog/ypha (jungmani Lyman, 1 878
= Ophiog/ypha Ihouleli Koehler, 1 896 madseni (Belyaev, 1 972) = Homalophiura madsen; (Belyaev, 1 972) (Ophiurog/ypha) mundata Koehler, 1 906 nilida Mortensen, 1 933 ophiura (Linnaeus, 1 758) = Asterias ophiura Linnaeus, 1 75 8
0. cWaris (Linnaeus, 1 766) = 0. ciliala (Retzius, 1 783) non (O.F. MUlier, 1 776) = O. lacerlosa (Pennant, 1 777)
O. lexlurala de Lamarck, 1 8 1 6 robllsla (Ayres, 1 8 5 1 ) = Ophiolepis robus/a Ayres, 1 8 5 1
O. /ascicu/ala Forbes, 1 854 = O. squamosa LUtken, 1 859 sarsii LUtken, 1 858 violaine (Cherbonnier & Sibuet, 1 972) = Homalophiura vio/aine Cherbonnier & Sibuet 1 972
Ophiurolepis '
inornata (Lyman, 1 878) Paterson, 1 985 = Homalophiura inornala (Lyman, 1 878)
Ophioglypha indivisa LUtken & Mortensen, 1 899 = Ophioglypha inornata Lyman, 1 878
Slegophiura nodosa (LUtken, 1 858) Matsumoto, 1 9 1 7 = Ophiura nodosa Liltken, 1 858 stuwilzii (LUtken, 1 858) Matsumoto, 1 9 1 7 = Ophiura slllwilzii LUtken, 1 85 8
Uriopha ios Paterson, 1 980
Family Ophioleucidae Ophiernus
adspersus Lyman, 1 883 vallincola Lyman, 1 878
O. abyssalis Koehler, 1 896 Ophiocrates
interval/us Madsen, 1 947 secundus Koehler, 1 906
Ophiopyren striatum Mortensen, 1 933
Family Ophionereididae Ophiochiton
- 345 -
Costello, Emblow and White (eds), 200 1
solutus Koehler, 1 906 lern;spinus Lyman, 1 883 = 0. grandis Verrill, 1 884
O. tenuispinus Hoyle, 1884 Ophionereis
reticulata (Say, 1 825) = Ophiura reticulata Say, 1 825
Family Ophiodermatidae Bathypectinura
heros (Lyman, 1 879) = B. elata (Koehler, 1 906) = Pectinura elala Koehler, 1 906 = Pectinura heros Lyman, 1 879
Cryptopelta brcvispina (Ludwig, 1 879)
Ophioconis brevispina Ludwig, 1 879 Ophiarachnella
scmicincla (Studer, 1 8??) = Peetinura semicincta Studer, 1 8??
Ophioconis forbesi (Heller, 1 863)
Pectinuraforbesi Heller, 1 863 vivipara Mortensen, 19??
Ophioderma longicauda (Retzius, 1 805)
Asterias longicauda Retzius, 1 805 = O. lacertosum Lamarek, 1801
Ophiura laevis Lyman, 1 865 Pectinura
vestita Forbes, 1 843
Class Echinoidea
Order Cidaroida
Family Cidaridac Cidaris
cidaris (Linnaeus, 1 758) = C. borealis DUben & Koren, 1 846 = C. papil/ala Leske, 1778 = Cidarites hyslrix de Lamarck, 1 8 1 6 = Dorocidaris abyssicola A . Agassiz, 1 8697 = Echinus cidaris Linnaeus, 1758 Iribu/oides de Lamarck, 1 8 1 6
Poriocidaris purpurala (Wyville Thomson, 1875) = Porocidaris gracilis Siaden, 1891 = Porocidaris purpurata Wyville Thomson, 1875
Siereocidaris ingolfiana Mortensen, 1 903
Stylocidaris affinis (Philippi, 1 845) = Cidaris affinis Philippi, 1 845
Cidaris stokesii L. Agassiz & DesoT, 1 846-47
Order Ecbinotburoida
Family Echinotburidae Calveriolloma
feneslralum (Wyville Thomson, 1872) = Araeosoma violaceum Mortensen, 1 903 = Calveriafeneslrala Wyville Thomson, 1 872 hystrix (Wyville Thomson, 1 872) = Aslhenosoma reynoldsii A. Agassiz, 1880
Calveria hyslrix Wyvil le Thomson, 1 872 Hygrosoma
pelersii (A. Agassiz, 1880) = Phormosoma luculenlum Koehler, 1 896, non A.
Agassiz, 1 8?? = Phormosoma peters;; A. Agassiz, 1 880
Phormosoma uranus Koehler, 1898, non Wyville Thomson, 1 877
uranus (Wyville Thomson, 1 877) = Phormosoma uranus Wyville Thomson, 1 877
Phormosoma placenta Wyville Thomson, 1 874
P. sigsbei A. Agassiz, 1 883 Sperosoma
grima/dii Koehler, 1 897 Tromikosoma
koehleri Mortensen, 1 903
Order Diadematoida
Family Aspidodiadematidae Aspidodiadema
jacobyi A. Agassiz, 1 883 Plesiodiadema
anlil/arum (A. Agassiz, 1 883) = Aspidodiadema antil/arum A. Agassiz, 1883
Family Diadematidae Cenlrostephanus
longispinus (Philippi, 1 845) Cidaris (Diadema) longispina Philippi, 1 845
Diadema anlil/arum Philippi, 1 845
Cidaris {Diadema} antil/arum Philippi, 1 845
Order Pedinoida
Family Pedinidae Caenopedina
cubensis A. Agassiz, 1 869
Order Salenoida
Family Saleniidae Sa/enia
patterson; A. Agassiz Salenocidaris
profundi subsp. profundi (Duncan, 1 877) = ?Sa/enia hastigera A. Agassiz
Salenia hastigera subsp. aeuminala Mortensen, 1935 = Sa/enia profundi subsp. oec/usa Mortensen, 1934 = Sa/enia profundi Duncan, 1 877 profundi subsp. brevispina Markov, 1988 varispina A. Agassiz, 1 869
Order Arbacioida
Family Arbaciidae Arbaeia
lau/a (Linnaeus, 1 758) Cidaris puslu/osa Leske, 1 778
= Echinus aequitubercu/atus de Blainville, 1 825 Echinus /ixu/a Linnaeus, 1 758
= Echinus neapolitanus Delle Chiaje, 1 825 Arbaciel/a
e/egans Mortensen, 19 10 Coe/op/eurus floridanus A. Agassiz, 1 872
Habrocidaris scutata (A. Agassiz, 1 880)
Podoeidaris scu/pla A. Agassiz
Order Temnopleuroida
Family Temnopleuridae Genocidaris
macula/a A. Agassiz, 1 869 = Arbacina pal/aryi Gauthier, 1 897
Temnechinus macula/us A. Agassiz, 1 869 Hypsiechinus
coronaWs Mortensen, 1903 Trigonocidaris
a/bid a A. Agassiz
Family Toxopneusfidae Sphaerechinus
granularis (de Lamarck, 1 8 1 6) Echinus (Toxopneusles) brevispinosus Risso, 1 826
= Echinlls granularis de Lamarck, 1 8 1 6 S. rosells Russo, 1 893
Order Ecbinoida
Family Echinidae Echinus
aculus de Lamarck, 1 8 16 ?Echinus depressus G.O. Sars, 1 871
- 346 -
?Echt'nus micros/oma Wyville Thomson, 1 875 � ?Echinus norvegiclIs Duben & Koren, 1 846 = ?Echinus rarispinus G.O. Sars, 1 872
E. flemingU Forbes, 1 84 1 � E. sardicus Delle Chiaje, 1 84 1 ajfinis Mortensen, 1 903 alexandri Danielssen & Koren, 1 883 elegans Duben & Koren, 1 846
? Echinus wallisi A. Agassiz, 1 883 esculentus Linnaeus. 1 75 8 � E. aurantiacus de Blainville. 1 82 5
E. globiformis d e Lamarck. 1 80 I = E. pseudomelo de Blainville. 1 825
E. quinqueangulatus de Blainville, 1 825 E. quinqueangulosus de Blainville, 1 834
� E. schwartz;; Nilsson & Holst. 1 8 1 7 E. sphaera O.F. MUlier, 1 776
= E. subangulosus de Blainville, 1 826 = E. violaceus de Blainville, 1 825 esculentus var. glacialis D'yakonov, 1 923 melD Lamarck, 1 8 1 6 tenuispinus Norman. 1 868
E. esculen/us var. tenuispinus Norman, 1 868 Paracenlrotus
lividus (de Lamarck, 1 8 1 6) = Echinus lividus de Lamarck, 1 8 I 6 = Echinus purpureus Risso, 1 826
Echinus saxatilis Tiedemann, 1 8 1 6 = Echinus vulgaris de Blainville, 1 825
Psammechinus microtuberculatus (de Blainville, 1 825) Heller. 1 86 8 � Echinus microtuberculatus d e Blainville, 1 825 miliaris (P.L.S , MUlier, 177 1 )
Echinus miliaris P.L.S. MUlier, 1 766 = Parechinus miliaris Mortensen, 1 903
Family Strongylocentrotidae Strongylocentrotus
droebachiensis (O.F. MUller, 1 776) = Echinus drobachiensis O.F. MUlier. 1 776 = Echinus granularis Say, 1 826 = Echinus neglec/us de Lamarck, 1 8 1 6
S. chlorocentrotus Brandt, 1 835 = Toxopneustes pictus Norman, 1 8?? pallidus (G.O. Sars, 1 87 1 ) = ? Strongylocentrotus granular;s (Say, 1 827) = S. echinoides A. Agassiz & H,L. Clark, 1 907
S. pic/us (Norman, 1 868) = S, sachalinicus Clark, 1 9 1 2
Order Clypeasteroida
Family Fibuillriidile Echinocyamus
grandiporus Mortensen. 1907 macros/om us Mortensen, 1907 pusillus (O.P, MOiler, 1 776) = E, angulosus Leske. 1 778 = E. parthenopaeus Costa, 1 869
E. speciosils Costa, 1 869 = Echinus minulus Omelin, 1 790 = Fibularia equina Aradas, 1 850
Fihularia tarentina de Lamarck, 1 8 1 6 = Spa/agus pusillus O.F. MUlier, 1 776
Order Cassiduloida
Family Neolampadidae Neolampas
rostellata A. Agassiz, 1 869
Order Pourtalesioida
Family Pourtalesiidae Echinosigra
phiale (Wyville Thomson. 1 873) Mortensen. 1 907 = E, paradoxa (Mortensen, 1 905) Mortensen. 1907 = Pourtalesia paradoxa Mortensen, 1 905 = Pourtalesia phiale Wyville Thomson, 1 873
Pourtalesia
European Register of Marine Species
alT. alcocki Koehler, 1 9 1 4 = P. miranda auctt . • non A . Agassiz, 1 869 jeffreysi Wyville Thomson. 1 873 wandeli Mortensen, 1 905
Family Urechinidae Urechinus
naresianus A. Agassiz, 1 88 1
Family Plexechinidae Plexechinus
hirsutus Mortensen, 1 905
Family Calymnidae Calymne
relicta Wyville Thomson, 1 877?
Order Spatangoida
Family Hemiasteridae Hemiaster
expergitus Loven, 1 874 = H. gibbosus A. Agassiz. 1 88 1
H. mentzii A . Agassiz. 1 883 = H. zonatus A. Agssiz, 1 881
Family Spatangidae Spatangus
inermis Mortensen, 1 9 1 3 purpureus (O.F. MUlier, 1 776) = Spatagus purpureus O.F. MUlier, 1 776
S. 'meridionalis Risso, J 825 = S. reginae Gray, 1 855 = S, spinosissimus Desor, 1 847 rasch/ Loven, 1 869 = S, meridionalis Norman, 1 8?? non Risso, 1 825
Family Schizasteridae Brisaster
fragilis (Dilben & Koren, 1 846) Brissusfragilis Duben & Koren, 1 846
= Schizaster fragilis (DUben & Koren) Schizaster
canaliferus (de Lamarck. 1 8 1 6) = Spatangus canaliferus de Lamarck, 1 8 1 6
Family Brissidae Brissopsis
atlantica Mortensen, 1 907 atlantica var. mediterranea Mortensen, 1 9 1 3 Iyrifera (Forbes, 1 84 1 ) = B. parma Val., 1 8??
Brissus Iyrifer Forbes, 1 84 1 = Brissus pulvinatus Philippi, 1 845 = Schizaster incer/us Aradas, 1 850
Brissus unicolor (Leske, 1 778) = B, brissus auctt.
B. scil/ae L. Agassiz, 1 836 = Spatangus brissus subsp. unieolor Leske, 1 778
Spatangus columbaris de Lamarck. 1 8 1 6 Plagiabrissus
ajfinis ? costai (Gasco. 1 876) = Metalia costae Gasco, 1 876 = Rhabdobrissus perrieri Bernard
Family Loveniidae
- 347 -
Echinocardium cordatum (Pennant, 1 777) = Amphidetus kiirtzii Girard. 1 852 = E. sebae Gray. 1 8??
Echinus cordatus Pennant, 1 777 = Spa/angus arcuarius de Lamarck, 1 8 1 6 fenauxi Pequignat, 1 963 flavescens (O.P. MUlier. 1 776)
= Amphidotus roseus Forbes, 1 84 1 = Spalagus flovescens O.F. MUlier. 1 776
Spatangus ovatus Leske, 1 778 mediterranellm (Forbes. 1 844)
Amphidetus mediterraneus Forbes, 1 844
Costello, Emblow and White (eds), 200 1
= E. gibbosus L. Agassiz & Desor, 1 847 marlenseni Thiery, 1 909 = ?Echinocardium laevigasler A. Agassiz, 1 8??
? Echinacardium pennatifidum Koehler, 1 898, nOli Nonnan, 1 868
= E. intermedium Mortensen, 1 907 pennatifidum Nonnan, 1 868 = E. gibbosus Barrett, 1 857 non L. Agassiz & Desor,
1 847
Family Aeropsidae Acesle
bellidifera Wyville Thomson, 1 877 Aeropsis
rastrata (Wyville Thomson, 1 877) Mortensen, 1 907 = Aerope rostra/a Wyville Thomson, 1 877
Family Asterostomatidae Palaeotropus
josephinae Loven, 1 8?? = Palaeobrissus hilgardi A. Agassiz = P. hirondellei Koehler
Peripatagus cinctus Koehler
Class Holothuroidea
Order Dendrochirotida
Family Psolidae Psolidium
complana/um Cherbonnier, 1 969 Psolus
fabricii (DUben & Koren, 1 846) Cuvieria /abricii DUben & Koren, 1 846
hypsinotus Heding, 1 942 nummularis R. Perrier, 1 899 opercula/us (Pourtales, 1 868)
Cuvieria operculala Pourtales, 1 868 phan/apus (Strussenfelt, 1 765) Jaeger, 1 833 = Hololhuria phan/apus Strussenfelt, 1 765 = P. granulalus Ayres, 1 85 1
P. laeviga/us Ayres, 1 85 1 = P. regalis Verrill, 1 866 pour/ales;; Theel, 1 886 squama/us (O.F. MUller, 1 776) L[Jtken, 1 857
Holo/huria squamala O.F. MOller, 1 776 = P. valvallls Ostergren, 1 904 lesselalus Koehler, 1 896
Family Phyllophoridae Ekmania
bar/hii (Troschel, 1 846) Orcula barlhii Troschel, 1 846
= Orcula luminosa Lampert, 1 885 Thyonidium pellucidum DUben & Koren, J 846
Euthyonidiella allantica (Ludwig & Heding, 1 935) = Pseudocucumis a/lanlica Ludwig & Heding, 1 935
Neocucumis arlantica (Ludwig & Heding, t 935) marioni (von Marenzeller, 1 878) = Cucl/maria marioni von Marenzeller, 1 878
Pseudocucumis marioni Ludwig. 1 874 Neopenladactyla
mix/a (Ostergren, 1 898) Deichmann, 1 944 = Pseudocucllmis cueno/i Koehler & Vancy, 1 905
Pseudocucumis mix/a Ostergren, 1 898 Phyllophorus
drachi Cherbonnier & Guille, 1 968 granulatus (Grube, 1 840)
Psolus granulalus Grube, 1 840 (Phyllophorus) pedinaequalis Cherbonnier, 1 969 (Phyllophorus) IIrna Grube, 1 840 = Holothuria penicillus Delle Chiaje, 1 828
Thyonidium hyalinum (Forbes, 1 84 1 ) = Holo/huriapellucida Fleming, 1 828, non Vahl, 1 806 drummondii (Thompson, 1 840)
Cucumaria communis Forbes, 1 84 1 = Cucumaria nobilis Ludwig, 1 874
Cucumaria perspicua Ludwig, 1 874 = Holo/huria drummondii Thompson, 1 840
Thyone porllocki Forbes. 1 84 1 = T. diibeni Nonnan, 1 869
Family Cucumariidae
- 348 -
Abyssocucumis abyssorum (Theel, 1 886) = Cucumaria abyssorum auctt., non Theel. 1 886
Cucumaria abyssorum Theel. 1 886 = Cucumaria sluiter; Ohshima, 1 9 1 5 = Staurocucumis ingolfl Clark & Deichmann, 1 936
Aslia le/evrei (Barrois, 1 882) = Cucumaria leJevrei Barrois, 1 882
Cucumaria normani Allen & Pace, 1 904 = Hololhl/ria monlagui Fleming, 1 828 (p.p.)
Cucumaria arcuala (Herouard, 1 92 1 ) = Psolidium arcua/um Herouard, 1 92 1 compressa (R. Perrier, 1 898)
Ocnl/Ii compressus R. Perrier, 1 898 /rondosa (Gunnerus, 1 767)
C. assimilis DOben & Koren, 1 846 = Holo/huria /rondosa Gunnerus, 1 767
Hololhuriafucicola Forbes & Goodsir, 1 839 = Holothuria grandis Forbes & Goodsir, 1 839
Hololhuria pen/acles Linnaeus, 1 767 paraglacialis Heding, 1 942
Havelockia inermis (Heller, 1 868)
Thyone elegans Nonnan, 1 869 = Thyone inermis Heller, 1 868
Urox;a auranliaea Costa, 1 869 Neocnus
incubans Cherbonnier, 1 972 Ocnus
glacialis (Ljungman, 1 879) = Cue umaria glacialis Ljungman, 1 879 grubei (von Marenzeller, 1 874) = Cucumaria dicquemarii M. Sacs. 1 857
Cucumaria grubei von Marenzeller. 1 874 = Pseudocnus grubei Panning, 1 949 koellikeri (Semper, 1 868) = Cucumaria Mllikeri Semper, 1 868
Pseudocnus dubiosus (Semper) Panning, 1 962 = Pseudocl1us kollikeri Panning. 1 949 lacleus (Forbes & Goodsir, 1 839) = Cucumaria mos/erensis Grieg, 1 889
Hololhuria laclea Forbes & Goodsir. 1 839 pe/iti (Cherbonnier, 1 958)
Ludwigia petiti Cherbonnier. 1 958 planei (Brandt, 1 835)
?Holothuria brunneus (W. Thompson, 1 840. ex Forbes MS), non Chamisso & Eysenhardt, 1 82 1 ?Ocnus brunneus Forbes & Goodsir, in Forbes, 1 84 1
= Cladodactyfa plonci Brandt, 1 835 syracusanus (Grube, 1 840) Panning, 1 949 = Cladodactyla syracusana Grube, 1 840
Cucumaria hyndmanni W. Thompson, 1 840 = Cucumaria syracusana auctl.
Pseudocnus syracusanus auctt. Paracucumaria
mauri/anica (Herouard, 1 929) = Cucumaria mauritanica Herouard, 1 929 thalassae Cherbonnier, 1 969
Pawsonia saxicola (Brady & Robertson, 1 87 1 ) = Colochirus l!lcazei Herouard
Cucumaria saxicola Brady & Robertson, 187 1 = Hololhuria mOlllagui Fleming, 1 828 (p.p.) aff. saxicola Garrido (email check-list ofCanarian echinoderms)
Penlamera
calcigera (Stimpson, 1 85 1) = Cueumaria koreni LOtken, 1 857
Penlacta calcigera Stimpson; J 85 1 Pseudolhyone
furnesli Cherbonnier, 1 969 raphanus (Dilben & Koren, 1 846)
Thyone poucheri Barrois, 1 882 = Thyone raphanus DUben & Koren, 1 846 seulponea Cherbonnier, 1 958 serrifera (Ostergren, 1898) Jensenius Madsen & Hansen, 1 994
= Thyone serrifera Ostergren, 1 898 Slereoderma
incerta Cherbonnier, 1 969 kirchsbergi (Heller, 1 868) = Cucumaria kirchsbergi Heller, 1868
Gcnus? kirchsbergi von Marenzeller, 1 874 Thyone
cherbonnieri Reys, 1960 fusus (O.F. MUller, 1776)
?Thyone flexus Hodge, 1 867 = Holo/huriafusus O.F. MOiler, 1776 = Holo/huria papillosa AbiidglUlrd, 1 789 = T. subvillosa Herouard, 1 890 fusus subsp. mcdilerranea Madsen, 1 94 1 gadeana R. Perrier, 1 898
T. wahrbergi Madsen, 1 94 1 roscovila Herouard, 1 890
Lep/open/acla elonga/a (DOben & Koren, 1 846)
?Cucumaria eucumis (Risso, 1 8??) = Cucumaria elonga/a Dilben & Koren, 1 846
Cueumaria pentacles Forbes, 1 84 1 non (Linnaeus, 1767)
/ergestina (M. Sars, 1 857) = Cucumaria incurvata Perrier, 1 902 = Cucumaria lergeslina M. Sars, 1 857
Order Dactylochirotida
Fllmily Ypsilolburiidae Echinocucumis
hispida (Barrett, 1 857) = Cucumaria hispida (Barrett, 1 857) = E. typica M. Sars, 1 859 = Eupyrgus hispidus Barrett, 1 857
Ypsilo/huria bifen/aculata subsp. atlenuata E. Perrier, 1 886 talisman; subsp. talismanl E. Perrier, 1 886 = ?Echinocucumis hispida var. abyssalis Koehler, 1 896
Order Aspidochirotida
Family Stichopodidae Eostichopus
regalis (Cuvier, 1 8 1 7) = Holothuria columnae Cuvier, 1 8 1 7
Holo/huria regalis Cuvier, 1 8 1 7 = Holothuria triquetra Delle Chiaje, [84 I = Sfichopus regalis (Cul/ier, 1 8 1 7)
Parastichopus tremulus: (Gunnerus, 1767) = ? Stichopus griegi ()stergren, 1 896
Holo/huria ecalcarea M. Sars, 1 859 = Holo/huria elegans O.F. MOiler, 1781
Holothuria tremula Gunnerus, 1767 = Slichopus richardi Herouard, 1 896
Stichopus tremulus (Gunnerus)
Family SynaUactidae Bathyplotes
natans (M. Sars, 1 868) ()stergren, 1 896 B. ass/mills Koehler & Vaney, 1 905
= B. helerostylides Heding, 1 942 B. ostergreni Ohshima, 1 9 1 5
= B . pap/Ilosus Koehler & V aney, 1 905 = B. palagia/us Fisher, 1 907 = Holothuria na/ans M. Sars, J 868
European Register of Marine Species
= Slichopus moseleyi Theel, 1 886 pourlalesii (Theel, 1 886) = B. bipartitus Herouard, 1 923 = Stichopus pourtalesii Theel, 1 886 lizardi (Theel, 1 882) = B.fallax Ostergren, 1 896 = B. replans R. Perrier, 1 902 = Stichopus lizardi Theel, 1 882
Benlhothuria funebris R. Perrier, 1 898
= ? Bentho/huria valdiviae Heding, 1 940 Gephyrothuria
alcocki Koehler & Vaney, 1905 = G. europeensis Herouard, 1 923
Mesothuria bifurcata Herouard, 1 906 candelabra Herouard, 1 923 cathedralis Heding, 1 940 connectens: (R. Perrier, 1 898) = Zygothuria connectens R. Perrier, 1 898 expectans R. Perrier, 1 898 intestinalis (Ascanius, 1 805) Ostergren, 1 896
Fistularia mollis M. Sars, 1 835 = Holothuria inteslinalis Ascanius, 1 805
Thyonidium scabrum M. Sars, 1 868 laelea (Theel, 1 886)
?Holothuria aspera Bell, 1 892 = Holothuria laclea Theel, 1 886 = Zygolhuria laclea (Theel, 1 886) maroccana Perrier, 1902 murrayi var. grandipes Herouard roulei (Koehler, 1 896)
Holo/huria roulei Koehler, 1 896 rugosa Herouard. 1 9 1 2 trirad/ata Heding, 1 942 verrilli (Theel. 1 886)
Holothuria verrilli Theel. 1 886 = M. oklaknemus Sluiter, 1 90 1
Meseres aeaudum Heding, 1 935 ingolji (Heding, 1940) = Plicaslichopus ingolji Heding, 1 942
Paelopolides atlantica Herouard, 1 902 grisea R. Perrier, 1 902
Paroriza pollens (Koehler, 1 896) = Stichopus pallens Koehler, 1 896 prahoi Herouard, 1 902
Pseudostichopus atlanticus R. Perrier, 1 898 depressus Herouard, 1 902 globigerinae Herouard, 1 902 lapidus Herouard marenzelleri Herouard, 1 902 oceultatus von Marenzeller, 1 893 villosus Theel, 1 886
ScolOthuria herringi Hansen, 1 978 = ?Dendrothuria simi/is Koehler & Vaney. 1 905
Synallactes crucifera R. Perrier, 1 898
Fllmily Holotburiidlle Gastroturia
- 349 -
limbata R. Perrier, 1 899 Holothuria
(Roweothuria) arguinensis Koehler & Vaney, 1 906 dakorensis Panning, 1 9?? forskali Delle Chiaje, 1 823 = H. catanensis Grube, 1 840 = H. nigra Peach, 1 845 = Slichopus selenkae B arrois, 1 882 helleri von Marenzeller, 1 878 = H. affinls Heller, 1 868, non Brandl, 1 83 5 (Thymiosicya) impatiens: (Forskal, 1 775)
Costello, Emblow and White (eds), 2001
= FislUiaria impatiens Forskal, 1 775 = H. botellus Selenka, 1 867
H. impatiens subsp. bicolor Clark, 1 93 8 = H. impatiens subsp. concolor Clark, 192 1
H. impatiens subsp. /utea Clark, 192 1 = H. impatiens subsp. pulchra Clark, 1 92 1 = H. truncata Lampert, 1 885 (I'aneyolhuria) lentiginosa von Marenzeller, 1 892 mammala Grube, 1 840 (Halodeima) mexicana Ludwig, 1 875 (Platy perona ) parvula (Selenka, 1 8??) polii Delle Chiaje, 1 823 sanelori Delle Chiaje, 1 823 = H. farcimen Selenka, 1 867 (Holothuria) stellali Delle Chiaje, 1 823 (Holothuria) tubulosa Gmclin, 1 788 = H. cavolin;; DeUe Chiaje, 1 84 1 = H. columnae Delle Chiaje, 1 823, non Cuvier, 1 8 1 7
H. maxima Delle Chiaje, 1 823 = H. pentagnae Delle Chiaje, 1 823
Order Elasipodida
Family Elpidiidae Ach/yonice
myriamae Gebruk, 1 997 Amperima
furcala (Herouard, 1 899) = Ko/ga furcala Herouard, 1 899 rosea (R. Perrier, 1 896) = Periamma rosea R. Perrier, 1 896
Ellipinion delagei (Herouard, 1 896) = Scoloplanes delagei Herouard, 1 896
Elpidia glacialis Theel, J 876 echinala (R. Perrier, 1 896)
TUlela echinala R. Perrier, 1 896 Trpa
abyssieola Danielssen & Koren, 1 878 Kolga
hyalina Danielssen & Koren, 1 879 Elpidia nana Theel, 1 879
Peniagone azorica von Marenzeller, 1 893 diaphana (Theel, 1 882) = Seoloonassa translucida Herouard, 1 899 gracilis (Ludwig, 1 894) islandica Deichmann, 1 930 porcella R. Perrier, 1900 purpurea (Theel, 1 882) = P. ferruginea Grieg, 192 1 = P. vexillum R. Perrier, 1900 w)'villii Theel, 1 882
Penifpidia IUdwigi (von Marenzeller, 1 893)
Kolga IUdwigi von Marenzeller, 1 893 Psychroplanes
obsolela (Herouard, 1 899) = Kolga foliacea Herouard, 19 12 = Kolga obsoleta Herouard, 1 899 = Peniagonefoliaeea (Herouard, 1 9 1 2) = Peniagone obsolela (Herouard, \ 899)
Family Deimatidae Deima
validulII subsp. valMum Theel, 1 879 = D. allanlicum Herouard, 1 898
D. blake! Theel, 1 886 Oneirophanta
mUlabilis Theel, 1 879 0. allernala R. Perrier, 1 900
Family Laetmogonidae Benthogone
rosea Koehler, 1 896 = B. quadrilineala Heding, 1 940
Laelmogone
violacea Theel, 1 879 = Cryodora spongiosa Theel, 1 879
L. brongniarli E. Perrier, 1 886 = L. jourdainii Pel it, 1 885 = L. wyvillelhomsoni Theel, 1 879
Pannyahia glutinosa Herouard, 1 902
Family Psychropotidae Benthodyles
janthina von Marenzeller, 1 893 = B. lingua R. Perrier, 1 896 typica Theel, 1 882 = B. glutinosa R. Perrier, 1 902
Psyehropoles depressa (Theel, 1 882) = Euphronides auricularia R. Perrier, 1 902
Euphronides com uta Verril l , 1 884 = Euphronides lalismani R. Perrier, 1 896 = Euphronides violacea Perrier, 1 902 grimaldii Herouard, 1 896 longicauda Theel, 1 882 = Neclolhuria translucida B elyaev & Vinogradov, 1969 = P. buglossa R. Perrier, 1 896 = P. furcola R. Perrier, 1 896 semperiana Theel, 1 882 = Euphronides anchora Herouard, 19 J 2 = Euphronides kerhervei (Herouard, 1 896) = P. kerhervei Herouard, 1 896
Family Pelagolhuridae Enypniastes
eximia Theel, 1 882 = Pelagothuria bouvieri Herouard, 1 906 = Planklothuria diaphana Gilchrist, 1 920
Order Apodida
Family SynaptidBe Epilomapla
- 350 -
roseola (Verrill , 1 874) Leplosynapta roseola Verrill, 1 874
Euapla lappa (J. MUlier, 1 850) = SYllapla lappa J. MUlier, 1 850
Labidop/ax buskii (M'lntosh, 1 866) = Synapla busk;; M'Intosh, 1 866 = Synapta lenera Norman, 1 865 (nomen nudum) digilala (Montagu, 1 8 1 5) = Hololhuria digilata Montagu, 1 8 1 5
L. (Oes/ergrenia) adrialiea Heding, 1 93 1 digitala. var. profundlcola Kemp, 1 905 media Ostergren, 1905 morlenseni Heding, 193 1 = L . (Oeslergrenia) morlenseni Heding, 1 93 1 similimedia Gage, 1 985 southwardorum Gage, 1 985 Ihomsoni (Herapalh, 1 865) = Synapta digilala var. Ihomsonii Herapath, 1 865
Leplosynapla bergensis (Ostergren, 1 905) = Synapla bergensis Ostergren, 1 905 decaria (Ostergren, 1905) = Synapla decaria Ostergren. 1 905 galiiennil (Herapath, 1 865) = Synapla gal/ienn;; Herapath, 1 865 = Synapla sarniensis Herapalh. 1 865 inhaerens (O.F. MUlier, 1 776) = Hololhuria inhaerens O.F. MUlier, 1776 = Synapla duvernaea de Quatrefages, I S?? makrankyra (Ludwig, 1 898) = Synapfa makrankyra Ludwig, 1 898 minula (Becher, 1 906) = Synapla mimlla Becher, 1 906
Prolankyra abyssicola (Theel, J 886) = Synapla abyssicola Theel, 1 886
Rhabdomolgus ruber Keferstein, 1 863
Synapfula reciproeans (Forskal, 1 775) = Synapta reciproeans Forsk�I, 1 775
Family Chiridotidae Achiridota
profunda Reding, 1 935 Chiridota
abyssicola von Marenze"er, 1 893 laevis (0. Fabricius, 1 780)
C. groenlandiea Heding, 1 935 � C. spirourna Heding, 1 935 = Hololhuria laevis O. Fabricius, 1 780
Hololhuria pellucida Vahl, 1 806 Trochodola furcipraedila Sa1vini Plawen, 1 972 venusta (Semon, 1 887) = Chiridola venusla Semon, 1 887
Family Myriotrocbidac Acanfhofrochus
mirabilis Danielssen & Koren, 1 879 Myriotrochus
balhybius H.L. Clark, 1 920 clarki Gage & Billett, 1 986 eurycyclus Heding, 1 935 geminiradialus Sa1vini-Plawen, 1 977 giganteus Clark, 1920 rinkii Steenstrup, 1 85 1 Iheeli Ostergren, 1 905 vilreu8 (M. Sars, 1 872) = Chiridofa brevis Huxley, 1 852 = Chiridola laevis C.GJ. Petersen, 1 898 = Oligolrochus vitreus M. Sars, 1 872
Parvolrochu8 belyaevi Gage & Billett, 1 986
Siniofrochus myriodontus Gage & B illett, 1 986
Trochoderma elegans Theel, 1 877
Order Molpadiida
Family Molpadiidae Cherbonniera
ulriculus Sibuet, 1 974 Molpadia
blakei (Theel, 1 886) = Paralrochosloma spiniferum Heding, 1 932 = Trochostoma blakei var. excenlrica Herouard
Trochos/oma blakei Theel, 1 886 = Trochosloma grossularia Herouard borealis M. Sars, 1 859 = Ankyroderma affine Danielssen & Koren, 1 882 = Ankyrodermajeffreysii Danielssen & Koren, 1 878
Haplodaetyla arc/iea von Marenzeller, 1877 = M (Paramofpadia) dipfoa Heding, 1 935 = M oolitica Mortensen, 1 9 1 4 non Pourtali:s, 1 85 1
Troehos/oma arc/iea Danie1ssen & Koren, 1 880 non (von Marenzeller, 1 877)
= Trochostoma boreale (M. Sars) Trochosloma Thomsonii Danielssen & Koren, 1 879
foricala (R. Perrier, 1 898) = Ankyroderma forica/urn Perrier, 1 898 maroccana (Perrier, 1 898) = Ankyroderma rnaroceana Perrier, 1 898 musculus Risso, 1 826 = Ankyroderrna danielsseni Theel, 1 886 = Ankyroderma hispanicum Petit
Ankyroderma perrieri Petit = Eumolpadia asaphes Heding, 1 935
Haplodaclyla mediterranea Grube, 1 840 = Trochostoma violaceum Theel, 1 886 ooli/ica (Pourtales, 1 SS I )
Family Eupyrgidae Eupyrgus
European Register of Marine Species
scaber LUtken, 1 857 = Echino.mma hispidum Semper, 1 868
Hedingia
- 3 5 1 -
albicans (Theel, 1 886) Deichrnann, 1 938 = Molpadia albicans auett.
Trochos/oma albieans Theel, 1 886 mediterranea (Bartolini Baldelli, 1 9 14) Tortonese,
1965 Trochostoma mediterraneum Bartolini Baldelli, 1 9 1 4
