Insect Fauna of Koreawebbook.me.go.kr/DLi-File/099/006/005/5580279.pdf · Insect Fauna of Korea Kyu-Tek Park The Korean Academy of Science and Technology Volume 16, ... any form or

Insect Fauna of Korea

Flora and Fauna of Korea

National Institute of Biological ResourcesMinistry of Environment

Volume 16, Number 9Arthropoda: Insecta: Lepidoptera: Gelechiidae

Gelechiidae II

Insect Fauna of KoreaVol. 16, N

o. 9G

elechiidaeII

NIB

R


China

PB

JG

YG

HB

HN

PN

HWB

GW

GG

CB

CNGB

GB(Ulleung-do)

GN

JN

JJ

JB

HWN

Russia

East Sea

Yellow Sea

South Sea

CB Chungcheongbuk-doCN Chungcheongnam-doGB Gyeongsangbuk-doGG Gyeonggi-doGN Gyeongsangnam-doGW Gangwon-doHB Hamgyeongbuk-doHN Hamgyeongnam-doHWB Hwanghaebuk-doHWN Hwanghaenam-doJB Jeollabuk-doJG Jagang-doJJ Jeju-doJN Jeollanam-doPB Pyeonganbuk-doPN Pyeongannam-doYG Yanggang-do



Gelechiidae II

2013



Kyu-Tek ParkThe Korean Academy of Science and Technology


Gelechiidae II

Copyright ⓒ 2013 by the National Institute of Biological Resources

Published by the National Institute of Biological ResourcesEnvironmental Research Complex, Hwangyeong-ro 42, Seo-guIncheon, 404-708, Republic of Koreawww.nibr.go.kr

All rights reserved. No part of this book may be reproduced, stored in a retrieval system, or transmitted, inany form or by any means, electronic, mechanical, photocopying, recording, or otherwise, without the priorpermission of the National Institute of Biological Resources.

ISBN : 9788968110290-96470Government Publications Registration Number 11-1480592-000543-01

Printed by Junghaengsa, Inc. in Korea on acid-free paper

Publisher : Sang Pal LeeProject Staff : Ki Gyoung Kim, Hong-Yul Seo, Sang-Hoon Han

Published on October 30, 2013

The Flora and Fauna of Korea logo was designed to represent six major target groups of the project includingvertebrates, invertebrates, insects, algae, fungi, and bacteria. The book cover and the logo were designed byJee-Yeon Koo.

A Korean translation of this issue is simultaneously published for Korean speaking readers. ThisEnglish version therefore should be regarded as an original publication that has nomenclatural priority.

Preface

In 2010, the 10th Conference of the Parties of the Convention on Biological Diversity (CBD) adoptedthe Nagoya Protocol on access and benefit-sharing of biological resources. Since then, the nationaland international environment of the use and management of biological resources has been rapidlychanged. Therefore, it is imperative to identify indigenous biological species in details and to buildscientific data worthy of international recognition in order to take the initiative in bio-industry.

The National Institute of Biological Resources of the Ministry of Environment has been publishingthe Flora and Fauna of Korea to generally manage biological resources and to enhance national com-petitiveness by setting the foundation for the sovereignty over biological resources. Professionalresearch group consisting of professors of taxonomy and related experts has systematically examineda variety of and wide range of taxa for the last 6 years since 2006.

As a result, 90 issues of the Flora and Fauna of Korea, both in Korean and in English, covering atotal of 8,888 species and 2 issues of World Monograph covering 216 species were published. And30 issues of the Flora and Fauna of Korea, both in Korean and in English, covering 1,665 species ofinvertebrates, insects and algae are additionally published this year.

These efforts serve to identify indigenous species living in Korea, to investigate biota, to improvethe quality of national biological resources management and to provide the opportunity to lay thegroundwork for the biotechnological industrialization of biological resources.

I would like to express my sincere appreciation for those who spared no effort to publish the bio-logical magazines; Dr. Kyu-Tek Park of The Korean Academy of Science and Technology, ProfessorJong Kyun Park of Kyungpook National University, Dr. Jinyoung Park of Kyungpook National Uni-versity, Professor Yang-Seop Bae of University of Incheon, Mr. Uljizarkal and Ms. Bo-Seon Park ofUniversity of Incheon, Dr. Hong-Sik Won of Jeohyeon Highschool, Dr. Jeong-Kyu Kim of HanseoUniversity, Professor Jong-Wook Lee of Yeungnam University, Dr. Jong-Chul Jeong of Korea NationalPark Service, Dr. Jin-Kyung Choi and Mr. Ki-Beom Kim of Yeungnam University, Dr. Heung-Sik LeeAnimal Plant and Quartine Agency, Professor Dong-Pyo Ryu of Sangji University, Dr. Boo-Hee Jungof Korea University, Dr. Young Bok Cho of Hannam University, Professor Sei-Woong Choi of MokpoNational University, Dr Kyungduk Han of Korea University, Dr. Sangwook Park of Research Insti-tute of Forest Insects Diversity, Professor Ki-Jeong Hong of Sunchon National University, andProfessor See-Ae Bae of Shinheung College. And I hope they will help to discover useful biologicalresources in Korea and to create high value-added activities including natural product, gene resourceand medical substance development.

Sang-pal LeePresidentNational Institute of Biological Resources

1

Contents

List of Taxa 2

Introduction 4

Materials and Methods 5

Taxonomic Notes 9

Family Gelechiidae 9Subfamily Gelechiinae 9

1. Teleiopsis Sattler 92. Pseudotelphusa Janse 113. Concubina N. Omelko and M. Omelko 134. Altenia Sattler 14

Subfamily Anacampsinae 165. Syncopacma Meyrick 166. Aproaerema Durrant 187. Anacampsis Curtis 208. Prolita Leraut 249. Mesophleps Hübner 26

Subfamily Dichomeridinae 3010. Helcystogramma Zeller 3111. Dichomeris Hübner 3712. Neofaculta Gozmány 7013. Encolapta Meyrick 7114. Hypatima Hübner 7515. Tornodoxa Meyrick 7916. Faristenia Ponomarenko 8117. Dendrophilia Ponomarenko 9318. Bagdadia Amsel 9919. Anarsia Zeller 102

Literature Cited 118

Plates 132

Index to Scientific Names 142

List of Taxa

Checklist of Gelechiidae in the Korean Peninsula

2

Order LepidopteraFamily Gelechiidae

Subfamily Gelechiinae Genus Teleiopsis Sattler, 1960

106. motleella Ponomarenko and Park, 2007

Genus Pseudotelphusa Janse, 1958 Sattleria Capu e, 1968 Klaussattleria Capu e, 1968

107. acrobrunella Park, 1992 fulvosella N. Omelko and M. Omelko,1993

Genus Concubina N. Omelko and M. Omelko,2004

108. trigonalis Park and Ponomarenko, 2007

Genus Altenia Sattler, 1960 109. inscriptella (Christoph, 1882)

Subfamily AnacampsinaeTribe Anacampsini

Genus Syncopacma Meyrick, 1925110. wormiella (Worff, 1958)

Genus Aproaerema Durrant, 1897111. anthyllidella (Hübner, [1813])

sparsiciliella Barrett, 1891

Genus Anacampsis Curtis, 1827 112. anisogramma (Meyrick, 1927) 113. lignaria (Meyrick, 1926)

homoplasta Meyrick, 1932114. solemnella (Christoph, 1882)

metagramma Meyrick, 1918

Genus Prolita Leraut, 1993115. sexpunctella (Fabricius, 1794)

virgella Thunberg, 1794longicornis Curtis, 1827histrionella Geyer, [1832]

zebrella Treitschke, 1833longicornella Doubleday, 1859petulans Braun, 1925

Genus Mesophleps Hübner, [1825]116. albilinella (Park, 1990) 117. sublutiana (Park, 1990) 118. acutunca Li and Sattler, 2012

Subfamily DichomeridinaeTribe Dichomeridini

Genus Helcystogramma Zeller, 1877119. fuscomarginatum Ueda, 1995 120. ineruditum (Meyrick, 1926)

inerudita Meyrick, 1926lutatella sensu Omelko, 1999

121. perelegans (N. Omelko and M. Omelko,1993)

122. triannulella (Herrich-Schäffer, 1854) sepiella Streudel, 1866macroscopa Meyrick, 1932

Genus Dichomeris Hübner, 1818123. atomogypsa (Meyrick, 1932) 124. autometra (Meyrick, 1934) 125. chinganella (Christoph, 1882)

fareasta Park, 1994126. christophi Ponomarenko and Mey, 2002

sparsella sensu Park, 2006127. cuspis Park, 1994 128. derasella ([Denis and Schiffermüller],

1775) fasciella Hübner, 1796coreanus Matsumura, 1931paranthes Meyrick, 1936unguiculatus Fabricius, 1798

129. harmonias Meyrick, 1922 130. heriguronis (Matsumura, 1931)

iothalles Forbes, 1939picrocarpa sensu auct.

List of Taxa 3

131. horoglypta Meyrick, 1932 132. issikii (Okada, 1961)

synergastis Ponomarenko and Park,1996

133. lespedezae Park, 1994 harmonias sensu auct.

134. litoxyla Meyrick, 1937 135. lutilinea Ponomarenko and Park, 1996136. minutia Park, 1994 137. mitteri Park, 1994 138. oceanis (Meyrick, 1920)

limitellus Caradja, 1920yanagawanus Matsumura, 1931

139. polypunctata Park, 1994 140. praevacua Meyrick, 1922

quercicola sensu auct.141. rasilella (Herrich-Schäffer, 1854) 142. sparsella (Christoph, 1882)

strictella Park, 1994143. tostella Stringer, 1930

kawamurai Matsumura, 1931144. ustalella (Fabricius, 1794)

capuinella Hübner, 1796cornutus Fabricius, 1798ustatus Fabricius, 1798

145. anisacuminata Li and Zheng, 1996146. japonicella (Zeller, 1877)

Tribe ChelariiniGenus Neofaculta Gozmány, 1955

147. taigana Ponomarenko, 1998

Genus Encolapta Meyrick, 1913148. catarina (Ponomarenko, 1994) 149. subtegulifera (Ponomarenko, 1994) 150. tegulifera (Meyrick, 1932)

Genus Hypatima Hübner, [1825]151. excellentella Ponomarenko, 1991

silvestris sensu auct.152. venefica Ponomarenko, 1991

Genus Tornodoxa Meyrick, 1921153. longiella Park, 1993 154. tholochorda Meyrick, 1921

Genus Faristenia Ponomarenko, 1991155. acerella Ponomarenko, 1991 156. atrimaculata Park, 1993 157. furtumella Ponomarenko, 1991 158. geminisignella Ponomarenko, 1991 159. jumbongae Park, 1993160. maritimella Ponomarenko, 1991 161. omelkoi Ponomarenko, 1991

nigriella Park, 1993162. quercivora Ponomarenko, 1991 163. ussuriella Ponomarenko, 1991

Genus Dendrophilia Ponomarenko, 1993164. leguminella Ponomarenko, 1993 165. mediofasciana (Park, 1991)

brunneofasciella Ponomarenko, 1993166. neotaphronoma Ponomarenko, 1993 167. unicolorella Ponomarenko, 1993

Genus Bagdadia Amsel, 1949Capidentalia Park, 1995

168. claviformis (Park, 1993) 169. eucalla (Li and Zheng, 1998) 170. gnomia Ponomarenko, 1995

Tribe AnarsiiniGenus Anarsia Zeller, 1839

Ananarsia Amsel, 1959171. bipinnata (Meyrick, 1932) 172. ulneongensis Park and Ponomarenko,

1996173. gajiensis (Park and Ponomarenko, 1996)174. asymmetrodes Park, 2013175. lineatella (Zeller, 1839) 176. bimaculata Ponomarenko, 1989 177. nigricana Park, 1991 178. subnigricana Park and Ponomarenko,

1996

Additional records

Subfamily Gelechiinae179. Gnorimoschema streliciellum

(Herrich-Schäffer, 1854) 180. Stenolechia kodamai Okada, 1962181. Bryotropha similis (Stainton, 1854)

Introduction

In the Part II of the family Gelechiidae, four species belonging to four genera of Gelechiinae;Teleiopsis Sattler, Pseudotelphusa Janse, Concubina Omelko, and Altenia Sattler, which were not includ-ed in the Part I, are listed, and nine species of five genera of Anacampsinae and 60 species of 10genera of Dichomeridinae are included. In addition, three species of Gelechiinae; Gnorimoschemastreliciellum (Herrich-Schäffer), Stenolechia kodamai Okada, and Bryotropha similis (Stainton) are new-ly added. These three species are listed in the last few pages. Consequently, the number of knownspecies of Gelechiidae in the Korean Peninsula are enumerated at a total of 181 species. Accordingto Hodges (1986), the subfamily Anacampsinae is included in the subfamily Gelechiinae, but it isindependently treated to be a valid subfamily following the system given by Park and Ponomarenko(2007). The subfamily Dichomeridinae is divided into three tribes: Dichomeridini, Chelarini, andAnarsini in this Part II. The tribe Dichomeridini include Helcystogramma Zeller and DichomerisHübner. Dichomeris is one of the most diversed genus, with worldwide distribution: 81 species inthe North America (Hodges, 1986), 68 species in China and 27 species in Taiwan (Li 2002), and 24species in the Korean Peninsula. In Chelarini, seven genera (Neofaculta Gozmány, Encolapta Meyrick,Hypatima Hübner, Tornodoxa Meyrick, Faristenia Ponomarenko, Dendrophilia Ponomarenko, andBagdadia Amsel) are included, with 24 known species in Korea. The genus Capidentalia Park, 1995which was described by the author was synonymized with Bagdadia Amsel by Ponomarenko (2004).In this part, three species belonging to Gelechiinae: Gnorimoschema streliciellum (Herrich-Schäffer),Stenolechia kodamai Okada, and Bryotropha simils (Stainton) are additionally found. Consequently,the total number of the species of the family Gelechiidae in Korea is enummerated as a totalof 181species, and they are listed at the end.

4

Materials and Methods

Specimens included in this book are based on the collections which are preserved in the KoreaNational Arboretum. They were mostly collected by the author and his colleagues during the lastdecades in Korea. Some of them were collected from North Korea provided by individuals fromeastern European countries including Hungary and Poland since 1970 (Hungarian National Museumof Natural History, Budapest and Polish Academy of Science, Cracow).

Gelechiids were usually collected by light traps, with the use of a 220 v/200 W mercury vaporlamp. Moths were killed in jars containing plaster soaked with ethyl acetate (100%) and/or potas-sium cyanide, and kept at low temperatures. After wings were spreaded and fully dried, the speci-mens have been stored in air tight boxes with fumigant. Wing expanse was measured betweenapices of forewings. For morphological study, wing patterns, venation, and external morphologi-cal characters of the head, thorax, and genitalia of both sexes were carefully examined under anOlympus SZ6045, stereomicroscope, and photos of genitalia were taken with a Carl Zeiss Sv8 stereo-microscope equipment with a digital camera.

The genital preparation technique used for gelechiids is generally identical to the standard pre-paration of Microlepidoptera genitalia (Robinson, 1976). The aedeagus is usually separated fromthe genital capsule, but note that sometimes, especially in most Teleiodini, it is strongly fused withparts of the saccus. In this case, they are not easy to separate from each other and are thus leftintact. The genitalia preparation includes the following main steps:- abdomen removed from the body and immersed in a test tube containing 10% KOH for 1-2 hours

in an incubator at 70-80°C.-washed in 50% ETOH and scales removed using a flexible dissecting needle.- after the initial cleaning, stained with chlorazol black or mercurochrome.- placed the stained abdomen in 75% ETOH and remaining scales removed by abrading with needle.

The genitalia are then pulled out from the abdomen and cleaned again.- after final cleaning, dehydrated in absolute alcohol for about 1-2 minutes and hardened suffi-

ciently for mounting. - cleaned in Euparal Essence.-mounted in Euparal.- labeled with species name, data, and given individual slide numbers following the preparation

numbering system. Sometimes genitalia are stored in 70% glycerin vials, and likewise labeled with numbers corre-

sponding to the pinned adult.The number assigned to each species corresponds with the numbers in the checklist, the text, and

the color plates. All the available synonyms of the genera and species are listed under each spe-cific description. The valid subspecies names are marked “as a subspecies” after the name andreferences.

In the description of the species, a full synonymy is given, including original combination andsource of description. The type locality (TL) for the species is given, with few exceptions when itsinformation is not available. The ADULT section does not include a full description but rather ageneral diagnosis or some major characters. Figures of the genitalia are illustrated by author orcited from the original figure by author and co-authors. When the figures were cited from others,the original sources were indicated in the figure’s explanation. The SPECIMENS EXAMINED section

5

includes a locality, date, and gender of available speci-mens from various collections in Korea as well as frominstitutions in other countries such as the HungarianNational Museum of Natural History, Budapest, Hun-gary and the Polish Academy of Science, Cracow,Poland, and from some other personal collections,especially in the case of specimens from N. Korea.Some of the data were cited courtesy of the Korea Bio-diversity Information System of the Korea NationalArboretum. KOREAN RECORDS are based on specimendata, including information on specimens reported injournals, reports or books by local workers with theindication of author and year. The HOST PLANTS sec-tion includes all host plants associated with larvaefrom adjacent countries and Europe, including FarEastern Russia, China and Japan, obtained from litera-tures, and the original resources were cited in paren-theses. The REMARKS includes information on thefirst record, flight period of adults and hibernationstage where available. The DISTRIBUTION refers tocountries or regions in which the taxa are found. Thesection for KOREA includes local distribution, ie. North(N. Korea), Central (the central part of S Korea: GW,GG, CB, CN, GB, JB), South (the southern part: GNand JN), and JJ. The terminology on the genital struc-tures generally follows Klots (1970) with the considera-tion of the established homology of genital sclerites(Ponomarenko, 2005).

Abbreviations for the institutions/organizations:BMNH: The Natural History Museum, London, United Kingdom.CIS: Center for Insect Systematics, Kangwon National University, Chuncheon, Korea.HNHM: Hungarian Natural History Museum, Budapest, Hungary. IZAS: Institute of Zoology. Chinese Academy of Science, Beijing, China.KNA: Korea National Arboretum, Pocheon, Korea.MGAB: Muzeul National de Istorie Naturala, “Grigore Antipa”, Bucharest, Rumania.NMNH: National Museum of Natural History, Leiden, Netherlands.OPU: Osaka Prefecture University, Osaka, Japan.USNM: National Museum of Natural History, Smithsonian Institution, Washington, U.S.A.ZFMK: Zoologisches Forschungsinstitut und Museum, Alexander Koenig, Bonn, Germany.ZIRAS: Zoological Institute of Russian Academy of Sciences, St.-Petersburg, Russia.ZMHB: Museum für Naturkunde, Humboldt-Universität, Berlin, Germany. ZMUC: Zoological Museum, University of Copenhagen, Denmark.ZMUH: Zoological Museum, University of Helsinki, Finland. ZSM: Zoologische Sammlung des Bayerischen Staates, München, Germany.

6 Insect Fauna of Korea·Gelechiidae II

Text-fig. 1. Map with abbreviations forprovinces in Korea. GG: Gyeonggi-do(Seoul included); GW: Gangwon-do; CB:Chungcheungbuk-do; CN: Chungcheong-nam-do; JB: Jeollabug-do; JN: Jeollanam-do; GB: Gyeongsangbug-do; GN: Gyeong-sangnam-do; JJ: Jeju-do.

Acknowledgments

I am particularly grateful to Dr. M.G. Ponomarenko, Institute of Biology and Soil Science, Far EastBranch of Russian Academy of Sciences, Vladivostock, Russia, who collaborated for the compre-hensive publication, ‘Gelechiidae of the Korean Peninsula and Adjacent Territories (2007),’ whichplayed an important role and provided basic sources for this publication. I would like to thank thefollowing colleagues who helped in collecting material, preparation of specimens and genitalia,and photographing: Prof. Y.S. Bae, Incheon University, Incheon; Prof. B.K. Byun, Hannam Univer-sity, Daejeon; S.R. Kim and M.Y. Kim, who are graduate students in the Seoul National University,Seoul, Korea; E.M. Ji, Gyeonggi-do Forestry Environment Research Center, Osan, Korea. I grateful-ly appreciate to Dr. S. Lee, Arizona State University, USA, for her careful reading and corrections ofthe manuscript. This work was supported by the National Institute of Biological Resources (NIBR)under the Ministry of Environment, Korea. Finally, we thank Kyo-Haeng Cho, president of Jung-haengsa, for her heartful contribution in printing this publication.

Materials and Methods 7

Taxonomic Notes

Order Lepidoptera

Family Gelechiidae Stainton, 1854

Subfamily Gelechiinae (A part of the subfamily)

Genus Teleiopsis Sattler, 1960

Teleiopsis Sattler, 1960, Dtl. ent. Z., N.F., 7: 16. Type-species: Recurvaria diffinis Haworth, 1828: 551.

Genus Teleiopsis is known to the Holarctic Region and its greatest species richness is known in thewestern Palaearctic (Pitkin, 1988). The genus belongs to one of the most specialized tribes Litini (=Teleiodini), including 12 species, of which three species: T. sophistica (Meyrick), T. diffinis (Haworth)and T. motleella Ponomarenko and Park, are recorded from East Asia (Sattler, 1982; Hodges, 1983;Huemer and Karsholt, 1999; Park and Ponomarenko, 2007). The male genitalia are characterizedby the elongated uncus, the tongue-shaped gnathos, the reduced cucullus, the glandiductors fusedwith sacculus basally, and the slender aedeagus. Host plants are known for nine species that belongto three families Anacardiaceae (Pistacia terebinthus, Rhus coriaria, Toxicodendron diversilobum), Poly-gonaceae (Rumex acetosella, R. lunaria, R. scutatus, Oxyria digyna), and Juglandaceae (Juglans regia).

106. Teleiopsis motleella Ponomarenko and Park, 2007

Teleiopsis motleella Ponomarenko and Park, 2007, Zootaxa, 1437: 55; Park and Ponomarenko, 2007:116.

Teleiopsis sophistica (nec Meyrick); Park, 2004: 64 (misid.).TL: Taebek, Korea.

ADULT: Wingspan 12 mm. The species is similar to Teleiopsis brevivalva Pitkin and T. sophistica(Meyrick) in the male genitalia. However, it differs from T. brevivalva by the uncus with round apexand the sacculus bilobed at right. It can be distinguished from T. sophistica by the asymmetric sac-culus, especially the left one, with the inner lobe as an indistinct small knob. In addition, the abdo-minal tergite of T. motleella has the posterior lobe slightly concave laterally and lacks prominentlateral lobes.

Male genitalia (Figs. 106A, B). See also Ponomarenko and Park (2007, Fig. 4); Park and Ponoma-

9

renko (2007, Pl. 43, Fig. 142). Uncus relatively broad, narrowed distally, with hook-shaped projec-tion at apex. Gnathos not reaching apex of uncus. Tegumen dilated basally, with broad anterior,lateral parts. Cucullus reduced. Sacculi asymmetric: right sacculus distinctly bilobed, its innerlobe about 1/3 as long as outer one; left sacculus with inner lobe represented by small knob; lengthof both sacculi from the glandiductor’s base to the apex longer than half the uncus. Glandiductorfused with sacculus basally, with distinct glandular channel. Aedeagus uniformly narrowed distal-ly, about four times longer than basal width, fused with anterior margin of vinculum by short tube.Straight part of ductus ejaculatori shorter than aedeagus. Abdominal tergite VIII with lateralbunches of strong spines, many of which have hooklike apices, arising from indistinctly separatedlobes; sternite VIII trapezoidal.

Female genitalia (Fig. 106C). Apophyses anteriores as long as 2/3 length of apophyses posteriors.Ostium bursae cup-shaped. Ductus bursae thin, membranous, about 1.5 times length of corpusbursae. Corpus bursae large; signum sickle-shaped, as long as half length of corpus bursae.

SPECIMENS EXAMINED: [GW] 1♂, Taebek, 7.ix.2000; [JN] 1♂, Mt. Peakwoon-san, Sooncheon, 19.viii.1996.

DISTRIBUTION: Korea (Central, South), Japan.


Fig. 106. Teleiopsis motleella. A. Male genitalia; B. Eighth abdominal sternite & tergite; C. Femalegenitalia with corpus bursae.

A C

B

S

T

Genus Pseudotelphusa Janse, 1958

Pseudotelphusa Janse, 1958, Moths S. Afr., 6: 68. Type-species: Telphusa probata Meyrick, 1909: 11.

Genus Pseudotelphusa is superficially similar to Carpatolechia Capuse, and the gnathos in male isabsent or reduced. It is characterized by the narrow and elongated uncus with the tapered apex;the gnathos absent; the glandiductor thin and pointed apically; the processes of juxta usually long,digitate or clavate apically. The genus has a Holarctic distributional range: 14 species are known tooccur in Europe, Asia, and eastern North America (Sattler, 1960, 1982; Huemer and Karsholt, 1999;Park and Ponomarenko, 2007; Lee and Brown, 2011), but only two species are represented in theEastern Palaearctic Region. The host plants of the species belong to Musci, Lichenes, Fagaceae,Salicaceae, Elaeagnaceae, Betulaceae, Rosaceae, Myricaceae and Berberidaceae (Sattler, 1980; Pisku-nov, 1981; Zhang, 1994; Huemer and Karsholt, 1999; Robinson et al., 2002; Lee and Brown, 2008).

107. Pseudotelphusa acrobrunella Park, 1992

Pseudotelphusa acrobrunella Park, 1992, Ins. Koreana, 9: 15; Omelko, 1999: 160; Li, 2002: 123; Park,2004: 59; Park and Ponomarenko, 2007: 116.

Pseudotelphusa fulvosella N. Omelko and M. Omelko, 1993, Biol. Invest. Gornot. Stn.: 220.TL: Yangyang, Korea.

ADULT: Wingspan 9-13 mm. This species is similar to the European species, P. scalella Scopoli,but it can be distinguished by the followings: the forewing ground color yellowish brown; theglandiductor shorter, about 2/3 length of genitalia and uncus bent ventrally in the male genitalia,whereas nearly straight in P. scalella. Head ochreous. Thorax yellowish brown. Second segment oflabial palpus roughly scaled, yellowish white, suffused with fuscous scales in basal half and sub-apical portion on outer surface; third segment slender, as long as second, with two dark stripes, oneat 1/3 and the other broad one preapically; apex white, sharply pointed. Forewings ground coloryellowish brown; basal fascia with three distinct scale tufts, at basal 1/5, 2/5, and 3/5; postmedianfascia convex outwardly, followed by yellowish white zone, beyond it dark fuscous scales scatter-ed. Hindwing gray. Abdominal tergite VIII with semicircular emargination on anterior margin;length of sternite VIII about 1/4 width.

Male genitalia (Figs. 107A-C). See also Park (1992, Figs. 14, 14-1); Omelko (1993, Fig. 11; 1999, Fig.24); Li (2002, Fig. 120); Park and Ponomarenko (2007, Pl. 43, Fig. 143). Uncus elongated, slightlydilated beyond the middle; cucullus reduced. Glandiductor inflated basally, slender in distal 3/4 oflength, with pointed apex. Processes of juxta dilated distally, nearly equal to glandiductor in length.

Female genitalia (Fig. 107D). See also Park (1992, Figs. 32, 32b, c); Omelko (1993, Fig. 12; 1999, Fig.25); Li (2002, Fig. 121); Park and Ponomarenko (2007, Pl. 84, Fig. 143). Apophyses posteriores abouttwice longer than length of apophyses anteriores. Ostium bursae surrounded by heavily sclerotiz-ed plate dorsally, forming hat-shaped, strongly emarginated at middle of posterior margin. Ductusbursae narrow. Corpus bursae longer than ductus bursae; signum hexagonal, with finely serratededges.

Lepidoptera: Gelechiidae: Gelechiinae 11

SPECIMENS EXAMINED: [GG] 1♂, Geomdan-san, 1.viii.2000; 2♂, Suweon, 1.vii.1987, 2.viii.1989; 1♂,Icheon, 1.vi.1980; 4♂, Gwangleung, 27.vi.1986, 8.vi.1987, 27.vi.1986; 4♀, Gwangleung, 8.vi.1977,10.vi.1998, 22.v.1998; 1♀, Mt. Yumyoung-san, 17.vi.1990; 1♀, Mt. Suri-san, 26.vii.1997; 1♀, 1♂, Mt.Myoungji-san, 23.v.1991, 25.v.1990; 1♀, Mt. Chelma-san, 14.vi.1996; 1♂, Mt. Cheonggye-san, 23.vii.1996; 1♀, Mt. Cheonggye-san, 25.vii.1996; 1♀, 1♂, Mt. Soyo-san, 17.v.1997; 1♂, Isl. Yongyudo,11.vii.1997; 1♂, Mt. Cheonma-san, 29.vii.1997; 1♀, 3♂, Mt. Hwaya-san, 4.viii.1997; 1♂, Icheon,1.vi.1980; [GW] 1♂, Mt. Gumgang-san, 23.vii.1982; 3exs, Mt. Gubong-san, 8.vi.1998, 20.vi.1998; 32♀,28♂, Chuncheon, 29.vii.1983, 10.vi.1985, 12.viii.1988, 11.vi.1989, 1.v.1989, 7.v.1989, 9.v.1989, 16.v.1989,26.v.1989, 29.v.1989, 13.vii.1989, 18.vii.1989, 28.v.1990, 2.vii.1990, 13.vi.1989, 16.v.1989, 19.vi.1990,15.viii.1992, 11.vi.1989, 16.v.1989, 29.v.1989, 29.v.1989, 19.vi.1990, 15.viii.1992; 1♂, Mt. Samak-san,19.vii.1989; 1♀, Dunnae, 7.vii.1990; 1♂, Hwengseong, 16.vii.1994; 1♀, Yangyang, 1.vii.1987; 2♂,Mt. Chiak-san, 23.vi.1987; 2♀, 1♂, Mt. Deokga-san, 16.vi.1997; 2♀, 1♀, Pyoungchang, 6.vi.1996,31.vii.1991; 2♂, 1♀, Yangyang, 4.vi.1987, 1.vii.1987, 1.vii.1987; 1♀, Mt. Seolak-san, 9.viii.1989; 1♂,Mt. Taebek-san, 27.vi.1996; 1♀, Mt. Daeduk-san, 11.vii.1997; 1♀, 1♂, Mt. Jeombong-san, 11.vii.1997,5.viii.1997; [CB] 1♀, Mt. Ingyeong-san, 8.vi.1997; [CN] 1♀, 1♂, Mt. Gaya-san, Yesan, 24.viii.1997;[JB] 2ex, Mt. Tonggo-san, 26-31.vi.2000; 2♀, Mt. Jiri-san, 22.vii.1981, 22.vii.1985; 1♂, Mt. Mireuk-san, 14.vi.1997; [JN] 5♀, Mt. Bekun-san, 19.viii.1992; [GN] 2♀, Namhae, 2.vi.1994; 1♀, Geochang,23.viii.1997; 1♀, Swamp Upo, Changyeong, 26.vii.1997; [JJ] 1♀, 2♂, Seongpanak, 23.viii.1992; 1♀,Yongsil, 22.viii.1992.

DISTRIBUTION: Korea (Central, South), Russian Far East (Primorsk, Kuril Isl.), Japan. HOST PALNTS: Fagaceae- Quercus sp. (Park, 1992).


Fig. 107. Pseudotelphusa acrobrunella. A, B. Male genitalia; C. Eighth abdominal sternite & tergite;D. Female genitalia with corpus bursae.

A B D

C

S

T

REMARKS: Larvae feed on leaves of Quercus sp. Moths appear from the early of May to the late ofAugust, and seem to be bivoltine in Korea.

Genus Concubina N. Omelko and M. Omelko, 2004

Concubina N. Omelko and M. Omelko, 2004, Biol. Invest. Gornot. Stn., 9: 193. Type-species: Concubina subita N. Omelko and M. Omelko, 2004: 193.

Genus Concubina includes two species in the Eastern Palaearctic. The male genitalia are charac-terized by the uncus elongate, pointed; the gnathos with mesial projection club-shaped, the ventralsclerite (fused vinculum and sacculi) with pair of processes.

108. Concubina trigonalis Park and Ponomarenko, 2007

Concubina trigonalis Park and Ponomarenko, 2007, Proc. Ent. Soc. Wash., 109: 807.TL: Mt. Seolak-san, Korea.

ADULT: Wingspan 14-15 mm. This species is similar to C. euryzeucta (Meyrick) which is distri-buted in the Russian Far East, but it is distinguished by the gray forewing ground color with dark-gray basal obliqua fascia and two small costal spots: one before the middle and the other beyondthe middle of the margin, and with indistinct fuscous spots on the tornus and apex, whereas in thelatter, the forewing ground color is white gray with a large triangular black sport at middle of costalmargin. Vein R3 separated from R4+5; R4 and R5 stalked at middle; R5 to costa; M1 arising from cellseparately; M2 close to M3 at base; distance M3-CuA1 about 1/2 of CuA1-CuA2. Hind wing gray;apex acute; termen strongly sinuate; fringe gray. Veins Sc and Rs connected with a crossvein atbasal 1/3 of Sc; Rs and M1 separated beyond cell; M2 absent; M3 and CuA1 separated.

Male genitalia (Figs. 108A-C). See also Park and Ponomarenko (2007, Pl. 44, Fig. 146). Abdominaltergite VIII subtriangular, rather small with round apex, anterior margin roundly concave, shorterthan length of sernite VIII. Abdominal sternite VIII trapezoidal, relatively short; anterior marginconvex medially. Uncus slightly narrowed towards bilobed apex. Tegumen broad, divided intotwo relatively wid parts anteriorly, with deep emargination on anterior margin reaching to one halflength, with two triangular lobes on distal part. Gnathos with mesial projection club-shaped, sub-equal in length with uncus. Ventral sclerite formed by fused sacculi and vinculum, relatively wide,with two triangular plates posteriorly; saccus absent. Aedeagus stout, tubelike, much wider basal-ly, narrowed gradually towards apex.

Female genitalia (Fig. 108D). See also Park and Ponomarenko (2007, Pl. 85, Fig. 146). Ovipositorlong, membrane between segment VIII and IX, more than 5.5 times as long as papillae anales.Apophyses posteriors about 3 times as long as apophyses anteriores. Abdominal segment VIIIannular, with longitudinal gutter near antrum along anterior margin. Antrum sclerotized laterally,narrowed near middle. Ductus bursa membranous, about twice as long as corpus bursae, withdilated anterior half; ductus seminalis arising from distal part. Corpus bursa oval, membranous,


with dense spicules on inner surface; signum rhomboidal, large, with serrated margins and a trans-verse furrow medially.

SPECIMENS EXAMINED: [GG] 2♀, Hanam, 25.vii.1996; [GW] 1♂, Mt. Seolak-san, 10.viii.1989; 1♂,Youngwol, 14.vii.1998; 2♀, 1♂, Mt. Taewha-san, 14.vii.1998, 1.viii.2000.

DISTRIBUTION: Korea (Central).REMARKS: Concubina trigonalis is similar to C. euryzeucta Meyrick which was described in the

genus Telphusa Chambers from China, in the structure of the male and female genitalia, but it differsfrom the latter species in having the uncus slightly narrowed distally and bilobed at its apex, theventral sclerite with triangular plates posteriorly instead of plates rounded on distal margin of eury-zeucta Meyrick in the male genitalia, and longer sternite VIII in the female genitalia.

Genus Altenia Sattler, 1960

Altenia Sattler, 1960, Dt. ent. Z. (N.F.), 7: 16. Type-species: Gelechia perspersella Wocke, 1862: 236.


Fig. 108. Concubina trigonalis. A. Male genitalia; B. Aedeagus; C. Eighth abdominal sternite &tergite; D. female genitalia.

C D

B

A

Genus Altenia has the Palaearctic distributional range, and only one species is known from theEast Asia. The genus is characterized by the bifid uncus with very long and narrow lobes; the gna-thos and the cucullus reduced; the glandiductor, if present, arched and slender; the processes ofjuxta short; the aedeagus stout. The host plants of the species belong to Empetraceae, Anacardia-ceae and Aceraceae (Huemer and Karsholt, 1999).

109. Altenia inscriptella (Christoph, 1882)

Teleia inscriptella Christoph, 1882, Bull. Soc. Nat. Mosc., 57(1): 25. Telphusa inscriptella: Meyrick, 1925: 71; Gaede, 1937: 127; Park, 1992: 17.Altenia inscriptella: Park, 2004: 63; Park and Ponomarenko, 2007: 120.

TL: Radde, Askold Isl., Primorsk Territory, Russia.

ADULT: Wingspan 12-16 mm. This species is close to the European species, A. scriptella in thesuperficial and genital characters, but it is separable by the creamy white ground color of the fore-wings, whereas ground color of the latter is yellowish white, especially yellowish beyond medianfascia.

Male genitalia (Figs. 109A-C). See also Park (1992, Figs. 19, 19-1); Park and Ponomarenko (2007,Pl. 45, Fig. 147). Abdominal sternit VIII very short and wide, about 1/3 length of width, distal


Fig. 109. Altenia inscriptella. A, B. Male genitalia; C. Eighth abdominal sternite & tergite; D. Femalegenitalia with corpus bursae.

A C D

B

T

S

margin slightly emarginated medially; tergite VIII elongate, with round distal margin; anteriormargin deeply emarginated. Uncus deeply bifurcated; lateral arms sickle-shaped, far from eachother at base. Processes of juxta digitate, about half length of arms of uncus. Glandiductor absent.Aedeagus very stout, narrowed beyond half, with sharp distal end.

Female genitalia (Fig. 109D). See also Park (1992, Figs. 37, 37c); Park and Ponomarenko (2007, Pl.85, Fig. 146). Apophyses posteriores slightly longer than apophyses anteriores. Ostium bursaecrescent. Ductus bursae long. Corpus bursae about half length of ductus bursae; signum large,with narrow groove centrally.

SPECIMENS EXAMINED: [GG] 1♀, Suweon, 15.v.1989; 3♂, Gwangleung, 8.vi.1977, 10.v.1994; 1♀, 2♂,Mt. Myoungji-san, 25.v.1990, 23.v.1991; 4♀, 3♂, Isl. Yeongjongdo, 26.v.1995, 21.vii.1995; 4♀, 2♂,Mt. Soyo-san, 5.viii.1996, 17.v.1997; 1♂, Mt. Hwaya-san, 18.vii.1997; 1♀, Mt. Cheonma-san, 29.vii.1997; [GW] 1♂, Mt. Hyinbau-san, 23.v.2000; 2ex, Yangyang, 30.v.1987, 4.vi.1987; 1♀, 1♂, Chun-cheon, 3.v.1995, 28. iv.1998; 1♂, Yangyang, 30.v.1989; 2♀, Yangyang, 4.vi.1987; 2♂, Mt. Odae-san,22.v.1989; 3♀, 3♂, Mt. Jeombong-san, 20.vi.1992, 22.vi.1992; 1♂, Pyoungchang, 6.vi.1996; 1♂, Mt.Gyebang-san, 14.vi.1995; 1♀, 1♂, Mt. Gwangdeok-san, 23.vi.1994, 3.vi.1995; 1♂, Mt. Gyebang-san,21.vi.1996; 3♀, 1♂, Mt. Taebek-san, 27.vi.1996; 1♀, Mt. Daeduk-san, 11.vii.1997. [CB] 2♀, 1♂, Mt.Weolak-san, 7.vi.1997; 2♀, Mt. Ingyeong-san, 23.viii.1997; [JB] 1♂, Mt. Tonggo-san, 26-29.vi.2000;[JJ] 3♀, Youngsil, 30.IV.1994, 2.vii.1994.

DISTRIBUTION: Korea (Central, South, Jeju), Russian Far East (Primorsk), Japan. HOST PLANT: Aceraceae- Acer ginnala (Park, 1992).

Subfamily Anacampsinae

Anacampsinae Bruand, 1850, Mém. Soc. Emul. Doubs., (1)3(5-6): 40.

Tribe Anacampsini

Anacampsini Bruand, 1850, Mém. Soc. Emul. Doubs., (1)3(5-6): 40.Type-genus: Anacampsis Curtis, 1827.

Genus Syncopacma Meyrick, 1925

Syncopacma Meyrick, 1925, In Wytsman, Genera Insects., 184: 14.Type-species: Telphusa acrophylla Meyrick, 1911: 65.

Genus Syncopacma includes more than 20 Palaearctic species. Among them, 16 species are repre-sented in the East Asia. In China, Li (2002) reviewed five species, but Ponmarenko et al. (2006)


reported S. ochrofasciella (Toll, 1936) and S. wormiella (Wolf, 1958) from Mt. Channgbai-shan, Jilin,which is located in border China and N. Korea. Thus, these two species probably occur in No.Korea. Male genitalia are characterized by followings: the short uncus with long setae, often flaton the caudal margin; the valva usually with distinct lobe on the ventral margin; the sacculus well-developed with the different shape of apex; the aedeagus usually with cornutus. The host plantsbelong to Fabaceae and Linaceae (Piskunov, 1981; Elsner et al., 1999).

110. Synacompa wormiella (Worff, 1958)

Stomopteryx wormiella Worff, 1958, Entomol. Med., 26: 244.Synacompa wormiella; Piskunov, 1981: 738; Karsholt and Riedl, 1996: 119; Elsner et al., 1999: 51;

Omelko, 1999: 165; Ponomarenko et al., 2006: 112; Park and Ponomarenko, 2007: 149.

ADULT: Wingspan 13 mm. This species is superficially similar to S. ochrofasciella (Toll) which alsooccurs in the same area with this species, but can be distinguished by the narrower postmedianband of the forewing and the male genitalia. The specimen could not be compared by the genitalia,because of missing abdomen, but it is no doubt its conspecific by the superficial characters. Femaleis unknown.

Male genitalia (Figs. 110A, B). See also Ponomarenko and Park (2006, Fig. 30); Park and Ponomarenko

Lepidoptera: Gelechiidae: Anacampsinae 17

Fig. 110. Syncopacma wormiella. A. Male genitalia; B. Female genitalia with corpus bursae.

A B

(2007, Pl. 45. Fig. 151). Cucullus dilated distally; sacculus shorter than length of tegumen+uncus,whereas it is heavily sclerotized, long, curved medially; vinculum with large median process.

Female genitalia (Fig. 110C). See also Ponomarenko and Park (2006, Fig. 40); Park and Ponomarenko(2007, Pl. 85. Fig. 151).

SPECIMENS EXAMINED: [GW]- 1♂ (Mt. Taewha-san, Youngwol: 27.v.1998 (abdomen missing).DISTRIBUTION: Korea (Central), China (Jilin), Russia (European part and Russian Far East), Europe. REMARKS: This species was also reported from Mt. Changbai-shan which is located in border

between the northerneast part of China and N. Korea (Ponomarenko et al., 2006).

Genus Aproaerema Durrant, 1897

Aproaerema Durrant, 1897, Entomol. month. Mag., 33: 221. Type-species: Tinea anthyllidella Hübner, [1813]: pl. 48, Fig. 330.SYNONYM: Schuetzeia Spuler, 1910.

Genus Aproaerema comprises three species, with a Holarctic distributio. The male genitalia arecharacterized by the uncus bilobed; the cucullus arched dorsally; the aedeagus straight tube-likewith long basal ansiform process, which longer than aedeagus; and the saccus large, gutter-like.The host plants belong to Fabaceae (Piskunov, 1981; Elsner et al., 1999).

111. Aproaerema anthyllidella (Hübner, [1813])

Tinea anthyllidella Hübner, 1813, Samml. Eur. Schmett., Lepid., VIII: t. 43, Fig. 330. Gelechia (Anacampsis) sparsiciliella Barrett, 1891, Ent. Mon. Mag., 27: 7. Anacampsis anthyllidella: Herrich-Schäffer, 1855: 195.Stomopteryx anthyllidella: Meyrick, 1925: 111; Pierce and Metcalfe, 1935: 18; Gaede, 1937: 322; Wolff,

1958: 262. Aproaerema anthyllidella: Durrant, 1897: 221; Issiki, 1957: 44; Piskunov, 1981: 742; Moriuti, 1982, I: 281,

II: 214; Park, 1983a: 87; Park, 1983b: 497; Park, 1991c: 121; Karsholt and Riedl, 1996: 119; Elsner etal., 1999: 52; Bland et al., 2002: 206; Park, 2004: 67; Park and Ponomarenko, 2007: 123.TL: Europe.

ADULT: Wingspan 8-11 mm. Forewing elongate, with yellowish white costal patch beyond 3/4;narrower towards apex beyond middle; ground color yellowish brown; apex acute. Hindwingaswide as forewing; apex sharply produced; termen falcate.

Male genitalia (Fig. 111A). See also Pierce and Metcalfe (1935, Pl. X); Piskunov (1981, Figs. 670: 4,5); Park (1983, Fig. 158); Elsner et al. (1999, Taf. 35, Fig. 308); Omelko (1999, Fig. 86: 2); Bland et al.(2002, Fig. 16g); Park and Ponomarenko (2007, Pl. 45. Fig. 151). Uncus bilobed with rounded apex;cucullus arched dorsally, slightly dilated at the middle; sacculus small triangular. Aedeagus nar-rowed towards apex with longer rod-like process.


Female genitalia (Fig. 111B). See also Pierce and Metcalfe (1935, Pl. X); Elsner et al. (1999, Taf. 77,Fig. 308); Bland et al. (2002, Fig. 40e).

SPECIMENS EXAMINED: [N. Korea] 1♀, Sariwon, N. Hwanghae, 28.ix.1978; 1♂, Pyoungyang, 15.ix.1979; [GG] 1♀, Suweon, 12.vi.1974; 1♀, Gwangleung, 40 km NE Seoul, 13.vii.1975; 1♀, 1♂, Gwang-leung, 10.vii.1982; [GW] 1♀, Chuncheon, 15.viii.1992; 1♀, Bongmyoungri, 20 km SE Chuncheon,5.vi.1995; 1♀, Chuncheon, 3.vi.1997; 1♀, Mt. Samak-san, Chuncheon, 19.vii.1989; 1♂, Mt. Gubong-san, Chuncheon, 20.vi.1998; 1♀, Mt. Bangtae-san, Inje, 23.vi.1996; 1♀, Mt. Obong-san, 1.vii.1993; 2♂,Mt. Gyebang-san, 21.vi.1996; [JB] 1♀, Mt. Mireuk-san, 14.vi.1997; [JN] 2♂, Mt. Bekun-san, 19.viii.1992.

DISTRIBUTION: Korea (North, Central, South), Japan (Honshu), China (Shaanxi, Ningxia, Qinghai,Xinjiang), Russia (European part, Caucasus, S. Ural, Altai, Irkutsk, Transbaikalia, Primorsk), AsiaMinor, Europe, N. America.

HOST PALNTS: Leguminosae- Medicago sp., Anthyllis polyphylla, A. vulneraria, Anthyllis spp., Trifoliumpratense, Trifolium sp., Onobrychis viciifolia, O. arenaria, Onobrychis sp., Ononis spinosa, O. campestris,O. repens, Glycine sp., Galega officinalis, Lathyrus, Lotus, Melilotus, Oxytropis pilosa, Phaseolus vulgaris,Trigonella monspeliaca, Arachis sp., Amoria repens, and includes Chamaecytisus, Chrysapsis, Coronilla,and Dorycnium (Piskunov, 1981; Park, 1983; Emmet, 1988; Elsner et al., 1999).

REMARKS: This species was first reported from Korea by Park (1983). Larva makes white blotch-mines on the leaves of host plants, and on the blossom and seed-pods of Anthyllis and perhaps theother food plants. Pupation takes place in detritus on the ground in May.


Fig. 111. Aproaerema anthyllidella. A. Male genitalia; B. Female genitalia with corpus bursae.

A B

Genus Anacampsis Curtis, 1827

Anacampsis Curtis, 1827, Brit. Ent.: 189. Type-species: Phalaena populella Clerck, 1759, pl. 11, Fig. 5.SYNONYMS: Tachyptilia Heinemann, 1870; Agriastis Meyrick, 1914.

Genus Anacampsis comprises about 50 species and widely distributed in Old and New World. Thegenus is well-known in N. America, but it is poorly known in the eastern Asia, with eleven knownspecies. The host plants of the species belong to Salicacea, Fagaceae, Betulaceae, Aceraceae, Rosa-ceae, Cistaceae, Lamiaceae, Fabaceae, Rosaceae (Piskunov, 1981; Elsner et al., 1999).

Key to the species of the genus Anacampsis Curtis, based on the male genitalia

1. Forewing with well-developed white postmedian line. Valva of male genitalia exceeding apexof uncus····································································································································2

- Forewing with postmedian line not presented. Valva of male genitalia not exceeding apex ofuncus··························································································································A. lignaria

2. Postmedian line of forewing concave on outer margin; white costal patch developed. Aedeagusof male genitalia more or less slender beyond basal 2/5 length; apex acute········A. anisogramma

- Postmedian line of forewing concave on outer margin; white costal patch developed. Aedeagusof male genitalia slender beyond basal 2/5 length; apex truncate····························A. solemnella

112. Anacampsis anisogramma (Meyrick, 1927)

Compsolechia anisogramma Meyrick, 1927, Exot. Microlep., 3: 353; Gaede, 1937: 348; Caradja, 1938:111; Issiki, 1957: 43; Arita, 1967: 79; Moriuti, 1982, I: 280, II: 213; Park, 1983a: 87; Park, 1983b: 496;Omelko, 1999: 161.

Anacampsis anisogramma: Park, 1988: 142; ESK and KSAE, 1994: 328; Omelko, 1999: 161; Li, 2002: 198;Park, 2004: 66; Park and Ponomarenko, 2007: 125.TL: Shanghai, China.

ADULT: Wingspan 12.5-16.5 mm. The species is characterized by the brownish gray forewingground color with blackish postmedian fascia followed by an orange-white costal patch.

Male genitalia (Figs. 112A, B). See also Arita (1967, Fig. 3); Park (1983, Fig. 156; 1988, Figs. 19-21);Omelko (1999, Fig. 82: 1); Li (2002, Fig. 229). Uncus short, more or less crown-shaped, broader thandistal part of tegument. Gnathos without median hook; Valva long, slender, smoothly bent at mid-dle, slightly narrower towards base, longer than total length tegumen+uncus; sacculus triangularwith rounded apex. Aedeagus inflated in basal part, slightly bent at middle, blunted at apex, witha long furrow-like tube situated closer to ventral than dorsal wall.

Female genitalia (Fig. 112C). See also Arita (1967, Fig. 4); Park (1988, Fig. 23); Li (2002, Fig. 230).Ostial plate narrow, extending posteriorly. Ductus bursae narrow, slender, longer than corpusbursae. Corpus bursae with arched serrated signum. Distal margin of sternite VII emarginatedmedially.


SPECIMENS EXAMINED: [GG] 1♂, Suweon, 15.vi.1969; [GW] 2♀, 9♂, Chuncheon, 21.v.-10.vi.1983.DISTRIBUTION: Korea (Central), Russian Far East (Primorsk), China (Shanghai, Jiangxi, Shan-

dong, Shaanxi, Sichuan, Guizhou, Fujian), Japan (Hokkaido, Honshu, Shikoku, Kyushu, Ryuku),Taiwan.

HOST PALNTS: Rosaceae- Prunus mume, P. salicin, P. persica, P. armeniaca var. ansu, P. serrulata var.spontanea, P. yedoensis Matsumura, Pyrus avium L. Cerasus tomentosa, C. glandulosa, Armeniaca mand-shurica, Amelanchier, Aronia melanocarpa (Arita, 1967; Park, 1988; Omelko, 1999).

REMARKS: Larva rolls the leaves of host plants horizontally and feeds on them.

113. Anacampsis lignaria (Meyrick, 1926)

Compsolechia lignaria Meyrick, 1926, Exot. Microlep., 3: 292; Gaede, 1937: 352; Clarke, 1969: 489. Anacampsis lignaria: Park, 1988: 147; Omelko, 1991: 146; ESK and KSAE, 1994: 328; Omelko, 1999: 161;

Park, 2004: 66; Park and Ponomarenko, 2007: 126.Compsolechia homoplasta Meyrick, 1932: 197; Issiki, 1957: 44; Arita, 1967: 82; Clarke, 1969, 6: 486;

Saito, 1969: 115; Moriuti, 1982, I: 280, II: 213; Omelko, 1999: 163. TL: Khabarovsk, Russian Far East.


Fig. 112. Anacampsis anisogramma. A. Male genitalia; B. Aedeagus; C. Female genitalia with corpusbursae.

A B C

ADULT: Wingspan 14-19 mm. There are considerable variations in wing size and color pattern.Head, thorax and ground color of forewing grayish ochreous in the pale form and somewhat darkbrown in the dark form. Costal edge of forewing yellowish ochreous except beyond 3/4. Two blackmarginal dots on costa around apex and 3-4 others along termen; discal stigmata dark fuscous,indistinct. Hindwing gray or dark gray.

Male genitalia (Figs. 113A, B). See also Arita (1967, Fig. 5); Park (1988, Figs. 33, 34); Omelko (1991,Fig. 17; 1999, Figs. 83: 1, 2). Uncus broader than distal part of tegument. Valva shorter than totallength tegumen+uncus; sacculus triangular. Aedeagus large, much dilated in basal half, slightlylonger than total length of genitalia.

Female genitalia (Fig. 113C). See also Arita (1967, Fig. 6); Clarke (1969(6), pl. 242: Figs. 1a, b); Park(1988, Fig. 36); Omelko (1999, Figs. 83: 3, 4). Ostial plate semioval and elongated distally, corpusbursae without signum.

SPECIMENS EXAMINED: [GW] 1♀, 2♂, Mt. Odae-san, 26.vi.1989; 1♀, Yangyang, 25.vii.1987; 2♀,Mt. Seolak-san, 10.viii.1989; 1♀, Mt. Yaksu-san, 9.viii.1989; 1♀, Pyeongchang, 31.vii.1991; [GN] 1♀,1♂, Mt. Geum-san, Namhae, 25.vii.1985, reared from Rhododendron sp.).

DISTRIBUTION: Korea (Central, South), Japan (Honshu), Russia (Amursk, Khabarovsk, Primorsk). HOST PLANT: Ericaceae- Rhododendron spp. (Park, 1988). REMARKS: Clarke (1969) indicated that the specimen marked ‘Type’ in the British Museum (Nat.

Hist.) was a male, but erroneously recorded as a female by Meyrick (1926).


Fig. 113. Anacampsis lignaria. A. Male genitalia; B. Aedeagus; C. Female genitalia.

A B C

114. Anacampsis solemnella (Christoph, 1882)

Tachyptilia solemnella Christoph, 1882, Bull. Soc. Imp. Nat. Moscou, 57(1): 27. Compsolechia agrimoniella var. solemnella: Meyrick, 1925: 123; Gaede, 1937: 347; Issiki, 1957: 44.Comsolechia solemnella: Moriuti, 1982, I: 281, II: 213; Park, 1983a: 87; Park, 1983b: 496; Omelko, 1999:

163. Anacampsis solemnella: Park, 1988: 144; Park, 1991c: 121; ESK and KSAE, 1994: 328; Li, 2002: 201;

Park, 2004: 67; Park and Ponomarenko, 2007: 128.Compsolechia metagramma Meyrick, 1918, Exot. Microlep., 2: 138.

TL: Amur, Russia.

ADULT: Wingspan 10-15 mm. This species is superficially similar to A. anisogramma Meyrick, butcan be distinguished by the well developed postmedian on the forewing which is clearly extendedto dorsum, curved inwardly, and much paler beyond the line.

Male genitalia (Figs. 114A, B). See also Park (1983, Fig. 157; 1988, Figs. 24, 25); Omelko (1999, Figs.84: 5, 6); Li (2002, Fig. 232). Valva exceed apex of uncus; sacculus narrowed towards pointed apex,slightly hollowed on ventral margin. Aedeagus narrowed towards apex, arched in distal 2/3 oflength; apex pointed.

Female genitalia (Fig. 114C). See also Park (1988, Fig. 27); Omelko (1999, Fig. 84: 7); Li (2002, Fig.


Fig. 114. Anacampsis solemnella. A. Male genitalia; B. Aedeagus; C. Female genitalia with corpusbursae.

A B C

233). A sclerotized plate beyond tergite VII differs from those of allied species, with lateral flapsfolding together, with deep emargination. Corpus bursae large, without signum.

SPECIMENS EXAMINED: [N. Korea] 1♀, Samjiyeon, Mt. Pektu-san, 19.vii.1977; [GG] 2♀, 1♂, Mt.Cheonma-san, 5.ix.1995, 3.ix.1996; 1♀, Mt. Myoungji-san, 5.v.1996.

DISTRIBUTION: Korea (North, Central), China (Beijing, Heilongjiang, Liaoning, Shaanxi, Sichuan,Anhui, Henan, Jiangsu, Zhejiang), Russia (Altai, Irkutsk, Buryatia, Transbaikalia, Amursk, Khaba-rovsk, Primorsk, Kuril Isl.), Japan (Hokkaido, Honshu).

HOST PALNTS: Rosaceae- Spiraea nipponica, S. betulifolia,, Spiraea sp. (Park, 1988).

Genus Prolita Leraut, 1993

Prolita Leraut, 1993, Alexanor Paris, 18(3): 182. Type-species: Lita zebrella Treitschke, 1833: 82.SYNONYM: Lita Treitschke, 1833.

Genus Prolita includes more than 20 species which are known in Holarctic Region. The hostplants of the species belong to Ericaceae, Rosaceae and Papilionaceae (Huemer and Karsholt, 1999).

115. Prolita sexpunctella (Fabricius, 1794)

Tinea sexpunctella Fabricius, 1794, Ent. Syst., 1794: 313.Tinea virgella Thunberg, 1794, D.D. Dissert. ent. sist. Ins. Svecica, 7: 92.Anacampsis longicornis Curtis, 1827, Brit. Ent., 4: N 189.[Tinea] histrionella Geyer, [1832], in Hübner, Samm. eur. Schmett., 8: pl. 70, Fig. 464.Lita zebrella Treitschke, 1833, Schmett. Ent., 9: 82.Gelechia longicornella Doubleday, 1859, Zool. syn. list. Br. Butterflies and Moths, 2: 30.Gelechia longicornis var. alpico Frey, 1867, Mitt. schweiz. ent. Gaz., 2: 299.Gelechia petulans Braun, 1925, Can. Ent., 57: 125.Prolita sexpunctella: Karsholt and Riedl, 1996: 113; Elsner et al., 1999: 39; Huemer and Karsholt, 1999:

172; Bland et al., 2002: 148; Park and Ponomarenko, 2007: 129.TL: Italy, Europe.

ADULT: Wingspan 17-18 mm. Head and thorax brownish mottled with light gray. Antennablackish. Labial palpus dark brown, more or less mottled with whitish. Forewing elongate, blackishbrown mottled with brownish; a black spot near base, one in fold two in cell; four light gray trans-verse fasciae. Hindwing dark gray. A considerable variations with regard to forewing color.

Male genitalia (Fig. 115A). See also Piskunov (1981, Figs. 673: 4, 5); Elsner et al. (1999, Taf. 20, Fig.178); Huemer and Karsholt (1999: Fig. 140); Bland et al. (2002, Fig. 10f). Uncus broad, with roundedapex; cucullus slender, clavate distally; sacculi fused with vinculum, extended distally, aedeaguswith long cecum, slightly curved and with small thorn before apex.


Female genitalia. See Elsner et al. (1999, Taf. 63, Fig. 178); Huemer and Karsholt (1999, Fig. 140);Bland et al. (2002, Fig. 32a).Antrum moderately long; coliculum small, ring-shaped. Externallysimilar to P. solitutella (Zeller) which is known in Europe, with weak differences in the shape of theantrum and colliculum.

SPECIMENS EXAMINED: [N. Korea] 1♂, Samjiyeon, Mt. Pektu-san, 1400 m, 16.vi.1988; 1♂, samelocality, 21.vi.1988.

DISTRIBUTION: Korea (North), Russia (European part, Altai, Tuva, Irkutsk region, Trasbaikalia,NW Yakutia, Khabarovsk), Europe, N. America.

HOST PALNTS: Ericaceae- Calluna vulgaris, Calluna sp., Erica cinerea, Erica herbacea, Vaccinium uligi-nosum, V. myrtillus. Rosaceae- Dryas octopetata (Piskunov, 1981; Huemer and Karsholt, 1999; Elsneret al., 1999).

REMARKS: This species was reported for the first time from Korea by Park and Ponomarenko(2007), based on specimens collected in Mt. Pektu-san, N. Korea.


Fig. 115. Prolita sexpunctella. A. Male genitalia; B. Male genitalia with aedeagus (after Elsner et al.,1999).

A B

Genus Mesophleps Hübner, [1825]

Mesophleps Hübner, [1825], Verz. bek. Schmett.: 406. Type-species: Tinea silacella Hübner, 1796: 37. SYNONYMS: Brachyacma Meyrick, 1886; Lathontogenus Walsingham, 1897; Paraspistes Meyrick, 1905;

Lipatia Busck, 1910.

Genus Mesophleps comprises 37 species, which are mostly distributed in the Palaearctic Region,three of them known from South Africa, India, Sri Lanka and Seychelles and one exists in CentralAmerica. The adults are average-sized gelechiids with wingspan of 10-25 mm, and the larvae areseed feeders in fruits and pods of several plant families but predominantly Leguminosae (Li andSattler, 2012). The genus was established as monophyletic by at least two synapomorphies: thetransverse bands of microtrichia on the anterior margin of abdominal tergites IV-VIII in both sexesand the double gnathos in the male genitalia. Park (1990) described two species belonging to Bra-chyachma Meyrick for the first time from the eastern Asia, and these species were transferred to thegenus Mesophleps. The genusis native to temperate and tropical parts of the Old world. It remainsuncertain whether M. adustipennis from southern North America, Central and South America. Thehost plants of the genus include Cistaceae, Cupressaceae, Dipterocarpaceae, Cruciferae, Legumino-sae, Rubiaceae (Li and Sattler, 2012).

Key to the species of the genus Mesophleps Hübner

1. Gnathos of male genitalia with two small median hooks; cucullus slender and dilated towardsrounded apex; aedeagus longer than cucullus·························································M. albilinella

- Gnathos of male genitalia with two lateral process···································································22. Uncus large, tongue-shaped; lateral processes of gnathos digitate, heavily sclerotized, shorter

than uncus·············································································································M. sublutiana- Uncus rather small, triangular; lateral processes of gnathos conic, heavily sclerotized, as long as

uncus·······················································································································M. acutunca

116. Mesophleps albilinella (Park, 1990)

Brachyacma albilinella Park, 1990, Kor. J. Appl. Ent., 29(2): 136. Mesophleps albilinella: Li and Zheng, 1995: 28; Li, 2002: 214; Park, 2004: 68; Li and Sattler, 2012: 42.

TL: Chuncheon, Korea.

ADULT: Wingspan 11-14 mm. Labial palpus recurved, swecond segment blackish, third segmentblackish in basal half, white to yellowish white distally; apex grayish black. Antenna grayish white,with light-brown annulations, blackish apically. Thorax and tegula creamy white. Forewing rightorange to yellowish white; costal margin dark, dark area wider in distal 3/5, interrupted at 3/4 bydistinct oblique white line directed towards termen; black discal and plical spots present; innermargin densely scattered with brown scales; fringe grayish white. Hindwing dark gray. Abdomenlight gray to yellowish brown.


Male genitalia (Figs. 116A, B). See also Park (1990, Figs. 6-8); Li and Sattler (2002, Fig. 76) Uncustriangular, with rounded apex. Gnathos with two sclerotized lateral hook-shaped processes. Valvaslender, dilated towards apex. Aedeagus inflated in basal 1/3, then slender, longer than length ofvalva.

Female genitalia (Fig. 116C). See also Park (1990, Fig. 9); Li and Sattler (2002, Fig. 105). Apophysesposteriores about twice length of apophyses anteriores. Ostium placed on level of anterior marginof abdominal sternite VIII. Ductus bursae about five times length of apophyses anteriores; ductusseminalis thin, inception close to entrance of ductus bursae. Corpus bursae oval to sub-spherical.

SPECIMENS EXAMINED: [GG] 1♀, 2♂, Mt. Acha-san, 12.vi.1997; 1♀, Mt. Bulam-san, 22.vii.1998; 2♂,Mt. Surak-san, 22.vii.1998; 3♂, Mt. Bugak-san, 23.vii.1998; 1♂, Mt. Inwang-san, 23.vii.1998; 2♂,Ipori, 20.viii.1990; 1♂, Mt. Suri-san, Gunpo, 25.vii.1997; 1♀, 1♂, Mt. Soyo-san, 3.?.1992; 2♂, 1♀,Gwangleung, 4.viii.1986, 13.viii.1986, 8.viii.1992; 1♀, 1♂, Hongleung, 13.vi.1996; 1♀, 1♂, Hong-leung, 13.vi.1996; 3♀, Mt. Chelma-san, Incheon, 14.vi.1996; 1♀, Mt. Cheonma-san, 13.vii.1996; 9♀,16♂, Mt. Soyo-san, Dongduchon, 5.viii.1996; 2♀, Mt. Soyo-san, 7.ix.1996, 7.vii.1997; 1♂, Isl. Yong-yudo, Incheon, 28.viii.1997; 4♂, Mt. Hwaya-san, Gapyoung, 18.vii.1997, 4.viii.1997; 1♀, Mt. Cheong-gye-san, Yangso, 25.vii.1996; [GW] 1♀, 5♂, Chuncheon, 26.vi.1984, 31.viii.1984, 30.viii.1990, 15.viii.1992, 16.viii.1996; 3♂, Chugok, Chuncheon, 2.ix.1996; 3♀, 7♂, Chuncheon, 16.viii.1996, 12.vi.1998;1♀, 2♂, Mt. Gubong-san, 8.vi.1998; 3♂, 1♂, Mt. Samak-san, Chuncheon, 19.viii.1986, 13.vi.1990;


Fig. 116. Mesophleps albilinella. A. Male genitalia; B. Aedeagus. C. Female genitalia with corpusbursae.

A B C

20♂, Hongcheon, 5.vii.1990; 7♂, Mt. Palbong-san, Hongcheon, 3.ix.1990; 2♀, Mt. Gyejok-san,Youngweol, 27.v.1998; 1♀, Mt. Obong-san, 1.vii.1993; 1♂, Mt. Gyebang-san, Pyoungchang, 2.vii.1997; 2♂, Mt. Jeombong-san, 11.vii.1997; 1♀, Mt. Deokga-san, 24.vii.1997; 1♂, Mt. Gwangdeok-san, 20.vii.1996; 2♂, Yangyang, 25-26.vii.1987; [CB] 1♀, Mt. Weolak-san, 9.viii.1997; 1♀, SwampUpo, Changyeong, 28.vii.1997; [JB] 2♂, Mt. Mireuk-san, Iksan, 14.vi.1997, 23.viii.1997; [GN] 1♀,Mt. Gaji-san, 19.viii.1993.

DISTRIBUTION: Korea (Central, South), China (Shaanxi, Gansu, Henan), Japan. REMARKS: Moths appear mostly from the end of May to the early of September in Korea.

117. Mesophleps sublutiana (Park, 1990)

Brachyacma sublutiana Park, 1990, Kor. J. Appl. Ent., 29(2): 139. Mesophleps sublutiana: Li and Zheng, 1995: 28; Li, 2002: 212; Park, 2004: 69; Li and Sattler, 2012: 26.

TL: Suweon, Korea.

ADULT: Wingspan 11-14 mm. Labial palpus variable; second segment thin, wider distally, blackwith white band distally; third segment slightly shorter than 2nd, white, sometimes speckled withblack scales basally. Antenna alternating white and brown. Forewing yellowish to ochreous brown,distal 3/5 of costa black, interrupted by faint oblique line running from 4/5 towards termen, some-


Fig. 117. Mesophleps sublutiana. A. Male genitalia; B. Aedeagus; C. Female genitalia; D. Abdomen.

A

B

C

D

times posterior margin darker; black discal and plical spots variably developed; termen with 2-3small black spots near apex.

Male genitalia (Figs. 117A, B). See also Li and Sattler (2002, Fig. 62). Uncus trapezoid and with flatslightly hollowed apex; gnathos with two large median hooks; cucullus slender, dilated at 2/3 of itslength; aedeagus shorter than cucullus.

Female genitalia (Fig. 117C). See also Park (1990: Fig. 10); Li and Sattler (2002: Fig. 121). Dorso-posterior margin of segment VIII straight, medially not convex; apophyses anteriores about one-half length of apophyses posteriores; ostium bursae near posterior margin of VIII. Subostial platemore or less trapezoid, caudal margin weakly to distinctly concave; anterior margin slightly convex,plate with less strongly sclerotized oval ‘window’. Antrum shorter than subostial plate, sometimesindistinct. Ductus bursae thin, less than twice length of apophyses posteriores, entering corpusbursae postero-laterally. Corpus bursae pyriform, less than twice length of apophyses posteriors.

SPECIMENS EXAMINED: [GG] 1♀, Cheongryangri, 31.v.1996; 1♀, Suweon, 16.vi.1983; 1♀, Ipori,20.viii.1990; 3♀, 1♂, Mt. Chelma-san, Incheon, 14.vi.1996; [GW] 1♂, Chuncheon, 11.vi.1988; [JB] 1♀,2♂, Mt. Mireuk-san, 23.viii.1997.

DISTRIBUTION: Korea (Central, South), China (Shaanxi, Hebei, Shanxi, Shandong, Henan, Hubei,Guizhou, Jiangsu, Anhui).

HOST PALNTS: Leguminosae- Cajanus cajan, Indigofera, Robinia pseudoacacia, Parkinsonia aculeate (Liand Sattler, 2012).

118. Mesophleps acutunca Li and Sattler, 2012

Mesophleps acutunca Li and Sattler, 2012, Zootaxa, 3373: 32.TL: Zhangjijie, Hunan, China.

ADULT: Wingspan 10-11 mm. This species is similar to the preceeding species, Mesophleps sublu-tiana, but can be distinguished by the smaller size, the forewing with small brownish costal patch,instead of a long brownish streak along costa. Head light yellow. Second segment of labial palpusrelatively thin, dark brown and white in distal part; third segment slightly shorter than secondsegment, white. Antenna with alternating white and dark brown rings. Forewing grayish orange,scattered with brownish scales, with small brownish costal patch medially; costal streak in distal3/5 indistinct, interrupted by oblique white line running from distal quarter to middle of termen;several black scales along termen; blackish discal spot sometimes indistinct and a blackish plicalspot distinct. This species differs from all other members in the male genitalia, with the unusualuncus and gnathos. Female is unknown in Korea.

Male genitalia (Figs. 118A, B). See also Li and Sattler (2012, Fig. 66). Uncus small, bud-shaped.Gnathos with conical. Sclerotized lateral processes, as long as uncus. Valva narrow, dilated inapical part. Tegumen moderate. Vinculum narrow; saccal region broad, about 1/4 length of vincu-lum, medially notched. Aedeagus globular, with short distal portion.

Female genitalia. See Li and Sattler (2012, Fig. 95).

SPECIMENS EXAMINED: [GG]1♂, Mt. Teawha-san, 24.vi 2006.DISTRIBUTION: Korea (Central), China (Hunan, Henan).


Subfamily Dichomeridinae Hampson, 1918

The subfamily Dichomeridinae are a medium-sized micromoths, which are known to occur on allcontinents except Antarctica. The number of known species in the world has not been enumerated,but assumed to be more than 550 species (Kristensen, 1999). Many different classifications of thesubfamilies in Gelechiidae have been proposed by authors: Hodges (1986) included at least sixgenera in this subfamily, including Dichomeris Hübner, Helystogramma Zeller, and Scodes Hodges,and Hodges (1986) synonymized five genera: Gaesa Walker, Acangthophila Heinemann, CarbatinaMeyrick, Uliaria Dumont, and Gomphocrates Meyrick with Dichomeris. Karsholt and Razowski(1996) included Dichomeris, Anasphaltis Meyrick, Brachmia Hübner, Helystogramma, Acompsia Hüb-ner, and Telephila Meyrick. Hodges (1999) included Brachmia, Dichomeris, Helystogramma, and Scodes;and Ponomarenko (1992) included the tribes Dichomeridini, Chelarini and Anarsini in the sub-family Dichomeridinae. Especially, the phylogenetic analysis proposed by Ponomarenko (1992)was with the use of these characters: muscle m4 being separated from the tegument and arisingfrom the parategumental sclerite. In this book, the author followed the classification given byPonomarenko (1992) and Ponomarenko and Park (2007), which was divided into three tibes Dicho-meridini, Chelarini and Anarsini.

The subfamily is defined by following apomorphic characters: 1) pupa with raised, setose mesial/


Fig. 118. Mesophleps acutunca. A. Male genitalia; B. Chinese specimen with aedeagus (after Li andSattler, 2012); C. Abdomen.

A B

C

submesial prominences on terga I, II, III, and IV; 2) corpus bursae with prominent secondary bursae;3) abdominal support system on 2nd sternum a pair of venulae, anterior end of each venula notdeveloped as an apodeme; 4) female with secondary bursae arising from corpus bursae; 5) fore-wing usually with CuA1 and CuA2 stalked or connate and directed posteriorly from end of cell.

Most larvae are leaf-tires, and some are leaf-rollers on various plant families, including Cruciferae,Eriaceae, Fagaceae, Gramineae, Leguminosae, Pinaceae, and Rosaseae.

Tribe Dichomeridini Hampson, 1918

Dichomeridini Hampson, 1918, Novit. zool., 25: 386. Type genus: Dichomeris Hübner, 1818.

Genus Helcystogramma Zeller, 1877

Helcystogramma Zeller, 1877, Horae Soc. ent. ross., 13: 369. Type species: Gelechia obseratella Zeller, 1877: 371. SYNONYMS: Ceratophora Heinemann, 1870; Teuchophanes Meyrick, 1914; Schemataspis Meyrick, 1918;

Parelectra Meyrick, 1925; Psamathoscopa Meyrick, 1937; Anathyrsotis Meyrick, 1939; ParelectroidesClarke, 1952.

Genus Helcystogramma comprises more than 100 described species in the world, and is mostlyknown to occur in the Oriental and Neotropical Regions. Among them, less than 10 species areknown in the Palaearctic Region. The genus is superficially very similar to the genus BrachmiaHübner, but it can be distinguished by the secondary radial retinaculum on the underside of fore-wing and lack a juxta in the male. The genus is characterized by the male genitalia: the sacculussmall, triangular; the valvella finger-like; the aedeagus with rounded inflated basal part and hook-like apex. In the female genitalia, the ostial area has priostial plates; the antrum narrow; the ductusbursae very short, often with small sclerotization; the corpus bursae membranous. The larvae feedon the herbaceous plants from Asteraceae, Convolvulaceae, and Poaceae (Piskunov, 1981; Ponoma-renko, 1999).

Key to the species of the genus Helcystogramma Zeller, based on the male and female genitalia

1. Aedeagus with large, lateral heavily sclerotized process·······························H. fuscomarginatum- Aedeagus without lateral heavily sclerotized process·······························································22. Valva narrowed medially; aedeagus about 1/2 length of cucullus. Female genitalia with ostium

placed medially on sternal part of 8th segment; corpus bursae with numerous thorns················································································································································H. perelegans

- Valva not narrowed medially; aedeagus nearly equal length to cucullus or longer. Female geni-talia with ostium placed near anterior margin of sternal part of 8th segment; corpus bursae witha small area of thorns on lateral side or on bottom····································································3

Lepidoptera: Gelechiidae: Dichomeridinae 31

3. Gnathos almost straight; saccus semioval. Female genitalia with ring-shaped ostial plate······································································································································H. flavilineolella

- Gnathos arched; saccus more or less triangular. Female genitalia with flat ostial plate bearingnarrow slit or consists of two plates··························································································4

4. Sacculus narrowed before triangular, dilated apical part. Female genitalia with ductus bursaenot narrowed beyond antrum···································································································5

- Sacculus smoothly narrowed towards apex. Female genitalia with ductus bursae narrowedbeyond antrum·····························································································H. compositaepictum

5. Cucullus with straight ventral margin; parategminal sclerite with narrow band-like plate; aedea-gus heavily sclerotized ventrally············································································H. ineruditum

- Cucullus hollowed on ventral margin before apex; parategminal sclerite with triangular plate;aedeagus weakly sclerotized ventrally··································································H. triannulella

119. Helcystogramma fuscomarginatum Ueda, 1995

Helcystogramma fuscomarginatum Ueda, 1995, Jpn. J. Ent., 63(2): 385; Ponomarenko, 1997a: 6; Park,2004: 87; Park and Ponomarenko, 2007: 134.TL: Kyushu, Japan.

ADULT: Wingspan 12-14 mm. This species is similar to H. contrubatum (Meyrick) in Africa, but it


Fig. 119. Helcystogramma fuscomarginatum (after Ueda, 1995). A. Male genitalia; B. Aedeagus; C.Female genitalia with accessory bursae.

A B C

can be easily distinguished by the female genitalia with the sclerotized dorsal side of ductus bursae,instead of membrane.

Male genitalia (Figs. 119A, B). See also Ueda (1995, Fig. 9d); Park and Ponomarenko (2007, Pl. 49,Fig. 166). Uncus broadened distally, with round apex. Gnathos heavily sclerotized. Valva elon-gate, slightly dilated towards apex; apex rounded. Saccus large, as long as 1/3 of valva. Aedeagusglobular basally, narrowed towards apex from half, with small spine near apex.

Female genitalia (Fig. 119C). See also Ueda (1995, Fig. 9d); Park and Ponomarenko (2007, Pl. 87, Fig.166). Apophyses anteriores short, less than 1/3 length of apophyses posteriores. Ductus bursaevery short, shorter than corpus bursae.

SPECIMENS EXAMINED: [GG] 1♂, Mt. Cheonggye-san, 23.vii.1996. DISTRIBUTION: Korea (Central), Japan (Honshu, Kyushu).HOST PLANT: Poaceae- Oplismenus undulatifolius (Ueda, 1995).

120. Helcystogramma ineruditum (Meyrick, 1926)

Brachmia inerudita Meyrick, 1926, Exot. Microlep., 3: 290. Brachmia inerudita: Gaede, 1937: 538; Clarke, 1969(6): 365. Helcystogramma ineruditum: Hodges, 1986: 122; Ponomarenko, 1997a: 7; 1999: 207; Park and Pono-


Fig. 120. Helcystogramma ineruditum. A. Male genitalia; B. Aedeagus; C. Female genitalia withcorpus bursae.

B C

A

marenko, 2007: 135.Helcystogramma lutatella: Omelko, 1999: 193 (misid.).

TL: Khabarovsk, Russia.

ADULT: Wingspan 14-15 mm. The forewing ground color is grayish-brown with three black dotson the cell, six dots along apex and outer margin, and light curved fascia at about 2/3 of length.

Male genitalia (Figs. 120A, B). See also Clarke (1969(6), 365, pl. 181, Figs. 4a, b); Omelko (1999, Fig.104, 3); Ponomarenko (1999, Fig. 116-8). Gnathos arched; cucullus not narrowed at middle, withstraight ventral margin; parategminal sclerite with a narrow band-like plate. Aedeagus almost equalto cucullus in length, with heavily sclerotized ventral side.

Female genitalia (Fig. 120C). See also Park and Ponomarenko (2007, Fig. 50). Ostial plate is extend-ed anteriorly with narrow split.

SPECIMENS EXAMINED: [CN] 1♀, Mt. Daedun-san, Nonsan, 13.vii.2000.DISTRIBUTION: Korea (Central), Russian Far East (Khabarovsk, Primorsk), Transbaikalia.

121. Helcystogramma perelegans (N. Omelko and M. Omelko, 1993)

Tricyanaula perelegans N. Omelko and M. Omelko, 1993, Biol. issled. estest. kult. ecosyst. Primorsk.kraya: 218.


Fig. 121. Helcystogramma perelegans. A. Male genitalia; B. Aedeagus; C. Female genitalia withcorpus bursae (after Omelko, 1993).

A B C

Tricyanaula perelegans: Omelko, 1999: 194. Helcystogramma perelegans: Ponomarenko, 1997a: 8; Ponomarenko, 1999: 207; Park, 2004: 86; Park

and Ponomarenko, 2007: 135.TL: Primorye, Russia.

ADULT: Wingspan 10-11 mm. The forewing has two white costal streaks on the middle, two bluetransverse fascias and two obliqua blue lines at basal half. In male genitalia, the cucullus narrowedat middle; and the aedeagus two times as short as cucullus. In female genitalia, the ostium placedon the middle of sternite VIII; and the corpus bursae with numerous thorns.

Male genitalia (Figs. 121A, B). See also N. Omelko and M. Omelko (1993, Fig. 4; 1999: Fig. 105, 3);Ponomarenko (1999, Figs. 115, 4-7); Park and Ponomarenko (2007: Pl. 50, Fig. 168).

Female genitalia. See N. Omelko and M. Omelko, 1993: Figs. 7, 8; Omelko, 1999: Figs. 106, 1, 2).

SPECIMENS EXAMINED: [GG] 1♀, Mt. Cheonggye-san, 19.vii.1976; 1♀, Gwangleung, 3.vi.1982; 3♂,Mt. Soyo-san, 9.vi.1996; [GW] 1♂, 2♂, Chuncheon, 12.viii.1988, 3.vi.1997; 1♂, Mt. Gubong-san,8.vi.1988; 1♂, Yangyang, 4.vi.1987; 1♂, Goseong, 12.vi.1987.

DISTRIBUTION: Korea (Central), Russian Far East (Primorsk), Japan.

122. Helcystogramma triannulella (Herrich-Schäffer, 1854)

Anacampsis triannulella (Herrich-Schäffer), 1854, Schmett. Eur., 5: 201, Fig. 458. Gelechia sepiella Streudel, 1866, Ent. Ztg. Stett., 27: 312.Brachmia macroscopa Meyrick, 1932, Exot. Microlep., 4: 206; 1935: 75.Brachmia triannulella: Rebel, 1901: 157; Gaede, 1937: 545; Piskunov, 1981: 735; Park, 1991a: 121. Brachmia macroscopa: Clarke, 1969: 366; Liu et al., 1981: 18.Brachmia triannulella macroscopa: Moriuti, 1982, I: 287, II: 215.Helcystogramma triannulella: Park and Hodges, 1995b: 230; Karsholt and Riedl, 1996: 121; Ponoma-

renko, 1997: 9; Elsner et al., 1999: 56; Omelko, 1999: 193; Ponomarenko, 1999: 209; Park, 2004: 86;Park and Ponomarenko, 2007: 135.

Helcystogramma triannulella macroscopum: Ueda, 1995: 380; Ueda et al., 1995: 150.TL: Swiss, Europe.

ADULT: Wingspan 16-20 mm. The forewing ground color is brown with three black dots circledby pale grey in the cell, eight dots along apex and outer margin, and with light arched fascia at 2/3of length.

Male genitalia (Figs. 122A, B). See also Clarke (1969(6), pl. 182, Figs. 2a, b); Liu et al. (1981, Fig. 67);Piskunov (1981, Fig. 665, 3); Moriuti (1982, II: pl. 259, Figs. 1, 1a); Park and Hodges (1995b: Figs. 24,25); Ueda (1995, Figs. 7a, b); Elsner et al. (1999, t. 40, Fig. 346); Ponomarenko (1999, Figs. 115, 10, 11).Similar to those of H. arotraeum (Meyrick), but differs from the latter by the shorter uncus, valvemuch more exceeding apex of uncus, and saccal region shorter. Valva hollowed on ventral marginbefore apex; the parategminal sclerite with triangular plate, and the aedeagus with evenly sclero-tized ventral side.

Female genitalia (Fig. 122C). See also Park and Hodges (1995b: Fig. 27); Ueda (1995: Fig. 7d); Elsneret al. (1999: t. 82, Fig. 346); Ponomarenko (1999: Fig. 117, 5). Ostial plate consists of two plates.


Ductus bursae dilated towards the corpus bursae. Corpus bursae elongate, about six times as longas ductus bursae; signum. small, crescent, with serrated edges.

SPECIMENS EXAMINED: [N. Korea] 1♂, Sariwon, N. Hwanghae Prov., 28.ix.1977; [GG] 5♀, 10♂,Mulwang Res., Siheung, 24.vii.1996, 30.viii.1996; [GG] 1♀, Isl. Yongyudo, Incheon, 17.ix.1997; 1♂,Isl. Daebudo, Ansan, 30.viii.1997; [GW] 1♂, Mt. Gyebang-san, Pyoungchang, 2.vii.1997; 1♂, Mt.Deokga-san, Wonju, 24.vii.1997; 1♂, Mt. Gyebang-san, 7.viii.1997; 1♀, 1♂, Mt. Daedeok-san, 8.viii.1997; 1♀, Mt.Gwangdeok-san, Hwacheon, 19.viii.1997; [CB] 1♂, Taean, 12.vii.1996; [CN] 1♂, Mt.Gaya-san, Yesan, 24.viii.1997; [JB] 2♀, 1♂, Mt. Mireuk-san, Iksan, 23.viii.1997; [GN] 3♀, SwampUpo, Changnyeong, 28.vii.1997; 2♂, Geochang, 23.viii.1997.

DISTRIBUTION: Korea (North, Central, South), Russia (European part, Caucasus, SW Siberia, Pri-morsk), West Kazakhstan, Central Asia, Europe, N India, China (Tianjin, Shaanxi, Henan, Sichuan,Shandong, Jiangsu, Jiangxi), Taiwan, Japan (Hokkaido, Honshu, Shikoku, Kyushu, Izu, Ryukyu Isl.).

HOST PALNTS: Convolvulaceae- Ipomoea batatas, I. aquatica, Hibiscus syriacus, Convolvulus arvensis,Calystegia sepium, C. japonica, Pharbitis nil, Calongction aculeatum, and Calystegia sepium (Elsner et al.,1999).


Fig. 122. Helcystogramma triannulella (after Ponomarenko, 1999). A. Male genitalia with aedeagus;B. Ditto, vincullum; C. Female genitalia with corpus bursae.

A C

B

Genus Dichomeris Hübner, 1818

Dichomeris Hübner, 1818, Zutr. Samml. exot. Schmett., 1: 25. Type species: Dichomeris ligulella Hübner, 1818: 25. SYNONYMS: Oxybelia Hübner [1825]; Rhinosia Treitschke, 1833; Anorthosia Clemens, 1860; Tricho-

taphe Clemens, 1860; Rhobonda Walker, 1864; Vazugada Walker, 1864; Gaesa Walker, 1864; Tocmia Wal-ker, 1864; Carna Walker, 1864; Noeza Walker, 1866; Sagaritis Chambers, 1872; Begoe Chambers, 1872;Epicorthylis Zeller, 1873; Malacotricha Zeller, 1873; Atasthalistis Meyrick, 1886; Zalithia Meyrick, 1894;Pappophorus Walsingham, 1897; Arotria Meyrick, 1904; Croesopola Meyrick, 1904; Hypelictis Meyrick,1905; Paristhmia Meyrick, 1909; Hylograptis Meyrick, 1910; Schematistis Meyrick, 1911; Paranoea Wal-singham, 1911; Plocamosaris Meyrick, 1912; Machlotricha Meyrick, 1912; Holaxyra Meyrick, 1913;Carbatina Meyrick, 1913; Zomeutis Meyrick, 1913; Ilingiotis Meyrick, 1914; Pachysaris Meyrick, 1914;Deimnestra Meyrick, 1918; Daemonarcha Meyrick, 1918; Macrozancla Turner, 1919; Euryzancla Turner,1919; Rhadinophylla Turner, 1919; Eurysara Turner, 1919; Iochares Meyrick, 1921; Uliaria Dumont, 1920;Eporgastis Meyrick, 1921; Aphnogenes Meyrick, 1921; Taphrosaris Meyrick, 1922; Prophoraula Meyrick,1922; Acribologa Meyrick, 1923; Ageliarchis Meyrick, 1923; Myrophila Meyrick, 1923; Brochometis Mey-rick, 1923; Neochrista Meyrick, 1923; Semiomeris Meyrick, 1923; Mythographa Meyrick, 1923; Cymotri-cha Meyrick, 1923; Sathrogenes Meyrick, 1923; Cotyloscia Meyrick, 1923; Sirogenes Meyrick, 1923;Musurga Meyrick, 1923; Thelyasceta Meyrick, 1923; Oxysactis Meyrick, 1923; Rhynchotona Meyrick,1923; Gomphocrates Meyrick, 1925; Catoptristis Meyrick, 1925; Cymatoplex Meyrick, 1925; CatelaphrisMeyrick, 1925; Cerycangela Meyrick, 1925; Hyperecta Meyrick, 1925; Epicharta Meyrick, 1926; Orso-dytis Meyrick, 1926; Prasodryas Meyrick, 1926; Xenorrhythma Meyrick, 1926; Thyrsomnestis Meyrick,1929; Desmophylax Meyrick, 1935; Ereboscaeas Meyrick, 1937; Chthonogenes Meyrick, 1938; Cymatopli-cella Fletcher, 1940; Neopachnistis Janse, 1954; Leuropalpa Janse, 1960; Picroptera Janse, 1960; Parabra-chmia Janse, 1960; Deltolophos Janse, 1960; Virgula Omelko, 1999; Resupina Omelko, 1999.

Genus Dichomeris comprises more than 550 species in the word. It is known as a New Worldgenus, with abundance in the tropical region. It was introduced to the Old World (Hodges, 1986)from the New World. The male genitalia have more or less triangular parategminal sclerites; thecucullus weakly sclerotized and shifted dorsally; the sacculus fused with arms of vinculum; theaedeagus, the sica, and the vinculum are often ankylozed into one sclerite. The female genitaliahave a wide antrum; the ductus bursae very short and broad; ductus seminalis usually with sclero-tized ring neae base; the corpus bursae with well-developed accessory bursae. Host plants of thespecies belong to different botanic families, the larvae of representatives of Asian fauna feed onplants from Fabaceae, Rosaceae, Fagaceae, Boraginaceae, Juglandaceae, Corylaceae, Asteraceae, Aceraceae,Betulaceae (Saito, 1969; Ponomarenko, 1993; Park and Hodges, 1995; Li and Zheng, 1996; Elsner etal., 1999; Li, 2002; Park and Ponomarenko, 2007).

Key to the species of the genus Dichomeris Hübner, based on the male genitalia(exception of D. tostella Stringer)

1. Saccus present·······································································································D. japonicella- Saccus absent··························································································································22. Vinculum band-like, not divided····························································································3- Vinculum divided into two arms···························································································12


3. Juxta long and narrow, extended caudally, without processes·················································4- Juxta with basal plate and distal processes··············································································54. Sica digitate·················································································································D. cuspis- Sica extended as plate-like distally············································································D. minutia5. Sicae more or less symmetric, with two straight processes······················································6- Sicae asymmetric, with curved processes················································································86. Processes of sicae digitate, shorter or equal to basal plate in length·········································7- Processes of sicae bifurcate apically respectively··················································D. heriguronis7. Processes of sicae symmetric, long, divided beyond 3/5 length································D. lutilinea- Processes of sicae nearly symmetric, right one is slightly shorter; basal part shorter than pro-

cesses························································································································D. rasilella8. Sacculus absent·······················································································································9- Sacculus present, sclerotized·································································································109. Sacculus absent; processes of juxta curved to right··············································D. atomogypsa- Sacculus membranous; processes of juxta curved to left···············································D. issikii

10. Sacculus symmetric·····························································································D. chinganella- Sacculus asymmetric·············································································································11

11. Right sacculus nearly five times as long as left one, wedge-shaped, curved medially; juxta andanellus ankylosed into one sclerite with seven processes········································D. bulawskii

- Right sacculus two times as long as left one, truncated at the apex, curved medially, and withsmall thorns; juxta and anellus ankylosed into one sclerite with five processes······D. christophi

12. Processes of sica equal in length····························································································13- Processes of sica differ in length····························································································19

13. Processes of sica narrowed towards apex··············································································14- Processes of sica with projections before apex or dilated distally···········································18

14. Anellus with processes··········································································································15- Anellus without proces·········································································································16

15. Anellus with two processes, every of them longer than half of total length of aedeagus, aedea-gus with four needle-like cornuti···············································································D. litoxyla

- Anellus with one process, it shorter than half of total length of aedeagus, aedeagus withoutneedle-like cornuti·································································································D. autometra

16. Processes of sica nearly same width along their length···············································D. mitteri- Processes of sica more or less dilated basally·········································································17

17. Aedeagus with thick cornutus at about 1/3 of its length·······································D. horoglypta- Aedeagus with needle-like cornutus, longer than half of its length·······················D. consertella

18. Anellus with two asymmetric lateral processes: beak-like left one and larger triangular rightone; aedeagus with one cornutus···············································································D. oceanis

- Anellus with two symmetric subulate lateral processes; aedeagus without cornutus···············································································································································D. lespedezae

19. Right process of sica longer than left one···············································································20- Right process of sica shorter than left one··············································································21

20. Processes of sica bifurcate on the apex, left one slightly shorter than right one; dorsal processesof anellus longer than cornutus of aedeagus······················································D. polypunctata

- Processes of sica not bifurcate at apex, left one about two times as short as right one; dorsalprocesses of anellus shorter than cornutus of aedeagus···········································D. sparsella


21. Processes of sica serrate on ventral margin, dilated at middle, with pointed apex, curved oneto other····················································································································D. derasella

- Processes of sica not serrate on ventral margin, not dilated at the middle, with beak-like apex··············································································································································22

22. Valva dilated toward base, deeply hollowed before apex·······································D. praevacua- Valva narrowed toward base, without hollow before apex····················································23

23. Arms of vinculum narrow, band-like; sacculus finger-like········································D. ustalella- Arms of vinculum dilated ventro-laterally; sacculus not finger-like·······················D. harmonias

123. Dichomeris atomogypsa (Meyrick, 1932)

Gaesa atomogypsa Meyrick, 1932, Exot. Microlep., 4: 202. Gaesa atomogypsa: Gaede, 1937: 447; Issiki, 1957: 41; Clarke, 1969(7): 103; Saito, 1969: 113; Moriuti,

1982, I: 285, II: 215.Dichomeris atomogypsa: Hodges, 1986: 72; Park and Hodges, 1995a: 41; Park and Ponomarenko,

1996: 346; Park, 2004: 93; Park and Ponomarenko, 2007: 141.TL: Hasimoto, Kii, Japan.


Fig. 123. Dichomeris atomogypsa. A. Male genitalia; B. Aedeagus; C. Female genitalia with corpusbursae.

A B C

ADULT: Wingspan 23-27 mm. This species is superficially similar to D. chinganella Christoph, butcan be distinguished by the male genitalia: distal processes of juxta divided into two arms whichcurved to right. It is also close to the North American species D. ventrella (Fitch) in the externalcharacter.

Male genitalia (Figs. 123A, B). See also Park and Hodges (1995a, Figs. 40, 41); Park and Ponoma-renko (2007, Pl. 50, Fig. 171). Uncus relatively short, with a pair of long setae dorsally and shortones densely spaced on ventral surface. Culticula strongly convex medially. Valva dilated distally;valvella very long, about 2/5 length of valva. Sicae asymmetrical, fused at basal 1/3; left lobelonger than right one, bent preapically; apex acute. A secondary sclerotized lobe complex stronglyankylosed with sicae and aedeagus. Aedeagus nearly equal to the length of genitalia; cornutusheavily sclerotized, about 1/5 length of aedeagus.

Female genitalia (Fig. 123C). Clarke (1969, pl. 51, Figs. 1a, b); Park (1994, Fig. 7c); Park and Hodges(1995a, Fig. 86); Park and Ponomarenko (2007, Pl. 88, Fig. 171). Antrum extremely broad. Ductusbursae with complex of several sclerotized plates and ridges; ductus seminalis arsing from dorsalsurface of antrum. Corpus bursae long; accessory bursae arising from ventral surface; signumabsent.

SPECIMENS EXAMINED: [GW] 2♀, 2♂, Chuncheon, 13.vii.1980, 25.iii.1991, 26.iv.1989, 22.vii.1991. DISTRIBUTION: Korea (Central), Japan (Honshu, Shikoku).HOST PALNTS: Fagaceae- Quercus acutissima, Q. dentata, Q. serrata (Saito, 1969).

124. Dichomeris autometra (Meyrick, 1934)

Cymotricha autometra Meyrick, in Caradja and Meyrick, 1934, Dt. ent. Z., Iris, 48: 34. Cymotricha autometra: Gaede, 1937: 455; Clarke, 1969(7): 522.Dichomeris autometra: Park and Hodges, 1995a: 53; Li and Zheng, 1996: 256; Park, 2004: 99; Park and

Ponomarenko, 2007: 141.TL: Kwanhsien, China.

ADULT: Wingspan 12.5-13 mm. Antenna with sensory scales ventrally in male. Second segmentof labial palpus thickened, with erect scale-tuft dorsally. Forewing with R4 and R5 stalked; CuA1

and CuA2 short-stalked.Male genitalia (Figs. 124A, B). See also Clarke (1969(7), pl. 260, Figs. 1a, b); Park and Hodges (1995a,

Figs. 64, 65); Park and Ponomarenko (2007, Pl. 51, Fig. 172). Valva exceeding apex of uncus; valvellashort, stout. Vinculum divided into two arms; anellus with one process which shorter than half oftotal length of aedeagus; the processes of sica equal in length, smoothly narrowed towards apex.Aedeagus with heavily sclerotized slender lateral lobe; internal lobe weakly developed, short; cor-nutus absent.

Female genitalia (Fig. 124C). See also Park and Hodges (1995a, Fig. 85); Park and Ponomarenko(2007, Pl. 88, Fig. 172). Ductus bursae with irregular shaped ridges on dorsolateral wall; ductusseminalis arsing from distal part of ductus bursae. Corpus bursae large, ovate, membranous; sig-num absent.

SPECIMENS EXAMINED: [GW] 1♀, Pyoungchang, 6.vi.1996; [JB] 1♂, Muju, 13.viii.1975.


DISTRIBUTION: Korea (Central, South), China (Sichuan), Taiwan.HOST PLANT: Boraginaceae- Lithospermum spp. (Park and Hodges, 1995).

125. Dichomeris chinganella (Christoph, 1882)

Nothris chinganella Christoph, 1882, Bull. Soc. Nat. Mosc., 57(1): 32; Meyrick, 1925: 98; Gaede, 1937:295.

Dichomeris chinganella: Park, 1996: 65; Ponomarenko, 1997: 17; Park, 2004: 94; Park and Ponomarenko,2007: 142.

Dichomeris fareasta Park, 1994: 15.Dichomeris fareasta: Ponomarenko, 1999, 219, 229; Li and Wang, 2000: 48 (misid.).

TL: Raddevka (Radde), Russian Far East.

ADULT: Wingspan 19-22 mm. This species is superficially similar to D. atomogypsa (Meyrick), butcan be distinguished by the male genitalia, especially the processes of juxta asymmetric: the leftone slender, curved; the right one bifurcated.

Male genitalia (Figs. 125A, B). See also Park (1994, Figs. 7a, b); Ponomarenko (1999, Figs. 124-1, 2);Li and Wang (2000: 48); Park and Ponomarenko (2007, Pl. 51, Fig. 173). Uncus with rounded distalmargin. Valva slightly dilated near middle, greatly exceeding apex of uncus; valvella large, distal


Fig. 124. Dichomeris autometra (after Park and Hodges, 1995). A. Male genitalia; B. Aedeagus; C.Female genitalia.

A B C

half digitate, basal half broad. Sicae asymmetrical, left lobe strongly curved ventrally, bifurcateapically, ventral arms longer with two long setae near apex; right one simple, digitate, with severallong setae on inner margin; a sclerotized lobe complex tightly fused with sicae at base. Aedeagusabout 3/4 length of genitalia, basal part narrowed, with heavily sclerotized pointed lobe on rightside; cornutus absent.

Female genitalia (Fig. 125C). See also Ponomarenko (1999, Fig. 132-2); Park and Ponomarenko (2007,Pl. 88, Fig. 173). Ductus bursae long, not well distinguished from corpus bursae, with variablesclerotized bands at posterior part ventrally; ductus seminalis arising from dorsal surface of ductusbursae posteriorly, with densely spiculous appendix bursae. Corpus bursae without signum; acces-sory bursae arising from near middle ventrally.

SPECIMENS EXAMINED: [GG] 1♂, Mt. Yaksu-san, 9.viii.1989; 1♂, Mt. Myoungji-san, 14.iv.1992; 1♂,Mt. Cheonggye-san, Gwacheon, 23.vii.1996; 3♀, 1♂, Mt. Hwaya-san, 18.vii.1997; 1♂, Mt Soyo-san,5.viii.1996; [GW] 17♀, 19♂, Chuncheon, 10.iv.1985, 7.vii.1987, 7.vii.1987, 9.ix.1988, 26.iv.1989, 26.iv.1989, 1.v.1989, 13.vii.1989, 21.iii.1990,10.iv.1990, 6.v.1990, 7.v.1990, 24.iv.1991, 22.vii.1991; 5♀, Mt.Samak-san, 8.v.1989, 19.vii.1989, 25.ix.1989; 1♂, Mt. Odae-san, 22.v.1989.

DISTRIBUTION: Korea (Central), Russian Far East (Primorsk), China (Henan).HOST PLANT: Fagaceae- Quercus mongolica.


Fig. 125. Dichomeris chinganella. A. Male genitalia; B. Aedeagus; C. Female genitalia.

A B C

126. Dichomeris christophi Ponomarenko and Mey, 2002

Dichomeris christophi Ponomarenko and Mey, 2002, Tinea, 17(2): 78. Gaesa sparsella(us): Issiki, 1957: 41; Saito, 1969: 113; Liu et al., 1981: 19; Moriuti, 1982, I: 285; II: 215;

Park, 1987: 177, 178 (misid.)Dichomeris sparsella(us): Hodges, 1986: 72; Park, 1994: 14; Park and Hodges, 1995: 40; Ueda et al.,

1995: 150; Li and Zheng, 1996: 236; Ponomarenko, 1997: 31; 1999: 221, 230; misidentification.Dichomeris christophi: Park and Ponomarenko, 2007: 143.

TL: Gornotajezhnoe, Ussuri, Russian Far East.

ADULT: Wingspan 21-23 mm. This species is superficially similar to D. bulawskii Ponomarenkoand Park, but it can be easily distinguished by the male and female genitalia. Forewing almoststraight costa, yellowish white along anterior margin; ground color light brown, with pale orangetransverse median line near end of cell; CuA1 and CuA2 relatively long stalk.

Male genitalia (Figs. 126A, B). See also Park (1987, Figs. 8, 9); Park and Hodges (1995, Figs. 38, 39);Ponomarenko (1999, Figs. 125, 1, 2); Ponomarenko and Mey (2002, Figs. 9, 10); Park and Ponoma-renko (2007, Pl. 51, Fig. 175). Uncus relatively small, short. Valva rather broad, exceeding apex ofuncus; valvella digitate, as long as gnathos. Sicae asymmetrical, left lobe longer, sickle-shaped;right one bent outwardly. Aedeagus with narrowed basal zone, surrounded by five processes ofankylosed sicae and anellus; dorsal lobe slender, sharply pointed; left one horn-shaped; right onevery short, spatulate; cornutus absent.


Fig. 126. Dichomeris christophi. A. Male genitalia; B. Aedeagus; C. Female genitalia with corpusbursae.

A B C

Female genitalia (Fig. 126C). See also Park and Hodges (1995, Fig. 84); Ponomarenko, 1999, Fig.134, 2); Ponomarenko and Mey (2002, Figs. 11, 12); Park and Ponomarenko (2007, Pl. 88, Fig. 175).Antrum heavily sclerotized, with several complicated longitudinal ridges, fusing with ductus bur-sae. Ductus bursae with large, sac-like sclerite on left side and plicated one on right side; ductusseminalis arising from middle of ductus bursae dorsally. Corpus bursae ovate; accessory bursaearising from near anteroventral end of corpus bursae.

SPECIMENS EXAMINED: [GW] 1♀, 2♂, Chuncheon, 12.vii.1989, 7.x.1986, 1.xi.1986; 1♂, Mt. Samak-san, 8.v.1989, 19.vii.1989.

DISTRIBUTION: Korea (Central), Russian Far East (Primorsk), China (Heilongjiang), Japan (Honshu,Kyushu).

HOST PLANT: Juglandaceae- Pterocarya rhoifolia, Juglans ailanthifolia, J. mandshurica, and J. regia (Li,2002).

REMARKS: This species has been misidentified as D. sparcella(us) until Ponomarenko and Mey(2002) described this species.

127. Dichomeris cuspis Park, 1994

Dichomeris cuspis Park, 1994, Ins. Koreana, 11: 19. Dichomeris cuspis: Li and Zheng, 1996: 236; Ponomarenko, 1997: 18; 1999: 216, 229; Li, 2002: 339; Park,

2004: 96; Park and Ponomarenko, 2007: 144.


Fig. 127. Dichomeris cuspis. A. Male genitalia; B. Female genitalia with corpus bursae.

A B

Virgula cuspis: Omelko, 1999: 106.TL: Hongcheon, Korea.

ADULT: Wingspan 14.5-16 mm. The species is superficially similar to the Indian species, D. enop-trias Meyrick or Japanese species, D. hoplocrates Meyrick, but it has less metalic color in the fore-wing. Head and thorax dark fuscous. Forewing elongate;ground color uniformly dark fuscous,shiny and blue-leaden metallic, with dash-like yellowish-white strigula at 3/4 on anterior margin.Hindwing broad, dark gray.

Male genitalia (Fig. 127A). See also Park (1994, Fig. 9a); Li and Zheng (1996, Figs. 27, 28); Omelko(1999, Fig. 52, 5); Ponomarenko (1999, Figs. 122, 1, 2); Li (2002, Fig. 389); Park and Ponomarenko(2007, Pl. 52, Fig. 177). Uncus short, broad, rather triangular with bround distal margin. Valvaapproximately attaining apex of uncus, basal half nearly parallel sided, dilated beyond middle.Valvella as long as half of cucullus; sacculus long, slender, rounded at apex; vinculum band-like,not divided into two arms. Sica single, small, slender, with clavate apex.

Female genitalia (Fig. 127B). See also Park (1994, Fig. 9b); Li and Zheng (1996, Fig. 29); Ponomarenko(1999, Fig. 133, 1); Li (2002, Fig. 390); Park and Ponomarenko (2007, Pl. 89, Fig. 177). Antrum withheavily sclerotized, irregular-shaped dentes distallyl. Ductus bursae broad, short, with complicatedsclerites. Corpus bursae large.

SPECIMENS EXAMINED: [GG] 1♂, Gwangleung, 27.vi.1986, paratype; 3♂, Mt. Soyo-san, 7.vii.1996;[GW] 1♀, Chuncheon, 12.vi.1998; 1♂, Mt. Gubong-san, 19.vi.1998; 1♂, Jiamri, Chuncheon, 19.vii.1995; 1♀, Hongcheon, 14.viii.1987, holotype; 1♂, Mt. Deokga-san, Wonju, 24.vii.1997.

DISTRIBUTION: Korea (Central), Russian Far East (Primorsk), China (Shaanxi).HOST PLANT: Fagaceae- Quercus acuteserrata (Li and Zheng, 1996).

128. Dichomeris derasella ([Denis and Schiffermüller], 1775)

Tinea derasella [Denis and Schiffermüller], 1775, Ank. syst. Werk. Schmett.: 140. Dichomeris derasella: Koçak, 1984: 149; Karsholt and Riedl, 1996: 121; Ponomarenko, 1997: 19; Elsner

et al., 1999: 55; Ponomarenko, 1999: 225, 227; Li, 2002: 343; Park, 2004: 91; Ponomarenko, 2007:145.

Tinea fasciella Hübner, 1796, Eur. Schmett., Tineen, pl. 16, Fig. 111.Dichomeris fasciella: Piskunov, 1981: 731; Park, 1994: 9; Li and Zheng, 1996: 241 (misid.).Dichomeris coreanus Matsumura, 1931, 6000 Ill. Ins. Japan: 1082. Dichomeris paranthes Meyrick, 1936, Exot. Microlep., 5: 47. Ypsolophus unguiculatus Fabricius, 1798: 247.

TL: Austria, Europe.

ADULT: Wingspan 18-25 mm. This species had been misidentified as D. fasciella Hübner in Korea.Head and thorax whitish ochreous. Forewing elongate, slightly dilated, costa faintly sinuate;ground color light brownish ochreous, slightly strigulated gray; costal area suffused gray costalstreak from base to near 3/4 length; some obscure gray suffusion in discal cell, and an obtuselyangulated fascia of gray suffusion at 3/4. Hindwing gray.

Male genitalia (Figs. 128A, B). See also Piskunov (1981, Fig. 662, 5); Park (1994, Figs. 4a, b); Elsner


et al. (1999, t. 38, Fig. 333); Ponomarenko (1999, Figs. 128, 5, 6); Li (2002: Fig. 393); Park and Pono-marenko (2007, Pl. 52, Fig. 178). Uncus relatively short. Gnathos stout. Valva exceeding apex ofuncus. Vinculum divided into two arms; processes of sica differ in size, right process shorter thanleft one, serrate partly on ventral margin, dilated at middle, narrowed towards pointed apex, slight-ly curved inwardly.

Female genitalia (Fig. 128C). See also. Park (1994, Fig. 4c); Elsner et al. (1999, t. 81, Fig. 333); Pono-marenko (1999, Fig. 131); Li (2002, Fig. 394). Park and Ponomarenko (2007, Pl. 89, Fig. 178). Antrumbroad, short. Ductus bursae with two sclerotized plates distally. Corpus bursae with complex ofsclerotized ridge; accessory bursae arising from wall of signum.

SPECIMENS EXAMINED: [N. Korea] 1♂, Mt. Keumgang-san, 28.vi.1985; [GG] 2♂, Suweon, 15.viii.1986, 12.iv.1985; 1♀, Yongin, 21.v.1989; 1♂, Incheon, Isl. Yeongjongdo, 13.viii.1996; 1♂, Mt. Hwaya-san, Gapyeong, 23.v.1997; 1♀, Mt. Soyo-san, 7.vii.1996; [GW] 1♂, Mt. Jeombong-san, 5.viii.1997; 3♀,8♂, Mt. Taebek-san, 27.vi.1996; 2♂, Chuncheon, 3.vi.1997, 1.viii.1986; 1♂, Sogeumgang, 8.viii.1988;1♀, Mt. Seolak-san, 16.vi.1993; [JN] 1♀, Mt. Baekun-san, 19.viii.1992.

DISTRIBUTION: Korea (North, Central, South), China (Shaanxi, Henan, Shandong, Zhejiang, Qing-hai, Ningxia), Russia (European part, Caucasus, S. Ural, S. Krasnoyarsk, Altai, Transbaikalia, Pri-morsk), Transcaucasia, Mediterranean region, Asia Minor, Europe.

HOST PALNTS: Rosaceae- Crataegus sp., Malus sp., Cerasus sp., Rubus sp., Malus sylvestris, Prunusspinosa, Rubus caesius, and R. fruticosus (Elsner et al., 1999; Li, 2002; Park and Ponomarenko, 2007).


Fig. 128. Dichomeris derasella. A. Male genitalia; B. Aedeagus; C. Female genitalia with appendixbursae.

A B

C

REMARKS: Matsumura (1931) described D. coreana, based on the Korean material (Korea, Seoul,19.viii.1918, coll. E. Gallois) and the type is deposited in the Institute of Entomology, HokkaidoUniversity, Japan.

129. Dichomeris harmonias Meyrick, 1922

Dichomeris harmonias Meyrick, 1922, Exot. Microlep., 2: 504. Dichomeris harmonias: Meyrick, 1925: 176; Gaede, 1937: 270; Park, 1994: 6; Park and Hodges, 1995a:

12; Li and Zheng, 1996: 232; Ponomarenko, 1997: 21; 1999: 225, 230; Li, 2002: 415; Park, 2004: 88;Park and Ponomarenko, 2007: 145.TL: Shanghai, China.

ADULT: Wingspan 13-15 mm. The species is superficially similar to D. lespedezae Park, but theycan be easily separable each other by the male genitalia.

Male genitalia (Figs. 129A, B). See also Park (1994, Figs. 2a, b); Ponomarenko (1999, Figs. 127: 3, 4);Park and Ponomarenko (2007, Pl. 52, Fig. 178). Uncus short, broad. Gnathos strongly bent; culcitulaconic, densely spiculous. Valva slightly exceeding apex of uncus. Vinculum divided into two arms,which dilated in ventral and lateral parts; sacculus arched, curved dorsally, with serrate on outermargin; processes of sica very long, almost equal to cucullus in length, right process slightly shorterthan left one, both with beak-like apices. Aedeagus large, with triangular protrusion beyond zone;


Fig. 129. Dichomeris harmonias. A. Male genitalia; B. Aedeagus; C. Female genitalia.

A B C

basal zone narrowed; lateral lobe weak, free, about 3/4 length of aedeagus. Female genitalia (Fig. 129C). See also Park (1994, Fig. 2c); Ponomarenko (1999, Fig. 133, 4). Antrum

broad, almost straight on distal margin. Ductus bursae heavily wrineckled in narrowed posteriorpart ventrally, then broadened, with two large, triangular sclerites posteriorly; ductus seminalisarising from anterior part. Corpus bursae ovate, rather small; accessory bursae arising from medi-ally.

SPECIMENS EXAMINED: [N. Korea] 2♂, Mt. Keumgan-san, 4.viii.1975, 23.vii.1982; [GG] 4♂, Mt.Cheonggye-san, 19.vii.1976; 1♀, 2♂, Mt. Cheonggye-san, 25.vii.1996; 1♂, Suweon, 7.ix.1982; 1♀,Gwangleung, 3.vi.1988; [GW] 1♀, Chuncheon, 4.vi.1998; 1♀, Yangyang, 25.vii.1987; [CB] 1♀, 2♂,Mt. Weolak-san, 20.vi.1984, 9.viii.1997.

DISTRIBUTION: Korea (North, Central), Russian Far East (Primorsk), China (Beijing, Shanghai,Shandong), Japan, Taiwan.

130. Dichomeris heriguronis (Matsumura, 1931)

Nothris heriguronis Matsumura, 1931, 6000 Ill. Ins. Japan: 1084. Nothris heriguronis: Moriuti, 1982, I: 286, II: 215 (as synonym of D. oceanis Meyrick).Trichotaphe iothalles Forbes, 1939, Jour. New York Ent. Soc., 47: 159. Carbatina picrocarpa: Issiki, 1957: 41; Saito, 1969: 112; Moriuti, 1982, 1: 286, 2: 215; Park, 1983: 502

(misid.).Dichomeris picrocarpa: Hodges, 1986: 119; Park, 1991: 123; 1994: 20; Park and Hodges, 1995: 46; Ueda

et al., 1995: 150; Li and Zheng, 1996: 249; Ponomarenko, 1997: 29; 1999: 218, 230; Li, 2002: 375(misid.).

Dichomeris heriguronis: Ponomarenko, 2004: 22; Park and Ponomarenko, 2007: 146.TL: Japan.

ADULT: Wingspan 15-17 mm. The species is superficially similar to the Indian species (D. picro-carpa Meyrick) and Korean species has been misidentified as the latter, but it was reconfirmed thatthe Japanese species differ from the latter by Ponomarenko (2004).

Male genitalia (Figs. 130A, B) See also Moriuti (1982, II: pl. 259, Figs. 3, 3a); Hodges (1986, Figs. 27:a, b); Ponomarenko (1999, Figs. 123: 1, 2; 2004: Fig. 10); Li (2002, Fig. 425); Park and Ponomarenko(2007, Pl. 52, Fig. 180). Valva dilated distally; sacculus long, serrate on ventral margin, curved dor-sally; vinculum band-like, not divided; sica bifurcate, processes about 1/2 length of the basal plate,bifid apically. Aedeagus stout, with short lateral processes and well-developed lobes from zone;cornutus absent.

Female genitalia (Fig. 130C) See also Hodges (1986: Fig. 27: c); Ponomarenko (1999: Fig. 134, 3; Li,2002: Fig. 426). Park and Ponomarenko (2007, Pl. 89, Fig. 180). Antrum broad; basal part of corpusbursae heavily sclerotized, lacking sclerotized ridges, with many inwardly directed spinules onwall near origin of accessory bursae.

SPECIMENS EXAMINED: [N. Korea] 1♂, Mt. Myohyang-san, N. Pyoungan Prov., 16.vii.1982; [GG]1♀, Suweon, 7.vii.1976; 2♂, Gwangleung, 40 km NE Seoul, 10.vii.1990, 17.vi.1994; 1♀, 1♂, Mt.Cheonggye-san, 23.vii.1996; 1♂, Mt. Cheonggye-san, 25.vii.1996; 1♂, Mt. Suri-san, 9.vii.1997; 5♀,


1♂, Mt. Mani-san, Isl. Ganghwado, 16.vii.1995, 26.vii.1997; 6♀, 10♂, Mt. Soyo-san, 9.vi.1996, 7.vii.1996; [GW] 2♂, Chuncheon, 26.vi.1984, 13.vi.1989; 1♂, Gajeongri, 22.vii.1995; 1♂, Mt. Gubong-san,20.vi.1998; 1♂, Mt. Palbong-san, Hongcheon, 5.vii.1990; 1♂, Mt. Daeam-san, 28.vii.1988; 1♀, MtOdae-san, 6.vii.1989; 1♂, Mt. Taehwa-san, Youngweol, 14.vii.1998; 1♂, Mt. Jeombong-san, 11.vii.1997; 2♂, Mt. Chiak-san, Wonju, 12.vii.1997, 13.vii.1997; 3♂, Mt. Deokga-san, Wonju, 24.vii.1997; 1♀,Mt. Gyebang-san, 7.viii.1997; [JB] 1♀, Muju, 13.viii.1975; [JJ] 1♀, 1♂, Youngsil, 22.viii.1992, 2.vii.1994; 1♀, Seongpanak, 1.vii.1994.

DISTRIBUTION: Korea (North, Central, South, Jeju), Russian Far East (Pimorsk), China (Heilong-jiang, Shanxi, Henan, Guizhou, Hubei, Zhejiang, Jiangxi, Yunnan), Japan (Hokkaido, Honshu, Shi-koku, Kyushu), Taiwan, N. America.

HOST PALNTS: Rosaceae- Prunus yedoensis, P. persica, P. pseudocerasus, and P. mume (Li, 2002; Parkand Ponomarenko, 2007).

REMARKS: The species has been known as D. picrocarpa (Meyrick) in Korea and the adjacent coun-tries, including Japan and China. The latter was described from Khasis, N. India. The lectotype inBMNH lacks an abdomen. Hodges (1986) identified the American species as D. picrocarpa andnoted that the Japanese species is identical to as D. picrocarpa, with illustration of the male genitalia,based on Japanese specimens. He noted that the American species was probably introduced fromthe origin country. Ponomarenko (2004) identified the Japanese species, D. heriguronis Matsumuradiffer from D. picrocarpa and separated it from the latter. Thus, it is considered that the Americanspecies is probably D. heriguronis Matsumura.


Fig. 130. Dichomeris heriguronis (Matsumura). A. Male genitalia; B. Aedeagus. C. Female genitaliawith posterior part; D. Ditto, ductus bursae+corpus bursae.

A B D

C

131. Dichomeris horoglypta Meyrick, 1932

Dichomeris horoglypta Meyrick, 1932, Exot. Microlep., 4: 202. Dichomeris horoglypta: Gaede, 1937: 434; Issiki, 1957: 42; Clarke, 1969: 24; Park, 1994: 17; Park and

Hodges, 1995a: 37; Li and Zheng, 1996: 232; Ponomarenko, 1997: 22; Li, 2002: 353; Park, 2004: 96.TL: Hasimoto, Kii, Japan.

ADULT: Wingspan 14 mm. The forewing ground color ochreous, scattered with brownish scalesalong costa and the inner margin; a dark brown costal patch near 3/4 length; a broad brownishfascia well-developed along the outer margin. The distinct brown fascia along outer margin of theforewing is a good separable character D. horoglypta from its allies.

Male genitalia (Figs. 131A, B). See also Clarke (1969, pl. 12, Figs. 4a, b); Park and Hodges (1995a,Figs. 34, 35); Li and Zheng (1996: Figs. 7, 8); Li (2002: Fig. 404); Park and Ponomarenko (2007, Pl. 53,Fig. 181). Gnathos relatively strong; culcitula with round distal margin. Valva exceeding apex ofuncus; valvella rather short, digitate.Vinculum divided into two arms; processes of sica equal inlength, more or less dilated to the base. Aedeagus with thick cornutus about 1/3 of its length;anellus without processes.

Female genitalia (Fig. 131C). See also Park (1994: Fig. 8d); Li and Zheng (1996: Fig. 9); Li (2002: Fig.405); Park and Ponomarenko (2007, Pl. 53, Fig. 181).

SPECIMENS EXAMINED: [JN] 1♀, Mt. Jiri-san, 19.vii.1981; 2♀, Mt. Bekun-san, 19.viii.1992; [JJ] 1♀,Mt. Halla-san, 5.viii.1986.


Fig. 131. Dichomeris horoglypta. A. Male genitalia; B. Aedeagus; C. Female genitalia.

A C

B

DISTRIBUTION: Korea (South, JJ), China (Shaanxi, Anhui), Japan (Honshu, Shikoku).HOST PLANT: Leguminosae- Indigofera pseudotenctoria (Li, 2002).

132. Dichomeris issikii (Okada, 1961)

Telephila issikii Okada, 1961, Pub. ent. Lab. Univ. Osaka Pref., 6: 47. Telephila issikii: Moriuti, 1982, I: 285, II: 215.Dichomeris issikii: Park and Hodges, 1995a: 59; Park, 2004: 99; Park and Ponomarenko, 2007: 147.Dichomeris synergastis Ponomarenko and Park, 1996, Korean J. Appl. Entomol., 35(2): 116.

TL: Japan.

ADULT: Wingspan 20-22 mm. This species is similar to D. atomogypsa (Meyrick) in the forewingcolor pattern, but the forewing is more narrow and elongated. The species was described as a newspecies by Ponomarenko and Park (1996), but it was synonymized with this species. Male withouthair-pencils on mesothoracic anepisternum.

Male genitalia (Figs. 132A, B). See also Okada (1961: Figs. 5, 6); Ponomarenko and Park (1996: Figs.9-11). Park and Ponomarenko (2007, Pl. 53, Fig. 182). Valva slightly curved at middle dorsally anddilated towards apex. Vinculum narrow, band-like, without lateral arms; sica with broad basal plate


Fig. 132. Dichomeris issikii (after Okada , 1961). A. Male genitalia; B. Aedeagus; C. Female genitaliawith corpus bursae.

A B C

and two hook-like processes curved to left. Aedeagus about equal to genitlaia in length, with apair of well-developed lobes from zone laterodorsally, right one almost straight, left one bent nearbase; cornutus single, bar-shaped.

Female genitalia (Fig. 132C). See also Okada (1961: Figs. 7-9). Park and Ponomarenko (2007, Pl. 89,Fig. 182).

SPECIMENS EXAMINED: [GG] 3♂, Yongin, 21.v.1989; [GW] 1♂, Mt. Odae-san, 22.v.1989; 1♂, Mt.Gaejuk-san, Youngwol, 3.viii.1989.

DISTRIBUTION: Korea (Central), Japan (Honshu, Kyushu).REMARKS: Dichomeris synergastis Ponomarenko and Park was synonymized with this species

(Park and Ponomarenko, 2007).

133. Dichomeris lespedezae Park, 1994

Dichomeris lespedezae Park, 1994, Ins. Koreana, 11: 4. Dichomeris lespedezae: Park and Hodges, 1995a: 25; Ponomarenko, 1997: 23; 1999: 225, 231; Park, 2004:

88; Park and Ponomarenko, 2007: 148.Dichomeris harmonias: Issiki, 1957: 40; Okano, 1959: 270; Moriuti, 1982: I: 285, II: 215; Park, 1983: 504;

1991a: 121 (misid.).TL: Mt. Halla, JJ, Korea.

ADULT: Wingspan 13-14 mm. The species is superficially similar to D. harmonias and was oftenmisidentified as the latter, but they can be distinguished by followings: second segment of labialpalpus with rather trapezoidal scale-tuft directed anteriorly; forewing less elongate than that of D.harmonias; ground color of forewing light orange whereas brownish orange of the latter; postmedianfascia of forewing broad, while narrowed and incurved in D. harmonias. More distinct separablecharacters are found in the structure of the male and female genitalia.

Male genitalia (Figs. 133A, B). See also Park (1994: Figs. 1a, b); Ponomarenko (1999: Figs. 127: 1, 2);Park and Ponomarenko (2007, Pl. 54, Fig. 184). Valva greatly exceeding apex of uncus; valvella digi-tate, slightly dilated distally. Vinculum divided into two arms which dilated ventrally; processesof sica almost equal in length, the left one gutter-like apically and the right one with ventral knobbefore apex; anellus with two symmetric subulate lateral processes. Aedeagus long, about equal tothe length of genitalia, narrowed towards base; with two slender lateral lobes; cornutus absent.

Female genitalia (Fig. 133C). See also Park (1994: Fig. 1c); Ponomarenko (1999: Fig. 135: 2); Parkand Ponomarenko (2007, Pl. 90, Fig. 184). Antrum very wide, with broad sclerites along lateral sides.Ductus bursae with heavily sclerotized ridges on ventral surface, expanded anteriorly forming anovate memberanous bursae. Corpus bursae round; signum consisting of two oval plates bearingnumerous short spines around margin; accessory bursae arising from area of signum.

SPECIMENS EXAMINED: [GG] 1♂, Hongleung, 8.vii.1996; 1♂, Gwangleung, 13.viii.1986; 1♂, Mt.Daeduk-san, 11.vii.1997; 1♂, Mt. Yaksu-san, 9.viii.1989; [GW] 2 ex, Chuncheon, 12.viii.1988, 21.vii.1992; 3 exs, Chuncheon, 5.v.1994; 1♂, Mt. Samak-san, 25.ix.1989; 1♀, Mt. Jeombong-san, 10.viii.1992;1♂, 1♀, Pyoungchang, 31.vii.1991; 2 exs, Mt. Seolak-san, 25.viii.1989, 15.vi.1993; [CN] 1♂, Mt.Gyeryong-san, 5.vii.1980; [JN] 2 exs, Mt. Paekun-san, 19.viii.1992, 20.viii.1996; [JJ] 1♂, Mt. Halla-


san, 5.vii.1986. DISTRIBUTION: Korea (Central, South, Jeju), Russia (Primorsk), Japan (Honshu, Kyushu). HOST PALNTS: Fagaceae- Lespedeza sp.; Trifolium repens (Ponomarenko, 1993; Park, 1994b; Park and

Hodges, 1995b).

134. Dichomeris litoxyla Meyrick, 1937

Dichomeris litoxyla Meyrick, 1937, Exot. Microlep., 5: 123. Dichomeris litoxyla: Clarke, 1969: 28; Park, 1994: 12; Ponomarenko, 1999: 222, 231; Park, 2004: 92;

Park and Ponomarenko, 2007: 149.TL: Yakovlevka, Primorsk Territory, Russia.

ADULT: Wingspan 21-25 mm. The color pattern of the forewing is very similar to D. derasella[Denis and Schiffermüller], the forewing broader, slightly dilated distally, a dark brown fasciadeveloped along termen.

Male genitalia (Figs. 134A, B). See also Clarke (1969: pl. 14, Figs. 4a, b); Park (1994: Figs. 6a, b);Ponomarenko (1999: Figs. 126: 3, 4); Park and Ponomarenko (2007, Pl. 54, Fig. 185). Valva greatlyexceeding apex of uncus; sacculus broad at base, digitate, extending beyond 4/5 length of valva;


Fig. 133. Dichomeris lespedezae. A. Male genitalia; B. Aedeagus; C. Female genitalia.

A B C

valvella digitate, rather slender. Vinculum divided into two arms; processes of sica equal in length,serrate on ventral margin, with pointed apex. Aedeagus with four needle-like cornuti.

Female genitalia (Fig. 134C). See also Park (1994: Fig. 6c); Ponomarenko (1999: Fig. 135, 1). Parkand Ponomarenko (2007, Pl. 90, Fig. 185). Antrum weakly sclerotized on dorsal surface, with irre-gular distal margin. Signum composed of minute spicules on anterior part of corpus bursae; acces-sory bursae arising from wall of spiculous patch.

SPECIMENS EXAMINED: [GG] 1♂, Is. Yeonjong, 13.viii.1996; [GW] 3♀, 2♂, Mt. Daeam-san, 28.vii.1988.

DISTRIBUTION: Korea (Central), Russia (Irkutsk, Transbaikalia, Amursk, Khabarovsk, Primorsk),China (Heilongjiang).

135. Dichomeris lutilinea Ponomarenko and Park, 1996

Dichomeris lutilinea Ponomarenko and Park, 1996, Korean J. Appl. Entomol., 35(2): 118. Dichomeris lutilinea: Ponomarenko, 1997: 25; Park, 2004: 97; Park and Ponomarenko, 2007: 149.

TL: Chuncheon, Korea.


Fig. 134. Dichomeris litoxyla. A. Male genitalia; B. Aedeagus; C. Female genitalia.

A B

C

ADULT: Wingspan 15.5 mm. The color pattern of the forewing is very similar to that of D. cuspisPark, but it can be easily distinguished by the male genitalia: valvella long, almost equal to the halfof cucullus; vinculum not divided into two arms; sica with basal extended plate, bifurcate into twolobe apically, lobes shorter than basal plate; sacculus long and slender, with a spine at apex. Femaleis unknown.

Male genitalia (Figs. 135A, B). See also Ponomarenko and Park (1996: Figs. 12-14); Park andPonomarenko (2007, Pl. 54, Fig. 186). Valva gradually dilated to apex. Valvella (Fig. 134B) as longas 1/2 length of valva. Vinculum thin, band-shaped; lateral lobes longer than valvella, curved in S-shape. Aedeagus with two lateral lobes, longer one nearly twice than shorter one.

SPECIMENS EXAMINED: [GW] 1♂, Chuncheon, 15.vi.1992. DISTRIBUTION: Korea (Central).

136. Dichomeris minutia Park, 1994

Dichomeris minutia Park, 1994, Ins. Koreana, 11: 21. Dichomeris minutia: Ponomarenko, 1997: 26; Park, 2004: 97; Park and Ponomarenko, 2007: 150.

TL: Suweon, Korea.


Fig. 135. Dichomeris lutilinea (after Ponomarenko and Park, 1996). A. Male genitalia; B. Sicae; C.Aedeagus.

A C

B

ADULT: Wingspan 7.5-9.5 mm. Head and thorax ochreous. Forewing elongate; ground colorpale ochreous, dark-brown scales scattered throughout, more dense near apex and termen; threedistinct dark-brown discal spots presented; costa gently arched beyond 3/4 length; apex pointed.Hindwing pale gray. Female is unknown.

Male genitalia (Figs. 136A, B). See also Park (1994: Figs. 10a, b); Park and Ponomarenko (2007, Pl.54, Fig. 187). Valva not reaching apex of uncus; valvella very slender, about 1/3 length of valva.Vinculum narrow; lateral lobes slender, bar-shaped with short, spine-like process at base, arisingfrom middle, about 2/3 length of vinculum; sica not developed. Aedeagus slender, with two long,needle-like lateral lobes; dorsal lobe arising from zone heavily sclerotized with obtuse apex.

SPECIMENS EXAMINED: [GG] 1♀, 10♂, Suweon, 11.vi.1974, 11.iv.1974, 12.vi.1974, 2.viii.1974, 11.iv.1976, 16.ix.1976, 31.v.1982, 13.vii.1982, 21.ix.1982, 22.ix.1982, 27.viii.1983; 1♀, 1♂, Mt. Cheonggye-san, 12.viii.1976, 23.vii.1996; 3♂, Gwangleung, 10.vii.1982; 1♂, Mt. Soyo-san, 3.vi.1997; 1♂, Isl.Daebudo, 28.vi.1997; 1♀, Mt. Cheonma-san, 29.vii.1997; 1ex, Mt. Godae, Yeonchon, 27.v.2000; [GW]21♀, 18♂, Chuncheon, 9.v.1980, 29.v.1980, 5.vi.1985, 21.vi.1985, 28.v.1990, 7.vi.1990, 22.iv.1985,22.iv.1986, 15.viii.1992, 5.vi.1995, 7.vi.1997, 4.vi.1998; 4♀, Mt. Gubong-san, 31.v.1998; 1♂, Mt. Samak-san, 19.vii.1989, paratype; 1♂, Mt. Palbong-san, Hongcheon, 5.vii.1990; 1♂, Mt. Gwangdeok-san,Hwacheon, 19.viii.1997; 1♀, Mt. Jeombong-san, 11.vii.1997; [CB] 1♂, Mt. Ingyeong-san, Cheungju,23.viii.1997; [CN] 1♂, Mt. Daedun-san, 18.v.1990; [JB] 8exs, Mt. Daedun-san, 22.v.1992; [JN] 1ex,Mt. Cheongwan-san, Janghyun, 27.v.2000.

DISTRIBUTION: Korea (Central, South), Russian Far East (Primorsk).


Fig. 136. Dichomeris minutia. A. Male genitalia; B. Aedeagus.

A B

137. Dichomeris mitteri Park, 1994

Dichomeris mitteri Park, 1994, Ins. Koreana, 11: 17. Dichomeris mitteri: Park and Hodges, 1995a: 34; Li and Zheng, 1996: 245; Ponomarenko, 1997: 27; Li,

2002: 364; Park, 2004: 97; Park and Ponomarenko, 2007: 150.TL: Chuncheon, Gangweon, Korea.

ADULT: Wingspan 13.5-14 mm. This species is similar to the Oriental species, D. ferruginosaMeyrick, but it can be easily separated by the second segment of labial palpus rather rectangular,whereas it is porrected with long hairlike scales in the latter. The genital character is much closerto D. horoglypta in general appearance, but the aedeagus differs from that of the latter, by lacking acornutus.

Male genitalia (Figs. 137A, B). See also Park (1994: Figs. 8a, b); Li (2002: Fig. 414); Park and Pono-marenko (2007, Pl. 54, Fig. 188). Gnathos with large, conic culcitula. Valva short, not reaching toapex of uncus; valvella digitate, broad at base. Vinculum narrow, shorter than tegumen+uncus,lacking lateral lobes; processes of sica long, slender, nearly equal in length, weakly serrated onventral margin. Sacal region nearly flat. Aedeagus simple, stout, lacking lateral processes andinternal lobe.

Female genitalia (Fig. 137C). See also Park (1994: Fig. 8c). Park and Ponomarenko (2007, Pl. 90, Fig.188). Antrum weakly sclerotized, in W-shaped. Ductus bursae about 2/3 length of corpus bursae,with two heavily sclerotized ridges arising from near middle: left one extended broadly and sepa-


Fig. 137. Dichomeris mitteri. A. Male genitalia; B. Aedeagus; C. Female genitalia.

A B C

rated into two plates surrounding signa; right one shorter, slender; accessory bursae arising fromnear middle of two signa. Signa double.

SPECIMENS EXAMINED: [GW] 2♀, 1♂, Chuncheon, 27.viii.1984, 7.v.1989, 9.v.1989; 1♂, Jiamri, 15km N. Chuncheon, 6.v.1994; 1♀, Yangyang, 30.v.1987.

DISTRIBUTION: Korea (Central), China (Shaanxi), Japan.

138. Dichomeris oceanis (Meyrick, 1920)

Dichomeris oceanis Meyrick, 1920, Exot. Microlep., 2: 306. Dichomeris oceanis: 1925: 177; Gaede, 1937: 441; Clarke, 1969: 43; Liu et al., 1981: 19; Kuznetzov,

Stekolnikov, 1984: 38; Ponomarenko, 1992: 170; Park, 1994: 9; Park and Hodges, 1995a: 14; Ueda etal., 1995: 150; Li and Zheng, 1996: 230; Ponomarenko, 1997: 27; 1999: 224, 228; Li, 2002: 368; Park,2004: 89; Park and Ponomarenko, 2007: 1510.

Ypsolophus limitellus Caradja, 1926, Dt. ent. Z., Iris, 34: 113.Dichomeris yanagawanus Matsumura, 1931: 1083.

TL: Japan.

ADULT: Wingspan 16-22 mm. The species is characterized by the forewing having a large dark-brown patch in discal cell and dark brown fascia along termen, which are connected in a S-shape.


Fig. 138. Dichomeris oceanis. A. Male genitalia; B. Aedeagus; C. Female genitalia.

A B C

Male genitalia (Figs. 138A, B). See also Kuznetcov, Stekolnikov (1984, Fig. 16, Б); Ponomarenko(1992: Figs. 20-23); Park and Hodges (1995a, Figs. 1, 1a, 2); Ponomarenko (1999: Figs. 111: 1, 2); Li(2002: Fig. 418); Park and Ponomarenko (2007, Pl. 55, Fig. 189). Gnathos small; culcitula with rounddistal margin. Valva greatly exceeding apex of uncus; valvella short stout; vinculum divided intotwo arms from middle; sica stout; lobes equal in length with triangular lateral knobs before apex.Aedeagus with triangular lateral processes; cornutus needle-like.

Female genitalia (Fig. 138C). See also Clarke (1969: Figs. 5a, b); Park and Hodges (1995a: Fig. 72);Ponomarenko (1999: Fig. 132: 1); Li (2002: Fig. 419); Park and Ponomarenko (2007, Pl. 90, Fig. 189).Antrum extremely broad, with pouch-like free flaps on caudal margin laterally, width about twicelonger than length. Ductus bursae with a large U-shaped sclerite in conjunction with corpus bur-sae, with two other large triangular ridges laterally; ductus seminalis arising from conjunction.Corpus bursaewith two weakly sclerotized foot-shaped signa; accessory bursae originates fromsigna, posterior 2/3 denselyspiculous.

SPECIMENS EXAMINED: [GG] 2♂, Gwangleung, 13.viii.1994, 22.ix.1994; 1♀, 1♂, Mt. Cheunggye-san, 19.viii.1976; 1♂, Mt. Suri-san, Gunpo, 25.vii.1997; [GW] 1♂, Mt. Gwangdeok-san, Hwacheon,19.viii.1997; [JB] 1♂, Mt. Mireuk-san, Iksan, 23.viii.1997.

DISTRIBUTION: Korea (Central, South), Russian Far East (Primorsk), China (Heilongjiang, Beijing,Gansu, Shaanxi, Henan, Shandong, Anhui, Zhejiang, Fujian), Japan (Honshu, Shikoku, Kyushu),Taiwan.

HOST PALNTS: Leguminosae- Wisteria floribunda, W. japonica, W. sinensis, W. brachybotrys, and Mil-lettia japonica (Li, 2002; Park and Ponomarenko, 2007).

139. Dichomeris polypunctata Park, 1994

Dichomeris polypunctata Park, 1994, Ins. Koreana, 11: 16. Dichomeris polypunctata: Li and Zheng, 1996, 24: 239; Ponomarenko, 1997: 29; 1999: 225, 230; Park,

2004: 95; Park and Ponomarenko, 2007: 152. TL: Chuncheon, [GW], Korea.

ADULT: Wingspan 20-21 mm. The species was described, based on a female. It is very similar tothe European species, D. limosellus (Schlager), but the forewing ground color is more yellowishorange.

Male genitalia (Figs. 139A, B). See also Emeljanov, Piskunov (1982: Fig. 59); Ponomarenko (1999:Figs. 128: 3, 4); Park and Ponomarenko (2007, Pl. 55, Fig. 191). Sacculus serrated on ventral margin,with curved apex ventrally; vinculum with small median knobs and sickle-shaped process apically,slightly dilated ventrally; processes of sica asymmetrical in size, bifurcated apically, right processslightly longer than left one. Aedeagus with slender, dorsal processes, longer than cornutus; cor-nutus slender, slightly arched.

Female genitalia (Fig. 139C). See also Park (1994: Fig. 10c); Ponomarenko (1999: Fig. 134: 1). Parkand Ponomarenko (2007, Pl. 91, Fig. 191). Antrum broad, sclerotized, with a large lobe at right side;caudal margin irregularly dentate; anterior margin concave in W-Shape. Ductus bursae with num-erous ridges on ventral surface; accessory bursae arising from right side of junction with ductusbursae.


SPECIMENS EXAMINED: [GW] 1♀, Chuncheon, 12.vi.1989. DISTRIBUTION: Korea (Central), Russian Far East (Primorsk, Kuril Isl.), Transbaikalia, China (Hei-

longjiang), Mongolia.

140. Dichomeris praevacua Meyrick, 1922

Dichomeris praevacua Meyrick, 1922, Exot. Microlep., 2: 504. Dichomeris praevacua: Meyrick, 1925: 176; Gaede, 1937: 442; Li and Zheng, 1996: 233; Park and Pono-

marenko, 2007: 152. Dichomeris quercicola: Issiki, 1957: 42; Moriuti, 1982, I: 284, II: 215; Emeljanov, Piskunov, 1982: 394;

Kostyuk et al., 1994: 10; Park, 1994: 7; Li and Zheng, 1996: 240; Ponomarenko, 1999: 225, 228; Li,2002: 384; Park, 2004: 89 (misid.).TL: Shanghai, China.

ADULT: Wingspan 12-13 mm. Superficially similar to the Indian species, D. quercicola Meyrickwhich was described from N. India, and has been misidentified as the latter by various authors.

Male genitalia (Figs. 140A, B). See also Emeljanov, Piskunov (1982: Fig. 58); Park (1994: Figs. 3a, b);Ponomarenko (1999: Figs. 129: 3, 4); Li (2002: Fig. 436); Park and Ponomarenko (2007, Pl. 56, Fig. 192).Valva extending to apex of uncus, with a protrusion near 3/4 on anterior margin, deeply hollowed


Fig. 139. Dichomeris polypunctata. A. Male genitalia; B. Aedeagus; C. Female genitalia.

A C

B

before apex; valvella slender, with round apex. Vinculum narrow, with small, knob-shaped lateralprocess. Sica nearly symmetrical with slightly shorter right process. Aedeagus stout, large, withslender, needle-like lateral processes.

Female genitalia (Fig. 140C). See also Park (1994: Fig. 3c); Ponomarenko (1999: Fig. 131, 3); Li (2002:Fig. 437); Park and Ponomarenko (2007, Pl. 91, Fig. 192). Antrum widely opened distally. Ductusbursae with various sclerotized ridges and wrinkles ventrally. Corpus bursae not distinguishedfrom ductus bursae; accessory bursae aring from area of signum, densely spiculous.

SPECIMENS EXAMINED: [GG] 1♀, Suweon, 10.ix.1974; [CN] 1♀, Mt. Gyeryong-san, 20.vi.1980; [JN]1♂, Mt. Bekun-san, 19.viii.1992l; [JJ] 1♂, Ipseok, 17.viii.1994.

DISTRIBUTION: Korea (Central, South, Jeju), Russian far East (Khabarovsk, Primorsk), Transbaikalia,China (Beijing, Gansu, Shaanxi, Henan, Hunan, Anhui, Jiangxi), Japan (Honshu), Mongolia.

HOST PLANT: Leguminosae- Lespedeza cyrtobotrya, Quercicus sp. (Li, 2002).

141. Dichomeris rasilella (Herrich-Schäffer, 1854)

Anacampsis rasilella Herrich-Schäffer, 1854, Syst. Bear. Schmett. Eur., 5: 191; Meyrick, 1925: 249.Gomphocrates rasilella: Caradja, 1931: 68; Gaede, 1937: 546.Uliaria rasilella: Zerny, 1927: 479; Meyrick, 1935: 73; Piskunov, 1981: 731; Moriuti, 1982, I: 286, II: 215;


Fig. 140. Dichomeris praevacua. A. Male genitalia; B. Aedeagus; C. Female genitalia.

A B C

Park, 1983: 505; Ponomarenko, 1992: 171; Budashkin and Kostjuk, 1994: 20; Ponomarenko, 1997:11; 1999: 212, 228.

Dichomeris rasilella: Hodges, 1986: 12; Park, 1994: 16; Park and Hodges, 1995a: 52; Karsholt andRiedl, 1996: 121; Li and Zheng, 1996: 240; Elsner et al., 1999: 55; Li, 2002: 386; Park, 2004: 94; Parkand Ponomarenko, 2007: 153. TL: Europe.

ADULT: Wingspan 13-16 mm. Forewing modelately broad. Male without scale-tuft on metatho-racic anepisternum; second segment of labial palpus thickened ventrally, with dorsal scale-tuft.Hindwing with cubital pectin.

Male genitalia (Figs. 141A, B). See also Piskunov (1981, Fig. 663, 2); Moriuti (1982, II: pl. 259, Fig.4); Ponomarenko (1992: Fig. 24); Park and Hodges (1995a, Figs. 56, 57); Elsner et al. (1999, t. 39, Fig.336); Ponomarenko (1999: Figs. 120, 5, 6) Li (2002, Fig. 438); Park and Ponomarenko (2007, Pl. 56,Fig. 193). Valva greatly exceeding apex of uncus. Vinculum slender, expanding before middle andnarrowed basally, lacking lateral lobes. Sica small, asymmetrical, right lobe slightly longer. Aedea-gus stout, about 4/5 length of genitalia, with a straight, long rod-like lateral lobe, about 2/3 lengthof aedeagus; cornutus absent.

Female genitalia (Fig. 141C). See also Park and Hodges (1995a, Fig. 92); Elsner et al. (1999, t. 81,Fig. 336); Ponomarenko (1999, Fig. 130, 3) Li (2002, Fig. 439). Ductus bursae relatively short, about1/2 length of corpus bursae, with a large and a small triangular sclerite ventrally; ductus seminalisarising from posterior part of ductus bursae. Corpus bursae oblong; accessory bursae originatingfrom anterior 3/4; signum absent.


Fig. 141. Dichomeris rasilella. A. Male genitalia; B. Aedeagus; C. Female genitalia.

A B C

SPECIMENS EXAMINED: [GG] 1♂, Gwangleung, 10.vii.1990; 1♂, Mulwang Res., Siheung, 24.viii.1996; 1♂, Mt. Gwangdeok-san, 20.vii.1996; 1♂, Mt. Suri-san, 28.viii.1997; 1♀, Isl. Daebudo, 30.viii.1997; 2♂, Incheon, Isl. Yongyudo, 28.viii.1997; 1♂, Mt. Hwaya-san, 18.vii.1997; [GW] 4♂, Chun-cheon, 12.vi.1989, 19.vi.1990, 4.vi.1998; 1♂, 1♀, Mt. Gubong-san, 31.v.1998, 20.vi.1998: 2♂, Hweng-seung, Dunnae, 15.viii.1992, 7.vii.1990; 1♀, Pyoungchang, Mahari, 6.vi.1996; 1♀, 2♂, Mt. Bangtae-san, Inje, 23-24.vi.1996, 8.ix.1996; 1♂, Mt. Namseolak-san, 12.vii.1997; [CB] 1♀, Mt. Weolak-san,7.vi.1997; [JJ] 1♂, Oradong, 10.vi.1993; 1♀, Ibseokdong, 30.vi.1994.

DISTRIBUTION: Korea (Central, South, Jeju), Russia (European part, Caucasus, South Ural, Altai,Tuva, Irkutsk region, Buryatia, Transbaikalia, Amursk, Khabarowsk, Primorsk, Kuril Isl.), China(Heilongjiang, Qinghai, Shanxi, Henan, Sichuan, Guizhou, Hubei, Anhui, Zhejiang, Jiangxi, Fujian),Japan (Honshu, Kyushu), Central Asia, Europe, Taiwan.

HOST PALNTS: Asteraceae- Centaurea spp., Acosta rhenana, Artemisia vulgaris, A. princeps var. orien-talis, A. pontica (Piskunov, 1981; Moriuti, 1982; Elsner et al., 1999; Li, 2002).

142. Dichomeris sparsella (Christoph, 1882)

Ypsolophus sparsellus Christoph, 1882, Bull. Soc. Nat. Mosc., 57(1): 29. Gaesa sparsella(us): Meyrick, 1925: 179; Gaede, 1937: 449.


Fig. 142. Dichomeris sparsella. A. Male genitalia; B. Aedeagus; C. Female genitalia with appendixbursae.

A B C

Dichomeris strictella Park, 1994: 11; Park, 1996: 65; Ponomarenko, 1997: 32; Ponomarenko, 1999: 225,227 (misid.).

Dichomeris sparsella: Ponomarenko and Mey, 2002: 74; Park and Ponomarenko, 2007: 155. TL: Raddefka (Radde), Russian Far East.

ADULT: Wingspan 20-23 mm. This species is superficially similar to the European species, D.babella Denis and Schiff., but it can be separated by following characters: more elongate forewingwith rather acute apex, with more distinct dark-brown costal patch and median fascia.

Male genitalia (Figs. 142A, B). See also Park (1994: Figs. 5a, b); Ponomarenko (1999: Figs. 128: 1, 2);Park and Ponomarenko (2007, Pl. 56, Fig. 194). Valva exceeding apex of uncus. Vinculum dividedinto two arms, dilated ventrally; Sica asymmetriacal; processes bar-shaped, left one about 2/3length of right one. Aedeagus moderately slender, about same length of genital capsule; left laterallobes exceeding beyond apex of dorsal lobe from zone, with a small tooth near middle; a largetriangular protrusion before middle; dorsal lobe with obtuse apex; cornutus horn-shaped, heavilysclerotized, about 1/2 length of aedeagus.

Female genitalia (Fig. 142C). See also Park (1994, Fig. 5c); Ponomarenko (1999, Fig. 131: 2); Parkand Ponomarenko (2007, Pl. 91, Fig. 193). Antrum broad, heavily sclerotized, divided into twoparts. Ductus bursae long, with heavily sclerotized plate dorsally and with several sclerotizedridges on ventral surface. Corpus bursae extended posteriorly and anteriorly; accessory bursaeoriginating from wall of signum; signum with rounded distal margin.

SPECIMENS EXAMINED: [GG] 1♀, Mt. Soyo-san, 5.viii.1996; [GW] 2♀, 2♂, Chuncheon, 24.iv.1991,1.v.1989, 26.iv.1989, 23.iv.1992; 1♂, Yangyang, 30.v.1987, holotype; [CB] 1♀, 1♂, Mt. Weolak-san,9.viii.1997.

DISTRIBUTION: Korea (Central), Russian Far East (Primorsk).REMARKS: The Korean species was described as D. strictella Park, 1994, but the latter was erro-

nously synonymized with D. chinganella (Christoph) by Park (1996) and it was synonymized againwith his species by Ponomarenko (2002).

143. Dichomeris tostella Stringer, 1930

Dichomeris tostella Stringer, 1930, Ann. Mag. nat. Hist., (10)6: 415. Dichomeris tostella: Gaede, 1937: 444; Inoue, 1954: 70; Issiki, 1957: 42; Okano, 1959: 270; Moriuti, 1982,

I: 284, II: 215; Park, 1983: 504; 1994: 9; Park and Hodges, 1995a: 18; Ueda et al., 1995: 150; Ponoma-renko, 1997: 33; Park, 2004: 91; Park and Ponomarenko, 2007: 155.

Dichomeris kawamurai Matsumura, 1931, 6000 Ill. Ins. Japan: 1082. TL: Japan.

ADULT: Wingspan 18-23 mm. The species is somewhat similar to D. ustalella (Fabricius), but theforewing ground color is yellowish ochreous, instead of brownish ochreous in D. ustalella.

Male genitalia (Figs. 143A, B). See also Park and Hodges (1995a, Figs. 5, 6); Park and Ponomarenko(2007, Pl. 57, Fig. 196). Valva exceeding apex of uncus; valvella short, spatulate. Vinculum pointedapically, lacking lateral lobes. Sica asymmetrical; left lobe as long as valva with a small branchnear basal 1/3, serrated along ventral margin; right one similar to left one, but shorter. Aedeagus


very stout, with well-developed lateral lobes; cornutus short, heavily sclerotizedFemale genitalia (Fig. 143C). See also Park and Hodges (1995a, Fig. 74). Antrum broad, with mem-

branous sacs. Ductus bursae with several irregular sclerotized plates; ductus seminalis arising fromright side of corpus bursae, with sclerotized ring near base. Corpus bursae with dense spiculesdistally; accessory bursae arising from ventral surface anteriorly; signum absent.

SPECIMENS EXAMINED: [GG] 1♀, Mt. Chelma-san, 14.vi.1996; 1♂, Mt. Soyo-san, 7.vii.1996; [GW] 2♀,7♂, Mt. Taebek-san, 27.vi.1996.

DISTRIBUTION: Korea (Central), Japan (Honshu, Shikoku, Kyushu).HOST PALNTS: Rosaceae- Malus pumila var. dulcissima, Prunus mume (Moriuti, 1982).

144. Dichomeris ustalella (Fabricius, 1794)

Tinea ustalella Fabricius, 1794, Ent. Syst., 1794: 307. Tinea capucinella Hübner, 1796, Eur. Schemett. Tin.: 159.Ypsolophus cornutus Fabricius, 1798, Suppl. Ent. Syst.: 505.Ypsolophus ustulatus Fabricius, 1798, Suppl. Ent. Syst.: 506.


Fig. 143. Dichomeris tostella. A. Male genitalia; B. Aedeagus; C. Female genitalia with apendixbursae.

A C

B

Rhinosia ustulella: Treitschke, 1833: 11.Hypsolopha ustulella: Herrich-Schäffer, 1855: 154.Hypsolophus ustulella: Frey, 1856: 139.Ypsolophus ustulella: Heinemann, 1870: 338.Dichomeris ustulella: Meyrick, 1925: 177.Dichomeris ustalella: Gaede, 1937: 444; Issiki, 1957: 42; Okano, 1959: 270; Povolny, 1978: 140; Pisku-

nov, 1981: 731; Moriuti, 1982, I: 284, II: 215; Park, 1983: 503; 1991a: 121; 1994: 9; Simpson, 1989: 17;Park and Hodges, 1995a: 17; Ueda et al., 1995: 150; Karsholt and Riedl, 1996: 120; Li and Zheng,1996: 235; Ponomarenko, 1997: 34; Elsner et al., 1999: 55; Ponomarenko, 1999: 225, 227; Li, 2002:398; Park, 2004: 90; Park and Ponomarenko, 2007: 155.TL: Italy, Europe.

ADULT: Wingspan 17-24 mm. This species is characterized by the brownish ochreous groundcolor of the forewing and brownish gray of the hindwing.

Male genitalia (Figs. 144A, B). See also Povolny (1978, Figs. 11, 12); Piskunov (1981, Fig. 662-1);Park and Hodges (1995a, Figs. 7, 8); Elsner et al. (1999, t. 38, Fig. 332); Ponomarenko (1999, Figs. 129,1, 2); Li (2002, Fig. 450); Park and Ponomarenko (2007, Pl. 57, Fig. 197). Valva narrowed toward base;valvella digitate, curved dorsally. Vinculum relatively slender; lateral lobes arising from beforemiddle, slender. Sica with asymmetrical lobes; left lobe about 2/3 length of vinculum; right oneshorter, with dilated hairly apex. Aedeagus stout, with a large flap-like lateral lobe on left side;cornutus horn-shaped, 1/3 length of aedeagus.


Fig. 144. Dichomeris ustalella. A. Male genitalia; B. Aedeagus; C. Female genitalia.

A

B C

Female genitalia (Fig. 144C). See also Park and Hodges (1995a: Figs. 75, 75a); Elsner et al. (1999, t.81, Fig. 332); Ponomarenko (1999: Fig. 130, 2) Li (2002: Fig. 451); Park and Ponomarenko (2007, Pl.92, Fig. 197). Ductus bursae simple, with parallel lateral walls; ductus seminbalis arising from pos-terior part of corpus bursae, with a long secondary bursae. Corpus bursae ovate; signum large, V-shaped sclerite near middle of corpus bursae.

SPECIMENS EXAMINED: [N. Korea] 1♂, Mt. Keumgan-san, 27.v.1985; [GG] 1♀, Suweon, 2.viii.1974;2♀, 4♂, Gwangleung, 20.v.1981, 17.vi.1994, 27.vi.1986, 20.v.1987, 8.vii.1992; 1♂, Mt. Suri-san,25.vii.1997; 1♀, Mt. Myoungji-san, 26.vi.1992; 3♂, Mt. Mani-san, Isl. Ganghwado, 20.vi.1997; 1♂,Mt. Soyo-san, Dongducheon, 17.v.1997; [GW] 1♀, Chuncheon, 15.vi.1995; 1♂, Jiamri, 15 km N.Chuncheon, 7.vi.1997; 1♀, 1♂, Mt. Gubong-san, 8.vi.1998; 1♀, Bongmyoungri, Chuncheon, 23.vii.1992; 1♂, Goseung, 16.v.1994; 1♂, Mt. Jeombong-san, 5.viii.1997; 2♂, 1♂, Mt. Daedeok-san, 11.vii.1997, 8.viii.1997; 6 exs, Mt. Taebek-san, 27.vi.1996; 1♂, Mt. Seolak-san, Temp. Baekdam, 23.vi.1997;[CN] 1♂, Mt. Daedun-san, 13.vii.2000; [JB] 6♂, Mt. Mireuk-san, Iksan, 14.vi.1997, 23.viii.1997; [JN]1♂, Mt. Wolchul-san, 13.viii.2000; [JJ] 1♂, Ibseokdong, 30.vi.1994; 1♂, 2♀, Seongpanak, 1.vii.1994,3.vii.1994; 2♂, Youngsil, 2.vii.1994; 2♂, 1♂, Mokuje, 15.vi 1993, 16.vi.1993.

DISTRIBUTION: Korea (North Central, South, Jeju), Russia (European part, Caucasus, Primorsk,Kuril Isl.), China (Gansu, Henan, Zhejiang, Jiangxi), Japan (Hokkaido, Honshu, Kyushu), Transcau-casia, Europe.

HOST PALNTS: Betulaceae- Corylus heterophylla var. thunbergii, Carpinus spp. Fagaceae- Fagus silva-tica, Quercus serrata. Aceraceae- Acer spp. (Moriuti, 1982; Li, 2002; Park and Ponomarenko, 2007).

145. Dichomeris anisacuminata Li and Zheng, 1996

Dichomeris anisacuminata Li and Zheng, 1996, SHILAP Revta. Lipid., 24(95): 231; Li, 2002: 331; Sohn,2007: 22. TL: Jiangxi, China.

ADULT: Wing span, 11-12 mm. This species is superficially similar to D. quercicola Meyrick des-cribed from India, and D. praevacua Meyrick from China, but it can be distinguished by the malegenitalia, especially the long sica with symmetric arms. Female is unknown.

Male genitlia (Figs. 145A-C). See Li and Zheng (1996, Figs. 4-6); Sohn (2007, Figs. 56, 56a), Charac-terized by having long, symmetric processes of sica and the aedeagus with heavily sclerotized spine-like process apically and two different length of slender processes arsing from middle.

SPECIMENS EXAMINED: [CN] 1♂, Mt. Gyeryong-san, 24.ix.2000; [GB] 1♂, Isl. Ulneung-do, Nari-bunji, 19-20.vi.2006.

DISTRIBUTION: Korea (Central), China.REMARKS: This species was described in Jiangxi, China, based on two males and a female, and

reported for the first time from Korea by Sohn (2007). Larvae were reared from Quercus mongolica(Sohn, 2007).

HOST PLANT: Fagaceae- Quercus mongolica (Sohn, 2007).


146. Dichomeris japonicella (Zeller, 1877)

Ceratophora japonicella Zeller, 1877, Horae Soc. Ent. Ross., 13: 363. Brachmia japonicella: Meyrick, 1925: 250; Gaede, 1937: 539.Dichomeris japonicella: Li and Wang, 1999: 45; Li, 2002: 408; Park, 2004: 99; Park and Ponomarenko,

2007: 148.Cymotricha japonicella: Omelko, 1999: 106.

TL: Japan.

ADULT: Wingspan 14-17 mm. Forewingrelatively broad; ground color yellowish brown; discalstigmata black, small at middle and end of cell; small blackish dots scattered; apex obtuse; termenwith blackish dots along margin. Hindwing broader than forewing; apex acute.

Male genitalia (Figs. 146A, B). See also Moriuti (1982, Fig. 259: 1); Omelko (1999: Fig. 52: 3); Li (2002:Fig. 461); Park and Ponomarenko (2007, Pl. 53, Fig. 183). Uncus broad, short, with round caudalmargin. Gnathos sclerotized; culcitula triangular. Valva well sclerotized, long, narrowed towardsapex, much more exceeding apex of uncus,; apex round. Parategminal sclerites large, lobe-like, V-shaped in caudal margin. Sica absent. Saccus very broad, with rounded anterior margin. Aedea-gus shorter than valve, with tubular, narrow signum, as long as 1/2 length of aedeagus.

Female genitalia (Fig. 146C); See also Omelko (1999: Fig. 52-4; Li, 2002: Fig. 462). Park and Pono-


Fig. 145. Dichomeris anisacuminata. A. Male genitalia; B. Aedeagus; C. Close-up of uncus &gnathos.

A B

C

marenko (2007, Pl. 90, Fig. 183). Apophyses anteriores very short, less than 1/6 of apophyses pos-teriors. Ostial plate large, sclerotized. with broadly expanded caudal part. Ductus bursae as longas corpus bursae, with long sclerotized plate, as long as ductus bursae; ductus seminalis arisingfrom posterior portion. Corpus bursae large, ovate; signum rounded, densely spiculos on surface;accessory bursae arising from signum plate.

SPECIMENS EXAMINED: [GG] 5♂, Mt. Soyo-san, 7.vii.1996; 2♂, Gwangleung, 27.vi.1986, 17.vi.1994;1ex, Gwangleung, 8.vii.1992; 1♀, Mt. Cheonggye-san, 4.vi.1986; [GW] 11♀, 1♂, Chunchon, 21.vii.1992, 3.vii.1990; 3♂, Mt. Yonghwa-san, 29.vi.1995; 1♂, Bongmyungri, 30.vi.1992; 1♂, 2♂, Mt.Gubong-san, 19-20.vi.1998; 1♂, Mt. Samak-san, 19.vii.1989; [CN] 1♂, Mt. Kyeryong-san, 20.vi.1980,5.vii.1980; [CB] 1♂, Cheongju, 6.vi.1983; [JN] 1ex, Mt. Jiri-san, 23.vii.1985; [JJ] 1♀, Mt. Halla-san,5.vii.1986.

DISTRIBUTION: Korea (Central, South, Jeju), Russian Far East (Primorsk), China (Henan), Japan.REMARKS: The generic status of this species should be discussed in a further study, because the

male genitalia have some considerable differences from congeners and there are some differentopinions on it by authors (Omelko, 1999; Li, 2001; Park, 2004).


Fig. 146. Dichomeris japonicella. A. Male genitalia; B. Aedeagus; C. Female genitalia.

A

B C

Tribe Chelariini Le Marchand, 1947

Chelariini Le Marchand, 1947, Rev. Lépidopt., 11: 153. Type genus: Chelaria Haworth, 1828.

Genus Neofaculta Gozmány, 1955

Neofaculta Gozmány, 1955, Annls hist.-nat. Mus. natn. hung., 6: 308. Type species: Gelechia infernella Herrich-Schäffer, 1854: 162.SYNONYM: Haplovalva Janse, 1958: 32.

Genus Neofaculta comprises five species, mainly distributed in the Palaearctic Region. The genusis one of the Old Word genera and introcuded to the New World (Hodges, 1986). The male geni-talia have small, rounded parategminal sclerites; the valva more or less of same width along thelength; the sacculus lobe-like; the aedeagus with turbinal processes. In the female genitalia, theostial plate is placed at right angle to longitudinal axis. The host plants belong to Ericaceae, Vacci-niaceae, Betulaceae, and Asteraceae (Piskunov, 1981; Elsner et al., 1999).

147. Neofaculta taigana Ponomarenko, 1998

Neofaculta taigana Ponomarenko, 1998, Far East. entomol., 67: 14. Neofaculta taigana: Ponomarenko, 1999: 234; Park and Ponomarenko, 2007: 160.

TL: Primorsk Territory, Russian Far East.

ADULT: Wingspan 20-21 mm. Head, tegula, and thorax dark gray. Forewing dark gray, withindistinct blackish dot at middle, 5-6 concolorous dots along apex and external margin; every dotalong external margin with white scales before. Hindwing gray. Female is unknown.

Male genitalia (Fig. 147A). See also Ponomarenko (1998, Fig. 23; 1999, Fig. 138: 3); Park andPonomarenko (2007, Pl. 59, Fig. 205). Uncus straight, narrowed laterally. Tegumen dilated basally.Gnathos relatively small. Cucullus dilated towards base; valvella equal to sacculus in length; distalpart of sacculus wider than half of cucullus width at middle. Vinculum band-like. Juxta as seta-ceous plate surrounding aedeagus, with ventral process on anterior margin. Saccal region flat.Aedeagus large, long, more than 1.5 times length of genitalia.

SPECIMENS EXAMINED: [GW] 1♂, Sogeumgang, 24.v.1988.DISTRIBUTION: Korea (Central), Russia (Irkutsk, Transbaikalia, Magadansk, Primorsk). HOST PLANT: Ericaceae- Rhododendron spp. (Ponomarenko, 1999).REMARKS: This species has been known as infernella Herrich-Schäffer, 1854 in Korea.


Genus Encolapta Meyrick, 1913

Encolapta Meyrick, 1913, J. Bombay nat. Hist. Soc., 22: 167. Type species: Encolapta metorcha Meyrick, 1913: 167.SYNONYMS: Homoshelas Meyrick, 1935; Chelophoba Meyrick, 1935; Homochelas Clarke, 1969.

Genus Encolapta comprises 14 species, distributed mostly in the eastern Asia. The male genitaliaare characterized as follows: the uncus broadly elongated, separated into basal sclerite and elongat-ed distal part; the cucullus noticeably dilated into ellipitical or other broad plate; the valvella extend-ed ventrally. In the female genitalia, often 8th segment has membranous ventral part; the signumas flattened funnel. The host plants of the species belong to Fagaceae (Li, 2002; Park and Ponoma-renko, 2007).

Key to the species of the genus Encolapta Meyrick, based on the male genitalia

1. Uncus dilated medially; cucullus triangular in shape··················································E. catarina- Uncus narrowed, not inflated medially; cucullus elliptical························································22. Cucullus with a spine at apex; distal part of valvella narrower than distal part of aedeagus; saccus

2.5 times as short as aedeagus; aedeagus with gutter-like apically····························E. tegulifera- Cucullus without spine at apex; distal parts of valvella and aedeagus equal in width; saccus

more than 3 times as short as aedeagus; aedeagus with a rounded flat lobe apically················································································································································E. subtegulifera


Fig. 147. Neofaculta taigana (after Ponomarenko, 1998). A. Male genitalia with aedeagus.

A

148. Encolapta catarina (Ponomarenko, 1994)

Dactylethrella catarina Ponomarenko, 1994, Japan Heterocerists’ J., 176: 8. Dactylethrella catarina: Park and Ponomarenko, 1996b: 345; Li, 2002: 256. Ethmiopsis catarina: Ponomarenko, 1997a: 42; 1999: 238. Encolapta catarina: Ponomarenko, 2004: 76; Park and Ponomarenko, 2007: 161.

TL: Ryazanovka, Primorsk Territory, Russian Far East.

ADULT: Wingspan 15-17 mm. This species is similar to E. tegulifera (Meyrick) and E. subteguliferaPonomarenko, but can be distinguished from them by the genitalic character, especially in theshape of the valva and the valvella of the male and the ostial par of the female.

Male genitalia (Figs. 148A, B). See also Ponomarenko (1997, Figs. 3, 6; 1999, Fig. 140: 3); Li (2002,Fig. 294); Park and Ponomarenko (2007, Pl. 59, Fig. 206). Uncus dilated medially. Gnathos stronglycurved medially at right angle. Valva with well-pronounced neck and more or less triangularcucullus; ventral lobe (= valvella) with sharp apex in lateral view. Vinculum consisting of twosclerites joining ventrally. Sica flat. Aedeagus more or less slender, curved in S-shape: cornutusabsent.

Female genitalia (Fig. 148C). See also Ponomarenko (1994, Fig. 9; 1999, Fig. 145: 1); Li (2002, Fig. 295);Park and Ponomarenko (2007, Pl. 93, Fig. 205). Papillae anales sclerotized. Caudal margin of abdo-minal sternite VIII with deep emargination medially. Ostium bursae oval. Antrum narrow. Ductusbursae narrow, longer than corpus bursae. Corpus bursae ovate; signum equal to ostium in width.


Fig. 148. Encolapta catarina. A. Male genitalia; B. Aedeagus; C. Female genitalia with corpusbursae.

A B C

SPECIMENS EXAMINED: [GG] 1♀, Gapyoung, 15.vii.1986; 1♀, Mt. Mani-san, Isl. Ganghwado, 20.vi.1997; [GW] 1♂, Chuncheon, 21.vi.1985; 1♀, Mt. Gubong-san, 8.vi.1998; 1♂, Mt. Gwangdeug-san,near Cheolwon, 24.vi.1994; 1♀, Mt. Deokga-san, Wonju, 16.vi.1997; 1♂, Sogeumgang, 7.vii.1988.

DISTRIBUTION: Korea (Central), China (Shaanxi, Gansu), Russian Far East (Primorsk), Japan.REMARKS: This species was first reported from Korea by Park and Ponomarenko (1996).

149. Encolapta subtegulifera (Ponomarenko, 1994)

Dactylethrella subtegulifera Ponomarenko, 1994, Japan Het. J., 176: 7. Dactylethrella subtegulifera: Ueda et al., 1995: 149; Park and Ponomarenko, 1996b: 345; Li, 2002: 260. Ethmiopsis subtegulifera: Ponomarenko, 1997a: 43; 1999: 239.Encolapta subtegulifera: Ponomarenko, 2004: 76; Park and Ponomarenko, 2007: 161.

TL: Ussuri, Primorsk Territory, Russian Far East.

ADULT: Wingspan 15-16 mm. The species is closely related to E. tegulifera (Meyrick) in the fore-wing pattern and the male genital structure, but can be distinguished by the smaller size, the ovalshape of cucullus, and the width of distal part of vavella; in the female genitalia, the ostial plateabsent, 8th sternite membranous, and the signum larger than ostium in width.

Male genitalia (Fig. 149A). See also Ponomarenko (1994, Fig. 2: 5; 1999, Fig. 140: 2); Li (2002, Fig.299); Park and Ponomarenko (2007, Pl. 59, Fig. 206). Uncus narrowed distally. Cucullus oval, with-


Fig. 149. Encolapta subtegulifera (after Ponomarenko, 1994): A. Male genitalia with Aedeagus; B.Female genitalia.

A B

out nob at extremity; distal parts of valvella and aedeagus equal in width; saccus about 1/3 lengthof aedeagus. Aedeagus with rounded flat lobe in apical part.

Female genitalia (Fig. 149B). See also Ponomarenko (1994, Fig. 8; 1999, Fig. 141, 4); Park and Pono-marenko (2007, Pl. 93, Fig. 206). Ostium placed on weakly sclerotized trapezoidal plate, which isfused with anterior angles of sternite VIII, forming a pocket. Ductus bursae narrow, longer thancorpus bursae. Corpus bursae ovate; signum funnel-like, flattened.

SPECIMENS EXAMINED: [GG] 1♂, Gwangleung, 10.vi.1998; [GW] 3♂, Chuncheon, 4.vi.1998; 1♂,Mt. Gubong-san, Chuncheon, 31.v.1998; 1♀, Mt. Odae-san, 26.vi.1989.

DISTRIBUTION: Korea (Central), China (Hubei), Russian Far East (Primorsk), Japan (Honshu).REMARKS: This species was first reported from Korea by Park and Ponomarenko (1996).

150. Encolapta tegulifera (Meyrick, 1932)

Dactylethra tegulifera Meyrick, 1932, Exot. Microlep., 4: 201. Dactylethra tegulifera: Gaede, 1937: 419; Clarke, 1969: 3; Inoue, 1954: 7; Issiki, 1957: 43.Dactylethrella tegulifera: Moriuti, 1982, I: 283, II: 214; Park, 1983a: 88; Park, 1983b: 500; Ponomarenko,

1992: 168; Park, 1993d: 38; ESK and KSAE, 1994: 328; Ponomarenko, 1994: 7; Ueda et al., 1995: 149;Li and Zheng, 1998b: 59; Li, 2002: 259.

Ethmiopsis tegulifera: Ponomarenko, 1997a: 43; 1999: 239.Encolapta tegulifera: Ponomarenko, 2004: 76; Park and Ponomarenko, 2007: 162.

TL: Narva, Primorsk Territory, Russian Far East.

ADULT: Wingspan 15-16 mm. This species is externally similar to the two preceeding species,but can be distinguished by the oval cucullus with distinct prominence at extremity.

Male genitalia (Figs. 150A, B). See also Clarke (1969, 3, pl. 1, Fig. 3-3b); Ponomarenko (1994, Fig. 4;1999: Fig. 140: 1); Li (2002: Fig. 297); Park and Ponomarenko (2007, Pl. 60, Fig. 208). Uncus narroweddistally. Cucullus rounded with knob posteriorly. Distal parts of valvella about 1/2 width of aedea-gus distally. Saccus about 1/2.5 of aedeagus in length. Aedeagus gutter-like apically.

Female genitalia (Fig. 150C). See also Ponomarenko (1994, Fig. 7; 1999, Fig. 141, 3); Li (2002, Fig.298); Park and Ponomarenko (2007, Pl. 93, Fig. 208). Papillae anales membranous. Ostial plateabsent. Abdominal sternite VIII membranous. Ostium placed near anterior margin of sternite VIII.Corpus bursae elongate; signum larger than ostium in width.

SPECIMENS EXAMINED: [GG] 2♀, 12♂, Gwangleung, 27.vi.1986, 3.vi.1988, 10.vii.1990, 8.vii.1992,10.vi.1998, 29.vi.1998, 13.vii.1998; 1♂, Suweon, 9.vii.1974; 2♀, Yongin, 21.v.1989, 8.vi.1989; 1♂, Mt.Yumyoung-san, 17.vi.1990; 2♂, Mt. Suri-san, 15.vi.1990, 6.vii.1988; 1♂, Mt. Cheonggye-san, 25.vii.1996; 1♂, Mt. Doduram-san, Icheon, 19.v.1990; [GW] 2♀, 21♂, Chuncheon, 21.vi.1985, 17.vii.1985,7.vii.1987, 1.v.1989, 2.vii.1989, 11.vi.1989, 12.vi.1989, 13.vi.1989, 19.vi.1990, 21.vii.1992; 1♀, Deok-duweon, Chuncheon, 5.vii.1993; 1♀, Moraeje, 16.vi.1993; 1♀, 7♂, Mt. Gubong-san, 20.vi.1998; 1♂,Jiamri, 15 km N. Chuncheon, 15.vi.1995; 1♂, Bongmyoungri, Chuncheon, 30.vi.1992; 1♀, Temp.Weona-sa, 24.vi.1994; 2♂, Mt. Yonghwa-san, 29.vi.1995; 2♂, Hongcheon, Mt. Palbong-san, 5.vii.1990; 1♂, Hwacheon, 2.vii.1985; 1♀, Hwengseung, 7.vii.1990; 2♂, Yangyang Seomyun, 4.vi.1987,1.vii.1987; 1♂, Mt. Odae-san, 26.vi.1989; 11♂, Mt. Taehwa-san, Youngweol, 14.vii.1998; 2♂, Seolak-


dong, 6.vii.1995; 1♀, 3♂, Sogeumgang, 6.vii.1988; 2♂, Mt. Gyebang-san, 21.vi.1996, 7.viii.1997; 1♂,Mt. Jeombong-san, 11.vii.1997; 2♂, Mt. Chiak-san, Wonju, 12.vii.1997; 1♂, Taebek, 22.vi.2000; [CN]1♂, Mt. Daedun-san, 13.vii.2000; [JB] 1♀, Muju, 13.viii.1975; [JJ] 1♀, Mt. Halla-san, 5.vii.1986.

DISTRIBUTION: Korea (Central, South, Jeju), Russian Far East (Primorsk), China (Shanxi, Gansu,Guizhou), Japan (Hokkaido, Honshu, Shikoku, Kyushu).

HOST PALNTS: Fagaceae- Quercus serrata T., Q. acutissima C., Quercus sp. (Li, 2002; Park and Pono-marenko, 2007).

REMARKS: This species was first reported from Korea by Park (1983).

Genus Hypatima Hübner, [1825]

Hypatima Hübner, [1825] 1816, Verz. bekannter Schmett.: 45. Type species: Tinea conscriptella Hübner, [1805], 8: Pl. 41, Fig. 283.SYNONYMS: Chelaria Haworth, 1828; Allocota Meyrick, 1904; Cymatomorpha Meyrick, 1904; Deu-

teroptila Meyrick, 1904; Semodictis Meyrick, 1909; Allocotaniana Strand, 1913; Episacta Turner, 1919.

Genus Hypatima comprises about 100 species, with worldwide distribution. The male genitalia ischaracterized as follows: the uncus large, elongated, with separated basal sclerite; the cucullus


Fig. 150. Encolapta tegulifera. A. Male genitalia; B. Aedeagus; C. Female genitalia with corpusbursae.

A

B C

dilated distally, with distinct neck; the valvella with thorns on the apex. The female genitalia hasan ostial plate, often with honeycomb zones with microtrichia around; the signum usually large,rhomboidal. The host plans of the species belong to Betulaceae, Fagaceae and Salicacea (Ponoma-renko, 1999).

Key to the species of the genus Hypatima Hübner, based on genital structure

1. Valva narrowed from base to middle part, then gradually broadened distally; cucullus more orless triangular, not exceeding apex of uncus; valvella placed at same level with posterior marginof vinculum················································································································H. venefica

- Valva narrowed toward middle, then broadened distally; cucullus more or less rectangular,exceeding apex of uncus; valvella exceeding the posterior margin of vinculum························································································································································H. excellentella

151. Hypatima excellentella Ponomarenko, 1991

Hypatima excellentella Ponomarenko, 1991, Entomol. obozr., 70(3): 617. Hypatima excellentella: Park, 1993: 28; 1995: 71; Ueda et al., 1995: 149; Ponomarenko, 1997: 38; 1999:

236; Li, 2002: 304; Park and Ponomarenko, 2007: 164.Hypatima silvestris auct. (nec Meyrick, 1913): Park, 1983: 88 (misid.).

TL: Barabash-Levada, Primorsk Territory, Russian Far East.

ADULT: Wingspan 13-14 mm. The species is superficially similar to H. silvestris which was knownfrom N. India, however, Korean species had been identified as the latter.

Male genitalia (Figs. 151A, B). See also Ponomarenko (1991, Fig. 25); Park (1993, Fig. 43); Ponoma-renko (1999, Fig. 139: 1); Park and Ponomarenko (2007, Pl. 60, Fig. 209). Uncus corn-shaped, withround distal margin. Gnathos hook-shaped. Vinculum triangular in lateral view. Valvella slightlydilated in distal part, narrower than neck of cucullus medially, exceeded posterior margin of vin-culum. Aedeagus curved beyond middle, smoothly narrowed toward apex.

Female genitalia (Fig. 151C). See also Ponomarenko (1991, Fig. 37); Park (1993, Fig. 60; 1999, Fig.141: 1); Park and Ponomarenko (2007, Pl. 93, Fig. 209). Ostium placed near anterior margin of abdo-minal sternite VIII; ostial plate rhomboidal, with microtrichia; ductus falls into corpus bursae alongits longitudinal axis; apex of apophysis posterioris almost at same level with that of apophysisanterioris.

SPECIMENS EXAMINED: [GG] 2♀, 8♂, Gwangleung, 23.vii.1975, 13.viii.1986, 13.vi.1988, 17.vi.1994,10.v.1998; 2♂, Suweon, 10.ix.1974; 1♀, 1♂, Mt. Suri-san, 25.vii.1987, 15.vi.1990; 2♀, Mt. Soyo-san,3.xi.1992; 2♀, 4♂, Mt. Soyo-san, 9.vi.1996; 1♀, Ipori, 20.viii.1990; 1♂, Mt. Yumyoung-san, 17.vi.1990;1♂, Mt. Cheonma-san, 13.vii.1996; 2♂, Isl. Yongyudo, 11.vi.1997; 1♂, Mt. Mani-san, Isl. Ganghwa-do, 20.vi.1997; 1♀, Mt. Hwaya-san, Gapyoung, 12.ix.1997; 1ex, Mt. Geumdan-san, Hanam, 1.viii.2000; [GW] 1♀, 20♂, Chuncheon, 19.vi.1990, 22.vii.1991, 4.vi.1998, 29.v.1989, 2.x.1989, 3.vi.1992,15.viii.1992, 16.viii.1996, 3.vi.1997; 1♂, Chugok, 2.ix.1996; 6♀, 27♂, Mt. Gubong-san, 31.v.1998,20.vi.1998, 8.vi.1998, 19.vi.1998; 1♀, Mt. Samak-san, 22.vi.1989; 1♂, Temp. Cheongpyoung-sa, 5.vi.1989; 2♂, Mt. Gwangdeok-san, 23.vi.1994; 2♀, 1♂, Yangyang, Seomyun, 4.vi.1987; 1♂, Mt. Yaksu-


san, 8.viii.1989; 1♂, Mt. Bangtae-san, 8.xi.1996; 3♂, Mt. Seolak-san, 9.viii.1989; 1♂, Mt. Seolak-san,15.vi.1993; 3♀, 1♂, Mt. Gyebang-san, 2.viii.1989, 7.vii.1997; 4♂, Mt. Bangtae-san, Inje, 8.ix.1996;[CB] 1♀, 1♂, Mt. Ingyeong-san, Cheongwon, 23.viii.1997; [CN] 1♂, Mt. Daedun-san, 13.vii.2000; 1♂, Mt. Chilgap-san, Cheungyang, 12.vii.2000; [JB] 1♀, Mt. Mireuk-san, Iksan, 23.viii.1997; [JN] 1♂,Mt. Wolchul-san, 13.vii. 2000; [JJ] 1♂, Seonheulri, 1.viii.1993.

DISTRIBUTION: Korea (Central, South, Jeju), Russian Far East (Primorsk), Japan (Honshu), China(Shaanxi, Gansu, Henan, Jiangxi, Anhui), Taiwan, N. India.

HOST PLANT: Fagaceae- Quercus mongolica. Fisch. (Li, 2002; Park and Ponomarenko, 2007).REMARKS: This species was first reported from Korea by Park (1983), under the name of H. silves-

tris Meyrick. Most of specimens were collected from mid-June to mid-August in Korea.

152. Hypatima venefica Ponomarenko, 1991

Hypatima venefica Ponomarenko, 1991, Entomol. Obozr., 70(3): 616. Hypatima venefica: Park, 1993: 29; Ponomarenko, 1997: 41; 1999: 236; Li, 2002: 309; Ponomarenko,

2004: 50; Park and Ponomarenko, 2007: 166.TL: Barabash-Levada, Primorsk Territory, Russia.


Fig. 151. Hypatima excellentella. A. Male genitalia; B. Aedeagus; C. Female genitalia with corpusbursae.

A B C

ADULT: Wingspan 17-19 mm. Similar to the type species of the genus, H. conscriptella Hübnerand the Indian species, H. silvestris Meyrick. Head and thorax light brown. Forewing ochreous,speckled with light brown scales; dark-brown streak near base, middle, beneath and beyond cell;4-5 small brown fascia developed along costa beyond middle.

Male genitalia (Figs. 152A, B). See also Ponomarenko (1997, Fig. 26); Park (1993, Fig. 44); Pono-marenko (1999, Fig. 139, 2); Li (2002: 367); Park and Ponomarenko (2007, Pl. 61, Fig. 211). Closelyresembles the preceding species, H. excellentella. Base of uncus broader. Vinculum triangular inlateral view. Valvella slightly dilated in distal part, narrower than neck of cucullus medially; apexof valvella placed on the same level as posterior margin of vinculum. Aedeagus curved beyondthe middle, smoothly narrowed to the apex.

Female genitalia (Fig. 152C). See also. Ponomarenko (1991, Fig. 36); Park (1993: Fig. 61; 1999, Fig.141: 2); Park and Ponomarenko (2007, Pl. 93, Fig. 211). Apophysis posterioris not reach to end ofapophysis anterioris. Ostium triangular, placed near anterior margin of abdominal sternite VIII;ostial plate semioval, with microtrichia; ductus falls into corpus bursae slightly laterally.

SPECIMENS EXAMINED: [N. Korea] 1♂, Samjiyeon, Mt. Pekdu-san, 19.vii.1977; [GG] 1♂, Mt.Cheonggye-san, 25.vii.1996; 1♂, Mt. Gwangdeok-san, 20.vii.1996; 1♂, Mt. Soyo-san, 5.viii.1996; 3♂,Mt. Hwaya-san, Gapyoung, 4.viii.1997; [GW] 2♀, 1♂, Chuncheon, 30.vi.1992, 12.ix.1996; 1♂, Mt.


Fig. 152. Hypatima venefica. A. Male genitalia; B. Aedeagus; C. Female genitalia with corpusbursae.

A

B C

Samak-san, 19.vii.1989; 1♀, Moonsanri, Youngweol, 25.vii.1996; 1♀, 2♂, Mt. Gyebang-san, 2.viii.1989; 1♀, Mt. Yaksu-san, 9.viii.1989; 1♂, Mt. Odae-san, 6.viii.1989; 1♀, 1♂, Mt. Jeombong-san,5.viii.1997.

DISTRIBUTION: Korea (north, Central), Russian Far East (Primorsk), China (Henan), Japan (Honshu).HOST PLANT: Fagaceae- Quercus mongolica (Li, 2002; Park and Ponomarenko, 2007).

Genus Tornodoxa Meyrick, 1921

Tornodoxa Meyrick, 1921, Exot. Microlep., 2: 432. Type species: Tornodoxa tholochorda Meyrick, 1921: 432.

Genus Tornodoxa Meyrick comprises three species distributed in East Asia. In the male genitalia,the uncus is broad and rounded, with more or less transversally dilated distal part; the valve isstrongly dilated in distal part; the valvella is small, digitate, in some species with pointed apex. Inthe female genitalia, 8th segment has membranous sternal part; the ostial plate is absent; the ductusand corpus bursae are membranous; and the signum is flatenned as a funnel.

Key to the species of the genus Tornodoxa Meyrick, based on the male genitalia

1. Cucullus oval, without pointed bill-like prominence on ventral margin; valvella pointed at apex····································································································································T. longiella

- Cucullus quadrate, with pointed bill-like prominence on ventral margin; valvella digitate········································································································································T. tholochorda

153. Tornodoxa longiella Park, 1993

Tornodoxa longiella Park, 1993, Ins. Koreana, 10: 39. Tornodoxa longiella: ESK and KSAE, 1994: 330; Ueda et al., 1995: 149; Ponomarenko, 1997: 46; Park

and Ponomarenko, 2007: 167.TL: Mt. Gyebang-san, Korea.

ADULT: Wingspan 17-18 mm. This species is similar to H. tholochorda Meyrick in the male geni-talia, but can be distinguished by the forewing venation, with R1 arising before middle, R4 and R5

with a long stalk, M1 free; and the second segment of labial palpus is also separable character.Female is unknown.

Male genitalia (Figs. 153A, B). See also Park (1993, Fig. 58); Park and Ponomarenko (2007, Pl. 61,Fig. 212). Similar to those of the following species, but differs as followings: uncus shorter andwider; cucullus with rounded distal margin; valvella pointed apically. Aedeagus smaller.

SPECIMENS EXAMINED: [GG] 1♂, Suweon, 16.vii.1982; 1♂, Mt. Hwaya-san, Gapyoung, 4.viii.1997;[GW] 4♂, Mt. Gyebang-san, 20.vi.1980; [JB] 1♂, Mt. Deogyu-san, Muju, 13.viii.1985; [GN] 1♂, Nam-


hae, 1.vi.1994. DISTRIBUTION: Korea (Central, South), Japan (Honshu).

154. Tornodoxa tholochorda Meyrick, 1921

Tornodoxa tholochorda Meyrick, 1921, Exot. Microlep., 2: 432. Tornodoxa tholochorda: Meyrick, 1925: 162; 1935: 70; Gaede, 1937: 418; Clarke, 1969(7): 488; Park,

1993: 39; Ponomarenko, 1997: 46; Li, 2002: 311; Park and Ponomarenko, 2007: 167.TL: Tokyo, Japan.

ADULT: Wingspan 20-22 mm. The species is distinguished from the preceding species, T. longiellaPark as noted in the preceding species.

Male genitalia (Figs. 154A, B). See also Clarke (1969, pl. 244, Figs. 1c, d); Park (1993, Fig. 57); Li(2002, Fig. 368); Park and Ponomarenko (2007, Pl. 61, Fig. 213). Uncus broad, densely haired lateral-ly. Ganthos hook-shaped, strong, heavily sclerotized. Valva rather short, with fan-shaped cuculluspointed, with bill-like prominence on its ventral margin; valvella digitate. Aedeagus inflated inbasal half, then slender, bent at 5/6 length.

Female genitalia (Fig. 154C). See also Park (1993, Fig. 59); Li (2002, Fig. 369). Park and Ponoma-


Fig. 153. Tornodoxa longiella. A. Male genitalia; B. Aedeagus.

A

B

renko (2007, Pl. 93, Fig. 213). Ostium bursae with ridges along lateral sides. Corpus bursae verylarge; signum conic.

SPECIMENS EXAMINED: [GG] 1♀, 1♂, Gwangleung, 4.viii.1988; [GW] 2♂, Mt. Daebuk-san, 11.vii.1997.

DISTRIBUTION: Korea (Central), China (Gansu, Shaanxi, Sichuan, Zhejiang, Jiangxi, Guangdong),Japan (Hokkaido, Honshu, Shikoku, Kyushu), Taiwan.

Genus Faristenia Ponomarenko, 1991

Faristenia Ponomarenko, 1991, Entomol. Obozr., 70(3): 601. Type species: Faristenia omelkoi Ponomarenko, 1991: 603.

Genus Faristenia comprises more than 20 species, distributed mostly in East Asia. The male geni-talia are characterized by followings: the tegumen narrowed distally; the uncus short, rounded; the


Fig. 154. Tornodoxa tholochorda. A. Male genitalia; B. Aedeagus; C. Female genitalia with corpusbursae.

A

B

C

gnathos small, hook-like; the cucullus with long neck, moderately dilated distally; the valvellastrongly sclerotized, with different shape of distal part; the vinculum elongated; the aedeagusinflated in basal part. In the female genitalia, the 8th segment is rounded near posterior margin oftergal part, the ostium is placed on the ostial plate with various shapes, and the signum is gutter-like plate or flatenned funnel.

Key to the species of the genus Faristenia Ponomarenko, based on the male genitalia

1. Valva S-shaped with rounded lobe medially on ventral margin················································2- Valva not S-shaped on ventral margin······················································································32. Valvella flatenned on inner margin, with acute apex; vinculum with deep triangular hollow on

posterior margin, reaching base of saccus; basal part of aedeagus three times as wide as distalpart···························································································································F. ussuriella

- Valvella digitate with round apex; vinculum slightly hollowed on posterior margin; basal partof aedeagus four times as wide as distal part···························································F. quercivora

3. Valvella not curved ventrally before apex; cucullus not dilated distally····································4- Valvella curved ventrally before apex, cucullus dilated distally················································································································································F. geminisignella and F. atrimaculata

4. Valva with more or less developed lobe on ventral margin at place of curve; subulate setae, ifpresent, presented only on the lobe···························································································5

- Valva without lobe on ventral margin at place of curve; subulate setae presented along distalhalf of its ventral margin···········································································································7

5. Valva with lobe on ventral margin at middle or before middle·················································6- Valva with lobe on ventral margin presented beyond middle; valvella flatenned dorsoventrally;

vinculum with deep hollow on posterior margin, reaching base of saccus····················F. omelkoi6. Valvella digitate, with beak-like apex directed laterally; basal part of aedeagus four time as broad

as distal part·················································································································F. acerella- Valvella narrowed towards pointed apex; basal part of aedeagus three time as broad as distal

part··························································································································F. furtumella7. Tegumen longer than length of valva; basal part of valva longer than cucullus, lower corner of

ventral margin of cucullus gently angled at lower corner·······································F. maritimella- Tegumen shorter than length of valva; basal part of valva shorter than cucullus, lower corner of

ventral margin of cucullus gently angled at lower corner········································F. jumbongae

155. Faristenia acerella Ponomarenko, 1991

Faristenia acerella Ponomarenko, 1991, Entomol. Obozr., 70(3): 606. Faristenia acerella: Park, 1993: 33; Ponomarenko, 1997: 44; 1999: 244; Park and Ponomarenko, 2007:

169.TL: Barabash-Levada, Primorsk Territory, Russian Far East.

ADULT: Wingspan 15-16 mm. Head pale brownish gray, paler towards frons. Thorax brownishgray. Forewing brownish gray, with a dark-fuscous scale-tuft at basal 1/3 below cell; a large costal


patch presented near middle, with rather small one before and 2-3 small ones beyond it; apexobtuse.

Male genitalia (Figs. 155A, B). See also Ponomarenko (1991, Figs. 15, 16); Park (1993, Fig. 48); Pono-marenko (1999, Figs. 143, 5, 6). Park and Ponomarenko (2007, Pl. 62, Fig. 214). Distal margin ofuncus round. Valva with lobe on the ventral margin medially, subulate setae placed along marginof lobe; valvella digitate, with beak-like apex directed laterally. Vinculum with small rounded hol-low on the posterior margin. Basal part of aedeagus four times as broad as distal part.

Female genitalia (Fig. 155C). See also Ponomarenko (1991, Fig. 29); Park (1993, Fig. 65); Ponoma-renko (1999, Fig. 146: 3); Park and Ponomarenko (2007, Pl. 93, Fig. 214). Ostium bursae relativelysclerotized, with triangular plate around. Ductus bursae thin, very long. Corpus bursae semiovate;signum crescent.

SPECIMENS EXAMINED: [GG] 2♂, Gwangleung, 10.vii.1990, 8.vii.1992; 1♂, Mt. Cheonma-san,13.vii.1996; 1♀, Mt. Hwaya-san, 18.vii.1997; [GW] 1♂, Chuncheon, 19.vi.1990; 2♂, 1♂, Mt. Samak-san, 19.vi.1989, 22.vi.1989; 1♂, Backyangri, Chuncheon, 30.v.1986; 2♂, Mt. Gubong-san, 19.vi.1998;1♂, Hwacheon, 2.vii.1985; 2♀, 3♂, Mt. Obong-san, Hongcheon, 30.v.1985, 4.vi.1985; 2♀, 4♂, Mt.Chiak-san, Wonju, 12.vii.1997; 1ex, Hwengseung, 30.vi.2000.

DISTRIBUTION: Korea (Central), Russia (Primorsk), Japan.


Fig. 155. Faristenia acerella (after Ponomarenko, 1999). A. Male genitalia; B. Aedeagus; C. Femalegenitalia with corpus bursae.

A

B C

HOST PLANT: Aceraceae- Acer ginnala (Ponomarenko, 1991; Park, 1993; Li, 2002; Park and Pono-marenko, 2007).

156. Faristenia atrimaculata Park, 1993

Faristenia atrimaculata Park, 1993, Ins. Koreana, 10: 36. Faristenia atrimaculata: Ponomarenko, 1997: 44; Park and Ponomarenko, 2007: 169.

TL: Muju, JB, Korea.

ADULT: Wingspan 14-16 mm. The species is differentiated from its allies by the absence of theventral protrusion beyond middle of cucullus in the male genitalia. The female genitalia closelyresembles F. ussuriella, but can be distinguished by the longer apophysis anterioris, longer corpusbursae, and rather smaller and round signum. Female is unknown.

Male genitalia (Figs. 156A, B). See also Park (1993, Fig. 54); Park and Ponomarenko (2007, Pl. 62,Fig. 215). Uncus moderate with round distal margin. Gnathos slender, gently curved; apex blunt.Valva paddle-like, without ventral protrusion, broadened beyond middle; valvella spatulate. Sac-cus small. Aedeagus globular at basal half, then slender beyond.

SPECIMENS EXAMINED: [GG] 1♂, Suweon, 20.vi.1977; [GW] 1♂, Chuncheon, 28.vi.1998; [JB] 1♂,Muju, 13.viii.1975; 2♀, 1♂, Muju, 13.viii.1975.

DISTRIBUTION: Korea (Central), Japan.


Fig. 156. Faristenia atrimaculata. A. Male genitalia; B. Aedeagus.

A B

157. Faristenia furtumella Ponomarenko, 1991

Faristenia furtumella Ponomarenko, 1991, Entomol. Obozr., 70(3): 603. Faristenia furtumella: Park, 1993: 34; Ueda et al., 1995: 149; Ponomarenko, 1997: 44; 1999: 244; Park

and Ponomarenko, 2007: 169.TL: Gornotajezhnoe, Ussuri, Primorsk Territory, Russian Far East.

ADULT: Wingspan 15-16 mm. Head and thorax brownish gray. Forewing rather paler than thoseof the allied species; costal patch trapezoidal, scattered with dark-brown short streaks. Hindwinggray.

Male genitalia (Figs. 157A, B). See also Ponomarenko (1991, Figs. 13, 14; 1999, Figs. 143, 3, 4); Park(1993, Fig. 49); Park and Ponomarenko (2007, Pl. 62, Fig. 216). Distal margin of uncus round. Valvawith rather triangular expansion ventrally before middle; subulate setae presented on lobe; valvellanarrower towards apex. Basal part of aedeagus three times as broad as distal part.

Female genitalia (Fig. 157C). See also Ponomarenko (1991, Fig. 28; 1999, Fig. 136: 2); Park and Pono-marenko (2007, Pl. 94, Fig. 216).

SPECIMENS EXAMINED: [GG] 2♂, 1♂, Cheongyangri, 27.v.1996, 15.v.1997; 2exs, Cheongyangri,13.v.1999; 3♀, 33♂, Gwangleung, 8.vi.1977, 9.vi.1977, 27.vi.1986, 3.vi.1988, 10.vi.1998, 27.vi.1988; 2♂,Mt. Soyo-san, 2.vi.1996, 12.iv.1997; 1♀, Mt. Hwaya-san, 18.vii.1997; 1♀, Mt. Suri-san, 22.v.1998;


Fig. 157. Faristenia furtumella (after Ponomarenko, 1999). A. Male genitalia; B. Aedeagus; C. Femalegenitalia with corpus bursae.

A B C

[GW] 8♀, 21♂, Chuncheon, 29.v.1989, 11.vi.1989, 7.vi.1990, 19.vi.1990, 12.vi.1996, 7.vi.1997; 1♀, 1♂,Mt. Samak-san, 22.vi.1989, 13.vi.1990; 1♂, Temp. Cheongpyoung-sa, 5.vi.1989; 1♂, Mt. Daewoo,Yanggu, 14.vi.2000; 1♂, Mt. Gyebang-san, 2.vii.1989; 1♂, Hyangro-bong, 11.vi.1987; 1♂, Mt. Jeom-bong-san, 22.vi.1992; 3♂, Jungseon, 5.vi.1996; 1♂, Sogeumgang, 6.vii.1988; 1♂, Mt. Deokga-san,Wonju, 24.vii.1997; [GB] 2 exs, Uljin, 26-31.vi.2000.

DISTRIBUTION: Korea (Central), Russia (Primorsk), Japan (Honshu).HOST PLANT: Fagaceae- Quercus mongolica (Ponomarenko, 1991).

158. Faristenia geminisignella Ponomarenko, 1991

Faristenia geminisignella Ponomarenko, 1991, Entomol. obozr., 70(3): 614. Faristenia geminisignella: Ueda et al., 1995: 150; Park and Ponomarenko, 1996b: 345; Ponomarenko,

1997: 44; 1999: 244, 246; Li, 2002: 286; Park and Ponomarenko, 2007: 170.TL: Barabash-Levada, Primorsk Territory, Russian Far East.

ADULT: Wingspan 14-16 mm. Similar to the preceding species, but can be distinguished by themale genitalia.

Male genitalia (Fig. 158A). See also Ponomarenko (1991, Figs. 17, 18; 1999, Figs. 142: 1, 2); Li (2002,


Fig. 158. Faristenia geminisignella (after Ponomarenko, 1991). A. Male genitalia with aedeagus; B.Female genitalia with corpus bursae.

A

B

Fig. 338); Park and Ponomarenko (2007, Pl. 62, Fig. 217). Valva dilated towards apex beyond middle;valvella curved ventrally before apex.

Female genitalia (Fig. 158B). See also Ponomarenko (1991, Figs. 33-35; 1999, Fig. 146: 4); Li (2002,Fig. 339); Park and Ponomarenko (2007, Pl. 94, Fig. 2176). Ovipositor long, membrane betweenabdominal segments VIII and IX, three times as long as papillae anales; signum as doubled funnel.

SPECIMENS EXAMINED: [GG] 2♂, 1♀, Gwangleung, 8.vii.1992, 17.vi.1994; 1♂, Isl. Daebudo, 28.vi.1997; 1♀, Mt. Hwaya-san, 18.vii.1997; 4♂, Mt. Soyo-san, 7.vii.1996; [GW] 1♂, Mt. Gubong-san,Chuncheon, 20.vi.1998; 1♀, Mt. Gyebang-san, 15.viii.1996; 1♂, Mt. Chiak-san, 12.vii.1997; 1♂, 2♀,Mt. Jeombong-san, 11.vii.1997, 13.vii.1997.

DISTRIBUTION: Korea (Central), Russia (Primorsk), China (Shaanxi, Gansu, Jiangxi), Japan (Hon-shu).

HOST PLANT: Aceraceae- Acer mono (Ponomarenko, 1991).

159. Faristenia jumbongae Park, 1993

Faristenia jumbongae Park, 1993, Ins. Koreana, 10: 37; Ueda et al., 1995: 150; Ponomarenko, 1997: 44;Li, 2002: 288; Park and Ponomarenko, 2007: 171.TL: Mt. Jeombong-san, Korea.


Fig. 159. Faristenia jumbongae. A. Male genitalia with aedeagus; B. Aedeagus; C. Female genitaliawith corpus bursae.

A C

B

ADULT: Wingspan 12-14.5 mm. Head pale gray, darker posteriorly. Tegula and thorax grayishbrown. Forewing brownish orange, with creamy white scales before middle and beyond costalpatch, with several short streaks throughout. Hindwing gray.

Male genitalia (Figs. 159A, B). See also Park (1993, Fig. 55); Li (2002, Fig. 341); Park and Pono-marenko (2007, Pl. 63, Fig. 218). Uncus very short, broadened to a semicircular plate, indented atmiddle. Gnathos small, hook-shaped. Valva slender at basal half, strongly bent near middle;cucullus foot-shaped, with strong long setae rarely along ventral margin, expanded before apex,and sharpened towards apex. Aedeagus moderate, globular in basal 1/3, then narrowed towardapex.

Female genitalia (Fig. 159C). See also Park (1993, Fig. 71); Li (2002, Fig. 342); Park and Ponomarenko(2007, Pl. 94, Fig. 218). Ostium bursaed rather small. Antrum narrow, short. Ductus bursae long,about three times as long as corpus bursae. Corpus bursae elongate; signum large, elongate platewith dentates on surface.

SPECIMENS EXAMINED: [GG] 2♀, 3♂, Hongleung, 13.v.1996, 8.vi.1996, 27.vi.1996, 1.vii.1996; 3♀,4♂, Gwangleung, 27.vi.1986, 8.vii.1992; 1♂, Mt. Gwanak-san, 18.vi.1997; 2♀, Isl. Ganghwado,16.vii.1995; 1♂, Incheon, Isl. Yeongjongdo, 21.vii.1995; 2♀, 2♂, Mt. Cheonggye-san, 25.vii.1996; 1♂,Mt. Mani-san, Isl. Ganghwado, 20.vi.1997; 1♀, 2♂, Isl. Deokjeok, 24.vi.1997; 1♀, Mt. Hwaya-san,13.vi.1989; 3♀, 3♂, Mt. Hwaya-san, 18.vii.1997; [GW] 2♀, 2♂, Chuncheon, 13.vi.1989, 7.viii.1987,21.vii.1992; 3♀, 1♂, Chugok, 30.vii.1986, 3.vii.1999; 1♂, Mt. Gubong-san, 8.vi.1998; 1♀, 3♂, Bong-myoungri, 30.vi.1992, 21.vii.1998; 1♀, 1♂, Mt. Palbong-san, Hongcheon, 5.vii.1990; 1♂, Hweng-seong, 16.vii.1994; 1♂, Mt. Jeombong-san, 22.vi.1992; 1♂, Mt. Odae-san, 6.viii.1989; 3♂, Pyoung-chang, 24.vii.1996; 1♂, Mt. Chiak-san 12.vii.1997; 2♂, Yangyang, 1.vii.1987; 1♂, Youngwol, 14.vii.1998; 1♂, Jeongseon, 23.vii.1996.

DISTRIBUTION: Korea (Central), China (Shaanxi, Gansu), Japan (Honshu).

160. Faristenia maritimella Ponomarenko, 1991

Faristenia maritimella Ponomarenko, 1991, Entomol. Obozr., 70(3): 613. Faristenia maritimella: Park, 1994: 82; Ponomarenko, 1997: 44; Ponomarenko, 1999: 244; Park and

Ponomarenko, 2007: 171. TL: Andreevka vill., Primorsk Territory, Russian Far East.

ADULT: Wingspan 16-18 mm. This species is superficially similar to F. jumbongae Park, but canbe distinguished by the more blackish scales and the yellowish white area beyond middle abovecell. The male genitalia have good separable characters, especially the lack of lobe on ventral mar-gin of valve and larger valvella.

Male genitalia (Fig. 160A). See also Ponomarenko (1991, Figs. 19, 20; 1999, Figs. 142: 3, 4); Park andPonomarenko (2007, Pl. 63, Fig. 219). Valva without lobe on ventral margin; subulate setae presentedalong distal half of its ventral margin; valvella with rounded apex; processes of juxta exceeded pos-terior margin of vinculum.

Female genitalia (Fig. 160B). See also Ponomarenko (1991: Fig. 30; 1999, Fig. 145: 2); Park and Pono-marenko (2007, Pl. 94, Fig. 219).


SPECIMENS EXAMINED: [GW] 1♀, Chugok, Chuncheon, 30.vii.1986. DISTRIBUTION: Korea (Central), Russia (Primorsk).

161. Faristenia omelkoi Ponomarenko, 1991

Faristenia omelkoi Ponomarenko, 1991, Entomol. Obozr., 70(3): 603. Faristenia nigriella Park, 1993: 35.Faristenia omelkoi: Ponomarenko, 1997: 44; 1999: 244, 246; Li, 2002: 291; Ponomarenko, 2004: 50; Park

and Ponomarenko, 2007: 172. TL: Barabash-Levada, Primorsk Territory, Russian Far East.

ADULT: Wingspan 14-15 mm. Head and thorax pale gray. Antenna pale gray, dark-brown annu-latins on flagellum. Forewing fuscous, with several yellowish-white patches before and beyonddark-fuscous costal patch; several dark streaks irregularly scattered. Hindwing pale gray.

Male genitalia (Figs. 161A, B). See also Ponomarenko (1991, Figs. 11, 12; 1999, Figs. 143, 1, 2); Park(1993, Figs. 52, 53); Li (2002, Fig. 345); Park and Ponomarenko (2007, Pl. 63, Fig. 221). Similar tothose of F. ussuriella Ponomarenko, but can be distinguished by the caudal margin of uncus ratherconvex, the median expansion on the ventral margin of valva rather smaller, triangular, and dense-


Fig. 160. Faristenia maritimella (after Ponomarenko, 1991). A. Male genitalia with aedeagus; B.Female genitalia with corpus bursae.

A B

ly setosed medially, instead of the setae along margin in F. ussuriella.Female genitalia (Fig. 161C). See also Ponomarenko (1991, Fig. 27; 1999, Fig. 146, 1); Park (1993,

Fig. 69); Li (2002, Fig. 346); Park and Ponomarenko (2007, Pl. 94, Fig. 219).

SPECIMENS EXAMINED: [GG] 3♀, Mt. Bulam-san, 22.vii.1998; 1♂, Mt. Bugak-san, 23.vii.1998; 1♀,Gwangleung, 8.vii.1992; 4♀, 2♂, Mt. Soyo-san, 9.vi.1996, 7.ix.1996, 7.vii.1996, 5.viii.1996; 3♀, 2♂,Mt. Hwaya-san, Gapyoung, 18.vii.1997, 4.viii.1997; 2♀, Mt. Soyo-san, 9.vi.1996, 7.ix.1996; 1♀, 2♂,Mt. Soyo-san, 7.vii.1996; 1♀, 2♂, Mt. Soyo-san, 5.viii.1996; 3♀, 2♂, Mt. Hwaya-san, Gapyoung,18.vii.1997, 4.viii.1997; [GW] 3♂, Chuncheon, 15.vi.1992, 15.viii.1996; 1♀, 1♂, Mt. Gubong-san,19.vi.1998, 8.vi.1998; 5♀, 11♂, Mt. Bangtae-san, Inje, 8.ix.1996; 4♀, Mt. Gyebang-san, 15.viii.1996; 1♂, Mt. Jeombong-san, 11.vii.1997; 2♀, Mt. Daedeok-san, 8.viii.1997; [CB] 1♀, Mt. Gaya-san, Yesan,15.vi.1997; [JJ] 1♂, Mt. Halla-san, 700 m, 12.viii.1996.

DISTRIBUTION: Korea (Central, South, Jeju), Russian Far East (Primorsk), China (Sichuan, Shaanxi,Gansu), Japan (Honshu).

HOST PLANT: Fagaceae- Quercus mongolica (Ponomarenko, 1991).


Fig. 161. Faristenia omelkoi. A. Male genitalia with aedeagus; B. Aedeagus; C. Female genitalia withcorpus bursae.

A

B

C

162. Faristenia quercivora Ponomarenko, 1991

Faristenia quercivora Ponomarenko, 1991, Entomol. obozr., 70(3): 615. Faristenia quercivora: Ponomarenko, 1992: 168; Park, 1993d: 34; Park and Byun, 1995: 138; Ponoma-

renko, 1997: 45; 1999: 243, 246; Li, 2002: 294; Ponomarenko, 2004: 26; Park and Ponomarenko, 2007:173.TL: Barabash-Levada, Primorsk Territory, Russian Far East.

ADULT: Wingspan 15-17 mm. Head brownish gray. Thorax dark fuscous. Forewing pale brown-ish gray, speckled with creamy-white scales; several dark-fuscous streaks irregularly scattered.Hindwing pale gray.

Male genitalia (Figs. 162A, B). See also Ponomarenko (1991, Figs. 21, 22; 1999, Figs. 142, 5, 6); Park(1993, Fig. 50); Li (2002, Fig. 349); Park and Ponomarenko (2007, Pl. 63, Fig. 222). Valva S-shaped,with a small rounded lobe on ventral margin, directed medially; valvella with rounded apex, slight-ly curved ventrally. Vinculum slightly hollowed on the posterior margin. Aedeagus moderate,global in basal part, distal part narrowed towards apex.

Female genitalia (Fig. 162C). See also Ponomarenko (1991, Fig. 31; 1999, Fig. 145, 3); Park (1993, 68);Li (2002: Fig. 350); Park and Ponomarenko (2007, Pl. 94, Fig. 222). Ostium oval, surrounded by quad-rate ostial plate. Ductus bursae very long, narrow. Corpus bursae large; signum hat-shaped.


Fig. 162. Faristenia quercivora. A. Male genitalia with aedeagus; B. Aedeagus; C. Female genitaliawith corpus bursae.

A

B

C

SPECIMENS EXAMINED: [GG] 1♂, Isl. Gulipdo, 11.viii.1995; [GW] 1♀, 2♂, Chuncheon, 5.vi.1989,11.vi.1989, 12.vi.1998; 5♀, 22♂, Mt. Gubong-san, 31.v.1998, 8.vi.1998, 20.vi.1998, 19.vi.1998; 1♂, MtSeolak-san, 10.viii.1989; 1♀, 2♂, Mt. Gyebang-san, 2.vii.1989, 15.viii.1996, 7.viii.1997; 1♀, Mt. Odae-san, 6.viii.1989; 1♂, Yangyang, 4.vi.1987.

DISTRIBUTION: Korea (Central), Russian Far East (Primorsk), China (Jiangxi, Shaanxi, Gansu),Japan (Honshu).

HOST PLANT: Fagaceae- Quercus mongolica (Ponomarenko, 1991; Li, 2002; Park and Ponomarenko,2007).

163. Faristenia ussuriella Ponomarenko, 1991

Faristenia ussuriella Ponomarenko, 1991, Entomol. Obozr., 70(3): 615. Faristenia ussuriella: Park, 1993d: 35; ESK and KSAE, 1994: 329; Ponomarenko, 1997: 45; 1999: 239;

Li, 2002: 297; Park and Ponomarenko, 2007: 173. TL: Gornotajezhnoe, Ussuri, Primorsk Territory, Russian Far East.

ADULT: Wingspan 14-15 mm. Head grayish brown. Thorax rather dark fuscous. Forewing gray-ish brown, with well-developed costal patch; several dark-fuscous streaks developed near base


Fig. 163. Faristenia ussuriella Ponomarenko: A. Male genitalia with aedeagus; B. Aedeagus; C.Female genitalia with corpus bursae.

A

B

C

along antemedian fascia and below costal patch. Hindwing pale gray; apex rather obtuse; termenslightly sinuated.

Male genitalia (Figs. 163A, B). See also Ponomarenko (1991, Figs. 23, 24; 1999, Figs. 144, 1, 2); Park(1993d: Fig. 51); Li (2002: Fig. 353); Park and Ponomarenko (2007, Pl. 64, Fig. 223). Similar to thoseof F. quercivora, but can be distinguished by the broader uncus and larger ventral expansion of valvaat middle. Aedeagus rather small.

Female genitalia (Fig. 163C). See also Ponomarenko (1991, Fig. 32; 1999, Fig. 145, 4); Park (1993d,Fig. 66); Li (2002, Fig. 354). Park and Ponomarenko (2007, Pl. 95, Fig. 223). Ostium bursae withstrongly convex apex and with a small semicircular flap at middle. Antrum long. Ductus bursaeshort. Corpus bursae ovate; signum round, densely dentate.

SPECIMENS EXAMINED: [GG] 1♀, Mt. Inwang-san, 29.v.1998; 1♂, Mt. Suri-san, 15.vi.1990; 1♂, Mt.Myounggi-san, 26.vi.1992; [GW] 9♀, 15♂, Chuncheon, 21.vii.1992, 15.vi.1992, 11.vi.1989, 13.vi.1989,19.vi.1990, 3.vi.1997, 12.vi.1998; 5♀, 11♂, Mt. Gubong-san, 8.vi.1998, 19.vi.1998; 1♂, Mt. Samak-san, 22.vi.1989; 1♂, Jiamri, 7.vi.1998; 1♂, Yangyang, 4.vi.1987; 1♂, Mt. Gyejok-san, 27.v.1998; 1♂,Mt. Odae-san, 26.vi.1989; 1♀, Hwacheon, 15.vi.1992; 1♂, Mt. Seolak-san, 15.vi.1993; 1♂, Mt. Gye-bang-san, 15.viii.1996.

DISTRIBUTION: Korea (Central), Russian Far East (Khabarovsk, Primorsk), China (Jiangxi, Gansu,Shaanxi), Japan.

HOST PLANT: Fagaceae- Quercus mongolica Fisch. (Ponomarenko, 1991; Li, 2002; Park and Pono-marenko, 2007).

Genus Dendrophilia Ponomarenko, 1993

Dendrophilia Ponomarenko, 1993, Zool. Zhurn., 72(4): 59. Type species: Nothris albidella Snellen, 1884: 171.

Genus Dendrophilia Ponomarenko comprises more than 20 species, mainly distributed in EastAsia. The male genitalia are characterized by the ovate or kernel-shaped uncus with sharp medianprojection, the long saccus, and the elongated valva with dilated cucullus. In the female genitalia,the abdominal sternite VIII nearly divided into two parts laterally, with triangular or quadrateplates; the antrum is variously modified, sclerotized, usually tube-like; the ductus bursae is narrow,usually as long as corpus bursae; the corpus bursae is elongate, with a signum.

Key to the species of the genus Dendrophilia Ponomarenko, based on the male genitalia

1. Valva with ventral margin nearly straight; gnathos inflated before apex···············D. leguminella- Valva with ventral margin concaved in distal part; gnathos not inflated before apex················22. Cucullus concaved on dorsal margin before apex·····································································3- Cucullus not concaved on dorsal margin before apex·······························································43. Vinculum trapezoidal in lateral view, with angular posterior margin; aedeagus narrowed before

apex···················································································································D. mediofasciana- Vinculum triangular in lateral view, with round posterior margin; aedeagus not narrowed before


apex······················································································································D. unicolorella4. Cucullus elongate; dorsal margin nearly S-shaped in distal part, with curved apex ventrally;

vinculum trapezoidal in lateral view; aedeagus with round apex·······························D. albidella- Cucullus semiovate, dorsal margin nearly straight; vinculum triangular in lateral view; aedeagus

with pointed apex····························································································D. neotaphronoma

164. Dendrophilia leguminella Ponomarenko, 1993

Dendrophilia leguminella Ponomarenko, 1993, Zool. Zhurn., 72(4): 68. Dendrophilia leguminella: Ponomarenko, 1997: 47; 1999: 250; Park and Ponomarenko, 2007: 175.

TL: Barabash-Levada, Pogranichnyi distr., Primorsk Territory, Russia.

ADULT: Wingspan 12-13 mm. The species is superficially similar to D. caragnella Ponomarenkowhich was described from Primorye, Russian Far East, but it can be distinguished by the malegenitalia, especially in the neary straight ventral margin of cucullus.

Male genitalia (Fig. 164A). See also Ponomarenko (1993, Figs. 2, 5, 6; 1999, Figs. 148: 3, 4); Park andPonomarenko (2007, Pl. 64, Fig. 225). Gnathos inflated before apex; cucullus large, with nearlystraight ventral margin.


Fig. 164. Dendrophila leguminella (after Ponomarenko, 1993). A. Male genitalia with aedeagus; B.Female genitalia with corpus bursae.

A

B

Female genitalia (Fig. 164B). See also Ponomarenko (1993, Fig. 3, 1; 1999, Fig. 150: 1). Park andPonomarenko (2007, Pl. 95, Fig. 225).

SPECIMENS EXAMINED: [GW] 1♂, Moonsanri, Youngweol, 7.vi.1996; [JN] 1♂, Mt. Baekun-san,Gwangyang, 18.viii.1995.

DISTRIBUTION: Korea (Central), Russian Far East (Primorsk), Japan.HOST PLANT: Fabaceae- Lespedeza bicolor (Ponomarenko, 1991; Li, 2002; Park and Ponomarenko,

2007).

165. Dendrophilia mediofasciana (Park, 1991)

Hypatima mediofasciana Park, 1991, Annls. hist. nat. Mus. natn. hung., 78: 119.Hypatima mediofasciana: Park, 1993: 30.Dendrophilia brunneofasciella Ponomarenko, 1993, Zool. Zhurn., 72(4): 67.Dendrophilia mediofasciana: Park and Ponomarenko, 1996b: 345; Ueda et al., 1995: 149; Ponomarenko,

1997: 47; 1999: 250; Li, 2002: 263; Park and Ponomarenko, 2007: 175.TL: Kaesung, N. Korea.

ADULT: Wingspan 14-15 mm. The species is very close to D. neotaphronoma Ponomarenko super-


Fig. 165. Dendrophila mediofasciana. A. Male genitalia; B. Aedeagus; C. Female genitalia with corpusbursae.

A B C

ficially, but it can be distinguished by the well-developed dark-brown rectangular median fascia inthe forewing.

Male genitalia (Figs. 165A, B). See also Park (1991, Figs. 5-7; 1993, Fig. 45); Ponomarenko (1993,Figs. 2, 3; 1999, Fig. 148: 1); Li (2002, Fig. 300); Park and Ponomarenko (2007, Pl. 65, Fig. 226). Gna-thos modelate, not inflated before apex. Valva with concaved dorsal margin. Vinculum trapezoidalin lateral view, with angular posterior margin. Aedeagus narrowed before apex.

Female genitalia (Fig. 165C). See also Park (1993, Fig. 62); Ponomarenko (1993, Figs. 3, 7; 1999, Fig.149: 3); Li (2002, Fig. 301). Park and Ponomarenko (2007, Pl. 95, Fig. 226).

SPECIMENS EXAMINED: [N. Korea] 1♂, Gaeseong, S. Hwanghae Prov., 30.vii.1982; 1♀, Mt. Myo-hyang, N. Pyoungan Prov., 18.vii.1982; [GG] 2♂, Suweon, 20.vii.1976, 29.vi.1976; 1♀, Gwangleung,20.vii.1982; 1♂, Mt. Yeogi-san, 17.vi.1983; 1♂, Mt. Cheonggye-san, 25.vii.1996; 1♂, Mt. Soyo-san,5.viii.1996; 2♂, Mt. Gwangdeok-san, 20.vii.1996; 1♀, Isl. Yongyudo, Incheon, 28.viii.1997; [GW] 4♀, 7♂, Chuncheon, 3.vi.1992, 20.vii.1987, 14.viii.1987, 2.ix.1988, 12.viii.1988, 3.vii.1990, 21.vii.1992;1♂, Bongmyoungri, Chuncheon, 30.vi.1992; 1♀, 1♂, Chugok, Chuncheon, 30.vii.1986; 1♀, Jiamri,Chuncheon, 24.vii.1995; 1♀, 1♂, Hongcheon, 14.viii.1987; 1♂, Mt. Palbong-san, Hongcheon, 5.vii.1986; 1♂, Hwenggye, 1.viii.1991; 7♂, Pyoungchang, 31.vii.1991; 5♀, 7♂, Seomyun, Yangyang, 1.vii.1987, 25.vii.1987, 26.vii.1987; 3♀, 2♂, Mt. Seolak-san, 10.viii.1989, 9.viii.1989, 17.viii.1993; 4♀, 7♂,Sogeumgang, 6.vii.1988, 7.vii.1988, 8.viii.1988, 9.viii.1988; 1♀, 3♂, Mt. Odae-san, 6.viii.1989; 3♀, 8♂,Mt. Jeombong-san, 10.viii.1992; 2♀, 7♂, Jeongseon, 30.vii.1991; 3♂, Mt. Gyebang-san, 15.viii.1996;1♂, Mt. Namseolak-san, 12.vii.1997; [JB] 1♂, Muju, 13.viii.1975; 1♂, Mt. Naebyun-san, 4-5.viii.1992.

DISTRIBUTION: Korea (North, Central, South), Russian Far East (Primorsk), China (Gansu, Shaanxi,Henan, Jiangxi), Japan (Honshu).

HOST PLANT: Fabaceae- Lespedeza bicolor Turcz. (Ponomarenko, 1991; Li, 2002; Park and Ponoma-renko, 2007).

166. Dendrophilia neotaphronoma Ponomarenko, 1993

Dendrophilia neotaphronoma Ponomarenko, 1993, Zool. zhurn., 72(4): 69. Hypatima obscurella Park, 1993, Ins. Koreana, 10: 30.Dendrophilia obscurella: Park, 1995b: 83.Dendrophilia neotaphronoma: Ponomarenko, 1997: 47; 1999: 250; 251; Li, 2002: 268; Park and Ponoma-

renko, 2007: 176. TL: Barabash-Levada, Primorsk Territory, Russian Far East.

ADULT: Wingspan 11-12 mm. The species is very close to D. mediofasciana Park superficially, butit can be distinguished by the forewing patterns, especially lack of distinct dark brown rectangularmedian fascia. The wing venation of this species differs from that of D. mediofasciana.

Male genitalia (Figs. 166A, B). See also Park (1993, Fig. 46); Ponomarenko (1993, Fig. 2: 7); Park(1995, Figs. 58, 59); Ponomarenko (1999, Fig. 147: 6); Li (2002, Fig. 306); Park and Ponomarenko(2007, Pl. 65, Fig. 227). Valva curved in basal part; cucullus large, more or less ovate, with almoststraight ventral margin. Vinculum triangular in the lateral view. Aedeagus pointed apically.

Female genitalia (Fig. 166C). See also Park (1993, Fig. 63); Ponomarenko (1993, Figs. 3, 4; 1999: Fig.150: 4); Park (1995, Fig. 83); Li (2002, Fig. 307); Park and Ponomarenko (2007, Pl. 95, Fig. 227).


SPECIMENS EXAMINED: [GG] 1♂, Mt. Surak-san, 22.vii.1998; 2♂, Mt. Cheonma-san, 13.vii.1996; 2♂,Mt. Cheonggye-san, 23.vii.1996, 25.vii.1996; 1♀, Isl. Yeongjongdo, Incheon, 26.v.1995; 1♀, 4♂, Isl.Yeongjongdo, Incheon, 26.v.1995, 7.ix.1995, 13.viii.1996, 11.vi.1997; 1♂, Mt. Gwangdeok-san,20.vii.1996; 1♂, Mt. Soyo-san, 28.viii.1997; 1♂, Mt. Suri-san, 28.viii.1997; 2♂, Isl. Yongyudo, Inchon,28.viii.1997, 17.ix.1997; 1♂, Mulwang reser., Siheung, 24.viii.1996; 1♂, Mt. Cheonggye-san, 19.viii.1976; 1♂, Mt. Soyo-san, 3.ix.1992; 1♀, Mt. Myoungji-san, 23.v.1991; [GW] 4♀, 5♂, Chuncheon,14.viii.1987, 2.ix.1988, 12.vi.1998, 2.vii.1989, 2 ix.1989, 16.viii.1996; 2♀, 13♂, Mt. Gubong-san, 8.vi.1998, 19.vi.1998, 20.vi.1998; 1♂, Jiamri, 19.vii.1995; 1♀, Hwaya-san, Chuncheon, 18.vii.1997; 1♂,Yangyang, 4.vi.1987; 1♂, Mt. Seolak-san, 9.viii.1989; 1♀, Mt. Bangtae-san, 24.vi.1996; 2♂, Mt. Odae-san, 26.vi.1980, 6.viii.1989; 1♂, Mt. Jeombong-san, 11.vii.1997; 1♂, Mt. Obong-san, 1.vii.1993; [CN]1♂, Mt. Jiri-san, 20.vii.1996; 1♂, Mt. Gaya-san, Yesan, 24.viii.1997; [JB] 2♂, Mt. Mireuk-san, Iksan,23.viii.1997; [GN] 1♂, Namhae, 2.vi.1994; 1♂, Geochang, 23.viii.1997; [JJ] 1♂, Oradong, 10.vi.1993.

DISTRIBUTION: Korea (Central, South, Jeju), Russian Far East (Primorsk), China (Shaanxi, Sichuan,Jiangxi, Fujian), Japan (Honshu), Taiwan, Japan.

HOST PLANT: Leguminosae- Lespedeza bicolor (Ponomarenko, 1991; Li, 2002; Park and Ponomarenko,2007).


Fig. 166. Dendrophila neotaphronoma. A. Male genitalia; B. Aedeagus; C. Female genitalia withcorpus bursae.

A B C

167. Dendrophilia unicolorella Ponomarenko, 1993

Dendrophilia unicolorella Ponomarenko, 1993, Zool. Zhurn., 72(4): 68. Dendrophilia unicolorella: Ponomarenko, 1997: 48; 1999: 250, 251; Li, 2002: 272; Park and Ponomarenko,

2007: 177.TL: Gornotajezhnoe, Primorsk Territory, Russian Far East.

ADULT: Wingspan 12-13 mm. This species is similar to D. mediofasciana and D. neotaphronoma,however it can be distinguished by the forewing without the drak-brown median fascia and thecucullus of the male genitalia concaved on dorsal margin.

Male genitalia (Fig. 167A). See also Ponomarenko (1993, Fig. 2: 4; 1999, Fig. 148: 2); Li (2002, Fig.313); Park and Ponomarenko (2007, Pl. 65, Fig. 229). Gnathos not inflated before apex; cucullus moreor less slender, concaved on dorsal margin; vinculum triangular, with rounded posterior margin.Aedeagus not narrowed before apex.

Female genitalia (Fig. 167B). See also Ponomarenko (1993, Figs. 3, 8, 9; 1999, Figs. 149: 4, 5); Parkand Ponomarenko (2007, Pl. 95, Fig. 229).

SPECIMENS EXAMINED: [GW] 1♀, Mt. Seolak-san, 15.vi.1993. DISTRIBUTION: Korea (Central), Russian Far East (Primorsk), China (Shaanxi, Gansu).HOST PLANT: Fabaceae- Lespedeza bicolor (Ponomarenko, 1991; Li, 2002; Park and Ponomarenko,

2007).


Fig. 167. Dendrophila unicolorella (after Ponomarenko, 1993). A. Male genitalia with aedeagus; B.Female genitalia with corpus bursae.

A B

Genus Bagdadia Amsel, 1949

Bagdadia Amsel, 1949, Bull. Soc. Fouad I Ent., 33: 321. Type species: Bagdadia irakella Amsel, 1949: 322.SYNONYM: Capidentalia Park, 1995.

Genus Bagdadia Amsel was established, being originally placed in Scythrididae, but it was trans-ferred to the Gelechiidae by Sattler (1973). The genus includes more than 10 species mainly distri-buted in East Asia: two species in the Russian Far East, four species in China, and two species inKorea. It is allied to the genus Hypatima Hubner or Dendrophilia Ponomarenko in the superficialcharacters. The male genitalia are characterized by having a crown-shaped uncus with doublesclerotized articulations connecting tegumen, and elongate valva. The genus Capidentalia Park,1995 was synonymized with Bagdadia by Ponomarenko (2004).

Key to the species of the genus Bagdadia Amsel, based on the male genitalia

1. Cucullus not dilated distally·····································································································2- Cucullus dilated distally······························································································B. gnomia2. Valvella with deep wedge-shaped hollow apically··················································B. claviformis- Valvella with not deep rounded hollow apically···························································B. eucalla

168. Bagdadia claviformis (Park, 1993)

Hypatima claviformis Park, 1993, Ins. Koreana, 10: 31; ESK and KSAE, 1994: 329.Capidentalia claviformis: Park, 1995b: 84; Ponomarenko, 1995: 47; 1999: 253; Li, 2002: 250.Bagdadia claviformis: Sattler, 1999: 235; Ponomarenko, 2004: 32; Park and Ponomarenko, 2007: 178.

TL: Mt. Deogyu-san, Muju, Korea.

ADULT: Wingspan 9-10 mm. Head creamy white, rarely speckled with gray scales. Antennasimple. Forewing ground color yellowish white, irrorated with dark-brown scales irregularly, withwell-developed scale-tuft near 1/4 and middle of costa and near base of center. Hindwing palegray, with strongly sinuate termen.

Male genitalia (Figs. 168A, B). See also Park (1993, Fig. 47); Ponomarenko (1995, Fig. 1; 1999, Fig.151: 1); Li (2002: Fig. 286); Park and Ponomarenko (2007, Pl. 65, Fig. 230). Uncus crown-shaped, short.Gnathos strong, hook-shaped, broadened triangularly at apical portion. Valva slender; taenioidslightly expanded near middle, much more exceeded to apex of uncus; valvella with deep wedge-shaped hollow apically.

Female genitalia (Fig. 168C). See also Park (1993, Fig. 64); Ponomarenko (1995, Fig. 18; 1999, Fig.152, 1); Li (2002: Fig. 287); Park and Ponomarenko (2007, Pl. 95, Fig. 230). Ductus bursae narrow, aslong as corpus bursae; ductus seminalis arising from conjunction of corpus bursae. Corpus bursaelarge, semiovate; signum crescent-shaped.

SPECIMENS EXAMINED: [GG] 2♂, Suweon, 11.vi.1976, 20.vi.1977; [GW] 1♂, Chuncheon, 28.vi.1985;1♀, 2♂, Chuncheon, 15.viii.1992; 1♀, 1♂, Mt. Gubong-san, 20.vi.1998; 1♂, Mt. Chiak-san, 23.vi.


1977; 1♂, Mt. Jeombong-san, 11.vii.1997; [JB] 2♀, 2♂, Mt. Deogyu-san, 13.viii.1975; [GB] 3♂, Mt.Juwang-san, 18.viii.1993; [GN] 1♂, Mt. Gaji-san, 19.viii.1993.

DISTRIBUTION: Korea (Central, South), Russian Far East (Primorsk), China (Shaanxi, Gansu, Henan,Anhui), Japan.

169. Bagdadia eucalla (Li and Zheng, 1998)

Capidentalia eucalla Li and Zheng, 1998, Reich. Mus. Tierkd. Dresden, 32(45): 309; Li, 2002: 251.Bagdadia eucalla: Sattler, 1999: 238; Park and Ponomarenko, 2006: 281; Park and Ponomarenko, 2007:

178.TL: Mt. Xinjiashan, Shaanxi, China.

ADULT: Wingspan 14 mm. This species is similar to the preceeding species, but can be distin-guished by the reddish-brown ground color of two forewing and the male genitalia, especially inthe shape of gnathos without inflation before apex and the bifurcate valvella.Female is unknown.

Male genitalia (Figs. 169A, B). See also Li and Zheng (1998, Fig. 4); Park and Ponomarenko (2006,Figs. 37, 37a); Li (2002, Fig. 288); Park and Ponomarenko (2007, Pl. 65, Fig. 230). Gnathos strong,


Fig. 168. Bagdadia claviformis (after Ponomarenko, 1995). A. Male genitalia with aedeagus; B.Aedeagus; C. Female genitalia with corpus bursae.

A C

B

without inflation before apex. Valva slender, not dilated distally; valvella bifurcate, with triangularprocesses.

SPECIMENS EXAMINED: [GN] 2♂, Geojedo, 14.vii.1999.DISTRIBUTION: Korea (South), China (Shaanxi, Guizhou).

170. Bagdadia gnomia Ponomarenko, 1995

Bagdadia gnomia Ponomarenko 1995, Actias, 2: 50. Capidentalia gnomia: Ponomarenko, 1997: 49; 1999: 254.Bagdadia gnomia: Sattler, 1999: 238; Park and Ponomarenko, 2007: 179.

TL: Barabash-Levada, Primorsk Territory, Russian Far East.

ADULT: Wing span, 11-12 mm. This species is externally similar to the preceeding species, B.eucalla (Li and Zheng), but the forewing gound color is not reddish brown. The male genitalia arecharacterized by the modified apical part of valva with digitate apical process. Female is unknown.

Male genitalia (Figs. 170A-C). Similar to those of B. claviformis (Park) and B. eucalla (Li and Zheng),but can be distinguished by the valva, with dilated distal part having a difitate process apically andthe stout valvella bifurcated apically.


Fig. 169. Bagdadia eucalla. A. Male genitalia; B. Aedeagus.

A B

SPECIMENS EXAMINED: [GB] 4♂, Isl. Ulneung-do, Naribunji, 19-20.vi.2006.DISTRIBUTION: Korea (Isl. Ulneung), Russian Far East (Primorisk), Japan.REMARKS: This species was recently reported from Isl. Ulneung-do, Korea (Park et al., 2013).

Tribe Anarsiini Amsel, 1977

Anarsiini Amsel, 1977, Beitr. naturk. Forsch. SW Dtl., 36: 234.Type-genus: Anarsia Zeller, 1839.

Genus Anarsia Zeller, 1839

Anarsia Zeller, 1839, Isis Oken, Leipzig, 1839: 190.Type species: Tinea spartiella Schrank, 1802: 104. SYNONYM: Ananarsia Amsel, 1959.


Fig. 170. Bagadia gnomia. A. Adult; B. Aedeagus; C. Male genitalia (Ventral view); D. Male genitalia(Lateral view).

A

B C

D

Genus Anarsia comprises more than 100 species, with a worldwide distribution; mostly distribut-ed in Ethiopian and Indo-Malay regions, and about half of the known species are represented inEast Asia. The male genitalia are asymmetrical, usually with uncus bearing more or less a medianknob-like process and rounded socii; the cucullus usually inflated with modified setae mediallyand with strong sclerotized processes on the ventral margin; the aedeagus usually slender, sinuousor curved. The female genitalia have often a glandular sac in sternal part or the tergal part of seg-ment VIII is gutter-like, hollowed, with a round process on the anterior margin. The host plant ofspecies belong to Elaeagnaceae, Rosaceae, Fabaceae, and Aceraceae.

Ponomarenko (1989) separated the genera Anarsia Zeller, 1839 and Ananarsia Amsel, 1959 andPark and Ponomarenko (2007) followed him, but auther treat the latter as a synonym of Anarsia,following Elsner et al. (1999) and Li (2002) in this book.

Key to the species of the genus Anarsia Zeller, based on the male genitalia

1. Valva nearly symmetric; socius large, fan-shaped; Aaedeagus nearly straight··························2- Valva asymmetric; socius small; Aaedeagus slender, curved·····················································3 2. Valva with membranous process on inner surfacen medially; cucullus semiovate·····A. gajiensis- Valva without membranous process on inner surface medially; cucullus more or less triangular·····························································································································A. ulneongensis

3. Valva asymmetric, but similar in length, without bandlike long process···································4- Valva asymmetric, left one broadly expanded and right one narrowed in middle, with but similar

in length, with bandlike long process·······················································································54. Valva with triangular, membranous process on inner surface; terminal processes of cucullus asym-

metric··················································································································A. asymmetrode - Valva without triangular, membranous process on inner surface; terminal processes of cucullus

symmetric················································································································A. bipinnata5. Left valva semiovate; cucullus poorly defined; bandlike process longer than valva, strongly

arched·······································································································································6- Left valva ovate; cucullus well defined; bandlike process shorter than valva, slightly arched

medially····················································································································A. lineatella6. Right cucullus with tongue-shaped process····························································A. bimaculata- Right cucullus without tongue-shaped process·········································································7 7. Length of left valva about twice of its width, dorsal surface of right cucullus nearly straight;

apex not well produced····························································································A. nigricana- Length of left valva longer than twice of its width, dorsal surface of right cucullus slightly arch-

ed; apex well produced························································································A. subnigricana

171. Anarsia bipinnata (Meyrick, 1932)

Chelaria bipinnata Meyrick, 1932, Exot. Microlep., 4: 200. Chelaria bipinnata: Gaede, 1937: 409; Clarke, 1969(6): 409.Anarsia bipinnata: Inoue, 1954: 69; Issiki, 1957: 43; Amsel, 1967: 25; Moriuti, 1982, I: 28, II: 214; Park,

1991b: 492; Park and Byun, 1995: 138; Park and Ponomarenko, 1996c: 74; Ueda, 1997: 82; Li, 2002:230.


Ananarsia bipinnata: Ponomarenko, 1997a: 51; 1999: 255; Park and Ponomarenko, 2007: 180.TL: Gifu, Japan.

ADULT: Wingspan 17-19 mm. Both sexes of this species resemble to A. euphorodes Meyrick or A.aspera Park which are known in China or Taiwan in the male genitalia, but it can be distinguishedfrom the latters by the apical process of cucullus and the shapes of the membranous lobes in themiddle of valva. It is also close to A. gajiensis Park and Ponomarenko.

Male genitalia (Figs. 171A, B). See also Amsel (1967, Pl. 8, Fig. 19); Park (1991b, Figs. 8, 9); Ueda(1997, Figs. 21b-c); Ponomarenko (1999, Figs. 153: 1, 2); Li (2002: Fig. 266); Park and Ponomarenko(2007, Pl. 66, Fig. 233). Uncus short, with a narrow, small hook-shaped process. Gnathos absent.Valva rather slender, both sides asymmetrical; left valva widely bulged at basal half, then narrow-ed distally, with a triangular plate; terminal portion widely expanded; right valva with straightcosta, almost even in width; terminal portion similar to left one. Juxta emarginate at middle, withshort lobes laterally. Aedeagus slender, strongly tapered near middle, narrowed toward apex.

Female genitalia (Figs. 171C, D). See also Amsel (1967, Pl. 8, Fig. 15); Clarke (1969, pl. 203, Fig. 2a,b); Ueda (1997, Fig. 21d); Li (2002, Fig. 267). Park and Ponomarenko (2007, Pl. 96, Fig. 233). Ostiumfunnel-shaped, posterior margin wide. Ductus bursae narrow, forming narrow stripe with finewrinkles along inner lateral surface to beyond middle. Corpus bursae large, with a small crescentsignum.


Fig. 171. Anarsia bipinnata. A. Male genitalia; B. Aedeagus; C. Female genitalia with corpusbursae.

A

B

C

SPECIMENS EXAMINED: [GG] 3♀, 2♂, Suweon, 29.vi.1976, 20.vi.1977, 5.vi.1981, 28.vii.1982, 15.vi.1990; 1♀, 3♂, Gwangleung, 27.vi.1986, 7.viii.1986, 17.vi.1994, 22.v.1998; 1♂, Isl. Daebudo, Ansan,28.vi.1997; 1♂, Isl. Deokjeokdo, 24.vi.1997; 1♂, Mt. Hwaya-san, 18.vii.1997; 1♂, Mt. Suri-san,Gunpo, 28.viii.1997; 2♀, 1♂, Isl. Yeongjongdo, Incheon, 31.iv.1995; 2♂, Mt. Cheonma-san, 6.v.1995;1♂, Mt. Soyo-san, 2.vi.1996; 1♂, Siheung, 30.viii.1996; [GW] 1♂, Chuncheon, 6.viii.1990; 1♂, Mt.Gubong-san, 8.vi.1998; 1♂, Yangyang, 4.vi.1987; 1♂, Mt. Taehwa-san, 27.v.1998; 1♂, Mt. Baekun-san, 5.viii.1994; 1♂, Mt. Odae-san, 26.vi.1989; 1♀, Mt. Jeombong-san, 13.vii.1997; [CB] 2♂, Mt.Weolak-san, Danyang, 7.vi.1997; 1♂, Mt. Ingyeong, Cheungju, 8.vi.1997; 1♂, Mt. Gaya-san, Yesan,15.vi.1997; [JB] 1♂, Muju, 13.viii.1975; 1♂, Mt. Obong-san, 4.vi.1985; 2♀, Mt. Yeogi-san, 5.vi.1985;2♂, Mt. Mireuk-san, 14.vi.1997; [JJ] 1♂, Seoguipo, 15.vi.1985; 2♀, 7♂, Oradong, 10.vi.1993; 3♂,Seonheul, 1.vii.1993, 1.viii.1993; 2♀, Siheung, near Hamdok, 11.viii.1993; 1♀, 3♂, Mt. Halla-san,5.vii.1986, 25.v.1988, 28.v.1992, 30.v.1987; 4♂, Mokseokweon, 30.v.1992; 1♀, 1♂, Seongpanak,23.viii.1992; 1♀, Bijarim, 15.vii.1993; 1♀, Temp. Gwanum-sa, 13.vii.1993; 1♂, Youngsil, 2.vii.1994.

DISTRIBUTION: Korea (Central, South, Jeju), Russian Far East (Primorsk), China (Shaanxi, Gansu,Qinghai, Anhui), Japan (Honshu, Izu Is., Kyushu).

HOST PALNTS: Elaeagnaceae- Elaeagnus umbellata. Asteraceae- Ageratum houstoniaum M., Acerginnala. Fagaceae- Quercus sp. (Park, 1991; Li, 2002).

172. Anarsia ulneongensis Park and Ponomarenko, 1996

Anarsia ulneongensis Park and Ponomarenko, 1996b, Kor. J. Ent., 26(4): 343.Ananarsia ulneongensis; Park and Ponomarenko, 2007: 182.


Fig. 172. Anarsia ulneongensis. A, B. Male genitalia with aedeagus.

A B

TL: Isl. Ulnungdo, Korea.

ADULT: Wingspan 15 mm. This species is similar to A. bipinata (Meyrick) in external and malegenital characters, but it differs by the following: more dense bunch of hairs at the base of the veinCu under surface of the forewing; hindwing lacking pectin on the base of R-M veins; stronglyemarginated between socii distally, valva without ventral narrow lobe on the triangular distal part,and stout, straight aedeagus in the male genitalia. Female is unknown.

Male genitalia (Figs. 172A, B). See also Park and Ponomarenko (1996, Fig. 4); Park and Ponoma-renko (2007, Pl. 67, Fig. 236). Similar to those of A. bipinata, but it differs from the latter by the tri-angulary expanded distal part without absence of apical spines, and the stouter aedeagus.

SPECIMENS EXAMINED: [GB] 2♂, Dodong, Isl. Ulnungdo, 200 km E. Donghae, 6.viii.1995. DISTRIBUTION: Korea (South), Japan.

173. Anarsia gajiensis (Park and Ponomarenko, 1996)

Anarsia gajiensis Park and Ponomarenko, 1996a, Act. Zool. hung., 42(1): 75.Ananarsia gajiensis: Ponomarenko, 1997: 52; Park and Ponomarenko, 2007: 180.

TL: Mt. Gaji-san, [GN], Korea.


Fig. 173. Ananarsia gajiensis. A, B. Male genitalia with aedeagus; C. Female genitalia with corpusbursae.

A C

B

ADULT: Wingspan 12-13 mm. This species closely resembles A. euphorodes Meyrick and A. asperaPark in the pattern of the forewing and the male genitalia, but can be distinguished by the mem-branous lobes on both valvae in the male genitalia and by the oval plate on the 8th tergite in thefemale genitalia.

Male genitalia (Figs. 173A, B). See also Park and Ponomarenko (1996, Fig. 2). Park and Ponoma-renko (2007, Pl. 66, Fig. 234). Distinguished from those of A. bipinata by the symmetric valva withheavily sclerotized, sharply extended apex of cucullus, and the symmetric membranous lobes inthe middle of valva. Aedeagus canal-shaped, shortly dilated basally, slightly narrowed towardsapex beyond 1/4 length.

Female genitalia (Fig. 173C). See also Park and Ponomarenko (1996, Fig. 3). Park and Ponoma-renko (2007, Pl. 96, Fig. 234). Ostium at same level with anterior margin of abdominal sternite VIII.Antrum goblet-shaped. Ductus bursae narrow, as long as corpus bursae. Corpus bursae oval, witha small crescent signum.

SPECIMENS EXAMINED: [GG] 5♀, 3♂, Isl. Yongyudo, Incheon, 11.vi.1997, 28.viii.1997; 1♂, Deok-jeokdo, 24.vi.1997; 1♂, Isl. Daebudo, Ansan, 30.viii.1997; [GW] 1♂, Munsanri, Pyoungchang, 7.vi.1996; 1♂, Mt. Jeombong-san, 5.viii.1997; [CN] 1♂, Muju, 13.viii.1995; [GB] 1♀, 1♂, Mt. Juwang-san, 18.viii.1993; [GN] 1♂, Mt. Gaji-san, 19.viii.1993, holotype; [JJ] 1♂, Seonheulri, 1.viii.1993; 1♂,Oradong, 10.vi.1993; 1♂, Temp. Gwanum-sa, 24.viii.1992.

DISTRIBUTION: Korea (Central, South, Jeju).

174. Anarsia asymmetrodes Park, 2013

Anarsia asymmetrodes Park, 2013. Ent. Res. 43: (in print).

ADULT: Wingspan, 14-15 mm. Head and thorax pale orange gray, speckling with dark-brownscales dorsally. Labial palpus of male (Fig. 2a): second segment with quadrate scale-tufts beneath,dark brown on outer surface, with yellowish-white dorsoanterior part, whitish on upper 1/3 andmixed with more dark-brown scales in lower part on inner surface; 3rd segment vestigial; female-second segment similar to that of male; 3rd segment well-developed, slender, as long as 2nd seg-ment, yellowish white with two dark-brown band at middle and beyond. Antenna orange gray;annulations brownish, paler towards apex. Forewing rather broader, gradually narrowed beyond3/4 length towards apex; apex more or less acute; ground color pale orange gray, speckling withdark-brown scales; basal short blackish streak on costa; median costal patch blackish, large, trape-zoidal, upper margin about 1.5 times of lower margin, inner margin very oblique, outer marginrounded, reaching about 1/4 distance across wing; 2-3 small, dark-brownish marks representedbefore and beyond median costal patch; a long hair-pencils well developed near base of discal cellunder surface of wing, longer than half length of forewing; venation with R1 arising from middleof cell, distance between R1 and R2 about twice of R2 and R3, R3 free, R4 and R5 stalked before middle,M1 free, nearly parallel to R4+5; distance M2 and M3 at base about 1.5 times of M3 and CuA1, CuA1

free, arising from near lower corner of cell; CuA2 free; fringe orange gray, more darker before apexand near tornus. Hindwing evenly grayish, broader than forewing, with broad anterior expansionextended to basal 3/5 of costa; Rs approximated to Sc, Rs and M1 short-stalked, M2 and M3 nearlyparallel; cubital pecten well-developed; apex obtuse; termen slightly sinuate; fringe grayish. Ventral


surface of fore and mid-legs dark brown; hind tibia with orange white hair-like scales above.Female is unknown.

Male genitalia (Fig. 174B). Uncus inflated medially, with hook-shaped apical process; sociistretched distally, with round caudal margin, bearing long hairs densely. Tegumen long, inflatedmedially, dilated towards base. Valvae asmymmetric; left valva with larger membranous medianprocess, cucullus with heavily sclerotized, elongated, distally rounded apical process; right valvawith shorter median rocess, cucullus with heavily sclerotized, spine-like apical process. Saccusmore or less triangular. Aedeagus as long as valva, taenioid, slightly inflated in median part, sharp-ly narrowed apically. The male genitalia are similar to those of A. largimacularis Li and Zheng, 1998,which was described on the basis of a male from Shaanxi, China, but it differ from the latter byhaving the uncus inflated medially, nearly same length of the valvae with well-developed medianprocess on dorsal margin of valva.

Female genitalia (Fig. 174C). Eighth segment sclerotized with a pair of processes as long as 1/3length of apophyses anteriores on anterior margin, posterior margin convex ventrally. Apophysesanteriores thick, as long as apophyses posteriors. Ostium plate heavily sclerotized, nearly triangu-lar. Corpus bursae semiovate, large, longer than ductus bursae, with dense particles on surface;signum a short transverse ridge, as wide as middle part of ductus bursae.

SPECIMENS EXAMINED: [GG] 1♂, 1♀, Is. Baengnyeong-do, 15.viii.2006.DISTRIBUTION: Korea (Central: Is. Baengnyeong-do).


Fig. 174. Anarsia asymmetrodes. A. Adult; B. Male genitalia with aedeagus; C. Female genitalia.

A

B C

175. Anarsia lineatella (Zeller, 1839)

Anarsia lineatella Zeller, 1839, Isis: 190; Gaede, 1937: 402; Amsel, 1967: 24; Liu et al., 1981: 18; Pisku-nov, 1981: 718; Karsholt and Riedl, 1996: 120; Elsner et al., 1999: 54; Park et al., 2000: 243; Li, 2002:237.

Anarsia (Ananarsia) lineatella: Ponomarenko, 1989: 637. Ananarsia lineatella heratella Amsel, 1967: 20.Ananarsia lineatella tauricella Amsel, 1967: 20.Ananarsia lineatella: Ponomarenko, 1997: 52; Ponomarenko, 2004: 56; Beljaev and Ponomarenko,

2005: 2; Park and Ponomarenko, 2007: 181.TL: Germany, Europe.

ADULT: Wingspan 11-14 mm. Head light gray. Forewing light gray, alternating white and grayscales; two dark, nearly black spots distinct at costal margin of wing; one in middle and other closerto base; gray stripe connecting three short transverse streaks extending from costal margin towardsapex, with longitudinal dark streaks on the light gray ground color.

Male genitalia (Figs. 175A, B). See also Amsel (1967, pl. 7, Fig. 9); Piskunov (1981, Figs. 654, 3, 4);Ponomarenko (1989, Fig. 13); Elsner et al. (1999, t. 37, Fig. 321); Park et al. (2000, Figs. 5, 6); Li (2002,Fig. 274); Beljaev and Ponomarenko (2005, Figs. 1, 2): Park and Ponomarenko (2007, Pl. 67, Fig. 235).Uncus with small median pointed knob; cucullus inflated: left one in proximal 2/3 part and right


Fig. 175. Anarsia lineatella. A. Male genitalia with aedeagus; B. Aedeagus; C. Female genitalia withcorpus bursae.

A C

B

one in basal part and at 2/3 of length, every with strong sclerotized long process on ventral margin.Female genitalia (Fig. 175C). See also Amsel (1967, pl. 10, 26; Ponomarenko (1989, Fig. 14); Elsner et

al. (1999, t. 79, Fig. 32); Li (2002, Fig. 275); Beljaev and Ponomarenko (2005: Fig. 3). Park and Pono-marenko (2007, Pl. 96, Fig. 235). Apophyses anteriores very short, slightly entering segment VII,ends directed dorsally; apophyses posteriors terminating before bent anterior margin of tergiteVIII. Signum forming irregular rectangle, anterior and posterior margins concave.

SPECIMENS EXAMINED: [N. Korea] 1♂, Duman River, 31.vii.2000.DISTRIBUTION: Korea (North), Russia (European part, S. Ural, Primorsk), China (Shaanxi, Xinjiang),

Europe; Caucasus; Transcaucasien region; Central Asia, N. Africa, Asia Minor, Iran, Afghanistan,India, N. America, Australia.

HOST PALNTS: Rosaceae- Prunus domestica, P. Spinosa, Prunus spp., Malus sylvestris, Malus spp.,Armeniaca vulgaris, Persica vulgaris, Eriophyes phloecoptes, Amygdalus communis, Amygdalus spp.Ebenaceae- Diospyros spp. Aceraceae- Acer campestre, A. tataricum (Piskunov, 1981; Elsner et al.,1999).

REMARKS: The species is one of the quarantined pests and it was first reported from the Koreanpeninsula by Park et al. (2000). It was collected at Chongshan, a small village at border of Chinaand N. Korea, but the moth flewed from the Korean side of Duman-River because the light trap wasplaced towards a small mountain in the N. Korean side, just across the river.

176. Anarsia bimaculata Ponomarenko, 1989

Anarsia bimaculata Ponomarenko, 1989, Entomol. Obozr., 68(3): 635. Anarsia bimaculata: Park, 1991b: 496; Ponomarenko, 1997: 54; Ueda, 1997: 86; Ponomarenko, 1999:

257; Ponomarenko et al., 2006: 113; Park and Ponomarenko, 2007: 183.TL: Gornotajezhnoe, Ussuri, Primorsk Territory, Russian Far East.

ADULT: Wingspan 14-15 mm. This species is distinguished from the following two species bythe right cucullus of the male genitalia, which has a tongue-shaped lobe on the ventral margin ofdilated distal part beyond the slender sclerotized process.

Male genitalia (Figs. 176A-C). See also Ponomarenko (1989, Figs. 19, 20); Park (1991b, Figs. 16, 17);Ueda (1997, Figs. 23b, c); Ponomarenko (1999, Figs. 153, 5, 6). Park and Ponomarenko (2007, Pl. 67,Fig. 237). Similar to those of A. nigricana Park, but can be distinguished by following characters:Tegumen without distinct dilation laterally; left valva broader at base, band-like slender processarsing from ventral margin; right valva with with tounge-shaped anal projection in distal portionventrally; juxta with shorter laterocaudal lobes.

Female genitalia (Fig. 176D). See also Ponomarenko (1989, Fig. 21); Ueda (1997, Fig. 23d); Ponoma-renko (1999, Fig. 154, 3). Park and Ponomarenko (2007, Pl. 96, Fig. 237). Ostium near anterior mar-gin of sternite VIII and ostial funnel shifted dextrally. Anterior margin of tergite VIII with processuniformally narrowed towards apex, margins turned inward; tergite with opening in center withmargins turned inward.

SPECIMENS EXAMINED: [GG] 3♀, 1♂, Suweon, 20.vi.1977, 24.v.1997; [GW] 1♂, Mt. Odae-san,6.viii.1989; 2♀, Jiamri, Chuncheon, 7.vi.1997, 13.vi.1997; 1♂, Inje, 23.vi.1996; 1♂, Yangyang, 25.vii.


1987; [JJ] 1♀, Seongpanak, 3.vii.1994. DISTRIBUTION: Korea (Central, Jeju), Russian Far East (Primorsk), China (Jilin), Japan (Hokkaido,

Honshu).HOST PLANT: Leguminosae- Maackia amurensis (Ponomarenko, 1989; Park and Ponomarenko, 2007).

177. Anarsia nigricana Park, 1991

Anarsia nigricana Park, 1991, Jpn J. Ent., 59(3): 494. Anarsia nigricana: Ponomarenko, 1997: 55; Park and Ponomarenko, 2007: 184.

TL: Suweon, GG, Korea.

ADULT: Wingspan 16-18 mm. This species is distinguished from A. bimaculata by the right cucul-lus of the male genitalia, which has a tongue-like lobe on the ventral margin of dilated distal partbeyond the slender sclerotized process; and also distinguished from the following species, A. sub-nigricana by the dorsal margin not hollowed before apex.

Male genitalia (Fig. 177A). See also Park (1991, Figs. 12, 13); Park and Ponomarenko (2007, Pl. 67,Fig. 238). Uncus narrow, with pointed apex. Tegumen long, narrowed towards distal end, dilatedat distal 3/4 length. Valva asymmetrical: left valva nearly semiovate, slightly angled near apex;right valva broadened at basal 1/5, forming narrow neck, then gradually widened towards distal


Fig. 176. Anarsia bimaculata. A, C. Male genitalia; B. Aedeagus; D. Female genitalia with corpusbursae.

A B

CD

end. Juxta with two caudal lobes laterally. Saccus short. Aedeagus slender, basal 1/3 dilated, andthen narrowed, tapered apically.

Female genitalia (Fig. 177B). See also Park (1991, Fig. 15); Park and Ponomarenko (2007, Pl. 96, Fig.238). Ostium moderate, not very widened distally. Ductus bursae narrow, nearly three times longerthan length of corpus bursae. Corpus bursae ovate, with a samll crescent signum.

SPECIMENS EXAMINED: [GG] 1♂, Suweon, 7.vii.1976: 2♂, Suweon, 7.vi.1976, 29.vi.1976; 2♂, Suweon,20.vi.1977; 1♀, Suweon, 12-13.vii.1977; [GW] 1♀, Mt. Chiak-san, 23.vi.1977.

DISTRIBUTION: Korea (Central).HOST PLANT: Leguminosae- Glycine max (Park, 1991b).

178. Anarsia subnigricana Park and Ponomarenko, 1996

Anarsia subnigricana Park and Ponomarenko, 1996b, Kor. J. Ent., 26(4): 344; Park and Ponomarenko,2007: 184. TL: Hongcheon, Korea.


Fig. 177. Anarsia nigricana. A. Male genitalia with aedeagus; B. Aedeagus; C. Female genitalia withcorpus bursae.

A B

ADULT: Wingspan 14-15 mm. This species is distinguished from the preceding species, A. nigri-cana by the right cucullus of the male genitalia which has a protrusion on dorsal margin and hol-lowed before apex.

Male genitalia (Fig. 178A). See also Park and Ponomarenko (1996, Figs. 5, 5a-b); Park and Pono-marenko (2007, Pl. 67, Fig. 239). Uncus with long, hook-shaped process, with acute apex. Valvaasymmetrical: left valva narrow, dilated at base and 3/4, with a long narrow process at 2/3 onventral margin; right valva with angular distal corner, with a long S-shaped process, arising fromnear 1/3 on outer surface. Aedeagus curved dorsally at middle and to right before apex, with twosclerotized ventral plates.

Female genitalia (Fig. 178B). See also Park and Ponomarenko (1996, Figs. 6, 6a); Park and Ponoma-renko (2007, Pl. 96, Fig. 239). Antrum arising laterally. A long, broad gutter-like sclerite on tergiteVIII. Ductus bursae extremely narrow, long, about four times length of corpus bursae. Corpusbursae with a small signum.

SPECIMENS EXAMINED: [GG] 1♂, Suweon, 20.vi.1977; 1♂, Mt. Suri-san, 15.vi.1990; [GW] 1♂, Jiamri,Chuncheon, 7.vi.1997; 1♂, Hongcheon, 15.vi.1989.

DISTRIBUTION: Korea (Central).


Fig. 178. Anarsia subnigricana. A. Male genitalia with aedeagus; B. Female genitalia with corpusbursae.

A

B

Additional Records

Subfamily Gelechiinae

179. Gnorimoschema streliciellum (Herrich-Schäffer, 1854)

Gelechia streliciellum Herrich-Schäffer, 1854, Schmett. Europe, 5: 171.Gnorimoschema streliciellum: Povolny, 1966: 393-395; Emeljanov and Piskunov, 1982: 377; Povolny,

1992: 230-232; Elsner, Huemer and Tokar, 1999: 41.Povolny, 2002: 25.

TL: Europe.

ADULT: Wingspan 13 mm. The species is similar to G. valesiella (Staudinger) which is distributedin Europe, from central to North, and Alaska, in external and male genital characters. The later has


Fig. 179. Gnorimoschema streliciellum. A. Adult; B. Labial palpus; C. Male genitalia; D. aedeagus; E.Abdomen.

A B

C D E

darker wing with only few red-brown scales (pers. comm. Karsholt, 2012). It is distributed inEurope and extended to Mongolia, Transbaikal, and Amur territory. Four subspecies have beenknown: the nominate subspecies in northern Europe; subsp. hoefneri (Rebel, 1909) in northern Italy;subsp. mongolorum Povolny, 1969 in Mongolia; subsp. cinctipunctellum (Erschoff, 1877) in Amur.The Korean species is treated as subsp. cinctipunctellum (Erschoff, 1877) in the zoogeographical view,but the subspecific status will be discussed in a future study when more material is available,because there are some differences in the male genitalia as followings.

Male genitalia (Figs. 179C, D). As compared with those of subsp. cinctipunctellum (Erschoff), theapical part of valva is less dilated; saccus broadly rounded anteriorly; aedeagus without preapicalspine. Uncus shortly developed, with triangularly acuted apex. Tegumen narrowed towards apex,weakly sclerotized. Valva broad basally, then narrow, elongate, gently curved inwardly; apex blunt-ed; basal process strongly curved at apex. Vinculum extremely broad. Saccus well-developed, aslong as tegumen. Aedeagus globular at basal 1/3, then elongate, as long as valve.

SPECIMENS EXAMINED: [GW] 1♂, Mt. Bannon-san, 37°37′N-128°45′E, Jeongseon, 14.ix.2010.DISTRIBUTION: Korea (Central), Amur, Mongolia, Europe.REMARKS: Genus Gnorimoschema Busk is a Holarctic genus, and more than 20 species are known

in the Palaearctic Region. In Korea, a single species, G. radkovitchi Piskunov has been known. G. streliciellumis is the species which was listed as unidentified species, Gnorimoschema sp. in thepart I.

180. Stenolechia kodamai Okada, 1962

Stenolechia kodamai Okada, 1962, Publs. Ent. Lab. Univ. Osaka Pref., 7: 32; Moriuti, 1982, part 1: 278,part 2: 213.TL: Japan.

ADULT (Fig. 180A): Wingspan, 10 mm. This species is superficially similar to S. bathrodyas Mey-rick, but slightly larger. It can be distinguished by the male genitalia: the shorter process of valva,about 1/2 length of tegumen, whereas nearly same length in S. bathrodyas.

Male genitalia (Figs. 180B-E). See Okada (1962, Figs. 19-21). Uncus spatulate; caudal marginslightly emmarginate medially. Ganthos well-developed, with large median process. Tegumenlong, about 4 times longer than uncus. Valva slender, about 1/2 length of tegumen. Process of juxtaheavily sclerotized, curved, longer than 1/2 length of aedeagus. Aedeagus long, taenioid, sharplyacuted apically.

Female genitalia. See also Okada (1962, Figs. 29-31). As figured, posterior part of ductus bursaenarrowed; antrum sclerotized. Anterior part of ductus bursae and corpus bursae missed.

SPECIMENS EXAMINED: [GG] 7♂, Is. Yeonpyeong-do, 37.40N-125.42E, 31.viii.2010.DISTRIBUTION: Korea (Central), Japan.


181. Bryotropha similis (Stainton, 1854)

Gelechia similis Stainton, 1854, Ins. Brit. Tin.: 115. Gelechia thuleella Zeller, 1857, Ent. Ztg. Stett, 18: 276.Gelechia similella Doubleday, 1859, Zool. Syn. List. Br. Lepid: 30.Gelechia pullifimbriella Clemens, 1863, Pro. Ent. Soc. Philad., 2: 120.Gelechia confinis Stainton, 1871, Ent. Annua: 98.Gelechia obscurecinerea Nolcken, 1871, Fauna Livld., 2: 573.Gelechia stolidella Morris, 1872, Horris Brit. Moths, pl. 108, fig. 1.Gelechia fuliginosella Snellen, 1882, Vlind. V. Nederld., 2(2): 645.Duvita tahavusella Forbes, 1922, Ent. News, 33: 103.Gelechia clandestina Meyrick, 1923, Exot. Microlepid., 3: 19.Bryotropha dufraneella Joannis, 1928, Bull. Soc. Ent. Fr., 1928: 195.Bryotropha novisimilis Li and Zheng, 1997, Acta Zootax. Sinica, 22(4): 398.


Fig. 180. Stenolechia kodamai. A. Adult; B. Male genitalia; C. Male genitalia; D. Close-up uncus; E.Close-up aedeagus.

A D

B C E

Bryotropha similis: Pierce and Metcalfe, 1935: 10; Svensson, 1962: 63; Piskunov, 1981: 714; Karsholtand Riedl, 1996: 109; Elsner et al., 1999: 31; Rutten, 1999: 88; Bland et al., 2002: 110; Li, 2002: 56;Karsholt and Rutten, 2005: 141; Park and Ponomarenko, 2007: 49.TL: Surrey, England.

ADULT: Wingspan, 11 mm. Superficially similar to B. affinis (Haworth) and B. senectella (Zeller),but can be distinguished by the male genitalia. This species was reported from Mt. Changbai-san,at border between the Korean peninsula and N. China (Park and Ponomarenko, 2007). The shapeof the signum in the female genitalia shows a bit difference from that of the European specimen,but it is tentatively identified as B. similis. A further critical examination of the male genitalia isneeded, comparing with those of the European specimen, when the male specimen is available.

SPECIMENS EXAMINED: [GG] 1♀, Isl. Decheong-do, 14.viii.2006.DISTRIBUTION: Korea (Central): New record, RFE, China, Japan, Europe.REMARKS: This species is reported for the first time from Korea.


Fig. 181. Bryotropha similis. A. Adult; B. Female genitalia (Corpus bursae missed).

A B

Literature Cited

Amsel HG (1939 (1938)) In Hartig and Amsel. Contributo alla conoscenza della fauna entomologica dellaSardegna. Noove forme di Lepidopteri. Mem. Soc. Ent. Ital. 17: 72

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132

Plates

106 107

108 109

110 111

112 113

Plates 133

114 115

116 117

118 119

120 121


122 123

124 125

126 127

128 129

Plates 135

130 131

132 133

134 135

136 137


138 139

140 141

142 143

144 145

Plates 137

146 147

148 149

150 151

152 153


154 155

156 157

158 159

160 161

Plates 139

162 163

164 165

166 167

168 169


170 171

172 173

174 175

176 177

Plates 141

178 179

180 181

A

Altenia 14inscriptella 15

Anacampsinae 16Anacampsini 16Anacampsis 20

anisogramma 20lignaria 21solemnella 23

Anarsia 102bimaculata 110bipinnata 103gajiensis 106lineatella 109nigricana 111asymmetrodes 107 subnigricana 112ulneongensis 105

Anarsiini 102Aproaerema 18

anthyllidella 18

B

Bagdadia 99claviformis 99eucalla 100gnomia 101

Bryotropha 116simils 116

C

Chelariini 70Concubina 13

trigonalis 13

D

Dendrophilia 93leguminella 94

mediofasciana 95neotaphronoma 96unicolorella 98

Dichomeridinae 30Dichomeridini 31Dichomeris 37

anisacuminata 67atomogypsa 39autometra 40chinganella 41christophi 43cuspis 44derasella 45harmonias 47heriguronis 48horoglypta 50issikii 51japonicella 68lespedezae 52litoxyla 53lutilinea 54minutia 55mitteri 57oceanis 58polypunctata 59praevacua 60rasilella 61sparsella 63tostella 64ustalella 65

E

Encolapta 71catarina 72subtegulifera 73tegulifera 74

F

Faristenia 81acerella 82

142

Index to Scientific Names

atrimaculata 84furtumella 85geminisignella 86jumbongae 87maritimella 88omelkoi 89quercivora 91ussuriella 92

G

Gelechiinae 9Gnorimoschema 114

streliciellum 114

H

Helcystogramma 31fuscomarginatum 32ineruditum 33perelegans 34triannulella 35

Hypatima 75excellentella 76venefica 77

M

Mesophleps 26acutunca 29

albilinella 26sublutiana 28

N

Neofaculta 70taigana 70

P

Prolita 24sexpunctella 24

Pseudotelphusa 11acrobrunella 11

S

Stenolechia 115kodamai 115

Syncopacma 16wormiella 17

T

Teleiopsis 9Motleella 9

Tornodoxa 79longiella 79tholochorda 80

Index to Scientific Names 143

China

PB

JG

YG

HB

HN

PN

HWB

GW

GG

CB

CNGB

GB(Ulleung-do)

GN

JN

JJ

JB

HWN

Russia

East Sea

Yellow Sea

South Sea

CB Chungcheongbuk-doCN Chungcheongnam-doGB Gyeongsangbuk-doGG Gyeonggi-doGN Gyeongsangnam-doGW Gangwon-doHB Hamgyeongbuk-doHN Hamgyeongnam-doHWB Hwanghaebuk-doHWN Hwanghaenam-doJB Jeollabuk-doJG Jagang-doJJ Jeju-doJN Jeollanam-doPB Pyeonganbuk-doPN Pyeongannam-doYG Yanggang-do


Flora and Fauna of Korea



Gelechiidae II

Insect Fauna of KoreaVol. 16, N

o. 9G

elechiidaeII

NIB

R


Documents

Insect Fauna of Koreawebbook.me.go.kr/DLi-File/099/006/005/5580279.pdf · Insect Fauna of Korea Kyu-Tek Park The Korean Academy of Science and Technology Volume 16, ... any form or