Inventory of Paleozoic radiolarian species (1880-2016)...Paleozoic radiolarians. Here we present a comprehensive list of all the Paleozoic radiolarian species of which we are aware

533GEODIVERSITAS bull 2017 bull 39 (3) copy Publications scientifiques du Museacuteum national drsquoHistoire naturelle Paris wwwgeodiversitascom

Jonathan C AITCHISON School of Earth and Environmental Sciences

University of Queensland St Lucia Qld 4072 (Australia)jonauqeduau (corresponding author)

Noritoshi SUZUKIDepartment of Earth Science Graduate School of Science

Tohoku University 6-3 Aoba Aramaki Aoba-ku Sendai 980-8578 (Japan)norinorimtohokuacjp

Luis OrsquoDOGHERTYFacultad de Ciencias del Mar Universidad de Cadiz

E-11510 Puerto Real (Spain)luisodoghertyucaes

Published on 29 September 2017

Inventory of Paleozoic radiolarian species (1880-2016)

MOTS CLEacuteSPaleacuteozoiumlque

radiolairespolycystines

inventaire

KEY WORDS Paleozoic

radiolarians polycystina

inventory

urnlsidzoobankorgpub9072CEF3-2F28-415D-A239-24E6A55DC3E6

Aitchison J C Suzuki N amp OrsquoDogherty L 2017 mdash Inventory of Paleozoic radiolarian species (1880-2016) in Danelian T Caridroit M Noble P amp Aitchison J C (eds) Catalogue of Paleozoic radiolarian genera Geodiversitas 39 (3) 533-637 httpsdoiorg105252g2017n3a6

ABSTRACTThis paper presents for the first time a complete list of Paleozoic radiolarian species (mainly Poly-cystina) described between 1880 and early 2016 It records 2323 names of taxa described originally as new species or subspecies that have been assigned to radiolarians This list attempts to only pro-vide an objective record without revision of taxonomic status and thus includes invalid taxa junior objective or subjective synonyms nomina dubia and nomina nuda

REacuteSUMEacuteInventaire des espegraveces de radiolaires du Paleacuteozoiumlque (1880-2016)Cet article preacutesente pour la premiegravere fois une liste complegravete drsquoespegraveces de radiolaires du Paleacuteozoiumlque (principalement des polycystines) deacutecrites entre 1880 et deacutebut 2016 Il contient 2323 noms de taxons deacutecrits agrave lrsquoorigine comme des nouvelles espegraveces ou sous-espegraveces affecteacutees aux radiolaires Cette liste vise donc agrave en fournir un eacutetat objectif sans reacutevision de leur statut taxonomique et comprend donc des taxons valides des synonymes juniors objectifs ou subjectifs des nomina dubia et nomina nuda

534 GEODIVERSITAS bull 2017 bull 39 (3)

Aitchison J C et al

OBJECTIVES

Following the publication of the Inventory of Mesozoic radiolarian species (OrsquoDogherty 2009) there was general agreement of participants in workshops held by the Paleozoic Working Group that it would be useful for the community of researchers if a similar document could be produced for Paleozoic radiolarians

Here we present a comprehensive list of all the Paleozoic radiolarian species of which we are aware that have been described between the late 1800s and early 2016 The inten-tion is to provide in a single file the names of all Paleozoic species that have been published All names listed are the original names given in the original publication and do not reflect any subsequent taxonomic revision or evalua-tion as to their validity and assignment at higher taxonomic levels The list records 2323 taxa described originally as either new species or subspecies Although every attempt has been made to be as complete as possible it is perhaps inevitable that some taxa may be missing The purpose of this list is to provide information on 1) the original Linnean combination used when a species was erected as new in Paleozoic material 2) the author(s) and page(s) on which it is described 3) the age of the holotype or of the name-bearing type as originally published (note in some cases there may have been considerable revision of the original age determination) and 4) the order to which the species or sub-species belongs (the numbers of which are indicated in parentheses with each order) Al Albaillellaria (363) Ar Archaeospicularia (154) E Entactinaria (726) L Latentifistularia (194) N possible Nassellaria (111) S Spumellaria (207) IS incertae sedis within orders (72) incertae sedis in both order and family (35) with 368 names that can be regarded as nomina dubium (nd) and 158 names introduced as nomina nuda (nn) Forty-four taxa are clearly not radiolarians (nr) with 15 of these Calcarea and five others that can be assigned as sponge spicules (ss) also identified Eleven names are invalid Type species (T valid type species Td types that are nomina dubia Th types that are homonyms Ti type species that are invalid names and Ts types that are synonyms) are indicated

COMPILATION OF SPECIES NAMES

The list of taxa has been compiled following a detailed search of gt1250 publications dealing with Paleozoic radiolarians of which 259 introduce new taxa Synonyms and species descriptions are not included because this is beyond the scope of the project and would require several years of work by many research groups The general agreement of participants at the workshops was that it would be useful to take this opportunity to make a step towards address-ing species-level taxonomy of species once the revision of genera has been fully achieved Such a work would have the additional benefit of providing a better-defined starting point for future taxonomists

FUTURE WORK

Objective evaluation of each species with respect to its taxo-nomic status and the subjective assignment to valid genera is the next step towards attaining a full proper standard of taxonomy and stratigraphy for Paleozoic radiolarians Evalu-ation of each species unless one is personally familiar with it is a time-consuming task depending on the availability of literature and given that there are 2323 species names to be evaluated is well beyond the scope of this particular work which is meant to focus on the genera However we empha-size that the species treatment is a logical future task of the Paleozoic Working Group It is our intention herein to simply provide this list as a useful starting point for any such study

AcknowledgementsWe thank all our fellow radiolarian researchers who have in various ways contributed to the accumulation of the body of data that has allowed development of this biozonation JA thanks Sarah Kachovich for her help in editing this list and he also acknowl-edges financial support towards investigation of Early Paleozoic radiolarian evolution in the form of a grant from the Australian Research Council (ARC DP 1501013325) We also gratefully acknowledge reviews by Taniel Danelian and Paula Noble

REFERENCES

Aberdeen e 1940 mdash Radiolarian fauna of the Caballos Formation Marathon Basin Texas Journal of Paleontology 14 (2) 127-139 httpwwwjstororgstable1298566

AfAnAsievA M s 1986 mdash Radiolarians of the family Pylenton-emidae Paleontological Journal 3 22-34

AfAnAsievA M s 1993 mdash New data on the Early Paleozoic radio-larian genus Caspiaza Paleontological Journal 4 115-118

AfAnAsievA M s 1997a mdash Early Frasnian Radiolarians and their biostrati-graphical significance Doklady Akademii Nauk 355 (2) 217-222

AfAnAsievA M s 1997b mdash Moscovistella victoralis the victorious star of Moscow Izvestija Vysshikh Uchebnykh Zavedeniy Geologiya i Razvedka 5 38-41

AfAnAsievA M s 2000a mdash Atlas of Paleozoic Radiolaria of the Rus-sian Platform Scientific World Moscow 477 p

AfAnAsievA M s 2000b mdash New radiolarians of the superfam-ily Entactinoidea from the Upper Devonian of Timan-Pechora Province Russia Paleontological Journal 34 (2) 131-146

AfAnAsievA M s 2000c mdash New radiolarians of the orders Aculearia and Sphaerellaria from the Upper Devonian of the Timan-Pechora Province (Russia) Paleontological Journal 34 (4) 359-376

AfAnAsievA M 2011 mdash Revision of the genus Entactinosphaera Foreman 1963 and the new genus Retientactinosphaera gen nov (Paleozoic radiolaria) Paleontological Journal 45 (2) 117-129 httpsdoiorg101134S003103011102002X

AfAnAsievA M s amp AMon e 2008 mdash New radiolarians from the Devonian of the southern Ural Mountains 1 Early-Middle Devonian (Late Emsian-Early Eifelian) Paleontological Journal 42 (2) 451-467

AfAnAsievA M s amp AMon e o 2009a mdash New Radiolarians from the Devonian of the Southern Ural Mountains 2 MiddlendashLate Devonian Paleontological Journal 43 (1) 34-49

AfAnAsievA M s amp AMon e o 2009b mdash Evolution of Devonian biodiversity of radiolarians with two porous spheres and one main spine Paleontological Journal 43 (5) 483-498 httpsdoiorg101134S0031030109050037



AfAnAsievA M s amp AMon e o 2011 mdash Devonian Radiolarians of Russia Paleontological Journal 45 (11) 1313-1532 httpsdoiorg101134S0031030111110013

AfAnAsievA M s amp AMon e o 2012 mdash New Middle Devonian radiolarians from western Mugodzhary Paleontological Journal 46 (6) 549-551 httpsdoiorg101134S0031030112060020

AfAnAsievA M s amp AMon e o 2016 mdash New radiolarian genera and species from the Lower Permian of the Southern Urals and Northern Mugodzhary Paleontological Journal 50 (3) 209-221 httpsdoiorg101134S0031030116020027

Aitchison J c 1993a mdash Albaillellaria from the New England orogen Eastern NSW Australia Marine Micropaleontology 21 (4) 353-367 httpsdoiorg1010160377-8398(93)90026-T

Aitchison J c 1993b mdash Devonian (Frasnian) Radiolarians from the Gogo Formation Canning Basin Western Australia Palaeonto-graphica Abteilung A Paleozoologie-Stratigraphie 228 (4-6) 105-128

Aitchison J c 1998 mdash A Lower Ordovician (Arenig) radiolarian fauna from the Ballantrae Complex Scotland Scottish Journal of Geology 34 (1) 73-81 httpsdoiorg101144sjg34010073

Aitchison J c hAdA s irelAnd t amp YoshikurA s 1996 mdash Ages of Silurian radiolarians from the Kurosegawa terrane South-west Japan constrained by UPb shrimp data Journal of Southeast Asian Earth Sciences 14 (1-2) 53-70 httpsdoiorg101016S0743-9547(96)00045-1

Aitchison J c flood P G amp MAlPAs J 1998 mdash Lowermost Ordovician (basal Tremadoc) radiolarians from the Little Port Complex western Newfoundland (Lower Ordovician radiolar-ians Newfoundland) Geological Magazine 135 (3) 413-419 httpsdoiorg101017S001675689800867X

Aitchison J c dAvis A M strAtford J M c amp sPiller f c P 1999 mdash Lower and Middle Devonian radiolarian biozonation of the Gamilaroi terrane New England Orogen eastern Australia Micropaleontology 45 (2) 138-162 httpsdoiorg1023071486110

AMon e o amp brAun A 1994 mdash Radiolarians from lower Per-mian deposits of the Belskaya depression Bashkiria (west slope of southern Urals Artinskian stage Burtsevsky horizon) in nAirn A e M amp koroteev A V (eds) Contributions to Eurasian Geology Permian Conference Papers Occasional Publications ESRI Columbia USA 9 1-7

AMon e o brAun A amp chuvAshov b i 1990 mdash Lower Per-mian (Artinskian) Radiolaria from the Sim type section Southern Urals Geologica et Palaeontologica 24 115-137

AMon e o brAun A amp ivAnov k s 1995 mdash Upper Silurian radio-larians from the southern Urals Geologica et Palaeontologica 29 1-18

belYAnskY G s nikitA A P amp rudenko v s 1984 mdash About Sebuchaz suite of Primorye in PoYArkovA Z N (ed) New Data an Detail Biostratigraphy of Phanerozoic of Far East DVNC Akademii Nauk SSSR Vladivostok 43-57

benGtson s 1986 mdash Siliceous microfossils from the Upper Cam-brian of Queensland Alcheringa 10 (3) 195-216 httpsdoiorg10108003115518608619155

boundY-sAnders s Q sAndberG A A MurcheY b l amp hAr-ris A G 1999 mdash A late Frasnian (Late Devonian) radiolarian sponge spicule and conodont fauna from the Slaven Chert northern Shoshone Range Roberts Mountains allochton Nevada Micropaleontology 45 (1) 62-68 httpsdoiorg1023071486203

brAun A 1989 mdash Neue unterkarbonische Radiolarien-Taxa aus Kieselschiefer-Gerollen des unteren maintales be Frankfurt a M Geologica et Palaeontologica 23 83-99

brAun A 1990 mdash Radiolarien aus dem Unter-Karbon Deutschlands Courier Forschungs Institut Senckenberg 133 1-177

brAun A MAAss r amp schMidt-effinG r 1992 mdash Oberdevonische Radiolarien aus dem Breuschtal (Nord-Vogesen Elsas) und ihr regionaler und stratigraphischer Zusammenhang Neues Jahrbuch fuumlr Geologie und Palaumlontologie Abhandlungen 185 (2) 161-178

bYkovA e v 1955 mdash Foraminifery i radiolyarii devona volgo-uralskoi oblasti i tsentralnogo devonskogo polya i iz znachenie dlya stratigrafii

[Foraminifera and radiolarians of the Devonian of the Volga-Ural area] Proceedings of the All Union Petroleum Scientific Research Institute for Geological Survey (VNIGRI) new series 87 5-189

cAridroit M amp de Wever P 1984 mdash Description de quelques nouvelles espegraveces de Follicucullidae et drsquoEntactinidae (Radiolaires polycystines) du Permien du Japon Geobios 17 (5) 639-644 httpsdoiorg101016S0016-6995(84)80035-2

cAridroit M amp de Wever P 1986 mdash Some late Permian radio-larians from pelitic rocks of the Tatsunoformation (Hyogo Pre-fecture) Southwest Japan Marine Micropaleontology 11 55-90 httpsdoiorg1010160377-8398(86)90005-8

cAridroit M dAneliAn t orsquodoGhertY l cuvelier J Aitch-ison J c Pouille l noble P duMitricA P suzuki n kuWAhArA k MAletz J amp fenG Q 2017 mdash An illustrated catalogue and revised classification of Paleozoic radiolarian genera in dAneliAn t cAridroit M noble P amp Aitchison J C (eds) Catalogue of Paleozoic radiolarian genera Geodiversitas 39 (3) 363-417 httpsdoiorg105252g2017n3a3

cAtAlAno r di stefAno P amp kozur h 1989 mdash Lower Permian Albaillellacea (Radiolaria) from Sicily and their stratigraphic and paleogeographic significance Rendiconto dellrsquoAccademia delle Scienze fisiche e matematiche Serie IV 56 1-24

chAPMAn f 1923 mdash Report on fossils from an Upper Cambrian horizon at Loyola near Mansfield Bulletin of the Geological Survey of Victoria 46 34-45

chenG Y-n 1986 mdash Taxonomic Studies on Upper Paleozoic Radio-laria National Museum of Natural Science Taiwan Special Publication vol 1 310 p

conWAY Morris s amp chen M 1990 mdash Blastulospongia pol-ytreta nsp an enigmatic organism from the Lower Cambrian of Hubei China Journal of Paleontology 64 (1) 26-30 httpsdoiorg101017S0022336000042207

cordeY f 1998 mdash Radiolaires des complexes drsquoaccreacutetion de la Cordillegravere Canadienne (Colombie-Britannique) Geological Survey of Canada Bulletin 509 1-209

cornell W c amp siMPson r d 1985 mdash New Permian albail-lellid radiolarians from West Texas Micropaleontology 31 (3) 271-279 httpsdoiorg1023071485545

cornell W c amp siMPson r d 1986 mdash Nabespecha leonar-dia n gen n sp An unusual radiolarian from the Permian of West Texas Micropaleontology 32 (3) 286-288 httpsdoiorg1023071485623

dAneliAn t amp floYd J d 2001 mdash Progress in describing Ordovi-cian siliceous biodiversity from the Southern Uplands (Scotland UK) Transactions of the Royal Society of Edinburgh Earth Sciences 91 (3-4) 489-498 httpsdoiorg101017S0263593300008336

dAneliAn t amp PoPov l 2003 mdash Ordovician radiolarian biodi-versity insights based on new and revised data from Kazakhstan Bulletin de la Socieacuteteacute geacuteologique de France Seacuterie VIII 174 (4) 325-335 httpsdoiorg1021131744325

dAneliAn t PoPov l e tolMAchevA t Y GhobAdi Pour M G neYevin A v amp MikolAichuk A v 2011 mdash Ordovician radiolaria and conodonts from the peri-Gondwanan Karatau-Naryn microplate (Sarydzhaz eastern Kyrgyzstan) Geobios 44 (6) 587-599 httpsdoiorg101016jgeobios201102006

de Wever P amp cAridroit M 1984 mdash Description de quelques nouveaux Latentifistulidae (radiolaires polycystines) Paleacuteozoiumlques du Japon Revue de Micropaleacuteontologie 27 (2) 98-106

deflAndre G 1952 mdash Albaillella nov gen radiolaire fossile du Carbonifegravere infeacuterieur type drsquoune ligneacutee aberrante eacuteteinte Comptes Rendus hebdomadaires des Seacuteances de lrsquoAcadeacutemie des Sciences (Paris) Seacuterie D Sciences naturelles 234 872-874 httpgallicabnffrark12148bpt6k3186w

deflAndre G 1953 mdash Radiolaires fossiles in GrAsseacute P P (ed) Traiteacute de Zoologie Masson Paris 389-436

deflAndre G 1955 mdash Palaeocryptidium ng cayeuxi nsp microor-ganismes incertae sedis des phtanites brioveacuteriens bretons Comptes Rendus sommaire de la Socieacuteteacute geacuteologique de France 9-10 182-185


Aitchison J C et al

deflAndre G 1958 mdash Lapidopiscum nov gen type nouveau de Radiolaire viseacuteen famille des Lapidopiscidae fam nov de lrsquoordre des Albaillellidae Defl 1953 Comptes Rendus heb-domadaires des Seacuteances de lrsquoAcadeacutemie des Sciences (Paris) Seacuterie D Sciences naturelles 246 2278-2280 httpgallicabnffrark12148bpt6k723q

deflAndre G 1960 mdash Agrave propos du deacuteveloppement des recherches sur les Radiolaires fossiles Revue de Micropaleacuteontologie 2 (4) 212-218

deflAndre G 1963 mdash Pylentonema nouveau genre de Radi-olaire du Viseacuteen Sphaerellaire ou Nassellaire Comptes Rendus hebdomadaires des Seacuteances de lrsquoAcadeacutemie des Sciences (Paris) Seacuterie D Sciences naturelles 257 3981-3984 httpgallicabnffrark12148bpt6k4009k

deflAndre G 1964 mdash La famille des Popofskyellidae fam nov et le genre Popofskyellum Defl Radiolaires viseacuteens de la Montagne Noire Comptes Rendus hebdomadaires des Seacuteances de lrsquoAcadeacutemie des Sciences (Paris) Seacuterie D Sciences naturelles 259 3055-3058 httpgallicabnffrark12148bpt6k4015m

deflAndre G 1972a mdash Le systegraveme trabeacuteculaire interne chez les Pylentoneacutemideacutes et les Popofskyellideacutes Radiolaires du Paleo-zoique Phylogenegravese des Nassellaires Comptes Rendus hebdo-madaires des Seacuteances de lrsquoAcadeacutemie des Sciences (Paris) Seacuterie D Sciences naturelles 274 (26) 3535-3540 httpgallicabnffrark12148bpt6k5682741m

deflAndre G 1972b mdash Remarques compleacutementaires sur la morpho logie et la nomenclature de quelques genres de Radiolaires du Paleacuteozoiumlque Comptes Rendus hebdomadaires des Seacuteances de lrsquoAcadeacutemie des Sciences (Paris) Seacuterie D Sciences naturelles 275 (1) 13-16 httpgallicabnffrark12148bpt6k57786873

deflAndre G 1973a mdash Sur quelques nouveaux types de radi-olaires Polycystines viseacuteens drsquoattribution systeacutematique ambigueuml certains eacutevoquant agrave la fois des Plectellaires et des Spumellaires Comptes Rendus hebdomadaires des Seacuteances de lrsquoAcadeacutemie des Sci-ences (Paris) Seacuterie D Sciences naturelles 276 289-293 httpgallicabnffrark12148bpt6k5803214k

deflAndre G 1973b mdash Compleacutements historiques et taxinom-iques sur les Radiolaires viseacuteens Remarques critiques sur les Plectellaires Comptes Rendus hebdomadaires des Seacuteances de lrsquoAcadeacutemie des Sciences (Paris) Seacuterie D Sciences naturelles 276 (1) 497-500 httpgallicabnffrark12148bpt6k5803214k

deflAndre G 1973c mdash Observations et remarques sur les Radiolaires Sphaerellaires du Paleacuteozoiumlque agrave propos drsquoune nouvelle espece viseacuteenne du genre Foremaniella Defl parfait intermediaire entre les Peacuteriaxoplastidieacutes et les Pylentoneacutemideacutes Comptes Rendus hebdomadaires des Seacuteances de lrsquoAcadeacutemie des Sci-ences (Paris) Seacuterie D Sciences naturelles 276 (1) 1147-1151 httpgallicabnffrark12148bpt6k5803214k

deflAndre G 1973d mdash Sur quelques nouvelles espegraveces drsquoArchocyrtium radiolaires Pylentonemidae du Viseacuteen de Cabriegraveres Comptes Rendus hebdomadaires des Seacuteances de lrsquoAcadeacutemie des Sci-ences (Paris) Seacuterie D Sciences naturelles 277 149-152 httpgallicabnffrark12148bpt6k5474901n

deflAndre G amp deflAndre-riGAud M 1958 mdash Donneacutees paleacuteontologiques sur lrsquoontogenegravese de la coque des radiolaires Sphaerellaires Conseacutequences taxonomiques Comptes Rendus hebdomadaires des Seacuteances de lrsquoAcadeacutemie des Sciences (Paris) Seacuterie D Sciences naturelles 246 (6) 968-970 httpgallicabnffrark12148bpt6k3198s

deMAnet f 1938 mdash La faune des Couches de passage du Dinan-tien au Namurien dans le synclinorium de Dinant Meacutemoires du Museacutee royal drsquoHistoire naturelle de Belgique 84 1-201

donG X P knoll A h amp liPPs J h 1997 mdash Late Cambrian Radiolaria from Hunan China Journal of Paleontology 71 (5) 753-758 httpsdoiorg101017S002233600003571X

fenG Q 1992 mdash Permian and Triassic radiolarian biostratigraphy in south and southwest China Journal of China University of Geosciences 3 (1) 51-62

fenG Q amp liu b 1992 mdash One new species of early Devonian radiolarian from west Yunnan Science in China Series D Earth Sciences 35 (5) 549-553

fenG Q amp liu b 1993 mdash Permian radiolarian in southwest Yun-nan Earth Science Journal of China University of Geosciences 18 (5) 540-564

fenG Q amp Ye M 1996 mdash Radiolarian stratigraphy of Devonian through Middle Triassic in southwestern Yunnan in fAnG n amp fenG Q (eds) Devonian to Triassic Tethys in Western Yunnan China China University of Geosciences Press Wuhan 15-22

fenG Q amp Gu s 2002 mdash Uppermost Changxingian (Permian) radiolarian fauna from southern Guizhou southwestern China Journal of Paleontology 76 (5) 797-809 httpsdoiorg101017S0022336000037483

fenG Q Mei Y amp zhAnG z 1997 mdash Early Carboniferous radiolarians from western Yunnan Acta Micropalaeontologica Sinica 14 (1) 79-92

fenG Q fAnG n zhAnG z amp huAnG J 1998 mdash Uppermost Permian Radiolaria from southwestern China Journal of China University of Geosciences 9 (3) 238-245

fenG Q Gu s JiAnG M amp Jin Y 2004a mdash Two new radiolarian genera from the uppermost Permian of southern China Revue de Micropaleacuteontologie 47 (3) 135-143 httpsdoiorg101016S0035-1598(04)00033-9

fenG Q helMcke d chonGlAkMAni c inGAvAt-helMcke r amp liu b 2004b mdash Early Carboniferous radiolarians from north-west Thailand palaeogeographical implications Palaeontology 47 (2) 377-393 httpsdoiorg101111j0031-0239200400370x

fenG Q he W Gu s Jin Y amp MenG Y 2006a mdash Latest Per-mian Spumellaria and Entactinaria (Radiolaria) from South China Revue de Micropaleacuteontologie 49 (1) 21-43 httpsdoiorg101016jrevmic200511003

fenG Q he W zhAnG s amp Gu s 2006b mdash Taxonomy of order Latentifistularia (Radiolaria) from the Latest Permian in Southern Guangxi China Journal of Paleontology 80 (5) 826-848 httpsdoiorg1016660022-3360(2006)80[826TOOLRF]20CO2

fenG Q MenG Y he W amp Gu s 2006c mdash A new genus of Entactiniidae (Radiolaria) from the Upper Permian of South China Eclogae geologicae Helvetiae 99 (supplement 1) S67-S78 httpsdoiorg101007s00015-006-0608-z

fenG Q Gu s he W amp Jin Y 2007 mdash Latest Permian Entactinaria (Radiolaria) from southern Guangxi China Journal of Micro-palaeontology 26 (1) 19-40 httpsdoiorg101144jm26119

fenG Q Mei Y amp crAsQuin s 2009 mdash Latest Permian Pal-aeolithocycliidae (Radiolaria) from South China Revue de Micropaleacuteontologie 52 (2) 141-148 httpsdoiorg101016jrevmic200708008

foreMAn h P 1963 mdash Upper Devonian Radiolaria from the Huron member of the Ohio shale Micropaleontology 9 (3) 267-304 httpsdoiorg1023071484751

furutAni h 1983 mdash Middle Palaeozoic Palaeoscenidiidae (Radio-laria) from Mt Yokokura Shikoku Japan Part 1 Transactions and Proceedings of the Palaeontological Society of Japan New Series 130 96-116 httpsdoiorg1014825prpsj19511983130_96

furutAni h 1990 mdash Middle Paleozoic radiolarians from Fukuji Area Gifu Prefecture central Japan Journal of Earth Sciences Nagoya University 37 1-56

GoodbodY Q h 1986 mdash Wenlock Palaeoscenidiidae and Entac-tiniidae (Radiolaria) from the Cape Phillips Formation of the Canadian Arctic Archipelago Micropaleontology 32 (2) 129-157 httpsdoiorg1023071485627

Goto h uMedA M amp ishiGA h 1992 mdash Late Ordovician Radiolarians from the Lachlan Fold Belt Southeastern Australia Memoirs of the Faculty of Science Shimane University 26 145-170

GourMelon f 1986 mdash Eacutetude des radiolaires drsquoun nodule phos-phateacute du Carbonifegravere inferieur de Bareilles Hautes-Pyrenees France Geobios 19 (2) 179-206 httpsdoiorg101016S0016-6995(86)80044-4



GourMelon f 1987 mdash Les Radiolaires tournaisiens des nodules phosphateacutes de la Montagne Noire et des Pyreacuteneacutees centrales Biostratigraphie du Paleacuteozoiumlque 6 1-172

Gui b fenG Q amp YuAn A 2009 mdash Late Changhsingian (Latest Permian) radiolarians from Chaohu Anhui Journal of Earth Science 20 (5) 797-810 httpsdoiorg101007s12583-009-0069-1

hAo Y amp shu d 1987 mdash The oldest known well-preserved Phaeo-darina (Radiolaria) from southern Shensi Geoscience (Beijing) 1 (3-4) 301-310

he W fenG Q Gu s amp Jin Y 2005 mdash Changxingian (Upper Permian) radiolarian fauna from Meishan D Section Changx-ing Zhejiang China and its possible paleoecological signifi-cance Journal of Paleontology 79 (2) 209-218 httpsdoiorg1016660022-3360(2005)079lt0209CUPRFFgt20CO2

he W zhAnG Y zhAnG Q zhAnG k YuAn A amp fenG Q 2011 mdash A latest Permian radiolarian fauna from Hushan South China and its geological implications Alcheringa 35 (4) 471-496 httpsdoiorg101080031155182010536649

hinde G J 1890 mdash Notes on Radiolaria from the Lower Palaeo-zoic rocks (Llandeilo-Caradoc) of the South of Scotland Annals and Magazine of Natural History sixth series 6 40-59 httpsdoiorg10108000222939008693993

hinde G J 1899a mdash On the Radiolaria in the Devonian rocks of New South Wales Quarterly Journal of the Geological Society of London 55 38-64 httpsdoiorg101144GSLJGS189905501-0406

hinde G J 1899b mdash On Radiolaria in chert from Chypons Farm Mullion Parish (Cornwall) Quarterly Journal of the Geological Society of London 55 214-219 httpsdoiorg101144GSLJGS189905501-0414

hinde G J amp foX h 1895 mdash On a well-marked horizon of radiolarian rocks in the Lower Culm Measures of Devon Cornwall and West Somerset Quarterly Journal of the Geological Society of London 51 609-668 httpsdoiorg101144GSLJGS189505101-0445

hinz-schAllreuter i amp schAllreuter r 2003 mdash Radiolarians from the Sularp shale (Ordovician of Scania) Geschiebekunde Aktuell 19 (3) 69-85

holdsWorth b k 1966 mdash A preliminary study of the palae-ontology and palaeoenvironment of some Namurian limestone lsquobullionsrsquo The Mercian Geologist 1 (4) 315-337

holdsWorth b k 1969 mdash Namurian Radiolaria of the genus Ceratoikiscum from Staffordshire and Derbyshire England Micro-paleontology 15 (2) 221-229 httpsdoiorg1023071484921

holdsWorth b k amp Jones d l 1980 mdash Preliminary radi-olarian zonation for late Devonian through Permian time Geology 8 (6) 281-285 httpsdoiorg1011300091-7613(1980)8lt281PRZFLDgt20CO2

holdsWorth b k amp MurcheY b l 1988 mdash Paleozoic radio-larian biostratigraphy of the northern Brooks Range Alaska in GrYc G (ed) Geology and Exploration of the National Petroleum Reserve in Alaska 1974 to 1982 United States Geological Survey professional Paper Report 1399 777-797

isAkovA t n amp nAzArov b b 1986 mdash Late Carboniferous-Early Permian stratigraphy and microfauna of the Southern Urals Doklady Akademii Nauk SSSR 402 1-183

ishiGA h 1982 mdash Late Carboniferous and Early Permian radiolarians from the Tamba Belt Southwest Japan Earth Science Journal of the Association for the Geological Collaboration in Japan 36 (6) 333-339

ishiGA h 1983 mdash Morphological change in the Permian Radiolaria Pseudoalbaillella scalprata in Japan Transactions and Proceedings of the Palaeontological Society of Japan New Series 129 1-8 httpsdoiorg1014825prpsj19511983129_1

ishiGA h 1988 mdash Middle and Late Paleozoic Radiolarian Biostratig-raphy of Japan Geological Report Shimane University 7 69-76 httpirlibshimane-uacjp6303

ishiGA h 1991 mdash Description of a new Follicucullus species from southwest Japan Memoirs of the Faculty of Science Shimane Uni-versity 25 107-118

ishiGA h 1992 mdash Middle Paleozoic radiolarians of the genus Ceratoikiscum from Japan in ishizAki k amp sAiti T (eds) Cen-tenary of Japanese Micropaleontology Terra Scientific Publishing Company Tokyo 389-397

ishiGA h amp iMoto n 1980 mdash Some Permian radiolarians in the Tamba District Southwest Japan Earth Science Journal of the Association for the Geological Collaboration in Japan 34 (6) 333-345

ishiGA h kito t amp iMoto n 1982a mdash Late Permian radio-larian assemblages in the Tamba District and an adjacent area Southwest Japan Earth Science Journal of the Association for the Geological Collaboration in Japan 36 (1) 10-22

ishiGA h kito t amp iMoto n 1982b mdash Middle Permian radio-larian assemblages in the Tamba District and an adjacent area Southwest Japan Earth Science Journal of the Association for the Geological Collaboration in Japan 36 (5) 272-281

ishiGA h iMoto n YoshidA M amp tAnAbe t 1984 mdash Early Permian radiolarians from the Tamba Belt Southwest Japan Earth Science Journal of the Association for the Geological Col-laboration in Japan 38 (1) 44-54

ishiGA h WAtAse h amp nAkA t 1986 mdash Permian radiolar-ians from Nishiki Group in Sangun-Chugoku Belt Southwest Japan Earth Science Journal of the Association for the Geological Collaboration in Japan 40 (2) 124-136

ishiGA h leitch e c nAkA t WAtAnAbe t amp iWAsAki M 1987 mdash Late Devonian Paleoscenidiidae from the Hastings Block New England Fold Belt NSW Australia Earth Sci-ence Journal of the Association for the Geological Collaboration in Japan 41 (6) 297-302

Jin Y fenG Q MenG Y he W amp Gu s 2007 mdash Albaillellidae (Radiolaria) from the latest Permian in southern Guangxi China Journal of Paleontology 81 (1) 9-18 httpsdoiorg1016660022-3360(2007)81[9ARFTLP]20CO2

Jones M k amp noble P J 2006 mdash Sheinwoodian (uppermost Lower Silurian) Radiolaria from the Cape Phillips Formation Nunavut Canada Micropaleontology 52 (4) 289-315 httpsdoiorg102113gsmicropal524289

khAbAkov A v 1932 mdash O nakhodke fauny radiolyarii v kremn-istykh slantsakh i yashmakh paleozoya Kavkazskogo khrebta i ee paleogeograficheskom znachenii [On the discovery of radiolarians in the Paleozoic cherts and jaspers of the Caucasus Range and its paleogeographic significance] Bulletin of the United Geological and Prospecting Service of USSR 51 (12) 219-225

kiesslinG W amp trAGelehn h 1994 mdash Devonian Radiolarian Faunas of Conodont-dated localities in the Frankenwald (Northern Bavaria Germany) Abhandlungen der geologisches Bundesanstalt in Wien 50 219-255

kozur h 1980 mdash Ruzhencevispongidae eine neue Spumellaria Familie aus dem oberen Kungurian (Leonardian) und Sakmarian des Vorurals Geologisch Palaumlontologische Mitteilungen Innsbruck 10 (6) 235-242

kozur h 1981 mdash Albaillellidae (Radiolaria) aus dem Unterperm des Vorurals Geologisch Palaumlontologische Mitteilungen Innsbruck 10 (8) 263-274

kozur h 1993 mdash Upper Permian Radiolarians from the Sosio Valley Area Western Sicily (Italy) and from the Uppermost Lamar Limestone of West Texas Jahrbuch der geologischen Bundesanstalt Wien 136 (1) 99-123

kozur h 1997 mdash First discovery of Muellerisphaerida (inc sedis) and Eoalbaillella (Radiolaria) in Turkey and the age of the silici-clastic sequence (clastic series) in Karaburun Peninsula Freiberger Forschungshefte C Geowissenschaften Geologie 4 33-59

kozur h amp Mostler h 1989 mdash Radiolarien und schwammskleren aus dem Unterperm des Vorurals Geologisch Palaumlontologische Mitteilungen Innsbruck Sonderband 2 147-275

kozur h W Mostler h amp rePetski J e 1996 mdash Well-preserved Tremadocian primitive Radiolaria from the Windfall Formation of the Antelope Range Eureka County Nevada USA Geologisch Palaumlontologische Mitteilungen Innsbruck 21 245-271


Aitchison J C et al

kurihArA t 2007 mdash Uppermost Silurian to Lower Devonian radiolarians from the Hitoegane area of the Hida-Gaien terrane central Japan Micropaleontology 53 (3) 221-237 httpsdoiorg102113gsmicropal533221

kurihArA t amp sAshidA k 2000 mdash Taxonomy of Late Silurian to Middle Devonian radiolarians from the Kuzuryu Lake district of the Hida Gaien Belt Fukui Prefecture central Japan Micro-paleontology 46 (1) 51-71 httpwwwjstororgstable1486025

kuWAhArA k 1999 mdash Phylogenetic Lineage of Late Permian Albaillella (Albaillellaria Radiolaria) Journal of Geosciences Osaka City University 42 (6) 85-101

kuWAhArA k amp sAkAMoto M 1992 mdash Late Permian Albaillella (Radiolaria) from a bedded chert section in the Gujo-hachiman area of the Mino Belt central Japan preliminary report on morphometry and cluster analysis Journal of Geosciences Osaka City University 35 (3) 33-52

li h 1991 mdash First discovery of Middle Silurian Radiolaria fossils in Xinjiang Scientia Geologica Sinica 1 75

li h 1994 mdash Middle Silurian radiolarians from Keerhada Xinjiang Acta Micropalaeontologica Sinica 11 (2) 259-272

li h 1995 mdash New genera and species of middle Ordovician Nas-sellaria and Albaillellaria from Baijingsi Quilian Mountains China Scientia Geologica Sinica 4 (3) 331-346

li Y X amp WAnG Y J 1991 mdash Upper Devonian (Frasnian) radiolarian fauna from the Liukiang Formation eastern and southeastern Guangxi Acta Micropalaeontologica Sinica 8 (4) 395-404

linG h Y amp forsYthe r d 1987 mdash Late Paleozoic pseudo-albaillellid radiolarians from southernmost Chile and their geological significance in Mckenzie G D (ed) Gonwanda Six Structure Tectonics and Geophysics American Geophysical Union (Geophysical Monograph vol 40) 253-260 httpsdoiorg101029GM040p0221

liu h 1992 mdash Fossil radiolarians from the turbidite of the Middle Carboniferous east Tianshan Mountain Xinjiang Journal of Northwest University Natural Science Education 22 123-130 169-170

liu Y amp hAo s 2006 mdash Evolutionary significance of pylenton-emid radiolarians and their Late Devonian species from south-western Tianshan China Acta Geologica Sinica 80 (5) 647-655

luo h Aitchison J c amp WAnG Y J 2002 mdash Devonian (upper Emsian-lower Givetian) radiolarians from the Tanhe Formation Nanning Guangxi Southwest China Micropaleontology 48 (supplement 1) 113-127 httpwwwjstororgstable1486015

MAcdonAld e W 1998 mdash Llandovery Secuicollactinae and Rotasphaeridae (Radiolaria) from the Cape Phillips Formation Cornwallis Island Arctic Canada Journal of Paleontology 72 (4) 585-604 httpsdoiorg101017S0022336000040324

MAcdonAld e W 1999 mdash Insolitignum ngen and Palaeoephip-pium Goodbody 1986 (Radiolaria) from the Lower Silurian of the Cape Phillips Formation Arctic Canada Canadian Journal of Earth Science 36 (12) 2051-2057 httpsdoiorg101139e99-101

MAcdonAld e W 2004 mdash Palaeoscenidiidae (Radiolaria) from the Lower Silurian ff the Cape Phillips Formation Cornwallis Island Nunavut Canada Journal of Paleontology 78 (2) 257-274 httpsdoiorg1016660022-3360(2004)078lt0257PRFTLSgt20CO2

MAcdonAld e W 2006 mdash Haplotaeniatumidae and Inanigutti-dae (Radiolaria) from the Lower Silurian of the Cape Phillips Formation Cornwallis Island Nunavut Canada Journal of Paleontology 80 (1) 19-37 httpsdoiorg1016660022-3360(2006)080[0019HAIRFT]20CO2

MAldonAdo A l amp noble P J 2010 mdash Radiolarians from the upper Guadalupian (Middle Permian) Reef Trail Member of the Bell Canyon Formation West Texas and their biostratigraphic implications Micropaleontology 56 (1-2) 69-115 httpswwwjstororgstable40607077

MAletz J 2007 mdash The Early Ordovician Beothuka terranova (Radiolaria) faunal assemblage in western Newfoundland Pal-aeontologische Zeitschrift 81 (1) 71-82 httpsdoiorg101007BF02988380

MAletz J amp bruton d l 2005 mdash The Beothuka terranova (Radiolaria) assemblage and its importance for the understanding of early Ordovician radiolarian evolution Geological Magazine 142 (6) 711-721 httpsdoiorg101017S0016756805001391

MAletz J amp bruton d l 2007 mdash Lower Ordovician (Chew-tonian to Castlemainian) radiolarians of Spitsbergen Journal of Systematic Palaeontology 5 (3) 245-288 httpsdoiorg101017S1477201907002039

MAletz J amp bruton d l 2008 mdash The Middle Ordovician Proventocitum procerulum radiolarian assemblage of Spitsbergen and its biostratigraphic correlation Palaeontology 51 (5) 1181-1200 httpsdoiorg101111j1475-4983200800803x

nAzArov b b 1973a mdash First occurrences of Radiolaria of the Entac-tiniidae and Ceratoikiscidae families in the upper Devonian of the Southern Urals Doklady Akademii Nauk SSSR 210 (3) 696-699

nAzArov b b 1973b mdash Radiolaria from the lower horizons of the Cambrian of Bazhenev Ridge in Problems of paleontology and biostratigraphy of the Lower Cambrian of Siberia and the Far Eastern USSR Transactions of the Institute of Geology and Geophysics Siberian Branch of the Academy of Sciences 49 5-13

nAzArov b b 1974a mdash Albaillellidae and Paleoscenidiidae from the Upper Devonian deposits of the Southern Urals in zhAMoidA A I (ed) Biostratigraphy and Paleontology of the Lower Cambrian of Europe and Middle Asia Publication of the All-Union Institute of Geology new series Leningrad vol 226 41-47

nAzArov b b 1974b mdash Problematic siliceous formations from the lower Paleozoic of Kazakhstan in zhAMoidA A I (ed) Biostratigraphy and Paleontology of the Lower Cambrian of Europe and Middle Asia Publication of the All-Union Institute of Geol-ogy new series Leningrad vol 226 110-112

nAzArov b b 1975 mdash Lower and Middle Paleozoic radiolar-ians of Kazakhstan (methods of investigation systematics and stratigraphic significance) in rAAben M E (ed) Transactions of the Academy of Sciences of the USSR Geological Institute Izdatelstvo Nauka Moscow vol 275 1-203

nAzArov b b 1977 mdash A new radiolarian family from the Ordo-vician of Kazakhstan Paleontological Journal 11 (2) 165-171

nAzArov b b 1980 mdash Class Sarcodina subclass Radiolaria in APollonov M k bAndAletov s M amp nikitin i F (eds) The Ordovician-Silurian Boundary in Kazakhstan Nauka Alma-Ata Kazakhstan USSR 22-24

nAzArov b b 1988 mdash Paleozoic radiolaria in zhAMoidA A I (ed) Practical Manual of Microfauna of the USSR Nedra Len-ingrad vol 2 232 p

nAzArov b b amp shkolnik e l 1974 mdash On the question of the age of the siliceous-volcanogenic sequences in the Ud-Selemzhin interfluve in sYssoiev V (ed) Biostratigraphy and Paleontol-ogy of the Lower Cambrian of Europe and Central Asia Nauka Moscow 104-109

nAzArov b b amp kruYcheck s A 1977 mdash Upper Devonian Radiolaria of the Russian Platform Transaction of Doklady of the Russian Academy of Science Earth Science Sections 6 (237) 1445-1448

nAzArov b b amp rudenko v s 1981 mdash Some radiolarians with bilateral symmetry of the upper Paleozoic of the Southern Urals Systematics and Morphology of Microorganisms Problems of Micropaleontology 24 129-139

nAzArov b b amp nolvAk J 1983 mdash Radiolarians from the Upper Ordovician of Estonia Izvestiya Akademii Nauk Estonskoy SSR Geologiya 32 (1) 1-8

nAzArov b b amp orMiston A 1983a mdash Upper Devonian (Frasnian) radiolarian fauna from the Gogo Formation West-ern Australia Micropaleontology 29 (4) 454-466 httpsdoiorg1023071485519



nAzArov b b amp orMiston A 1983b mdash A new superfamily of stauraxon polycystine Radiolaria from the Late Paleozoic of the Soviet Union and North America Senckenbergiana Lethaea 64 (2-4) 363-379

nAzArov b b amp orMiston A 1984 mdash Tentative system of Paleozoic Radiolaria in PetrushevskAYA M G amp stePAnJAnts S D (eds) Morphology Ecology and Evolution of Radiolarians Material from the IV Symposium of European Radiolarists EURO-RAD IV Academiya Nauk SSSR Zoological Institute Leningrad USSR 64-87

nAzArov b b amp orMiston A r 1985 mdash Radiolaria from the Late Paleozoic of the Southern Urals USSR and West Texas USA Micropaleontology 31 (1) 1-54 httpsdoiorg1023071485579

nAzArov b b amp orMiston A r 1987 mdash A new Carboniferous radiolarian genus and its relation to the multishelled entactiniids Micropaleontology 33 (1) 66-73 httpsdoiorg1023071485527

nAzArov b b amp orMiston A r 1989 mdash New radiolarian species from the Permian of the Southern Urals and Nevada Paleontological Journal 23 (2) 10-20

nAzArov b b amp orMiston A r 1990 mdash The biostratigraphic potential of radiolarians from the Palaeozoic in chuvAshov B I zhAMoidA A i amp AMon E O (eds) Radiolaria in Biostratigraphy Collected Proceedings Uralrsquoskoe otdelenie AN SSSR Sverdlovsk 3-25 121 125 126

nAzArov b b amp orMiston A r 1993 mdash New biostratigraphically important Paleozoic Radiolaria of Eurasia and North America in blueford J r amp MurcheY B L (eds) Radiolaria of Giant and Subgiant Fields in Asia Nazarov Memorial Volume Micro-paleontology Special Publication 6 22-60

nAzArov b b amp PoPov l Y 1976 mdash Radiolarians ecardine brachiopods and organisms of uncertain systematic position from the Middle Ordovician of eastern Kazakhstan Paleontological Journal 4 407-416

nAzArov b b amp PoPov l Y 1980 mdash Stratigraphy and fauna of the siliceous-carbonate sequence of the Ordovician of Kazakh-stan (Radiolaria and inarticulate brachiopods) Transactions of the Geological Institute of the Soviet Akademy of Sciences 331 1-192

nAzArov b b PoPov l Y amp APollonov M 1975 mdash Lower Paleozoic Radiolaria in Kazakhstan Izvestija Akademii Nauk SSSR serija geologičeskaja 10 96-105

nAzArov b b PoPov l Y amp APollonov M 1977 mdash Lower Paleozoic radiolarians of Kazakhstan International Geology Review 19 (8) 913-920 httpsdoiorg10108000206817709471089

nAzArov b b tkAchenko v i amp shulGinA v s 1981 mdash Radio-laria and age of terrigenous siliceous beds of the Kolyma region Izvestija Akademii Nauk SSSR serija geologičeskaja 10 79-89

nAzArov b b cockbAin A e amp PlAYford P e 1982 mdash Late Devonian Radiolaria from the Gogo Formation Canning Basin Western Australia Alcheringa 6 (3) 161-173 httpsdoiorg10108003115518208565408

nestell G P amp nestell M k 2010 mdash Late Capitanian (latest Guadalupian Middle Permian) radiolarians from the Apache Mountains West Texas Micropaleontology 56 (1-2) 7-68 httpswwwjstororgstable40607076

nestell G P PoPe J P amp nestell M k 2012 mdash Middle Penn-sylvanian (Desmoinesian) Radiolaria from the Midcontinent of North America Micropaleontology 58 (3) 217-257 httpswwwjstororgstable23250632

nishiMurA k amp ishiGA h 1987 mdash Radiolarian biostratigraphy of the Maizuru Group in Yanahara area Southwest Japan Memoirs of the Faculty of Science Shimane University 21 169-188

nitecki M h 1963 mdash Catalogue of type specimens in the Walker Museum of Paleontology and Chicago Natural History Museum ndash Radiolaria and Eurypterida Fieldiana Geology (New Series) 13 (3) 163-183 httpbiodiversitylibraryorgpage4260689

noble P J 1994 mdash Silurian Radiolarian Zonation for the Caballos Novaculite Marathon Uplift West Texas Bulletins of American Pale-ontology 106 (2) 1-55 httpbiodiversitylibraryorgpage10684913

noble P amp renne P 1990 mdash Paleoenvironmental and biostratigraphic significance of siliceous microfossils of the Permo-Triassic Redding Sec-tion Eastern Klamath Mountains California Marine Micropaleontology 15 (3-4) 379-391 httpsdoiorg1010160377-8398(90)90021-D

noble P J amp MAletz J 2000 mdash Radiolaria from the Telychian (Llandovery Early Silurian) of Dalarna Sweden Micropaleontol-ogy 46 (3) 265-275 httpsdoiorg102113463265

noble P J amp lenz A c 2007 mdash Upper Wenlock Ceratoikis-cidae (Radiolaria) from the Cape Phillips Formation Arctic Canada Journal of Paleontology 81 (5) 1044-1052 httpsdoiorg101666pleo05-0531

noble P J amp WebbY b d 2009 mdash Katian (Ordovician) radiolar-ians from the Malongulli Formation New South Wales Australia a re-examination Journal of Paleontology 83 (4) 548-561 httpsdoiorg10166608-179R1

noble P J amp Jin Y 2010 mdash Radiolarians from the Lamar Lime-stone Guadalupe MountainsWest Texas Micropaleontology 56 (1-2) 117-147 httpswwwjstororgstable40607078

noble P J ketner k b amp McclellAn W 1997 mdash Early Silurian Radiolaria from Northern Nevada USA Marine Micro-paleontology 30 (1-3) 215-223 httpsdoiorg101016S0377-8398(96)00026-6

noble P J tekin u k Gedik i amp sukru P 2008 mdash Middle to Upper Tournasian Radiolaria of the Baltalimani Formation Istanbul Turkey Journal of Paleontology 82 (1) 37-56 httpsdoiorg10166606-0461

orsquodoGhertY l 2009 mdash Inventory of Mesozoic radiolarian spe-cies (1867-2008) Geodiversitas 31 (2) 371-481 httpsdoiorg105252g2009n2a6

obut o t amp iWAtA k 2000 mdash Lower Cambrian Radiolaria from the Gorny Altai (southern West Siberia) Novosti Paleontologii i Stratigrafii 2-3 33-37

obut o t amp shcherbAnenko t A 2008 mdash Late Devonian radiolarians from the Rudny Altai (SW Siberia) Bulletin of Geosciences 83 (4) 371-382 httpsdoiorg103140bullgeo-sci200804371

orMiston A r amp lAne h r 1976 mdash A unique radiolarian fauna from the Sycamore Limestone (Mississippian) and its biostratigraphic significance Palaeontographica Abteilung A Paleozoologie-Stratigraphie 154 158-180

orMiston A r amp bAbcock l 1979 mdash Follicucullus new radio-larian genus from the Guadalupian (Permian) Lamar limestone of the Delaware Basin Journal of Paleontology 53 (2) 328-334 httpwwwjstororgstable1303874

PArk i-Y amp Won M-z 2012 mdash Tropical radiolarian assemblages from the Lower Carboniferous Delle Phosphatic Member of the Woodman Formation of Utah USA Journal of the Paleontologi-cal Society of Korea 28 (1-2) 29-101

Pickett J amp Jell P A 1983 mdash Middle Cambrian Sphinctozoa (Porifera) from New South Wales Memoirs of the Association of Australian Palaeontologists 1 85-92

Pouille l dAneliAn t Pour M G amp PoPov l e 2013 mdash New and Revised Inaniguttid Radiolaria and Associated Trilobites from the Upper Darriwilian (Ordovician) Shundy Formation of Kazakhstan Journal of Paleontology 87 (6) 1143-1159 httpsdoiorg10166612-106

Pouille l dAneliAn t amp PoPov l e 2014 mdash A diverse Upper Darriwilian radiolarian assemblage from the Shundy Formation of Kazakhstan insights into late Middle Ordovician radiolar-ian biodiversity Journal of Micropalaeontology 33 (2) 149-163 httpsdoiorg101144jmpaleo2014-008

PulfreY W 1932 mdash The occurrence of radiolarian-bearing nodules at the base of the Edale Shales near Calver Sough North Der-byshire Proceedings of the Geologistsrsquo Association 43 (2) 192-198 httpsdoiorg101016S0016-7878(32)80027-1

renz G W 1990 mdash Late Ordovician (Caradocian) radiolarians from Nevada Micropaleontology 36 (4) 367-377 httpsdoiorg1023071485476


Aitchison J C et al

rothPletz A 1880 mdash Radiolarien Diatomaceen und Spharoso-matiten im silurischen Kieselschiefer von Langenstriegis Sachsen Zeitschrift der deutschen geologischen Gesellschaft 32 447-467

rudenko v s amp PAnAsenko e s 1990a mdash A new findings of the Upper Permian radiolarians in Primorye region in zAkhArov Y d belYAevA G v amp nikitinA A P (eds) New Data on Palaeozoic and Mesozoic Biostratigraphy of the South Far East (IUGS Project 272) Far Eastern Branch of the USSR Academy of Sciences Vladivostok 117-124

rudenko v s amp PAnAsenko e s 1990b mdash Permian Albail-lellaria (Radiolaria) of the Pantovian sequence in Primorye in zAkhArov Y d belYAevA G v amp nikitinA A P (eds) New Data on Palaeozoic and Mesozoic Biostratigraphy of the South Far East (IUGS Project 272) Far Eastern Branch of the USSR Academy of Sciences Vladivostok 181-193

ruedeMAnn r amp Wilson t Y 1936 mdash Eastern New York Ordovician cherts Geological Society of America Bulletin 74 (10) 1535-1586 httpsdoiorg101130GSAB-47-1535

ruumlst d 1892 mdash Beitraumlge zur Kenntniss der fossilen Radiolar-ien aus Gesteinen der Trias und der palaeozoischen Schichten Palaeonto graphica 38 107-179 httpbiodiversitylibraryorgpage33193870

sAshidA k amp tonishi k 1985 mdash Permian radiolarians from the Kanto Mountains central Japan some Upper Permian Spumel-laria from Itsukaichi western part of Tokyo Prefecture Science Reports of the Institute of Geoscience University of Tsukuba Section B Geological Sciences 6 1-19

sAshidA k amp tonishi k 1986 mdash Upper Permian stauraxon polycystine Radiolaria from Itsukaichi western part of Tokyo Prefecture Science Reports of the Institute of Geoscience University of Tsukuba Section B Geological Sciences 7 1-13

sAshidA k amp tonishi k 1988 mdash Additional note on Upper Permian radiolarian fauna from Itsukaichi western part of Tokyo Prefecture central Japan Transactions and Proceedings of the Palaeontological Society of Japan New Series 151 523-542 httpsdoiorg1014825prpsj19511988151_523

sAshidA k amp tonishi k 1991 mdash An Upper Permian coiled radiolarian from Itsukaichi central Japan Micropaleontology 37 (1) 86-94 httpsdoiorg1023071485747

sAshidA k AdAchi s iGo h nAkornsri n amp AMPornMAhA A 1997 mdash Middle to Upper Permian and Middle Triassic radiolarians from Eastern Thailand Science Reports of the Insti-tute of Geoscience University of Tsukuba Section B Geological Sciences 18 1-17

sAshidA k iGo h AdAchi s ueno k kAJiWArA Y nAkornsri n amp sArdsud A 2000a mdash Late Permian to Middle Triassic radiolarian faunas from Northern Thailand Journal of Paleontology 74 (5) 789-811 httpsdoiorg101017S0022336000033011

sAshidA k nAkornsri n ueno k amp sArdsud A 2000b mdash Carboniferous and Triassic radiolarian faunas from the Saba Yoi area southernmost part of peninsular Thailand and their paleogeographic significance Science Reports of the Institute of Geoscience University of Tsukuba Section B Geological Sciences 21 71-99

sAshidA k sAlYAPonGse s amp nAkornsri n 2000c mdash Latest permian radiolarian fauna from Klaeng eastern Thailand Micro-paleontology 46 (3) 245-263 httpsdoiorg102113463245

schWArtzAPfel J A amp holdsWorth b k 1996 mdash Upper Devo-nian and Mississippian radiolarian zonation and biostratigraphy of the Woodford Sycamore Caney and Goddard Formations Oklahoma Cushman Foundation for Foraminiferal Research Special Publication 33 1-275

seo e-h amp Won M-z 2009 mdashReview of the genus Polyen-tactinia and the family Polyentactiniidae Micropaleontology 55 (1) 61-74 httpswwwjstororgstable20627972

shAnG Q-h cAridroit M amp WAnG Y J 2001 mdash Radiolar-ians from the uppermost Permian Changhsingiang of southern Guangxi Acta Micropalaeontologica Sinica 18 (3) 229-240

shenG J z amp WAnG Y J 1982 mdash Fossil Radiolaria of the Mid-dle Devonian of Qiziqiao Formation Xintian Hunan Acta Palaeontologica Sinica 21 (1) 58-66

shenG J z amp WAnG Y J 1985 mdash Fossil Radiolaria from Kufeng Formation at Longtan Nanjing Acta Palaeontologica Sinica 24 (2) 171-183

shu d amp chen l 1989 mdash Discovery of Early Cambrian Radio-laria and its significance Science in China Series B Chemistry 32 (8) 986-994

siveter d J Aitchison J c siveter d J amp sutton M d 2007 mdash The Radiolaria of the Herefordshire Konservat-Lager-staumltte (Silurian) England Journal of Micropalaeontology 26 (1) 86-95 httpsdoiorg101144jm26187

sPiller f c P 2002 mdash Radiolarian biostratigraphy of peninsu-lar Malaysia and implications for regional palaeotectonics and palaeogeography Palaeontographica Abteilung A Paleozoologie-Stratigraphie 266 (1-3) 1-91

strAtford J M c amp Aitchison J c 1997 mdash Lower to Middle Devonian radiolarian assemblages from the Gamilaroi Terrane Glenrock Station NSW Australia Marine Micropaleontology 30 (1-3) 225-250 httpsdoiorg101016S0377-8398(96)00027-8

tAkeMurA A amp nAkAseko k 1981 mdash A new Permian radiolarian genus from the Tamba Belt Southwest Japan Transactions and Proceedings of the Palaeontological Society of Japan New Series 124 208-214 httpsdoiorg1014825prpsj19511981124_208

tAkeMurA A MoriMoto t AitA Y hori r s hiGuchi Y sPoumlrli k b cAMPbell h J kodAMA k amp sAkAi t 1999 mdash Permian Albaillellaria (Radiolaria) from a limestone lens at the Arrow Rocks in the Waipapa Terrane (Northland New Zealand) Geodiversitas 21 (4) 751-765

ters M amp deflAndre G 1966 mdash Sur lrsquoacircge cambro-silurien des terrains anciens de la Vendeacutee Littorale (ex-Brioveacuterien) Comptes Rendus hebdomadaires des Seacuteances de lrsquoAcadeacutemie des Sciences (Paris) Seacuterie D Sciences naturelles 262 (3) 339-342 httpgallicabnffrark12148bpt6k5489696x

tetArd M noble P J dAneliAn t Monnet c amp lenz A c 2015 mdash A new Gorstian radiolarian fauna from the upper Silurian of the Cape Phillips Formation Cornwallis and Bathurst islands Canadian Arctic Canadian Journal of Earth Sciences 52 (10) 863-879 httpsdoiorg101139cjes-2015-0028

uMedA M 1997 mdash Late Silurian and Early Devonian radiolarians from the Konomori area in the Kurosegawa Terrane southwest Japan Earth Science Journal of the Association for the Geological Collaboration in Japan 51 413-432

uMedA M 1998a mdash Early to Middle Devonian Ceratoikiscidae (Radiolaria) from the Yokokurayama Group in the Kurosegawa Terrane Southwest Japan Paleontological Research 2 (2) 96-107 httpsdoiorg102517prpsj296

uMedA M 1998b mdash Some Late Silurian characteristic radiolar-ians from the Yokokurayama Group in the Kurosegawa Terrane southwest Japan Earth Science Journal of the Association for the Geological Collaboration in Japan 52 (3) 203-209

uMedA M amp suzuki Y 2005 mdash Aeronian (Llandovery Early Silurian) Radiolarians from the Kallholn Formation in Siljan district Sweden Micropaleontology 51 (1) 83-92 httpsdoiorg10211351183

vishnevskAYA v s 1998 mdash The domanikoid facies of the Rus-sian platform and basin paleogeography in crAsQuin-soleAu s amp bArrier Eacute (eds) Peri-Tethys Memoir 3 Stratigraphy and Evolution of Peri-Tethyan Platforms Museacuteum national drsquoHistoire naturelle Paris (Meacutemoires du Museacuteum national drsquoHistoire naturelle 177) 45-69

WAkAMAtsu h suGiYAMA k amp furutAni h 1990 mdash Silurian and Devonian radiolarians from the Kurosegawa Tectonic Zone southwest Japan Journal of Earth Sciences Nagoya University 37 157-192

WAnG n W 1989 mdash Micropaleontological study of Lower Pal-aeozoic siliceous sequences of the Yangtze Platform and Eastern



Qinling Range Journal of Southeast Asian Earth Sciences 3 (1-4) 141-161 httpsdoiorg1010160743-9547(89)90018-4

WAnG r J 1993a mdash Fossil Radiolaria from Kufeng formation of Chaohu Anhui Acta Palaeontologica Sinica 32 (4) 442-457

WAnG r J 1993b mdash Morphological change of the Ruzhence-vispongus uralicus in Gufeng Formation (Lower Permian) from Jiangsu and Anhui provinces Shanghai Geology 4 5-12

WAnG r J 1995 mdash Radiolarian fauna from Gufeng Formation (Lower Permian) in Hushan area of Nanjing Jiangsu Province Scientia Geologica Sinica 30 (2) 139-148

WAnG Y J 1993c mdash Middle Ordovician radiolarians from the Pingliang Formation of Gansu Province China Micropaleon-tology Special Publication 6 98-114

WAnG Y J1997 mdash An Upper Devonian (Famennian) radio-larian fauna from carbonate rocks northern Xinjiang Acta Micropalaeontologica Sinica 14 (2) 149-160

WAnG Y J amp shAnG Q 2001 mdash Discovery of the Neoalbail-lella radiolarian fauna in the Shaiwa Group of Ziyun District Guizhou Acta Micropalaeontologica Sinica 18 (2) 111-121

WAnG Y J chenG Y n amp YAnG Q 1994 mdash Biostratigraphy and systematics of Permian Radiolarians in China in YuGAn J uttinG J amp WArdlAW B R (eds) Permian Stratigraphy Environments and Resources Palaeoworld Special Issue 4 172-202

WAnG Y J fAnG z YAnG Q zhou z chenG Y duAn Y amp XiAo Y 2000 mdash Middle-Late Devonian strata with cherty facies and radiolarian faunas in western Yunnan China Acta Micropalaeontologica Sinica 17 (3) 235-254

WAnG Y J Aitchison J c amp luo h 2003 mdash Devonian radiolarian faunas from south China Micropaleontology 49 (2) 127-145 httpsdoiorg102113492127

WAnG Y J chenG J f amp zhAnG Y d 2008 mdash New radio-larian genera and species of Heituao Formation (Ordovician) in the Kuruktag Region Xinjiang Acta Palaeontologica Sinica 47 (4) 393-404

WAnG Y J chenG J f amp zhAnG Y d 2010 mdash Fossil preser-vation modes and functional discussion of the basic structures of a new radiolarian genus Gansuceratoikiscum Wang gen nov Acta Palaeontologica Sinica 49 (4) 472-476

WAnG Y J amp kuAnG G 1993 mdash Early Carboniferous radiolar-ians from Qinzhou southeastern Guangxi Acta Micropalae-ontologica Sinica 10 (3) 275-287

WAnG Y J amp li J X 1994 mdash Discovery of the Follicucullus bipartitus-F charveti radiolarian assemblage zone and geological significance Acta Micropalaeontologica Sinica 11 (2) 201-212

WAnG Y J amp luo h 2009 mdash Upper Devonian (Frasnian) Hele-nifore robustum Radiolarian Fauna from the Bazhai Village in Ziyun County Guizhou Province Acta Micropalaeontologica Sinica 26 (2) 129-138

WAnG Y J luo h amp YAnG Q 2012 mdash Late Paleozoic Radio-larians in the Qinfang Area Southeast Guangxi University of Science and Technology of China Press Anhui 123 p

WebbY b d amp bloM W 1986 mdash The first well-preserved radio-larians from the Ordovician of Australia Journal of Paleontology 60 (1) 145-157 httpsdoiorg101017S0022336000021594

Won M-z 1983 mdash Radiolarien aus dem Unterkarbon des Rheinischen Schiefergebirges (Deutschland) Palaeontographica Abteilung A Paleozoologie-Stratigraphie 182 (4-6) 116-175

Won M-z 1990 mdash Lower Carboniferous radiolarians from Riescheid (Germany) Journal of the Paleontological Society of Korea 6 (2) 111-143

Won M-z 1991a mdash Phylogenetic study of some species of genus Albaillella Deflandre 1952 and a radiolarian zonation in the Rheinische Schiefergebirge West Germany Journal of the Paleontological Society of Korea 7 (1) 13-25

Won M-z 1991b mdash Lower Carboniferous radiolarians from siliceous boulders in Western Germany Journal of the Paleon-tological Society of Korea 7 (1) 77-106

Won M-z 1992 mdash The biostratigraphic age of the transitional beds of the Riescheid section Germany by means of the radiolarian fauna Journal of Science Pusan National University 53 195-210

Won M-z 1997a mdash Review of family Entactiniidae (Radiolaria) and taxonomy and morphology of Entactiniidae in the late Devo-nian (Frasnian) Gogo Formation Australia Micropaleontology 43 (4) 333-369 httpsdoiorg1023071485930

Won M-z 1997b mdash The proposed new radiolarian subfamily Retentactiniinae (Entactiniidae) from the late Devonian (Frasnian) Gogo Formation Australia Micropaleontology 43 (4) 371-418 httpsdoiorg1023071485931

Won M-z 1998 mdash A Tournaisian (Lower Carboniferous) radio-larian zonation and radiolarians of the A pseudoparadoxa zone from Oese (Rheinische Schiefergebirge) Germany Journal of the Korean Earth Science Society 19 (2) 216-259

Won M-z amp beloW r 1999 mdash Cambrian Radiolaria from the Georgina Basin Queensland Australia Micropaleontology 45 (4) 325-363 httpsdoiorg1023071486119

Won M-z blodGett r b amp nestor v 2002 mdash Llandover-ian (Early Silurian) radiolarians from the Road River Formation of east-central Alaska and thenew family Haplotaeniatumidae Journal of Paleontology 76 (6) 941-964 httpsdoiorg101017S0022336000057796

Won M-z amp iAMs W J 2002 mdash Late Cambrian radiolarian faunas and biostratigraphy of the Cow Head Group western Newfoundland Journal of Paleontology 76 (1) 1-33 httpsdoiorg101017S0022336000017315

Won M-z amp seo e-h 2010 mdash Lower Carboniferous radiolar-ian biozones and faunas from Bergishes land Germany Journal of the Palaeontological Society of Korea 26 (2) 193-269

Won M-z amp iAMs W J 2011 mdash Earliest Arenig Radiolarians from the Cow Head Group Western Newfoundland Journal of Paleontology 85 (1) 156-177 httpsdoiorg10166610-1021

Won M-z amp iAMs W J 2013 mdash Early Ordovician (early Arenig) radiolarians from the Cow Head Group and review of the Lit-tle Port Complex fauna Western Newfoundland Palaeoworld 22 (1-2) 10-31 httpsdoiorg101016jpalwor201211001

Won M-z amp iAMs W J 2015a mdash EarlyMiddle Arenig (Late Floian) Radiolarian Faunal Assemblages from Cow Head Group Western Newfoundland Palaeontographica Abteilung A 304 (1-6) 1-63

Won M-z amp iAMs W J 2015b mdash Review of the Beothuka terranova assemblage and characteristics of the middle Arenig (Ordovician latest Floian) radiolarian assemblage from the Cow Head Group Newfoundland Neues Jahrbuch fuumlr Geolo-gie und Palaumlontologie-Abhandlungen 278 (1) 1-21 httpsdoiorg101127njgpa20150513

Won M-z iAMs W J amp reed k 2005 mdash Earliest Ordovician (Early to Middle Tremadocian) radiolarian faunas of the Cow Head Group Western Newfoundland Journal of Paleontology 79 (3) 433-459 httpsdoiorg1016660022-3360(2005)079lt0433EOETMTgt20CO2

Won M-z iAMs W J amp reed k 2007 mdash Late Tremado-cian Radiolarian Faunas and Biostratigraphy of the Cow Head Group Western Newfoundland Canada Journal of the Korean Earth Science Society 28 (4) 497-540 httpsdoiorg105467JKESS2007284497

WonGAnAn n amp cAridroit M 2005 mdash Middle and Upper Devonian radiolarian faunas from Chiang Dao area Chiang Mai province northern Thailand Micropaleontology 51 (1) 39-57 httpsdoiorg10211351139

Wu h r amp li h s 1989 mdash Carboniferous and Permian Radiolaria in the Menglian area Western Yunnan Acta Micropalaeontologica Sinica 6 (4) 337-343

Wu J fenG Q Gui b amp liu G 2010 mdash Some New Radiolarian Species and Genus from Upper Permian in Guangxi Province South China Journal of Paleontology 84 (5) 879-894 httpsdoiorg10166609-0571


Aitchison J C et al

XiAn W amp zhAnG n 1998 mdash Early to Middle Permian radiolar-ians from the Kuhfeng Formation in southeastern Guangxi South China Earth Science Journal of the Association for the Geological Collaboration in Japan 52 (3) 188-202

YAo A amp kuWAhArA k 1999 mdash Permian and Triassic radiolarian assemblages from the Yangzi Platform in YAo A ezAki Y hAo W amp WAnG X (eds) Biotic and Geological Development of the Paleo-Tethys in China Peking University Press 1-16

zhAnG k Wu s amp liu Y 1992 mdash Radiolarians and Conodonts from the Dalong Formation at Hushan of Nanjing and their Facieological significance Earth Science Journal of China Uni-versity of Geosciences 17 (3) 295-300

zhAnG n henderson c M XiA W WAnG G amp shAnG h

2010 mdash Conodonts and radiolarians through the Cisuralian-Guadalupian boundary from the Pingxiang and Dachongling sections Guangxi region South China Alcheringa 34 (2) 135-160 httpsdoiorg10108003115510903523292

zhAnG n XiA W donG Y amp shAnG h 2008 mdash Conodonts and radiolarians from pelagic cherts of the Frasnian-Famennian boundary interval at Bancheng Guangxi China Global recogni-tion of the upper Kellwasser event Marine Micropaleontology 67 (1-2) 180-190 httpsdoiorg101016jmarmicro200801011

zhurAlevA i t 1975 mdash Kembriyskaya Fauna i Flora khrebta Dzhagdyi (Dalrsquoniy Vostok) [The Cambrian fauna and flora of the Dzahagda Ridge] Trudyi Instituta Geologii i Geogiziki Sibiriskoe Otlodelenie Akademiya Nauk SSSR 226 52-56

Submitted on 25 October 2016 accepted on 17 March 2017

published on 29 September 2017



Original combination Ord

er

Stat

usTy

pe Original description(authorships of cited taxa)

Age originally assigned to n-b type Stage

Stratigraphic unit from which n-b type was described

AcanthopyleAcanthopyle antiqua E nn ndash Deflandre 1960 216 Lower

CarboniferousVisean Cabriegraveres Heacuterault France

Acanthopyle antiqua E ndash T Deflandre 1963 3981 Lower Carboniferous

Visean Cabriegraveres Heacuterault France

AcanthosphaeraAcanthosphaera antiqua ndash nd ndash Hinde 1890 51 Middle

OrdovicianDarriwilian Southern Uplands Peeblesshire

Scotland UKAcanthosphaera australis ndash nd ndash Hinde 1899a 48 Middle Devonian Givetian Yarramie Formation Tamworth

Belt northern NSW AustraliaAcanthosphaera

cyclocerandash nn ndash Deflandre 1960 216 Lower


Acanthosphaera dodecaspinosa

ndash nd ndash Aberdeen 1940 137 Upper Devonian Famennian Caballos Formation Marathon Basin Texas USA

Acanthosphaera entactinia

ndash nd ndash Ruumlst 1892 147 Lower Carboniferous

ndash Kieselschiefer Harz Germany

Acanthosphaera etheridgei

ndash nd ndash Hinde 1899a 49 Middle Devonian Givetian Yarramie Formation Tamworth Belt northern NSW Australia

Acanthosphaera grandispinosa


Acanthosphaera hirsuta ndash nd ndash Aberdeen 1940 138 Upper Devonian Famennian Caballos Formation Marathon Basin Texas USA

Acanthosphaera laxa ndash nd ndash Hinde amp Fox 1895 637 Carboniferous Serpukhovian-Bashikirian

Codden Hill bed Devon England UK

Acanthosphaera macracantha

ndash nd ndash Ruumlst 1892 147 Lower Silurian ndash Cabriegraveres Heacuterault France

Acanthosphaera microspinosa


Acanthosphaera minuta ndash nd ndash Ruedemann amp Wilson 1936 1571

Lower Ordovician Floian Deepkill shale Mt Merino Columbia County New York USA

Acanthosphaera perspinosa

ndash nd ndash Ruedemann amp Wilson 1936 1571


Acanthosphaera polygonophora

ndash nn ndash Deflandre 1960 216 Lower Carboniferous


Acanthosphaera robusta ndash nd ndash Ruedemann amp Wilson 1936 1571

Lower Ordovician Floian Normanskill chert Ghent Columbia County New York USA

Aciferopylorum (254)Aciferopylorum

admirandumE ndash ndash Nazarov amp Ormiston 1990 22 Lower Silurian ndash Sakmarsk Formation Bolrsquoshoy

Abimevo Village Bashkortostan Region S Urals Russia

Aciferopylorum admirandum

S inv ndash Nazarov amp Ormiston 1993 43 Lower Silurian ndash Sakmarsk Formation Bolrsquoshoy Abimevo Village Bashkortostan Region S Urals Russia


E nn Th Ts

Nazarov 1988 71 Middle Silurian Homerian Maksyutov Complex Tarangul River 10 km N of Kosistek Village S Urals Russia

AcrosphaeraAcrosphaera xinhui ndash nn ndash Li 1995 332 Middle

OrdovicianDapingian Qingshuigou-Baijingsi Complex

Baijingsi Quilian County Qinghai China

Acrosphaera glitzii ndash nd ndash Ruumlst 1892 140 Lower Carboniferous


Acrosphaera herzynica ndash nd ndash Ruumlst 1892 140 Upper Devonian ndash SchaebenholzHarz Mountains Elbingerode Germany

ActinommaActinomma

schaebenholzianumndash nd ndash Ruumlst 1892 148 Upper Devonian ndash SchaebenholzHarz Mountains

Elbingerode Germany

APPENDIX

Appendix 1 mdash Inventory of Paleozoic radiolarian species (1880-2016) Abbreviations orders Al Albaillellaria Ar Archaeospicularia E Entactinaria L Latentifis-tularia N possible Nassellaria S Spumellaria IS incertae sedis Status nd nomina dubium nn nomina nuda nr not radiolarians ss sponge spicules inv invalid Types species T valid type species Td types that are nomina dubia Th types that are homonyms Ti type species that are invalid names Ts types that are synonyms n-b type name-bearing type The numbers in brackets after genus names correspond to the figurersquos numbers in Caridroit et al 2017 (this issue) XLS version of this list is available on the website of Geodiversitas


Aitchison J C et al


er

Stat

usTy




Adamas (289)Adamas cathedrarius IS

(OampF)nc T

ThAfanasieva 2000a 85 Upper Devonian Famennian Zadonsk Formation W

Lekkeyaginsk-65 Saremboi-Lekkeyaga Timan-Pechora Basin Russia

Afanasievella (76)Afanasievella apachensis E ndash T Nestell amp Nestell 2010 23 Middle Permian Capitanian Bell Canyon Formation Apache

Mountains Culberson County W Texas USA

Aitchisonellum (48)Aitchisonellum aspinosus Ar ndash Ts Won in Won amp Below 1999 356 Middle Cambrian ndash Inca Formation Georgina Basin

Queensland Australia

Albaillella (1)Albaillella amplificata Al ndash Ts Nazarov amp Ormiston 1985 46 Upper Carboniferous Gzhelian E of Saraktash Nikol Village Ural

River Bashkirsk Region Urals Russia

Albaillella amygdalaspinosa

Al ndash ndash Schwartzapfel amp Holdsworth 1996 59

Lower Carboniferous

Visean Sycamore Limestone southern Arbuckle Mountains Murray County Oklahoma USA

Albaillella angusta Al ndash ndash Kuwahara 1999 91 Upper Permian Wuchia pingian Funafuseyama Unit Tamba-Mino-Ashio Belt Mino Valley Seki City Gifu Japan

Albaillella apporrecta Al ndash ndash Nazarov amp Ormiston 1985 47 Lower Permian Artinskian Kandurov Formation Donskoye Ural River Orenburgskaya Region S Urals Russia

Albaillella asymmetrica Al ndash ndash Ishiga amp Imoto in Ishiga et al 1982b 276

Lower Permian Kungurian Funafuseyama Unit Tamba-Mino-Ashio Belt Ashimi-dani section Ukyo-ku Kyoto Japan

Albaillella bialata Al ndash ndash Wu amp Feng in Wu et al 2010 882Upper Permian Changhsingian Dalong Formation Dongpan section Dongpan Village Guangxi Zhuang Region China

Albaillella bipennata Al ndash ndash Cheng 1986 50 Upper Carboniferous

Bashkirian Johns Valley Shale Pittsburg County eastern Oklahoma USA

Albaillella brauni Al ndash ndash Won 1998 224 Lower Carboniferous

Tournaisian Oese Rheinische Schiefergebirge Germany

Albaillella brushensis Al ndash ndash Cheng 1986 51 Upper Carboniferous


Albaillella cartalla Al ndash Ts Ormiston amp Lane 1976 171 Lower Carboniferous

Tournaisian Sycamore Limestone southern Arbuckle Mountains Oklahoma USA

Albaillella cavitata Al ndash ndash Kuwahara 1999 90 Upper Permian Wuchiapingian Funafuseyama Unit Tamba-Mino-Ashio Belt Mino Valley Seki City Gifu Japan

Albaillella cornuta Al ndash ndash Deflandre 1952 872 Lower Carboniferous

Tournaisian Cabriegraveres Heacuterault France

Albaillella crenulata Al ndash ndash Won 1991a 18 Lower Carboniferous

Visean Rheinisches Schiefergebirge Frankenwald Germany

Albaillella cylindra Al ndash ndash Schwartzapfel amp Holdsworth 1996 53

Lower Carboniferous

Visean Caney Shale Arbuckle Mountains Phillips Creek Section Carter County Oklahoma USA

Albaillella deflandrei Al ndash ndash Gourmelon 1987 85 Lower Carboniferous


Albaillella dellensis Al ndash ndash Park amp Won 2012 42 Lower Carboniferous

Tournaisian Woodman Formation South Lakeside Mts Tooele County Utah USA

Albaillella demenita Al ndash ndash Nazarov in Isakova amp Nazarov 1986 112

Upper Carboniferous

Gzhelian Saraktash Nikol Village Southern Urals Russia

Albaillella eliasi Al ndash ndash Schwartzapfel amp Holdsworth 1996 56

Lower Carboniferous

Serpukhovian Goddard Formation Oil Creek section Johnston County Oklahoma USA

Albaillella excelsa Al ndash ndash Ishiga Kito amp Imoto 1982a 17 Upper Permian Changhsingian Funafuseyama Unit Tamba-Mino-Ashio Belt Ubara section Fukuchiyama City Kyoto Japan

Albaillella fida Al ndash ndash Jin amp Feng in Jin et al 2007 14 Upper Permian Changhsingian Dalong Formation Dongpan section Dongpan Village Guangxi Zhuang Region China

Albaillella flabellata Al ndash ndash Jin amp Feng in Jin et al 2007 11 Upper Permian Changhsingian Dalong Formation Dongpan section Dongpan Village Guangxi Zhuang Region China

Appendix 1 mdash Continuation of the inventory of Paleozoic radiolarian species (1880-2016)




er

Stat

usTy




Albaillella (1) continuationAlbaillella flexa Al ndash ndash Kuwahara in Kuwahara amp

Sakamoto 1992 39Upper Permian Changhsingian Funafuseyama Unit Tamba-Mino-

Ashio Belt Mino Valley Seki City Gifu Japan

Albaillella flexiloqua Al ndash ndash Nazarov in Isakova amp Nazarov 1986 111

Upper Carboniferous


Albaillella foremanae Al ndash ndash Cornell amp Simpson 1985 273 Middle Permian Roadian Bone Spring Formation Guadalupe Mts Culbertson County W Texas USA

Albaillella furcata Al ndash ndash Won 1983 126 Lower Carboniferous


Albaillella gordoni Al ndash ndash Schwartzapfel amp Holdsworth 1996 77

Lower Carboniferous


Albaillella graciliforma Al ndash ndash Won amp Seo 2010 235 Lower Carboniferous

Tournaisian Bergisches Land N Westfalia Germany

Albaillella groessensi Al ndash ndash Schwartzapfel amp Holdsworth 1996 64

Lower Carboniferous


Albaillella harrisae Al ndash ndash Schwartzapfel amp Holdsworth 1996 70

Lower Carboniferous


Albaillella hassi Al ndash ndash Schwartzapfel amp Holdsworth 1996 80

Lower Carboniferous


Albaillella higginsi Al ndash ndash Schwartzapfel amp Holdsworth 1996 83

Lower Carboniferous


Albaillella hushanensis Al ndash ndash He amp Feng in He et al 2011 477

Upper Permian Changhsingian Talung Formation Hushan section Nanjing Jiangsu Region China

Albaillella indensis Al ndash ndash Won 1983 127 Lower Carboniferous


Albaillella indensis ambigua

Al ndash ndash Braun 1989 86 Lower Carboniferous

upper Tournaisian

Pebbles of siliceous shale from the lower Main-valley near Frankfurt a M Germany

Albaillella indensis brauni Al ndash ndash Feng amp Ye 1996 20 Lower Carboniferous

Tournaisian Yiliu Formation Menxin section S of Menglian SW Yunnan China

Albaillella inferioalata Al ndash ndash Nazarov in Isakova amp Nazarov 1986 113

Upper Carboniferous


Albaillella inflata Al ndash ndash Cheng 1986 52 Upper Carboniferous

Bashkirian Wesley Formation Turnpike section Pittsburg County Oklahoma USA

Albaillella ishigai Al ndash ndash Cheng 1986 53 Lower Carboniferous

Serpukhovian Sand Branch Member of Caney Formation Pittsburg County Oklahoma USA

Albaillella kayai Al ndash ndash Noble Tekin Gedik amp Pehlivan 2008 46

Lower Carboniferous

Tournaisian Baltalimani Formation Istanbul Turkey

Albaillella ladarezensis Al ndash ndash Gourmelon 1987 88 Lower Carboniferous


Albaillella lanceolata Al ndash ndash Schwartzapfel amp Holdsworth 1996 61

Lower Carboniferous


Albaillella lauta Al ndash ndash Kuwahara in Kuwahara amp Sakamoto 1992 40

Upper Permian Wuchiapingian Funafuseyama Unit Tamba-Mino-Ashio Belt Mino Valley Seki City Gifu Japan

Albaillella levis Al ndash Ts Ishiga Kito amp Imoto 1982a 17 Upper Permian Changhsingian Funafuseyama Unit Tamba-Mino-Ashio Belt Ubara section Fukuchiyama City Kyoto Japan

Albaillella longoriai Al ndash ndash Schwartzapfel amp Holdsworth 1996 58

Lower Carboniferous


Albaillella macrocephalata

Al ndash ndash Cheng 1986 54 Upper Carboniferous


Albaillella magnifica Al ndash ndash Cheng 1986 55 Upper Devonian Famennian Johns Valley Shale Pittsburg County eastern Oklahoma USA

Albaillella mameti Al ndash ndash Schwartzapfel amp Holdsworth 1996 63

Lower Carboniferous


Albaillella minuta Al ndash ndash Won amp Seo 2010 235 Lower Carboniferous


Albaillella mucronulata Al ndash ndash Nestell Pope amp Nestell 2012 230

Upper Carboniferous

Moscovian Mouse Creek Formation Excello Shale Member south-central Iowa USA



Aitchison J C et al


er

Stat

usTy




Albaillella (1) continuationAlbaillella multisegmenta Al ndash ndash Wang in Wang et al 2012 107 Lower

CarboniferousVisean Shijia Formation Bancheng

Qinzhou area Guangxi Zhuang Region SW China

Albaillella murcheyae Al ndash ndash Schwartzapfel amp Holdsworth 1996 84

Lower Carboniferous


Albaillella nazarovi Al ndash ndash Cheng 1986 55 Upper Carboniferous


Albaillella nigriniae Al ndash ndash Cheng 1986 56 Upper Carboniferous


Albaillella ouachitaensis Al ndash ndash Cheng 1986 57 Lower Carboniferous

Bashkirian Jones Valley Shale Le Flore County eastern Oklahoma USA

Albaillella palmeri Al ndash ndash Schwartzapfel amp Holdsworth 1996 67

Lower Carboniferous


Albaillella paradoxa var globosa

Al ndash ndash Deflandre 1952 872 Lower Carboniferous


Albaillella paradoxa Al ndash T Deflandre 1952 872 Lower Carboniferous


Albaillella pennata Al ndash ndash Holdsworth 1966 323 Upper Carboniferous

Bashkirian Upper Dove Valley SW Derbyshire England UK

Albaillella perforata Al ndash ndash Won 1991b 85 Lower Carboniferous

Tournaisian Boulders in Quaternary Rhine Terrace near Oberkassel Bonn

Albaillella perforata perforata

Al ndash ndash Won 1991b 86 Lower Carboniferous


Albaillella perforata uniramosa



Albaillella pessagnoi Al ndash ndash Cheng 1986 58 Upper Carboniferous


Albaillella philipsensis Al ndash ndash Schwartzapfel amp Holdsworth 1996 66

Lower Carboniferous


Albaillella prava Al ndash ndash Cheng 1986 59 Upper Carboniferous


Albaillella procera Al ndash ndash Cheng 1986 60 Upper Carboniferous


Albaillella protoforemanae

Al ndash ndash Zhang Henderson amp Xia in Zhang et al 2010 154

Lower Permian Kungurian Dachongling section Qinzhou City Guangxi S China

Albaillella protolevis Al ndash ndash Kuwahara 1999 90 Upper Permian Wuchiapingian Funafuseyama Unit Tamba-Mino-Ashio Belt Mino Valley Seki City Gifu Japan

Albaillella pseudoparadoxa

Al ndash ndash Won 1991a 21 Lower Carboniferous

Tournaisian Rheinisches Schiefergebirge Frankenwald Germany

Albaillella pseudoparadoxa spongiosa

Al ndash ndash Won 1998 225 Lower Carboniferous


Albaillella ramsbottomi Al ndash ndash Schwartzapfel amp Holdsworth 1996 74

Lower Carboniferous


Albaillella riescheidensis Al ndash ndash Won 1990 115 Lower Carboniferous

Visean Riescheid Section Wuppertal-Barmen Germany

Albaillella robusta Al ndash ndash Cheng 1986 60 Upper Carboniferous


Albaillella rockensis Al ndash ndash Cheng 1986 61 Lower Carboniferous


Albaillella saltatoria Al ndash ndash Cheng 1986 62 Upper Carboniferous


Albaillella sandbergi Al ndash ndash Won 1991a 23 Lower Carboniferous


Albaillella saundersi Al ndash ndash Schwartzapfel amp Holdsworth 1996 82

Lower Carboniferous


Albaillella shijaensis Al ndash ndash Wang in Wang et al 2012 107 Lower Carboniferous

Tournaisian Shijia Formation Bancheng Qinzhou area Guangxi Zhuang Region SW China





er

Stat

usTy




Albaillella (1) continuationAlbaillella sinuata Al ndash ndash Ishiga amp Watase in Ishiga et al

1986 126Middle Permian Capitanian Nishiki Group Sangun-Chugoku

Belt Muikaichi area JapanAlbaillella sinuosa Al ndash ndash Won amp Seo 2010 241 Lower

CarboniferousTournaisian Bergisches Land N Westfalia

Germany

Albaillella spinosa Al ndash ndash Cheng 1986 63 Upper Carboniferous


Albaillella tela Al ndash ndash Schwartzapfel amp Holdsworth 1996 73

Lower Carboniferous


Albaillella thomasi Al ndash ndash Braun 1990 92 Lower Carboniferous

upper Tournaisian

Rheinisches Schiefergebirge Frankenwald Germany

Albaillella tooelensis Al ndash ndash Park amp Won 2012 44 Lower Carboniferous


Albaillella triangularis Al ndash ndash Ishiga Kito amp Imoto 1982a 17 Upper Permian Changhsingian Funafuseyama Unit Tamba-Mino-Ashio Belt Ubara section Fukuchiyama City Kyoto Japan

Albaillella tuboforma Al ndash ndash Won 1991b 91 Lower Carboniferous


Albaillella tumida Al ndash ndash Cheng 1986 64 Upper Carboniferous


Albaillella turgida Al ndash ndash Cheng 1986 65 Upper Carboniferous


Albaillella u-forma reflexa Al nc ndash Ling amp Forsythe 1987 257 Lower Permian Sakmarian Denaro Complex Magallanes y Antartica Chilena Chile

Albaillella uncus Al ndash ndash Won 1983 127 Lower Carboniferous


Albaillella undulata Al ndash ndash Deflandre 1952 872 Lower Carboniferous


Albaillella unusalata Al ndash ndash Cheng 1986 66 Upper Carboniferous


Albaillella utahensis Al ndash ndash Park amp Won 2012 44 Lower Carboniferous


Albaillella xiaodongensis Al ndash ndash Wang in Wang et al 1994 185 Lower Permian Kungurian Dachongling section Qinzhou City Guangxi S China

Albaillella yamakitai Al ndash ndash Kuwahara 1999 89 Upper Permian Wuchiapingian Funafuseyama Unit Tamba-Mino-Ashio Belt Mino Valley Seki City Gifu Japan

Albaillella yaoi Al ndash ndash Kuwahara 1999 92 Upper Permian Changhsingian Funafuseyama Unit Tamba-Mino-Ashio Belt Mino Valley Seki City Gifu Japan

Albaillella yaoi longa Al ndash ndash Jin amp Feng in Jin et al 2007 11 Upper Permian Changhsingian Dalong Formation Dongpan section Dongpan Village Guangxi Zhuang Region China

Albaillella protractosegmentata

Al ndash ndash Nazarov in Isakova amp Nazarov 1986 113

Upper Carboniferous


Albaillella zhui Al ndash ndash Feng amp Liu 1993 555 Upper Permian Changhsingian Jinghong Formation NE of Jinghong City Xishuang-bannadai County SW Yunnan China

Altaiesphaera (49)Altaiesphaera

acanthophora Ar ndash T Obut amp Iwata 2000 35 Lower Cambrian Stage 4 Shashkunar Formation Gorny

Altai SW Siberia RussiaAltaiesphaera

sparsispinosaAr ndash ndash Obut amp Iwata 2000 35 Lower Cambrian Stage 4 Shashkunar Formation Gorny

Altai SW Siberia Russia

AmphibrachiumAmphibrachium bacillum ndash nd ndash Ruumlst 1892 169 Upper Devonian ndash SchaebenholzHarz Mountains

Elbingerode GermanyAmphibrachium

desecatumndash nd ndash Ruumlst 1892 169 Upper Permian ndash Sosio Valley Area Western Sicily

ItalyAmphibrachium

devoniensendash nd ndash Ruumlst 1892 169 Upper Devonian ndash SchaebenholzHarz Mountains

Elbingerode Germany



Aitchison J C et al


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Amphibrachium (continuation)Amphibrachium

inaequalendash nd ndash Ruumlst 1892 169 Lower Devonian ndash Southern Urals Russia

Amphibrachium pulchellum

ndash nd ndash Ruumlst 1892 169 Upper Devonian ndash SchaebenholzHarz Mountains Elbingerode Germany

AmphymeniumAmphymenium alienum E nd T Ruumlst 1892 170 Upper Devonian ndash SchaebenholzHarz Mountains

Elbingerode GermanyAmphymenium krautii E ndash ndash Ruumlst 1892 170 Upper Devonian ndash SchaebenholzHarz Mountains

Elbingerode Germany

AmpullaAmpulla tubulata nr is Ti Rudenko amp Panasenko 1990a

123Upper Permian Changhsingian Yastrebov Formation Taukha

Terrane Primoriya Russia

Anakrusa (335)Anakrusa conspersa nr ss ndash Nazarov 1977 169 Middle

OrdovicianDarriwilian Bestomaksk Formation Chagan

River Chingiz Range Kazakhstan

Anakrusa miriacantha nr ss T Nazarov 1977 169 Middle Ordovician

Darriwilian Bestomaksk Formation Chagan River Chingiz Range Kazakhstan

Antygopora (234)Antygopora bella S ndash ndash Maletz amp Bruton 2007 271 Lower Ordovician Floian Valhallfonna Formation

Buldrebreen arm Ny Frieslan Spitsbergen Norway

Antygopora compacta S ndash ndash Maletz amp Bruton 2007 271 Lower Ordovician Floian Valhallfonna Formation Buldrebreen arm Ny Frieslan Spitsbergen Norway

Antygopora intermediata S ndash ndash Won amp Iams 2015b 10 Lower Ordovician Floian Cow Head Group Western Newfoundland Canada

Antygopora irregularia S ndash ndash Maletz 2007 80 Lower Ordovician Floian Cow Head Group Western Newfoundland Canada

Antygopora labyrinthina S ndash ndash Maletz amp Bruton 2007 273 Lower Ordovician Floian Valhallfonna Formation Buldrebreen arm Ny Frieslan Spitsbergen Norway

Antygopora littleportensis S ndash ndash Won amp Iams 2013 15 Lower Ordovician Floian Little Port Complex Newfoundland Canada

Antygopora littleportensis littleportensis

S ndash ndash Won amp Iams 2013 15 Lower Ordovician Floian Little Port Complex Newfoundland Canada

Antygopora littleportensis sigillata


Antygopora microspina S ndash ndash Maletz amp Bruton 2007 271 Lower Ordovician Floian Valhallfonna Formation Buldrebreen arm Ny Frieslan Spitsbergen Norway

Antygopora ordovicica S ndash T Maletz amp Bruton 2005 5 Lower Ordovician Floian Valhallfonna Formation Buldrebreen arm Ny Frieslan Spitsbergen Norway

Apophysiactinia (77)Apophysiactinia

duplospinulanr Calc-

areandash Won 1997a 362 Upper Devonian Frasnian Gogo Formation Pillara Range

Canning Basin Western Australia

Apophysiactinia iamsi nr Calc-area

ndash Park amp Won 2012 64 Lower Carboniferous


Apophysiactinia testacea nr Calc-area

ndash Won 1997a 364 Upper Devonian Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Apophysiactinia trispinula nr Calc-area

Tnr Won 1997a 364 Upper Devonian Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Apophysisphaera (78)Apophysisphaera

elegansE ndash ndash Won 1997b 374 Upper Devonian Frasnian Gogo Formation Pillara Range


Apophysisphaera parva E ndash ndash Won 1997b 375 Upper Devonian Frasnian Gogo Formation Pillara Range Canning Basin Western Australia





er

Stat

usTy




Apophysisphaera (78) continuationApophysisphaera

membranaceaE ndash ndash Won 1997b 376 Upper Devonian Frasnian Gogo Formation Pillara Range


Apophysisphaera rosae E ndash ndash Won 1997b 376 Upper Devonian Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

ArbustrumArbustrum congregatum nr Calc-

areandash Wang 1989 155 Lower Cambrian ndash Niutitang Formation Sandu

district Guizhou Province China

Arbustrum inflata nr Calc-area

ndash Wang 1989 146 Lower Cambrian ndash Damiao Formation Erlongping Group Eastern Qinling ophiolite belt Henan Province China

Arbustrum luxurians nr Calc-area


Arbustrum strigosum nr Calc-area

Tnr Wang 1989 146 Lower Cambrian ndash Damiao Formation Erlongping Group Eastern Qinling ophiolite belt Henan Province China

Archaeocenosphaera (50)Archaeocenosphaera

muricata Ar ndash Ts Obut amp Iwata 2000 34 Lower Cambrian Stage 4 Shashkunar Formation Gorny


Archaeopyramisa (203)Archaeopyramisa

concavaL ndash ndash Cheng 1986 185 Lower

CarboniferousBashkirian Jones Valley Shale Le Flore

County eastern Oklahoma USAArchaeopyramisa

convexaL ndash ndash Cheng 1986 186 Lower

CarboniferousSerpukhovian Sand Branch Member of Caney

Formation Pittsburg County Oklahoma USA

Archaeopyramisa globosa

L ndash ndash Cheng 1986 187 Lower Carboniferous


Archaeopyramisa haeckeli

L ndash Ts Cheng 1986 186 Lower Carboniferous


Archaeopyramisa robusta L ndash ndash Cheng 1986 187 Lower Carboniferous


Archaeopyramisa talihinaensis



ArchaeospongoprunumArchaeospongoprunum

mengiS ndash ndash Feng in Feng et al 2006a 35 Upper Permian Changhsingian Dalong Formation Dongpan

section Dongpan Village Guangxi Zhuang Region China

Archeoentactinia (43)Archeoentactinia

angulataAr ndash ndash Won in Won amp Below 1999 332 Middle Cambrian ndash Inca Formation Georgina Basin

Queensland AustraliaArcheoentactinia delicata Ar ndash ndash Won in Won amp Below 1999 332 Middle Cambrian ndash Inca Formation Georgina Basin

Queensland AustraliaArcheoentactinia

hexactiniaAr ndash ndash Won in Won amp Below 1999 333 Middle Cambrian ndash Inca Formation Georgina Basin


incaensisAr ndash T Won in Won amp Below 1999 333 Middle Cambrian ndash Inca Formation Georgina Basin


tetractiniaAr ndash ndash Won in Won amp Below 1999 335 Middle Cambrian ndash Inca Formation Georgina Basin


Archeoproventocitum (280)Archeoproventocitum

nudiformumIS ndash T Won Iams amp Reed 2007 534 Lower Ordovician lower-middle

TremadocianCow Head Group Western

Newfoundland CanadaArcheoproventocitum

retiformumIS ndash ndash Won Iams amp Reed 2007 534 Lower Ordovician lower-middle


Newfoundland Canada

ArchidiscusArchidiscus lens ndash nd ndash Ruumlst 1892 166 Lower

Carboniferousndash Kieselschiefer Harz Germany



Aitchison J C et al


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Archocyrtium (219)Archocyrtium amoenus N ndash ndash Afanasieva amp Amon 2011 1513 Upper Devonian lower

FamennianPolar Urals Palnik Yu River Russia

Archocyrtium angulosum N ndash ndash Deflandre 1973d 150 Lower Carboniferous


Archocyrtium breviora N ndash ndash Park amp Won 2012 61 Lower Carboniferous


Archocyrtium callimorphum

N ndash ndash Braun 1989 90 Lower Carboniferous

upper Tournaisian


Archocyrtium castuligerum

N ndash Deflandre 1972a 3539 Lower Carboniferous



N nn Ts Deflandre 1972b 15 Lower Carboniferous


Archocyrtium clinoceros N ndash ndash Deflandre 1973d 151 Lower Carboniferous


Archocyrtium coronaesimile

N ndash ndash Won 1983 128 Lower Carboniferous


Archocyrtium delicatum N ndash ndash Cheng 1986 123 Lower Carboniferous

Tournaisian Woodford Formation TI Valley section Pittsburg County Oklahoma USA

Archocyrtium diductum N ndash ndash Deflandre 1973d 150 Lower Carboniferous


Archocyrtium dilatipes N ndash ndash Deflandre 1973d 150 Lower Carboniferous


Archocyrtium effingi N ndash ndash Kiessling amp Tragelehn 1994 231Upper Devonian Famennian Frankenwald north Bavaria Germany

Archocyrtium eupectum N ndash ndash Braun 1989 90 Lower Carboniferous

upper Tournaisian


Archocyrtium ferreum N ndash ndash Braun 1989 90 Lower Carboniferous

upper Tournaisian


Archocyrtium formosum N ndash ndash Cheng 1986 124 Lower Carboniferous


Archocyrtium lagabriellei N ndash ndash Gourmelon 1987 115 Lower Carboniferous


Archocyrtium ludicrum N ndash ndash Deflandre 1973d 150 Lower Carboniferous


Archocyrtium medium N ndash ndash Liu amp Hao 2006 649 Upper Devonian Famennian Tielimaitidaban section SW Tianshan Xinjiang China

Archocyrtium menglianense

N ndash ndash Wu in Wu amp Li 1989 341 Lower Carboniferous

Visean Banshun section Menglian County Yunnan Province China

Archocyrtium obesum N ndash ndash Cheng 1986 124 Lower Carboniferous


Archocyrtium ormistoni N ndash ndash Cheng 1986 125 Upper Devonian Famennian Woodford Formation TI Valley section Pittsburg County Oklahoma USA

Archocyrtium parvispina N ndash ndash Park amp Won 2012 62 Lower Carboniferous


Archocyrtium parvum N ndash ndash Deflandre 1972a 3536 Lower Carboniferous

Visean Breacutetignolles-sur-mer Vendeacutee Pays de la Loire France

Archocyrtium parvum N nn ndash Deflandre 1972b 15 Lower Carboniferous


Archocyrtium petrushevskaye

N ndash ndash Deflandre 1973d 151 Lower Carboniferous


Archocyrtium procerum N ndash ndash Cheng 1986 126 Upper Devonian Famennian Woodford Formation TI Valley section Pittsburg County Oklahoma USA

Archocyrtium pulchrum N ndash ndash Braun 1990 126 Lower Carboniferous

upper Tournaisian


Archocyrtium riedeli N nn T Deflandre 1960 216 Lower Carboniferous


Archocyrtium riedeli N nn ndash Deflandre 1972a 3539 Lower Carboniferous






er

Stat

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Archocyrtium (219) continuationArchocyrtium riedeli N ndash ndash Deflandre 1972b 15 Lower


Archocyrtium sashidai N ndash ndash Feng in Feng et al 2004b 383 Lower Carboniferous

Visean Shan-Thai terrane Chiang Mai NW Thailand

Archocyrtium shijiaensis N ndash ndash Wang in Wang et al 2012 112 Lower Carboniferous


Archocyrtium strictum N ndash ndash Deflandre 1973d 151 Lower Carboniferous


Archocyrtium tersae N ndash Deflandre 1972a 3536 Lower Carboniferous


Archocyrtium tersae N nn ndash Deflandre 1972b 15 Lower Carboniferous


Archocyrtium typicum N ndash ndash Cheng 1986 126 Upper Devonian Famennian Woodford Formation TI Valley section Pittsburg County Oklahoma USA

Archocyrtium validum N ndash ndash Cheng 1986 127 Lower Carboniferous


Archocyrtium venustum N ndash ndash Cheng 1986 128 Lower Carboniferous


Archocyrtium wonae N ndash ndash Cheng 1986 128 Lower Carboniferous


Archocyrtium babini N ndash ndash Gourmelon 1986 189 Lower Carboniferous

Tournaisian Hautes-Pyreacuteneacutees France

Arcoclathrata (131)Arcoclathrata alekseevi E ndash ndash Afanasieva amp Amon 2016 219 Lower Permian Sakmarian Kondurovka section Sakmara

River Southern Urals Russia

Areolicaudatus (181)Areolicaudatus elongatus L nn ndash Feng 1992 55 Upper Permian Changhsingian Papai Formation of Changning-

Menglian Belt Papai Village Cangyuan County Yunnan China

Areolicaudatus semiglobosus

L nn ndash Feng 1992 55 Upper Permian Changhsingian Papai Formation of Changning-Menglian Belt Papai Village Cangyuan County Yunnan China


L ndash T Feng amp Liu 1993 545 Upper Permian Changhsingian Muyinhe Formation Nanpan Lancang County Yunnan China

Arrectoalatus (276)Arrectoalatus bicorniger IS ndash ndash Nazarov in Isakova amp Nazarov

1986 124Upper

CarboniferousGzhelian Saraktash Nikol Village Southern

Urals RussiaArrectoalatus cernuus IS ndash T Nazarov amp Ormiston 1985 49 Upper

CarboniferousGzhelian E of Saraktash Nikol Village Ural


Arrectoalatus eximius IS ndash ndash Nazarov in Isakova amp Nazarov 1986 123

Upper Carboniferous


Arrectoalatus uncinatus IS ndash ndash Rudenko amp Panasenko 1990a 122

Upper Permian Changhsingian Yastrebov Formation Taukha Terrane Primoriya Russia

Arrhiniella (132)Arrhiniella surrecta ndash nn

-ncndash Li 1995 332 Middle



Aspiculum (284)Aspiculum angulatum IS ndash ndash Won Iams amp Reed 2005 441 Lower Ordovician lower-middle


Newfoundland CanadaAspiculum densum IS ndash ndash Won Iams amp Reed 2007 507 Lower Ordovician lower-middle


Newfoundland CanadaAspiculum eccentricum IS ndash T Won Iams amp Reed 2005 441 Lower Ordovician lower-middle


Newfoundland CanadaAspiculum gigantium IS ndash ndash Won Iams amp Reed 2007 511 Lower Ordovician lower-middle


Newfoundland CanadaAspiculum jamesi IS ndash ndash Won Iams amp Reed 2007 513 Lower Ordovician lower-middle


Newfoundland Canada



Aitchison J C et al


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Aspiculum (284) continuationAspiculum jamesi

parajamesiIS ndash ndash Won amp Iams 2011 163 Lower Ordovician Floian Cow Head Group Western

Newfoundland CanadaAspiculum multistratum IS ndash ndash Won Iams amp Reed 2007 515 Lower Ordovician lower-middle


Newfoundland Canada

Astroentactinia (79)Astroentactinia

biaciculataE ndash ndash Nazarov 1975 84 Upper Devonian Frasnian Egindinsk Formation Aitpaika

River N Mugodazhar S Urals Kazakhstan

Astroentactinia capitanensis

E ndash ndash Nestell amp Nestell 2010 20 Middle Permian Capitanian Bell Canyon Formation Apache Mountains Culberson County W Texas USA

Astroentactinia crassata E ndash ndash Nazarov 1975 85 Upper Devonian Frasnian Egindinsk Formation Aitpaika River N Mugodazhar S Urals Kazakhstan

Astroentactinia deorsiacus

E ndash ndash Nazarov amp Ormiston 1993 33 Upper Devonian Famennian Aitpaika River Aktyubinsk region Southern Urals Russia


E nn ndash Nazarov 1988 129 Upper Devonian Frasnian Egindinsk Formation Aitpaika River N Mugodazhar S Urals Kazakhstan

Astroentactinia digitosa E ndash ndash Braun 1990 101 Lower Carboniferous

upper Tournaisian


Astroentactinia erinacea E ndash ndash Nazarov in Nazarov amp Popov 1980 45

Middle Ordovician


Astroentactinia formosa E ndash ndash Wang in Wang et al 2012 110 Lower Carboniferous


Astroentactinia inscita E ndash ndash Nazarov in Isakova amp Nazarov 1986 67

Upper Carboniferous


Astroentactinia luxuria E ndash ndash Nazarov amp Ormiston 1985 23 Lower Permian Artinskian Kandurov Formation Donskoye Ural River Orenburgskaya Region S Urals Russia

Astroentactinia mendosa E ndash ndash Nazarov in Isakova amp Nazarov 1986 68

Upper Carboniferous


Astroentactinia pauxilla E ndash ndash Nazarov 1975 86 Upper Devonian Frasnian Egindinsk Formation Aitpaika River N Mugodazhar S Urals Kazakhstan

Astroentactinia porosa E ndash ndash Maldonado amp Noble 2010 84 Middle Permian Capitanian Bell Canyon Formation Apache Mountains Culberson County W Texas USA

Astroentactinia praeacudura

E nn ndash Nazarov 1988 130 Upper Devonian Famenian Zapadno Valavskaya borehole 1-R Pripyat Depression Belarus

Astroentactinia radiata E ndash ndash Braun 1990 102 Lower Carboniferous

upper Tournaisian


Astroentactinia ramificans

E ndash T Nazarov 1975 87 Middle Ordovician


Astroentactinia rusaevi E nn ndash Afanasieva 1997a 221 Upper Devonian lower Frasnian Domanik Formation outcrop 1904 Lyaiol River Timan-Pechora Basin Russia

Astroentactinia rusaevi E ndash ndash Afanasieva 2000b 140 Upper Devonian middle Frasnian

Domanik Formation borehole Shuda-Yag 1003 SW of Uktha Timan-Pechora Basin Russia

Astroentactinia spatiosa E ndash ndash Braun 1990 103 Lower Carboniferous

upper Tournaisian


Astroentactinia speciosa E ndash ndash Amon amp Braun 1994 2 Lower Permian Artinskian Burtsevesky horizon Dalny Tulkas Rill Bashkortostan Region S Urals Russia

Astroentactinia stellata E ndash T Nazarov 1975 82 Upper Devonian Frasnian Egindinsk Formation Aitpaika River N Mugodazhar S Urals Kazakhstan

Astroentactinia tantilla E ndash ndash Nazarov 1975 85 Upper Devonian Frasnian Egindinsk Formation Aitpaika River N Mugodazhar S Urals Kazakhstan

Astroentactinia tikhomirovi

E ndash ndash Afanasieva 2000b 142 Upper Devonian middle Frasnian


Astroentactinia valentiniae

E ndash ndash Nazarov in Nazarov et al 1981 87

Upper Devonian Famennian Duksundinsk Formation along Duksunda River Magadan Russia





er

Stat

usTy




Astroentactinia (79) continuationAstroentactinia

vishnevskayae E ndash ndash Afanasieva 2000b 142 Upper Devonian middle

FrasnianDomanik Formation borehole

Ukhtinskaya-3B SW of Uktha Timan-Pechora Basin Russia

Astroentactinia crubellata

E ndash ndash Nazarov 1975 86 Lower Silurian ndash Sakmarsk Formation Bazyam Sakmar Orenburgskaya Region S Urals Russia

Astroentactinia echinosimilis

E ndash ndash Won 1991b 95 Lower Carboniferous


Astroentactinia mirousi E ndash ndash Gourmelon 1986 184 Lower Carboniferous


Astroentactinia miscella E ndash ndash Nazarov in Nazarov amp Popov 1980 47

Middle Ordovician


Astroentactinia radiata E ndash ndash Aitchison 1993b 118 Upper Devonian lower Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Astroentactinia stellaesimilis



AstrophacusAstrophacus cingulatus ndash nd ndash Aberdeen 1940 139 Upper Devonian Famennian Caballos Formation Marathon

Basin Texas USA

Auliela (336)Auliela annula nr ss ndash Li 1995 332 Middle



Auliela aspersa nr ss Ts Nazarov 1977 170 Middle Ordovician


Auliela taplowensis nr ss ndash Webby amp Blom 1986 151 Upper Ordovician Katian Malongulli Formation Belubula River E of Canowindra central NSW Australia

AxellipsisAxellipsis longitudinalis ndash nd ndash Ruumlst 1892 152 Upper Permian ndash Sosio Valley Area Western Sicily

Italy

Azyrtalia (337)Azyrtalia dilata nr Calc-

areaTnr Nazarov 1973b 12 Lower Cambrian Stage 2 Bograd Mountains Betenevskiy

area Khakasiya Region Russia

Batoballa (338)Batoballa communis nr Calc-

areaTnr Wang in Wang et al 2008 400 Middle

OrdovicianDapingian Heituao Formation Tarim Basin

Kuruktag region Xinjiang ChinaBatoballa longiovata nr Calc-

areandash Wang in Wang et al 2008 401 Middle


Kuruktag region Xinjiang China

Belowea (80)Belowea crassitesta E ndash ndash Won 1983 131 Lower

CarboniferousVisean Rheinisches Schiefergebirge

Frankenwald GermanyBelowea hexaculeata E ndash ndash Won 1990 121 Lower

CarboniferousVisean Riescheid Section Wuppertal-

Barmen GermanyBelowea tenuistesta E ndash ndash Won 1983 133 Lower


Frankenwald Germany

Beothuka (235)Beothuka aitchisoni S ndash ndash Won amp Iams 2011 161 Lower Ordovician Floian Cow Head Group Western

Newfoundland CanadaBeothuka echinata S ndash ndash Won amp Iams 2013 16 Lower Ordovician Floian Cow Head Group Western

Newfoundland CanadaBeothuka grosmornensis S ndash ndash Won amp Iams 2011 162 Lower Ordovician Floian Cow Head Group Western

Newfoundland CanadaBeothuka longispiniforma S ndash ndash Wang in Wang et al 2008 397 Middle


Kuruktag region Xinjiang ChinaBeothuka maletziana S ndash ndash Won amp Iams 2015b 12 Lower Ordovician Floian Cow Head Group Western

Newfoundland CanadaBeothuka multiaculeata S ndash ndash Won amp Iams 2015a 26 Lower Ordovician Floian Cow Head Group Western

Newfoundland CanadaBeothuka robusta

bispinosaS ndash ndash Won amp Iams 2013 17 Lower Ordovician Floian Cow Head Group Western

Newfoundland Canada



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Beothuka (235) continuationBeothuka robusta

intermediataS ndash ndash Won amp Iams 2013 17 Lower Ordovician Floian Cow Head Group Western

Newfoundland CanadaBeothuka spissa S ndash ndash Won amp Iams 2015a 26 Lower Ordovician Floian Cow Head Group Western

Newfoundland CanadaBeothuka spongiosa S ndash ndash Won amp Iams 2013 17 Lower Ordovician Floian Cow Head Group Western

Newfoundland CanadaBeothuka spongiosa

spongiosaS ndash ndash Won amp Iams 2013 19 Lower Ordovician Floian Cow Head Group Western


variabilisS ndash ndash Won amp Iams 2013 19 Lower Ordovician Floian Cow Head Group Western

Newfoundland CanadaBeothuka stougei S ndash ndash Won amp Iams 2011 162 Lower Ordovician Floian Cow Head Group Western

Newfoundland CanadaBeothuka terranova S ndash T Aitchison Flood amp Malpas

1998 417Lower Ordovician Tremadocian Little Port Complex at Winter

House Brook Lookout Newfoundland Canada

Beothuka terranova dimunata

S ndash ndash Won amp Iams 2015a 25 Lower Ordovician Floian Cow Head Group Western Newfoundland Canada

Beothuka terranova intermediata


Beothuka terranova mutispinosa

S ndash ndash Won amp Iams 2011 163 Lower Ordovician Floian Cow Head Group Western Newfoundland Canada

Beothuka concreta S ndash ndash Won amp Iams 2013 21 Lower Ordovician Floian Cow Head Group Western Newfoundland Canada

Beothuka stellata S ndash ndash Won amp Iams 2011 163 Lower Ordovician Floian Cow Head Group Western Newfoundland Canada

Bientactinosphaera (81)Bientactinosphaera

maslakovaeE ndash ndash Afanasieva 2000a 51 Upper Devonian middle


Shuda-Yag 1003 SW of Uktha Timan-Pechora Basin Russia

Bientactinosphaera miletenkoi

E ndash ndash Afanasieva 2000c 372 Upper Devonian middle Frasnian


Bientactinosphaera morozovi

E ndash ndash Afanasieva 2000a 53 Upper Devonian middle Frasnian


Bientactinosphaera pinica



Bientactinosphaera spinofoliacea

E ndash ndash Nazarov amp Afanasieva in Afanasieva 2000c 54

Upper Devonian Famennian Zadonsk Formation W Lekkeyaginsk-65 Saremboi-Lekkeyaga Timan-Pechora Basin Russia

Bientactinosphaera zuraevi E ndash ndash Afanasieva amp Amon 2011 1501 Upper Devonian Frasnian Polar Urals Palnik Yu River Russia

Bipylospongia (238)Bipylospongia rudosa S ndash T Noble 1994 37 Upper Silurian Gorstian Caballos Novaculite Formation

East Bourland Mts Marathon Basin W Texas USA

Bisphaera (82)Bisphaera beniguna E ndash ndash Won 1997a 344 Upper Devonian Frasnian Gogo Formation Pillara Range


Bisphaera cribrisimilis E ndash ndash Won 1997a 345 Upper Devonian Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Bisphaera cribrisimilis crassa

E ndash ndash Won 1997a 345 Upper Devonian Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Bisphaera cribrisimilis cribrisimilis


Bisphaera dissimilicortex E ndash ndash Won 1997a 346 Upper Devonian Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Bisphaera solidispinosa E ndash Th Ts

Won 1997a 346 Upper Devonian Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Bisphaera uniprocera E ndash ndash Won 1997a 346 Upper Devonian Frasnian Gogo Formation Pillara Range Canning Basin Western Australia





er

Stat

usTy




BissulentactiniaBissulentactinia

diaphoraxisndash nn ndash Li 1995 332 Middle



Bissylentactinia (133)Bissylentactinia bifida E ndash ndash Nazarov in Nazarov amp Popov

1980 64Middle



BissylentactinIa penita E ndash ndash Nazarov 1975 95 Upper Devonian Frasnian Egindinsk Formation Aitpaika River N Mugodazhar S Urals Kazakhstan

Bissylentactinia pilifera E ndash ndash Nazarov in Nazarov amp Popov 1980 64

Middle Ordovician


Bissylentactinia rudicula E ndash Ts Nazarov 1975 94 Upper Devonian Frasnian Egindinsk Formation Aitpaika River N Mugodazhar S Urals Kazakhstan

BistarkumBistarkum martiali S ndash ndash Feng in Feng et al 2006a 37 Upper Permian Changhsingian Dalong Formation Dongpan


BlastulospongiaBlastulospongia

mindyallicanr Calc-

areandash Bengtson 1986 205 Upper Cambrian Paibian Mungerebar Limestone Georgina

Basin Queensland AustraliaBlastulospongia

monothalamosnr Calc-

areaTnr Pickett amp Jell 1983 87 Middle Cambrian ndash Mootwingee New South Wales

Blastulospongia polytreta nr Calc-area

ndash Conway Morris amp Chen 1990 26

Lower Cambrian Stage 2-Stage 3

Quarry in Taishanmiao section N of Yichang City Hubei Province China

Borisella (83)Borisella bykovae E nn ndash Afanasieva 1997a 221 Upper Devonian middle

FrasnianDomanik Formation outcrop 1904

Lyaiol River Timan-Pechora Basin Russia

Borisella bykovae E ndash ndash Afanasieva 2000b 144 Upper Devonian middle Frasnian

Domanik Formation outcrop 1904 Lyaiol River Timan-Pechora Basin Russia

Borisella dunhilli E ndash ndash Noble amp Webby 2009 555 Upper Ordovician Katian Malongulli Formation Cliefden Caves area NSW Australia

Borisella invisitata E ndash ndash Afanasieva 2000a 45 Upper Devonian middle Frasnian


Borisella maksimovae E nn ndash Afanasieva 1997a 221 Upper Devonian middle Frasnian


Borisella maksimovae E ndash T Afanasieva 2000b 144 Upper Devonian middle Frasnian


Borisella mariae E nn ndash Afanasieva 1997a 221 Upper Devonian middle Frasnian


Borisella mariae E ndash ndash Afanasieva 2000b 145 Upper Devonian middle Frasnian


Borisella primitiva E ndash ndash Afanasieva 2000a 47 Upper Devonian middle Frasnian


Brianellium (182)Brianellium buckense L ndash ndash Cheng 1986 181 Upper

CarboniferousBashkirian Wesley Formation Turnpike section

Pittsburg County Oklahoma USA

Brianellium gordoni L ndash ndash Cheng 1986 181 Upper Carboniferous


Brianellium holdsworthi L ndash Ts Cheng 1986 182 Upper Carboniferous


Brianellium ishigai L ndash ndash Cheng 1986 182 Upper Carboniferous




Aitchison J C et al


er

Stat

usTy




Brianellium (182) continuationBrianellium laxum L ndash ndash Cheng 1986 183 Upper

CarboniferousBashkirian Wesley Formation Turnpike

section Pittsburg County Oklahoma USA

Brianellium medium L ndash ndash Cheng 1986 183 Upper Carboniferous


Brianellium obesum L ndash ndash Cheng 1986 184 Upper Carboniferous


Brianellium ruestae L ndash ndash Cheng 1986 184 Upper Carboniferous


Callela (84)Callela conispinosa E ndash ndash Won 1983 134 Lower


Frankenwald GermanyCallela hexactina E ndash ndash Won 1983 134 Lower


Frankenwald GermanyCallela parvispinosa E ndash ndash Won 1983 134 Lower


Frankenwald GermanyCallela stellaesimilis E ndash T Won 1983 135 Lower


Frankenwald GermanyCallela tetravalidaculeata E ndash ndash Won 1990 132 Lower


Barmen GermanyCallela variaculeata E ndash ndash Park amp Won 2012 66 Lower

CarboniferousTournaisian Woodman Formation South

Lakeside Mts Tooele County Utah USA

Campanulithus (9)Campanulithus falcatus Al ndash T Nazarov amp Rudenko 1981 137 Lower Permian Artinskian Aktastyi River Orenburgskaya

region Southern Urals RussiaCampanulithus insuetus Al ndash ndash Nestell amp Nestell 2010 14 Middle Permian Capitanian Bell Canyon Formation Apache


Camptoalatus (277)Camptoalatus benignus IS ndash ndash Nazarov amp Ormiston 1985 49 Upper



Camptoalatus monopterygius

IS ndash T Nazarov amp Rudenko 1981 138 Lower Permian Sakmarian Kandurov Formation Donskoye Ural River Orenburgskaya Region S Urals Russia

Camptoalatus volaticus IS ndash ndash Maldonado amp Noble 2010 80 Middle Permian Capitanian Bell Canyon Formation Apache Mountains Culberson County W Texas USA

Camptoalatus aretinus IS ndash ndash Nazarov in Isakova amp Nazarov 1986 121

Upper Carboniferous


Cancellientactinia (291)Cancellientactinia acifera IS

(OampF)ndash T Obut amp Shcherbanenko 2008

374Upper Devonian Frasnian Gryaznuka Formation Gryaznukha

Brook Rudny Altai southern West Siberia Russia

Cancellisphaera (51)Cancellisphaera kozuri Ar ndash ndash Won amp Iams 2015a 20 Lower Ordovician Floian Cow Head Group Western

Newfoundland Canada

Cangyuanella (10)Cangyuanella deflandrei Al nn ndash Feng 1992 55 Upper Permian Changhsingian Papai Formation of Changning-


Cangyuanella deflandrei Al ndash T Feng amp Liu 1993 550 Upper Permian Changhsingian Muyinhe Formation Nanpan Lancang County Yunnan China

Cariver (29)Cariver dorsoconvexus Al ndash ndash Kozur 1993 109 Upper Permian upper

ChanghsingianSosio Valley Area Torrente San

Calogero Western Sicily Italy





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CarposphaeraCarposphaera

aberdeenaendash nd ndash Riedel in Nitecki 1963 164 Upper Devonian Famennian Caballos Formation Marathon

Basin Texas USACarposphaera equalis ndash nd ndash Aberdeen 1940 133 Upper Devonian Famennian Caballos Formation Marathon

Basin Texas USACarposphaera infracrinita ndash nd ndash Ruumlst 1892 136 Lower


Carposphaera jejuna ndash nd ndash Ruumlst 1892 136 Lower Carboniferous


Carposphaera macractinia



Carposphaera magna ndash nd ndash Aberdeen 1940 133 Upper Devonian Famennian Caballos Formation Marathon Basin Texas USA

Carposphaera magna ndash nd ndash Ruumlst 1892 136 Lower Carboniferous


Carposphaera nana ndash nd ndash Hinde 1899b 215 Middle Devonian Givetian Chypons Farm Mullion Parrish Cornwall England UK

Carposphaera pygmaea ndash nd ndash Ruumlst 1892 135 Lower Carboniferous


Carposphaeridium (307)Carposphaeridium

cambrensendash nd Td Chapman 1923 37 Upper Cambrian ndash Lachlan Fold Belt Victoria

Australia

CaryosphaeraCaryosphaera groddeekii ndash nd ndash Ruumlst 1892 139 Upper Devonian ndash SchaebenholzHarz Mountains

Elbingerode Germany

Caspiaza (292)Caspiaza aculeata IS

(OampF)ndash ndash Afanasieva 1986 32 Lower

CarboniferousSerpukhovian Tyan-Shan Karachaganak area

Pre-Caspinsk Batys Qazaqstan Kazakhstan

Caspiaza calva IS(OampF)

ndash T Afanasieva 1986 31 Lower Carboniferous

Serpukhovian Tyan-Shan Karachaganak area Pre-Caspinsk Batys Qazaqstan Kazakhstan

Caspiaza collaricostulata IS(OampF)

ndash ndash Afanasieva 1993 117 Upper Devonian Famennian Polar Urals Yamalo-Nenetskiy Region Russia

Caspiaza spinifera IS(OampF)


Caspiaza urceus IS(OampF)

ndash ndash Afanasieva 1986 32 Lower Carboniferous


Cauletella (170)Cauletella constricta L ndash ndash Feng in Feng et al 2006b 833 Upper Permian Changhsingian Dalong Formation Dongpan


Cauletella paradoxa L ndash ndash Shang Caridroit amp Wang 2001 233

Upper Permian Changhsingian Changshing Formation Liuqiao section Fusui County southern Nanning Guangxi China

Cauletella porosa L ndash ndash Feng in Feng et al 2006b 833 Upper Permian Changhsingian Dalong Formation Dongpan section Dongpan Village Guangxi Zhuang Region China

Cavasphaera (285)Cavasphaera diversa IS ndash T Won amp Iams 2015a 30 Lower Ordovician Floian Cow Head Group Western

Newfoundland CanadaCavasphaera inaequalis IS ndash ndash Won amp Iams 2015a 30 Lower Ordovician Floian Cow Head Group Western

Newfoundland Canada

CecryphaliumCecryphalium

infundibulumndash nd ndash Ruumlst 1892 184 Upper Permian ndash Sosio Valley Area Western Sicily

ItalyCecryphalium

oligoporumndash nd ndash Ruumlst 1892 183 Upper Permian ndash Novgorod Russia

CenellipsisCenellipsis areolata ndash nd ndash Ruumlst 1892 151 Lower Devonian ndash Southern Urals RussiaCenellipsis cepaeformis ndash nd ndash Ruumlst 1892 152 Upper Devonian ndash SchaebenholzHarz Mountains

Elbingerode GermanyCenellipsis citrus ndash nd ndash Ruumlst 1892 151 Lower Devonian ndash Southern Urals RussiaCenellipsis curvatoporata ndash nd ndash Ruumlst 1892 151 Lower Devonian ndash Southern Urals Russia



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Cenellipsis (continuation)Cenellipsis cypridina ndash nd ndash Ruumlst 1892 152 Lower Devonian ndash Southern Urals RussiaCenellipsis diversipora ndash nd ndash Ruumlst 1892 151 Lower Silurian ndash Cabriegraveres Heacuterault FranceCenellipsis favosa ndash nd ndash Hinde 1899b 216 Middle Devonian Givetian Chypons Farm Mullion Parrish

Cornwall England UKCenellipsis multiplex ndash nd ndash Ruumlst 1892 152 Carboniferous ndash Buumlkk-Gebirge Inner Western

Carpathians HungaryCenellipsis perovalis ndash nd ndash Ruumlst 1892 151 Lower


Cenellipsis rectipora ndash nd ndash Ruumlst 1892 151 Lower Devonian ndash Southern Urals RussiaCenellipsis reticulosa ndash nd ndash Ruumlst 1892 152 Lower


Cenellipsis scitula ndash nd ndash Hinde 1899b 216 Middle Devonian Givetian Chypons Farm Mullion Parrish Cornwall England UK

Cenellipsis setosa ndash nd ndash Hinde 1899b 216 Middle Devonian Givetian Chypons Farm Mullion Parrish Cornwall England UK

CenodiscusCenodiscus intermedius ndash nd ndash Ruumlst 1892 162 Lower


Cenodiscus nummularis ndash nd ndash Ruumlst 1892 162 Lower Carboniferous


Cenodiscus primordialis ndash nd ndash Ruumlst 1892 161 Lower Silurian ndash Cabriegraveres Heacuterault France

CenosphaeraCenosphaera

aberdeenaeAr ndash ndash Furutani 1990 51 Upper Devonian Famennian Caballos Formation Marathon

Basin Texas USACenosphaera affinis ndash nd ndash Hinde 1899a 44 Middle Devonian Givetian Yarramie Formation Tamworth

Belt northern NSW AustraliaCenosphaera antiqua ndash nd ndash Ruedemann amp Wilson

1936 1567Lower Ordovician Floian Deepkill shale Mt Merino

Columbia County New York USA

Cenosphaera apiaria ndash nd ndash Ruumlst 1892 135 Lower Devonian ndash Southern Urals RussiaCenosphaera

cabrierensisndash nd ndash Deflandre 1973c 1150 Lower


Cenosphaera carbonica ndash nd ndash Ruumlst 1892 134 Lower Carboniferous


Cenosphaera castanea ndash nd ndash Ruumlst 1892 134 Upper Devonian ndash SchaebenholzHarz Mountains Elbingerode Germany

Cenosphaera hexagonalis


Cenosphaera hexata ndash nd ndash Pulfrey 1932 194 Lower Carboniferous

Serpukhovian Calver Sough Derbyshire England UK

Cenosphaera ingens ndash nd ndash Ruumlst 1892 134 Lower Silurian ndash Cabriegraveres Heacuterault FranceCenosphaera macropora ndash nd ndash Ruumlst 1892 134 Lower Silurian ndash Cabriegraveres Heacuterault FranceCenosphaera

marathonensisndash nd ndash Riedel in Nitecki 1963 165 Upper Devonian Famennian Marathon Basin Texas USA

Cenosphaera rossica ndash nd ndash Ruumlst 1892 134 Lower Devonian ndash Southern Urals RussiaCenosphaera scitula ndash nd ndash Hinde 1899a 43 Middle Devonian Givetian Yarramie Formation Tamworth

Belt northern NSW AustraliaCenosphaera

semiequalisndash nd ndash Aberdeen 1940 133 Upper Devonian Famennian Caballos Formation Marathon

Basin Texas USACenosphaera uralensis ndash nd ndash Ruumlst 1892 134 Lower Devonian ndash Southern Urals RussiaCenosphaera variabilis ndash nd ndash Aberdeen 1940 133 Upper Devonian Famennian Caballos Formation Marathon

Basin Texas USA

CentrocubusCentrocubus giganteus E ndash ndash Won 1983 135 Lower


Frankenwald Germany

CentroloncheCentrolonche obscura ndash nn ndash Deflandre 1953 404 Lower

CarboniferousTournaisian Cabriegraveres Heacuterault France

Cerarchocyrtium (220)Cerarchocyrtium

ambiguumN ndash T Deflandre 1973d 151 Lower


Cerarchocyrtium dirum N ndash ndash Cheng 1986 129 Upper Devonian Famennian Woodford Formation TI Valley section Pittsburg County Oklahoma USA





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Cerarchocyrtium (220) continuationCerarchocyrtium

singulariumN ndash ndash Cheng 1986 130 Upper Devonian Famennian Woodford Formation TI Valley


Cerarchocyrtium tinnulum



Ceratoclathrulum (221)Ceratoclathrulum

ambiguumndash nn Tn Deflandre 1960 216 Lower


Ceratoikiscum (11)Ceratoikiscum

acatangulatumAl ndash ndash Nazarov in Nazarov et al 1975

103Middle



Ceratoikiscum acatangulatum

Al ndash ndash Nazarov in Nazarov et al 1977 919

Middle Ordovician


Ceratoikiscum araneosum

Al ndash ndash Afanasieva 2000c 367 Upper Devonian middle Frasnian


Ceratoikiscum ardmorensis


Upper Devonian upper Famennian

Woodford Formation Criner Hills 2A section Carter County Oklahoma USA

Ceratoikiscum armiger Al ndash ndash Furutani 1990 51 Upper Silurian Ludfordian Hitoegane Formation Hida-gaien Belt E of Hitoegane Fukuji area Takayama City Japan

Ceratoikiscum astrum Al ndash ndash Cheng 1986 76 Upper Devonian Famennian Woodford Formation TI Valley section Pittsburg County Oklahoma USA

Ceratoikiscum asymmetricum



Ceratoikiscum avimexpectans

Al ndash T Deflandre 1953 409 Lower Carboniferous


Ceratoikiscum avimexpectans nonspongiosum



Ceratoikiscum berggreni Al ndash ndash Gourmelon 1987 96 Lower Carboniferous


Ceratoikiscum bicancellatum

Al ndash ndash Holdsworth 1969 228 Upper Carboniferous

Bashkirian Waterhouses Staffordshire England UK

Ceratoikiscum bujugum Al ndash ndash Foreman 1963 288 Upper Devonian Famennian Huron member Ohio Shale Ohio USA

Ceratoikiscum calvum Al ndash ndash Stratford amp Aitchison 1997 245 Lower Devonian Emsian Gamilaroi terrane Glenrock Station New England orogen NSW Australia

Ceratoikiscum canningense

Al ndash ndash Aitchison 1993b 108 Upper Devonian lower Famennian

Gogo Formation Pillara Range Canning Basin Western Australia

Ceratoikiscum carinatum Al ndash ndash Won 1998 234 Lower Carboniferous


Ceratoikiscum chengi Al ndash ndash Schwartzapfel amp Holdsworth 1996 113



Ceratoikiscum chengi Al ndash ndash Won 1998 235 Lower Carboniferous


Ceratoikiscum coroniferum

Al ndash ndash Luo Aitchison amp Wang 2002 115

Middle Devonian Givetian Tanhe Formation Wuxiangling section S of Nanning Guangxi Zhuang Region SW China

Ceratoikiscum costaciculare

Al ndash ndash Nazarov 1974a 44 Upper Devonian Frasnian Egindinsk Formation Aitpaika River N Mugodazhar S Urals Kazakhstan

Ceratoikiscum crinerensis Al ndash ndash Schwartzapfel amp Holdsworth 1996 106


Woodford Formation Locality 1 Carter County Oklahoma USA

Ceratoikiscum delawarensis


Lower Carboniferous

Visean Caney Shale Arbuckle Mountains Sand Branch Section Johnston County Oklahoma USA

Ceratoikiscum delicatum Al ndash ndash Cheng 1986 78 Upper Devonian Famennian Woodford Formation TI Valley section Pittsburg County Oklahoma USA



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Ceratoikiscum (11) continuationCeratoikiscum echinatum Al ndash ndash Aitchison 1993b 109 Upper Devonian lower

FamennianGogo Formation Pillara Range


Ceratoikiscum echinocostatum

Al ndash ndash Nazarov 1975 102 Upper Devonian Frasnian Egindinsk Formation Aitpaika River N Mugodazhar S Urals Kazakhstan

Ceratoikiscum elmensis Al ndash ndash Cheng 1986 80 Upper Carboniferous

Bashkirian Sand Branch Member of Caney Formation Pittsburg County Oklahoma USA

Ceratoikiscum extraordinarium

Al ndash ndash Cheng 1986 80 Lower Carboniferous


Ceratoikiscum famenium Al ndash ndash Nazarov amp Ormiston 1990 23 Upper Devonian Famenian Zapadno Valavskaya borehole 1-R Pripyat Depression Belarus

Ceratoikiscum famenium Al nn ndash Nazarov 1988 130 Upper Devonian Famenian Zapadno Valavskaya borehole 1-R Pripyat Depression Belarus

Ceratoikiscum famenium E nn ndash Nazarov 1988 132 Upper Devonian Famenian Zapadno Valavskaya borehole 1-R Pripyat Depression Belarus

Ceratoikiscum famennium

Al inv ndash Nazarov amp Ormiston 1993 50 Upper Devonian Famenian Zapadno Valavskaya borehole 1-R Pripyat Depression Belarus

Ceratoikiscum fenestratum

Al ndash ndash Won amp Seo 2010 245 Lower Carboniferous

Visean Bergisches Land N Westfalia Germany

Ceratoikiscum formosum Al ndash ndash Braun 1990 96 Lower Carboniferous

upper Tournaisian


Ceratoikiscum fragile Al ndash ndash Aitchison 1993b 109 Upper Devonian lower Famennian


Ceratoikiscum goodbodyi



Ceratoikiscum herkommeri




Ceratoikiscum hexaspina Al ndash ndash Won 1990 133 Lower Carboniferous


Ceratoikiscum hexastellatum

Al ndash ndash Cheng 1986 81 Upper Devonian Famennian Johns Valley Shale Ouachita Mountains eastern Oklahoma USA

Ceratoikiscum ichinotaniense

Al ndash ndash Ishiga 1992 395 Upper Silurian Pridoli Yoshiki Formation Hida-gaien Belt Ichinotani Valley Fukuji area Takayama City Gifu Japan

Ceratoikiscum incomptum


Ceratoikiscum izumiensis Al ndash ndash Kurihara amp Sashida 2000 64 Lower Devonian Emsian Shibasudani Formation Shibasudani Valley Ohno City Fukui Prefecture Japan

Ceratoikiscum jucundum Al ndash ndash Noble Tekin Gedik amp Pehlivan 2008 46

Lower Carboniferous


Ceratoikiscum kochiense Al ndash ndash Umeda 1997 419 Upper Silurian Pridoli Jyoru Formation Kurosegawa Belt Mt Konomori Kochi City Shikoku Japan

Ceratoikiscum konomoriense

Al ndash ndash Ishiga 1988 75 Upper Devonian middle Frasnian

Jyoru Formation Kurosegawa Belt Konomori section Kochi City Shikoku Japan

Ceratoikiscum kurosegum

Al ndash ndash Aitchison Hada Ireland amp Yoshikura 1996 59

Upper Silurian Pridoli Unit G4 Jingamori Kurosegawa terrane S Mt Konomori Kochi City Shikoku Japan

Ceratoikiscum labyrintheum

Al ndash ndash Cheng 1986 82 Upper Devonian Famennian Woodford Formation TI Valley section Pittsburg County Oklahoma USA

Ceratoikiscum lorum Al ndash ndash Holdsworth 1969 224 Upper Carboniferous


Ceratoikiscum lyratum Al ndash ndash Ishiga 1988 74 Upper Devonian middle Frasnian


Ceratoikiscum marginatum

Al ndash ndash Aitchison 1993b 109 Upper Devonian lower Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Ceratoikiscum mertsi Al ndash ndash Vishnevskaya 1998 65 Upper Devonian Frasnian Domanik Formation Uktha section Domanik creek Russia





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Ceratoikiscum (11) continuationCeratoikiscum mirum Al ndash ndash Cheng 1986 83 Lower

CarboniferousTournaisian Woodford Formation TI Valley


Ceratoikiscum neosurculum



Ceratoikiscum omicron Al ndash ndash Ormiston amp Lane 1976 172 Lower Carboniferous


Ceratoikiscum orbistellerae


Ceratoikiscum paragyium Al ndash ndash Stratford amp Aitchison 1997 244 Middle Devonian Eifelian Gamilaroi terrane Glenrock Station New England orogen NSW Australia

Ceratoikiscum patagiatum


Ceratoikiscum perittacanthinum

Al ndash ndash Foreman 1963 292 Upper Devonian Famennian Huron member Ohio Shale Ohio USA

Ceratoikiscum pillaraense Al ndash ndash Aitchison 1993b 110 Upper Devonian lower Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Ceratoikiscum planistellare


Ceratoikiscum quinquestellatum


Ceratoikiscum rectum Al ndash ndash Nazarov 1973a 698 Upper Devonian Frasnian Egindinsk Formation Aitpaika River N Mugodazhar S Urals Kazakhstan

Ceratoikiscum rectum Al ndash ndash Nazarov 1975 102 Upper Devonian Frasnian Egindinsk Formation Aitpaika River N Mugodazhar S Urals Kazakhstan

Ceratoikiscum regalinodus

Al ndash ndash Stratford amp Aitchison 1997 244 Middle Devonian Eifelian Gamilaroi terrane Glenrock Station New England orogen NSW Australia

Ceratoikiscum riedeli Al ndash ndash Cheng 1986 84 Upper Devonian Famennian Woodford Formation TI Valley section Pittsburg County Oklahoma USA

Ceratoikiscum robustum Al ndash ndash Aitchison 1993b 110 Upper Devonian lower Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Ceratoikiscum rotundum Al ndash ndash Braun 1990 97 Lower Carboniferous

upper Tournaisian


Ceratoikiscum sandbergi Al ndash ndash Cheng 1986 84 Upper Devonian Famennian Woodford Formation TI Valley section Pittsburg County Oklahoma USA

Ceratoikiscum simplum Al ndash ndash Cheng 1986 85 Upper Devonian Famennian Woodford Formation TI Valley section Pittsburg County Oklahoma USA

Ceratoikiscum spiculatum


Ceratoikiscum spinosiarcuatum


Ceratoikiscum spinosum Al ndash ndash Cheng 1986 85 Lower Carboniferous


Ceratoikiscum stellatum Al ndash ndash Aitchison 1993b 111 Upper Devonian lower Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Ceratoikiscum surculum Al ndash ndash Schwartzapfel amp Holdsworth 1996 99



Ceratoikiscum torale Al ndash ndash Aitchison 1993b 111 Upper Devonian lower Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Ceratoikiscum triactina Al ndash ndash Park amp Won 2012 47 Lower Carboniferous


Ceratoikiscum triangulatum





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Ceratoikiscum (11) continuationCeratoikiscum

tricancellatumAl ndash ndash Holdsworth 1969 225 Upper

CarboniferousBashkirian Upper Dove Valley SW Derbyshire

England UKCeratoikiscum turgidum Al ndash ndash Umeda 1998a 98 Middle Devonian Eifelian Nakahata Formation Section B

Ochi Town Shikoku JapanCeratoikiscum ukhtensis Al nn ndash Afanasieva 1997a 221 Upper Devonian middle



Ceratoikiscum ukhtensis Al ndash ndash Afanasieva 2000c 367 Upper Devonian middle Frasnian


Ceratoikiscum umbraculum



Ceratoikiscum umbraculum bisulcatum


upper Tournaisian


Ceratoikiscum vimenum Al ndash ndash Nazarov amp Ormiston 1983a 462 Upper Devonian Frasnian Sadler Range SE Cap Creek Gap Canning Basin Western Australia

Ceratoikiscum vimineum Al ndash ndash Wakamatsu Sugiyama amp Furutani 1990 179

Lower Devonian Emsian Nakahata Formation SW of Mt Yokokurayama Ochi Town Shikoku Japan

Ceratoikiscum wonae Al ndash ndash Cheng 1986 86 Upper Carboniferous


Ceratoikiscum xinjangense

Al ndash ndash Liu 1992 130 Upper Carboniferous

Bashkirian Houxia Formation Ewirgol-Houxia districts of Ulumuqi Xinjiang China

Ceratoikiscum acantangulatum


Middle Ordovician


Ceratoikiscum apertum Al nn ndash Deflandre 1960 216 Lower Carboniferous


Ceratoikiscum evolutum Al nn ndash Deflandre 1960 216 Lower Carboniferous


Ceratoikiscum reductum

Al nn ndash Deflandre 1960 216 Lower Carboniferous


Ceratoikiscum speciosum



Cessipylorum (256)Cessipylorum

admirandumE nn ndash Nazarov 1988 122 Upper Silurian Pridoli Maksyutov Complex Tarangul

River 10 km N of Kosistek Village S Urals Russia

Cessipylorum robuspinatum

E nn ndash Li 1995 332 Middle Ordovician

Dapingian Qingshuigou-Baijingsi Complex Baijingsi Quilian County Qinghai China

ChoenicosphaeraChoenicosphaera

brevispinandash nd ndash Ruedemann amp Wilson 1936

1568Lower Ordovician Floian Deepkill shale Mt Merino


Choenicosphaera multispoinosa



Circulaforma (12)Circulaforma admissarius Al ndash ndash Stratford amp Aitchison 1997 245 Lower Devonian Emsian Gamilaroi terrane Glenrock

Station New England orogen NSW Australia

Circulaforma annula Al ndash ndash Won amp Seo 2010 246 Lower Carboniferous


Circulaforma davidi Al ndash ndash Aitchison et al 1999 158 Lower Devonian Emsian Gamilaroi terrane Glenrock Station New England orogen NSW Australia

Circulaforma delicata Al ndash T Cheng 1986 87 Lower Carboniferous


Circulaforma oilensis Al ndash ndash Schwartzapfel amp Holdsworth 1996 122

Lower Carboniferous


Circulaforma robusta Al ndash ndash Cheng 1986 89 Upper Carboniferous






er

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ClavataClavata lancangensis ndash nd ndash Feng 1992 57 Upper Permian Changhsingian Muyinhe Formation Nanpan

Lancang County Yunnan ChinaClavata lancangensis ndash nn ndash Feng amp Liu 1993 545 Upper Permian Changhsingian Muyinhe Formation Nanpan

Lancang County Yunnan China

Copiconulus (293)Copiconulus acanthicus IS

(OampF)ndash ndash Feng in Feng et al 2006a 31 Upper Permian Changhsingian Dalong Formation Dongpan


Copiconulus solidus IS(OampF)

ndash T Feng in Feng et al 2006a 31 Upper Permian Changhsingian Dalong Formation Dongpan section Dongpan Village Guangxi Zhuang Region China

Copicyntra (239)Copicyntra acilaxa S ndash T Nazarov in Isakova amp Nazarov

1986 70Upper


Urals RussiaCopicyntra acilaxa

trigonaS ndash ndash Nazarov in Isakova amp Nazarov

1986 71Upper


Urals RussiaCopicyntra acilongata S ndash ndash Nazarov in Isakova amp Nazarov

1986 71Upper


Urals RussiaCopicyntra akikawaensis S ndash ndash Sashida amp Tonishi 1988 530 Upper Permian Changhsingian Ohirayama Unit South Chichibu

Belt Kashiwara Akigawa River Akiruno City Tokyo Japan

Copicyntra brevidentata S ndash ndash Kozur amp Mostler 1989 217 Lower Permian Kungurian Koshelev Foramtion Alogazovo Village Movsovic Region Russia

Copicyntra cuspidata S ndash ndash Nazarov amp Ormiston 1985 25 Lower Permian Artinskian Aktastyi River Orenburgskaya region Southern Urals Russia

Copicyntra erinacea S ndash ndash Nestell amp Nestell 2010 24 Middle Permian Capitanian Bell Canyon Formation Apache Mountains Culberson County W Texas USA

Copicyntra fragilispinosa S ndash ndash Kozur amp Mostler 1989 217 Lower Permian Sakmarian Sarabil Formation Sakmarskiy Orenburgskaya Region S Urals Russia

Copicyntra irregulata S ndash ndash Maldonado amp Noble 2010 100 Middle Permian Capitanian Bell Canyon Formation Apache Mountains Culberson County W Texas USA

Copicyntra leviuscula S ndash ndash Amon amp Braun 1994 5 Lower Permian Artinskian Burtsevesky horizon Dalny Tulkas Rill Bashkortostan Region S Urals Russia

Copicyntra multispinosa S ndash ndash Kozur amp Mostler 1989 217 Lower Permian Sakmarian Sarabil Formation Sakmarskiy Orenburgskaya Region S Urals Russia

Copicyntra phymatodonta

S ndash ndash Nazarov amp Ormiston 1985 25 Lower Permian Artinskian Aktastyi River Orenburgskaya region Southern Urals Russia

Copicyntra robustodentata

S ndash ndash Kozur amp Mostler 1989 217 Lower Permian Sakmarian Sarabil Formation Sakmarskiy Orenburgskaya Region S Urals Russia

Copicyntra ruzhencevi S ndash ndash Kozur amp Mostler 1989 218 Lower Permian Sakmarian Sarabil Formation Sakmarskiy Orenburgskaya Region S Urals Russia

Copicyntra ruzhencevi gracilispinosa


Copicyntra ruzhencevi ruzhencevi


Copicyntra shaiwaensis S ndash Ts Wang in Wang amp Shang 2001 118

Upper Permian Wuchiapingian Shaiwa Group Shaiwa Ziyun County Guizhou China

Copicyntra spinosa S ndash ndash Maldonado amp Noble 2010 100 Middle Permian Capitanian Bell Canyon Formation Apache Mountains Culberson County W Texas USA

Copicyntra tongi S ndash ndash Gui amp Feng in Gui et al 2009 804

Upper Permian Changhsingian Dalong Formation Pingdingshan section N Chaou Anhui Region China

Copicyntra ziyunensis S ndash T Feng amp Gu 2002 803 Upper Permian Changhsingian Qiaoma Formation Shaiwa section Ziyun County Guizou China

Copicyntra nuda S ndash ndash Wakamatsu Sugiyama amp Furutani 1990 169

Upper Silurian Pridoli Jyoru Formation Kurosegawa Belt S Mt Yokokurayama Ochi Town Shikoku Japan

Copicyntra simulens S ndash ndash Nazarov amp Ormiston 1985 25 Middle Permian Capitanian Bell Canyon Formation Guadalupe Mts Culbertson County W Texas USA



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Copicyntroides (240)Copicyntroides

asteriformisS ndash T Nazarov amp Ormiston 1985 27 Middle Permian Roadian Bell Canyon Formation Guadalupe

Mts Culbertson County W Texas USA

Copicyntroides nazarovi S ndash ndash Maldonado amp Noble 2010 108 Middle Permian Capitanian Bell Canyon Formation Apache Mountains Culberson County W Texas USA

Copicyntroides parvulus S ndash ndash Feng in Feng et al 2006a 39 Upper Permian Changhsingian Dalong Formation Dongpan section Dongpan Village Guangxi Zhuang Region China

Copicyntroides stellatus S ndash ndash He amp Feng in He et al 2011 486


Copiellintra (241)Copiellintra bispina S ndash ndash Nazarov amp Afanasieva in

Afanasieva 2000c 91Lower Permian Sakmarian station 13 Karachaganak area

Batys Qazaqstan KazakhstanCopiellintra diploacantha S ndash T Nazarov amp Ormiston 1985 26 Lower Permian Sakmarian Sarabil Formation Sakmarskiy

Orenburgskaya Region S Urals Russia

Copiellintra elongata S ndash ndash Feng in Feng et al 2006a 37 Upper Permian Changhsingian Dalong Formation Dongpan section Dongpan Village Guangxi Zhuang Region China

Copiellintra fastuosa S ndash ndash Nestell amp Nestell 2010 27 Middle Permian Capitanian Bell Canyon Formation Apache Mountains Culberson County W Texas USA

Copiellintra ferula S ndash ndash Noble amp Jin 2010 134 Middle Permian Capitanian Bell Canyon Formation Guadalupe Mts Culberson County W Texas USA

Copiellintra laurelae S ndash ndash Noble amp Jin 2010 134 Middle Permian Capitanian Bell Canyon Formation Guadalupe Mts Culberson County W Texas USA

Copiellintra orbiculata S ndash ndash Nestell amp Nestell 2010 26 Middle Permian Capitanian Bell Canyon Formation Apache Mountains Culberson County W Texas USA

Cornellus (189)Cornellus sakmaraensis L ndash Ts Kozur amp Mostler 1989 212 Lower Permian Sakmarian Sarabil Formation Sakmarskiy


Cornum (278)Cornum mittereri IS ndash T Schwartzapfel amp Holdsworth

1996 246Lower

CarboniferousVisean Caney Shale Arbuckle Mountains

Phillips Creek Section Carter County Oklahoma USA

Cornum mullerae IS ndash ndash Schwartzapfel amp Holdsworth 1996 238

Lower Carboniferous


Cornum olssonae IS ndash ndash Schwartzapfel amp Holdsworth 1996 242

Lower Carboniferous


Cornum repetskii IS ndash ndash Schwartzapfel amp Holdsworth 1996 232

Lower Carboniferous


CornutannaCornutanna dilatata ndash nd ndash Ruumlst 1892 180 Upper Permian ndash Sosio Valley Area Western Sicily

ItalyCornutanna macropora ndash nd ndash Ruumlst 1892 180 Upper Permian ndash Sosio Valley Area Western Sicily

Italy

Corythoecia (279)Corythoecia dichoptera IS ndash T Foreman 1963 298 Upper Devonian Famennian Huron member Ohio Shale Ohio

USACorythoecia

loxosegmentataIS ndash ndash Nazarov in Isakova amp Nazarov

1986 119Upper


Urals Russia

Costaentactinia (86)Costaentactinia delicata E ndash ndash Won amp Seo 2010 247 Lower

CarboniferousVisean Bergisches Land N Westfalia

GermanyCostaentactinia

foremanaeE ndash T Seo amp Won 2009 67 Upper Devonian Famennian Huron member Ohio Shale Ohio

USA





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Stat

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Cowheadia (281)Cowheadia duplextesta IS ndash T Won Iams amp Reed 2007 535 Lower Ordovician lower-middle


Newfoundland Canada

CromyodruppaCromyodruppa prunulina ndash nd ndash Ruumlst 1892 157 Upper Permian ndash Sosio Valley Area Western Sicily

Italy

CromyommaCromyomma

grandaevumndash nd ndash Ruumlst 1892 149 Lower Silurian ndash Cabriegraveres Heacuterault France

CromyosphaeraCromyosphaera alternans ndash nd ndash Ruumlst 1892 139 Upper Permian ndash Sosio Valley Area Western Sicily

ItalyCromyosphaera distans ndash nd ndash Ruumlst 1892 139 Lower Devonian ndash Southern Urals RussiaCromyosphaera eminens ndash nd ndash Ruumlst 1892 139 Lower


Cromyosphaera frequens ndash nd ndash Ruumlst 1892 138 Lower Carboniferous


Cromyosphaera petschorae

ndash nd ndash Ruumlst 1892 138 Lower Devonian ndash Southern Urals Russia

Cromyosphaera prisca ndash nd ndash Ruumlst 1892 138 Lower Silurian ndash Cabriegraveres Heacuterault FranceCromyosphaera radiata ndash nd ndash Ruumlst 1892 138 Lower


Cromyosphaera rotiforme ndash nd ndash Pulfrey 1932 194 Lower Carboniferous


CromyostylusCromyostylus variabilis E ndash Ts Ormiston amp Lane 1976 170 Lower

CarboniferousTournaisian Sycamore Limestone southern

Arbuckle Mountains Oklahoma USA

CrucidiscusCrucidiscus praecursor ndash nd ndash Ruumlst 1892 163 Lower Silurian ndash Cabriegraveres Heacuterault France

CubaxoniumCubaxonium antiquorum S ndash ndash Sheng amp Wang 1982 60 Middle Devonian Givetian Qizigiao Formation Matangwuo

Xintian district Hunan ChinaCubaxonium

octaedrospongiosumS ndash ndash Won 1983 139 Lower


Frankenwald Germany

CubosphaeraCubosphaera significans ndash nd ndash Deflandre amp Deflandre-Rigaud

1958 969Lower


Curvechidnina (52)Curvechidnina fusa Ar ndash ndash Won amp Iams 2002 23 Upper Cambrian Stage 10 Cow Head Group Western

Newfoundland CanadaCurvechidnina kozuri Ar ndash ndash Won amp Iams 2002 23 Upper Cambrian Stage 10 Cow Head Group Western

Newfoundland CanadaCurvechidnina

longispinosaAr ndash ndash Won amp Iams 2002 24 Upper Cambrian Stage 10 Cow Head Group Western


multiramosaAr ndash ndash Won Iams amp Reed 2005 446 Lower Ordovician lower-middle


Newfoundland CanadaCurvechidnina tetractina Ar ndash T Won amp Iams 2002 24 Upper Cambrian Stage 10 Cow Head Group Western

Newfoundland Canada

Cyclocarpus (87)Cyclocarpus tubiformis E ndash Tn Li amp Wang 1991 398 Upper Devonian Frasnian Liukiang Formation Etang section

Hexian County Guangxi China

CyphantaCyphanta piscis ndash nd ndash Ruumlst 1892 160 Lower Devonian ndash Southern Urals RussiaCyphanta quiniseriata ndash nd ndash Ruumlst 1892 161 Lower Devonian ndash Southern Urals Russia

Cyrtentactinia (229)Cyrtentactinia

cibdelosphaera N ndash T Foreman 1963 285 Upper Devonian Famennian Huron member Ohio Shale Ohio

USACyrtentactinia foremanae N ndash ndash Cheng 1986 160 Upper Devonian Famennian Woodford Formation TI Valley




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Cyrtentactinia (229) continuationCyrtentactinia formosa N ndash ndash Cheng 1986 161 Upper Devonian Famennian Woodford Formation TI Valley


Cyrtentactinia macrocephala

N ndash ndash Cheng 1986 161 Upper Devonian Famennian Woodford Formation TI Valley section Pittsburg County Oklahoma USA

Cyrtentactinia pessagnoi N ndash ndash Schwartzapfel amp Holdsworth 1996 141



Cyrtentactinia petrushevskayae


Cyrtentactinia primotica N ndash T Foreman 1963 285 Upper Devonian Famennian Huron member Ohio Shale Ohio USACyrtentactinia splendida N ndash ndash Cheng 1986 162 Upper Devonian Famennian Woodford Formation TI Valley


Cyrtisphaeractenium (222)Cyrtisphaeractenium

crassumN ndash ndash Cheng 1986 130 Upper Devonian Famennian Woodford Formation TI Valley


Cyrtisphaeractenium delicatum

N ndash ndash Cheng 1986 131 Lower Carboniferous


Cyrtisphaeractenium mendax

N nn ndash Deflandre 1960 216 Lower Carboniferous



N ndash ndash Deflandre 1972a 3538 Lower Carboniferous



N ndash T Deflandre 1972b 15 Lower Carboniferous


Cyrtisphaeractenium rurae



Cyrtisphaeractenium sandbergi

N ndash ndash Schwartzapfel amp Holdsworth 1996 170



Cyrtisphaeractenium shengi


Cyrtisphaeractenium spinosum



Cyrtisphaeractenium fluegeli

N ndash ndash Kiessling amp Tragelehn 1994 232 Upper Devonian Famennian Frankenwald north Bavaria Germany

Cyrtisphaeronemium (223)Cyrtisphaeronemium

prudentigerumN nn ndash Deflandre 1960 216 Lower


Cyrtisphaeronemium prudentigerum






CyrtocalpisCyrtocalpis obtusa ndash nd ndash Ruumlst 1892 180 Lower Devonian ndash Southern Urals RussiaCyrtocalpis prima ndash nd ndash Ruumlst 1892 180 Lower


Cyrtocalpis serieporata ndash nd ndash Ruumlst 1892 180 Lower Devonian ndash Southern Urals Russia

Deflandrella (171)Deflandrella manica L ndash T

Ts Th

De Wever amp Caridroit 1984 99 Middle Permian Capitanian Oi Formation Ultra-Tamba Belt Yanogawa River Yamasakicho-Kusune Shisou City Japan

Deflandrellium (224)Deflandrellium georgesi N ndash T Cheng 1986 134 Upper Devonian Famennian Woodford Formation TI Valley


Deflandrellium minutum N ndash ndash Cheng 1986 135 Upper Devonian Famennian Woodford Formation TI Valley section Pittsburg County Oklahoma USA

Deflandrellium pronum N ndash ndash Cheng 1986 135 Lower Carboniferous






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Deflantrica (142)Deflantrica furutanii E ndash ndash Kurihara amp Sashida 2000 60 Lower Devonian Emsian Shibasudani Formation

Shibasudani Valley Ohno City Fukui Prefecture Japan

Deflantrica solidum E ndash ndash Wakamatsu Sugiyama amp Furutani 1990 168

Lower Devonian Pragian Jyoru Formation Kurosegawa Belt Konomori area Kochi City Shikoku Japan

Devoniglansus (242)Devoniglansus brachiatus S ndash ndash Umeda 1998b 205 Upper Silurian Pridoli Mt Yokokurayama area Kochi

Prefecture JapanDevoniglansus unicus S ndash T Wakamatsu Sugiyama amp

Furutani 1990 172Upper Silurian Pridoli Jyoru Formation Kurosegawa

Belt S Mt Yokokurayama Ochi Town Shikoku Japan

DictyocephalusDictyocephalus

ventricosusndash nd ndash Ruumlst 1892 182 Upper Permian ndash Sosio Valley Area Western Sicily

Italy

DictyomitraDictyomitra magnifica ndash nd ndash Ruumlst 1892 188 Upper Permian ndash Sosio Valley Area Western Sicily

ItalyDictyomitra micropora ndash nd ndash Ruumlst 1892 188 Upper Permian ndash Sosio Valley Area Western Sicily

ItalyDictyomitra pumilio ndash nd ndash Ruumlst 1892 187 Upper Devonian ndash SchaebenholzHarz Mountains

Elbingerode Germany

DictyophimusDictyophimus dubius ndash nd ndash Ruumlst 1892 181 Upper Permian ndash Sosio Valley Area Western Sicily

Italy

Diparvapila (69)Diparvapila hicocki Ar ndash T MacDonald 1998 595 Lower Silurian Telychian Cape Phillips Formation

Cornwallis Island Canadian Arctic Archipelago

Diparvapila larseni Ar ndash ndash MacDonald 1998 595 Lower Silurian Telychian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago

Diparvapila pygmaea Ar ndash ndash Won Blodgett amp Nestor 2002 957

Lower Silurian Telychian Road River Formation subsidiary channel of Tatonduck River east-central Alaska USA

Diparvapila saintrochae Ar ndash ndash MacDonald 1998 595 Lower Silurian Telychian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago

Diploplegma (308)Diploplegma cinctum ndash nd Td Hinde 1890 48 Middle


Scotland UKDiploplegma

schonensisndash nd ndash Hinz-Schallreuter amp Schallreuter

2003 82Upper Ordovician Katian Gisloumlvshammar Scania Sweden

DistospiculaDistospicula delicata nr is Tnr Wang 1989 146 Lower Cambrian ndash Niutitang Formation Sandu


Distospicula vigorum nr is ndash Wang 1989 146 Lower Cambrian ndash Damiao Formation Erlongping Group Eastern Qinling ophiolite belt Henan Province China

DistriactisDistriactis vetusta E nd ndash Hinde 1899a 53 Middle Devonian Givetian Yarramie Formation Tamworth

Belt northern NSW Australia

DizoniumDizonium laeve ndash nd ndash Ruumlst 1892 174 Upper Permian ndash Sosio Valley Area Western Sicily

Italy

Dorydictyum (309)Dorydictyum magnum ndash nd ndash Ruedemann amp Wilson 1936



Dorydictyum minutum ndash nd ndash Ruedemann amp Wilson 1936 1574


Dorydictyum simplex ndash nd Td Hinde 1890 54 Middle Ordovician

Darriwilian Southern Uplands Peeblesshire Scotland UK



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DorylonchidiumDorylonchidium

antiquumndash nn ndash Deflandre 1953 391 Lower


Doryplegma (310)Doryplegma armatum ndash nd ndash Ruedemann amp Wilson 1936

1575Lower Ordovician Floian Normanskill chert Fly Summit

Washington County New York USA

Doryplegma gracile ndash nd ndash Hinde 1890 54 Middle Ordovician


Doryplegma nasutum ndash nd Td Hinde 1890 53 Middle Ordovician


Doryplegma nux ndash nd ndash Ruedemann amp Wilson 1936 1575


Doryplegma priscum ndash nd ndash Ruedemann amp Wilson 1936 1574

Lower Ordovician Floian Normanskill chert Glenmont Albany County New York USA

Dorysphaera (311)Dorysphaera

domanicensisndash nd ndash Bykova 1955 68 Upper Devonian Frasnian Domanik Formation Tatarstan

RussiaDorysphaera echinata ndash nd ndash Hinde 1899a 45 Middle Devonian Givetian Yarramie Formation Tamworth


Dorysphaera laxa ndash nd ndash Hinde 1890 53 Middle Ordovician


Dorysphaera nucula ndash nd ndash Hinde 1890 53 Middle Ordovician


Dorysphaera reticulata ndash nd Td Hinde 1890 52 Middle Ordovician


Druppalonche (88)Druppalonche clavigera E ndash Ts Hinde 1899b 217 Middle Devonian Givetian Chypons Farm Mullion Parrish

Cornwall England UKDruppalonche ovata E ndash ndash Hinde 1899b 218 Middle Devonian Givetian Chypons Farm Mullion Parrish

Cornwall England UK

DruppatractusDruppatractus

dictyococcusndash nd ndash Ruumlst 1892 158 Lower


DruppulaDruppula amygdalina ndash nd ndash Ruumlst 1892 155 Lower Devonian ndash Southern Urals RussiaDruppula andreana ndash nd ndash Ruumlst 1892 155 Lower


Druppula angustiporata ndash nd ndash Ruumlst 1892 156 Carboniferous ndash Buumlkk-Gebirge Inner Western Carpathians Hungary

Druppula berberis ndash nd ndash Ruumlst 1892 156 Upper Permian ndash Sosio Valley Area Western Sicily Italy

Druppula cembra ndash nd ndash Ruumlst 1892 155 Lower Devonian ndash Southern Urals RussiaDruppula cornus ndash nd ndash Ruumlst 1892 156 Carboniferous ndash Buumlkk-Gebirge Inner Western

Carpathians HungaryDruppula crucifera ndash nd ndash Ruumlst 1892 156 Lower


Druppula endechinata ndash nd ndash Ruumlst 1892 155 Lower Carboniferous


Druppula fixata ndash nd ndash Ruumlst 1892 156 Upper Permian ndash Sosio Valley Area Western Sicily Italy

Druppula silurica ndash nd ndash Ruumlst 1892 154 Lower Silurian ndash Cabriegraveres Heacuterault FranceDruppula simplex ndash nd ndash Ruedemann amp Wilson 1936

1576Lower Ordovician Floian Normanskill chert Flint Mine Hill


Duodecimentactinia (89)Duodecimentactinia

amplaE ndash ndash Won 1997b 377 Upper Devonian Frasnian Gogo Formation Pillara Range

Canning Basin Western AustraliaDuodecimentactinia

araneaE ndash T Won 1997b 378 Upper Devonian Frasnian Gogo Formation Pillara Range


mediaE ndash ndash Won 1997b 378 Upper Devonian Frasnian Gogo Formation Pillara Range






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Duosphaera (90)Duosphaera hebes E ndash ndash Park amp Won 2012 55 Lower



Duplexia (312)Duplexia spinocurva E nd ndash Afanasieva 2000a 62 Upper Devonian Famennian Zadonsk Formation W


Duplexia parviperforata E nd ndash Won 1990 134 Lower Carboniferous


Durahelenifore (13)Durahelenifore robustum Al ndash Ts Boundy-Sanders amp Murchey in

Boundy-Sanders et al 1999 64


Slaven Chert Northern Shoshone Range Roberts Mountains Allochthon Nevada USA

DystympaniumDystympanium

carbonicumndash nd ndash Ruumlst 1892 177 Lower


Echidnina (53)Echidnina bengtsoni Ar ndash ndash Kozur Mostler amp Repetski 1996

249Lower Ordovician Tremadocian Windfall Formation Antelope

Range Eureka County Nevada USA

Echidnina conexa Ar ndash ndash Won Iams amp Reed 2005 447 Lower Ordovician lower-middle Tremadocian

Cow Head Group Western Newfoundland Canada

Echidnina conjuncta Ar ndash ndash Won amp Iams 2002 25 Upper Cambrian Stage 10 Cow Head Group Western Newfoundland Canada

Echidnina curvata Ar ndash ndash Won amp Iams 2002 25 Upper Cambrian Stage 10 Cow Head Group Western Newfoundland Canada

Echidnina irregularis Ar ndash ndash Won amp Iams 2002 25 Upper Cambrian Stage 10 Cow Head Group Western Newfoundland Canada

Echidnina laxa Ar ndash ndash Won Iams amp Reed 2005 447 Lower Ordovician lower-middle Tremadocian


Echidnina runnegari Ar ndash T Bengtson 1986 202 Upper Cambrian Paibian Mungerebar Limestone Georgina Basin Queensland Australia

Echidnina semiconexa Ar ndash ndash Won Iams amp Reed 2005 447 Lower Ordovician lower-middle Tremadocian


Echidnina severedeformis Ar ndash ndash Won Iams amp Reed 2005 449 Lower Ordovician lower-middle Tremadocian


Echidnina immanis Ar ndash ndash Won Iams amp Reed 2005 449 Lower Ordovician lower-middle Tremadocian


Echidnina monactina Ar ndash ndash Won amp Iams 2002 26 Upper Cambrian Stage 10 Cow Head Group Western Newfoundland Canada

Echidnina stevensi Ar ndash ndash Won amp Iams 2002 26 Upper Cambrian Stage 10 Cow Head Group Western Newfoundland Canada

EchinommaEchinomma

oligacanthumndash nd ndash Ruumlst 1892 149 Upper Permian ndash Sosio Valley Area Western Sicily

Italy

Ectoactinia (313)Ectoactinia tulufanensis ndash nd Td Liu 1992 130 Upper

CarboniferousBashkirian Houxia Formation Ewirgol-Houxia

districts of Ulumuqi Xinjiang China

Ehrenbergia (314)Ehrenbergia baijingensis ndash nd Td Li 1995 335 Middle



EllipsidiumEllipsidium aculeatum ndash nd ndash Ruumlst 1892 152 Lower


Ellipsidium castanea ndash nd ndash Hinde 1899a 51 Middle Devonian Givetian Yarramie Formation Tamworth Belt northern NSW Australia

Ellipsidium spinosum ndash nd ndash Ruumlst 1892 153 Lower Carboniferous




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Ellipsocopicyntra (243)Ellipsocopicyntra acantha S ndash ndash Feng in Feng et al 2006a 33 Upper Permian Changhsingian Dalong Formation Dongpan


Ellipsocopicyntra oviformis

S ndash T Kozur amp Mostler 1989 218 Lower Permian Sakmarian Sarabil Formation Sakmarskiy Orenburgskaya Region S Urals Russia

Ellipsocopicyntra sakmaraensis


Ellipsocopicyntra wani S ndash ndash Feng in Feng et al 2006a 33 Upper Permian Changhsingian Dalong Formation Dongpan section Dongpan Village Guangxi Zhuang Region China

Ellipsostigma (163)Ellipsostigma australe E nd Td Hinde 1899a 51 Middle Devonian Givetian Yarramie Formation Tamworth


EllipsostylusEllipsostylus obliquus ndash nd ndash Ruumlst 1892 153 Upper Permian ndash Sosio Valley Area Western Sicily

Italy

EllipsoxiphusEllipsoxiphus hystrix ndash nd ndash Ruumlst 1892 153 Lower


Ellipsoxiphus procerus ndash nd ndash Ruumlst 1892 153 Lower Carboniferous


Entactinia (91)Entactinia akdymensis E ndash ndash Nazarov 1975 53 Middle



Entactinia americana E ndash ndash Park amp Won 2012 51 Lower Carboniferous


Entactinia aperticuva E ndash ndash Aitchison 1993b 112 Upper Devonian lower Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Entactinia atypica E ndash ndash Nazarov 1975 53 Upper Cambrian Furongian Burudaytalrsquosk Formation SW Pribalkhash Kazakhstan

Entactinia austrouralica E ndash ndash Nazarov in Isakova amp Nazarov 1986 64

Upper Carboniferous


Entactinia bella E ndash ndash Afanasieva amp Amon 2011 1498 Upper Devonian Frasnian Polar Urals Palnik Yu River RussiaEntactinia bifida E ndash ndash Afanasieva 2000c 369 Upper Devonian middle



Entactinia bogdanovi E ndash ndash Afanasieva 2000c 371 Upper Devonian middle Frasnian

Domanik Formation borehole Ukhtinskaya-3B SW of Uktha Timan-Pechora Basin Russia

Entactinia carbonica E ndash ndash Liu 1992 128 Upper Carboniferous


Entactinia cometes E ndash ndash Foreman 1963 271 Upper Devonian Famennian Huron member Ohio Shale Ohio USA

Entactinia complanata E ndash ndash Nazarov 1975 56 Middle Ordovician


Entactinia complanata E ndash ndash Nazarov in Nazarov et al 1977 917

Middle Ordovician


Entactinia consociata E ndash ndash Nazarov 1975 50 Upper Devonian Frasnian Egindinsk Formation Aitpaika River N Mugodazhar S Urals Kazakhstan

Entactinia crustescens E ndash ndash Foreman 1963 272 Upper Devonian Famennian Huron member Ohio Shale Ohio USA

Entactinia dasysa E ndash ndash Nazarov in Nazarov amp Popov 1976 408

Middle Ordovician


Entactinia densa E ndash ndash Won 1998 236 Lower Carboniferous


Entactinia densissima E ndash ndash Nazarov amp Ormiston 1985 18 Lower Permian Sakmarian Sarabil Formation Sakmarskiy Orenburgskaya Region S Urals Russia





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Entactinia (91) continuationEntactinia dianae E ndash ndash Nestell Pope amp Nestell 2012

236Upper

CarboniferousMoscovian Mouse Creek Formation Excello

Shale Member south-central Iowa USA

Entactinia dimidiata E ndash ndash Nazarov 1975 48 Upper Devonian Frasnian Egindinsk Formation Aitpaika River N Mugodazhar S Urals Kazakhstan

Entactinia dissora E ndash ndash Nazarov 1975 48 Upper Devonian Frasnian Egindinsk Formation Aitpaika River N Mugodazhar S Urals Kazakhstan

Entactinia diversita E ndash ndash Nazarov 1973a 697 Upper Devonian Frasnian Egindinsk Formation Aitpaika River N Mugodazhar S Urals Kazakhstan

Entactinia dolichoacus E ndash ndash Nazarov in Isakova amp Nazarov 1986 62

Upper Carboniferous


Entactinia dolichoacus dolichoacus

E ndash ndash Nazarov in Isakova amp Nazarov 1986 63

Upper Carboniferous


Entactinia dolichoacus praematura


Upper Carboniferous


Entactinia duksundiensis E ndash ndash Nazarov in Nazarov et al 1981 82


Entactinia eiklaensis E ndash ndash Nazarov in Nazarov amp Nolvak 1983 4

Upper Ordovician Hirnantian Saursquoyask Formation Bore hole Eikla Saaremaa Estonia

Entactinia elongata E ndash ndash Nazarov 1975 55 Middle Ordovician


Entactinia exilispina E ndash ndash Foreman 1963 273 Upper Devonian Famennian Huron member Ohio Shale Ohio USA

Entactinia faceta E ndash ndash Amon Braun amp Chuvashov 1990 121

Lower Permian Artinskian Shelyvaginsk Formation around Sim Town Chalyabinskaya region Urals Russia

Entactinia foveolata E ndash ndash Nazarov 1975 51 Upper Devonian Frasnian Egindinsk Formation Aitpaika River N Mugodazhar S Urals Kazakhstan

Entactinia gogoense E ndash ndash Aitchison 1993b 112 Upper Devonian lower Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Entactinia haeckeliana E ndash ndash Won 1997a 347 Upper Devonian Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Entactinia herculea E ndash T Foreman 1963 271 Upper Devonian Famennian Huron member Ohio Shale Ohio USA

Entactinia hindeiana E ndash ndash Won 1997a 348 Upper Devonian Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Entactinia hirsuta E ndash ndash Nazarov in Nazarov amp Popov 1980 33

Middle Ordovician

Darriwilian Kizyilzarsk Formation area between Moyynty and Zhamishinsk Rivers Kazakhstan

Entactinia hoxtolgayensis E ndash ndash Wang 1997 152 Upper Devonian Famennian Gennaren Formation S Utubulak NW Jungar Basin Xinjiang China

Entactinia hystricuosa E ndash Aitchison 1993b 113 Upper Devonian lower Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Entactinia iriensis E ndash ndash Nazarov in Nazarov amp Shkolnik 1974 108

Middle Cambrian ndash Dzhagdish between Nim and Dzharovdik River Khabarovsk Russia

Entactinia itsukaichiensis E ndash ndash Sashida amp Tonishi 1985 9 Upper Permian Changhsingian Ohirayama Unit South Chichibu Belt Kashiwara Akigawa River Akiruno City Tokyo Japan

Entactinia leptura E ndash ndash Goto Umeda amp Ishiga 1992 153

Upper Devonian upper Frasnian Malongulli Formation Lachlan Fold Belt eastern NSW Australia

Entactinia mediforma E ndash ndash Won 1997a 348 Upper Devonian Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Entactinia meishanensis E ndash ndash He Feng amp Gu 2005 210 Upper Permian Changhsingian Changxing Formation Meishan D section Changxing Zhejiang China

Entactinia micropora E ndash ndash Goto Umeda amp Ishiga 1992 153

Upper Ordovician Katian Malongulli Formation Lachlan Fold Belt 30 km NW of Taralga NSW Australia



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Entactinia (91) continuationEntactinia micula E ndash ndash Foreman 1963 272 Upper Devonian Famennian Huron member Ohio Shale Ohio

USAEntactinia minuta E ndash ndash Feng in Feng et al 2007 23 Upper Permian Changhsingian Dalong Formation Dongpan


Entactinia mixta E ndash ndash Park amp Won 2012 51 Lower Carboniferous


Entactinia modesta E ndash ndash Goto Umeda amp Ishiga 1992 151


Malongulli Formation Lachlan Fold Belt eastern NSW Australia

Entactinia modesta E ndash ndash Sashida amp Tonishi 1985 10 Upper Permian Changhsingian Ohirayama Unit South Chichibu Belt Kashiwara Akigawa River Akiruno City Tokyo Japan

Entactinia monalloea E ndash ndash Foreman 1963 273 Upper Devonian Famennian Huron member Ohio Shale Ohio USA

Entactinia octaparvispina E ndash ndash Won 1990 137 Lower Carboniferous


Entactinia octaradiata E ndash ndash Won 1997a 349 Upper Devonian Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Entactinia ormistoni E ndash ndash Won 1983 142 Lower Carboniferous


Entactinia oumonhaoensis

E ndash ndash Wang 1997 152 Upper Devonian Famennian Gennaren Formation S Utubulak NW Jungar Basin Xinjiang China

Entactinia pantosompha E ndash ndash Foreman 1963 273 Upper Devonian Famennian Huron member Ohio Shale Ohio USA

Entactinia parapycnoclada

E ndash ndash Nazarov amp Ormiston 1985 19 Middle Permian Roadian Bell Canyon Formation Guadalupe Mts Culbertson County W Texas USA

Entactinia parva E ndash ndash Won 1983 143 Lower Carboniferous


Entactinia parviporosa E ndash ndash Won 1990 136 Lower Carboniferous


Entactinia patorovaria E ndash ndash Afanasieva 2000a 43 Upper Devonian middle Frasnian


Entactinia paula E ndash ndash Foreman 1963 271 Upper Devonian Famennian Huron member Ohio Shale Ohio USA

Entactinia perplexa E ndash ndash Nazarov in Zhuravleva 1975 55 Upper Cambrian Furongian Dzhagdy River area Khabarovsk Russia

Entactinia pillaraense E ndash ndash Aitchison 1993b 113 Upper Devonian lower Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Entactinia pinrasensis E ndash ndash Afanasieva amp Amon 2016 216 Lower Permian Artinskian Northern Mugodzhary Aktasty River Kazakhstan

Entactinia postadditiva E ndash ndash Nazarov amp Ormiston 1993 32 Upper Carboniferous

Moscovian Nyatvinsk Formation Popovka River Prikolyma Region Magadan Region NE Russia

Entactinia praenuntia E ndash ndash Nazarov amp Ormiston 1993 30 Upper Devonian Famenian Zapadno Valavskaya borehole 1-R Pripyat Depression Belarus

Entactinia praenuntia ndash nn ndash Nazarov 1988 190 Upper Devonian Famenian Zapadno Valavskaya borehole 1-R Pripyat Depression Belarus

Entactinia praepycnoclada

E ndash ndash Nazarov amp Ormiston 1993 32 Upper Carboniferous



ndash nn ndash Nazarov 1988 191 Upper Carboniferous

Moscovian Popovka River in Prikolyma area Magadan Region Russia

Entactinia proceraspina E ndash ndash Aitchison 1993b 113 Upper Devonian lower Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Entactinia prodigialis E ndash ndash Nazarov 1975 52 Upper Devonian Frasnian Egindinsk Formation Aitpaika River N Mugodazhar S Urals Kazakhstan

Entactinia profundisulcus E ndash Ts Aitchison 1993b 114 Upper Devonian lower Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Entactinia pycnoclada E ndash ndash Nazarov amp Ormiston 1985 19 Lower Permian Sakmarian Sarabil Formation Kondurovska Orenburgskaya Region S Urals Russia





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Entactinia (91) continuationEntactinia quantilla E ndash ndash Foreman 1963 273 Upper Devonian Famennian Huron member Ohio Shale Ohio

USAEntactinia reticulata E ndash ndash Sashida amp Tonishi 1985 10 Upper Permian Changhsingian Ohirayama Unit South Chichibu


Entactinia rostriformis E ndash ndash Afanasieva amp Amon 2008 465 Lower Devonian Emsian Kamennaya Gora Tanalyk River Southern Ural Russia

Entactinia sashidai E ndash ndash Feng in Feng et al 2007 23 Upper Permian Changhsingian Dalong Formation Dongpan section Dongpan Village Guangxi Zhuang Region China

Entactinia sexradiata E ndash ndash Won 1997a 348 Upper Devonian Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Entactinia somphorhips E ndash ndash Foreman 1963 272 Upper Devonian Famennian Huron member Ohio Shale Ohio USA

Entactinia spinosa E ndash ndash Liu 1992 129 Upper Carboniferous


Entactinia spongites E ndash ndash Foreman 1963 272 Upper Devonian Famennian Huron member Ohio Shale Ohio USA

Entactinia suave E ndash ndash Nazarov in Nazarov amp Kruycheck 1977 1447

Upper Devonian Famennian Eletsk horizon Valavsk Belarus

Entactinia subulata E ndash ndash Webby amp Blom 1986 149 Upper Ordovician Katian Malongulli Formation Belubula River E of Canowindra central NSW Australia

Entactinia tenuiacerosa E ndash ndash Nazarov 1975 52 Upper Devonian Frasnian Egindinsk Formation Aitpaika River N Mugodazhar S Urals Kazakhstan

Entactinia tyrrelli E ndash ndash Nazarov amp Ormiston 1985 19 Middle Permian Capitanian Bell Canyon Formation Guadalupe Mts Culbertson County W Texas USA

Entactinia unica E ndash ndash Nazarov 1975 54 Middle Ordovician


Entactinia unica E ndash T Nazarov in Nazarov et al 1977 917

Middle Ordovician


Entactinia vulgaris E ndash ndash Won 1983 144 Lower Carboniferous


Entactinia vulgaris microporata

E ndash ndash Braun 1989 88 Lower Carboniferous

upper Tournaisian


Entactinia wangi E ndash ndash Feng in Feng et al 2007 21 Upper Permian Changhsingian Dalong Formation Dongpan section Dongpan Village Guangxi Zhuang Region China

Entactinia xinjiangensis E ndash ndash Wang 1997 153 Upper Devonian Famennian Gennaren Formation S Utubulak NW Jungar Basin Xinjiang China

Entactinia additiva E ndash ndash Foreman 1963 273 Upper Devonian Famennian Huron member Ohio Shale Ohio USA

Entactinia brilonensis E ndash ndash Won 1983 144 Lower Carboniferous


Entactinia delvolvei E ndash ndash Gourmelon 1987 45 Lower Carboniferous


Entactinia inaequoporosa

E ndash ndash Won 1983 145 Lower Carboniferous


Entactinia multispinosa E ndash ndash Won 1983 145 Lower Carboniferous


Entactinia octaculeata E ndash ndash Won 1983 146 Lower Carboniferous


Entactinia pantotolma E ndash ndash Braun 1989 88 Lower Carboniferous

upper Tournaisian


Entactinia sheeprockensis

E ndash ndash Park amp Won 2012 54 Lower Carboniferous


Entactinia spinifera E ndash ndash Amon Braun amp Chuvashov 1990 121




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Entactinia (91) continuationEntactinia spongia E ndash ndash Renz 1990 369 Upper Ordovician Katian Hanson Creek Formation N Martin

Ridge Eureka County Nevada USA

Entactinia strena E ndash ndash Feng amp Ye 1996 19 Middle Devonian Givetian Lailei Formation Lila Village Ximeng County SW Yunnan China

Entactinia sychnacanthina

E ndash ndash Foreman 1963 274 Upper Devonian Famennian Huron member Ohio Shale Ohio USA

Entactinia unispina E ndash ndash Won 1991b 98 Lower Carboniferous


Entactinia ximengensis E ndash ndash Feng amp Ye 1996 19 Middle Devonian Givetian Lailei Formation Lila Village Ximeng County SW Yunnan China

Entactinosphaera (92)Entactinosphaera

aculeataE ndash ndash Nazarov 1975 70 Upper Ordovician Hirnantian Otyzbes area Qostanay Central

KazakhstanEntactinosphaera

aculeatissimaE ndash ndash Aitchison 1993b 114 Upper Devonian lower Frasnian Gogo Formation Pillara Range


Entactinosphaera aenigma


Upper Carboniferous


Entactinosphaera aitpaiensis

E ndash ndash Nazarov 1973a 697 Upper Devonian Frasnian Egindinsk Formation Aitpaika River N Mugodazhar S Urals Kazakhstan

Entactinosphaera aksakensis



Entactinosphaera assidera

E ndash ndash Nazarov 1975 64 Upper Devonian Frasnian Egindinsk Formation Aitpaika River N Mugodazhar S Urals Kazakhstan

Entactinosphaera aubouini

E ndash ndash Caridroit amp De Wever 1986 75 Upper Permian Wuchiapingian Oi Formation Ultra-Tamba Belt Yanogawa River Yamasakicho-Kusune Shisou City Japan

Entactinosphaera australis

E ndash ndash Aitchison 1993b 115 Upper Devonian lower Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Entactinosphaera baragensis



Entactinosphaera bellula E ndash ndash Wang 1997 154 Upper Devonian Famennian Gennaren Formation S Utubulak NW Jungar Basin Xinjiang China

Entactinosphaera brevispinosa

E ndash ndash Sashida amp Tonishi 1988 529 Upper Permian Changhsingian Ohirayama Unit South Chichibu Belt Kashiwara Akigawa River Akiruno City Tokyo Japan

Entactinosphaera calthrata


Upper Carboniferous


Entactinosphaera cancellicula


Entactinosphaera carssispira

E ndash ndash Liu 1992 129 Upper Carboniferous


Entactinosphaera cimelia E ndash ndash Nazarov amp Ormiston 1985 22 Middle Permian Capitanian Bell Canyon Formation Guadalupe Mts Culbertson County W Texas USA

Entactinosphaera conglobata


Entactinosphaera crassiclathrata

E ndash ndash Nazarov amp Ormiston 1985 20 Lower Permian Artinskian Kandurov Formation Donskoye Ural River Orenburgskaya Region S Urals Russia

Entactinosphaera dystactotata


Entactinosphaera egindyensis


Entactinosphaera esostrongyla

E ndash T Foreman 1963 274 Upper Devonian Famennian Huron member Ohio Shale Ohio USA

Entactinosphaera euthlasta






er

Stat

usTy




Entactinosphaera (92) continuationEntactinosphaera

excurrensE ndash ndash Nazarov in Nazarov amp Popov

1980 40Upper Ordovician Hirnantian Ulrsquokuntassk Member along Oy-Su

River KazakhstanEntactinosphaera

excurrensndash inv ndash Nazarov 1980 23 Upper Ordovician Hirnantian Ulrsquokuntassk Member along Oy-Su


explicataE ndash ndash Nazarov 1975 68 Middle

OrdovicianDarriwilian Erzhansk Formation Lake

Sasykser NE Central Kazakhstan

Entactinosphaera foremanae

E ndash Td Ormiston amp Lane 1976 165 Lower Carboniferous


Entactinosphaera fredericki


Entactinosphaera grandis E ndash ndash Nazarov 1975 65 Upper Devonian Frasnian Egindinsk Formation Aitpaika River N Mugodazhar S Urals Kazakhstan

Entactinosphaera guangxiensis

E ndash ndash Li amp Wang 1991 400 Upper Devonian Frasnian Liukiang Formation Etang section Hexian County Guangxi China

Entactinosphaera hystricosa


Entactinosphaera inconstans

E ndash ndash Nazarov 1975 68 Middle Ordovician


Entactinosphaera inpercepta

E ndash ndash Nazarov 1975 70 Upper Ordovician Hirnantian Otyzbes area Qostanay Central Kazakhstan

Entactinosphaera inusitata


Entactinosphaera liquidambarfructa

E ndash ndash Ormiston amp Lane 1976 166 Lower Carboniferous


Entactinosphaera menneri

ndash nn ndash Afanasieva 1997a 221 Upper Devonian middle Frasnian


Entactinosphaera neocimelia

E ndash ndash Feng Fang Zhang amp Huang 1998 240

Upper Permian Changhsingian Papai Formation of Changning-Menglian Belt Papai Village Cangyuan County Yunnan China

Entactinosphaera palimbola


Entactinosphaera paracimela

E ndash ndash Feng amp Liu 1993 558 Upper Permian Changhsingian Laba Group SW Yunnan China

Entactinosphaera pseudocimelia


Entactinosphaera pulcherima

E nn ndash Nazarov amp Ormiston 1990 16 Upper Devonian Famenian Zapadno Valavskaya borehole 1-R Pripyat Depression Belarus


E ndash ndash Nazarov amp Ormiston 1993 32 Upper Devonian Famenian Zapadno Valavskaya borehole 1-R Pripyat Depression Belarus


ndash nn ndash Nazarov 1988 130 Upper Devonian Famenian Zapadno Valavskaya borehole 1-R Pripyat Depression Belarus

Entactinosphaera riedeli E ndash Ts Foreman 1963 275 Upper Devonian Famennian Huron member Ohio Shale Ohio USA

Entactinosphaera ruesti E ndash ndash Won 1991b 99 Lower Carboniferous


Entactinosphaera strangulata

E ndash T Nazarov amp Ormiston 1985 20 Lower Permian Artinskian Kandurov Formation Donskoye Ural River Orenburgskaya Region S Urals Russia

Entactinosphaera symphypora


Entactinosphaera tretactinia


Entactinosphaera unimana

E ndash ndash Nazarov in Nazarov amp Kruycheck 1977 1448


Entactinosphaera variacanthina

E ndash Ts Foreman 1963 278 Upper Devonian Famennian Huron member Ohio Shale Ohio USA

Entactinosphaera verrucula

E ndash ndash Nazarov in Nazarov amp Popov 1976 408

Middle Ordovician


Entactinosphaera wuhanensis

ndash nn ndash Feng 1992 59 Middle Permian Capitanian Gufeng Formation Hubei China



Aitchison J C et al


er

Stat

usTy




Entactinosphaera (92) continuationEntactinosphaera almae E ndash ndash Won 1983 147 Lower


Frankenwald GermanyEntactinosphaera

altasulcataE ndash ndash Won 1983 147 Lower



brevihebesE ndash ndash Won 1983 148 Lower



crassispinosaE ndash ndash Sashida amp Tonishi 1985 12 Upper Permian Changhsingian Ohirayama Unit South Chichibu


Entactinosphaera diplostraca


Entactinosphaera elegans

E ndash ndash Furutani 1990 41 Upper Silurian Gorstian Hitoegane Formation Hida-gaien Belt E of Hitoegane Fukuji area Takayama City Japan

Entactinosphaera erebenna


Entactinosphaera gengmaensis

ndash nn ndash Feng 1992 55 Upper Permian Changhsingian Papai Formation of Changning-Menglian Belt Papai Village Cangyuan County Yunnan China

Entactinosphaera hapala


Entactinosphaera hirta E ndash ndash Nazarov in Nazarov amp Popov 1980 39

Middle Ordovician


Entactinosphaera octospina



Entactinosphaera orientalis


Entactinosphaera polyacanthina


Entactinosphaera riedelii

ndash nn ndash Li 1995 332 Middle Ordovician


Entactinosphaera robusta


Entactinosphaera trendalli



Eoalbaillella (14)Eoalbaillella lilaensis Al ndash T Feng amp Liu 1992 550 Middle Devonian Givetian Lailei Formation Lila Village

Ximeng County Yunnan ChinaEoalbaillella turkensis Al ndash ndash Kozur 1997 49 Middle Devonian Givetian Gerence Karaburun peninsula

Izmir Region Turkey

Eoconcharium (339)Eoconcharium chinense nr is Th Hao amp Shu 1987 309 Lower Cambrian Terreneuvian Shuijingtuo Formation Shaanxi

Province Zhenba County ChinaEoconcharium chinense nr inv ndash Shu amp Chen 1989 992 Lower Cambrian Stage 3 Dengying Formation Xiaoyang

Village Zhenba Shaanxi Province Zhenba County China

Eoconcharium ovalnm nr is ndash Hao amp Shu 1987 309 Lower Cambrian Terreneuvian Shuijingtuo Formation Shaanxi Province Zhenba County China

Eoconcharium rhomboidale

nr is ndash Hao amp Shu 1987 309 Lower Cambrian Terreneuvian Shuijingtuo Formation Shaanxi Province Zhenba County China

Eoconcharium spiniferum nr is ndash Hao amp Shu 1987 309 Lower Cambrian Terreneuvian Shuijingtuo Formation Shaanxi Province Zhenba County China

EoscenidiumEoscenidium

hexaradiatumnr is ndash Wang 1989 156 Lower Cambrian ndash Niutitang Formation Sandu district

Guizhou Province ChinaEoscenidium longicrurum nr is ndash Wang 1989 156 Lower Cambrian ndash Damiao Formation Erlongping

Group Eastern Qinling ophiolite belt Henan Province China

Eoscenidium longiradiatum

nr is ndash Wang 1989 155 Lower Cambrian ndash Damiao Formation Erlongping Group Eastern Qinling ophiolite belt Henan Province China

Eoscenidium solidescens nr is Tnr Wang 1989 155 Lower Cambrian ndash Niutitang Formation Sandu district Guizhou Province China





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Stat

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Eostylodictya (136)Eostylodictya eccentrica E ndash Ts Ormiston amp Lane 1976 171 Lower



Eostylodictya palaeoeccentrica

E ndash ndash Won amp Seo 2010 252 Lower Carboniferous


Eostylodictya pericinctus spinosa



Eostylodictya speculum E ndash Ts Won 1983 152 Lower Carboniferous


Etymalbaillella (294)Etymalbaillella renzii IS

(OampF)ndash ndash Li 1995 338 Middle



Etymalbaillella toriterminolis

IS(OampF)

ndash ndash Li 1995 338 Middle Ordovician


Etymalbaillella yennienii IS(OampF)

ndash T Li 1995 338 Middle Ordovician


Follicucullus (31)Follicucullus bipartitus Al ndash ndash Caridroit amp De Wever 1984 640 Middle Permian Capitanian Oi Formation Ultra-Tamba Belt

Yanogawa River Yamasakicho-Kusune Shisou City Japan

Follicucullus bispinosus Al ndash ndash Kozur 1993 105 Upper Permian upper Changhsingian

Lamar Limestone Guadalupe Mts Culberson County West Texas USA

Follicucullus crenulatus Al ndash ndash Spiller 2002 26 Middle Permian ndash Lower Chert Member Semanggol Formation Pokok Sena NW Peninsular Malaysia

Follicucullus dactylinus Al ndash ndash Rudenko in Rudenko amp Panasenko 1990b 188

Upper Permian Wuchiapingian Pantov Member Tauka Terrane Tumanovk Pantov area Primorskiy Kray Russia

Follicucullus dilatatus Al ndash ndash Rudenko in Belyansky et al 1984 54

Upper Permian Capitanian Sebuchar Formation of Taukha Belt Dalrsquonny-Kut Bolrsquoshoy Ussurka Primorskiy Kray Russia

Follicucullus elongatus Al ndash ndash Spiller 2002 27 Middle Permian ndash Lower Chert Member Semanggol Formation Pokok Sena NW Peninsular Malaysia

Follicucullus falx Al ndash ndash Caridroit amp De Wever 1984 641 Middle Permian Capitanian Oi Formation Ultra-Tamba Belt Yanogawa River Yamasakicho-Kusune Shisou City Japan

Follicucullus furca Al ndash ndash Caridroit amp De Wever 1986 64 Upper Permian Wuchiapingian Oi Formation Ultra-Tamba Belt Yanogawa River Yamasakicho-Kusune Shisou City Japan

Follicucullus guangxiensis

Al ndash ndash Wang in Wang et al 2012 108 Upper Permian upper Changhsingian

Bancheng Formation SE Guangxi China

Follicucullus hamatus Al ndash ndash Caridroit amp De Wever 1984 642 Middle Permian Capitanian Oi Formation Ultra-Tamba Belt Yanogawa River Yamasakicho-Kusune Shisou City Japan

Follicucullus hamatus uncinatus

Al ndash ndash Caridroit amp De Wever 1984 642 Middle Permian Capitanian Oi Formation Ultra-Tamba Belt Yanogawa River Yamasakicho-Kusune Shisou City Japan

Follicucullus japonicus Al ndash ndash Ishiga 1991 108 Upper Permian Changhsingian Imajo Unit NW of the Nabejiri-yama area Taga Town Inukami-gun Shiga Japan

Follicucullus lagenarius Al ndash ndash Rudenko in Belyansky et al 1984 55


Follicucullus latior Al ndash ndash Feng amp Liu 1993 557 Upper Permian Changhsingian Laba Group SW Yunnan ChinaFollicucullus

monacanthusAl ndash ndash Ishiga amp Imoto in Ishiga et al

1982b 276Lower Permian Kungurian Funafuseyama Unit Tamba-Mino-

Ashio Belt Ashimi-dani section Ukyo-ku Kyoto Japan

Follicucullus orthogonus Al ndash Ts Caridroit amp De Wever 1984 642 Middle Permian Capitanian Oi Formation Ultra-Tamba Belt Yanogawa River Yamasakicho-Kusune Shisou City Japan

Follicucullus porrectus Al ndash ndash Rudenko in Belyansky et al 1984 55




Aitchison J C et al


er

Stat

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Follicucullus (31) continuationFollicucullus quadrataris Al ndash ndash Wang in Wang et al 1994 187 Lower Permian Kungurian Dachongling section Qinzhou City

Guangxi S ChinaFollicucullus scholasticus Al ndash Ts Ormiston amp Babcock 1979 333 Middle Permian Capitanian Bell Canyon Formation Delaware

Basin Culberson County W Texas USA

Follicucullus sphaericus Al ndash ndash Takemura in Takemura et al 1999 760

Middle Permian Capitanian Oruatemanu Formation Waipapa Terrane Arrow Rocks North Island New Zealand

Follicucullus ventricosus Al ndash T Ormiston amp Babcock 1979 332 Middle Permian Capitanian Bell Canyon Formation Delaware Basin Culberson County W Texas USA

Follicucullus whangaroaensis

Al ndash ndash Takemura in Takemura et al 1999 763


Follicucullus charveti Al ndash ndash Caridroit amp De Wever 1984 641 Middle Permian Capitanian Oi Formation Ultra-Tamba Belt Yanogawa River Yamasakicho-Kusune Shisou City Japan

Foremanconus (32)Foremanconus

postsakmaraensisAl ndash ndash Kozur amp Mostler 1989 180 Lower Permian Artinskian Sasayama-cho Takigun Hyogo

Prefecture Japan

Foremanhelena (172)Foremanhelena circula L ndash ndash Shang Caridroit amp Wang 2001

233Upper Permian Changhsingian Changshing Formation Liuqiao

section Fusui County southern Nanning Guangxi China

Foremanhelena intermixta

L ndash ndash Feng in Feng et al 2006b 839 Upper Permian Changhsingian Dalong Formation Dongpan section Dongpan Village Guangxi Zhuang Region China

Foremanhelena robusta L ndash ndash Feng in Feng et al 2006b 839 Upper Permian Changhsingian Dalong Formation Dongpan section Dongpan Village Guangxi Zhuang Region China

Foremanhelena triangula L ndash T De Wever amp Caridroit 1984 106 Upper Permian Wuchiapingian Oi Formation Ultra-Tamba Belt Yanogawa River Yamasakicho-Kusune Shisou City Japan

Foremaniella (225)Foremaniella helenae N ndash ndash Deflandre 1973c 1150 Lower


Franklinia (268)Franklinia dipulvisphaera S ndash ndash Jones amp Noble 2006 312 MIddle Silurian Sheinwoodian Cape Phillips Formation


Franklinia tricae S ndash T Th

Jones amp Noble 2006 310 MIddle Silurian Sheinwoodian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago

Fukujius (143)Fukujius yamakoshii E ndash Td Furutani 1990 47 Upper Silurian Pridoli Yoshiki Formation Hida-gaien Belt

Ichinotani Valley Fukuji area Takayama City Gifu Japan

Fungomacula (295)Fungomacula barbatula S ndash T Won in Won amp Below 1999 336 Middle Cambrian ndash Inca Formation Georgina Basin


Fusalfanus (257)Fusalfanus bilateralis S ndash ndash Tetard Noble amp Danelian in

Tetard et al 2015 871Upper Silurian Gorstian Cape Phillips Formation


Fusalfanus osobudaniensis

S ndash T Furutani 1990 38 Upper Silurian Gorstian Hitoegane Formation Hida-gaien Belt E of Hitoegane Fukuji area Takayama City Japan

Fusalfanus konomoriensis

S ndash ndash Aitchison Hada Ireland amp Yoshikura 1996 66


Fusuconcharium (340)Fusuconcharium

minimumnr is ndash Hao amp Shu 1987 309 Lower Cambrian Terreneuvian Shuijingtuo Formation Shaanxi

Province Zhenba County ChinaFusuconcharium typicum nr is Tnr Hao amp Shu 1987 309 Lower Cambrian Terreneuvian Shuijingtuo Formation Shaanxi

Province Zhenba County China





er

Stat

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Futobari (258)Futobari morishitai S ndash ndash Furutani 1990 35 Upper Silurian Pridoli Yoshiki Formation Hida-gaien Belt


Futobari solidus S ndash T Furutani 1990 34 Upper Silurian Pridoli Yoshiki Formation Hida-gaien Belt Ichinotani Valley Fukuji area Takayama City Gifu Japan

Futobari jingamoriensis S ndash ndash Aitchison Hada Ireland amp Yoshikura 1996 65


Futobari tosaensis S ndash ndash Aitchison Hada Ireland amp Yoshikura 1996 65


Gansuceratoikiscum (15)Gansuceratoikiscum

guanzhuangensisAl ndash T Wang in Wang et al 2010 472 Upper Ordovician Sandbian Pinglian Formation Guanzhuang

section Pinglang City Gansu China

Gedauia (137)Gedauia bibrachialis E ndash Ts Won 1983 153 Lower


Frankenwald Germany

Gigantospiculum (54)Gigantospiculum

dumitricaiAr ndash T Won amp Iams 2015a 17 Lower Ordovician Floian Cow Head Group Western

Newfoundland Canada

Glanta (16)Glanta crinerensis Al ndash ndash Schwartzapfel amp Holdsworth

1996 129Upper Devonian upper

Famennian Woodford Formation Locality 1

Carter County Oklahoma USA Glanta fragilis Al ndash T Wakamatsu Sugiyama amp

Furutani 1990 180Lower Devonian Emsian Nakahata Formation SW of Mt

Yokokurayama Ochi Town Shikoku Japan

Glanta triangularis Al ndash ndash Wakamatsu Sugiyama amp Furutani 1990 182


Glanta yokokurayamaensis

Al ndash ndash Umeda 1998a 101 Middle Devonian Eifelian Nakahata Formation Section C Ochi Town Shikoku Japan

Goodbodium (144)Goodbodium elegans E ndash ndash Furutani 1990 43 Upper Silurian Gorstian Hitoegane Formation Hida-gaien

Belt E of Hitoegane Fukuji area Takayama City Japan

Goodbodium nishiyamai E ndash ndash Furutani 1990 44 Upper Silurian Pridoli Yoshiki Formation Hida-gaien Belt Ichinotani Valley Fukuji area Takayama City Gifu Japan

Gracilentactinia (93)Gracilentactinia gracilis E ndash T Kozur amp Mostler 1989 193 Lower Permian Kungurian Koshelev Foramtion Alogazovo

Village Movsovic Region Russia

Gracilentactinia sphaerica

E ndash ndash Kozur amp Mostler 1989 193 Lower Permian Sakmarian Sarabil Formation Sakmarskiy Orenburgskaya Region S Urals Russia

Grandetortura (200)Grandetortura nipponica L ndash T Sashida amp Tonishi 1991 92 Upper Permian Changhsingian Ohirayama Unit South Chichibu


Grosmorneus (55)Grosmorneus globosus Ar ndash T Won amp Iams 2002 31 Upper Cambrian Stage 10 Cow Head Group Western

Newfoundland Canada

Guangxitrisphaera (315)Guangxitrisphaera

wanyaoensisndash nd Td Wang amp Kuang 1993 281 Lower

CarboniferousTournaisian Shijia Formation Shijia Resevoir


Gustefana (316)Gustefana

obliqueannulatandash nd Td Kozur 1993 111 Upper Permian Wuchiapingian Sosio Valley Area Torrente San




Aitchison J C et al


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Gyrosphaera (250)Gyrosphaera cavea S ndash ndash Jones amp Noble 2006 304 Middle Silurian Sheinwoodian Cape Phillips Formation


Gyrosphaera primigena S ndash ndash Umeda amp Suzuki 2005 87 Lower Silurian Aeronian Kallholn Formation Solberga Quarry of Boda Siljan area Dalarna Sweden

Gyrosphaera raneatela S ndash T Noble amp Maletz 2000 267 Lower Silurian Telchyian Kallholn Formation Solberga Quarry of Boda Siljan area Dalarna Sweden

Gyrosphaera siljanensis S ndash ndash Noble amp Maletz 2000 267 Lower Silurian Telchyian Kallholn Formation Solberga Quarry of Boda Siljan area Dalarna Sweden

HalicalyptraHalicalyptra aculeata ndash nd ndash Ruumlst 1892 179 Lower Devonian ndash Southern Urals RussiaHalicalyptra ambulans ndash nd ndash Ruedemann amp Wilson 1936



Halicalyptra similis ndash nd ndash Ruedemann amp Wilson 1936 1578

Lower Ordovician Floian Normanskill chert Fly Summit Washington County New York USA

HalicapsaHalicapsa megapora ndash nd ndash Ruumlst 1892 181 Upper Permian ndash Sosio Valley Area Western Sicily

Italy

HaliommaHaliomma antiquum ndash nd ndash Ruedemann amp Wilson 1936



Haliomma cornutum ndash nd ndash Hinde 1890 51 Middle Ordovician


Haliomma paternum ndash nn ndash Deflandre 1960 216 Lower Carboniferous


Haliomma penrosei ndash nd ndash Ruedemann amp Wilson 1936 1573


Haliomma perfecta ndash nd ndash Aberdeen 1940 138 Upper Devonian Famennian Caballos Formation Marathon Basin Texas USA

Haliomma speciosum ndash nd ndash Ruumlst 1892 148 Upper Permian ndash Sosio Valley Area Western Sicily Italy

Haliomma stigmophorum ndash nd ndash Ruumlst 1892 148 Lower Carboniferous


Haliomma vetustum ndash nd ndash Hinde 1890 51 Middle Ordovician


Haplentactinia (134)Haplentactinia alekseevi E ndash ndash Afanasieva 2000c 374 Upper Devonian middle



Haplentactinia armillata E ndash ndash Nazarov in Nazarov amp Popov 1980 55

Middle Ordovician


Haplentactinia armista E ndash ndash Siveter Aitchison Siveter amp Sutton 2007 91

Middle Silurian Homerian Wenlock Series Herefordshire England UK

Haplentactinia arrhinia E ndash Ts Foreman 1963 270 Upper Devonian Famennian Huron member Ohio Shale Ohio USA

Haplentactinia attenuata E ndash ndash Goto Umeda amp Ishiga 1992 156


Haplentactinia baltica E ndash ndash Nazarov in Nazarov amp Nolvak 1983 5


Haplentactinia barskovi E ndash ndash Afanasieva 2000c 375 Upper Devonian middle Frasnian


Haplentactinia bicepsispina



Haplentactinia bornazi E ndash ndash Afanasieva 2000a 31 Upper Devonian middle Frasnian






er

Stat

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Haplentactinia (134) continuationHaplentactinia flagellifera E ndash ndash Nazarov amp Ormiston 1993 40 Upper Devonian Famenian Zapadno Valavskaya borehole 1-R

Pripyat Depression BelarusHaplentactinia flagellifera ndash nn ndash Nazarov 1988 129 Upper Devonian Famenian Zapadno Valavskaya borehole 1-R

Pripyat Depression BelarusHaplentactinia inaudata ndash nn ndash Nazarov 1988 129 Upper Devonian Famenian Zapadno Valavskaya borehole 1-R

Pripyat Depression BelarusHaplentactinia infida E ndash ndash Nazarov in Nazarov et al 1975

102Middle



Haplentactinia infida E ndash ndash Nazarov in Nazarov et al 1977 919

Middle Ordovician


Haplentactinia jinzhangii ndash nn ndash Li 1995 332 Middle Ordovician


Haplentactinia juncta E ndash ndash Nazarov 1975 58 Middle Ordovician


Haplentactinia kuzminae E ndash ndash Afanasieva amp Amon 2011 1504 Upper Devonian middle Frasnian

Pechora Basin Ukhtinskii District Ukhta River Russia

Haplentactinia minia ndash nn ndash Li 1995 332 Middle Ordovician


Haplentactinia rhinophyusa


Haplentactinia silurica E ndash ndash Nazarov amp Ormiston 1990 18 Lower Silurian ndash Sakmarsk Formation Bolrsquoshoy Abimevo Village Bashkortostan Region S Urals Russia


Haplentactinia silurica E ndash ndash Nazarov 1988 122 Lower Silurian ndash Sakmarsk Formation Bolrsquoshoy Abimevo Village Bashkortostan Region S Urals Russia

Haplentactinia simplex E ndash ndash Wang in Wang amp Luo 2009 136 Upper Devonian Frasnian Xiangshuidong Formation Bazhai Ziyun County Guizhou China

Haplentactinia vilvaensis E ndash ndash Afanasieva amp Amon 2011 1502 Upper Devonian lower Famennian

Vilva River of the Perm Region Middle Urals Russia

Haplentactinia ichikawai E ndash ndash Caridroit amp De Wever 1984 643 Middle Permian Capitanian Oi Formation Ultra-Tamba Belt Yanogawa River Yamasakicho-Kusune Shisou City Japan

Haplentactinia ozawai E ndash ndash Furutani 1990 40 Upper Silurian Gorstian Hitoegane Formation Hida-gaien Belt E of Hitoegane Fukuji area Takayama City Japan

Haplentactinia pentochela

E ndash ndash Li 1994 263 Upper Silurian Gorstian Keerhada Mayila Mts Xinjiang China

Haploaxon (190)Haploaxon melorax ndash nd ndash Cordey 1998 53-54 Lower Permian Artinskian SW of Aspen Grove British

Columbia CanadaHaploaxon merrittensis ndash nd Ts Cordey 1998 53 Lower Permian Artinskian SW of Aspen Grove British

Columbia Canada

Haplodiacanthus (33)Haplodiacanthus

anfractusAl ndash Ts Nazarov amp Rudenko 1981 133 Lower Permian Artinskian Kandurov Formation Donskoye

Ural River Orenburgskaya Region S Urals Russia

Haplodiacanthus circinatus

Al ndash ndash Nazarov amp Ormiston 1985 47 Upper Carboniferous

Gzhelian E of Saraktash Nikol Village Ural River Bashkirsk Region Urals Russia

Haplodiacanthus levitoflexus


Upper Carboniferous


Haplodiacanthus levitoflexus fenistratus


Upper Carboniferous




Aitchison J C et al


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Stat

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HaploplusHaploplus qunii ndash nn ndash Li 1995 332 Middle



HaplopolarisHaplopolaris dipolaris ndash nn ndash Li 1991 75 Upper Silurian Gorstian Mayila Complex Keerhada S

Mayila Mt Xinjiang China

Haplopolus (269)Haplopolus dipolus S ndash Ts Li 1994 263 Upper Silurian Gorstian Keerhada Mayila Mts Xinjiang

China

Haplotaeniatum (251)Haplotaeniatum

cathenatumS ndash ndash Nazarov amp Ormiston 1993 41 Lower Silurian ndash Sakmarsk Formation Bolrsquoshoy


Haplotaeniatum cathenatum

ndash nn ndash Nazarov 1988 122 Lower Silurian ndash Sakmarsk Formation Bolrsquoshoy Abimevo Village Bashkortostan Region S Urals Russia

Haplotaeniatum circulus S ndash ndash Pouille amp Danelian in Pouille et al 2014 157

Middle Ordovician

upper Darriwilian

Shundy Formation Balkhash region Kazakhstan

Haplotaeniatum fenestratum

S ndash ndash Goto Umeda amp Ishiga 1992 158


Haplotaeniatum fissura S ndash ndash MacDonald 2006 20 Middle Silurian Sheinwoodian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago

Haplotaeniatum labyrintheum

S ndash Th Nazarov amp Ormiston 1990 20 Lower Silurian ndash Sakmarsk Formation Bolrsquoshoy Abimevo Village Bashkortostan Region S Urals Russia


ndash inv ndash Nazarov amp Ormiston 1993 41 Lower Silurian ndash Sakmarsk Formation Bolrsquoshoy Abimevo Village Bashkortostan Region S Urals Russia



Haplotaeniatum leptum ndash nn ndash Li 1995 332 Middle Ordovician


Haplotaeniatum nunavutensis

S ndash ndash MacDonald 2006 23 Middle Silurian Sheinwoodian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago

Haplotaeniatum ovatum S ndash ndash Noble amp Webby 2009 557 Upper Ordovician Katian Malongulli Formation Cliefden Caves area NSW Australia

Haplotaeniatum prolatum S ndash ndash Noble amp Webby 2009 557 Upper Ordovician Katian Malongulli Formation Cliefden Caves area NSW Australia

Haplotaeniatum spinatum S ndash ndash Goto Umeda amp Ishiga 1992 157


Haplotaeniatum tegimentum

S ndash ndash Nazarov amp Ormiston 1990 20 Lower Silurian ndash Sakmarsk Formation Bolrsquoshoy Abimevo Village Bashkortostan Region S Urals Russia





Haplotaeniatum vertigospongum

S ndash ndash Jones amp Noble 2006 302 MIddle Silurian Sheinwoodian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago

Haplotaeniatum aperturatum

S ndash ndash Noble Ketner amp McClellan 1997 721

Lower Silurian Rhuddanian Cherry Spring Chert Garden Pass Roberts Mts Allochthon Nevada USA

Haplotaeniatum giganteum

S ndash ndash Pouille amp Danelian in Pouille et al 2014 160

Middle Ordovician

upper Darriwilian


HapolpolusHapolpolus ruedemannii ndash nn ndash Li 1995 332 Middle







er

Stat

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Hegleria (270)Hegleria agnusiforma S ndash ndash Noble amp Jin 2010 140 Middle Permian Capitanian Bell Canyon Formation Guadalupe

Mts Culberson County W Texas USA

Hegleria globosa S ndash ndash Noble amp Jin 2010 141 Middle Permian Capitanian Bell Canyon Formation Guadalupe Mts Culberson County W Texas USA

Hegleria mammifera S ndash T Nazarov amp Ormiston 1985 22 Middle Permian Capitanian Bell Canyon Formation Guadalupe Mts Culbertson County W Texas USA

Helenifore (17)Helenifore gogoense Al ndash ndash Aitchison 1993b 111 Upper Devonian lower Frasnian Gogo Formation Pillara Range


Helenifore laticlavium Al ndash T Nazarov amp Ormiston 1983a 464 Upper Devonian Frasnian Sadler Range SE Cap Creek Gap Canning Basin Western Australia

Helenifore pilosidiscus Al ndash ndash Aitchison et al 1999 158 Middle Devonian Eifelian Gamilaroi terrane Silver Gully Nundle NSW Australia

Helenifore planus Al ndash ndash Umeda 1997 418 Upper Silurian Gorstian Jyoru Formation Kurosegawa Belt Konomori area Kochi City Shikoku Japan

Helenifore quadrispina Al ndash ndash Noble amp Lenz 2007 1050 Middle Silurian Homerian Cape Phillips Formation Rookery Creek Cornwallis Island Canadian Arctic Archipelago

Helenifore fasciola Al ndash ndash Nazarov amp Ormiston 1993 52 Middle Silurian Homerian Maksyutov Complex Tarangul River 10 km N of Kosistek Village S Urals Russia

HelenoforeHelenofore fasciola ndash nn ndash Nazarov 1988 122 Upper Silurian Pridoli Maksyutov Complex Tarangul


HeliodiscusHeliodiscus acucinctus ndash nd ndash Ruumlst 1892 165 Lower Silurian ndash Cabriegraveres Heacuterault FranceHeliodiscus saturnalis ndash nd Td Ruumlst 1892 165 Lower


Helioentactinia (94)Helioentactinia aster E ndash ndash Aitchison 1993b 118 Upper Devonian lower Frasnian Gogo Formation Pillara Range


Helioentactinia biexosphaera


Upper Carboniferous


Helioentactinia circumtexta


Helioentactinia gudymovae



Helioentactinia ikka E ndash ndash Nazarov amp Ormiston 1990 16 Lower Permian Sakmarian Dalny Tulkas Rill Belskaya depression Bashkortostan Region S Urals Russia


Helioentactinia ikka ndash nn ndash Nazarov 1988 156 Lower Permian Artinskian Dalny Tulkas Rill Belskaya depression Bashkortostan Region S Urals Russia

Helioentactinia nazarovi E ndash ndash Sashida amp Tonishi 1985 12 Upper Permian Changhsingian Ohirayama Unit South Chichibu Belt Kashiwara Akigawa River Akiruno City Tokyo Japan

Helioentactinia perjucunda

E ndash ndash Nazarov amp Ormiston 1983a 460 Upper Devonian Frasnian Sadler Range SE Cap Creek Gap Canning Basin Western Australia

Helioentactinia secutrix E ndash ndash Nazarov 1975 90 Upper Devonian Frasnian Egindinsk Formation Aitpaika River N Mugodazhar S Urals Kazakhstan

Helioentactinia stellaepolus




Aitchison J C et al


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Stat

usTy




Helioentactinia (94) continuationHelioentactinia uralica E ndash ndash Kozur amp Mostler 1989 193 Lower Permian Sakmarian Sarabil Formation Sakmarskiy


Helioentactinia valavica E ndash ndash Nazarov amp Ormiston 1993 34 Upper Devonian Famenian Zapadno Valavskaya borehole 1-R Pripyat Depression Belarus

Helioentactinia valavica ndash nn ndash Nazarov 1988 130 Upper Devonian Famenian Zapadno Valavskaya borehole 1-R Pripyat Depression Belarus

Helioentactinia yunnaensis

E ndash ndash Wang in Wang et al 2000 248 Middle Devonian Givetian Changyucun Formation Shaijingpo section Xianyun County Yunnan China

Helioentactinia asymmetrica

E ndash ndash Nazarov 1975 88 Middle Cambrian ndash Agyirsksk Formation Mukhyr River Chingiz Mountain range eastern Kazakhstan

Helioentactinia bakanasensis

E ndash T Nazarov 1975 89 Upper Cambrian Furongian Ushkyzyilrsquosk Formation Ushkyzyl Mts Gora Ushkyzyl Gora AkshataQyzylord E Kazakhstan

Helioentactinia prismspinosa

E ndash ndash Wakamatsu Sugiyama amp Furutani 1990 168


HeliosestrumHeliosestrum nigrum E ndash ndash Hinde 1899a 54 Middle Devonian Givetian Yarramie Formation Tamworth


HeliosomaHeliosoma echinatum E ndash ndash Hinde 1899a 50 Middle Devonian Givetian Yarramie Formation Tamworth

Belt northern NSW AustraliaHeliosoma mojsisovicsi E ndash ndash Ruumlst 1892 148 Lower


Heliosoma paronae E ndash ndash Hinde 1899a 50 Middle Devonian Givetian Yarramie Formation Tamworth Belt northern NSW Australia

Heliosoma roemeri ndash nd ndash Ruumlst 1892 148 Lower Silurian ndash Cabriegraveres Heacuterault France

HeliosphaeraHeliosphaera alternata ndash nd ndash Aberdeen 1940 138 Upper Devonian Famennian Caballos Formation Marathon

Basin Texas USAHeliosphaera bardanum ndash nd ndash Ruumlst 1892 147 Lower


Heliosphaera clavata ndash nd ndash Hinde 1899a 50 Middle Devonian Givetian Yarramie Formation Tamworth Belt northern NSW Australia

Heliosphaera fenestrata ndash nd ndash Hinde 1899a 49 Middle Devonian Givetian Yarramie Formation Tamworth Belt northern NSW Australia

Heliosphaera haeckeli ndash nd ndash Ruedemann amp Wilson 1936 1572


Heliosphaera kjrulfi ndash nd ndash Ruumlst 1892 147 Lower Carboniferous


Heliosphaera macrospinosa


Heliosphaera micropora ndash nd ndash Ruedemann amp Wilson 1936 1572


Heliosphaera robusta E ndash ndash Hinde 1899a 49 Middle Devonian Givetian Yarramie Formation Tamworth Belt northern NSW Australia

Heliosphaera ruesti ndash nd nc

ndash Ruedemann amp Wilson 1936 1572


Heliosphaera tamworthi E ndash ndash Hinde 1899a 49 Middle Devonian Givetian Yarramie Formation Tamworth Belt northern NSW Australia

Heliosphaera venusta ndash nd ndash Ruedemann amp Wilson 1936 1572


Helminentactinia (259)Helminentactinia nazarovi ndash nn Td Li 1991 75 Upper Silurian Gorstian Mayila Complex Keerhada S

Mayila Mt Xinjiang ChinaHelminentactinia nazarovi ndash nd ndash Li 1994 268 Upper Silurian Gorstian Keerhada Mayila Mts Xinjiang

ChinaHelminentactinia nazarovi

longispinandash nn ndash Li 1991 75 Upper Silurian Gorstian Mayila Complex Keerhada S






er

Stat

usTy




HemisphaeraHemisphaera diflandrei ndash nn ndash Li 1995 332 Middle



HeptacladusHeptacladus permicus E ndash ndash Kozur amp Mostler 1989 184 Lower Permian Sakmarian Sarabil Formation Sakmarskiy


HexacladusHexacladus speciosus ndash nn ndash Deflandre 1960 216 Lower


HexalastrumHexalastrum infans ndash nd ndash Ruumlst 1892 172 Carboniferous ndash Buumlkk-Gebirge Inner Western

Carpathians Hungary

HexaloncheHexalonche palaeozoica E ndash ndash Ruumlst 1892 146 Upper Devonian ndash SchaebenholzHarz Mountains

Elbingerode GermanyHexalonche valida E ndash ndash Ruumlst 1892 146 Upper Permian ndash Sosio Valley Area Western Sicily

Italy

HexastylidiumHexastylidium variatum ndash nd ndash Aberdeen 1940 136 Upper Devonian Famennian Caballos Formation Marathon

Basin Texas USA

HexastylusHexastylus basiporosus ndash nd ndash Aberdeen 1940 136 Upper Devonian Famennian Caballos Formation Marathon

Basin Texas USAHexastylus ferox ndash nn ndash Deflandre 1960 216 Lower


Hexastylus montisnigrae ndash nn ndash Deflandre 1960 216 Lower Carboniferous


Holdisphaera (95)Holdisphaera furutanii E ndash T Kozur amp Mostler 1989 187 Lower Permian Sakmarian Sarabil Formation Sakmarskiy


Holdisphaera laevis E ndash ndash Kozur amp Mostler 1989 187 Lower Permian Sakmarian Sarabil Formation Sakmarskiy Orenburgskaya Region S Urals Russia

Holdsworthella (34)Holdsworthella perforata Al ndash ndash Kozur 1981 269 Lower Permian Sakmarian Sarabil Formation Kondurovska


Holdsworthella permica Al ndash Ts Kozur 1981 268 Lower Permian Sakmarian Sarabil Formation Kondurovska Orenburgskaya Region S Urals Russia

Holdsworthum (145)Holdsworthum japonicus E ndash T Furutani 1990 45 Upper Silurian Pridoli Yoshiki Formation Hida-gaien Belt


Holoeciscus (18)Holoeciscus auceps Al ndash T Foreman 1963 294 Upper Devonian Famennian Huron member Ohio Shale Ohio

USAHoloeciscus brevis Al ndash ndash Cheng 1986 90 Upper Devonian Famennian Woodford Formation TI Valley


Holoeciscus elongatus Al ndash ndash Kiessling amp Tragelehn 1994 230Upper Devonian Famennian Frankenwald north Bavaria Germany

Holoeciscus foremanae Al ndash ndash Cheng 1986 91 Upper Devonian Famennian Woodford Formation TI Valley section Pittsburg County Oklahoma USA

Holoeciscus longus Al ndash ndash Schwartzapfel amp Holdsworth 1996 126



Holoeciscus quasiauceps Al ndash ndash Wang 1997 158 Upper Devonian Famennian Gennaren Formation S Utubulak NW Jungar Basin Xinjiang China

Holoeciscus renzae Al ndash ndash Schwartzapfel amp Holdsworth 1996 128





Aitchison J C et al


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Huasha (2)Huasha densa Al ndash ndash Cheng 1986 71 Lower



Huasha holdsworthi Al ndash Ts Cheng 1986 73 Lower Carboniferous


Huasha magnifica Al ndash ndash Cheng 1986 74 Lower Carboniferous


Huasha quinquecostata Al ndash ndash Cheng 1986 74 Lower Carboniferous


Inaequalientactinia (96)Inaequalientactinia

deflandreiE ndash ndash Won 1991b 101 Lower

CarboniferousTournaisian Boulders in Quaternary Rhine

Terrace near Oberkassel BonnInaequalientactinia

gourmelonaeE ndash ndash Won 1991b 102 Lower


Terrace near Oberkassel BonnInaequalientactinia parva E ndash ndash Won 1991b 103 Lower


Terrace near Oberkassel BonnInaequalientactinia typica E ndash Ts Won 1991b 103 Lower



wuppertalensisE ndash ndash Won 1992 203 Lower

CarboniferousTournaisian Riescheid Section Wuppertal-

Barmen Germany

Inanibigutta (260)Inanibigutta goodbodyii S nn ndash Li 1995 332 Middle



Inanibigutta ishigea S nn ndash Li 1995 332 Middle Ordovician


Inanibigutta keerhadaensis

S nn ndash Li 1991 75 Upper Silurian Gorstian Mayila Complex Keerhada S Mayila Mt Xinjiang China


S ndash ndash Li 1994 261 Lower Devonian Emsian Keerhada Mayila Mts Xinjiang China

Inanibigutta llanvirniana S nn ndash Li 1995 332 Middle Ordovician


Inanibigutta maletzi S ndash ndash Pouille amp Danelian in Pouille et al 2013 1154

Middle Ordovician

upper Darriwilian


Inanibigutta minuta S ndash ndash Wang 1993c 100 Upper Ordovician Sandbian Pinglian Formation Horizon Pg 3 Yindongguan village Pinglian County Gansu China

Inanibigutta minutia S nn ndash Li 1995 332 Middle Ordovician


Inanibigutta pinglianensis S ndash ndash Wang 1993c 100 Upper Ordovician Sandbian Pinglian Formation Horizon Pg 3 Yindongguan village Pinglian County Gansu China

InanigiguttaInanigigutta yaoi S nn ndash Li 1995 332 Middle



Inanigutta (261)Inanigutta gansuensis S ndash ndash Wang 1993c 99 Upper Ordovician Sandbian Pinglian Formation Horizon Pg 3

Yindongguan village Pinglian County Gansu China

Inanigutta hanquanii S nn ndash Li 1995 333 Middle Ordovician


Inanigutta haoroui S nn ndash Li 1995 333 Middle Ordovician


Inanigutta magnifica S ndash ndash Maletz amp Bruton 2008 1193 Middle Ordovician

lower Darriwilian

Valhallfonna Formation Buldrebreen arm Ny Frieslan Spitsbergen Norway





er

Stat

usTy




Inanigutta (261) continuationInanigutta yujingii S nn ndash Li 1995 333 Middle



Inanigutta zhiyuanii S nn ndash Li 1995 333 Middle Ordovician


Inanigutta kyrgyza S ndash ndash Danelian Popov amp Ghobadi Pour 2011 591

Lower Ordovician Floian Sarydzhaz eastern Kyrgyzstan

Inanihella (262)Inanihella duroacus S nn ndash Nazarov amp Ormiston 1993 37 Middle Silurian Homerian Maksyutov Complex Tarangul

River 10 km N Kosistek Village S Urals Russia

Inanihella duroacus S ndash ndash Nazarov 1988 122 Upper Silurian Pridoli Maksyutov Complex Tarangul River 10 km N of Kosistek Village S Urals Russia

Inanihella legincula S nn ndash Nazarov 1988 122 Middle Silurian Homerian Maksyutov Complex Tarangul River 10 km N of Kosistek Village S Urals Russia

Inanihella legiuncula S ndash ndash Nazarov amp Ormiston 1993 38 Middle Silurian Homerian Maksyutov Complex Tarangul River 10 km N Kosistek Village S Urals Russia

Inanihella perarmata S ndash ndash Nazarov amp Ormiston 1993 38 Middle Silurian Homerian Maksyutov Complex Tarangul River 10 km N Kosistek Village S Urals Russia

Inanihella perarmata S nn ndash Nazarov 1988 122 Middle Silurian Homerian Maksyutov Complex Tarangul River 10 km N of Kosistek Village S Urals Russia

Inanihella sagena S ndash ndash Siveter Aitchison Siveter amp Sutton 2007 89


Inanihella tarangulica S ndash ndash Nazarov amp Ormiston 1984 73 Middle Silurian Homerian Makyutov Complex Tarangul River Kosistek Village Southern Ural Kazakhstan

Inanihella vulnerata S nn ndash Nazarov 1988 122 Middle Silurian Homerian Maksyutov Complex Tarangul River 10 km N of Kosistek Village S Urals Russia

Inanihella akzhala S ndash ndash Danelian amp Popov 2003 333 Lower Ordovician Floian Akzhal Mts S of Betpak-Dala desert Kazakhstan

Induropilarius (317)Induropilarius

aenigmaticusE nn Tn Ters amp Deflandre 1966 340 Ordovician ndash Phtanites drsquoAngers Vendeacutee

France

Insolitignum (146)Insolitignum peranima E ndash ndash MacDonald 1999 2054 Lower Silurian Telychian Cape Phillips Formation


Insolitignum vivanima E ndash ndash MacDonald 1999 2055 Lower Silurian Telychian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago

Intracorpus (271)Intracorpus octaedron S ndash T Won 1997a 366 Upper Devonian Frasnian Gogo Formation Pillara Range


Involutentactinia (97)Involutentactinia

eccentraE ndash T Jones amp Noble 2006 297 MIddle Silurian Sheinwoodian Cape Phillips Formation


Involutentactinia eccentra var cupressa

E ndash ndash Jones amp Noble 2006 297 MIddle Silurian Sheinwoodian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago

Ishigaum (173)Ishigaum complanum L ndash ndash Feng in Feng et al 2006b 837 Upper Permian Changhsingian Dalong Formation Dongpan


Ishigaum craticula L ndash ndash Shang Caridroit amp Wang 2001 233




Aitchison J C et al


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Stat

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Ishigaum (173) continuationIshigaum fusinum L ndash ndash Feng in Feng et al 2006b 837 Upper Permian Changhsingian Dalong Formation Dongpan


Ishigaum klaengensis L ndash ndash Sashida in Sashida et al 2000b 254

Upper Permian Changhsingian Ratuburi Group Sukhothai Fold Belt section I Khao Wang Chik Rayong Thailand

Ishigaum labaensis L ndash ndash Feng amp Liu 1993 545 Upper Permian Changhsingian Muyinhe Formation Nanpan Lancang County Yunnan China

Ishigaum longispina L ndash ndash Feng in Feng et al 2006b 837 Upper Permian Changhsingian Dalong Formation Dongpan section Dongpan Village Guangxi Zhuang Region China

Ishigaum obesum L ndash ndash De Wever amp Caridroit 1984 100 Middle Permian Capitanian Oi Formation Ultra-Tamba Belt Yanogawa River Yamasakicho-Kusune Shisou City Japan

Ishigaum trifustis L ndash T De Wever amp Caridroit 1984 99 Middle Permian Capitanian Oi Formation Ultra-Tamba Belt Yanogawa River Yamasakicho-Kusune Shisou City Japan

Ishigaum tristylum L ndash ndash Feng in Feng et al 2006b 837 Upper Permian Changhsingian Dalong Formation Dongpan section Dongpan Village Guangxi Zhuang Region China

Kalimnasphaera (263)Kalimnasphaera

maculosaS ndash T Webby amp Blom 1986 152 Upper Ordovician Katian Malongulli Formation Belubula

River E of Canowindra central NSW Australia

Kantollum (230)Kantollum bicornum N ndash ndash Cheng 1986 163 Upper

CarboniferousBashkirian Johns Valley Shale Pittsburg

County eastern Oklahoma USAKantollum blancoensis N ndash ndash Cheng 1986 165 Upper Devonian Famennian Woodford Formation TI Valley


Kantollum crinerensis N ndash ndash Schwartzapfel amp Holdsworth 1996 149



Kantollum longicornutum N ndash ndash Wang in Wang et al 2012 112 Upper Devonian Famennian Shiti Reservoir Formation Shiwu road section Guangxi Zhuang Region SW China

Kantollum pittsburgense N ndash T Cheng 1986 164 Upper Devonian Famennian Woodford Formation TI Valley section Pittsburg County Oklahoma USA

Kantollum undulatum N ndash ndash Cheng 1986 164 Upper Devonian Famennian Woodford Formation TI Valley section Pittsburg County Oklahoma USA

Kappaforma (19)Kappaforma insecta Al ndash T Noble amp Lenz 2007 1052 Middle Silurian Homerian Cape Phillips Formation Rookery

Creek Cornwallis Island Canadian Arctic Archipelago

Kashiwara (98)Kashiwara magna E ndash T Sashida amp Tonishi 1985 15 Upper Permian Changhsingian Ohirayama Unit South Chichibu


Klaengspongus (244)Klaengspongus formosus S ndash ndash Feng in Feng et al 2009 147 Upper Permian Changhsingian Dalong Formation Dongpan


Klaengspongus planus S ndash ndash Nestell amp Nestell 2010 25 Middle Permian Capitanian Bell Canyon Formation Apache Mountains Culberson County W Texas USA

Klaengspongus spinosus S ndash T Sashida in Sashida et al 2000b 256


Klaengspongus umbilicatus

S ndash ndash Feng in Feng et al 2009 147 Upper Permian Changhsingian Dalong Formation Dongpan section Dongpan Village Guangxi Zhuang Region China

Konyrium (341)Konyrium varium nr is Tnr Nazarov in Nazarov amp Popov

1976 415Middle







er

Stat

usTy




Labyrinthia (236)Labyrinthia inexpectata S ndash T Maletz amp Bruton 2007 281 Lower Ordovician Floian Valhallfonna Formation


Labyrinthia robusta S ndash ndash Maletz 2007 76 Lower Ordovician Floian Cow Head Group Western Newfoundland Canada

Labyrinthosphaera (252)Labyrinthosphaera lancia S ndash ndash Jones amp Noble 2006 298 MIddle Silurian Sheinwoodian Cape Phillips Formation


Labyrinthosphaera macdonaldi

S ndash T Noble amp Maletz 2000 268 Lower Silurian Telchyian Kallholn Formation Solberga Quarry of Boda Siljan area Dalarna Sweden

Labyrinthosphaera lenzi S ndash ndash Jones amp Noble 2006 298 MIddle Silurian Sheinwoodian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago

Lacisus (342)Lacisus siphunculus nr is Tnr Kozur 1993 109 Upper Permian Wuchiapingian Sosio Valley Area Torrente San


Lapidopiscum (4)Lapidopiscum piri Al ndash ndash Schwartzapfel amp Holdsworth



Carter County Oklahoma USA Lapidopiscum piveteaui Al ndash T Deflandre 1958 2278 Lower


Lapidopiscum transversum




Latentibifistula (183)Latentibifistula

asperspongiosaL ndash ndash Sashida amp Tonishi 1986 7 Upper Permian Changhsingian Ohirayama Unit South Chichibu


Latentibifistula triacanthophora

L ndash T Nazarov amp Ormiston 1983b 374Lower Permian Sakmarian Maloik Formation Don Village Ural River Orenburgskaya Region S Urals Russia

Latentidiota (210)Latentidiota choripelata L nc ndash Nazarov in Isakova amp Nazarov

1986 82Upper


Urals RussiaLatentidiota clarisona L ndash ndash Nazarov in Isakova amp Nazarov

1986 79Upper


Urals RussiaLatentidiota fallacia L ndash ndash Nazarov in Isakova amp Nazarov

1986 82Upper


Urals RussiaLatentidiota medorensis L ndash ndash Nestell Pope amp Nestell 2012

240Upper



Latentidiota semilamina L ndash ndash Nazarov in Isakova amp Nazarov 1986 78

Upper Carboniferous


Latentidiota trigimena L ndash ndash Nazarov amp Ormiston 1984 81 Upper Carboniferous

Gzhelian Makyutov Complex Tarangul River Kosistek Village Southern Ural Kazakhstan

Latentidiota tripetalina L nc ndash Nazarov in Isakova amp Nazarov 1986 80

Upper Carboniferous


Latentidiota visenda L ndash T Nazarov amp Ormiston 1985 29 Upper Carboniferous

Gzhelian Maloik Formation Don Village Ural River Orenburgskaya Region S Urals Russia

Latentidiota ambigua L nc ndash Nazarov in Isakova amp Nazarov 1986 81

Upper Carboniferous


Latentifistula (184)Latentifistula actinoteres L ndash ndash Nazarov in Isakova amp Nazarov

1986 87Upper


Urals RussiaLatentifistula astricta L ndash ndash Nazarov in Isakova amp Nazarov

1986 88Upper


Urals RussiaLatentifistula astricta

astrictaL ndash ndash Nazarov in Isakova amp Nazarov

1986 89Upper



solidaL ndash ndash Nazarov in Isakova amp Nazarov

1986 90Upper


Urals Russia



Aitchison J C et al


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Stat

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Latentifistula (184) continuationLatentifistula

banchengensisL ndash ndash Wang in Wang et al 2012 109 Lower Permian Artinskian Bancheng Formation SE Guangxi

ChinaLatentifistula brauni L ndash ndash Wang in Wang et al 2012 109 Upper

CarboniferousGzhelian Shijia Formation Bancheng


Latentifistula crux L ndash T Nazarov amp Ormiston 1983b 372Upper Carboniferous


Latentifistula densa L ndash ndash Nazarov amp Ormiston 1985 34 Middle Permian Capitanian Bell Canyon Formation Guadalupe Mts Culbertson County W Texas USA

Latentifistula hetroextrema

L ndash ndash Nazarov in Isakova amp Nazarov 1986 90

Upper Carboniferous


Latentifistula mushroomformis

L ndash ndash Wang in Wang et al 2012 109 Lower Permian Artinskian Bancheng Formation SE Guangxi China

Latentifistula muyinheensis


Latentifistula neotenica L ndash T Nazarov amp Ormiston 1985 33 Middle Permian Roadian Bell Canyon Formation Guadalupe Mts Culbertson County W Texas USA

Latentifistula patagilaterala

L ndash ndash Nazarov amp Ormiston 1985 33 Middle Permian Capitanian Bell Canyon Formation Hegler Limestone Culberson County W Texas USA

Latentifistula similicutis L ndash ndash Caridroit amp De Wever 1986 78 Upper Permian Wuchiapingian Oi Formation Ultra-Tamba Belt Yanogawa River Yamasakicho-Kusune Shisou City Japan

Latentifistula texana L ndash ndash Nazarov amp Ormiston 1985 33 Lower Permian Artinskian Kandurov Formation Donskoye Ural River Orenburgskaya Region S Urals Russia

Latentifistula torulosa L ndash ndash Nazarov in Isakova amp Nazarov 1986 85

Upper Carboniferous


Latentifistula torulosa ampliata


Upper Carboniferous


Latentifistula triradiata L ndash ndash Wang 1993a 451 Lower Permian Kungurian Gufeng Formation Qiaotou Village Chaou City Anhui China

Latentifistula valdeinepta L ndash Ts Nazarov amp Ormiston 1985 33 Lower Permian Sakmarian Kandurov Formation Donskoye Ural River Orenburgskaya Region S Urals Russia

LiosphaeraLiosphaera devoniensis ndash nd ndash Ruumlst 1892 136 Lower Devonian ndash Southern Urals RussiaLiosphaera mera ndash nd ndash Ruumlst 1892 136 Lower Silurian ndash Cabriegraveres Heacuterault France

LithapiumLithapium claviformis IS ndash ndash Nazarov 1973b 10 Lower Cambrian Stage 2 Bograd Mountains Betenevskiy

area Khakasiya Region RussiaLithapium siluricum ndash nd ndash Ruumlst 1892 154 Lower Silurian ndash Cabriegraveres Heacuterault FranceLithapium tesiensis IS ndash ndash Nazarov 1973b 10 Lower Cambrian Stage 2 Bograd Mountains Betenevskiy

area Khakasiya Region RussiaLithatractus

Lithatractus brevispinus ndash nd ndash Ruumlst 1892 158 Upper Permian ndash Sosio Valley Area Western Sicily Italy

Lithatractus mejeri ndash nd ndash Ruumlst 1892 157 Lower Carboniferous


Lithatractus perforatus ndash nd ndash Ruumlst 1892 158 Upper Permian ndash Sosio Valley Area Western Sicily Italy

LitheliusLithelius difficilis ndash nd ndash Ruumlst 1892 175 Lower


Lithelius sinensis S ndash ndash Feng in Feng et al 2006a 35 Upper Permian Changhsingian Dalong Formation Dongpan section Dongpan Village Guangxi Zhuang Region China

LithocampeLithocampe lossenii ndash nd ndash Ruumlst 1892 189 Lower Devonian ndash Southern Urals RussiaLithocampe orenburgensis ndash nd ndash Ruumlst 1892 188 Lower Devonian ndash Southern Urals RussiaLithocampe pyramis ndash nd ndash Ruumlst 1892 189 Upper Permian ndash Sosio Valley Area Western Sicily

Italy





er

Stat

usTy




Lithocampe (continuation)Lithocampe

sphaerocephalicandash nd ndash Ruumlst 1892 189 Upper Permian ndash Sosio Valley Area Western Sicily

ItalyLithocampe stenostoma ndash nd ndash Ruumlst 1892 189 Upper Permian ndash Sosio Valley Area Western Sicily

ItalyLithocampe

tschernytschewiindash nd ndash Ruumlst 1892 189 Lower Devonian ndash Southern Urals Russia

Lithocampe tutata ndash nd ndash Ruumlst 1892 190 Carboniferous ndash Buumlkk-Gebirge Inner Western Carpathians Hungary

Lithocampe spinosa ndash nd ndash Ruedemann amp Wilson 1936 1579


Lithocannosphaeropsis (296)Lithocannosphaeropsis

fallaxndash nn Tn Deflandre 1960 216 Lower


Lithocannosphaeropsis taylori

ndash nd ndash Ormiston amp Lane 1976 175 Lower Carboniferous


LithocycliaLithocyclia devoniensis ndash nd ndash Hinde amp Fox 1895 639 Carboniferous Serpukhovian-

BashikirianCodden Hill bed Devon England

UKLithocyclia macrococcus E ndash T Ruumlst 1892 166 Upper Devonian ndash SchaebenholzHarz Mountains

Elbingerode GermanyLithocyclia ulrichi E ndash Ts Ruumlst 1892 166 Lower


LithomespilusLithomespilus bipolaris ndash nd ndash Ruumlst 1892 154 Lower


Lithomespilus hexacanthus

ndash nd ndash Ruumlst 1892 153 Lower Silurian ndash Langenstriegis Germany

Lithomespilus steinvorthi ndash nd ndash Ruumlst 1892 154 Upper Devonian ndash SchaebenholzHarz Mountains Elbingerode Germany

Lithosphaera (44)Lithosphaera oesensis Ar ndash Th Won 1998 257 Lower

CarboniferousTournaisian Oese Rheinische Schiefergebirge

Germany

LithostrobusLithostrobus wendlandi ndash nd ndash Ruumlst 1892 187 Upper Permian ndash Sosio Valley Area Western Sicily

Italy

Longtanella (36)Longtanella

mengshengensisAl nn ndash Feng 1992 54 Upper Permian Changhsingian Gufeng Formation Hubei China

Longtanella turgida Al ndash ndash Feng 1992 59 Middle Permian Capitanian Gufeng Formation Hubei ChinaLongtanella

zhengpanshanensisAl ndash Ts Sheng amp Wang 1985 175 179 Lower Permian Kungurian Gufeng Formation N Mt

Konghshan Hushan Nanjing China

Magnentactinia (99)Magnentactinia fragilis E ndash T Won 1997a 366 Upper Devonian Frasnian Gogo Formation Pillara Range


Magnentactinia ostentata E ndash ndash Jones amp Noble 2006 297 MIddle Silurian Sheinwoodian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago

Magnentactinia hexagonia



Magnisphaera (100)Magnisphaera

aitchisonianaE ndash ndash Won 1997a 367 Upper Devonian Frasnian Gogo Formation Pillara Range

Canning Basin Western AustraliaMagnisphaera gigantea E ndash T Won 1997a 368 Upper Devonian Frasnian Gogo Formation Pillara Range

Canning Basin Western AustraliaMagnisphaera imperfecta E ndash ndash Won 1997a 368 Upper Devonian Frasnian Gogo Formation Pillara Range




Aitchison J C et al


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Stat

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Megaporus (297)Megaporus jini IS

(OampF)ndash T Feng in Feng et al 2006c S68 Upper Permian Changhsingian Dalong Formation Dongpan


Megaporus unicum IS(OampF)

ndash ndash Feng in Feng et al 2006c S70 Upper Permian Changhsingian Dalong Formation Dongpan section Dongpan Village Guangxi Zhuang Region China

Megaporus yangi IS(OampF)


Megaporus yini IS(OampF)


Meschedea (101)Meschedea akcetensis E ndash ndash Noble Tekin Gedik amp Pehlivan

2008 53Lower

CarboniferousTournaisian Baltalimani Formation Istanbul

TurkeyMeschedea crassicortex E ndash ndash Won 1997a 360 Upper Devonian Frasnian Gogo Formation Pillara Range


Meschedea hirsuta E ndash ndash Won 1983 154 Lower Carboniferous


Meschedea permica E ndash ndash Sashida amp Tonishi 1985 15 Upper Permian Changhsingian Ohirayama Unit South Chichibu Belt Kashiwara Akigawa River Akiruno City Tokyo Japan

Meschedea pyramispinosa

E ndash Ts Won 1983 154 Lower Carboniferous


Meschedea endocarpa E ndash ndash Nazarov amp Ormiston 1993 38 Middle Permian Wordian Bone Spring Formation Guadalupe Mts Culbertson County W Texas USA

Microporosa (102)Microporosa aktastensis E ndash ndash Afanasieva amp Amon 2016 214 Lower Permian Artinskian Northern Mugodzhary Aktasty

River KazakhstanMicroporosa rozhnovi E ndash ndash Afanasieva amp Amon 2016 213 Lower Permian Artinskian Northern Mugodzhary Aktasty

River Kazakhstan

Moskovistella (103)Moskovistella allbori E nn ndash Afanasieva 1997a 221 Upper Devonian middle



Moskovistella allbori E nn ndash Afanasieva 1997b 40 Upper Devonian middle Frasnian


Moskovistella allbororum E ndash ndash Afanasieva 2000b 138 Upper Devonian middle Frasnian


Moskovistella baccata E ndash ndash Afanasieva 2000a 67 Upper Devonian middle Frasnian


Moskovistella khaini E nn ndash Afanasieva 1997a 221 Upper Devonian middle Frasnian


Moskovistella khaini E nn ndash Afanasieva 1997b 40 Upper Devonian middle Frasnian


Moskovistella khaini E ndash ndash Afanasieva 2000b 135 Upper Devonian middle Frasnian


Moskovistella lucet E nn ndash Afanasieva 1997b 40 Upper Devonian middle Frasnian


Moskovistella lucet E ndash ndash Afanasieva 2000a 68 Upper Devonian middle Frasnian


Moskovistella mira E nn ndash Afanasieva 1997b 40 Upper Devonian middle Frasnian


Moskovistella mira E ndash ndash Afanasieva 2000b 140 Upper Devonian middle Frasnian






er

Stat

usTy




Moskovistella (103) continuationMoskovistella octoradiata E ndash T Afanasieva 2000b 136 Upper Devonian middle



Moskovistella rozanovi E nn ndash Afanasieva 1997b 40 Upper Devonian middle Frasnian


Moskovistella rozanovi E ndash ndash Afanasieva 2000b 136 Upper Devonian middle Frasnian


Moskovistella sincera E nn ndash Afanasieva 1997b 40 Upper Devonian middle Frasnian


Moskovistella sincera E ndash ndash Afanasieva 2000b 135 Upper Devonian middle Frasnian


Moskovistella viatoria E nn ndash Afanasieva 1997b 40 Upper Devonian middle Frasnian


Moskovistella viatoria E ndash ndash Afanasieva 2000b 70 Upper Devonian middle Frasnian


Moskovistella victorialis E nn ndash Afanasieva 1997a 221 Upper Devonian middle Frasnian


Moskovistella victorialis E nn ndash Afanasieva 1997b 40 Upper Devonian middle Frasnian


Moskovistella victorialis E ndash ndash Afanasieva 2000b 138 Upper Devonian middle Frasnian


Mostlerispongus (204)Mostlerispongus

sosioensisL ndash T Kozur 1993 110 Upper Permian Wuchiapingian Sosio Valley Area Torrente San


Mostlerium (226)Mostlerium unicum N ndash Ts Cheng 1986 136 Lower



Multientactinia (104)Multientactinia

inconstansE ndash T Won 1997b 379 Upper Devonian Frasnian Gogo Formation Pillara Range


Multientactinia iniqua E ndash ndash Won 1997b 379 Upper Devonian Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Multientactinia iniqua cancellibasis

E ndash ndash Won 1997b 380 Upper Devonian Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Multientactinia iniqua iniqua


Multientactinia reedae E ndash ndash Won 1997b 380 Upper Devonian Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Multientactinia soluta E ndash ndash Won 1997b 380 Upper Devonian Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Multisphaera (164)Multisphaera impersepta E ndash T Nazarov amp Afanasieva in

Afanasieva 2000c 75Lower Permian Artinskian Kandurov Formation Donskoye


Nabespecha (191)Nabespecha leonardia L ndash Ts Cornell amp Simpson 1986 285 Middle Permian Roadian Bone Spring Formation

Guadalupe Mts Culbertson County W Texas USA

Nazaromistonella (20)Nazaromistonella

speciosusAl ndash Ts Furutani 1990 53 Upper Silurian Gorstian Hitoegane Formation Hida-gaien




Aitchison J C et al


er

Stat

usTy




Nazarovella (192)Nazarovella gracilis L ndash Ts De Wever amp Caridroit 1984 101 Middle Permian Capitanian Oi Formation Ultra-Tamba Belt


Nazarovella inflata L ndash ndash Sashida amp Tonishi 1986 10 Upper Permian Changhsingian Ohirayama Unit South Chichibu Belt Kashiwara Akigawa River Akiruno City Tokyo Japan

Nazarovella phlogides L ndash ndash Wang in Wang amp Li 1994 206 Middle Permian Capitanian Dazong Range Guangxi Xiadong Qinzhou area China

Nazarovella scalae L ndash ndash Caridroit amp De Wever 1986 83 Middle Permian Capitanian Oi Formation Ultra-Tamba Belt Yanogawa River Yamasakicho-Kusune Shisou City Japan

Nazarovispongus (211)Nazarovispongus

globosumL ndash ndash Nestell amp Nestell 2010 58 Middle Permian Capitanian Bell Canyon Formation Apache


Nazarovispongus pavlovi L ndash Ts Kozur 1980 238 Lower Permian Sakmarian Sarabil Formation Verchneozernaja Station Orenburgskaya Region Urals Russia

Nazarovispongus permicus

L ndash ndash Kozur 1980 238 Lower Permian Sakmarian Sarabil Formation Verchneozernaja Station Orenburgskaya Region Urals Russia

Nazarovites (21)Nazarovites aprelevkensis Al ndash ndash Afanasieva 2000c 360 Upper Devonian middle



Nazarovites bioculus Al ndash Ts Afanasieva 2000c 364 Upper Devonian middle Frasnian


Nazarovites mikhailovae Al nn ndash Afanasieva 1997a 221 Upper Devonian middle Frasnian


Nazarovites mikhailovae Al ndash ndash Afanasieva 2000c 365 Upper Devonian middle Frasnian


Nazarovites pinnula Al ndash ndash Afanasieva 2000c 365 Upper Devonian middle Frasnian


Neoalbaillella (5)Neoalbaillella antaixiangi Al ndash ndash Yao amp Kuwahara 1999 12 Upper Permian Changhsingian Dalong Formation Changjianggou

section N of Shangsi Sichuan China

Neoalbaillella camarata Al ndash ndash Wu amp Feng in Wu et al 2010 886

Upper Permian Changhsingian Dalong Formation Dongpan section Dongpan Village Guangxi Zhuang Region China

Neoalbaillella cephalota Al ndash ndash Wu amp Feng in Wu et al 2010 886


Neoalbaillella cribrosa Al ndash ndash Rudenko amp Panasenko 1990a 121


Neoalbaillella gracilis Al ndash ndash Takemura amp Nakaseko 1981 213

Upper Permian Changhsingian Funafuseyama Unit Tamba-Mino-Ashio Belt Mt Kurogara-dake Takatsuki City Osaka Japan

Neoalbaillella grypus Al ndash ndash Ishiga Kito amp Imoto 1982a 16 Upper Permian Changhsingian Funafuseyama Unit Tamba-Mino-Ashio Belt Ubara section Fukuchiyama City Kyoto Japan

Neoalbaillella minuta Al ndash ndash Jin amp Feng in Jin et al 2007 16 Upper Permian Changhsingian Dalong Formation Dongpan section Dongpan Village Guangxi Zhuang Region China

Neoalbaillella optima Al ndash ndash Ishiga Kito amp Imoto 1982a 16 Upper Permian Changhsingian Funafuseyama Unit Tamba-Mino-Ashio Belt Ubara section Fukuchiyama City Kyoto Japan

Neoalbaillella ornithoformis

Al ndash T Takemura amp Nakaseko 1981 211


Neoalbaillella pseudogrypus

Al ndash ndash Sashida amp Tonishi 1988 527 Upper Permian Changhsingian Ohirayama Unit South Chichibu Belt Kashiwara Akigawa River Akiruno City Tokyo Japan





er

Stat

usTy




Neoaspiculum (286)Neoaspiculum

cancelliumIS ndash ndash Won amp Iams 2011 165 Lower Ordovician Floian Cow Head Group Western

Newfoundland CanadaNeoaspiculum echinatum IS ndash ndash Won amp Iams 2015a 29 Lower Ordovician Floian Cow Head Group Western

Newfoundland CanadaNeoaspiculum headense IS ndash T Won amp Iams 2011 165 Lower Ordovician Floian Cow Head Group Western

Newfoundland CanadaNeoaspiculum headense

delicatumIS ndash ndash Won amp Iams 2011 168 Lower Ordovician Floian Cow Head Group Western


headenseIS ndash ndash Won amp Iams 2011 165 Lower Ordovician Floian Cow Head Group Western

Newfoundland CanadaNeoaspiculum laxum IS ndash ndash Won amp Iams 2011 168 Lower Ordovician Floian Cow Head Group Western

Newfoundland CanadaNeoaspiculum

transformumIS ndash ndash Won amp Iams 2011 168 Lower Ordovician Floian Cow Head Group Western

Newfoundland CanadaNeoaspiculum amplum IS ndash ndash Won amp Iams 2011 168 Lower Ordovician Floian Cow Head Group Western


maletzianaIS ndash ndash Won amp Iams 2015a 29 Lower Ordovician Floian Cow Head Group Western

Newfoundland Canada

Neoechidnina (56)Neoechidnina compacta Ar ndash T Won amp Iams 2015a 15 Lower Ordovician Floian Cow Head Group Western

Newfoundland Canada

Neoholoeciscus (22)Neoholoeciscus

cancermimusAl ndash T Ormiston amp Lane 1976 174 Lower



Neopalaeospiculum (65)Neopalaeospiculum

densum Ar ndash ndash Won Iams amp Reed 2007 527 Lower Ordovician lower-middle


Newfoundland CanadaNeopalaeospiculum

laxum Ar ndash T Won Iams amp Reed 2007 525 Lower Ordovician lower-middle



transformum Ar ndash ndash Won Iams amp Reed 2007 527 Lower Ordovician lower-middle


Newfoundland Canada

Noblella (61)Noblella tremadociensis Ar ndash Th

TsKozur Mostler amp Repetski 1996



NodocapsaNodocapsa multipartita nr Is Tnr Wang 1989 145 Lower Cambrian ndash Niutitang Formation Sandu


Nyfrieslandia (287)Nyfrieslandia complicata IS ndash ndash Maletz amp Bruton 2007 281 Lower Ordovician Floian Valhallfonna Formation


Nyfrieslandia crassispinosa

IS ndash ndash Won amp Iams 2013 30 Lower Ordovician Floian Cow Head Group Western Newfoundland Canada

Nyfrieslandia sphaeroidea

IS ndash T Maletz amp Bruton 2007 278 Lower Ordovician Floian Valhallfonna Formation Buldrebreen arm Ny Frieslan Spitsbergen Norway

Octatormentum (201)Octatormentum

babcockaeL ndash ndash Nazarov amp Ormiston 1985 43 Middle Permian Capitanian Bell Canyon Formation Guadalupe


Octatormentum cornelli L ndash T Nazarov amp Ormiston 1985 42 Middle Permian Roadian Bell Canyon Formation Guadalupe Mts Culbertson County W Texas USA

Octatormentum floriferum

L ndash ndash Sashida amp Tonishi 1988 533 Upper Permian Changhsingian Ohirayama Unit South Chichibu Belt Kashiwara Akigawa River Akiruno City Tokyo Japan

Octatormentum yaoi L ndash ndash Wu amp Feng in Wu et al 2010 889




Aitchison J C et al


er

Stat

usTy




OdontosphaeraOdontosphaera

echinocactusndash nd ndash Ruumlst 1892 140 Lower Silurian ndash Cabriegraveres Heacuterault France

Orbiculopylorum (253)Orbiculopylorum

adobensisS ndash ndash Noble Ketner amp McClellan

1997 721Lower Silurian Rhuddanian Cherry Spring Chert Garden

Pass Roberts Mts Allochthon Nevada USA

Orbiculopylorum granti S ndash ndash MacDonald 2006 29 Middle Silurian Sheinwoodian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago

Orbiculopylorum marginatum

S ndash T Noble Ketner amp McClellan 1997 719


Orbiculopylorum splendens



Oriundogutta (264)Oriundogutta bella S ndash ndash Wang 1993c 101 Upper Ordovician Sandbian Pinglian Formation Horizon Pg 3


Oriundogutta litterula S ndash ndash Nazarov amp Ormiston 1990 18 Lower Silurian ndash Sakmarsk Formation Bolrsquoshoy Abimevo Village Bashkortostan Region S Urals Russia

Oriundogutta litterula S ndash ndash Nazarov amp Ormiston 1993 36 Middle Silurian Homerian Maksyutov Complex Tarangul River 10 km N Kosistek Village S Urals Russia

Oriundogutta litterula S nn ndash Nazarov 1988 122 Middle Silurian Homerian Maksyutov Complex Tarangul River 10 km N of Kosistek Village S Urals Russia

Oriundogutta miscella minuta

S ndash ndash Wang 1993c 101 Upper Ordovician Sandbian Pinglian Formation Horizon Pg 3 Yindongguan village Pinglian County Gansu China

Oriundogutta nazarovi S ndash ndash Wang 1993c 102 Upper Ordovician Sandbian Pinglian Formation Horizon Pg 3 Yindongguan village Pinglian County Gansu China

Oriundogutta qilianenesis S nn ndash Li 1995 333 Middle Ordovician


Oriundogutta kingi S ndash ndash Noble 1994 31 Upper Silurian Gorstian Caballos Novaculite Formation Payne Hills Bourland Mts Marathon Basin W Texas USA

Oriundogutta varispina S ndash ndash Noble 1994 31 Upper Silurian Gorstian Caballos Novaculite Formation Monument Ck Bourland Mts Marathon Basin W Texas USA

Ormistonella (193)Ormistonella adhaerens L ndash ndash Feng in Feng et al 2006b 845 Upper Permian Changhsingian Dalong Formation Dongpan


Ormistonella hickoryensis L ndash ndash Schwartzapfel amp Holdsworth 1996 216

Lower Carboniferous

Visean Caney Shale Arbuckle Mountains Hickory Creek section Carter County Oklahoma USA

Ormistonella robusta L ndash T De Wever amp Caridroit 1984 100 Middle Permian Capitanian Oi Formation Ultra-Tamba Belt Yanogawa River Yamasakicho-Kusune Shisou City Japan

Ormistonella turgida ndash nn ndash Feng 1992 55 Upper Permian Changhsingian Papai Formation of Changning-Menglian Belt Papai Village Cangyuan County Yunnan China

Ormistonella cylindrata L ndash ndash Feng in Feng et al 2006b 845 Upper Permian Changhsingian Dalong Formation Dongpan section Dongpan Village Guangxi Zhuang Region China

Ormistonia (299)Ormistonia pteracaena ndash nn Td Li 1991 75 Upper Silurian Gorstian Mayila Complex Keerhada S

Mayila Mt Xinjiang ChinaOrmistonia pteracaena ndash nd Th

TdLi 1994 271 Upper Silurian Gorstian Keerhada Mayila Mts Xinjiang

China





er

Stat

usTy




Ornatoentactinia (106)Ornatoentactinia agarkovi E ndash ndash Afanasieva 2000a 56 Upper Devonian middle



Ornatoentactinia beljaevorum



Ornatoentactinia klevtsovae



Ornatoentactinia solita E ndash Ts Afanasieva 2000c 373 Upper Devonian middle Frasnian

Domanik Formation Locality 4 Ukhta River Timan-Pechora Basin Komi Region Russia

Ornatoentactinia spartaci E ndash ndash Afanasieva 2000c 374 Upper Devonian middle Frasnian


Ornatoentactinia spartaki E nn ndash Afanasieva 1997a 221 Upper Devonian middle Frasnian


Ornatoentactinia spinisica



Ouaka (185)Ouaka asymmetrica L ndash Ts Cheng 1986 176 Upper



Ouaka ruesti L ndash ndash Cheng 1986 177 Upper Carboniferous


Pactarentinia (147)Pactarentinia holdsworthi E ndash T Furutani 1983 109 Lower Devonian Emsian Nakahata Formation S Mt

Yokokurayama Ochi Town Kochi Shikoku Japan

Pactarentinia igoi E ndash ndash Kurihara amp Sashida 2000 61 Lower Devonian Emsian Shibasudani Formation Shibasudani Valley Ohno City Fukui Prefecture Japan

Pactarentinia intermedia E ndash ndash Kurihara amp Sashida 2000 61 Lower Devonian Emsian Shibasudani Formation Shibasudani Valley Ohno City Fukui Prefecture Japan

Palacantholithus (40)Palacantholithus crux Al nn ndash Deflandre 1973a 291 Lower


Palacantholithus crux Al ndash ndash Deflandre 1973b 499 Lower Carboniferous


Palacantholithus curvativus

Al ndash ndash Afanasieva 2000a 120 Upper Devonian middle Frasnian


Palacantholithus stellatus Al nn ndash Deflandre 1973a 291 Lower Carboniferous


Palacantholithus stellatus Al ndash T Deflandre 1973b 499 Lower Carboniferous


Palaeoactinosphaera (300)Palaeoactinosphaera

anticaIS

(OampF)ndash T Noble 1994 34 Upper Silurian Gorstian Caballos Novaculite Formation


Palaeoactinosphaera asymmetrica

IS(OampF)

ndash ndash Noble 1994 34 Middle Silurian Homerian Caballos Novaculite Formation Payne Hills Bourland Mts Marathon Basin W Texas USA

Palaeoactinosphaera barricki

IS(OampF)

ndash ndash Noble 1994 34 Upper Silurian Gorstian Caballos Novaculite Formation Payne Hills Bourland Mts Marathon Basin W Texas USA

Palaeoactinosphaera elegantissima

IS(OampF)


Palaeoactinosphaera crucispina

IS(OampF)


Palaeoactinosphaera octaspina

IS(OampF)

ndash ndash Noble 1994 35 Middle Silurian Homerian Caballos Novaculite Formation East Bourland Mts Marathon Basin W Texas USA



Aitchison J C et al


er

Stat

usTy




PalaeocryptidiumPalaeocryptidium cayeuxi ndash nn ndash Deflandre 1955 184 Cambrian ndash Ville-au-Roi-en-Maroueacute lamballe

Brittany France

Palaeodecaradium (148)Palaeodecaradium

gordoniE ndash ndash MacDonald 2004 268 Middle Silurian Sheinwoodian Cape Phillips Formation


Palaeodecaradium umbelliforme

E ndash T Goodbody 1986 136 Lower Silurian Telychian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago

Palaeodiscaleksus (333)Palaeodiscaleksus

saturniformisnr ndash ndash Afanasieva amp Amon 2011 1511 Middle Devonian Givetian Shipunikha Formation

Zmeinogorskii District Shipunikha River Rudny Altai West Siberia Russia

Palaeodiscaleksus shuldakensis

nr ndash ndash Afanasieva amp Amon 2012 550 Middle Devonian Givetian Shuldak Formation Shuldak River western Mugodzhary Kazakhstan

Palaeodiscaleksus tumefactus

nr ndash ndash Afanasieva amp Amon 2008 462 Middle Devonian Upper Eifelian Kariyukmas Mountain NW Staryi Sibai Southern Ural Russia

PalaeoellipsoidesPalaeoellipsoides

convexocylindratusL nd ndash Afanasieva amp Amon 2012 550 Middle Devonian Givetian Shuldak Formation Shuldak

River western Mugodzhary Kazakhstan

Palaeoellipsoides planoconvexus

L nd ndash Afanasieva amp Amon 2012 550 Middle Devonian Givetian Shuldak Formation Shuldak River western Mugodzhary Kazakhstan

Palaeoephippium (149)Palaeoephippium adraini E ndash ndash MacDonald 1999 2056 Lower Silurian Telychian Cape Phillips Formation


Palaeoephippium aranea E ndash ndash Goodbody 1986 138 Middle Silurian Sheinwoodian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago

Palaeoephippium bifurcum

E ndash T Goodbody 1986 139 Middle Silurian Sheinwoodian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago

Palaeoephippium dissimile


Palaeoephippium echinatum

E ndash ndash Goodbody 1986 140 Lower Silurian Telychian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago

Palaeoephippium fimbriatum


Palaeoephippium fukujiensis

E ndash ndash Furutani 1990 46 Upper Silurian Pridoli Yoshiki Formation Hida-gaien Belt Osobudani Valle Fukuji area Takayama City Gifu Japan

Palaeoephippium irregulum



Palaeoephippium multiramosum


Palaeoephippium octaramosum

E ndash ndash Renz 1990 370 Upper Ordovician Katian Hanson Creek Formation N Martin Ridge Eureka County Nevada USA

Palaeoephippium pariradiatum

E ndash ndash Goodbody 1986 143 Middle Silurian Sheinwoodian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago

Palaeoephippium pariramosum


Palaeoephippium plattum E nc ndash Goto Umeda amp Ishiga 1992 164


Palaeoephippium radices E ndash ndash Goodbody 1986 144 Middle Silurian Sheinwoodian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago





er

Stat

usTy




Palaeoephippium (149) continuationPalaeoephippium

ramipendentesE ndash ndash Goodbody 1986 146 Middle Silurian Sheinwoodian Cape Phillips Formation


Palaeoephippium reteforme


Palaeoephippium spinosum


Palaeoephippium tricorne


Palaeoephippium unipariradiatum



Palaeolithochytris (318)Palaeolithochytris lethaea ndash nd ndash Li 1995 336 Middle



Palaeolithochytris olenus ndash nd Td Li 1995 336 Middle Ordovician


Palaeolithocyclia (139)Palaeolithocyclia pilata E ndash ndash Feng in Feng et al 2009 143 Upper Permian Changhsingian Dalong Formation Dongpan


Palaeolithocyclia platta E ndash ndash Feng in Feng et al 2009 143 Upper Permian Changhsingian Dalong Formation Dongpan section Dongpan Village Guangxi Zhuang Region China

Palaeopentactinorbis (107)Palaeopentactinorbis

longispinousE ndash Ts Kozur amp Mostler 1989 195 Lower Permian Sakmarian Sarabil Formation Sakmarskiy


Palaeopyramidium (150)Palaeopyramidium

arcuatumE ndash ndash Amon Braun amp Ivanov 1995 6 Upper Silurian Gorstian Maksyutov Complex Tarangul


Palaeopyramidium furutanium



Palaeopyramidium ramosum

E ndash ndash Kurihara amp Sashida 2000 62 Lower Devonian Emsian Shibasudani Formation Shibasudani Valley Ohno City Fukui Prefecture Japan

Palaeopyramidium spinosum


Palaeorubus (343)Palaeorubus hastingensis nr Calc-

areaTnr Ishiga in Ishiga et al 1987 300 Upper Devonian Frasnian Yarras Hastings block New

England fold belt NSW Australia

Palaeoscenidium (151)Palaeoscenidium

apertumE ndash ndash Goodbody 1986 150 Middle Silurian Sheinwoodian Cape Phillips Formation


Palaeoscenidium appendium



Palaeoscenidium bicorne E nn ndash Deflandre 1960 216 Lower Carboniferous


Palaeoscenidium bispina E nn ndash Li 1995 333 Middle Ordovician


Palaeoscenidium cancellatum


Palaeoscenidium cladophorum

E ndash T Deflandre 1953 408 Lower Carboniferous


Palaeoscenidium daktylethra




Aitchison J C et al


er

Stat

usTy




Palaeoscenidium (151) continuationPalaeoscenidium

delicatum E ndash ndash Aitchison 1993b 122 Upper Devonian lower Frasnian Gogo Formation Pillara Range


Palaeoscenidium deweveri

E ndash ndash Gourmelon 1987 127 Lower Carboniferous


Palaeoscenidium echinatum


Palaeoscenidium flammatum

E ndash Ts Goodbody 1986 152 Middle Silurian Sheinwoodian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago

Palaeoscenidium fragilis E ndash ndash Kurihara amp Sashida 2000 60 Lower Devonian Emsian Shibasudani Formation Shibasudani Valley Ohno City Fukui Prefecture Japan

Palaeoscenidium hakogasensis


Palaeoscenidium ishigai E ndash ndash Wakamatsu Sugiyama amp Furutani 1990 166


Palaeoscenidium kuriharai

E ndash ndash Umeda amp Suzuki 2005 85 Lower Silurian Aeronian Kallholn Formation Solberga Quarry of Boda Siljan area Dalarna Sweden

Palaeoscenidium nudum E ndash ndash Aitchison 1993b 122 Upper Devonian lower Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Palaeoscenidium phalangium


Palaeoscenidium rarispinosum


Palaeoscenidium robustum


Palaeoscenidium scaurum



Palaeoscenidium schaafi E ndash ndash Gourmelon 1987 128 Lower Carboniferous


Palaeoscenidium simplum


Palaeoscenidium tabernaculum


Palaeoscenidium venustum


Palaeoscenidium quadriramosum


Palaeosphaera (70)Palaeosphaera micra Ar ndash Ts Renz 1990 372 Upper Ordovician Katian Hanson Creek Formation N Martin


Palaeospiculum (66)Palaeospiculum

arcussimileAr ndash ndash Won in Won amp Below 1999 340 Middle Cambrian ndash Inca Formation Georgina Basin

Queensland AustraliaPalaeospiculum

burkensisAr ndash T Won in Won amp Below 1999 340 Middle Cambrian ndash Inca Formation Georgina Basin

Queensland AustraliaPalaeospiculum curvum Ar ndash ndash Won Iams amp Reed 2005 451 Lower Ordovician lower-middle


Newfoundland CanadaPalaeospiculum

dendroeidesAr ndash ndash Won in Won amp Below 1999 341 Middle Cambrian ndash Inca Formation Georgina Basin


devoncourtensisAr ndash ndash Won in Won amp Below 1999 341 Middle Cambrian ndash Inca Formation Georgina Basin


georginaensisAr ndash ndash Won in Won amp Below 1999 341 Middle Cambrian ndash Inca Formation Georgina Basin






er

Stat

usTy




Palaeospiculum (66) continuationPalaeospiculum

greenpointensisAr ndash ndash Won amp Iams 2002 27 Upper Cambrian Stage 10 Cow Head Group Western


grosmornensisAr ndash ndash Won amp Iams 2002 27 Upper Cambrian Stage 10 Cow Head Group Western


hexaradiatumAr ndash ndash Won amp Iams 2002 28 Upper Cambrian Stage 10 Cow Head Group Western


martinpointensisAr ndash ndash Won amp Iams 2002 28 Upper Cambrian Stage 10 Cow Head Group Western


multifurcatumAr ndash ndash Won Iams amp Reed 2005 451 Lower Ordovician lower-middle



neofurcatumAr ndash ndash Won Iams amp Reed 2005 451 Lower Ordovician lower-middle


Newfoundland CanadaPalaeospiculum parvum Ar ndash ndash Won in Won amp Below 1999 346 Middle Cambrian ndash Inca Formation Georgina Basin

Queensland AustraliaPalaeospiculum radiatum Ar ndash ndash Won in Won amp Below 1999 348 Middle Cambrian ndash Inca Formation Georgina Basin

Queensland AustraliaPalaeospiculum reedae Ar ndash ndash Won in Won amp Below 1999 350 Middle Cambrian ndash Inca Formation Georgina Basin

Queensland AustraliaPalaeospiculum solutum Ar ndash ndash Won amp Iams 2015a 13 Lower Ordovician Floian Cow Head Group Western


tetractinumAr ndash ndash Won Iams amp Reed 2005 453 Lower Ordovician lower-middle



tetraradiatumAr ndash ndash Won amp Iams 2002 28 Upper Cambrian Stage 10 Cow Head Group Western

Newfoundland CanadaPalaeospiculum variabile Ar ndash ndash Won amp Iams 2015a 13 Lower Ordovician Floian Cow Head Group Western


discordiosumAr ndash ndash Won amp Iams 2015a 14 Lower Ordovician Floian Cow Head Group Western

Newfoundland Canada

Palaeothalomnus (152)Palaeothalomnus

antiquusIS

(OampF)nn ndash Deflandre 1973a 290 Lower


Palaeothalomnus antiquus

IS(OampF)

ndash T Deflandre 1973b 498 Lower Carboniferous


Palaeothalomnus piletocladus

IS(OampF)

ndash ndash Nazarov amp Ormiston 1993 50 Upper Devonian Famenian Zapadno Valavskaya borehole 1-R Pripyat Depression Belarus


IS(OampF)

nn ndash Nazarov 1988 130 Upper Devonian Famenian Zapadno Valavskaya borehole 1-R Pripyat Depression Belarus

Palaeothalomnus timokhini

IS(OampF)

nn ndash Afanasieva 1997a 221 Upper Devonian middle Frasnian



IS(OampF)

ndash ndash Afanasieva 2000c 366 Upper Devonian middle Frasnian


Palaeotrifidus (153)Palaeotrifidus ballator E ndash Ts Renz 1990 374 Upper Ordovician Katian Hanson Creek Formation N Martin


Palaeotrifidus imbifurcus E ndash ndash Renz 1990 374 Upper Ordovician Katian Hanson Creek Formation N Martin Ridge Eureka County Nevada USA

Palaeotripus (154)Palaeotripus cancellatus E ndash ndash Goodbody 1986 153 Middle Silurian Sheinwoodian Cape Phillips Formation


Palaeotripus gogoense E ndash ndash Aitchison 1993b 124 Upper Devonian lower Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Palaeotripus impariramosus


Palaeotripus monospinosus


Palaeotripus nudus E ndash T Goodbody 1986 154 Middle Silurian Sheinwoodian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago



Aitchison J C et al


er

Stat

usTy




Palaeotripus (154) continuationPalaeotripus ramalinus E ndash ndash Goto Umeda amp Ishiga 1992

163Upper Ordovician Katian Malongulli Formation Lachlan

Fold Belt 30 km NW of Taralga NSW Australia

Palaeotripus sexabrachiatus


Palaeotripus simplum E nc ndash Goto Umeda amp Ishiga 1992 165


Palaeotripus tetractinos ndash nn ndash Li 1995 333 Middle Ordovician


Palaeoumbraculum (155)Palaeoumbraculum

expandumE ndash T Amon Braun amp Ivanov 1995 8 Upper Silurian Gorstian Maksyutov Complex Tarangul


Palaeoumbraculum hidense


Palaeoxyphostylus (108)Palaeoxyphostylus

variospinaE ndash T Won 1983 156 Lower


Frankenwald Germany

Palamphimorphium (319)Palamphimorphium

speciosumndash nn Tn Ters amp Deflandre 1966 340 Ordovician ndash Phtanites drsquoAngers Vendeacutee

France

Paleocenosphaera (320)Paleocenosphaera

magnandash nd Td Nazarov 1973b 8 Lower Cambrian Stage 2 Bograd Mountains Betenevskiy

area Khakasiya Region RussiaPaleocenosphaera parva ndash nd ndash Nazarov 1973b 9 Lower Cambrian Stage 2 Bograd Mountains Betenevskiy


Paleoxiphosphaera (109)Paleoxiphosphaera

erbiensisIS ndash Ts Nazarov 1973b 9 Lower Cambrian Stage 2 Bograd Mountains Betenevskiy


Palhindeolithus (321)Palhindeolithus

ambiguusIS nn ndash Deflandre 1973a 291 Lower


Palhindeolithus ambiguus

IS ndash T Deflandre 1973b 498 Lower Carboniferous


Palhindeolithus bibiceps IS nn ndash Li 1995 333 Middle Ordovician


Palhindeolithus bitrispinosus

IS nn ndash Deflandre 1973a 290 Lower Carboniferous



IS nd ndash Deflandre 1973b 498 Lower Carboniferous


Palhindeolithus cnidus IS nn ndash Li 1995 333 Middle Ordovician


Palhindeolithus diductus IS nn ndash Deflandre 1973a 291 Lower Carboniferous


Palhindeolithus diductus IS nd ndash Deflandre 1973b 498 Lower Carboniferous


Palhindeolithus ditetrablastus

IS nn ndash Li 1995 333 Middle Ordovician


Palhindeolithus pulcher IS nn ndash Deflandre 1973a 291 Lower Carboniferous


Palhindeolithus pulcher IS nd ndash Deflandre 1973b 498 Lower Carboniferous


Papinochium (322)Papinochium dubium ndash nn Tn Ters amp Deflandre 1966 340 Ordovician ndash Phtanites drsquoAngers Vendeacutee

France





er

Stat

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Parabeothuka (237)Parabeothuka

haeckelianaS ndash ndash Won amp Iams 2015a 28 Lower Ordovician Floian Cow Head Group Western

Newfoundland CanadaParabeothuka transformis S ndash T Won amp Iams 2015a 27 Lower Ordovician Floian Cow Head Group Western

Newfoundland Canada

Paracopicyntra (245)Paracopicyntra

longispinaS ndash ndash Feng in Feng et al 2006a 25 Upper Permian Changhsingian Dalong Formation Dongpan


Paracopicyntra puncta S ndash ndash Maldonado amp Noble 2010 104 Middle Permian Capitanian Bell Canyon Formation Apache Mountains Culberson County W Texas USA

Paracopicyntra simplex S ndash ndash Feng in Feng et al 2006a 25 Upper Permian Changhsingian Dalong Formation Dongpan section Dongpan Village Guangxi Zhuang Region China

Paracopicyntra snyderi S ndash ndash Nestell amp Nestell 2010 24 Middle Permian Capitanian Bell Canyon Formation Apache Mountains Culberson County W Texas USA

Parafollicucullus (37)Parafollicucullus cornelli Al ndash ndash Kozur amp Mostler 1989 175 Middle Permian Roadian Bone Spring Formation


Parafollicucullus fusiformis

Al ndash Ts Holdsworth amp Jones 1980 285 Middle Permian Roadian Mankomen Group Clearwater Mountains USGS MR loc 0435 Alaska Range USA

Parafollicucullus sakmarensis

Al ndash Ts Kozur 1981 266 Lower Permian Sakmarian Sarabil Formation Kondurovska Orenburgskaya Region S Urals Russia

Parafollicucullus nazarovi

Al ndash ndash Kozur 1981 266 Lower Permian Sakmarian Sarabil Formation Kondurovska Orenburgskaya Region S Urals Russia

Paraholoeciscus (23)Paraholoeciscus

bingaraensisAl ndash Ts Aitchison 1993a 363 Upper Devonian Famennian Djungati terrane Bingara region

New England orogen NSW Australia

Paramphibrachium (140)Paramphibrachium woni E ndash ndash Feng in Feng et al 2009 143 Upper Permian Changhsingian Dalong Formation Dongpan


Pararcheoentactinia (45)Pararcheoentactinia

parastillaAr ndash ndash Won amp Iams 2015a 11 Lower Ordovician Floian Cow Head Group Western

Newfoundland CanadaPararcheoentactinia

reedaeAr ndash T Won amp Iams 2002 13 Upper Cambrian Stage 10 Cow Head Group Western

Newfoundland CanadaPararcheoentactinia stilla Ar ndash ndash Won Iams amp Reed 2007 505 Lower Ordovician lower-middle



cowheadensisAr ndash ndash Won Iams amp Reed 2005 438 Lower Ordovician lower-middle



impeditaAr ndash ndash Won amp Iams 2002 17 Upper Cambrian Stage 10 Cow Head Group Western

Newfoundland Canada

Pararchocyrtium (227)Pararchocyrtium mirabile N nn ndash Deflandre 1972a 3539 Lower


Pararchocyrtium mirabile N ndash T Deflandre 1972b 15 Lower Carboniferous


Parasecuicollacta (71)Parasecuicollacta bipola Ar ndash Ts Won Blodgett amp Nestor 2002

953Lower Silurian Telychian Road River Formation subsidiary

channel of Tatonduck River east-central Alaska USA

Parasecuicollacta hexactina

Ar ndash ndash Won Blodgett amp Nestor 2002 953


Parasecuicollacta multispinosa



Parasecuicollacta nannoglobosa





Aitchison J C et al


er

Stat

usTy




Parechidnina (57)Parechidnina delicata Ar ndash ndash Won Iams amp Reed 2005 443 Lower Ordovician lower-middle


Newfoundland CanadaParechidnina densa Ar ndash ndash Maletz amp Bruton 2007 276 Lower Ordovician Floian Valhallfonna Formation


Parechidnina imperfecta Ar ndash ndash Won amp Iams 2015a 28 Lower Ordovician Floian Cow Head Group Western Newfoundland Canada

Parechidnina jamesi Ar ndash ndash Won amp Iams 2002 30 Upper Cambrian Stage 10 Cow Head Group Western Newfoundland Canada

Parechidnina nevadensis Ar ndash T Kozur Mostler amp Repetski 1996 250

Lower Ordovician Tremadocian Windfall Formation Antelope Range Eureka County Nevada USA

Parechidnina variospina Ar ndash ndash Won Iams amp Reed 2005 445 Lower Ordovician lower-middle Tremadocian


ParentactiniaParentactinia vetustum E ndash T Furutani 1983 108 Lower Devonian Emsian Nakahata Formation S Mt


ParonaellaParonaella impella L ndash Ts Ormiston amp Lane 1976 169 Lower



Paronaella turgida L ndash ndash Ormiston amp Lane 1976 169 Lower Carboniferous


Paronaella triporosa L ndash ndash Holdsworth amp Murchey 1988 788

Upper Carboniferous

Bashkirian Kuna Formation Nigu Bluff Northern Brooks Range Alaska USA

Parvalanapila (72)Parvalanapila

fleischerorumAr ndash Ts MacDonald 1998 599 Lower Silurian Telychian Cape Phillips Formation


PentalastrumPentalastrum primitivum ndash nd ndash Ruumlst 1892 172 Carboniferous ndash Buumlkk-Gebirge Inner Western

Carpathians Hungary

Perforentactinia (111)Perforentactinia excepta E ndash T Jones amp Noble 2006 298 MIddle Silurian Sheinwoodian Cape Phillips Formation


PeripanartusPeripanartus deficiens ndash nd ndash Ruumlst 1892 161 Lower Devonian ndash Southern Urals Russia

PhaenicosphaeraPhaenicosphaera

mammillaS ndash ndash Sheng amp Wang 1985 176 179-

180Lower Permian Kungurian Gufeng Formation N Mt


PhaenosceniumPhaenoscenium

excentricumndash nd ndash Ruumlst 1892 179 Lower


PhorticiumPhorticium macropylium ndash nd ndash Ruumlst 1892 175 Lower Silurian ndash Cabriegraveres Heacuterault France

PlagiacanthaPlagiacantha australis ndash nd ndash Hinde 1899a 56 Middle Devonian Givetian Yarramie Formation Tamworth

Belt northern NSW AustraliaPlagiocantha nana ndash nd ndash Bykova 1955 72 Upper Devonian Frasnian Domanik Formation Tatarstan

Russia

PlagoniscusPlagoniscus colligatus ndash nd ndash Hinde 1899a 56 Middle Devonian Givetian Yarramie Formation Tamworth

Belt northern NSW AustraliaPlagoniscus cristatus ndash nd ndash Hinde 1899a 56 Middle Devonian Givetian Yarramie Formation Tamworth

Belt northern NSW AustraliaPlagoniscus simplex ndash nd ndash Hinde 1899a 56 Middle Devonian Givetian Yarramie Formation Tamworth

Belt northern NSW AustraliaPlagoniscus vetustus ndash nd ndash Hinde 1899a 57 Middle Devonian Givetian Yarramie Formation Tamworth






er

Stat

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PlegmosphaeraPlegmosphaera spiculata ndash nd ndash Aberdeen 1940 134 Upper Devonian Famennian Caballos Formation Marathon

Basin Texas USA

Plenoentactinia (112)Plenoentactinia concreta E ndash ndash Won 1997b 381 Upper Devonian Frasnian Gogo Formation Pillara Range

Canning Basin Western AustraliaPlenoentactinia

gourmelonaeE ndash ndash Won 1998 244 Lower


GermanyPlenoentactinia pinguis E ndash ndash Won 1997b 381 Upper Devonian Frasnian Gogo Formation Pillara Range


sexangulaE ndash T Won 1997b 382 Upper Devonian Frasnian Gogo Formation Pillara Range

Canning Basin Western AustraliaPlenoentactinia stella E ndash ndash Won 1997b 382 Upper Devonian Frasnian Gogo Formation Pillara Range


Plenosphaera (113)Plenosphaera gigantea E ndash ndash Park amp Won 2012 57 Lower



Plenosphaera usitata E ndash ndash Park amp Won 2012 57 Lower Carboniferous


Plenosphaera tenuitesta E ndash ndash Park amp Won 2012 58 Lower Carboniferous


Pluristratoentactinia (114)Pluristratoentactinia

conspissataE ndash T Nazarov in Nazarov et al 1981

87Upper Devonian Famennian Duksundinsk Formation along

Duksunda River Magadan Russia

Pluristratoentactinia trisphaerata

E ndash ndash Afanasieva amp Amon 2011 1509 Upper Devonian lower Famennian

Polar Urals Palnik Yu River Russia

Plussatispila (265)Plussatispila aethra S ndash ndash Jones amp Noble 2006 308 MIddle Silurian Sheinwoodian Cape Phillips Formation


Plussatispila cornwallisensis


Plussatispila delicata S ndash ndash MacDonald 2006 34 Middle Silurian Sheinwoodian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago

Plussatispila magnilimax S ndash T MacDonald 2006 33 Middle Silurian Sheinwoodian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago

Plussatispila pellicia S ndash ndash MacDonald 2006 35 Middle Silurian Sheinwoodian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago

Polyedroentactinia (115)Polyedroentactinia

cisuralicaE ndash T Kozur amp Mostler 1989 194 Lower Permian Sakmarian Sarabil Formation Sakmarskiy


Polyedroentactinia guadalupensis


Polyedroentactinia quadrata

E ndash ndash Maldonado amp Noble 2010 86 Middle Permian Capitanian Bell Canyon Formation Apache Mountains Culberson County W Texas USA

Polyentactinia (161)Polyentactinia

amplictibrosaE ndash ndash Nazarov in Isakova amp Nazarov

1986 108Upper


Urals RussiaPolyentactinia applanata E ndash ndash Nazarov amp Ormiston 1989 18 Lower Permian Artinskian Kandurov Formation Donskoye


Polyentactinia aranaeosa E ndash ndash Nazarov amp Ormiston 1984 70 Lower Permian Artinskian Kandurov Formation Donskoye Ural River Orenburgskaya Region S Urals Russia

Polyentactinia aranea E ndash ndash Gourmelon 1987 77 Lower Carboniferous




Aitchison J C et al


er

Stat

usTy




Polyentactinia (161) continuationPolyentactinia centrata E ndash ndash Nazarov amp Ormiston 1985 43 Middle Permian Roadian Bell Canyon Formation Guadalupe


Polyentactinia circumretia

E ndash ndash Nazarov amp Ormiston 1990 22 Upper Devonian Frasnian Egindinsk Formation Aitpaika River N Mugodazhar S Urals Kazakhstan


E inv ndash Nazarov amp Ormiston 1993 46 Upper Devonian Frasnian Egindinsk Formation Aitpaika River N Mugodazhar S Urals Kazakhstan


E nn ndash Nazarov 1988 pl 14 fig 1 Upper Devonian Frasnian Egindinsk Formation Aitpaika River N Mugodazhar S Urals Kazakhstan

Polyentactinia craticulata E ndash T Foreman 1963 281 Upper Devonian Famennian Huron member Ohio Shale Ohio USA

Polyentactinia djagdiensis

E ndash ndash Nazarov in Nazarov amp Shkolnik 1974 106


Polyentactinia fiscina E ndash ndash Nazarov amp Ormiston 1989 17 Lower Permian Artinskian Kandurov Formation Donskoye Ural River Orenburgskaya Region S Urals Russia

Polyentactinia fragilis E ndash ndash Nazarov amp Ormiston 1989 17 Lower Permian Artinskian Kandurov Formation Donskoye Ural River Orenburgskaya Region S Urals Russia

Polyentactinia hunanensis

E ndash ndash Dong Knoll amp Lipps 1997 754 Upper Cambrian Paibian Bitiao Formation Paibi Village Huayuan County Hunan China

Polyentactinia invenusta E ndash ndash Aitchison 1993b 117 Upper Devonian lower Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Polyentactinia ishigai E ndash ndash Kozur amp Mostler 1989 195 Lower Permian Sakmarian Sarabil Formation Sakmarskiy Orenburgskaya Region S Urals Russia

Polyentactinia kossistekensis


Polyentactinia lautitia E ndash ndash Nazarov amp Ormiston 1985 43 Lower Permian Artinskian Aktastyi River Orenburgskaya region Southern Urals Russia

Polyentactinia leptosphaera


Polyentactinia miopora E ndash ndash Wang in Wang et al 2000 248 Middle Devonian Givetian Changyucun Formation Shaijingpo section Xianyun County Yunnan China

Polyentactinia multifida E ndash ndash Nazarov in Isakova amp Nazarov 1986 106

Upper Carboniferous


Polyentactinia multifora E ndash ndash Nazarov in Isakova amp Nazarov 1986 107

Upper Carboniferous


Polyentactinia njatvica E ndash ndash Nazarov amp Ormiston 1990 23 Upper Devonian Famenian Zapadno Valavskaya borehole 1-R Pripyat Depression Belarus

Polyentactinia nyatvica E inv ndash Nazarov amp Ormiston 1993 48 Upper Carboniferous


Polyentactinia nyatvica E nn ndash Nazarov 1988 139 Upper Carboniferous


Polyentactinia octupla E ndash ndash Nazarov in Isakova amp Nazarov 1986 105

Upper Carboniferous


Polyentactinia offerta E ndash ndash Nazarov in Nazarov amp Popov 1980 42

Middle Ordovician


Polyentactinia offirmata E ndash ndash Nazarov amp Ormiston 1989 18 Lower Permian Artinskian Kandurov Formation Donskoye Ural River Orenburgskaya Region S Urals Russia

Polyentactinia polygonia E ndash ndash Foreman 1963 281 Upper Devonian Famennian Huron member Ohio Shale Ohio USA

Polyentactinia propinqua E ndash ndash Nazarov 1975 79 Upper Devonian Frasnian Egindinsk Formation Aitpaika River N Mugodazhar S Urals Kazakhstan

Polyentactinia rudihispida E ndash ndash Nazarov amp Ormiston 1993 48 Upper Devonian Famenian Zapadno Valavskaya borehole 1-R Pripyat Depression Belarus

Polyentactinia rudihispida E nn ndash Nazarov 1988 132 Upper Devonian Famenian Zapadno Valavskaya borehole 1-R Pripyat Depression Belarus

Polyentactinia sexangulata

E ndash ndash Feng Mei amp Zhang 1997 89 Lower Carboniferous

Visean Changning-Menglian Zone SE of Gengma Yunnan China





er

Stat

usTy




Polyentactinia (161) continuationPolyentactinia spinulenta E ndash ndash Pouille amp Danelian

in Pouille et al 2014 156Middle

Ordovicianupper

DarriwilianShundy Formation Balkhash

region KazakhstanPolyentactinia tenera E ndash ndash Aitchison 1993b 117 Upper Devonian lower Frasnian Gogo Formation Pillara Range


Polyentactinia tortispina E ndash ndash Ormiston amp Lane 1976 166 Lower Carboniferous


Polyentactinia zhamoidai E nn ndash Afanasieva 1997a 221 Upper Devonian middle Frasnian


Polyentactinia zhamoidai E ndash ndash Afanasieva 2000a 38 Upper Devonian middle Frasnian


Polyentactinia coldinensis


Polyentactinia estonica E ndash ndash Nazarov in Nazarov amp Nolvak 1983 5


Polyentactinia helioformis


Polyentactinia leeorum E ndash ndash Goodbody 1986 156 Lower Silurian Telychian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago

Polyentactinia nazarovi E ndash ndash Kozur amp Mostler 1989 195 Lower Permian Sakmarian Sarabil Formation Sakmarskiy Orenburgskaya Region S Urals Russia

Polyentactinia perampla E ndash ndash Braun Maass amp Schmidt-Effing 1992 172

Upper Devonian Famennian Breuschtal northern Vosges Alsace France

Polyentactinia plecta E ndash ndash Foreman 1963 282 Upper Devonian Famennian Huron member Ohio Shale Ohio USA

Polyentactinia stelcki E ndash ndash Goodbody 1986 156 Middle Silurian Sheinwoodian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago

Polyentactinia udiensis E ndash ndash Nazarov in Nazarov amp Shkolnik 1974 107


Polyfistula (195)Polyfistula hexalobata L ndash ndash Nazarov amp Ormiston 1989 13 Lower Permian Artinskian Havallah Formation Cedar Mts

Lander County Nevada USAPolyfistula Iongiquitas L ndash T Nazarov amp Ormiston 1984 80 Lower Permian Artinskian Kandurov Formation Donskoye


Polyfistula novem L ndash ndash Noble amp Renne 1990 388 Middle Permian Roadian Dekkas Formation Eastern Klamath Mountains California USA

Polyfistula regularis L ndash ndash Feng in Feng et al 2006b 846 Upper Permian Changhsingian Dalong Formation Dongpan section Dongpan Village Guangxi Zhuang Region China

Polyfistula grantmackiei L ndash ndash Sashida in Sashida et al 2000a 85

Upper Permian Changhsingian Fang Chert Quarry of Ban Huai Tian Tang Chiang Mai Thailand

Popofskyellum (231)Popofskyellum

annulatumN ndash ndash Deflandre 1964 3056 Lower


Popofskyellum aspinosum



Popofskyellum bispina N ndash ndash Park amp Won 2012 60 Lower Carboniferous


Popofskyellum campanella



Popofskyellum conicum N ndash ndash Deflandre 1964 3056 Lower Carboniferous


Popofskyellum crinerensis




Popofskyellum daisyensis N ndash ndash Cheng 1986 153 Upper Devonian Famennian Woodford Formation TI Valley section Pittsburg County Oklahoma USA



Aitchison J C et al


er

Stat

usTy




Popofskyellum (231) continuationPopofskyellum deflandrei N ndash ndash Cheng 1986 154 Upper Devonian Famennian Woodford Formation TI Valley


Popofskyellum delicatum N ndash ndash Cheng 1986 154 Upper Devonian Famennian Woodford Formation TI Valley section Pittsburg County Oklahoma USA

Popofskyellum dumitricai N ndash ndash Cheng 1986 155 Lower Carboniferous


Popofskyellum elmense N ndash ndash Cheng 1986 156 Upper Devonian Famennian Woodford Formation TI Valley section Pittsburg County Oklahoma USA

Popofskyellum hendricksi N ndash ndash Cheng 1986 156 Upper Devonian Famennian Woodford Formation TI Valley section Pittsburg County Oklahoma USA

Popofskyellum obesum N ndash ndash Cheng 1986 157 Lower Carboniferous


Popofskyellum pulchrum N nn T Deflandre 1960 216 Lower Carboniferous


Popofskyellum pulchrum N nn ndash Deflandre 1964 3056 Lower Carboniferous


Popofskyellum robustum N ndash ndash Cheng 1986 157 Upper Devonian Famennian Woodford Formation TI Valley section Pittsburg County Oklahoma USA

Popofskyellum tardicarboniferum

N ndash ndash Nazarov in Isakova amp Nazarov 1986 118

Upper Carboniferous


Popofskyellum triportium N ndash ndash Won 1990 138 Lower Carboniferous


Popofskyellum turpiculum


Popofskyellum undulatum

N ndash ndash Deflandre 1964 3056 Lower Carboniferous


Popofskyellum wonae N ndash ndash Cheng 1986 159 Upper Devonian Famennian Woodford Formation TI Valley section Pittsburg County Oklahoma USA

Popofskyellum procerulum

N ndash T Nazarov in Nazarov amp Popov 1980 68

Middle Ordovician


PorodiscusPorodiscus brevis ndash nd ndash Hinde amp Fox 1895 640 Carboniferous Serpukhovian-


UKPorodiscus cabrierensis ndash nd ndash Ruumlst 1892 168 Lower Silurian ndash Cabriegraveres Heacuterault FrancePorodiscus clathratus ndash nd ndash Hinde amp Fox 1895 640 Carboniferous Serpukhovian-


UKPorodiscus intricatus ndash nd ndash Ruumlst 1892 168 Lower Devonian ndash Southern Urals RussiaPorodiscus lens ndash nd ndash Hinde amp Fox 1895 640 Carboniferous Serpukhovian-


UKPorodiscus percinctus ndash nd ndash Hinde amp Fox 1895 640 Carboniferous Serpukhovian-


UKPorodiscus rossicus ndash nd ndash Ruumlst 1892 168 Lower Devonian ndash Southern Urals Russia

Praedeflandrella (175)Praedeflandrella firmata L ndash ndash Maldonado amp Noble 2010 88 Middle Permian Capitanian Bell Canyon Formation Apache


Praedeflandrella prolata L ndash ndash Maldonado amp Noble 2010 88 Middle Permian Capitanian Bell Canyon Formation Apache Mountains Culberson County W Texas USA

Praesaturnalis (156)Praesaturnalis

octoaciculandash nn ndash Li 1991 75 Upper Silurian Gorstian Mayila Complex Keerhada S

Mayila Mt Xinjiang ChinaPraesaturnalis

octoaculeatusE ndash ndash Li 1994 262 Lower Devonian Emsian Keerhada Mayila Mts Xinjiang

ChinaPraesaturnalis

septemistellandash nn ndash Li 1991 75 Upper Silurian Gorstian Mayila Complex Keerhada S


septemistellatusE ndash Ts Li 1994 262 Upper Silurian Ludfordian Keerhada Mayila Mts Xinjiang

China





er

Stat

usTy




Praespongocoelia (301)Praespongocoelia

fusiformaIS

(OampF)ndash ndash Noble 1994 38 Upper Silurian Gorstian Caballos Novaculite Formation


Praespongocoelia robusta

IS(OampF)

ndash ndash Umeda 1997 423 Lower Devonian Lochkovian Jyoru Formation Kurosegawa Belt Mt Konomori Kochi City Shikoku Japan

Primaritripus (323)Primaritripus

buribayensisS nd ndash Afanasieva amp Amon 2008 465 Lower Devonian Emsian Kamennaya Gora Tanalyk River

Southern Ural RussiaPrimaritripus chuvashovi S nd ndash Afanasieva amp Amon 2008 465 Upper Devonian lower Frasnian Argagan Mountains

Sultantimirovo southern Urals Russia

Primaritripus kariukmasensis

S nd ndash Afanasieva amp Amon 2009a 45 Upper Devonian lower Frasnian Argagan Mountains Sultantimirovo southern Urals Russia

PriscilatentidiotaPriscilatentidiota kozurii ndash nn ndash Li 1995 333 Middle



PrismatiumPrismatium paradictyum ndash nd ndash Ruumlst 1892 178 Upper Permian ndash Sosio Valley Area Western Sicily

Italy

Procyrtis (157)Procyrtis qinglai E ndash ndash Li 1995 337 Middle



Procyrtis rustii E ndash T Li 1995 336 Middle Ordovician


Proholoeciscus (24)Proholoeciscus deweveri Al ndash ndash Li 1995 339 Middle



Proholoeciscus foremanii Al ndash Ts Li 1995 339 Middle Ordovician


Proholoeciscus holdsworthi

Al ndash ndash Li 1995 339 Middle Ordovician


Protoalbaillella (6)Protoalbaillella

anaiwanensisAl ndash ndash Aitchison 1993a 361 Lower

CarboniferousTournaisian Gundahl Complex Anaiwan

terrane Jackadgery NSW Australia

Protoalbaillella deflandrei Al ndash T Cheng 1986 68 Lower Carboniferous


Protoalbaillella formosa Al ndash ndash Cheng 1986 68 Upper Devonian Famennian Woodford Formation and Johns Valley Shale Ouachita Mtns Oklahoma USA

Protoalbaillella oilensis Al ndash ndash Schwartzapfel amp Holdsworth 1996 91

Lower Carboniferous


Protoalbaillella pinetopensis


Protoalbaillella talihinaensis



Protoalbaillella triangulata Al ndash ndash Won 1990 138 Lower Carboniferous


Protoalbaillella unispinosa



Protoalbaillella aprathensis



Protobiramus (302)Protobiramus biaxialus S ndash T Won in Won amp Below 1999 360 Middle Cambrian ndash Inca Formation Georgina Basin




Aitchison J C et al


er

Stat

usTy




Protoceratoikiscum (25)Protoceratoikiscum

arachnoidesAl nc ndash Goto Umeda amp Ishiga 1992



Protoceratoikiscum chinocrystallum

Al ndash T Goto Umeda amp Ishiga 1992 166


Protoceratoikiscum clarksoni

Al ndash ndash Danelian amp Floyd 2001 495 Lower Ordovician Floian Lower Crawford track Southern Uplands Scotland UK

Protoceratoikiscum crossingi

Al ndash ndash Noble amp Webby 2009 553 Upper Ordovician Katian Malongulli Formation Cliefden Caves area NSW Australia

Protoceratoikiscum similistellatum



Protoentactinia (63)Protoentactinia bifurcata Ar ndash ndash Maletz amp Bruton 2007 262 Lower Ordovician Floian Valhallfonna Formation


Protoentactinia deformis Ar ndash ndash Won Iams amp Reed 2005 455 Lower Ordovician lower-middle Tremadocian


Protoentactinia gracilispinosa

Ar ndash Ts Kozur Mostler amp Repetski 1996 253


Protoentactinia kozuriana Ar ndash ndash Won Iams amp Reed 2005 455 Lower Ordovician lower-middle Tremadocian


Protoentactinia latospinosa

Ar ndash ndash Kozur Mostler amp Repetski 1996 254


Protoentactinia minuta Ar ndash ndash Kozur Mostler amp Repetski 1996 254


Protoentactinia primigena

Ar ndash ndash Won Iams amp Reed 2005 456 Lower Ordovician lower-middle Tremadocian


Protoentactinia transformis



Protoholoeciscus (26)Protoholoeciscus hindea Al ndash T Aitchison 1993a 362 Upper Devonian Famennian Djungati terrane Upper Barnard

Valley New England orogen NSW Australia

Protoholoeciscus ochiensis


Protoholoeciscus spinosus


Protoproventocitum (282)Protoproventocitum

aitchisoniIS ndash T Won Iams amp Reed 2007 536 Lower Ordovician lower-middle


Newfoundland CanadaProtoproventocitum

nazaroviiIS ndash ndash Won Iams amp Reed 2007 537 Lower Ordovician lower-middle



floianumIS ndash ndash Won amp Iams 2015a 22 Lower Ordovician Floian Cow Head Group Western

Newfoundland Canada

Protosegmentum (324)Protosegmentum

xinjiangensisndash nd Td Wang in Wang et al 2008 397 Middle



Protospongentactinia (64)Protospongentactinia

spongiosa Ar ndash T Won Iams amp Reed 2007 532 Lower Ordovician lower-middle


Newfoundland Canada

Proventocitum (283)Proventocitum

cylindricumIS ndash ndash Won amp Iams 2011 173 Lower Ordovician Floian Cow Head Group Western

Newfoundland CanadaProventocitum

holdsworthiIS ndash ndash Aitchison 1998 79 Lower Ordovician Arenig Ballantrae Complex Bennane Lea

SW Scotland UKProventocitum piriforme IS ndash ndash Won amp Iams 2011 173 Lower Ordovician Floian Cow Head Group Western

Newfoundland Canada





er

Stat

usTy




Provisocyntra (116)Provisocyntra amplissima E ndash ndash Nazarov amp Ormiston 1987 72 Upper

CarboniferousMoscovian Nyatvinsk Formation Popovka

River Prikolyma Region Magadan Region NE Russia

Provisocyntra cassicula E ndash T Nazarov amp Ormiston 1987 72 Lower Carboniferous

Tournaisian Lodgepole Formation Amoco Daniel Anderson No 1 well Rolette County North Dakota USA

Provisocyntra conferta E ndash ndash Won 1997b 383 Upper Devonian Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Provisocyntra densa E ndash ndash Feng in Feng et al 2007 34 Upper Permian Changhsingian Dalong Formation Dongpan section Dongpan Village Guangxi Zhuang Region China

Provisocyntra ormistoni E ndash ndash Feng in Feng et al 2007 34 Upper Permian Changhsingian Dalong Formation Dongpan section Dongpan Village Guangxi Zhuang Region China

Provisocyntra pskemensis E ndash ndash Nazarov amp Ormiston 1987 73 Lower Carboniferous

Serpukhovian Pskemskiy Khrebet Pskem Range Tashkent Uzbekistan

Provisocyntra tenuitomenta



PrunulumPrunulum armeniacum ndash nd ndash Ruumlst 1892 157 Upper Devonian ndash SchaebenholzHarz Mountains

Elbingerode GermanyPrunulum murrayi ndash nd ndash Ruumlst 1892 157 Lower


Pseudoalbaillella (39)Pseudoalbaillella

(Pseudoalbaillella) eurasiatica

Al ndash ndash Kozur amp Mostler 1989 177 Lower Permian Kungurian Ashimi-dani section Ukyo-ku Kyoto Prefecture Japan

Pseudoalbaillella aidensis Al ndash ndash Nishimura amp Ishiga 1987 174 Middle Permian Wordian Maizuru Belt Mimasaka Chugoku District Japan

Pseudoalbaillella annulata

Al ndash ndash Ishiga in Ishiga et al 1984 48 Lower Permian Asselian Funafuseyama Unit Tamba-Mino-Ashio Belt Ohmori near Shuzan Kinki Prefecture Japan

Pseudoalbaillella apachensis

Al ndash ndash Nestell amp Nestell 2010 18 Middle Permian Capitanian Bell Canyon Formation Apache Mountains Culberson County W Texas USA

Pseudoalbaillella banchengensis

Al ndash ndash Xian amp Zhang 1998 194 Lower Permian Kungurian Bancheng Formation SE Guangxi China

Pseudoalbaillella bella Al ndash ndash Sheng amp Wang 1985 174 178-179

Lower Permian Kungurian Gufeng Formation N Mt Konghshan Hushan Nanjing China

Pseudoalbaillella boonevillensis

Al ndash ndash Nestell Pope amp Nestell 2012 232

Upper Carboniferous


Pseudoalbaillella bulbosa Al ndash ndash Ishiga 1982 335 Upper Carboniferous

Gzhelian Funafuseyama Unit Tamba-Mino-Ashio Belt Loc 1 Kurookagawa River Sasayama City Japan

Pseudoalbaillella chilensis

Al ndash ndash Ling amp Forsythe 1987 257 Upper Carboniferous

Gzhelian Denaro Complex Magallanes y Antartica Chilena Chile

Pseudoalbaillella cona Al ndash ndash Cornell amp Simpson 1985 276 Middle Permian Roadian Bone Spring Formation Guadalupe Mts Culbertson County W Texas USA

Pseudoalbaillella convexa Al ndash ndash Rudenko in Rudenko amp Panasenko 1990b 184


Pseudoalbaillella corniculata

Al ndash ndash Rudenko in Rudenko amp Panasenko 1990b 185


Pseudoalbaillella decurvata


Upper Carboniferous


Pseudoalbaillella delawarensis

Al ndash ndash Maldonado amp Noble 2010 82 Middle Permian Capitanian Bell Canyon Formation Apache Mountains Culberson County W Texas USA

Pseudoalbaillella desmoinesiensis


Upper Carboniferous




Aitchison J C et al


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Stat

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Pseudoalbaillella (39) continuationPseudoalbaillella elegans Al ndash ndash Ishiga amp Imoto 1980 337 Lower Permian Kungurian Funafuseyama Unit Tamba-Mino-

Ashio Belt Sasayama area Chugoku District Japan

Pseudoalbaillella elongata

Al ndash Ts Ishiga amp Imoto 1980 339 Lower Permian Kungurian Funafuseyama Unit Tamba-Mino-Ashio Belt Sasayama area Chugoku District Japan

Pseudoalbaillella excella Al ndash ndash Nestell Pope amp Nestell 2012 233

Upper Carboniferous


Pseudoalbaillella globosa Al ndash ndash Ishiga amp Imoto in Ishiga et al 1982b 275


Pseudoalbaillella internata

Al ndash ndash Wang in Wang et al 2012 108 Middle Permian Roadian Bancheng Formation SE Guangxi China

Pseudoalbaillella ishigai Al ndash ndash Wang in Wang et al 1994 181 Lower Permian Kungurian Dachongling section Qinzhou City Guangxi S China

Pseudoalbaillella lanceolata

Al ndash Ts Ishiga amp Imoto in Ishiga et al 1982b 275


Pseudoalbaillella lomentaria

Al ndash ndash Ishiga amp Imoto 1980 338 Lower Permian Kungurian Funafuseyama Unit Tamba-Mino-Ashio Belt Sasayama area Chugoku District Japan

Pseudoalbaillella longtanensis

Al ndash ndash Sheng amp Wang 1985 174 179 Lower Permian Kungurian Gufeng Formation N Mt Konghshan Hushan Nanjing China

Pseudoalbaillella longuscornis


Pseudoalbaillella nanjingensis

Al ndash ndash Sheng amp Wang 1985 174-175 179


Pseudoalbaillella nodosa Al ndash ndash Ishiga 1982 334 Upper Carboniferous


Pseudoalbaillella ornata Al ndash ndash Ishiga amp Imoto 1980 339 Lower Permian Kungurian Funafuseyama Unit Tamba-Mino-Ashio Belt Sasayama area Chugoku District Japan

Pseudoalbaillella papaiensis

Al nn ndash Feng 1992 55 Upper Permian Changhsingian Papai Formation of Changning-Menglian Belt Papai Village Cangyuan County Yunnan China

Pseudoalbaillella rhombothoracata


Pseudoalbaillella scalprata

Al ndash T Holdsworth amp Jones 1980 285 Lower Permian Artinskian Havallah Formation USGS MR 0203 Nevada USA

Pseudoalbaillella scalprata postscalprata

Al ndash ndash Ishiga 1983 3 Lower Permian Kungurian Fujioka-oku in the Fujiokagawa Valley Sasayama City Hyogo Prefecture SW Japan

Pseudoalbaillella scalprata praescalprata

Al ndash ndash Catalano Di Stefano amp Kozur 1989 92

Lower Permian Kungurian Fujioka-oku in the Fujiokagawa Valley Sasayama City Hyogo Prefecture SW Japan

Pseudoalbaillella scalprata scalprata


Pseudoalbaillella simplex Al ndash ndash Ishiga amp Imoto 1980 337 Lower Permian Kungurian Funafuseyama Unit Tamba-Mino-Ashio Belt Sasayama area Chugoku District Japan

Pseudoalbaillella triangularis

Al ndash ndash Wang in Wang et al 2012 108 Lower Permian Artinskian Bancheng Formation SE Guangxi China

Pseudoalbaillella u-forma Al nc Ts Holdsworth amp Jones 1980 285 Lower Permian Artinskian Big Delta quadrangle USGS MR loc 0349 east-central Alaska USA

Pseudoalbaillella yanaharensis

Al ndash ndash Nishimura amp Ishiga 1987 173 Middle Permian Wordian Maizuru Belt Mimasaka Chugoku District Japan

Pseudolithelius (202)Pseudolithelius permicus L ndash T Kozur amp Mostler 1989 186 Lower Permian Artinskian Koshelev Formation Alogazovo






er

Stat

usTy




Pseudorotasphaera (73)Pseudorotasphaera

communaAr ndash ndash Noble 1994 26 Middle Silurian Homerian Caballos Novaculite Formation


Pseudorotasphaera hispida

Ar ndash T Noble 1994 25 Upper Silurian Gorstian Caballos Novaculite Formation Payne Hills Bourland Mts Marathon Basin W Texas USA

Pseudorotasphaera Ianceolata

Ar ndash ndash Noble 1994 26 Upper Silurian Gorstian Caballos Novaculite Formation Beckwith Hills Marathon Basin W Texas USA

Pseudorotasphaera robustispina

Ar ndash ndash Noble 1994 27 Upper Silurian Gorstian Caballos Novaculite Formation Payne Hills Bourland Mts Marathon Basin W Texas USA

Pseudorotasphaera rotunda


Pseudospongoprunum (273)Pseudospongoprunum

parvispinaS ndash ndash Tetard Noble amp Danelian in



Pseudospongoprunum prototypum

S ndash ndash Umeda 1998b 207 Upper Silurian Pridoli Kurosegawa Belt Shikoku Japan

Pseudospongoprunum sagittatum

S ndash ndash Wakamatsu Sugiyama amp Furutani 1990 173

Lower Devonian Lochkovian G4 formation of Gion-yama Group Mt Gionyama area Kyushu Japan

Pseudospongoprunum tazukawaensis

S ndash T Wakamatsu Sugiyama amp Furutani 1990 173

Upper Silurian Gorstian SE Suberidani along Tazukawa River Katsuura Town Shikoku Japan

Pseudospongoprunum chiangdaoensis

S ndash ndash Sashida in Sashida et al 2000c 805


Pseudospongoprunum fontainei

S ndash ndash Sashida in Sashida et al 2000b 255


Pseudospongoprunum tauversi

S ndash ndash Noble 1994 28 Upper Silurian Gorstian Caballos Novaculite Formation Monument Ck Bourland Mts Marathon Basin W Texas USA

Pseudotormentus (176)Pseudotormentus

delawarensisL ndash ndash Schwartzapfel amp Holdsworth

1996 222Lower


Sand Branch Section Johnston County Oklahoma USA

Pseudotormentus kamigoriensis

L ndash T De Wever amp Caridroit 1984 101 Middle Permian Capitanian Oi Formation Ultra-Tamba Belt Yanogawa River Yamasakicho-Kusune Shisou City Japan

Pseudotormentus monoporus

L ndash ndash Wang in Wang et al 2012 110 Upper Permian upper Changhsingian


Pterocanium tenuecostatum

ndash nd ndash Ruumlst 1892 183 Upper Permian ndash Sosio Valley Area Western Sicily Italy

PygacapsaPygacapsa bilaminata nr ndash Tnr Wang 1989 145 Lower Cambrian ndash Niutitang Formation Sandu


Pylentonema (160)Pylentonema antiqua E ndash ndash Deflandre 1963 3982 Lower


Pylentonema aperta E ndash Ts Nazarov in Nazarov et al 1975 102

Middle Ordovician


Pylentonema aperta ndash inv ndash Nazarov in Nazarov et al 1977 918

Middle Ordovician


Pylentonema banchengensis

E ndash ndash Wang amp Kuang 1993 279 Lower Carboniferous

Tournaisian Shijia Formation Shijia Resevoir Qinzhou area Guangxi Zhuang Region SW China

Pylentonema eucosmeta E ndash ndash Braun 1989 91 Lower Carboniferous

upper Tournaisian


Pylentonema hindei E ndash ndash Cheng 1986 117 Lower Carboniferous




Aitchison J C et al


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Pylentonema (160) continuationPylentonema insueta E ndash Ts Nazarov 1975 97 Middle


River Chingiz Range KazakhstanPylentonema insueta E ndash ndash Nazarov in Nazarov et al 1977

918Middle



Pylentonema mira E ndash ndash Cheng 1986 118 Upper Devonian Famennian Woodford Formation TI Valley section Pittsburg County Oklahoma USA

Pylentonema racheboeufi E ndash ndash Gourmelon 1986 189 Lower Carboniferous


Pylentonema rimata E ndash ndash Nazarov in Nazarov amp Popov 1980 62

Middle Ordovician


Pylentonema robusta E ndash ndash Schwartzapfel amp Holdsworth 1996 154



Pylentonema transitorum E ndash ndash Liu amp Hao 2006 649 Upper Devonian Famennian Tielimaitidaban section SW Tianshan Xinjiang China

Pylentonema triangulata E ndash ndash Cheng 1986 119 Upper Devonian Famennian Woodford Formation TI Valley section Pittsburg County Oklahoma USA

Pylentonema typica E ndash ndash Cheng 1986 119 Upper Devonian Famennian Woodford Formation TI Valley section Pittsburg County Oklahoma USA

Pylentonema parviaperta



Pylentonema serrensis E ndash ndash Gourmelon 1987 107 Lower Carboniferous


Quadrapesus (162)Quadrapesus araneae E ndash ndash Schwartzapfel amp Holdsworth



Carter County Oklahoma USA Quadrapesus bakeri E ndash ndash Schwartzapfel amp Holdsworth


Famennian Woodford Formation Criner Hills

2A section Carter County Oklahoma USA

Quadrapesus dumitricai E ndash T Cheng 1986 121 Upper Devonian Famennian Woodford Formation TI Valley section Pittsburg County Oklahoma USA

Quadrapesus transitivus E ndash ndash Liu amp Hao 2006 649 Upper Devonian Famennian Tielimaitidaban section SW Tianshan Xinjiang China

Quadrapesus unicus E ndash ndash Cheng 1986 122 Upper Devonian Famennian Woodford Formation TI Valley section Pittsburg County Oklahoma USA

Quadrapesus conili E ndash ndash Schwartzapfel amp Holdsworth 1996 158



Quadrapesus jimisonae E ndash ndash Schwartzapfel amp Holdsworth 1996 159



Quadrapesus sixi E ndash ndash Schwartzapfel amp Holdsworth 1996 157



Quadratapora (344)Quadratapora

zhenbaensisnr is Tnr Hao amp Shu 1987 309 Lower Cambrian Terreneuvian Shuijingtuo Formation Shaanxi


Quadricaulis (196)Quadricaulis femoris L ndash Ts Caridroit amp De Wever 1986 79 Upper Permian Wuchiapingian Oi Formation Ultra-Tamba Belt


Quadrilobata (205)Quadrilobata

ephippiomorphaL ndash T Wang 1995 144 Middle Permian Capitanian Gufeng Formation Jiangsu China

Quadrilobata blomei L ndash ndash Nestell amp Nestell 2010 50 Middle Permian Capitanian Bell Canyon Formation Apache Mountains Culberson County W Texas USA

Quadriremis (197)Quadriremis flata L ndash ndash Wang 1993a 451 Lower Permian Kungurian Gufeng Formation Qiaotou Village

Chaou City Anhui ChinaQuadriremis gliptoacus L ndash T Nazarov amp Ormiston 1985 36 Lower Permian Artinskian Kandurov Formation Donskoye






er

Stat

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Quadriremis (197) continuationQuadriremis minima L ndash ndash Nazarov amp Ormiston 1985 36 Middle Permian Roadian Bell Canyon Formation Guadalupe


Quadriremis nevadensis L ndash ndash Nazarov amp Ormiston 1989 11 Middle Permian Capitanian Willow Canyon Formation Toquima Range Mill Canyon Nevada USA


Quasibeothuka (274)Quasibeothuka bifurcata S ndash ndash Wang in Wang et al 2008 400 Middle


Kuruktag region Xinjiang ChinaQuasibeothuka

ellipsoidalaS ndash ndash Wang in Wang et al 2008 400 Middle


Kuruktag region Xinjiang ChinaQuasibeothuka fusiforma S ndash Td Wang in Wang et al 2008 399 Middle



longifusiformaS ndash ndash Wang in Wang et al 2008 400 Middle


Kuruktag region Xinjiang ChinaQuasibeothuka ovata S ndash ndash Wang in Wang et al 2008 399 Middle



Quinqueremis (198)Quinqueremis arundinea L ndash ndash Nazarov amp Ormiston 1983b 375Lower Permian Artinskian Kandurov Formation Donskoye


Quinqueremis arundinea L ndash T Nazarov amp Ormiston 1985 37 Lower Permian Artinskian Kandurov Formation Donskoye Ural River Orenburgskaya Region S Urals Russia

Quinqueremis clathrolobulatus

L ndash ndash Amon amp Braun 1994 6 Lower Permian Artinskian Burtsevesky horizon Dalny Tulkas Rill Bashkortostan Region S Urals Russia

Quinqueremis robusta L ndash ndash Nazarov amp Ormiston 1985 37 Middle Permian Capitanian US Highway 180 Culbertson County Texas USA

Quinqueremis yunnanensis


Radiobisphaera (117)Radiobisphaera acuta E ndash ndash Won 1997a 351 Upper Devonian Frasnian Gogo Formation Pillara Range

Canning Basin Western AustraliaRadiobisphaera flamans E ndash ndash Won 1997a 351 Upper Devonian Frasnian Gogo Formation Pillara Range

Canning Basin Western AustraliaRadiobisphaera

magnifenestraE ndash ndash Won 1997a 352 Upper Devonian Frasnian Gogo Formation Pillara Range

Canning Basin Western AustraliaRadiobisphaera menneri E ndash ndash Afanasieva 2000a 60 Upper Devonian middle



Radiobisphaera multa E ndash ndash Won 1997a 353 Upper Devonian Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Radiobisphaera multiaculeata


Radiobisphaera multiaculeata multiaculeata


Radiobisphaera nazaroviana


Radiobisphaera rozanovi E ndash ndash Afanasieva amp Amon 2009b 495 Upper Devonian middle Frasnian

Kamenevskaya Formation Goryunovo Zolotukha River Goryunovo Rudnyi Altai Russia

Radiobisphaera variantia E ndash T Won 1997a 356 Upper Devonian Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Radiobisphaera variantia inaequalis


Radiobisphaera variantia variantia




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Ramuspiculum (67)Ramuspiculum curvatum Ar ndash ndash Won amp Iams 2002 29 Upper Cambrian Stage 10 Cow Head Group Western

Newfoundland CanadaRamuspiculum

multiramosumAr ndash T Won amp Iams 2002 29 Upper Cambrian Stage 10 Cow Head Group Western


trifurcatumAr ndash ndash Won amp Iams 2002 29 Upper Cambrian Stage 10 Cow Head Group Western

Newfoundland Canada

Raphidociclicus (27)Raphidociclicus gemellus Al ndash ndash Nazarov amp Rudenko 1981 136 Lower Permian Artinskian Aktastyi River Orenburgskaya

region Southern Urals RussiaRaphidociclicus gemellus

americanusAl ndash ndash Nazarov amp Ormiston 1985 52 Middle Permian Capitanian Bell Canyon Formation Guadalupe


Raphidociclicus hiulcus Al ndash T Nazarov amp Rudenko 1981 135 Lower Permian Sakmarian Kandurov Formation Donskoye Ural River Orenburgskaya Region S Urals Russia

Raphidociclicus scutum Al ndash ndash Maldonado amp Noble 2010 80 Middle Permian Capitanian Bell Canyon Formation Apache Mountains Culberson County W Texas USA

Rectotormentum (213)Rectotormentum fengi L ndash ndash Noble amp Jin 2010 143 Middle Permian Capitanian Bell Canyon Formation Guadalupe


Rectotormentum fornicatum

L ndash T Nazarov amp Ormiston 1985 41 Lower Permian Artinskian Kandurov Formation Donskoye Ural River Orenburgskaya Region S Urals Russia

Rectotormentum wardlawi

L ndash ndash Nestell amp Nestell 2010 42 Middle Permian Capitanian Bell Canyon Formation Apache Mountains Culberson County W Texas USA

Retentactinia (118)Retentactinia aspera E ndash ndash Won 1997b 384 Upper Devonian Frasnian Gogo Formation Pillara Range

Canning Basin Western AustraliaRetentactinia aspinosa E ndash ndash Won 1997b 384 Upper Devonian Frasnian Gogo Formation Pillara Range

Canning Basin Western AustraliaRetentactinia delicata E ndash ndash Won 1997b 384 Upper Devonian Frasnian Gogo Formation Pillara Range

Canning Basin Western AustraliaRetentactinia dellensus E ndash ndash Park amp Won 2012 58 Lower



Retentactinia impedita E ndash T Th

Won 1997b 385 Upper Devonian Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Retentactinia interreticulata


Retentactinia kelleri E ndash ndash Afanasieva 2000a 76 Upper Devonian middle Frasnian


Retentactinia levis E ndash ndash Won 1997b 386 Upper Devonian Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Retentactinia longa E ndash ndash Won 1997b 386 Upper Devonian Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Retentactinia longa densa


Retentactinia longa longa E ndash ndash Won 1997b 386 Upper Devonian Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Retentactinia longa pilosa


Retentactinia repleta E ndash T Won 1997b 390 Upper Devonian Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Retentactinia spongiosa E ndash ndash Seo amp Won 2009 72 Lower Carboniferous


Retientactinosphaera (303)Retientactinosphaera

clavataIS

(OampF)ndash ndash Afanasieva 2011 128 Upper Devonian middle



Retientactinosphaera magnifica

IS(OampF)

ndash T Afanasieva 2011 127 Upper Devonian lower Famennian

Zadonsk Formation W Lekkeyaginsk-65 Saremboi-Lekkeyaga Timan-Pechora Basin Russia





er

Stat

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Retisphaera (119)Retisphaera distratalis E ndash ndash Won 1997b 396 Upper Devonian Frasnian Gogo Formation Pillara Range


Retisphaera ovata E ndash ndash Won 1997b 398 Upper Devonian Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Retisphaera rota E ndash ndash Won 1997b 398 Upper Devonian Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

RhizoplegmaRhizoplegma betulinum ndash nd ndash Ruumlst 1892 150 Upper Permian ndash Sosio Valley Area Western Sicily

Italy

RhodosphaeraRhodosphaera crucifera E nd ndash Ruumlst 1892 137 Lower


Rhodosphaera devoniensis

E nd ndash Ruumlst 1892 137 Upper Devonian ndash SchaebenholzHarz Mountains Elbingerode Germany

Rhodosphaera idonea E nd ndash Ruumlst 1892 137 Lower Carboniferous


Rhodosphaera pulchra E nd ndash Ruumlst 1892 138 Upper Permian ndash Sosio Valley Area Western Sicily Italy

Rhodosphaera ruesti ndash nd ndash Hinde 1899a 44 Middle Devonian Givetian Yarramie Formation Tamworth Belt northern NSW Australia

RhopalastrumRhopalastrum

carbonicumL ndash ndash Ruumlst 1892 170 Lower


Rhopalastrum clavatum ndash nd ndash Hinde amp Fox 1895 641 Carboniferous Serpukhovian-Bashikirian


Rhopalastrum giganteum L ndash ndash Ruumlst 1892 171 Upper Permian ndash Sosio Valley Area Western Sicily Italy

Rhopalastrum ingens ndash nd ndash Hinde amp Fox 1895 641 Carboniferous Serpukhovian-Bashikirian


Rhopalastrum isselii L ndash ndash Ruumlst 1892 171 Upper Permian ndash Sosio Valley Area Western Sicily Italy

Rhopalastrum siculum L ndash ndash Ruumlst 1892 171 Upper Permian ndash Sosio Valley Area Western Sicily Italy

RhopalodictyumRhopalodictyum

astrictumndash nd ndash Ruumlst 1892 174 Upper Devonian ndash SchaebenholzHarz Mountains

Elbingerode Germany

Robotium (228)Robotium biauriculum N ndash ndash Cheng 1986 137 Lower



Robotium firmum N ndash ndash Cheng 1986 137 Lower Carboniferous


Robotium gordoni N ndash ndash Cheng 1986 138 Lower Carboniferous


Robotium ruesti N nc ndash Cheng 1986 139 Lower Carboniferous


Robotium validum N ndash Ts Cheng 1986 139 Lower Carboniferous


Robotium venustum N ndash ndash Cheng 1986 140 Lower Carboniferous


Rotasphaera (74)Rotasphaera

beckwithensisAr ndash ndash Noble 1994 20 Upper Silurian Gorstian Caballos Novaculite Formation

Beckwith Hills Marathon Basin W Texas USA

Rotasphaera delicata Ar ndash ndash Noble 1994 20 Upper Silurian Gorstian Caballos Novaculite Formation Payne Hills Bourland Mts Marathon Basin W Texas USA



Aitchison J C et al


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Rotasphaera (74) continuationRotasphaera

marathonensisAr ndash Ts Noble 1994 21 Middle Silurian Homerian Caballos Novaculite Formation

Payne Hills Bourland Mts Marathon Basin W Texas USA

Rotasphaera nuda Ar ndash ndash Noble 1994 21 Upper Silurian Gorstian Caballos Novaculite Formation East Bourland Mts Marathon Basin W Texas USA

Rotasphaera quadrata Ar ndash ndash Noble 1994 21 Middle Silurian Homerian Caballos Novaculite Formation East Bourland Mts Marathon Basin W Texas USA

Rotasphaera robertsorum Ar ndash ndash Noble 1994 22 Upper Silurian Gorstian Caballos Novaculite Formation Beckwith Hills Marathon Basin W Texas USA

Rotasphaera severa Ar ndash ndash MacDonald 1998 599 Lower Silurian Telychian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago

Russirad (304)Russirad kazintsovae IS

(OampF)ndash T Afanasieva 2000a 35 Upper Devonian middle



Ruzhencevispongus (214)Ruzhencevispongus

aktastiensisL ndash ndash Nazarov amp Ormiston 1993 44 Lower Permian Artinskian Aktastyi River Orenburgskaya

region Southern Urals RussiaRuzhencevispongus

aktastiensisL nn ndash Nazarov 1988 75 Lower Permian Artinskian Aktastyi River Orenburgskaya


apertusL ndash ndash Amon Braun amp Chuvashov

1990 126Lower Permian Artinskian Shelyvaginsk Formation around

Sim Town Chalyabinskaya region Urals Russia

Ruzhencevispongus cataphractus

L nn Ts Nazarov amp Ormiston 1984 pl 5 fig 11

Lower Permian Artinskian Kandurov Formation Donskoye Ural River Orenburgskaya Region S Urals Russia


L ndash ndash Nazarov amp Ormiston 1985 29 Lower Permian Artinskian Kandurov Formation Donskoye Ural River Orenburgskaya Region S Urals Russia

Ruzhencevispongus girtyi L ndash ndash Nazarov amp Ormiston 1985 32 Middle Permian Capitanian Bell Canyon Formation Guadalupe Mts Culbertson County W Texas USA

Ruzhencevispongus laqueus

L ndash ndash Nazarov amp Ormiston 1993 45 Lower Permian Artinskian Maloik Formation Don Village Ural River Orenburgskaya Region S Urals Russia

Ruzhencevispongus partilaminatus


Ruzhencevispongus retiporosus


Ruzhencevispongus rotundus


Ruzhencevispongus uralicus

L ndash T Kozur 1980 237 Lower Permian Sakmarian Sarabil Formation Verchneozernaja Station Orenburgskaya Region Urals Russia

Ruzhencevispongus uralicus subtriangulus

L ndash ndash Wang 1993b 9 Middle Permian Capitanian Gufeng Formation Jiangsu China

Ruzhencevispongus uralicus triradiatus


Ruzhencevispongus uralicus uralicus


Ruzhencevispongus cacumenireticulatus


Ruzhencevispongus emarginatus


Ruzhencevispongus plumatus

L ndash ndash Nazarov amp Ormiston 1985 30 Lower Permian Sakmarian Kandurov Formation Donskoye Ural River Orenburgskaya Region S Urals Russia

Ruzhencevispongus promiscuus






er

Stat

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Sanctipaulelum (288)Sanctipaulelum

biglobosumIS ndash ndash Won amp Iams 2015a 31 Lower Ordovician Floian Cow Head Group Western

Newfoundland CanadaSanctipaulelum pingue IS ndash ndash Won amp Iams 2015a 31 Lower Ordovician Floian Cow Head Group Western

Newfoundland CanadaSanctipaulelum

superpingueIS ndash ndash Won amp Iams 2015a 32 Lower Ordovician Floian Cow Head Group Western


contractumIS ndash ndash Won amp Iams 2015a 32 Lower Ordovician Floian Cow Head Group Western


indivisumIS ndash ndash Won amp Iams 2015a 33 Lower Ordovician Floian Cow Head Group Western

Newfoundland CanadaSanctipaulelum inusitata IS ndash ndash Won amp Iams 2015a 33 Lower Ordovician Floian Cow Head Group Western

Newfoundland Canada

Scharfenbergia (215)Scharfenbergia concava L ndash ndash Park amp Won 2012 68 Lower



Scharfenbergia plenospongiosa

L ndash ndash Won 1983 160 Lower Carboniferous


Scharfenbergia tailleurense

L ndash ndash Holdsworth amp Murchey 1988 787

Upper Carboniferous


Secuicollacta (75)Secuicollacta alaskensis Ar ndash ndash Won Blodgett amp Nestor 2002



Secuicollacta amoenitas Ar ndash ndash Nazarov amp Ormiston 1993 42 Upper Devonian Famenian Zapadno Valavskaya borehole 1-R Pripyat Depression Belarus

Secuicollacta amoenitas Ar nn ndash Nazarov 1988 130 Upper Devonian Famenian Zapadno Valavskaya borehole 1-R Pripyat Depression Belarus

Secuicollacta araneam Ar ndash ndash Aitchison 1993b 121 Upper Devonian lower Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Secuicollacta baijingensis Ar nn ndash Li 1995 333 Middle Ordovician


Secuicollacta cassa Ar ndash T Nazarov amp Ormiston 1984 75 Middle Silurian Homerian Makyutov Complex Tarangul River Kosistek Village Southern Ural Kazakhstan

Secuicollacta foliaspinella Ar ndash ndash Noble 1994 23 Middle Silurian Homerian Caballos Novaculite Formation East Bourland Mts Marathon Basin W Texas USA

Secuicollacta glaebosa Ar ndash ndash MacDonald 1998 592 Lower Silurian Telychian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago

Secuicollacta gliris Ar ndash ndash MacDonald 1998 589 Lower Silurian Telychian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago

Secuicollacta herrimani Ar ndash ndash MacDonald 1998 591 Lower Silurian Telychian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago

Secuicollacta horrida Ar ndash ndash Furutani 1990 50 Upper Silurian Gorstian Hitoegane Formation Hida-gaien Belt E of Hitoegane Fukuji area Takayama City Japan

Secuicollacta itoigawai Ar ndash ndash Furutani 1990 50 Upper Silurian Gorstian Hitoegane Formation Hida-gaien Belt E of Hitoegane Fukuji area Takayama City Japan

Secuicollacta labyrinthica Ar ndash ndash Aitchison 1993b 121 Upper Devonian lower Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Secuicollacta magnitesta Ar ndash ndash Won Blodgett amp Nestor 2002 949


Secuicollacta malevola Ar ndash ndash MacDonald 1998 589 Lower Silurian Telychian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago

Secuicollacta minuta Ar ndash ndash Goto Umeda amp Ishiga 1992 161




Aitchison J C et al


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Secuicollacta (75) continuationSecuicollacta nesymmetra Ar ndash ndash Li 1994 264 Upper Silurian Gorstian Keerhada Mayila Mts Xinjiang

ChinaSecuicollacta nesymmetris ndash nn ndash Li 1991 75 Upper Silurian Gorstian Mayila Complex Keerhada S

Mayila Mt Xinjiang ChinaSecuicollacta ornata Ar ndash ndash Goto Umeda amp Ishiga 1992



Secuicollacta parvitesta Ar ndash ndash Won Blodgett amp Nestor 2002 949


Secuicollacta resodiosae Ar ndash ndash MacDonald 1998 592 Lower Silurian Telychian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago

Secuicollacta sceptri Ar ndash ndash MacDonald 1998 589 Lower Silurian Telychian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago

Secuicollacta segari Ar ndash ndash MacDonald 1998 591 Lower Silurian Telychian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago

Secuicollacta silex Ar ndash ndash Goto Umeda amp Ishiga 1992 161


Secuicollacta solara Ar ndash ndash Noble 1994 24 Upper Silurian Gorstian Caballos Novaculite Formation Beckwith Hills Marathon Basin W Texas USA

Secuicollacta spissa Ar ndash ndash Nazarov amp Ormiston 1993 42 Lower Silurian ndash Sakmarsk Formation Bolrsquoshoy Abimevo Village Bashkortostan Region S Urals Russia

Secuicollacta spissa Ar nn ndash Nazarov 1988 122 Lower Silurian ndash Sakmarsk Formation Bolrsquoshoy Abimevo Village Bashkortostan Region S Urals Russia

Secuicollacta stelligera Ar ndash ndash Renz 1990 376 Upper Ordovician Katian Hanson Creek Formation N Martin Ridge Eureka County Nevada USA

Secuicollacta tatondukensis



Secuicollacta teli Ar ndash ndash MacDonald 1998 593 Lower Silurian Telychian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago

Secuicollacta vallipuellae Ar ndash ndash MacDonald 1998 592 Lower Silurian Telychian Cape Phillips Formation Cornwallis Island Canadian Arctic Archipelago

Secuicollacta vulgaris Ar ndash ndash Furutani 1990 49 Upper Silurian Gorstian Hitoegane Formation Hida-gaien Belt E of Hitoegane Fukuji area Takayama City Japan

Secuicollacta xinjiangensis

ndash nn ndash Li 1991 75 Upper Silurian Gorstian Mayila Complex Keerhada S Mayila Mt Xinjiang China

Secuicollacta exquisita Ar ndash ndash Wakamatsu Sugiyama amp Furutani 1990 170

Upper Silurian Ludfordian G3 formation of Gion-yama Group NE of Kyowa Gokase Town Kyushu Japan

Secuicollacta platyspina Ar ndash ndash Noble 1994 24 Middle Silurian Homerian Caballos Novaculite Formation East Bourland Mts Marathon Basin W Texas USA

SecuidcollactaSecuidcollacta hindei ndash nn ndash Li 1995 333 Middle



SethocapsaSethocapsa obstipa ndash nd ndash Ruumlst 1892 182 Lower Devonian ndash Southern Urals RussiaSethocapsa pytine ndash nd ndash Ruedemann amp Wilson 1936



SethocyrtisSethocyrtis excisa ndash nd ndash Ruumlst 1892 182 Lower


SethodiscusSethodiscus

chudleighensisndash nd ndash Hinde amp Fox 1895 639 Carboniferous Serpukhovian-


UK





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SethostaurusSethostaurus exsculptus ndash nd ndash Ruumlst 1892 165 Lower Silurian ndash Cabriegraveres Heacuterault France

Shangella (177)Shangella capitanensis L ndash ndash Nestell amp Nestell 2010 38 Middle Permian Capitanian Bell Canyon Formation Apache


Shangella longa L ndash T Feng in Feng et al 2006b 829 Upper Permian Changhsingian Dalong Formation Dongpan section Dongpan Village Guangxi Zhuang Region China

Shangella regularis L ndash ndash Feng in Feng et al 2006b 831 Upper Permian Changhsingian Dalong Formation Dongpan section Dongpan Village Guangxi Zhuang Region China

SinocyrtisSinocyrtis bellulus nr ndash Tnr Wang 1989 145 Lower Cambrian ndash Niutitang Formation Sandu


Sinocyrtis latobasalis nr ndash ndash Wang 1989 146 Lower Cambrian ndash Niutitang Formation Sandu district Guizhou Province China

Sinosphaera (120)Sinosphaera spinosa E ndash T Feng in Feng et al 2004a 137 Upper Permian Changhsingian Dalong Formation Dongpan


Siphonosphaera streptosiphonia



Somphoentactinia (165)Somphoentactinia cavata E ndash ndash Aitchison 1993b 119 Upper Devonian lower Frasnian Gogo Formation Pillara Range


Somphoentactinia gavrilovi



Somphoentactinia multisphaerata



Somphoentactinia saecularis

E ndash ndash Afanasieva amp Amon 2016 216 Lower Permian Artinskian Kondurovka section Sakmara River Southern Urals Russia

Sphaerodiscus (141)Sphaerodiscus rota E ndash Ts Won 1983 163 Lower


Frankenwald Germany

Sphaeroentactinia (46)Sphaeroentactinia

bispinosaAr ndash ndash Maletz amp Bruton 2007 265 Lower Ordovician Floian Valhallfonna Formation


Sphaeroentactinia hexaspinosa

Ar ndash T Maletz amp Bruton 2007 264 Lower Ordovician Floian Valhallfonna Formation Buldrebreen arm Ny Frieslan Spitsbergen Norway

Sphaeroentactinia integrata

Ar ndash ndash Maletz amp Bruton 2007 265 Lower Ordovician Floian Valhallfonna Formation Buldrebreen arm Ny Frieslan Spitsbergen Norway

Sphaeroentactinia robusta

Ar ndash ndash Won amp Iams 2015a 21 Lower Ordovician Floian Cow Head Group Western Newfoundland Canada

Sphaeroentactinia saintpaulensis


Sphaeroentactinia multiradiata

Ar ndash ndash Won amp Iams 2013 27 Lower Ordovician Floian Cow Head Group Western Newfoundland Canada

SphaeropyleSphaeropyle dreyeri ndash nd ndash Ruumlst 1892 141 Lower Devonian ndash Southern Urals RussiaSphaeropyle entostoma ndash nd ndash Ruumlst 1892 141 Upper Devonian ndash SchaebenholzHarz Mountains

Elbingerode GermanySphaeropyle eurystoma ndash nd ndash Ruumlst 1892 141 Upper Permian ndash Sosio Valley Area Western Sicily

ItalySphaeropyle laevis ndash nd ndash Ruumlst 1892 140 Lower Devonian ndash Southern Urals RussiaSphaeropyle

multangularisndash nd ndash Ruumlst 1892 141 Lower Devonian ndash Southern Urals Russia



Aitchison J C et al


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SphaerozoumSphaerozoum minutum ndash nd ndash Ruedemann amp Wilson 1936



Sphaerozoum patulum ndash nd ndash Hinde 1890 58 Middle Ordovician


Sphaerozoum priscum ndash nd ndash Hinde 1890 57 Middle Ordovician


Spinodeflandrella (8)Spinodeflandrella

acuminataAl ndash ndash Rudenko in Rudenko amp

Panasenko 1990b 192Upper Permian Wuchiapingian Pantov Member Tauka Terrane

Tumanovk Pantov area Primorskiy Kray Russia

Spinodeflandrella acutata Al ndash ndash Rudenko in Rudenko amp Panasenko 1990b 191


Spinodeflandrella bicornuta


Upper Permian Changhsingian Samarkinsk Formation Taukah Terrane Breevka Primorskiy Kray Russia

Spinodeflandrella obtusa Al ndash ndash Rudenko in Rudenko amp Panasenko 1990b 192


Spinodeflandrella tetraspinosa

Al ndash T Kozur 1981 267 Lower Permian Sakmarian Sarabil Formation Kondurovska Orenburgskaya Region S Urals Russia

Spinodeflandrella siciliensis


Lower Permian Kungurian Sosio Valley Area Torrente San Calogero Western Sicily Italy

SpirocapsaSpirocapsa singularis ndash nd Td Ruumlst 1892 192 Upper Permian ndash Sosio Valley Area Western Sicily

Italy

SpironiumSpironium haeckelii ndash nd ndash Ruumlst 1892 175 Lower


SpongatractusSpongatractus

pleurosigmandash nd ndash Ruumlst 1892 160 Upper Permian ndash Sosio Valley Area Western Sicily

Italy

Spongentactinella (166)Spongentactinella

abstrusa E ndash ndash Aitchison 1993b 120 Upper Devonian lower Frasnian Gogo Formation Pillara Range


Spongentactinella corynacantha


Spongentactinella faceta E ndash ndash Nazarov amp Ormiston 1993 34 Upper Devonian Famenian Zapadno Valavskaya borehole 1-R Pripyat Depression Belarus

Spongentactinella faceta E nn ndash Nazarov 1988 130 Upper Devonian Famenian Zapadno Valavskaya borehole 1-R Pripyat Depression Belarus

Spongentactinella intracata


Spongentactinella olafi E ndash ndash Afanasieva 2000a 82 Upper Devonian middle Frasnian


Spongentactinella windjanensis


Upper Devonian Frasnian Emanuel Range SW Longs Well Canning Basin Western Australia

Spongentactinia (121)Spongentactinia

concinna E ndash T Aitchison 1993b 116 Upper Devonian lower Frasnian Gogo Formation Pillara Range


Spongentactinia exquisita


Spongentactinia flexa E ndash ndash Won 1997b 404 Upper Devonian Frasnian Gogo Formation Pillara Range Canning Basin Western Australia





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Spongentactinia (121) continuationSpongentactinia fungosa E ndash T Nazarov 1975 75 Upper Devonian Frasnian Egindinsk Formation Aitpaika


Spongentactinia indisserta


Spongentactinia inusitata E ndash ndash Won 1997b 406 Upper Devonian Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Spongentactinia marina E ndash ndash Afanasieva amp Amon 2011 1507 Upper Devonian Famennian Zadonsk Formation W Lekkeyaginsk-65 Saremboi-Lekkeyaga Timan-Pechora Basin Russia

Spongentactinia nupera E ndash ndash Nazarov in Nazarov et al 1981 84


Spongentactinia olafi E nn ndash Afanasieva 1997a 221 Upper Devonian middle Frasnian


Spongentactinia polaris E ndash ndash Afanasieva amp Amon 2011 1505 Upper Devonian lower Famennian


Spongentactinia rigida E ndash ndash Amon Braun amp Chuvashov 1990 123


Spongentactinia shaijingpoensis


Spongentactinia subtiradiata

E ndash ndash Nazarov 1975 76 Lower Silurian ndash Sakmarsk Formation Bolrsquoshoy Abimevo Village Bashkortostan Region S Urals Russia

Spongentactinia ambigua


Spongentactinia incerta E ndash ndash Nestell Pope amp Nestell 2012 236

Upper Carboniferous


Spongentactinia mira E ndash ndash Won 1997b 410 Upper Devonian Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Spongentactinia simensis

E ndash ndash Amon Braun amp Chuvashov 1990 123


Spongocoelia (326)Spongocoelia citreum ndash nd Td Hinde 1899a 52 Middle Devonian Givetian Yarramie Formation Tamworth

Belt northern NSW AustraliaSpongocoelia

kamitakarensisIS ndash ndash Furutani 1990 48 Upper Silurian Gorstian Hitoegane Formation Hida-gaien


Spongocoelia oliva ndash nd ndash Hinde 1899a 52 Middle Devonian Givetian Yarramie Formation Tamworth Belt northern NSW Australia

Spongocoelia parvus IS ndash T Furutani 1990 47 Upper Silurian Gorstian Hitoegane Formation Hida-gaien Belt E of Hitoegane Fukuji area Takayama City Japan

SpongocyrtisSpongocyrtis

eurydictyumndash nd ndash Ruumlst 1892 181 Lower


SpongodictyonSpongodictyon

triquetrumndash nd ndash Ruumlst 1892 140 Lower Devonian ndash Southern Urals Russia

SpongodiscusSpongodiscus acinus ndash nd ndash Hinde 1899a 54 Middle Devonian Givetian Yarramie Formation Tamworth

Belt northern NSW AustraliaSpongodiscus cribrarius ndash nd ndash Hinde 1899a 54 Middle Devonian Givetian Yarramie Formation Tamworth

Belt northern NSW AustraliaSpongodiscus herzynicus ndash nd ndash Ruumlst 1892 172 Upper Devonian ndash SchaebenholzHarz Mountains

Elbingerode Germany



Aitchison J C et al


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Spongodiscus (continuation)Spongodiscus punctus ndash nd Tnr

ThHinde 1899a 54 Middle Devonian Givetian Yarramie Formation Tamworth

Belt northern NSW AustraliaSpongodiscus scutulatus IS ndash ndash Hinde 1899a 55 Middle Devonian Givetian Yarramie Formation Tamworth

Belt northern NSW AustraliaSpongodiscus

sphaereatusIS ndash ndash Liu 1992 129 Upper



Spongodiscus lenticulatus

IS ndash ndash Won 1983 165 Lower Carboniferous


SpongodruppaSpongodruppa

ornithopusndash nd ndash Ruumlst 1892 160 Lower


Spongodruppa triradiata ndash nd ndash Ruumlst 1892 159 Lower Carboniferous


SpongoentactinellaSpongoentactinella bella E ndash ndash Wang in Wang et al 2012 111 Upper Devonian Famennian Shiti Reservoir Formation Shiwu

road section Guangxi Zhuang Region SW China

Spongoentactinella borealia


Spongoentactinella micrococcua

E ndash ndash Wang in Wang et al 2012 111 Upper Devonian Frasnian Shiti Reservoir Formation Shiwu road section Guangxi Zhuang Region SW China

Spongoentactinella paracorynacantha


SpongoloncheSpongolonche lens E ndash ndash Hinde 1899a 55 Middle Devonian Givetian Yarramie Formation Tamworth


Spongomassa (47)Spongomassa gigantea Ar ndash T Won in Won amp Below 1999 337 Middle Cambrian ndash Inca Formation Georgina Basin

Queensland AustraliaSpongomassa

nannosphaeraAr ndash ndash Won in Won amp Below 1999 338 Middle Cambrian ndash Inca Formation Georgina Basin


SpongopilaSpongopila juniperina ndash nd ndash Ruumlst 1892 149 Upper Permian ndash Sosio Valley Area Western Sicily

Italy

SpongoplegmaSpongoplegma australe IS ndash ndash Hinde 1899a 44 Middle Devonian Givetian Yarramie Formation Tamworth

Belt northern NSW AustraliaSpongoplegma

brevisphaerandash nd ndash Aberdeen 1940 134 Upper Devonian Famennian Caballos Formation Marathon

Basin Texas USASpongoplegma

longispinosandash nd ndash Aberdeen 1940 134 Upper Devonian Famennian Caballos Formation Marathon

Basin Texas USASpongoplegma priscum ndash nd ndash Hinde 1890 48 Middle


Scotland UK

SpongoprunumSpongoprunum

holodictyosumndash nd ndash Ruumlst 1892 159 Lower


Spongoprunum oligoporum



Spongoprunum trispinosum


Spongosphaera (122)Spongosphaera

crasseptataE ndash ndash Wang in Wang et al 2012 112 Upper Devonian Famennian Shiti Reservoir Formation Shiwu


Spongosphaera grandispongiosa

E ndash T Th


Spongosphaera induta ndash nd ndash Ruumlst 1892 150 Lower Devonian ndash Southern Urals Russia





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Spongosphaera (122) continuationSpongosphaera

trendallianaE ndash ndash Won 1997b 412 Upper Devonian Frasnian Gogo Formation Pillara Range


Spongosphaera tritestacea

ndash nd ndash Rothpletz 1880 449 Lower Silurian Telchyian Langenstriegis FrankenbergSa Germany



Spongosphaera varia E ndash Ts Won 1997b 412 Upper Devonian Frasnian Gogo Formation Pillara Range Canning Basin Western Australia

Spongosphaeradiscus (247)Spongosphaeradiscus

shaiwaensisS ndash ndash Wang in Wang amp Shang 2001

118Upper Permian Wuchiapingian Shaiwa Group Shaiwa Ziyun

County Guizhou China

SpongotripusSpongotripus

concentricusL ndash Th

TsRuumlst 1892 173 Lower


Spongotripus fenestratus IS ndash ndash Hinde 1899a 55 Middle Devonian Givetian Yarramie Formation Tamworth Belt northern NSW Australia

Spongotripus patella IS ndash Td Hinde 1899a 55 Middle Devonian Givetian Yarramie Formation Tamworth Belt northern NSW Australia

Spongotripus ruestae L ndash ndash Ormiston amp Lane 1976 168 Lower Carboniferous


Spongotripus sinensis L ndash ndash Liu 1992 129 Upper Carboniferous


Spongotripus xintianensis

E ndash ndash Sheng amp Wang 1982 61 Middle Devonian Givetian Qizigiao Formation Matangwuo Xintian district Hunan China

SpongotrochusSpongotrochus

elongatusndash nd ndash Ruumlst 1892 173 Upper Permian ndash Sosio Valley Area Western Sicily

ItalySpongotrochus

primaevusndash nd ndash Ruedemann amp Wilson 1936



Spongotrochus discoidalis

S nc ndash Won 1983 164 Lower Carboniferous


SpongurusSpongurus lacunosus ndash nd ndash Ruumlst 1892 159 Upper Devonian ndash SchaebenholzHarz Mountains

Elbingerode GermanySpongurus plantaeformis ndash nd ndash Ruumlst 1892 159 Upper Permian ndash Sosio Valley Area Western Sicily

ItalySpongurus variabilis ndash nd ndash Ruumlst 1892 158 Lower Devonian ndash Southern Urals Russia

Srakaeosphaera (327)Srakaeosphaera minuta ndash nd Td Sashida in Sashida et al 1997

12Middle Permian Capitanian Sra Kaeo Group Ban Bo Rae Sra

Kaeo Thailand

StauracontiumStauracontium inaequale E ndash ndash Ruumlst 1892 145 Upper Permian ndash Sosio Valley Area Western Sicily

ItalyStauracontium

incompletumndash nd ndash Ruumlst 1892 145 Lower


Stauracontium xiphophorum

E ndash ndash Ruumlst 1892 145 Lower Carboniferous


StauralastrumStauralastrum aculeatum L ndash ndash Ruumlst 1892 171 Upper Permian ndash Sosio Valley Area Western Sicily

Italy

Staurentactinia (186)Staurentactinia nazarovi L ndash T Schwartzapfel amp Holdsworth



Carter County Oklahoma USA

StaurodorasStaurodoras brevispinosa ndash nd ndash Aberdeen 1940 136 Upper Devonian Famennian Caballos Formation Marathon

Basin Texas USAStaurodoras gracilis ndash nd ndash Hinde 1890 49 Middle


Scotland UK



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Staurodruppa (123)Staurodruppa foxii E ndash ndash Hinde 1899b 217 Middle Devonian Givetian Chypons Farm Mullion Parrish

Cornwall England UKStaurodruppa nana E ndash ndash Hinde 1899a 52 Middle Devonian Givetian Yarramie Formation Tamworth

Belt northern NSW AustraliaStaurodruppa nucula E ndash ndash Hinde 1899a 52 Middle Devonian Givetian Yarramie Formation Tamworth

Belt northern NSW AustraliaStaurodruppa praelonga E ndash Ts Hinde 1899a 51 Middle Devonian Givetian Yarramie Formation Tamworth

Belt northern NSW AustraliaStaurodruppa prolata E ndash ndash Foreman 1963 275 Upper Devonian Famennian Huron member Ohio Shale Ohio

USA

StauroloncheStaurolonche davidi E ndash ndash Hinde 1899a 46 Middle Devonian Givetian Yarramie Formation Tamworth

Belt northern NSW AustraliaStaurolonche insignis E ndash ndash Ruumlst 1892 144 Upper Permian ndash Sosio Valley Area Western Sicily

ItalyStaurolonche laterna E ndash ndash Hinde 1899a 47 Middle Devonian Givetian Yarramie Formation Tamworth

Belt northern NSW AustraliaStaurolonche

macracanthaE ndash ndash Ruumlst 1892 144 Upper Permian ndash Sosio Valley Area Western Sicily

ItalyStaurolonche micropora ndash nd ndash Ruumlst 1892 143 Lower Silurian ndash Cabriegraveres Heacuterault FranceStaurolonche scitula ndash nd ndash Hinde 1899a 47 Middle Devonian Givetian Yarramie Formation Tamworth

Belt northern NSW AustraliaStaurolonche spinigera ndash nd ndash Ruumlst 1892 144 Lower


Staurolonche tenella E ndash ndash Hinde 1899a 47 Middle Devonian Givetian Yarramie Formation Tamworth Belt northern NSW Australia

StaurolonchidiumStaurolonchidium coheni E ndash ndash Ruumlst 1892 144 Lower


Staurolonchidium obliquum


Stauroplegma (328)Stauroplegma barbatum ndash nd ndash Hinde 1890 50 Middle


Scotland UKStauroplegma brevispina ndash nd Td Hinde 1890 50 Middle


Scotland UKStauroplegma

compressumndash nd ndash Hinde 1890 50 Middle


Scotland UKStauroplegma diffusum ndash nd ndash Hinde 1890 50 Middle



houxiaensendash nd ndash Liu 1992 128 Upper



Stauroplegma lepidum S ndash ndash Sheng amp Wang 1982 60 Middle Devonian Givetian Qizigiao Formation Matangwuo Xintian district Hunan China

Stauroplegma nanjingensis

ndash nd ndash Zhang Wu amp Liu 1992 299 Upper Permian Changhsingian Dalong Formation Zhengpanshan near Longtan E Nanjing China

Stauroplegma pulcherrimum

S ndash ndash Sheng amp Wang 1982 60 Middle Devonian Givetian Qizigiao Formation Matangwuo Xintian district Hunan China

Stauroplegma robustospina


Stauroplegma tianshanense

ndash nd ndash Liu 1992 128 Upper Carboniferous


StaurosphaeraStaurosphaera

crassispinandash nd ndash Ruedemann amp Wilson 1936



Staurosphaera fragilis ndash nd ndash Ruumlst 1892 143 Lower Silurian ndash Cabriegraveres Heacuterault FranceStaurosphaera ornata E ndash ndash Hinde 1899a 46 Middle Devonian Givetian Yarramie Formation Tamworth

Belt northern NSW AustraliaStaurosphaera pusilla E ndash ndash Hinde 1899a 46 Middle Devonian Givetian Yarramie Formation Tamworth

Belt northern NSW AustraliaStaurosphaera

quadrangularisndash nd ndash Ruumlst 1892 143 Lower Silurian ndash Cabriegraveres Heacuterault France





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Staurosphaera (continuation)Staurosphaera rotunda ndash nd ndash Aberdeen 1940 135 Upper Devonian Famennian Caballos Formation Marathon

Basin Texas USAStaurosphaera sancta ndash nd ndash Ruedemann amp Wilson 1936



Staurosphaera setispina ndash nd ndash Ruumlst 1892 143 Lower Devonian ndash Southern Urals Russia

StaurosphaerellaStaurosphaerella

setispiniformisndash nd ndash Demanet 1938 29 Lower

CarboniferousVisean Dinant Synclinorium Belgium

StaurostylusStaurostylus caucasicus ndash nd ndash Khabakov 1932 224 Carboniferous ndash Left river bank of Kala-kulak

Elbrus Caucasus RussiaStaurostylus tenuispinus ndash nd ndash Ruumlst 1892 143 Lower


Staurostylus varispinatus ndash nd ndash Aberdeen 1940 135 Upper Devonian Famennian Caballos Formation Marathon Basin Texas USA

Staurostylus xiphophorus ndash nd Td Ruumlst 1892 143 Lower Devonian ndash Southern Urals Russia

StichocapsaStichocapsa anguillula ndash nd ndash Ruumlst 1892 192 Lower Devonian ndash Southern Urals RussiaStichocapsa biceps ndash nd ndash Ruumlst 1892 191 Upper Permian ndash Sosio Valley Area Western Sicily

ItalyStichocapsa bukkiana ndash nd ndash Ruumlst 1892 191 Carboniferous ndash Buumlkk-Gebirge Inner Western

Carpathians HungaryStichocapsa citriformis ndash nd ndash Ruumlst 1892 191 Upper Permian ndash Sosio Valley Area Western Sicily

ItalyStichocapsa ellipsoides ndash nd ndash Ruumlst 1892 191 Upper Permian ndash Sosio Valley Area Western Sicily

ItalyStichocapsa elpenor ndash nd ndash Ruumlst 1892 192 Lower Devonian ndash Southern Urals RussiaStichocapsa gravida ndash nd ndash Ruumlst 1892 191 Upper Permian ndash Sosio Valley Area Western Sicily

ItalyStichocapsa pupa ndash nd ndash Ruumlst 1892 191 Upper Permian ndash Sosio Valley Area Western Sicily

ItalyStichocapsa taenia ndash nd ndash Ruumlst 1892 192 Lower Devonian ndash Southern Urals Russia

StichoperaStichopera aculeata ndash nd ndash Ruumlst 1892 186 Upper Permian ndash Sosio Valley Area Western Sicily

Italy

StigmosphaeraStigmosphaera echinata ndash nd ndash Hinde 1899b 215 Middle Devonian Givetian Chypons Farm Mullion Parrish

Cornwall England UKStigmosphaera mira E ndash Td Ruumlst 1892 135 Lower


Stigmosphaera rothpletzii E ndash ndash Ruumlst 1892 135 Lower Carboniferous


Stigmosphaera suspecta E ndash ndash Ruumlst 1892 135 Lower Carboniferous


Stigmosphaerostylus (124)Stigmosphaerostylus

bazhaiensisE ndash ndash Wang in Wang amp Luo 2009 136 Upper Devonian Frasnian Xiangshuidong Formation Bazhai

Ziyun County Guizhou ChinaStigmosphaerostylus

cubicusE ndash ndash Luo Aitchison amp Wang 2002

116Middle Devonian Givetian Tanhe Formation Wuxiangling

section S of Nanning Guangxi Zhuang Region SW China

Stigmosphaerostylus dobruskinae


Stigmosphaerostylus favusa


Stigmosphaerostylus inaequalis

ndash nd ndash Hinde 1899b 216 Middle Devonian Givetian Chypons Farm Mullion Parrish Cornwall England UK

Stigmosphaerostylus istanbulensis

E ndash ndash Noble Tekin Gedik amp Pehlivan 2008 51

Lower Carboniferous


Stigmosphaerostylus leveni




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Stigmosphaerostylus (124) continuationStigmosphaerostylus

meyeniE ndash ndash Kozur amp Mostler 1989 189 Lower Permian Sakmarian Sarabil Formation Sakmarskiy


Stigmosphaerostylus notabilis

E ndash Ts Ruumlst 1892 142 Lower Carboniferous


Stigmosphaerostylus qinzhouensis

E ndash ndash Wang in Wang et al 2012 110 Lower Carboniferous


Stigmosphaerostylus sakmaraensis


Stigmosphaerostylus sakmaraensis elevatus


Stigmosphaerostylus sakmaraensis sakmaraensis


Stigmosphaerostylus wildei

E ndash ndash Noble amp Jin 2010 125 Middle Permian Capitanian Bell Canyon Formation Guadalupe Mts Culberson County W Texas USA

Stigmosphaerostylus yaoi E ndash ndash Kozur amp Mostler 1989 190 Lower Permian Sakmarian Sarabil Formation Sakmarskiy Orenburgskaya Region S Urals Russia

Stigmosphaerostylus coronatus


Stigmosphaerostylus variospinus


Stylactinosphaera (305)Stylactinosphaera prima IS

(OampF)ndash T Noble 1994 36 Middle Silurian Homerian Caballos Novaculite Formation


StylodictyaStylodictya polaris E ndash ndash Ruumlst 1892 169 Upper Devonian ndash SchaebenholzHarz Mountains

Elbingerode GermanyStylodictya zittelii ndash nd ndash Ruumlst 1892 168 Upper Devonian ndash SchaebenholzHarz Mountains

Elbingerode Germany

StylodiscusStylodiscus dictyosus E ndash ndash Ruumlst 1892 162 Lower


StylosphaeraStylosphaera acuta E ndash ndash Ruumlst 1892 142 Lower


Stylosphaera obtusa E ndash ndash Hinde 1899a 45 Middle Devonian Givetian Yarramie Formation Tamworth Belt northern NSW Australia

Stylosphaera quasiobtusa


Stylosphaera vetusta E ndash ndash Hinde 1899a 46 Middle Devonian Givetian Yarramie Formation Tamworth Belt northern NSW Australia

Stylosphaera magnaspina

S ndash ndash Noble 1994 39 Upper Silurian Gorstian Caballos Novaculite Formation Payne Hills Bourland Mts Marathon Basin W Texas USA

StylostaurusStylostaurus hindei ndash nd ndash Ruedemann amp Wilson 1936



StylotrochusStylotrochus dolfussi ndash nd ndash Ruumlst 1892 173 Upper Permian ndash Sosio Valley Area Western Sicily

Italy

StyptosphaeraStyptosphaera antiqua ndash nd ndash Hinde 1890 47 Middle


Scotland UK





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Subechidnina (58)Subechidnina delicata Ar ndash ndash Won amp Iams 2002 26 Upper Cambrian Stage 10 Cow Head Group Western

Newfoundland CanadaSubechidnina florea Ar ndash ndash Won amp Iams 2002 26 Upper Cambrian Stage 10 Cow Head Group Western

Newfoundland CanadaSubechidnina gracilis Ar ndash Ts Won amp Iams 2002 27 Upper Cambrian Stage 10 Cow Head Group Western

Newfoundland CanadaSubechidnina pressa Ar ndash ndash Won amp Iams 2002 27 Upper Cambrian Stage 10 Cow Head Group Western

Newfoundland Canada

Svalbardospiculum (68)Svalbardospiculum

arenigiumAr ndash T Maletz amp Bruton 2007 259 Lower Ordovician Floian Valhallfonna Formation


Syntagentactinia (135)Syntagentactinia afflicata E nn ndash Nazarov 1988 pl 11 fig 6 Lower Silurian ndash Sakmarsk Formation Bolrsquoshoy


Syntagentactinia afflicta E ndash ndash Nazarov amp Ormiston 1990 19 Lower Silurian ndash Sakmarsk Formation Bolrsquoshoy Abimevo Village Bashkortostan Region S Urals Russia

Syntagentactinia afflicta E inv ndash Nazarov amp Ormiston 1993 40 Lower Silurian ndash Sakmarsk Formation Bolrsquoshoy Abimevo Village Bashkortostan Region S Urals Russia

Syntagentactinia biocculosa

E ndash T Nazarov in Nazarov amp Popov 1980 56

Middle Ordovician


Syntagentactinia excelsa E ndash ndash Nazarov amp Ormiston 1993 40 Lower Silurian ndash Sakmarsk Formation Bolrsquoshoy Abimevo Village Bashkortostan Region S Urals Russia

Syntagentactinia excelsa E nn ndash Nazarov 1988 122 Lower Silurian ndash Sakmarsk Formation Bolrsquoshoy Abimevo Village Bashkortostan Region S Urals Russia

Syntagentactinia pauca E ndash ndash Nazarov in Nazarov amp Popov 1980 57

Middle Ordovician


Syntagentactinia pentarhachia


Syntagentactinia angulata

E ndash ndash Pouille amp Danelian in Pouille et al 2014 152

Middle Ordovician

upper Darriwilian


Tamonella (248)Tamonella aculeata S ndash ndash Feng in Feng et al 2006a 37 Upper Permian Changhsingian Dalong Formation Dongpan


Tetracircinata (206)Tetracircinata reconda L ndash Ts Nazarov amp Ormiston 1984 74 Lower Permian Artinskian Makyutov Complex Sim Village

Chalyabinskaya Region Kazakhstan

Tetraedroclathrum (158)Tetraedroclathrum

cabrierensendash nn Tn Deflandre 1960 216 Lower


Tetragregnon (207)Tetragregnon iowaensis L ndash ndash Nestell Pope amp Nestell 2012

239Upper



Tetragregnon japonicum L ndash ndash Sashida amp Tonishi 1985 13 Upper Permian Changhsingian Ohirayama Unit South Chichibu Belt Kashiwara Akigawa River Akiruno City Tokyo Japan

Tetragregnon longispinatum


Tetragregnon nitidus L ndash ndash Nazarov amp Ormiston 1985 23 Middle Permian Capitanian Bell Canyon Formation Guadalupe Mts Culbertson County W Texas USA

Tetragregnon pyramidatus


Upper Carboniferous


Tetragregnon scalpratus L ndash ndash Nazarov amp Ormiston 1985 23 Middle Permian Capitanian Bell Canyon Formation Guadalupe Mts Culbertson County W Texas USA



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Tetragregnon (207) continuationTetragregnon sphaericus L ndash ndash Nazarov in Isakova amp Nazarov

1986 72Upper


Urals RussiaTetragregnon

sycamorensisL ndash T Ormiston amp Lane 1976 167 Lower



Tetragregnon sycamorensis tricarinatus



Tetragregnon tunicatus L ndash ndash Nazarov in Isakova amp Nazarov 1986 74

Upper Carboniferous


Tetragregnon vimineum L ndash ndash Amon Braun amp Chuvashov 1990 124


TetrapyloniumTetrapylonium ovatum ndash nd ndash Ruumlst 1892 174 Upper Permian ndash Sosio Valley Area Western Sicily

Italy

TetraregnonTetraregnon permicus E ndash ndash Kozur amp Mostler 1989 185 Lower Permian Sakmarian Sarabil Formation Sakmarskiy


Tetrasphaera (329)Tetrasphaera

kuruktagensisS ndash Th

TsWang in Wang et al 2008 397 Middle



TetraspongentactiniaTetraspongentactinia

riescheidensisL ndash ndash Won amp Seo 2010 249 Lower


GermanyTetraspongentactinia

triporosaL ndash ndash Won amp Seo 2010 249 Lower


Germany

Tetraspongoactinia (208)Tetraspongoactinia

holdsworthiL ndash T Won 1998 257 Lower


Germany

TetraspongodiscusTetraspongodiscus

stauracanthusS ndash ndash Feng in Feng et al 2006a 39 Upper Permian Changhsingian Dalong Formation Dongpan


Tetraspongodiscus tetragonius

S ndash ndash Feng in Feng et al 2006a 41 Upper Permian Changhsingian Dalong Formation Dongpan section Dongpan Village Guangxi Zhuang Region China

Tetratormentum (209)Tetratormentum acutum L ndash ndash Sashida amp Tonishi 1988 532 Upper Permian Changhsingian Ohirayama Unit South Chichibu


Tetratormentum crateris L ndash ndash Nazarov in Isakova amp Nazarov 1986 102

Upper Carboniferous


Tetratormentum diremptum


Upper Carboniferous


Tetratormentum globiforme


Tetratormentum magniporosa

L ndash ndash Won amp Seo 2010 251 Lower Carboniferous


Tetratormentum nandanensis

L ndash ndash Wu amp Feng in Wu et al 2010 889


Tetratormentum narthecium

L ndash T Nazarov amp Ormiston 1985 42 Upper Carboniferous


Tetratormentum ormistoni


Tetratormentum condensum


Upper Carboniferous






er

Stat

usTy




Tetrentactinia (167)Tetrentactinia barysphaera E ndash T Foreman 1963 282 Upper Devonian Famennian Huron member Ohio Shale Ohio

USATetrentactinia

gracilispinosa E ndash ndash Foreman 1963 284 Upper Devonian Famennian Huron member Ohio Shale Ohio

USATetrentactinia incondita E ndash ndash Nazarov 1973a 698 Upper Devonian Frasnian Egindinsk Formation Aitpaika


Tetrentactinia inopia E ndash ndash Nazarov in Nazarov et al 1981 84


Tetrentactinia puncticulosa E ndash ndash Nazarov 1975 74 Lower Silurian ndash Sakmarsk Formation Bolrsquoshoy Abimevo Village Bashkortostan Region S Urals Russia

Tetrentactinia quadrispinosa


Tetrentactinia somphosphaera


Tetrentactinia somphozona


Tetrentactinia spinulosa E ndash ndash Kiessling amp Tragelehn 1994 238Upper Devonian Famennian Frankenwald north Bavaria GermanyTetrentactinia spongacea E ndash ndash Foreman 1963 284 Upper Devonian Famennian Huron member Ohio Shale Ohio

USATetrentactinia teuchestes E ndash ndash Foreman 1963 283 Upper Devonian Famennian Huron member Ohio Shale Ohio

USATetrentactinia veles E ndash Td Foreman 1963 283 Upper Devonian Famennian Huron member Ohio Shale Ohio

USA

Thecentactinia (126)Thecentactinia

indeterminataE ndash ndash Nazarov 1975 72 Lower Silurian ndash Sakmarsk Formation Bolrsquoshoy


ThecosphaeraThecosphaera

hexpenetratandash nd ndash Aberdeen 1940 134 Upper Devonian Famennian Caballos Formation Marathon

Basin Texas USAThecosphaera sexactis E ndash ndash Ruumlst 1892 137 Upper Permian ndash Sosio Valley Area Western Sicily

ItalyThecosphaera sicula ndash nd ndash Ruumlst 1892 137 Upper Permian ndash Sosio Valley Area Western Sicily

Italy

TheocampeTheocampe bulbosa ndash nd ndash Ruumlst 1892 185 Upper Permian ndash Sosio Valley Area Western Sicily

Italy

TheodiscusTheodiscus aculeatus ndash nd ndash Hinde amp Fox 1895 639 Carboniferous Serpukhovian-


UKTheodiscus

brachyacanthusndash nd ndash Ruumlst 1892 163 Lower


Theodiscus convexus ndash nd ndash Ruumlst 1892 163 Lower Carboniferous


Theodiscus gigas ndash nd ndash Ruumlst 1892 163 Lower Silurian ndash Cabriegraveres Heacuterault FranceTheodiscus hastatus E ndash ndash Hinde 1899a 53 Middle Devonian Givetian Yarramie Formation Tamworth

Belt northern NSW AustraliaTheodiscus planus ndash nd ndash Ruumlst 1892 163 Lower Silurian ndash Cabriegraveres Heacuterault France

TheosyringiumTheosyringium praecox ndash nd ndash Ruumlst 1892 184 Upper Permian ndash Sosio Valley Area Western Sicily

Italy

Tlecerina (159)Tlecerina exilis E ndash ndash Furutani 1983 111 Lower Devonian Emsian Nakahata Formation S Mt


Tlecerina fenestrata E ndash ndash Furutani 1983 111 Lower Devonian Emsian Nakahata Formation S Mt Yokokurayama Ochi Town Kochi Shikoku Japan

Tlecerina horrida E ndash T Furutani 1983 110 Lower Devonian Emsian Nakahata Formation S Mt Yokokurayama Ochi Town Kochi Shikoku Japan

Tlecerina isensis E ndash ndash Kurihara amp Sashida 2000 62 Lower Devonian Emsian Shibasudani Formation Shibasudani Valley Ohno City Fukui Prefecture Japan



Aitchison J C et al


er

Stat

usTy




Tormentum (216)Tormentum aequilateralis L ndash ndash Nazarov in Isakova amp Nazarov

1986 97Upper


Urals RussiaTormentum aequilateralis

orbiculatumL ndash ndash Nazarov in Isakova amp Nazarov

1986 97Upper


Urals RussiaTormentum circumfluum L ndash ndash Nazarov in Isakova amp Nazarov

1986 100Upper


Urals RussiaTormentum circumfusum L ndash ndash Nazarov amp Ormiston 1985 40 Lower Permian Sakmarian Maloik Formation Don Village Ural

River Orenburgskaya Region S Urals Russia

Tormentum coarctatum L ndash ndash Nazarov amp Ormiston 1993 44 Upper Carboniferous


Tormentum complicatum L ndash ndash Nazarov in Isakova amp Nazarov 1986 100

Upper Carboniferous


Tormentum delicatum L ndash ndash Nazarov amp Ormiston 1985 40 Middle Permian Roadian Bell Canyon Formation Guadalupe Mts Culbertson County W Texas USA

Tormentum orientalis L ndash ndash Nazarov amp Ormiston 1993 44 Upper Carboniferous


Tormentum pervagatum L ndash ndash Nazarov amp Ormiston 1985 40 Upper Carboniferous


Tormentum protei L ndash T Nazarov amp Ormiston 1983b 376Upper Carboniferous


Tormentum protei ndash inv ndash Nazarov amp Ormiston 1985 40 Upper Carboniferous


Tormentum protei pannosum


Upper Carboniferous


Tormentum rotundatum L ndash ndash Feng in Feng et al 2006b 831 Upper Permian Changhsingian Dalong Formation Dongpan section Dongpan Village Guangxi Zhuang Region China

Tormentum sertulum L ndash ndash Nazarov amp Ormiston 1985 41 Middle Permian Capitanian Bell Canyon Formation Hegler Limestone Culberson County W Texas USA

Tormentum subrotundum L ndash ndash Nazarov in Isakova amp Nazarov 1986 99

Upper Carboniferous


Tormentum inflatum L ndash ndash Nazarov amp Ormiston 1985 41 Middle Permian Capitanian Bell Canyon Formation Hegler Limestone Culberson County W Texas USA

Totollum (232)Totollum blomei N ndash T Schwartzapfel amp Holdsworth



Carter County Oklahoma USA Totollum undulatum N ndash ndash Schwartzapfel amp Holdsworth



Carter County Oklahoma USA Totollum ziegleri N ndash ndash Schwartzapfel amp Holdsworth




TriactiscusTriactiscus lanceola E ndash ndash Hinde 1899a 53 Middle Devonian Givetian Yarramie Formation Tamworth


Triactofenestrella (178)Triactofenestrella nicolica L ndash Ts Nazarov amp Ormiston 1984 79 Upper

CarboniferousGzhelian Makyutov Complex Tarangul River

Kosistek Village Southern Ural Kazakhstan

Triaenoentactinosphaera (169)Triaenoentactinosphaera

regulariaE ndash Ts Wang 1997 156 Upper Devonian Famennian Gennaren Formation S Utubulak

NW Jungar Basin Xinjiang China

Triaenosphaera (168)Triaenosphaera hebes E ndash ndash Won 1983 167 Lower


Frankenwald GermanyTriaenosphaera

longispinaE ndash ndash Sashida in Sashida et al 2000b

250Lower

CarboniferousTournaisian Khlong Kon Limestone along

Highway 42 Saba Yoi area SE Thailand





er

Stat

usTy




Triaenosphaera (168) continuationTriaenosphaera

megacanthaE ndash ndash Feng in Feng et al 2007 29 Upper Permian Changhsingian Dalong Formation Dongpan


Triaenosphaera minuscula


upper Tournaisian


Triaenosphaera minutus E ndash ndash Sashida amp Tonishi 1988 530 Upper Permian Changhsingian Ohirayama Unit South Chichibu Belt Kashiwara Akigawa River Akiruno City Tokyo Japan

Triaenosphaera sicarius ndash nn ndash Deflandre 1960 216 Lower Carboniferous


Triaenosphaera sicarius E ndash T Deflandre 1973c 1150 Lower Carboniferous


Triaenosphaera tetraculeata



Triaenosphaera variabilis E ndash ndash Feng in Feng et al 2007 29 Upper Permian Changhsingian Dalong Formation Dongpan section Dongpan Village Guangxi Zhuang Region China

Triaenosphaera bareillensis



Triaenosphaera leherissei



TricolocapsaTricolocapsa abdominalis ndash nd ndash Ruumlst 1892 186 Upper Permian ndash Sosio Valley Area Western Sicily

ItalyTricolocapsa anceps ndash nd ndash Ruumlst 1892 185 Upper Permian ndash Sosio Valley Area Western Sicily

ItalyTricolocapsa grandaeva ndash nd ndash Ruumlst 1892 185 Lower


Tricolocapsa thoracica ndash nd ndash Ruumlst 1892 186 Upper Permian ndash Sosio Valley Area Western Sicily Italy

Trifidospongus (179)Trifidospongus angustus L ndash ndash Noble amp Renne 1990 389 Middle Permian Roadian Dekkas Formation Eastern

Klamath Mountains California USA

Trifidospongus dekkasensis

L ndash Ts Noble amp Renne 1990 388 Middle Permian Roadian Dekkas Formation Eastern Klamath Mountains California USA

Trigonosphaera (306)Trigonosphaera calvata IS

(OampF)ndash T Feng in Feng et al 2004a 142 Upper Permian Changhsingian Dalong Formation Dongpan


Trilacertus (187)Trilacertus baumgartneri L ndash Ts Cheng 1986 178 Lower



Trilacertus delicatus L ndash ndash Cheng 1986 178 Upper Carboniferous


Trilacertus densus L ndash ndash Cheng 1986 179 Lower Carboniferous


Trilonche (127)Trilonche chiangdaoensis E ndash ndash Wonganan amp Caridroit 2005 46 Upper Devonian Famennian Chiang Dao chert along the Road

1150 for Phrao City Chiang Mai northern Thailand

Trilonche cimelia E ndash ndash Ormiston amp Lane 1976 167 Lower Carboniferous


Trilonche crassus E ndash ndash He amp Feng in He et al 2011 491


Trilonche dihelicis E ndash ndash Wonganan amp Caridroit 2005 46 Upper Devonian Famennian Chiang Dao chert along the Road 1150 for Phrao City Chiang Mai northern Thailand

Trilonche elegans E ndash ndash Hinde 1899a 48 Middle Devonian Givetian Yarramie Formation Tamworth Belt northern NSW Australia



Aitchison J C et al


er

Stat

usTy




Trilonche (127) continuationTrilonche nanningensis E ndash ndash Luo Aitchison amp Wang 2002



Trilonche parapalimbola E ndash ndash Wang in Wang et al 2003 133 Upper Devonian Famennian Shiti Reservoir Formation E Bancheng Guangxi Zhuang Region SW China

Trilonche pittmani E ndash ndash Hinde 1899a 48 Middle Devonian Givetian Yarramie Formation Tamworth Belt northern NSW Australia

Trilonche remosa E ndash ndash Luo Aitchison amp Wang 2002 119


Trilonche tanheensis E ndash ndash Luo Aitchison amp Wang 2002 119


Trilonche textilis E ndash ndash Feng in Feng et al 2007 25 Upper Permian Changhsingian Dalong Formation Dongpan section Dongpan Village Guangxi Zhuang Region China

Trilonche trifoliolata E ndash ndash Umeda 1997 422 Upper Silurian Pridoli Jyoru Formation Kurosegawa Belt Mt Konomori Kochi City Shikoku Japan

Trilonche vachardi E ndash ndash Wonganan amp Caridroit 2005 48 Upper Devonian Frasnian Chiang Dao chert along the Road 1150 for Phrao City Chiang Mai northern Thailand

Trilonche vetusta E ndash T Hinde 1899a 48 Middle Devonian Givetian Yarramie Formation Tamworth Belt northern NSW Australia

Trilonche xinpoensis E ndash ndash Luo Aitchison amp Wang 2002 120Middle Devonian Givetian Tanhe Formation Wuxiangling section S of Nanning Guangxi Zhuang Region SW China

Trilonche yaoi E ndash ndash Zhang Xia Dong amp Shang 2008 188

Lower Carboniferous

lower Tournaisian


TripilidiumTripilidium dissociatum ndash nd ndash Ruumlst 1892 178 Lower


Triplanospongos (180)Triplanospongos

musashiensisL ndash Ts Sashida amp Tonishi 1988 536 Upper Permian Changhsingian Ohirayama Unit South Chichibu


Triplococcus (266)Triplococcus acanthicus S ndash T Danelian amp Popov 2003 333 Lower Ordovician Floian Akzhal Mts S of Betpak-Dala

desert KazakhstanTriplococcus

aksuranensisS ndash ndash Pouille amp Danelian


Ordovicianupper


region Kazakhstan

Triposphaera (330)Triposphaera armata ndash nd ndash Hinde 1890 56 Middle


Scotland UKTriposphaera densa ndash nd ndash Hinde 1890 55 Middle


Scotland UKTriposphaera hastata ndash nd ndash Hinde 1890 55 Middle


Scotland UKTriposphaera maxima ndash nd ndash Ruedemann amp Wilson 1936



Triposphaera peachii ndash nd Td Hinde 1890 55 Middle Ordovician


Triposphaera permica E ndash ndash Kozur amp Mostler 1989 191 Lower Permian Sakmarian Sarabil Formation Sakmarskiy Orenburgskaya Region S Urals Russia

Triposphaera permica irregularis


Triposphaera permica permica


Triposphaera tetrasphaera






er

Stat

usTy




Triposphaera (330) continuationTriposphaera tribrachiata E ndash ndash Kozur amp Mostler 1989 191 Lower Permian Sakmarian Sarabil Formation Sakmarskiy


Triposphaera uralica E ndash ndash Kozur amp Mostler 1989 192 Lower Permian Sakmarian Sarabil Formation Sakmarskiy Orenburgskaya Region S Urals Russia

Triposphaera woni E ndash ndash Kozur amp Mostler 1989 192 Lower Permian Sakmarian Sarabil Formation Sakmarskiy Orenburgskaya Region S Urals Russia

Triposphaera dumitricai E ndash ndash Kozur amp Mostler 1989 190 Lower Permian Sakmarian Sarabil Formation Sakmarskiy Orenburgskaya Region S Urals Russia

TrissocircusTrissocircus primaevus ndash nd ndash Ruumlst 1892 177 Upper Permian ndash Sosio Valley Area Western Sicily

ItalyTrissocircus quadratus ndash nd ndash Ruumlst 1892 177 Lower Devonian ndash Southern Urals Russia

TrochodiscusTrochodiscus convexus E ndash ndash Ruumlst 1892 164 Lower


Trochodiscus diversispinus


Trochodiscus longispinus ndash nd ndash Ruumlst 1892 165 Lower Carboniferous


Trochodiscus nicholsoni E ndash ndash Ruumlst 1892 164 Lower Carboniferous


Trochodiscus planatus E ndash ndash Hinde 1899a 53 Middle Devonian Givetian Yarramie Formation Tamworth Belt northern NSW Australia

Trochodiscus proavus ndash nd ndash Ruumlst 1892 164 Lower Carboniferous


Trochodiscus recurvispinus


Trochodiscus serrula ndash nd ndash Ruumlst 1892 164 Lower Carboniferous


Trochodiscus dubius ndash nd ndash Hinde 1899b 218 Middle Devonian Givetian Chypons Farm Mullion Parrish Cornwall England UK

Tuscaritellum (233)Tuscaritellum imitatum N ndash ndash Deflandre 1972a 3536 Lower


Tuscaritellum imitatum N ndash Ts Deflandre 1972b 16 Lower Carboniferous


Uberinterna (128)Uberinterna

virgispinosumE ndash T Sashida amp Tonishi 1988 531 Upper Permian Changhsingian Ohirayama Unit South Chichibu


Ulcundia (331)Ulcundia incomperta ndash nd Td Nazarov 1974b 111 Middle



Varispiculum (59)Varispiculum

ectospiculatumAr ndash ndash Won amp Iams 2015a 17 Lower Ordovician Floian Cow Head Group Western

Newfoundland CanadaVarispiculum delicatum Ar ndash ndash Won amp Iams 2015a 18 Lower Ordovician Floian Cow Head Group Western

Newfoundland CanadaVarispiculum radiatum Ar ndash ndash Won amp Iams 2015a 19 Lower Ordovician Floian Cow Head Group Western

Newfoundland Canada

Wangia (332)Wangia biturbinata ndash nn ndash Feng 1992 57 Upper Permian Changhsingian Muyinhe Formation Nanpan

Lancang County Yunnan ChinaWangia biturbinata ndash nd Td Feng amp Liu 1993 544 Upper Permian Changhsingian Muyinhe Formation Nanpan

Lancang County Yunnan ChinaWangia elegans ndash nn ndash Feng 1992 57 Upper Permian Changhsingian Muyinhe Formation Nanpan

Lancang County Yunnan ChinaWangia elegans ndash nd ndash Feng amp Liu 1993 544 Upper Permian Changhsingian Muyinhe Formation Nanpan

Lancang County Yunnan ChinaWangia kangwuensis ndash nd ndash Feng Fang Zhang amp Huang

1998 240Upper Permian Changhsingian Papai Formation of Changning-




Aitchison J C et al


er

Stat

usTy




Westernbrookia (60)Westernbrookia cancella Ar ndash ndash Won Iams amp Reed 2007 521 Lower Ordovician lower-middle


Newfoundland CanadaWesternbrookia diversa Ar ndash T Won Iams amp Reed 2007 519 Lower Ordovician lower-middle


Newfoundland CanadaWesternbrookia incohota Ar ndash ndash Won amp Iams 2013 23 Lower Ordovician Floian Cow Head Group Western

Newfoundland CanadaWesternbrookia ovata Ar ndash ndash Won Iams amp Reed 2007 519 Lower Ordovician lower-middle


Newfoundland Canada

Wuyia (130)Wuyia dongpanica E ndash T Feng in Feng et al 2007 33 Upper Permian Changhsingian Dalong Formation Dongpan


Xiphachistrella (28)Xiphachistrella acipensis Al nn ndash Deflandre 1960 216 Lower


Xiphachistrella acipensis Al nn ndash Deflandre 1973a 292 Lower Carboniferous


Xiphachistrella acipensis Al ndash Ts Deflandre 1973b 500 Lower Carboniferous


Xiphocabrium (41)Xiphocabrium rigidum Al nn ndash Deflandre 1973a 293 Lower


Xiphocabrium rigidum Al ndash T Deflandre 1973b 500 Lower Carboniferous


Xiphocabrium aprelevkensis

Al nn ndash Afanasieva 1997a 221 Upper Devonian middle Frasnian


Xiphocladiella (42)Xiphocladiella cuja Al nn ndash Deflandre 1960 216 Lower


Xiphocladiella cuja Al nn ndash Deflandre 1973a 291 Lower Carboniferous


Xiphocladiella cuja Al ndash T Deflandre 1973b 499 Lower Carboniferous


XiphosphaeraXiphosphaera

brachyacanthandash nd ndash Ruedemann amp Wilson 1936



Xiphosphaera macracantha



Xiphosphaera macrostyla ndash nd ndash Ruumlst 1892 141 Lower Silurian ndash Cabriegraveres Heacuterault FranceXiphosphaera minax E ndash ndash Hinde 1899a 45 Middle Devonian Givetian Yarramie Formation Tamworth

Belt northern NSW AustraliaXiphosphaera parva ndash nd ndash Ruedemann amp Wilson 1936



Xiphosphaera spinifera ndash nd ndash Pulfrey 1932 195 Lower Carboniferous


XiphostylusXiphostylus ardea ndash nd ndash Ruumlst 1892 142 Lower


Xiphostylus asper ndash nd ndash Ruumlst 1892 142 Lower Silurian ndash Langenstriegis GermanyXiphostylus inclinatus ndash nd ndash Aberdeen 1940 135 Upper Devonian Famennian Caballos Formation Marathon

Basin Texas USA

YujingellaYujingella triacantha S ndash T Feng in Feng et al 2006a 33 Late Permian Changhsingian Dalong Formation Dongpan






er

Stat

usTy




Zadrappolus (267)Zadrappolus lunaris S ndash ndash Noble 1994 32 Upper Silurian Pridoli Caballos Novaculite Formation


Zadrappolus spinosus S ndash ndash Furutani 1990 37 Upper Silurian Gorstian Hitoegane Formation Hida-gaien Belt E of Hitoegane Fukuji area Takayama City Japan

Zadrappolus tenuis S ndash ndash Furutani 1990 36 Upper Silurian Pridoli Yoshiki Formation Hida-gaien Belt Ichinotani Valley Fukuji area Takayama City Gifu Japan

Zadrappolus yoshikiensis S ndash T Furutani 1990 35 Upper Silurian Pridoli Yoshiki Formation Hida-gaien Belt Ichinotani Valley Fukuji area Takayama City Gifu Japan

Zadrappolus hitoeganensis

S ndash ndash Furutani 1990 37 Upper Silurian Ludfordian Hitoegane Formation Hida-gaien Belt E of Hitoegane Fukuji area Takayama City Japan

Zadrappolus nudus S ndash ndash Kurihara 2007 230 Lower Devonian Emsian Hitoegane Formation Hitoegane Forest Road Hitoegane Takayama Japan

Zhuangodiscus (275)Zhuangodiscus

biacanthusS ndash ndash Wu amp Feng in

Wu et al 2010 887Upper Permian Changhsingian Dalong Formation Dongpan


Zhuangodiscus orthacanthus

S ndash T Wu amp Feng in Wu et al 2010 887


Zhuangodiscus pentacanthus

S ndash ndash Wu amp Feng in Wu et al 2010 889


ZonodiscusZonodiscus dentatus ndash nd ndash Ruumlst 1892 162 Lower Silurian ndash Cabriegraveres Heacuterault FranceZonodiscus macrozona ndash nd ndash Ruumlst 1892 162 Lower


ZygocircusZygocircus priscus ndash nd ndash Ruumlst 1892 176 Lower


Zygocircus simplicissimus


ZygosterhanusZygosterhanus

quadrupesndash nd ndash Ruumlst 1892 177 Upper Permian ndash Sosio Valley Area Western Sicily

Italy



Aitchison J C et al

OBJECTIVES

Following the publication of the Inventory of Mesozoic radiolarian species (OrsquoDogherty 2009) there was general agreement of participants in workshops held by the Paleozoic Working Group that it would be useful for the community of researchers if a similar document could be produced for Paleozoic radiolarians

Here we present a comprehensive list of all the Paleozoic radiolarian species of which we are aware that have been described between the late 1800s and early 2016 The inten-tion is to provide in a single file the names of all Paleozoic species that have been published All names listed are the original names given in the original publication and do not reflect any subsequent taxonomic revision or evalua-tion as to their validity and assignment at higher taxonomic levels The list records 2323 taxa described originally as either new species or subspecies Although every attempt has been made to be as complete as possible it is perhaps inevitable that some taxa may be missing The purpose of this list is to provide information on 1) the original Linnean combination used when a species was erected as new in Paleozoic material 2) the author(s) and page(s) on which it is described 3) the age of the holotype or of the name-bearing type as originally published (note in some cases there may have been considerable revision of the original age determination) and 4) the order to which the species or sub-species belongs (the numbers of which are indicated in parentheses with each order) Al Albaillellaria (363) Ar Archaeospicularia (154) E Entactinaria (726) L Latentifistularia (194) N possible Nassellaria (111) S Spumellaria (207) IS incertae sedis within orders (72) incertae sedis in both order and family (35) with 368 names that can be regarded as nomina dubium (nd) and 158 names introduced as nomina nuda (nn) Forty-four taxa are clearly not radiolarians (nr) with 15 of these Calcarea and five others that can be assigned as sponge spicules (ss) also identified Eleven names are invalid Type species (T valid type species Td types that are nomina dubia Th types that are homonyms Ti type species that are invalid names and Ts types that are synonyms) are indicated

COMPILATION OF SPECIES NAMES

The list of taxa has been compiled following a detailed search of gt1250 publications dealing with Paleozoic radiolarians of which 259 introduce new taxa Synonyms and species descriptions are not included because this is beyond the scope of the project and would require several years of work by many research groups The general agreement of participants at the workshops was that it would be useful to take this opportunity to make a step towards address-ing species-level taxonomy of species once the revision of genera has been fully achieved Such a work would have the additional benefit of providing a better-defined starting point for future taxonomists

FUTURE WORK

Objective evaluation of each species with respect to its taxo-nomic status and the subjective assignment to valid genera is the next step towards attaining a full proper standard of taxonomy and stratigraphy for Paleozoic radiolarians Evalu-ation of each species unless one is personally familiar with it is a time-consuming task depending on the availability of literature and given that there are 2323 species names to be evaluated is well beyond the scope of this particular work which is meant to focus on the genera However we empha-size that the species treatment is a logical future task of the Paleozoic Working Group It is our intention herein to simply provide this list as a useful starting point for any such study

AcknowledgementsWe thank all our fellow radiolarian researchers who have in various ways contributed to the accumulation of the body of data that has allowed development of this biozonation JA thanks Sarah Kachovich for her help in editing this list and he also acknowl-edges financial support towards investigation of Early Paleozoic radiolarian evolution in the form of a grant from the Australian Research Council (ARC DP 1501013325) We also gratefully acknowledge reviews by Taniel Danelian and Paula Noble

REFERENCES

Aberdeen e 1940 mdash Radiolarian fauna of the Caballos Formation Marathon Basin Texas Journal of Paleontology 14 (2) 127-139 httpwwwjstororgstable1298566

AfAnAsievA M s 1986 mdash Radiolarians of the family Pylenton-emidae Paleontological Journal 3 22-34

AfAnAsievA M s 1993 mdash New data on the Early Paleozoic radio-larian genus Caspiaza Paleontological Journal 4 115-118

AfAnAsievA M s 1997a mdash Early Frasnian Radiolarians and their biostrati-graphical significance Doklady Akademii Nauk 355 (2) 217-222

AfAnAsievA M s 1997b mdash Moscovistella victoralis the victorious star of Moscow Izvestija Vysshikh Uchebnykh Zavedeniy Geologiya i Razvedka 5 38-41

AfAnAsievA M s 2000a mdash Atlas of Paleozoic Radiolaria of the Rus-sian Platform Scientific World Moscow 477 p

AfAnAsievA M s 2000b mdash New radiolarians of the superfam-ily Entactinoidea from the Upper Devonian of Timan-Pechora Province Russia Paleontological Journal 34 (2) 131-146

AfAnAsievA M s 2000c mdash New radiolarians of the orders Aculearia and Sphaerellaria from the Upper Devonian of the Timan-Pechora Province (Russia) Paleontological Journal 34 (4) 359-376

AfAnAsievA M 2011 mdash Revision of the genus Entactinosphaera Foreman 1963 and the new genus Retientactinosphaera gen nov (Paleozoic radiolaria) Paleontological Journal 45 (2) 117-129 httpsdoiorg101134S003103011102002X

AfAnAsievA M s amp AMon e 2008 mdash New radiolarians from the Devonian of the southern Ural Mountains 1 Early-Middle Devonian (Late Emsian-Early Eifelian) Paleontological Journal 42 (2) 451-467

AfAnAsievA M s amp AMon e o 2009a mdash New Radiolarians from the Devonian of the Southern Ural Mountains 2 MiddlendashLate Devonian Paleontological Journal 43 (1) 34-49

AfAnAsievA M s amp AMon e o 2009b mdash Evolution of Devonian biodiversity of radiolarians with two porous spheres and one main spine Paleontological Journal 43 (5) 483-498 httpsdoiorg101134S0031030109050037









































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Aitchison J C et al










































































Aitchison J C et al











































































Aitchison J C et al













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E nn Th Ts








ActinommaActinomma


Elbingerode Germany

APPENDIX



Aitchison J C et al


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Stat

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(OampF)nc T











Lower Carboniferous





















Lower Carboniferous







Upper Carboniferous



Lower Carboniferous









er

Stat

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Upper Carboniferous






Lower Carboniferous





Lower Carboniferous



Lower Carboniferous



Lower Carboniferous



Lower Carboniferous








upper Tournaisian





Upper Carboniferous







Lower Carboniferous





Lower Carboniferous






Lower Carboniferous







Lower Carboniferous





Upper Carboniferous




Aitchison J C et al


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Stat

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Lower Carboniferous









Lower Carboniferous




















Lower Carboniferous

















Lower Carboniferous













Lower Carboniferous








er

Stat

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Germany




Lower Carboniferous



upper Tournaisian


























Upper Carboniferous











ItalyAmphibrachium


Elbingerode Germany



Aitchison J C et al


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Stat

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Elbingerode Germany









































er

Stat

usTy

























































Newfoundland Canada





Aitchison J C et al


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Stat

usTy












upper Tournaisian





















upper Tournaisian



upper Tournaisian


























upper Tournaisian










er

Stat

usTy


































1986 124Upper






Upper Carboniferous


















Newfoundland Canada



Aitchison J C et al


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Stat

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Newfoundland Canada












upper Tournaisian



Middle Ordovician





Upper Carboniferous




Upper Carboniferous







upper Tournaisian









upper Tournaisian















er

Stat

usTy
















Middle Ordovician







Basin Texas USA








Italy

















Frankenwald Germany










Newfoundland Canada



Aitchison J C et al


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Stat

usTy


























































er

Stat

usTy









1980 64Middle





Middle Ordovician






mindyallicanr Calc-






































Aitchison J C et al


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Stat

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Upper Carboniferous







Newfoundland Canada











er

Stat

usTy






















Australia


Elbingerode Germany






















Newfoundland Canada



ItalyCecryphalium






Aitchison J C et al


er

Stat

usTy







































Basin Texas USA



Frankenwald Germany











er

Stat

usTy















103Middle





Middle Ordovician















































Lower Carboniferous





Aitchison J C et al


er

Stat

usTy























upper Tournaisian





















Lower Carboniferous






















er

Stat

usTy































upper Tournaisian






















Aitchison J C et al


er

Stat

usTy


















upper Tournaisian












Middle Ordovician

































Lower Carboniferous








er

Stat

usTy













1986 70Upper




1986 71Upper



1986 71Upper































Aitchison J C et al


er

Stat

usTy























1996 246Lower




Lower Carboniferous



Lower Carboniferous



Lower Carboniferous




Italy


USACorythoecia


1986 119Upper


Urals Russia





USA





er

Stat

usTy






Newfoundland Canada


Italy

CromyommaCromyomma





















Frankenwald Germany


1958 969Lower










Newfoundland Canada










Aitchison J C et al


er

Stat

usTy

























































Ts Th











er

Stat

usTy














Italy




Elbingerode Germany


Italy


















Italy










Aitchison J C et al


er

Stat

usTy






























Cornwall England UK




























er

Stat

usTy





































EchinommaEchinomma


Italy














Aitchison J C et al


er

Stat

usTy














Italy













Upper Carboniferous













Middle Ordovician





Middle Ordovician









er

Stat

usTy





236Upper







Upper Carboniferous




Upper Carboniferous




Upper Carboniferous

















Middle Ordovician















Aitchison J C et al


er

Stat

usTy




















































er

Stat

usTy






















Middle Ordovician






upper Tournaisian

















upper Tournaisian









Aitchison J C et al


er

Stat

usTy



















Upper Carboniferous





















Upper Carboniferous
























er

Stat

usTy



































































Middle Ordovician






Aitchison J C et al


er

Stat

usTy


























Middle Ordovician



















Izmir Region Turkey



















er

Stat

usTy




















IS(OampF)




































Aitchison J C et al


er

Stat

usTy
















Prefecture Japan


































er

Stat

usTy
















Frankenwald Germany



Newfoundland Canada





















Newfoundland Canada










Aitchison J C et al


er

Stat

usTy















Italy




















Middle Ordovician




















er

Stat

usTy








102Middle




Middle Ordovician
























Columbia Canada









Upper Carboniferous




Upper Carboniferous




Aitchison J C et al


er

Stat

usTy










China







Middle Ordovician

upper Darriwilian

































Middle Ordovician

upper Darriwilian









er

Stat

usTy























Upper Carboniferous


















Aitchison J C et al


er

Stat

usTy





















































er

Stat

usTy












Carpathians Hungary



Italy


Basin Texas USA




























Aitchison J C et al


er

Stat

usTy


























Barmen Germany













Middle Ordovician

upper Darriwilian
















lower Darriwilian






er

Stat

usTy

























France

















Aitchison J C et al


er

Stat

usTy






































1976 415Middle







er

Stat

usTy





























1986 82Upper



1986 79Upper



1986 82Upper



240Upper




Upper Carboniferous





Upper Carboniferous





Upper Carboniferous



1986 87Upper



1986 88Upper




1986 89Upper




1986 90Upper


Urals Russia



Aitchison J C et al


er

Stat

usTy














Upper Carboniferous












Upper Carboniferous




Upper Carboniferous
















Italy





er

Stat

usTy







ItalyLithocampe























Germany


Italy



















Aitchison J C et al


er

Stat

usTy














2008 53Lower













River Kazakhstan




























er

Stat

usTy



















































Aitchison J C et al


er

Stat

usTy
















































er

Stat

usTy


















Newfoundland Canada


Newfoundland Canada














Newfoundland Canada























Aitchison J C et al


er

Stat

usTy
































Lower Carboniferous








China





er

Stat

usTy











































anticaIS




IS(OampF)



IS(OampF)



IS(OampF)



IS(OampF)



IS(OampF)




Aitchison J C et al


er

Stat

usTy





Brittany France

















































er

Stat

usTy



























































Aitchison J C et al


er

Stat

usTy




























































er

Stat

usTy






























Newfoundland Canada


antiquusIS




IS(OampF)




IS(OampF)



IS(OampF)



IS(OampF)




IS(OampF)
















Aitchison J C et al


er

Stat

usTy





















Frankenwald Germany



France




































France





er

Stat

usTy







Newfoundland Canada































Newfoundland Canada



















Aitchison J C et al


er

Stat

usTy






















Upper Carboniferous






Carpathians Hungary














Russia










er

Stat

usTy





Basin Texas USA








































1986 108Upper









Aitchison J C et al


er

Stat

usTy





























Upper Carboniferous



Upper Carboniferous








Upper Carboniferous



Middle Ordovician














er

Stat

usTy






Ordovicianupper



















































Aitchison J C et al


er

Stat

usTy




















Upper Carboniferous












Middle Ordovician


















ChinaPraesaturnalis




China





er

Stat

usTy





fusiformaIS




IS(OampF)












Italy






















Lower Carboniferous



















Aitchison J C et al


er

Stat

usTy




















































Newfoundland Canada








Newfoundland Canada






Newfoundland Canada





er

Stat

usTy


































Upper Carboniferous















Upper Carboniferous






Upper Carboniferous




Aitchison J C et al


er

Stat

usTy









Upper Carboniferous















































er

Stat

usTy





































1996 222Lower















Middle Ordovician



Middle Ordovician






upper Tournaisian






Aitchison J C et al


er

Stat

usTy







918Middle







Middle Ordovician















































er

Stat

usTy




















































Aitchison J C et al


er

Stat

usTy









Newfoundland Canada




































clavataIS





IS(OampF)







er

Stat

usTy









Italy





















Elbingerode Germany




















Aitchison J C et al


er

Stat

usTy

























































er

Stat

usTy














Newfoundland Canada









Upper Carboniferous

























Aitchison J C et al


er

Stat

usTy










































UK





er

Stat

usTy





























Frankenwald Germany




















Aitchison J C et al


er

Stat

usTy




























Italy





Italy

























er

Stat

usTy

























Upper Carboniferous




















Elbingerode Germany



Aitchison J C et al


er

Stat

usTy


































Italy








Scotland UK













E ndash T Th







er

Stat

usTy






























ItalySpongotrochus












Kaeo Thailand


ItalyStauracontium







Italy








Scotland UK



Aitchison J C et al


er

Stat

usTy









USA

















































er

Stat

usTy

























Italy






















Lower Carboniferous






Aitchison J C et al


er

Stat

usTy































Elbingerode Germany















Italy



Scotland UK





er

Stat

usTy








Newfoundland Canada










Middle Ordovician





Middle Ordovician






Middle Ordovician

upper Darriwilian










239Upper









Upper Carboniferous





Aitchison J C et al


er

Stat

usTy





1986 72Upper










Upper Carboniferous





Italy














Germany




Germany









Upper Carboniferous




Upper Carboniferous

















Upper Carboniferous






er

Stat

usTy





USATetrentactinia
















USA








Italy


Italy



UKTheodiscus








Italy








Aitchison J C et al


er

Stat

usTy





1986 97Upper




1986 97Upper



1986 100Upper







Upper Carboniferous













Upper Carboniferous





Upper Carboniferous

























250Lower







er

Stat

usTy









upper Tournaisian














































Aitchison J C et al


er

Stat

usTy



















Lower Carboniferous

lower Tournaisian











Ordovicianupper


region Kazakhstan























er

Stat

usTy









































Newfoundland Canada










Aitchison J C et al


er

Stat

usTy











Newfoundland Canada






































Basin Texas USA







er

Stat

usTy






























Italy










































Aitchison J C et al









































































Aitchison J C et al










































































Aitchison J C et al











































































Aitchison J C et al













er

Stat

usTy














































E nn Th Ts








ActinommaActinomma


Elbingerode Germany

APPENDIX



Aitchison J C et al


er

Stat

usTy





(OampF)nc T











Lower Carboniferous





















Lower Carboniferous







Upper Carboniferous



Lower Carboniferous









er

Stat

usTy








Upper Carboniferous






Lower Carboniferous





Lower Carboniferous



Lower Carboniferous



Lower Carboniferous



Lower Carboniferous








upper Tournaisian





Upper Carboniferous







Lower Carboniferous





Lower Carboniferous






Lower Carboniferous







Lower Carboniferous





Upper Carboniferous




Aitchison J C et al


er

Stat

usTy








Lower Carboniferous









Lower Carboniferous




















Lower Carboniferous

















Lower Carboniferous













Lower Carboniferous








er

Stat

usTy








Germany




Lower Carboniferous



upper Tournaisian


























Upper Carboniferous











ItalyAmphibrachium


Elbingerode Germany



Aitchison J C et al


er

Stat

usTy










Elbingerode Germany









































er

Stat

usTy

























































Newfoundland Canada





Aitchison J C et al


er

Stat

usTy












upper Tournaisian





















upper Tournaisian



upper Tournaisian


























upper Tournaisian










er

Stat

usTy


































1986 124Upper






Upper Carboniferous


















Newfoundland Canada



Aitchison J C et al


er

Stat

usTy








Newfoundland Canada












upper Tournaisian



Middle Ordovician





Upper Carboniferous




Upper Carboniferous







upper Tournaisian









upper Tournaisian















er

Stat

usTy
















Middle Ordovician







Basin Texas USA








Italy

















Frankenwald Germany










Newfoundland Canada



Aitchison J C et al


er

Stat

usTy


























































er

Stat

usTy









1980 64Middle





Middle Ordovician






mindyallicanr Calc-






































Aitchison J C et al


er

Stat

usTy




































Upper Carboniferous







Newfoundland Canada











er

Stat

usTy






















Australia


Elbingerode Germany






















Newfoundland Canada



ItalyCecryphalium






Aitchison J C et al


er

Stat

usTy







































Basin Texas USA



Frankenwald Germany











er

Stat

usTy















103Middle





Middle Ordovician















































Lower Carboniferous





Aitchison J C et al


er

Stat

usTy























upper Tournaisian





















Lower Carboniferous






















er

Stat

usTy































upper Tournaisian






















Aitchison J C et al


er

Stat

usTy


















upper Tournaisian












Middle Ordovician

































Lower Carboniferous








er

Stat

usTy













1986 70Upper




1986 71Upper



1986 71Upper































Aitchison J C et al


er

Stat

usTy























1996 246Lower




Lower Carboniferous



Lower Carboniferous



Lower Carboniferous




Italy


USACorythoecia


1986 119Upper


Urals Russia





USA





er

Stat

usTy






Newfoundland Canada


Italy

CromyommaCromyomma





















Frankenwald Germany


1958 969Lower










Newfoundland Canada










Aitchison J C et al


er

Stat

usTy

























































Ts Th











er

Stat

usTy














Italy




Elbingerode Germany


Italy


















Italy










Aitchison J C et al


er

Stat

usTy






























Cornwall England UK




























er

Stat

usTy





































EchinommaEchinomma


Italy














Aitchison J C et al


er

Stat

usTy














Italy













Upper Carboniferous













Middle Ordovician





Middle Ordovician









er

Stat

usTy





236Upper







Upper Carboniferous




Upper Carboniferous




Upper Carboniferous

















Middle Ordovician















Aitchison J C et al


er

Stat

usTy




















































er

Stat

usTy






















Middle Ordovician






upper Tournaisian

















upper Tournaisian









Aitchison J C et al


er

Stat

usTy



















Upper Carboniferous





















Upper Carboniferous
























er

Stat

usTy



































































Middle Ordovician






Aitchison J C et al


er

Stat

usTy


























Middle Ordovician



















Izmir Region Turkey



















er

Stat

usTy




















IS(OampF)




































Aitchison J C et al


er

Stat

usTy
















Prefecture Japan


































er

Stat

usTy
















Frankenwald Germany



Newfoundland Canada





















Newfoundland Canada










Aitchison J C et al


er

Stat

usTy















Italy




















Middle Ordovician




















er

Stat

usTy








102Middle




Middle Ordovician
























Columbia Canada









Upper Carboniferous




Upper Carboniferous




Aitchison J C et al


er

Stat

usTy










China







Middle Ordovician

upper Darriwilian

































Middle Ordovician

upper Darriwilian









er

Stat

usTy























Upper Carboniferous


















Aitchison J C et al


er

Stat

usTy





















































er

Stat

usTy












Carpathians Hungary



Italy


Basin Texas USA




























Aitchison J C et al


er

Stat

usTy


























Barmen Germany













Middle Ordovician

upper Darriwilian
















lower Darriwilian






er

Stat

usTy

























France

















Aitchison J C et al


er

Stat

usTy






































1976 415Middle







er

Stat

usTy





























1986 82Upper



1986 79Upper



1986 82Upper



240Upper




Upper Carboniferous





Upper Carboniferous





Upper Carboniferous



1986 87Upper



1986 88Upper




1986 89Upper




1986 90Upper


Urals Russia



Aitchison J C et al


er

Stat

usTy














Upper Carboniferous












Upper Carboniferous




Upper Carboniferous
















Italy





er

Stat

usTy







ItalyLithocampe























Germany


Italy



















Aitchison J C et al


er

Stat

usTy














2008 53Lower













River Kazakhstan




























er

Stat

usTy



















































Aitchison J C et al


er

Stat

usTy
















































er

Stat

usTy


















Newfoundland Canada


Newfoundland Canada














Newfoundland Canada























Aitchison J C et al


er

Stat

usTy
































Lower Carboniferous








China





er

Stat

usTy











































anticaIS




IS(OampF)



IS(OampF)



IS(OampF)



IS(OampF)



IS(OampF)




Aitchison J C et al


er

Stat

usTy





Brittany France

















































er

Stat

usTy



























































Aitchison J C et al


er

Stat

usTy




























































er

Stat

usTy






























Newfoundland Canada


antiquusIS




IS(OampF)




IS(OampF)



IS(OampF)



IS(OampF)




IS(OampF)
















Aitchison J C et al


er

Stat

usTy





















Frankenwald Germany



France




































France





er

Stat

usTy







Newfoundland Canada































Newfoundland Canada



















Aitchison J C et al


er

Stat

usTy






















Upper Carboniferous






Carpathians Hungary














Russia










er

Stat

usTy





Basin Texas USA








































1986 108Upper









Aitchison J C et al


er

Stat

usTy





























Upper Carboniferous



Upper Carboniferous








Upper Carboniferous



Middle Ordovician














er

Stat

usTy






Ordovicianupper



















































Aitchison J C et al


er

Stat

usTy




















Upper Carboniferous












Middle Ordovician


















ChinaPraesaturnalis




China





er

Stat

usTy





fusiformaIS




IS(OampF)












Italy






















Lower Carboniferous



















Aitchison J C et al


er

Stat

usTy




















































Newfoundland Canada








Newfoundland Canada






Newfoundland Canada





er

Stat

usTy


































Upper Carboniferous















Upper Carboniferous






Upper Carboniferous




Aitchison J C et al


er

Stat

usTy









Upper Carboniferous















































er

Stat

usTy





































1996 222Lower















Middle Ordovician



Middle Ordovician






upper Tournaisian






Aitchison J C et al


er

Stat

usTy







918Middle







Middle Ordovician















































er

Stat

usTy




















































Aitchison J C et al


er

Stat

usTy









Newfoundland Canada




































clavataIS





IS(OampF)







er

Stat

usTy









Italy





















Elbingerode Germany




















Aitchison J C et al


er

Stat

usTy

























































er

Stat

usTy














Newfoundland Canada









Upper Carboniferous

























Aitchison J C et al


er

Stat

usTy










































UK





er

Stat

usTy





























Frankenwald Germany




















Aitchison J C et al


er

Stat

usTy




























Italy





Italy

























er

Stat

usTy

























Upper Carboniferous




















Elbingerode Germany



Aitchison J C et al


er

Stat

usTy


































Italy








Scotland UK













E ndash T Th







er

Stat

usTy






























ItalySpongotrochus












Kaeo Thailand


ItalyStauracontium







Italy








Scotland UK



Aitchison J C et al


er

Stat

usTy









USA

















































er

Stat

usTy

























Italy






















Lower Carboniferous






Aitchison J C et al


er

Stat

usTy































Elbingerode Germany















Italy



Scotland UK





er

Stat

usTy








Newfoundland Canada










Middle Ordovician





Middle Ordovician






Middle Ordovician

upper Darriwilian










239Upper









Upper Carboniferous





Aitchison J C et al


er

Stat

usTy





1986 72Upper










Upper Carboniferous





Italy














Germany




Germany









Upper Carboniferous




Upper Carboniferous

















Upper Carboniferous






er

Stat

usTy





USATetrentactinia
















USA








Italy


Italy



UKTheodiscus








Italy








Aitchison J C et al


er

Stat

usTy





1986 97Upper




1986 97Upper



1986 100Upper







Upper Carboniferous













Upper Carboniferous





Upper Carboniferous

























250Lower







er

Stat

usTy









upper Tournaisian














































Aitchison J C et al


er

Stat

usTy



















Lower Carboniferous

lower Tournaisian











Ordovicianupper


region Kazakhstan























er

Stat

usTy









































Newfoundland Canada










Aitchison J C et al


er

Stat

usTy











Newfoundland Canada






































Basin Texas USA







er

Stat

usTy






























Italy



Aitchison J C et al









































































Aitchison J C et al










































































Aitchison J C et al











































































Aitchison J C et al













er

Stat

usTy














































E nn Th Ts








ActinommaActinomma


Elbingerode Germany

APPENDIX



Aitchison J C et al


er

Stat

usTy





(OampF)nc T











Lower Carboniferous





















Lower Carboniferous







Upper Carboniferous



Lower Carboniferous









er

Stat

usTy








Upper Carboniferous






Lower Carboniferous





Lower Carboniferous



Lower Carboniferous



Lower Carboniferous



Lower Carboniferous








upper Tournaisian





Upper Carboniferous







Lower Carboniferous





Lower Carboniferous






Lower Carboniferous







Lower Carboniferous





Upper Carboniferous




Aitchison J C et al


er

Stat

usTy








Lower Carboniferous









Lower Carboniferous




















Lower Carboniferous

















Lower Carboniferous













Lower Carboniferous








er

Stat

usTy








Germany




Lower Carboniferous



upper Tournaisian


























Upper Carboniferous











ItalyAmphibrachium


Elbingerode Germany



Aitchison J C et al


er

Stat

usTy










Elbingerode Germany









































er

Stat

usTy

























































Newfoundland Canada





Aitchison J C et al


er

Stat

usTy












upper Tournaisian





















upper Tournaisian



upper Tournaisian


























upper Tournaisian










er

Stat

usTy


































1986 124Upper






Upper Carboniferous


















Newfoundland Canada



Aitchison J C et al


er

Stat

usTy








Newfoundland Canada












upper Tournaisian



Middle Ordovician





Upper Carboniferous




Upper Carboniferous







upper Tournaisian









upper Tournaisian















er

Stat

usTy
















Middle Ordovician







Basin Texas USA








Italy

















Frankenwald Germany










Newfoundland Canada



Aitchison J C et al


er

Stat

usTy


























































er

Stat

usTy









1980 64Middle





Middle Ordovician






mindyallicanr Calc-






































Aitchison J C et al


er

Stat

usTy




































Upper Carboniferous







Newfoundland Canada











er

Stat

usTy






















Australia


Elbingerode Germany






















Newfoundland Canada



ItalyCecryphalium






Aitchison J C et al


er

Stat

usTy







































Basin Texas USA



Frankenwald Germany











er

Stat

usTy















103Middle





Middle Ordovician















































Lower Carboniferous





Aitchison J C et al


er

Stat

usTy























upper Tournaisian





















Lower Carboniferous






















er

Stat

usTy































upper Tournaisian






















Aitchison J C et al


er

Stat

usTy


















upper Tournaisian












Middle Ordovician

































Lower Carboniferous








er

Stat

usTy













1986 70Upper




1986 71Upper



1986 71Upper































Aitchison J C et al


er

Stat

usTy























1996 246Lower




Lower Carboniferous



Lower Carboniferous



Lower Carboniferous




Italy


USACorythoecia


1986 119Upper


Urals Russia





USA





er

Stat

usTy






Newfoundland Canada


Italy

CromyommaCromyomma





















Frankenwald Germany


1958 969Lower










Newfoundland Canada










Aitchison J C et al


er

Stat

usTy

























































Ts Th











er

Stat

usTy














Italy




Elbingerode Germany


Italy


















Italy










Aitchison J C et al


er

Stat

usTy






























Cornwall England UK




























er

Stat

usTy





































EchinommaEchinomma


Italy














Aitchison J C et al


er

Stat

usTy














Italy













Upper Carboniferous













Middle Ordovician





Middle Ordovician









er

Stat

usTy





236Upper







Upper Carboniferous




Upper Carboniferous




Upper Carboniferous

















Middle Ordovician















Aitchison J C et al


er

Stat

usTy




















































er

Stat

usTy






















Middle Ordovician






upper Tournaisian

















upper Tournaisian









Aitchison J C et al


er

Stat

usTy



















Upper Carboniferous





















Upper Carboniferous
























er

Stat

usTy



































































Middle Ordovician






Aitchison J C et al


er

Stat

usTy


























Middle Ordovician



















Izmir Region Turkey



















er

Stat

usTy




















IS(OampF)




































Aitchison J C et al


er

Stat

usTy
















Prefecture Japan


































er

Stat

usTy
















Frankenwald Germany



Newfoundland Canada





















Newfoundland Canada










Aitchison J C et al


er

Stat

usTy















Italy




















Middle Ordovician




















er

Stat

usTy








102Middle




Middle Ordovician
























Columbia Canada









Upper Carboniferous




Upper Carboniferous




Aitchison J C et al


er

Stat

usTy










China







Middle Ordovician

upper Darriwilian

































Middle Ordovician

upper Darriwilian









er

Stat

usTy























Upper Carboniferous


















Aitchison J C et al


er

Stat

usTy





















































er

Stat

usTy












Carpathians Hungary



Italy


Basin Texas USA




























Aitchison J C et al


er

Stat

usTy


























Barmen Germany













Middle Ordovician

upper Darriwilian
















lower Darriwilian






er

Stat

usTy

























France

















Aitchison J C et al


er

Stat

usTy






































1976 415Middle







er

Stat

usTy





























1986 82Upper



1986 79Upper



1986 82Upper



240Upper




Upper Carboniferous





Upper Carboniferous





Upper Carboniferous



1986 87Upper



1986 88Upper




1986 89Upper




1986 90Upper


Urals Russia



Aitchison J C et al


er

Stat

usTy














Upper Carboniferous












Upper Carboniferous




Upper Carboniferous
















Italy





er

Stat

usTy







ItalyLithocampe























Germany


Italy



















Aitchison J C et al


er

Stat

usTy














2008 53Lower













River Kazakhstan




























er

Stat

usTy



















































Aitchison J C et al


er

Stat

usTy
















































er

Stat

usTy


















Newfoundland Canada


Newfoundland Canada














Newfoundland Canada























Aitchison J C et al


er

Stat

usTy
































Lower Carboniferous








China





er

Stat

usTy











































anticaIS




IS(OampF)



IS(OampF)



IS(OampF)



IS(OampF)



IS(OampF)




Aitchison J C et al


er

Stat

usTy





Brittany France

















































er

Stat

usTy



























































Aitchison J C et al


er

Stat

usTy




























































er

Stat

usTy






























Newfoundland Canada


antiquusIS




IS(OampF)




IS(OampF)



IS(OampF)



IS(OampF)




IS(OampF)
















Aitchison J C et al


er

Stat

usTy





















Frankenwald Germany



France




































France





er

Stat

usTy







Newfoundland Canada































Newfoundland Canada



















Aitchison J C et al


er

Stat

usTy






















Upper Carboniferous






Carpathians Hungary














Russia










er

Stat

usTy





Basin Texas USA








































1986 108Upper









Aitchison J C et al


er

Stat

usTy





























Upper Carboniferous



Upper Carboniferous








Upper Carboniferous



Middle Ordovician














er

Stat

usTy






Ordovicianupper



















































Aitchison J C et al


er

Stat

usTy




















Upper Carboniferous












Middle Ordovician


















ChinaPraesaturnalis




China





er

Stat

usTy





fusiformaIS




IS(OampF)












Italy






















Lower Carboniferous



















Aitchison J C et al


er

Stat

usTy




















































Newfoundland Canada








Newfoundland Canada






Newfoundland Canada





er

Stat

usTy


































Upper Carboniferous















Upper Carboniferous






Upper Carboniferous




Aitchison J C et al


er

Stat

usTy









Upper Carboniferous















































er

Stat

usTy





































1996 222Lower















Middle Ordovician



Middle Ordovician






upper Tournaisian






Aitchison J C et al


er

Stat

usTy







918Middle







Middle Ordovician















































er

Stat

usTy




















































Aitchison J C et al


er

Stat

usTy









Newfoundland Canada




































clavataIS





IS(OampF)







er

Stat

usTy









Italy





















Elbingerode Germany




















Aitchison J C et al


er

Stat

usTy

























































er

Stat

usTy














Newfoundland Canada









Upper Carboniferous

























Aitchison J C et al


er

Stat

usTy










































UK





er

Stat

usTy





























Frankenwald Germany




















Aitchison J C et al


er

Stat

usTy




























Italy





Italy

























er

Stat

usTy

























Upper Carboniferous




















Elbingerode Germany



Aitchison J C et al


er

Stat

usTy


































Italy








Scotland UK













E ndash T Th







er

Stat

usTy






























ItalySpongotrochus












Kaeo Thailand


ItalyStauracontium







Italy








Scotland UK



Aitchison J C et al


er

Stat

usTy









USA

















































er

Stat

usTy

























Italy






















Lower Carboniferous






Aitchison J C et al


er

Stat

usTy































Elbingerode Germany















Italy



Scotland UK





er

Stat

usTy








Newfoundland Canada










Middle Ordovician





Middle Ordovician






Middle Ordovician

upper Darriwilian










239Upper









Upper Carboniferous





Aitchison J C et al


er

Stat

usTy





1986 72Upper










Upper Carboniferous





Italy














Germany




Germany









Upper Carboniferous




Upper Carboniferous

















Upper Carboniferous






er

Stat

usTy





USATetrentactinia
















USA








Italy


Italy



UKTheodiscus








Italy








Aitchison J C et al


er

Stat

usTy





1986 97Upper




1986 97Upper



1986 100Upper







Upper Carboniferous













Upper Carboniferous





Upper Carboniferous

























250Lower







er

Stat

usTy









upper Tournaisian














































Aitchison J C et al


er

Stat

usTy



















Lower Carboniferous

lower Tournaisian











Ordovicianupper


region Kazakhstan























er

Stat

usTy









































Newfoundland Canada










Aitchison J C et al


er

Stat

usTy











Newfoundland Canada






































Basin Texas USA







er

Stat

usTy






























Italy









































Aitchison J C et al










































































Aitchison J C et al











































































Aitchison J C et al













er

Stat

usTy














































E nn Th Ts








ActinommaActinomma


Elbingerode Germany

APPENDIX



Aitchison J C et al


er

Stat

usTy





(OampF)nc T











Lower Carboniferous





















Lower Carboniferous







Upper Carboniferous



Lower Carboniferous









er

Stat

usTy








Upper Carboniferous






Lower Carboniferous





Lower Carboniferous



Lower Carboniferous



Lower Carboniferous



Lower Carboniferous








upper Tournaisian





Upper Carboniferous







Lower Carboniferous





Lower Carboniferous






Lower Carboniferous







Lower Carboniferous





Upper Carboniferous




Aitchison J C et al


er

Stat

usTy








Lower Carboniferous









Lower Carboniferous




















Lower Carboniferous

















Lower Carboniferous













Lower Carboniferous








er

Stat

usTy








Germany




Lower Carboniferous



upper Tournaisian


























Upper Carboniferous











ItalyAmphibrachium


Elbingerode Germany



Aitchison J C et al


er

Stat

usTy










Elbingerode Germany









































er

Stat

usTy

























































Newfoundland Canada





Aitchison J C et al


er

Stat

usTy












upper Tournaisian





















upper Tournaisian



upper Tournaisian


























upper Tournaisian










er

Stat

usTy


































1986 124Upper






Upper Carboniferous


















Newfoundland Canada



Aitchison J C et al


er

Stat

usTy








Newfoundland Canada












upper Tournaisian



Middle Ordovician





Upper Carboniferous




Upper Carboniferous







upper Tournaisian









upper Tournaisian















er

Stat

usTy
















Middle Ordovician







Basin Texas USA








Italy

















Frankenwald Germany










Newfoundland Canada



Aitchison J C et al


er

Stat

usTy


























































er

Stat

usTy









1980 64Middle





Middle Ordovician






mindyallicanr Calc-






































Aitchison J C et al


er

Stat

usTy




































Upper Carboniferous







Newfoundland Canada











er

Stat

usTy






















Australia


Elbingerode Germany






















Newfoundland Canada



ItalyCecryphalium






Aitchison J C et al


er

Stat

usTy







































Basin Texas USA



Frankenwald Germany











er

Stat

usTy















103Middle





Middle Ordovician















































Lower Carboniferous





Aitchison J C et al


er

Stat

usTy























upper Tournaisian





















Lower Carboniferous






















er

Stat

usTy































upper Tournaisian






















Aitchison J C et al


er

Stat

usTy


















upper Tournaisian












Middle Ordovician

































Lower Carboniferous








er

Stat

usTy













1986 70Upper




1986 71Upper



1986 71Upper































Aitchison J C et al


er

Stat

usTy























1996 246Lower




Lower Carboniferous



Lower Carboniferous



Lower Carboniferous




Italy


USACorythoecia


1986 119Upper


Urals Russia





USA





er

Stat

usTy






Newfoundland Canada


Italy

CromyommaCromyomma





















Frankenwald Germany


1958 969Lower










Newfoundland Canada










Aitchison J C et al


er

Stat

usTy

























































Ts Th











er

Stat

usTy














Italy




Elbingerode Germany


Italy


















Italy










Aitchison J C et al


er

Stat

usTy






























Cornwall England UK




























er

Stat

usTy





































EchinommaEchinomma


Italy














Aitchison J C et al


er

Stat

usTy














Italy













Upper Carboniferous













Middle Ordovician





Middle Ordovician









er

Stat

usTy





236Upper







Upper Carboniferous




Upper Carboniferous




Upper Carboniferous

















Middle Ordovician















Aitchison J C et al


er

Stat

usTy




















































er

Stat

usTy






















Middle Ordovician






upper Tournaisian

















upper Tournaisian









Aitchison J C et al


er

Stat

usTy



















Upper Carboniferous





















Upper Carboniferous
























er

Stat

usTy



































































Middle Ordovician






Aitchison J C et al


er

Stat

usTy


























Middle Ordovician



















Izmir Region Turkey



















er

Stat

usTy




















IS(OampF)




































Aitchison J C et al


er

Stat

usTy
















Prefecture Japan


































er

Stat

usTy
















Frankenwald Germany



Newfoundland Canada





















Newfoundland Canada










Aitchison J C et al


er

Stat

usTy















Italy




















Middle Ordovician




















er

Stat

usTy








102Middle




Middle Ordovician
























Columbia Canada









Upper Carboniferous




Upper Carboniferous




Aitchison J C et al


er

Stat

usTy










China







Middle Ordovician

upper Darriwilian

































Middle Ordovician

upper Darriwilian









er

Stat

usTy























Upper Carboniferous


















Aitchison J C et al


er

Stat

usTy





















































er

Stat

usTy












Carpathians Hungary



Italy


Basin Texas USA




























Aitchison J C et al


er

Stat

usTy


























Barmen Germany













Middle Ordovician

upper Darriwilian
















lower Darriwilian






er

Stat

usTy

























France

















Aitchison J C et al


er

Stat

usTy






































1976 415Middle







er

Stat

usTy





























1986 82Upper



1986 79Upper



1986 82Upper



240Upper




Upper Carboniferous





Upper Carboniferous





Upper Carboniferous



1986 87Upper



1986 88Upper




1986 89Upper




1986 90Upper


Urals Russia



Aitchison J C et al


er

Stat

usTy














Upper Carboniferous












Upper Carboniferous




Upper Carboniferous
















Italy





er

Stat

usTy







ItalyLithocampe























Germany


Italy



















Aitchison J C et al


er

Stat

usTy














2008 53Lower













River Kazakhstan




























er

Stat

usTy



















































Aitchison J C et al


er

Stat

usTy
















































er

Stat

usTy


















Newfoundland Canada


Newfoundland Canada














Newfoundland Canada























Aitchison J C et al


er

Stat

usTy
































Lower Carboniferous








China





er

Stat

usTy











































anticaIS




IS(OampF)



IS(OampF)



IS(OampF)



IS(OampF)



IS(OampF)




Aitchison J C et al


er

Stat

usTy





Brittany France

















































er

Stat

usTy



























































Aitchison J C et al


er

Stat

usTy




























































er

Stat

usTy






























Newfoundland Canada


antiquusIS




IS(OampF)




IS(OampF)



IS(OampF)



IS(OampF)




IS(OampF)
















Aitchison J C et al


er

Stat

usTy





















Frankenwald Germany



France




































France





er

Stat

usTy







Newfoundland Canada































Newfoundland Canada



















Aitchison J C et al


er

Stat

usTy






















Upper Carboniferous






Carpathians Hungary














Russia










er

Stat

usTy





Basin Texas USA








































1986 108Upper









Aitchison J C et al


er

Stat

usTy





























Upper Carboniferous



Upper Carboniferous








Upper Carboniferous



Middle Ordovician














er

Stat

usTy






Ordovicianupper



















































Aitchison J C et al


er

Stat

usTy




















Upper Carboniferous












Middle Ordovician


















ChinaPraesaturnalis




China





er

Stat

usTy





fusiformaIS




IS(OampF)












Italy






















Lower Carboniferous



















Aitchison J C et al


er

Stat

usTy




















































Newfoundland Canada








Newfoundland Canada






Newfoundland Canada





er

Stat

usTy


































Upper Carboniferous















Upper Carboniferous






Upper Carboniferous




Aitchison J C et al


er

Stat

usTy









Upper Carboniferous















































er

Stat

usTy





































1996 222Lower















Middle Ordovician



Middle Ordovician






upper Tournaisian






Aitchison J C et al


er

Stat

usTy







918Middle







Middle Ordovician















































er

Stat

usTy




















































Aitchison J C et al


er

Stat

usTy









Newfoundland Canada




































clavataIS





IS(OampF)







er

Stat

usTy









Italy





















Elbingerode Germany




















Aitchison J C et al


er

Stat

usTy

























































er

Stat

usTy














Newfoundland Canada









Upper Carboniferous

























Aitchison J C et al


er

Stat

usTy










































UK





er

Stat

usTy





























Frankenwald Germany




















Aitchison J C et al


er

Stat

usTy




























Italy





Italy

























er

Stat

usTy

























Upper Carboniferous




















Elbingerode Germany



Aitchison J C et al


er

Stat

usTy


































Italy








Scotland UK













E ndash T Th







er

Stat

usTy






























ItalySpongotrochus












Kaeo Thailand


ItalyStauracontium







Italy








Scotland UK



Aitchison J C et al


er

Stat

usTy









USA

















































er

Stat

usTy

























Italy






















Lower Carboniferous






Aitchison J C et al


er

Stat

usTy































Elbingerode Germany















Italy



Scotland UK





er

Stat

usTy








Newfoundland Canada










Middle Ordovician





Middle Ordovician






Middle Ordovician

upper Darriwilian










239Upper









Upper Carboniferous





Aitchison J C et al


er

Stat

usTy





1986 72Upper










Upper Carboniferous





Italy














Germany




Germany









Upper Carboniferous




Upper Carboniferous

















Upper Carboniferous






er

Stat

usTy





USATetrentactinia
















USA








Italy


Italy



UKTheodiscus








Italy








Aitchison J C et al


er

Stat

usTy





1986 97Upper




1986 97Upper



1986 100Upper







Upper Carboniferous













Upper Carboniferous





Upper Carboniferous

























250Lower







er

Stat

usTy









upper Tournaisian














































Aitchison J C et al


er

Stat

usTy



















Lower Carboniferous

lower Tournaisian











Ordovicianupper


region Kazakhstan























er

Stat

usTy









































Newfoundland Canada










Aitchison J C et al


er

Stat

usTy











Newfoundland Canada






































Basin Texas USA







er

Stat

usTy






























Italy


Documents

Inventory of Paleozoic radiolarian species (1880-2016)...Paleozoic radiolarians. Here we present a comprehensive list of all the Paleozoic radiolarian species of which we are aware