k p 2008 – A p ` k x n o k p 2010 WWF ü ! 2 2011 · U b f l k o Z l t i b \ i ^ f n b b l f g Z m b l f o y n b f m k h x b ` ^ h x o b l k m n p ` g l f o f g Y b o ^ k p h f

$Page 1: k p 2008 – A p ` k x n o k p 2010 WWF ü ! 2 2011 · U b f l k o Z l t i b \ i ^ f n b b l f g Z m b l f o y n b f m k h x b ` ^ h x o b l k m n p ` g l f o f g Y b o ^ k p h f$
( ) 2008 – 2010

WWF , 2011

1

:

, PhD, ), [email protected]

, , PhD, [email protected]

«Winds of change in Dadia National Park»

WWF . , .

WWF ,

) .

: . & . 2011.

( ) 2008 – 2010, . 48. WWF , .

2

1. 2. 3.

3.1 , , 3.2 3.3 3.4

4. &

4.1 , , 4.2 4.3 4.4

5. 6. 7. 8. 9.

1. 8 2008 4 2010.

2. 3.

3

1.

, ( ).

« », (Thelander et al.

2007, Smallwood et al. 2008). ,

(de Lucas et al. 2004, Barrios & Rodriguez 2004, Smallwood et al. 2009, Masden et al. 2010).

, ( . lcalde 2003, Dürr & Bach 2004,

Brinknmann et al. 2006, Barclay et al. 2007, Kunz et al. 2007a, Arnett et al. 2008, Horn et al. 2008, Arnett et al. 2009).

(Brinkmann et al. 2006, Arnett et al. 2008). ,

Brinkmann et al. (2006)

(50 9 ). ,

,

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(Brinkmann et al. 2006, Arnett et al. 2008),

, . ,

(post monitoring),

, .

, . , ’ « »

(Cryan 2008). O ( , . )

,

4

,

(Brinkmann et al. 2006).

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2006, Baerwald et al. 2008).

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. ,

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.

, WWF 2008 – 2010. WWF , .

,

5

. , ,

, ,

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, -

, 11 500 . , « »

GR1110003 « – » GR1110010

» GR1130011. 11 ( 163

) 88 127 ( 1).

,

WWF (Carcamo et al. 2011).

1. ( ) ( 3/8/2009 10/8/2010).

%

D 8 8 (100%) D1-D8 42 21 (50%) 5-T13, T26- 36, 42

K 14 14 (100%) K1-K14 MA 3 3 (100%) 1- 3

MO 13 13 (100%) MO1-MO13 P 10 3 (30%) P8-P10

M 19 9 (47%) M3-M5, M7, M15-M19 X 5 4 (80%) X1-X4 S 13 13 (100%) S1-S13 127 88 (70%)

2008-2009 127 14 (Carcamo et al. 2011), 3/8/2009-10/10/2009 88 5

( ) , (11/10/2009-4/8/2010) 88

(Doutau et al. 2011). , 2009 – 2010,

6

, ( ), , . ,

3/8/2009-4/8/2010 251 . , ,

3/8/2009. 50m 2 .

, .

( 50m),

, . 88 , 2 2

33 45 4,5 .

30% 100% 88 50%.

, 50% 50% ( -

). , ( ) , ,

.

, , .

( )

WWF E (Carcamo et el. 2011, Doutau et al. 2011).

.

Dietz & Helversen (2004). ,

, . P. pipistrellus P. pygmaeus,

. , «Pipistrellus

pipistrellus/pygmaeus». Racey (1988),

( : 2-5 : 5 ), Anthony (1988).

(Correlation Analysis) SPSS 15. Excel 2007.

7

3.

3.1 , ,

( 8/7/2008 4/8/2010) 192 ( ) .

186 3/8/2009 4/8/2010, .

1. ,

Pipistrellus kuhlii 9/8/2009,

( 2).

Nyctalus leisleri (58 , 29.90% ) Pipistrellus pipistrellus/pygmaeus

(54 , 27.84% ) Pipistrellus nathusii Hypsugo savii (35 & 26 , 18.04% & 13.40%

, ), 1. N. noctula

(10 , 5.15% ), ,

.

1. ( 8/7/2008 4/8/2010), .

.

8

2. .

Red Data Book IUCN

2009

Red Data Book GR

2009

92/43/EEC Bern Convention

Bonn Convention

.

67/1981 Eptesicus serotinus (Schreber, 1774) LC LC Annex IV Annex II Appendix II - - Hypsugo savii (Bonaparte, 1837) LC LC Annex IV Annex II Appendix II Myotis mystacinus (Kuhl, 1819) LC DD Annex IV Annex II Appendix II Nyctalus lasiopterus (Schreber, 1780) NT VU Annex IV Annex II Appendix II Nyctalus leisleri (Kuhl, 1817) LC LC Annex IV Annex II Appendix II Nyctalus noctula (Schreber, 1774) LC DD Annex IV Annex II Appendix II Pipistrellus kuhlii (Kuhl, 1817) LC LC Annex IV Annex II Appendix II Pipistrellus nathusii (Keyserling & Blasius, 1839) LC DD Annex IV Annex II Appendix II Pipistrellus pipistrellus (Schreber, 1774) LC DD Annex IV Annex II Appendix II Pipistrellus pygmaeus (Leach 1825) LC DD Annex IV Annex II Appendix II - Vespertilio murinus Linnaeus, 1758 LC DD Annex IV Annex II Appendix II

Red Data Books VU = NT = LC = DD =

Habitats directive (92/43/EEC) Annex IV:

Bern Convention Annex II: Annex II :

Bonn Convention Appendix II: .

9

5/8/2009 4/8/2010, (173 , 89.18% )

. Hypsugo savii

, ( 2).

2. ( ) .

, , (66% 71% ),

( 3).

10

3. .

3.2

3/8/2009 4/8/2010,

(8 ) 14 . 3/8/2009 4/8/2010

5 , (5/8/2009

4/8/2010). 93% (173 ) 2009

2010. 2009 2010 ( 4 5). ,

( 6).

11

4. . 5/8/2009 4/8/2010,

.

5. . 2009 2010 ,

.

12

6. , .

2009 2010 , .

3.3

(194 )

(106 , 55%) . ( )

(53 , 27%) , , ( . , , ). 35

, , .

( 7 8),

. ,

.

13

7. .

8.

.

14

3.4

3/8/2009 4/8/2010 .

(66 , 35.48% 186 ), , ,

(25, 22, 20 13 , 13.44%, 11.83%, 10.75% 6.99% , ), 9.

9. .

186 .

( ),

. ,

6.25 , 4.71 ( 3).

15

3. .

. / )

.

(%)

. )

14/14 66 35.48% 4.71 4/5 25 13.44% 6.25

9/19 22 11.83% 2.44

8/8 20 10.75% 2.50

13/13 13 6.99% 1.00

3/10 9 4.84% 3.00

3/3 9 4.84% 3.00

13/13 9 4.84% 0.69

21/42 13 6.99% 0.62

(4.5% ), 26,34%

, 13 (14.8% ) ( 10).

43.55% ( 4).

4. ( ), .

K 14 19 10.22% 1.14%

X 04 14 17.74% 2.27%

K 03 9 22.58% 3.41%

D 08 7 26.34% 4.55%

K 13 7 30.11% 5.68%

K 02 5 32.80% 6.82%

K 12 5 35.48% 7.95%

M 03 5 38.17% 9.09%

M 16 5 40.86% 10.23%

X 01 5 43.55% 11.36%

16

10. .

.

.

, . ,

Nyctalus leisleri Pipistrellus pipistrellus/pygmaeus,

Pipistrellus pipistrellus/pygmaeus

Pipistrellus nathusii ( 11).

(Pearson

correlation, r = 0.60, p(one-tailed) < 0.05), (r = 0.53, p > 0.05).

17

11. , ( 3/8/2009 4/8/2010).

4. & 4.1 , ,

.

3/8/2009 4/8/2010 186 , 2.11 .

, . :

1. (searcher efficiency),

(Kunz 2004). ,

18

40% 84%, (Brinkmann et al. 2006).

2. (scavenger removal). ,

,

.

(Erickson 2004, Arnett et al. 2008). ,

. 3.

. ,

,

.

, .

,

. 4.

, .

5. 3/8/2009 10/10/2009,

2009 – 2010.

( . ).

( . Arnett et al. 2008 ).

, ,

19

(Cryan 2008, Kunz et al. 2007b). ,

(1-2 )

, (Brinkmann et al. 2006, Kunz et al.

2007a). .

, ( 2),

,

(Brinkmann et al. 2006, Rodrigues et al. 2008, Baerwald & Barclay 2009, Arnett et al. 2009). . (Bats and Wind Energy Cooperative – BWEC, http://www.batsandwind.org)

, ,

. 25 (Papadatou 2010, ), 10

11 ( 1), (Rodrigues et al. 2008). , 90%

(Nyctalus leisleri, Pipistrellus pipistrellus/pygmaeus, Pipistrellus nathusii Hypsugo savii),

. , ( Hypsugo savii)

(Dietz et al. 2008) « »

(Brinkmann et al. 2006, Arnett et al. 2008, Cryan 2008).

(Horn et al. 2008, Cryan & Barclay, 2009).

, ,

(Arnett et al. 2008, Baerwald & Barclay 2009) (Brinkmann et al. 2006).

, Nyctalus leisleri Pipistrellus nathusii (Dietz et al. 2008),

. , ,

20

(Hanák et al. 2001, O. Helversen, ).

, , ,

(Barclay et al. 2007, Grunwald & Schafer 2007). N. leisleri, N. noctula H. savii (Dietz et al. 2008),

.

. , , ( N. noctula, Papadatou 2010)

, . , »

. ) , ,

(Cryan & Barclay 2009). Nyctalus leisleri Pipistrellus

pipistrellus/pygmaeus 14 4

. ,

, , . ( 2),

(Anthony 1988). (Arnett et al. 2008, .

Brinkmann et al. 2006). ,

1 , ,

. ( ) ,

, ( 3). , (Arnett et al.

2008, . Brinkmann et al. 2006), , : 1) , Pipistrellus pipistrellus/pygmaeus Hypsugo savii

, (Papadatou 2010). ,

21

(Holzhaider & Zahn 2001, Russo 2002). ,

, . 2)

. , ( , )

Nyctalus P. nathusii , ,

(Hanák et al. 2001, O. v. Helversen ). 3)

.

(Cryan & Barclay 2009).

4.2

3/8/2009 4/8/2010 93% (173 ) ,

( 4 & 5). ,

, .

, ,

(Papadatou 2010). ,

( 6).

, , Pipistrellus nathusii

« » , ,

Nyctalus leisleri.

. (

) (Kunz et al. 2007a, 2007b, Arnett et al. 2008, Baerwald & Barclay 2009, .

Brinkmann et al. 2006).

(Kunz et al. 2007a, Arnett et al. 2008). ,

22

(Hanák et al. 2001, O. v. Helversen ),

. ,

N. leisleri P. nathusii ( . ). 4.3

(106 , 55% )

, (18%)

. 55% ,

, (Baerwald et al. 2008).

4.4

,

, .

, , , ( 9 & 3).

(Arnett et al. 2008),

. T

(Barclay 2007), . -

(Brinkmann et al.

2006), , ,

Pipistrellus pipistrellus, .

, , .

Nyctalus leisleri, Pipistrellus pipistrellus/pygmaeus P. nathusii

23

, ( 11 ), .

, (13

, 4 10),

( . ).

,

3,4 , 2004 (Johnson 2004).

, ( )

(Brinkmann et al. 2006).

. ,

( . ). 5.

, ,

( 2). ,

. ,

,

(Behr & v. Helversen 2006, Arnett et al. 2008),

(Brinkmann et al. 2006). ,

., 76 , Arnett et al. (2009), 73% 3,5 m/sec.

76 5

6.5 m/sec, ’ ,

24

3% 11% ( ) 0.3% 1% ( ) .

, ,

, . , (

) 4 m/sec ( )

2,5 (Baerwald et al. 2009).

, ,

( 5), . , ,

. 10

( ) ( ),

, ( , . 1):

13 14: 04: 03:

D08: 02: 12: 03: 16: 01:

6.5 m/sec.

(Horn et al. 2008), ,

. ,

. .

, , (

25

) (Rodrigues et al. 2008).

,

. 6.

.

, ,

. ,

- (Rodrigues et al. 2008).

, . ,

,

(Kunz 2004, Kunz et al. 2007a). , ,

,

(Arnett et al. 2008). ,

.

– (

) (Erickson 2004, Brinkmann et al. 2006, Arnett et al. 2008, Rodrigues et al. 2008).

(Arnett et al. 2008).

,

, ,

, , , ,

) ’ (Kunz 2004, Rodrigues et al. 2008). ,

26

,

,

(Barclay et al. 2007).

,

, , ,

, , ,

. ,

, (

).

(Kunz et al. 2007b). 7.

, , ,

( )

(Kunz et al. 2007b, Rodrigues et al. 2008).

, ,

,

(Kunz et al. 2007b, Rodrigues et al. 2008).

(Kunz 2004, Kunz et al. 2007b, Rodrigues et al. 2008): 1. ( ): )

.

.

27

, , ,

(Baewald & Barclay 2009). ,

( . ).

. ) , ,

. ,

.

. 2. ( –

, .) ( ).

. 3. ( ,

).

, ( . ),

(Brinkmann et al. 2006, Horn et al. 2008). 8. Alcalde, J.T. (2003): Impacto de los parques eólicos sobre las poblaciónes de

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Arnett, E. B., M. M. P. Huso, J. P. Hayes, and M. Schirmacher. 2009. Effectiveness of changing wind turbine cut-in speed to reduce bat fatalities at wind facilities. A final report submitted to the Bats and Wind

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31

9. 1. 8 2008 4 2010.

.

-

Myotis mystacinus 8/7/08 09 ? ? 5/9/08 P 02 ? ? 5/9/08 P 09 ? ? 16/9/08 14 ? ? 25/5/09 X 02

Pipistrellus pipistrellus/pygmaeus ? 30/5/09 P 01 Nyctalus leisleri 8/6/09 P 01 Hypsugo savii 19/6/09 11 Hypsugo savii ? ? 5/8/09 06 Hypsugo savii ? ? 5/8/09

X 04 . ,

Hypsugo savii ? ? 5/8/09 MA 03 . , Hypsugo savii 5/8/09 P 09 Hypsugo savii ? 5/8/09 MA 01 Nyctalus leisleri 5/8/09 06 Nyctalus leisleri 5/8/09

S 01 ,

Pipistrellus pipistrellus/pygmaeus ? 5/8/09

06 ,

) Pipistrellus pipistrellus/pygmaeus 5/8/09 33 Hypsugo savii 6/8/09 MA 03

32

.

-

' Pipistrellus pipistrellus/pygmaeus ? ? 6/8/09 D 07 . , Hypsugo savii ? ? 7/8/09 D 01 Hypsugo savii ? ? 7/8/09 D 02 Hypsugo savii ? 7/8/09

12 ,

Hypsugo savii ? 7/8/09 D 01 Pipistrellus kuhlii/pipistrellus / pygmaeus

? ? 7/8/09 X 01

Pipistrellus pipistrellus/pygmaeus ? 7/8/09 11 Hypsugo savii ? 10/8/09 03 Nyctalus leisleri 11/8/09 18 Pipistrellus pipistrellus/pygmaeus ? 12/8/09 42 Pipistrellus pipistrellus/pygmaeus 12/8/09 D 07 Hypsugo savii 14/8/09 16 Pipistrellus pipistrellus/pygmaeus 26/8/09 04 Hypsugo savii ? 31/8/09 S 11 Pipistrellus pipistrellus/pygmaeus 31/8/09 D 08 . , Nyctalus leisleri ? 3/9/09 13 , Nyctalus leisleri 4/9/09 12 Nyctalus leisleri 4/9/09 13 Nyctalus leisleri 4/9/09 05 Nyctalus leisleri 4/9/09 03 Nyctalus leisleri 4/9/09 05

33

.

-

Nyctalus leisleri 7/9/09 13 Nyctalus leisleri 7/9/09 05 Nyctalus noctula 7/9/09 09 Pipistrellus nathusii 7/9/09 P 08 Nyctalus leisleri 14/9/09 MA 01 Nyctalus leisleri 14/9/09

14 . ,

Nyctalus leisleri 14/9/09

14 . ,

Nyctalus leisleri 14/9/09 03 Nyctalus noctula 14/9/09

03 . ,

Pipistrellus nathusii ? 14/9/09 P 10 Pipistrellus nathusii 14/9/09

14 . ,

. Nyctalus leisleri 15/9/09

05 . ,

Nyctalus leisleri 15/9/09 13 Nyctalus noctula ? 15/9/09 28 Nyctalus noctula 16/9/09

S 01 . ,

Nyctalus noctula 16/9/09

05 . ,

Pipistrellus nathusii ? 17/9/09 X 04 Pipistrellus nathusii 17/9/09 07 Pipistrellus pipistrellus/pygmaeus 17/9/09 13

34

.

-

Nyctalus noctula 18/9/09 12 Nyctalus noctula ? 18/9/09 MO 12 Vespertilio murinus 18/9/09 05 Pipistrellus pipistrellus/pygmaeus 19/9/09 14 Nyctalus leisleri 24/9/09 03 Pipistrellus pipistrellus/pygmaeus ? 24/9/09 X 04 . Pipistrellus pipistrellus/pygmaeus 24/9/09

X 03 . ,

Pipistrellus nathusii 30/9/09 07 Pipistrellus pipistrellus/pygmaeus 1/10/09 14 Pipistrellus pipistrellus/pygmaeus 1/10/09 04 Nyctalus leisleri 5/10/09

06 ,

Nyctalus leisleri 5/10/09 05 Pipistrellus nathusii 7/10/09 18 Nyctalus leisleri 11/10/09 X 02 Pipistrellus pipistrellus/pygmaeus 11/10/09 X 01 Nyctalus leisleri 12/10/09 P 10 Nyctalus leisleri 12/10/09

K 10 . ,

Nyctalus leisleri 21/10/09 P 09 Pipistrellus pipistrellus/pygmaeus 22/10/09 X 04 Nyctalus leisleri 25/10/09 MA 01 Pipistrellus nathusii 29/10/09 X 01

35

.

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Pipistrellus pipistrellus/pygmaeus 29/10/09 X 02 ? ? 29/10/09 X 04

Nyctalus leisleri 15/11/09 16 Pipistrellus nathusii 16/11/09 MA 02 Pipistrellus nathusii 17/11/09 03 Pipistrellus pipistrellus/pygmaeus 23/3/10 06 Pipistrellus nathusii 24/3/10 M 15 Pipistrellus nathusii 8/4/10 MA 02 Pipistrellus nathusii 8/4/10 MO 04 Pipistrellus nathusii 16/4/10 02 Pipistrellus nathusii 19/4/10 05 Pipistrellus nathusii 21/4/10 D 07 Pipistrellus nathusii 22/4/10 17 Pipistrellus nathusii 25/4/10 16 Pipistrellus pipistrellus/pygmaeus 25/4/10 D 05 Pipistrellus pipistrellus/pygmaeus 3/5/10 MO 07 Nyctalus leisleri 4/5/10 K 01 Nyctalus leisleri 4/5/10 14 Pipistrellus nathusii 4/5/10 MO 06 Pipistrellus pipistrellus/pygmaeus 4/5/10 K 01 Nyctalus leisleri 5/5/10 02 Nyctalus leisleri 5/5/10 06 Nyctalus leisleri 7/5/10 02

36

.

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Nyctalus leisleri 11/5/10 14 Nyctalus leisleri 11/5/10 14 Nyctalus leisleri 11/5/10 14 Hypsugo savii 12/5/10 X 02 Pipistrellus pipistrellus/pygmaeus 12/5/10 D 08 Pipistrellus pipistrellus/pygmaeus 12/5/10 P 09 Hypsugo savii ? 13/5/10 MO 12 , Nyctalus leisleri 13/5/10 14 Nyctalus leisleri 13/5/10 14 Nyctalus leisleri 19/5/10 14 . Pipistrellus pipistrellus/pygmaeus ? 19/5/10 06 Hypsugo savii 30/5/10 S 02 Pipistrellus nathusii 31/5/10 13 Pipistrellus nathusii 1/6/10 P 09 , Pipistrellus pipistrellus/pygmaeus 1/6/10 26 . Nyctalus leisleri 3/6/10 14 Pipistrellus nathusii 4/6/10 09 Pipistrellus nathusii 4/6/10 D 06 Pipistrellus nathusii 8/6/10 M 15 Nyctalus leisleri 9/6/10 26 Pipistrellus nathusii 9/6/10 03 Pipistrellus nathusii 9/6/10 MO 01 Pipistrellus pipistrellus/pygmaeus 9/6/10 X 04

37

.

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Pipistrellus pipistrellus/pygmaeus 10/6/10 D 08 Nyctalus leisleri 11/6/10 D 03 Pipistrellus pipistrellus/pygmaeus 11/6/10 X 01 , Nyctalus leisleri 13/6/10 14 Nyctalus leisleri 13/6/10 14 Nyctalus noctula 13/6/10 S 13 Pipistrellus nathusii 13/6/10 07 Pipistrellus nathusii 13/6/10 X 04 Pipistrellus pipistrellus/pygmaeus 13/6/10 04 Nyctalus leisleri 14/6/10 14 Nyctalus leisleri 14/6/10 MO 03 Nyctalus noctula 14/6/10 D 07 Pipistrellus nathusii 14/6/10 13 Pipistrellus pipistrellus/pygmaeus 14/6/10 X 03 Pipistrellus pipistrellus/pygmaeus 14/6/10 05 Nyctalus leisleri 15/6/10 14 Pipistrellus pipistrellus/pygmaeus 15/6/10 MO 11 Pipistrellus pipistrellus/pygmaeus 15/6/10 MO 05 Nyctalus leisleri 16/6/10 MO 04 Pipistrellus pipistrellus/pygmaeus 16/6/10 12 Eptesicus serotinus 17/6/10 15 Hypsugo savii 17/6/10 05 . Nyctalus leisleri 17/6/10 14

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.

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Nyctalus lasiopterus 20/6/10 D 08 Nyctalus leisleri 21/6/10 14 Pipistrellus nathusii 21/6/10 16

? 29/6/10 MO 09 . , Nyctalus leisleri 30/6/10 D 08 Pipistrellus pipistrellus/pygmaeus /

1/7/10

30

Hypsugo savii 2/7/10 04 Pipistrellus pipistrellus/pygmaeus 2/7/10 03 Nyctalus leisleri 4/7/10 MO 06 Nyctalus noctula 5/7/10 D 02 Hypsugo savii 6/7/10 D 08 Nyctalus leisleri 7/7/10 D 06 Pipistrellus pipistrellus/pygmaeus 7/7/10 03 Hypsugo savii 11/7/10 07 Hypsugo savii 12/7/10 D 08 Nyctalus leisleri 12/7/10 K 01 Nyctalus leisleri 12/7/10 K 01 Nyctalus leisleri 12/7/10 02 Pipistrellus pipistrellus/pygmaeus 12/7/10 X 01 Pipistrellus pipistrellus/pygmaeus 13/7/10 D 06 Pipistrellus pipistrellus/pygmaeus 13/7/10 X 03 Pipistrellus pipistrellus/pygmaeus 13/7/10 16

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Hypsugo savii 14/7/10 X 04 Nyctalus leisleri 14/7/10 12 Pipistrellus nathusii 14/7/10 X 04 Pipistrellus nathusii 14/7/10 S 10 Pipistrellus pipistrellus/pygmaeus 14/7/10 S 09 Pipistrellus pipistrellus/pygmaeus 14/7/10 X 04 Pipistrellus pipistrellus/pygmaeus 14/7/10 X 04 Pipistrellus pipistrellus/pygmaeus ? 15/7/10 X 04 Pipistrellus pipistrellus/pygmaeus

15/7/10

X 04

Pipistrellus pipistrellus/pygmaeus 15/7/10 02 Pipistrellus pipistrellus/pygmaeus 16/7/10 MA 03 Pipistrellus pipistrellus/pygmaeus 16/7/10 03 Pipistrellus nathusii 18/7/10 03 Pipistrellus nathusii 21/7/10 03 Hypsugo savii 27/7/10 S 11 Nyctalus leisleri 27/7/10 03 Pipistrellus pipistrellus/pygmaeus 27/7/10 X 04 Pipistrellus nathusii 28/7/10 MA 01 Pipistrellus pipistrellus/pygmaeus 28/7/10 09 Pipistrellus pipistrellus/pygmaeus 28/7/10 MO 12 Pipistrellus pipistrellus/pygmaeus 30/7/10 03 Hypsugo savii 1/8/10 S 04

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Nyctalus leisleri 1/8/10 P 08 Hypsugo savii ? 4/8/10 P 08

: : : :

?: D: G: K: MA: MO: M: P: X: S:

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2.

1. , . Hypsugo savii, 01/08/2010 S 04 ( ).

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2. Vespertilio murinus, 18/09/2009 M 05 ( ).

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3. Pipistrellus pipistrellus P. pygmaeus, 05/08/2009 T 06 ).

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4. Pipistrellus nathusii, 09/06/2010 MO 01 ( ).

.

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5. Nyctalus noctula, 05/07/2010 D 02 ( ). .

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6. Nyctalus leisleri, 05/10/2009 K 06 ( ). .

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7. Hypsugo savii, 05.08.09 MA 03 ( ). .

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3.

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