35
, A .... .. Techn ic al Pap er No. 6/2002 6/2545 Study on Biology and Breeding of Orange-headed Temple Terrapin · Heosemys grandis (Gray) Wonnut Hirunchulha Chaiwat Ratanadadas Kamthon Jaroonsak Satun Inland Fisheries Station 91110 La-ngu, Satun Province, 91110 OJ <u 0-7478-1299 Tel. 0-7478-1299 2545 2002 . Q..o 1Cf,QQ..r 42- 42 -2-1573-01 - 1-492-070

 · 'l.I'VI FlIYl!i'fl m1~m::tl~'J':)'VltJ1LL~~m1L'Y'I1~'V~\.},ib~h'i-1'JltJ~lLU'Um1tlPmv'hmr~mil~'U~'El~t)1~tJ mr . U~~~n'l::tru~~'J':)'Vl . tJ1mr~1J9,'l\.li ~.;j1~bbri

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A

Technical Paper No 62002Leuroln~Tn1$lnl~lJ1JfI 62545

Study on Biology and Breeding of Orange-headed Temple Terrapin middot

Heosemys grandis (Gray)

Wonnut Hirunchulha

Chaiwat Ratanadadas

Kamthon Jaroonsak

Satun Inland Fisheries Station

il ~1 ~ ~~~ 91110 La-ngu Satun Province 91110 OJ ltu

tYI~ 0-7478-1299 Tel 0-7478-1299

2545 2002

Qo 1CfQQr ~

middotfVI ~t1~LUJ1~HH~OZJ t1 42-42-2-1573-01 -1-492-070

~f)mnfdlJlf11)uUUYI 62545 Technical Paper No 62002

n1ai3in~1~rl~VI~1 bb~~n1ab~1~rrU fiblPiTVIrl1~ q

Study on Biology and Breeding of Orange-headed Temple Terrapin

Heosemys grandis (Gray)

OJ 4JiIiIWWVI VIiIrulfl-fVIl Wonnut Hirunchulha b1lJ100lW rClWfil1 [Pl1M Chaiwat Ratanadadas

o ltU 4 n1fiil ~iIru~nfil Kamthon Jaroonsak

OJ W

Satun Inland Fisheries Station

Cl Inland Fisheries Division

Department of Fisheries

Ministry of Agriculture and Cooperatives

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16

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17

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18 ilW~ilJIflJ ~JWfll-nJiJmJ-J~lJ (tJflVi

20

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21

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24

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(nn)

34

32

30

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24

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25

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26

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3

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Nutaphand W 1 The Turtles Mitbhadung Press 222pp

  • การศกษาชววทยาและการเพาะพนธเตาหวาย
Page 2:  · 'l.I'VI FlIYl!i'fl m1~m::tl~'J':)'VltJ1LL~~m1L'Y'I1~'V~\.},ib~h'i-1'JltJ~lLU'Um1tlPmv'hmr~mil~'U~'El~t)1~tJ mr . U~~~n'l::tru~~'J':)'Vl . tJ1mr~1J9,'l\.li ~.;j1~bbri

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Study on Biology and Breeding of Orange-headed Temple Terrapin

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Satun Inland Fisheries Station

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Ministry of Agriculture and Cooperatives

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Nutaphand W 1 The Turtles Mitbhadung Press 222pp

  • การศกษาชววทยาและการเพาะพนธเตาหวาย
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21

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(nn)

34

32

30

28

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= 04 152

24

22

20

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25

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26

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ilf 48 (3)5-10

Nutaphand W 1 The Turtles Mitbhadung Press 222pp

  • การศกษาชววทยาและการเพาะพนธเตาหวาย
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q

1 gt I1YI~

4

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J iI cv i b~ tJ~ l)tJfl1jn~tJ nm

q

w1jnn

bull ltV

WflWWn q

m1~ tJ1) w1VlWn m1~ tJ1) ~1W)W~ rJWb~flWU

mJ~Wfi q

~1W)W ~n fYm1 rJWb~flWU ~tJb)~1 o

l1W)W

(nn) (TIlJ) (TIlJ) (nn) (1llJ) (1llJ) (nll~ ~)1l1~ 1~ bUWIil ytn ~YlmuWIi) (rJW) i~~fl~fll

(Ii) ) (Ylfll) (Ii)) () (Ii))

740 332 410 310 244 290 1 11242 2 50 6542 83 CD

790 354 420 320 270 295 ~ 4342 4 2 50 9642 96 2

560 320 330 X

470 300 340 X

320 241 284 X

240 280 310 X

765 343 415 370 276 308 333 300 3 50 895 100

plusmn035 plusmn156 plusmn071 plusmn120 plusmn308 plusmn227 plusmn141 plusmn650

bull bull

18 ilW~ilJIflJ ~JWfll-nJiJmJ-J~lJ (tJflVi

20

--~ ----~~----------~--~~----

tlfl1V~-Ib~m 7690 731 7381 1591

v eJn1j-I 774 009 9339 505

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y 41035 x + 3144 (R2

06295 pltO05)

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21

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nAl 275 8500

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bd-J tJ 295 8940

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~

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lJElU(9J1~lJ1lbUWl~JElU 1 l~ i1l il~ltJUWLb~Wm iI El1~tJ1nDEl~uWt-I~1 ~~~~lWViEl1 ~~W1J1-nil ~IO)W

n~lbti1lJW1i1~1 V~UfDiElWJh~~bt-Ii1d-JAd-JUi1~tJ1l U1blrub~lhIL~i11t1m~ ~m~lru~~ltJ ~~d-J U~blrulJ1

I I iI I

5 lIt1C911li LI711 bb~~lIWlC91~n l[~hVlYlf1fYf)f1ll n 1 li1Jel~bb~iWeuroVlVI(il~el~ b~Jl~JtIelTWlit1~llt1~l ~f1u~illln~JW14 (UelVl1) q q

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24

21

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15

13

y 41035 x + 3144

R 06295

11

26 28 3 32 34 36 38

(nn)

34

32

30

28

y 66996 x + 81453 26

= 04 152

24

22

20

3 31 32 33 34 35 36 37 38

25

11 i1UViflrlenffm~~~n1i~tWfni~lti1t1

ltOJ

l 1 9gt

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I $t I II ~

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26

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ilf 48 (3)5-10

Nutaphand W 1 The Turtles Mitbhadung Press 222pp

  • การศกษาชววทยาและการเพาะพนธเตาหวาย
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q

1 gt I1YI~

4

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J iI cv i b~ tJ~ l)tJfl1jn~tJ nm

q

w1jnn

bull ltV

WflWWn q

m1~ tJ1) w1VlWn m1~ tJ1) ~1W)W~ rJWb~flWU

mJ~Wfi q

~1W)W ~n fYm1 rJWb~flWU ~tJb)~1 o

l1W)W

(nn) (TIlJ) (TIlJ) (nn) (1llJ) (1llJ) (nll~ ~)1l1~ 1~ bUWIil ytn ~YlmuWIi) (rJW) i~~fl~fll

(Ii) ) (Ylfll) (Ii)) () (Ii))

740 332 410 310 244 290 1 11242 2 50 6542 83 CD

790 354 420 320 270 295 ~ 4342 4 2 50 9642 96 2

560 320 330 X

470 300 340 X

320 241 284 X

240 280 310 X

765 343 415 370 276 308 333 300 3 50 895 100

plusmn035 plusmn156 plusmn071 plusmn120 plusmn308 plusmn227 plusmn141 plusmn650

bull bull

18 ilW~ilJIflJ ~JWfll-nJiJmJ-J~lJ (tJflVi

20

--~ ----~~----------~--~~----

tlfl1V~-Ib~m 7690 731 7381 1591

v eJn1j-I 774 009 9339 505

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y 41035 x + 3144 (R2

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21

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nAl 275 8500

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~

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n~lbti1lJW1i1~1 V~UfDiElWJh~~bt-Ii1d-JAd-JUi1~tJ1l U1blrub~lhIL~i11t1m~ ~m~lru~~ltJ ~~d-J U~blrulJ1

I I iI I

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24

21

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R 06295

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26 28 3 32 34 36 38

(nn)

34

32

30

28

y 66996 x + 81453 26

= 04 152

24

22

20

3 31 32 33 34 35 36 37 38

25

11 i1UViflrlenffm~~~n1i~tWfni~lti1t1

ltOJ

l 1 9gt

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I $t I II ~

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26

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ilf 48 (3)5-10

Nutaphand W 1 The Turtles Mitbhadung Press 222pp

  • การศกษาชววทยาและการเพาะพนธเตาหวาย
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45

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q

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bull ltV

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(nn) (TIlJ) (TIlJ) (nn) (1llJ) (1llJ) (nll~ ~)1l1~ 1~ bUWIil ytn ~YlmuWIi) (rJW) i~~fl~fll

(Ii) ) (Ylfll) (Ii)) () (Ii))

740 332 410 310 244 290 1 11242 2 50 6542 83 CD

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560 320 330 X

470 300 340 X

320 241 284 X

240 280 310 X

765 343 415 370 276 308 333 300 3 50 895 100

plusmn035 plusmn156 plusmn071 plusmn120 plusmn308 plusmn227 plusmn141 plusmn650

bull bull

18 ilW~ilJIflJ ~JWfll-nJiJmJ-J~lJ (tJflVi

20

--~ ----~~----------~--~~----

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21

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24

21

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26 28 3 32 34 36 38

(nn)

34

32

30

28

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= 04 152

24

22

20

3 31 32 33 34 35 36 37 38

25

11 i1UViflrlenffm~~~n1i~tWfni~lti1t1

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I $t I II ~

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~ ElUW11 ~~1rlJdl~ ~Vi6l V1~U91lll lbi1 rllVi6l El1IJflnJ1U1Yi 2532 niJYit7J6lEll Co bull

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ilf 48 (3)5-10

Nutaphand W 1 The Turtles Mitbhadung Press 222pp

  • การศกษาชววทยาและการเพาะพนธเตาหวาย
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45

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790 354 420 320 270 295 ~ 4342 4 2 50 9642 96 2

560 320 330 X

470 300 340 X

320 241 284 X

240 280 310 X

765 343 415 370 276 308 333 300 3 50 895 100

plusmn035 plusmn156 plusmn071 plusmn120 plusmn308 plusmn227 plusmn141 plusmn650

bull bull

18 ilW~ilJIflJ ~JWfll-nJiJmJ-J~lJ (tJflVi

20

--~ ----~~----------~--~~----

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v eJn1j-I 774 009 9339 505

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21

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26 28 3 32 34 36 38

(nn)

34

32

30

28

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= 04 152

24

22

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3 31 32 33 34 35 36 37 38

25

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6(2)1 48

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ilf 48 (3)5-10

Nutaphand W 1 The Turtles Mitbhadung Press 222pp

  • การศกษาชววทยาและการเพาะพนธเตาหวาย
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nr~(i)f)1U~L1rull1nh

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Nutaphand W 1 The Turtles Mitbhadung Press 222pp

  • การศกษาชววทยาและการเพาะพนธเตาหวาย
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16

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17

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q q q

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740 332 410 310 244 290 1 11242 2 50 6542 83 CD

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560 320 330 X

470 300 340 X

320 241 284 X

240 280 310 X

765 343 415 370 276 308 333 300 3 50 895 100

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18 ilW~ilJIflJ ~JWfll-nJiJmJ-J~lJ (tJflVi

20

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v eJn1j-I 774 009 9339 505

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21

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bull bull

24

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17

15

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y 41035 x + 3144

R 06295

11

26 28 3 32 34 36 38

(nn)

34

32

30

28

y 66996 x + 81453 26

= 04 152

24

22

20

3 31 32 33 34 35 36 37 38

25

11 i1UViflrlenffm~~~n1i~tWfni~lti1t1

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I $t I II ~

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bllBlJUl (~lnnlj~BUmlJ iJb~tm~ni(eltJ tlmr)uu~bJrutn~Vlb6l~1WYlUbf91111u(j)

Nutaphand (1979) j

Jl1fCln6lIl bb6llllY4trfl bbf1~YlUV1UJbVlIlYlWl Wlbm~tJ bbf1StiL(j)ih~tJ ~l1~UllY4rnlUelelm~tJl

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n1JV1 bf91 lllulYlil~lYlmlJlru6ll bUf)lllniJillIlltirrU bf911ll U~mUU~mPl tif)n~lnilill

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~ ~

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  • การศกษาชววทยาและการเพาะพนธเตาหวาย
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16

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17

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q q q

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Il - shy

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I iI I V I

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11

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18 ilW~ilJIflJ ~JWfll-nJiJmJ-J~lJ (tJflVi

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34

32

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24

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25

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6(2)1 48

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ilf 48 (3)5-10

Nutaphand W 1 The Turtles Mitbhadung Press 222pp

  • การศกษาชววทยาและการเพาะพนธเตาหวาย
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18 ilW~ilJIflJ ~JWfll-nJiJmJ-J~lJ (tJflVi

20

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(nn)

34

32

30

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24

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25

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26

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560 320 330 X

470 300 340 X

320 241 284 X

240 280 310 X

765 343 415 370 276 308 333 300 3 50 895 100

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bull bull

18 ilW~ilJIflJ ~JWfll-nJiJmJ-J~lJ (tJflVi

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ilf 48 (3)5-10

Nutaphand W 1 The Turtles Mitbhadung Press 222pp

  • การศกษาชววทยาและการเพาะพนธเตาหวาย
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q q q

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ilf 48 (3)5-10

Nutaphand W 1 The Turtles Mitbhadung Press 222pp

  • การศกษาชววทยาและการเพาะพนธเตาหวาย
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ilf 48 (3)5-10

Nutaphand W 1 The Turtles Mitbhadung Press 222pp

  • การศกษาชววทยาและการเพาะพนธเตาหวาย
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3 31 32 33 34 35 36 37 38

25

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ilf 48 (3)5-10

Nutaphand W 1 The Turtles Mitbhadung Press 222pp

  • การศกษาชววทยาและการเพาะพนธเตาหวาย
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24

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26 28 3 32 34 36 38

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34

32

30

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24

22

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3 31 32 33 34 35 36 37 38

25

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Nutaphand W 1 The Turtles Mitbhadung Press 222pp

  • การศกษาชววทยาและการเพาะพนธเตาหวาย
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Nutaphand W 1 The Turtles Mitbhadung Press 222pp

  • การศกษาชววทยาและการเพาะพนธเตาหวาย
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Nutaphand W 1 The Turtles Mitbhadung Press 222pp

  • การศกษาชววทยาและการเพาะพนธเตาหวาย
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51

1

164

173

31 53

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44 50

51

10 w

315plusmn007 008 2977 plusmn 1 15

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24

21

17

15

13

y 41035 x + 3144

R 06295

11

26 28 3 32 34 36 38

(nn)

34

32

30

28

y 66996 x + 81453 26

= 04 152

24

22

20

3 31 32 33 34 35 36 37 38

25

11 i1UViflrlenffm~~~n1i~tWfni~lti1t1

ltOJ

l 1 9gt

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I $t I II ~

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Nutaphand (1979) j

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26

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27

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1 iJgf9ljlf11jJll1~ 6667 bUElfbJlUI7l bb~iJ~IWdU1~ 19 ~Ell iiJWbb~

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b1l~tJ 305 1JYl11b1l~b~tJ61 -e1f9lnm1Yfn bUElfbJlW17l JjWlf71~m7l1~U1ViUn

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n 1 JVl f71flEl~ fl1JU 611I-J1 Jtl UlI-J 1HUjLu-n ulWn 1 j l~ llYrWfb7l 1V1d lU1mb~JElj n 1 JElu1mfb ~ El flJrlJ 1I-J

28

f1)1tJ~ul~

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4 1

29

2529 n1J~nltl~UEll~UnljJllhj1lElH[hm~ellU 11j~Irn1J1hIJl

2539

39 V1ih

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melTW1WUtlV11 91 bYltJIJ ll6lV11 rnfiJjIJllI1fi 39(4)391-421

tJJbtJlIU bll1J~1~U tluWll ~~1rudl~ LLi1 lltlVifltf ~JruPln~ eJ6l1ltl~1 IlllJWJJIJ lb6l1rYi bbtl6l Co

nJtl1Wbb6l 1

nremJjIJlJ1~ f9l

21 -lUi

~~~tJ bVilY1rl lJrJlldtJ ~llIJUVl~ rlTVi6l tlrnIJVllbWIWIVl Lb6l q

~tH~tVHltlnin~I~WhnjEnW J1J~1JnljthIJl 637

bb~ 3-ntJ 2525 lL6lM)lJ~dW~W~uhJ1~f9l1W

6(2)1 48

~ ElUW11 ~~1rlJdl~ ~Vi6l V1~U91lll lbi1 rllVi6l El1IJflnJ1U1Yi 2532 niJYit7J6lEll Co bull

Yrn1li b[hmtl1W1un~Ellb~IJ jltJ~1tm1jtrIJIJW11l11 2533 mlJtlJIJ~

ilf 48 (3)5-10

Nutaphand W 1 The Turtles Mitbhadung Press 222pp

  • การศกษาชววทยาและการเพาะพนธเตาหวาย
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~n ~rulJell1)l bli1WJ ltJ btl~f)n1ll bb-lJ--1 A~r)tJ1~1ri lJUI (9) In ~b~tJl flU1ll1ri~WL~ell1W1 (9) nr~ I 1I

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320plusmn006 b6JlW~hWlI~ mld-JmJbil~tJ 543plusmn008 bDW~bd-JlI~ fl1d-J~~U bwi1lln~n~rumJ1mn

n~1 ~lJIJ 1W1~VlnyJellnyJillellnrflCleltJeltl bb6llJ fN b~ClJf11tJb W1~Cl1t1m1~ biJelVilHtI~d-Jlru q 11

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q q q

brlfl~ntJl~rmil~tJ 800+170 nYClnfd-J AJId-Jn11lmVl eJlbU~tJ 339+551 b6JlW~bWfl~ AJId-JtJ1J

Il - shy

m~Ellbil~tJ 425plusmn354 b6JlW~bWlI~ ~JWbVlflb~tJntJl~rmil~tJ 298plusmn048 nTClnfd-J mld-Jn11l

I iI I V I

bliI~JltJy)V1~Clellb~tJl~JtJernUl~lJ(UelVl 2) b~lbVlflerntJl~rmil~tJ 765+035 mClnfw mld-J q Il shy

n11l m(Jell bil~tJ 343plusmn156 b1W~ bW fl~ AJId-JtJ1J m~ell bil~tJ 415plusmn0 71 b1W~ bWll~ ~JWbrlfl ~~- v J A 11 d ~ltV

bwtJd-JWI~Wmil6ltJ 37plusmn113 nTClmd-J mlWnJIln~(9)EllbilCltJ 276plusmn308 b6JlWllbd-Jll~ AJId-JtJ1J

n~(9)ellbil~tJ 308+227 b1W~bd-JllJ bbw~Wrfb(h~JltJ~b~tJl~JtJell~I~1WUel~ 1 nnr~J1l1~ 4 llJ-

ln 69lJ ~llnnr~JIlhI66 7 bUelfb~Wilm~~WJW 19 ~ill ~1elmmrubil~tJ 475plusmn200 ~ell~il

bbW bb6lbbW~Weuro~b~tJl~JtJil1~I~lWtJil~ 2 Jll1ll 2 9lJ ~ln 6llJ ~mlnr~JIlhi 333 btlflfb~Wil n q

1ll~WJW 6 yJfll ~1flmmrubil~tJ 300plusmn141 ~illriilbbW ~llnnr~yJmUWllJbbCl~II~lnr~~flllll1tJ

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11

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(nn) (TIlJ) (TIlJ) (nn) (1llJ) (1llJ) (nll~ ~)1l1~ 1~ bUWIil ytn ~YlmuWIi) (rJW) i~~fl~fll

(Ii) ) (Ylfll) (Ii)) () (Ii))

740 332 410 310 244 290 1 11242 2 50 6542 83 CD

790 354 420 320 270 295 ~ 4342 4 2 50 9642 96 2

560 320 330 X

470 300 340 X

320 241 284 X

240 280 310 X

765 343 415 370 276 308 333 300 3 50 895 100

plusmn035 plusmn156 plusmn071 plusmn120 plusmn308 plusmn227 plusmn141 plusmn650

bull bull

18 ilW~ilJIflJ ~JWfll-nJiJmJ-J~lJ (tJflVi

20

--~ ----~~----------~--~~----

tlfl1V~-Ib~m 7690 731 7381 1591

v eJn1j-I 774 009 9339 505

tl~1V~-Ib~meJ~~rrnij-I 14 2157 192 9156 844

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y 41035 x + 3144 (R2

06295 pltO05)

dI ~ (f V ) V 1 V 0 V I wi ltV ~ trv

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iilJn1J Y = 66996 x + 81453 (R2=041 P lt 005) biJ)tJJPllllJ~lJvrWfi~lriel~lWJtKlJthW

nilljv-ielWjflj~lly]lrllJ~eltJfl 42 5 6 IlflitJ~ 9 10)lJ

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elrul1Jl~IU~tJ 290plusmn100 eljPllllili~tJii 11iIJPllllJ~W~lJvrVlfilU~tJ 881

21

~iIElW Elrut-lJli (oc) AYld-J~W~d-J~Ylfi()

nAl 275 8500

d-JA 285 8020

bd-J tJ 295 8940

Al A 300 9365

itJ 295 9245

~U~tJ plusmnSD 290 plusmn 100 8814 plusmn 555

b~El~ n Ll1 AJU rhVlWiI ~l~1~ nElEl nronnt ~ i1 n b~ll-ihh nniI~~~~c1WLt~Eln 1~ lO)Wbt~Eln~ ~

~

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b tiEl191 n iJ Lt~ El n 1 ~ bUWllU1 AU Vl~l11 nElEl n91 n l~m~ i1El~ bb~ElEl n n~11 U~bl ru~lWViEl~iJ tI~1~ b1JW

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lJElU(9J1~lJ1lbUWl~JElU 1 l~ i1l il~ltJUWLb~Wm iI El1~tJ1nDEl~uWt-I~1 ~~~~lWViEl1 ~~W1J1-nil ~IO)W

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51

1

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173

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44 50

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315plusmn007 008 2977 plusmn 1 15

bull bull

24

21

17

15

13

y 41035 x + 3144

R 06295

11

26 28 3 32 34 36 38

(nn)

34

32

30

28

y 66996 x + 81453 26

= 04 152

24

22

20

3 31 32 33 34 35 36 37 38

25

11 i1UViflrlenffm~~~n1i~tWfni~lti1t1

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l 1 9gt

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I $t I II ~

ntielll111 iSW bb~ d bf911l11d It Pl~lti6lj1ueltJ1II

tiUl bUBl~1 n bf911 n~wil~ltiL lIIruell ~tIt tiUl~ (j) ~B (j) Pl~B~ ~

YJll wnYlUf)~1tiUl1-l1nn~IUtiUn

n1J bbYl imlltJIlBl bf911l11dlt1 1tirrlWr~~ Wl~YlU U711Wn tI biYl1ml1 tJBtJVlnfil UlfliJ 1111

Wln~~~~fil ~dti~iilUlB~ti 1 b-DtifilUlBb~Bl VlltiL(j)ti vilbbYl fCntin1l11~j ~llllr)1 Lb6l

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Nutaphand (1979) j

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ilf 48 (3)5-10

Nutaphand W 1 The Turtles Mitbhadung Press 222pp

  • การศกษาชววทยาและการเพาะพนธเตาหวาย
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n

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b1l~tJ 305 1JYl11b1l~b~tJ61 -e1f9lnm1Yfn bUElfbJlW17l JjWlf71~m7l1~U1ViUn

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28

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nJtl1Wbb6l 1

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6(2)1 48

~ ElUW11 ~~1rlJdl~ ~Vi6l V1~U91lll lbi1 rllVi6l El1IJflnJ1U1Yi 2532 niJYit7J6lEll Co bull

Yrn1li b[hmtl1W1un~Ellb~IJ jltJ~1tm1jtrIJIJW11l11 2533 mlJtlJIJ~

ilf 48 (3)5-10

Nutaphand W 1 The Turtles Mitbhadung Press 222pp

  • การศกษาชววทยาและการเพาะพนธเตาหวาย
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560 320 330 X

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18 ilW~ilJIflJ ~JWfll-nJiJmJ-J~lJ (tJflVi

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v eJn1j-I 774 009 9339 505

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21

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26 28 3 32 34 36 38

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34

32

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= 04 152

24

22

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3 31 32 33 34 35 36 37 38

25

11 i1UViflrlenffm~~~n1i~tWfni~lti1t1

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ilf 48 (3)5-10

Nutaphand W 1 The Turtles Mitbhadung Press 222pp

  • การศกษาชววทยาและการเพาะพนธเตาหวาย
Page 24:  · 'l.I'VI FlIYl!i'fl m1~m::tl~'J':)'VltJ1LL~~m1L'Y'I1~'V~\.},ib~h'i-1'JltJ~lLU'Um1tlPmv'hmr~mil~'U~'El~t)1~tJ mr . U~~~n'l::tru~~'J':)'Vl . tJ1mr~1J9,'l\.li ~.;j1~bbri

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21

17

15

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y 41035 x + 3144

R 06295

11

26 28 3 32 34 36 38

(nn)

34

32

30

28

y 66996 x + 81453 26

= 04 152

24

22

20

3 31 32 33 34 35 36 37 38

25

11 i1UViflrlenffm~~~n1i~tWfni~lti1t1

ltOJ

l 1 9gt

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I $t I II ~

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Nutaphand (1979) j

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26

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27

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n

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bull Y

b1l~tJ 305 1JYl11b1l~b~tJ61 -e1f9lnm1Yfn bUElfbJlW17l JjWlf71~m7l1~U1ViUn

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n 1 JVl f71flEl~ fl1JU 611I-J1 Jtl UlI-J 1HUjLu-n ulWn 1 j l~ llYrWfb7l 1V1d lU1mb~JElj n 1 JElu1mfb ~ El flJrlJ 1I-J

28

f1)1tJ~ul~

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29

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39 V1ih

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melTW1WUtlV11 91 bYltJIJ ll6lV11 rnfiJjIJllI1fi 39(4)391-421

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21 -lUi

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6(2)1 48

~ ElUW11 ~~1rlJdl~ ~Vi6l V1~U91lll lbi1 rllVi6l El1IJflnJ1U1Yi 2532 niJYit7J6lEll Co bull

Yrn1li b[hmtl1W1un~Ellb~IJ jltJ~1tm1jtrIJIJW11l11 2533 mlJtlJIJ~

ilf 48 (3)5-10

Nutaphand W 1 The Turtles Mitbhadung Press 222pp

  • การศกษาชววทยาและการเพาะพนธเตาหวาย
Page 25:  · 'l.I'VI FlIYl!i'fl m1~m::tl~'J':)'VltJ1LL~~m1L'Y'I1~'V~\.},ib~h'i-1'JltJ~lLU'Um1tlPmv'hmr~mil~'U~'El~t)1~tJ mr . U~~~n'l::tru~~'J':)'Vl . tJ1mr~1J9,'l\.li ~.;j1~bbri

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21

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5 lIt1C911li LI711 bb~~lIWlC91~n l[~hVlYlf1fYf)f1ll n 1 li1Jel~bb~iWeuroVlVI(il~el~ b~Jl~JtIelTWlit1~llt1~l ~f1u~illln~JW14 (UelVl1) q q

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21

17

15

13

y 41035 x + 3144

R 06295

11

26 28 3 32 34 36 38

(nn)

34

32

30

28

y 66996 x + 81453 26

= 04 152

24

22

20

3 31 32 33 34 35 36 37 38

25

11 i1UViflrlenffm~~~n1i~tWfni~lti1t1

ltOJ

l 1 9gt

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I $t I II ~

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26

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6(2)1 48

~ ElUW11 ~~1rlJdl~ ~Vi6l V1~U91lll lbi1 rllVi6l El1IJflnJ1U1Yi 2532 niJYit7J6lEll Co bull

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ilf 48 (3)5-10

Nutaphand W 1 The Turtles Mitbhadung Press 222pp

  • การศกษาชววทยาและการเพาะพนธเตาหวาย
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26 28 3 32 34 36 38

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34

32

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28

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3 31 32 33 34 35 36 37 38

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6(2)1 48

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ilf 48 (3)5-10

Nutaphand W 1 The Turtles Mitbhadung Press 222pp

  • การศกษาชววทยาและการเพาะพนธเตาหวาย
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6(2)1 48

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ilf 48 (3)5-10

Nutaphand W 1 The Turtles Mitbhadung Press 222pp

  • การศกษาชววทยาและการเพาะพนธเตาหวาย
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Nutaphand W 1 The Turtles Mitbhadung Press 222pp

  • การศกษาชววทยาและการเพาะพนธเตาหวาย
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Nutaphand W 1 The Turtles Mitbhadung Press 222pp

  • การศกษาชววทยาและการเพาะพนธเตาหวาย
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Nutaphand W 1 The Turtles Mitbhadung Press 222pp

  • การศกษาชววทยาและการเพาะพนธเตาหวาย
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ilf 48 (3)5-10

Nutaphand W 1 The Turtles Mitbhadung Press 222pp

  • การศกษาชววทยาและการเพาะพนธเตาหวาย
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ilf 48 (3)5-10

Nutaphand W 1 The Turtles Mitbhadung Press 222pp

  • การศกษาชววทยาและการเพาะพนธเตาหวาย
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Nutaphand W 1 The Turtles Mitbhadung Press 222pp

  • การศกษาชววทยาและการเพาะพนธเตาหวาย
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Nutaphand W 1 The Turtles Mitbhadung Press 222pp

  • การศกษาชววทยาและการเพาะพนธเตาหวาย