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Morphometric Relationships of Marine Fishes Common toCentral California and the Southern California BightAuthor(s): Chelsea M. Williams, Jonathan P. Williams, Jeremy T. Claisse,Daniel J. PondellaII, Michael L. Domeier, and Laurel A. ZahnSource: Bulletin, Southern California Academy of Sciences, 112(3):217-227.2013.Published By: Southern California Academy of SciencesDOI: http://dx.doi.org/10.3160/0038-3872-112.3.217URL: http://www.bioone.org/doi/full/10.3160/0038-3872-112.3.217
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Morphometric Relationships of Marine Fishes Common toCentral California and the Southern California Bight
Chelsea M. Williams,1* Jonathan P. Williams,1 Jeremy T. Claisse,1
Daniel J. Pondella II.,1 Michael L. Domeier,2,3 and Laurel A. Zahn1
1Vantuna Research Group, Department of Biology, Occidental College, 1600 Campus
Road, Los Angeles, CA 90041, USA2California Department of Fish and Wildlife, 1416 9th Street, 12th Floor, Sacramento,
CA 95814, USA3(Current affiliation) Marine Conservation Science Institute, 68-1825 Lina Poepoe
Street, Waikoloa, HI 96738, USA
Length-weight relationships have several applications in fish stock assessments and
ecological studies (e.g., Ricker 1975, Newman et al. 2006). Particularly, they are
important for visual surveys of fish populations where the estimated total lengths are
converted to weights to estimate fish biomass (e.g., Hamilton et al. 2010, Sala et al. 2012).
The available information on length-weight relationships and length-length conversions
for marine fishes in California are mostly limited to commercial catch (RecFIN 2009) or
the occasional ecological survey (Miller et al. 2008), and a recent compilation of these
parameters (Cailliet et al. 2000) demonstrated many species are lacking this basic
information. Fishes used in this study were collected in various large- and small-scale
projects by the Vantuna Research Group, Occidental College and California State
University Northridge from 1984 to 2012. These included state-mandated programs
dedicated to assessing the biological and economic impacts of its stocking efforts
(ORHEP) and localized fisheries surveys (San Diego and Morro Bay) where a variety of
species were caught. Measurements of lengths and weights provide the opportunity to
generate information on morphometric relationships that will be useful to other
researchers. Here we provide standard length (SL) to total length (TL) conversions
(Table 1) for 32 near-shore marine fish species (Class Actinopterygii) and length-weight
equation parameters (Table 2) for 71 near-shore marine fish species (57 from Class
Actinopterygii and 14 from Subclass Elasmobranchii), common to central and southern
California (Miller and Lea 1972).
Fishes were collected by several methods. (White Seabass Gill Net Survey) Collections
using monofilament gill nets were made at 19 stations dispersed throughout the Southern
California Bight from 1995–2005 in shallow (5–14 m) depths at the edge of rocky reefs as
part of the Nearshore Gill Net Sampling Program for White Seabass (Age I-IV). For
detailed methods see Pondella and Allen (2000). (San Diego Bay Fisheries Inventory and
Utilization Surveys) Fish assemblages in San Diego Bay were assessed using a variety of
methods (large seine, small seine, square enclosure, purse seine, beam trawl and otter
trawl) (Allen et al. 2002). The bay is divided into four unique ecoregions that were
sampled in April and July of 2005, 2008 and 2012, and by purse seine and square
enclosure only in June 2009. (Morro Bay Fish Survey) Fish populations were surveyed in
Morro Bay using methods similar to the San Diego Bay Fisheries Inventory and
* Corresponding author: [email protected]
Bull. Southern California Acad. Sci.112(3), 2013, pp. 217–227
E Southern California Academy of Sciences, 2013
217
Table 1. Standard length (SL; in mm) to total length (TL; in mm) conversion parameters (see main text
for equation description), sample size (N) and length characteristics of the sampled population for 32 fish
species (Class Actinopterygii) common to southern and central California.
length characteristics parameters of the relationship
Scientific Name Survey N
Min.
(mm)
Max.
(mm) a b R2
Clupeiformes
Engraulidae - anchovies
Anchoa compressa (Girard, 1858) b 63 45 114 7.22 1.16 0.95
Anchoa delicatissima (Girard, 1854) b 633 19 67 2.42 1.16 0.91
Aulopiformes
Synodontidae - lizardfishes
Synodus lucioceps (Ayers, 1955) abc 20 68 193 21.66 1.15 0.99
Batrachoidiformes
Batrachoididae - toadfishes
Porichthys myriaster Hubbs & Schultz, 1939 ab 23 16 327 20.44 1.16 0.99
Atheriniformes
Atherinopsidae - New World silversides
Atherinops affinis (Ayers, 1860) abc 1745 11 193 20.02 1.19 . 0.99
Atherinopsis californiensis (Girard, 1854) abc 11 31 282 22.33 1.22 . 0.99
Leuresthes tenuis (Ayers, 1860) bc 16 62 166 1.36 1.15 0.99
Beloniformes
Hemiramphidae - halfbeaks
Hyporhamphus rosae (Jordan & Gilbert, 1880) b 14 24 127 20.17 1.13 0.99
Belonidae - needlefishes
Strongylura exilis (Girard, 1854) ab 3 51 337 1.22 1.08 . 0.99
Cyprinodontiformes
Fundulidae - topminnows
Fundulus parvipinnis Girard, 1854 bc 195 14 78 1.91 1.11 0.99
Gasterosteiformes
Syngnathidae - pipefishes
Syngnathus leptorhynchus Girard, 1854 bc 799 33 248 1.20 1.02 . 0.99
Scorpaeniformes
Scorpaenidae - scorpionfishes
Scorpaena guttata Girard, 1854 abd 3 85 230 1.15 1.23 . 0.99
Cottidae - sculpins
Leptocottus armatus Girard, 1854 abc 763 11 119 0.63 1.16 0.99
Perciformes
Polyprionidae – wreckfishes
Stereolepis gigas Ayers, 1859 ag 35 336 1450 210.87 1.21 0.99
Serranidae – sea basses
Paralabrax clathratus (Girard, 1854) abcd 76 19 165 1.31 1.19 0.99
Paralabrax maculatofasciatus
(Steindachner, 1868) abd 348 32 306 1.71 1.22 0.99
Paralabrax nebulifer (Girard, 1854) abd 154 43 180 4.77 1.15 0.98
Haemulidae - grunts
Anisotremus davidsonii (Steindachner, 1876) abde 623 20 328 5.78 1.23 0.99
Sciaenidae – drums and croakers
Atractoscion nobilis (Ayers, 1860) abf 6513 71 1220 11.29 1.15 0.99
218 SOUTHERN CALIFORNIA ACADEMY OF SCIENCES
Utilization Surveys in April, August and November of 2005–2007 and in May of 2008.
(Cryptic reef fish collections from King Harbor, Redondo Beach and Agua Hedionda, San
Diego) Collections of cryptic benthic fishes in King Harbor, Redondo Beach have been
made periodically (1–12 times per year) since 1984 by divers using anesthetic and air lifts
(Stephens et al. 1994). A similar collection was made from Agua Hedionda Lagoon in
2005. (Heat Treat and Impingement Surveys) Samples were also collected during heat
treatments in 2005 at Encina Generating Station, Cabrillo Power Plant, and Huntington
Beach Generating Station. For detailed methods see Pondella et al. (2008). Some white
seabass (Atractoscion nobilis) specimens were also collected opportunistically by hook
and line or spear. Additionally, data for giant sea bass (Stereolepis gigas) collected by
hook and line was included (Michael Domeier, pers. comm.). While fishes caught during
some of these studies were batch weighed by species, all individuals used here were
measured individually: TL and/or SL or disc width (DW) were typically recorded to the
nearest millimeter (mm) or occasionally centimeter (cm) and weight was recorded to the
length characteristics parameters of the relationship
Scientific Name Survey N
Min.
(mm)
Max.
(mm) a b R2
Cheilotrema saturnum (Girard, 1858) ab 6 35 178 1.40 1.19 . 0.99
Roncador stearnsii (Steindachner, 1876) abe 493 138 542 11.85 1.20 0.99
Embiotocidae – sea perches
Cymatogaster aggregata Gibbons, 1854 abc 828 26 112 1.83 1.24 0.98
Micrometrus minimus (Gibbons, 1854) abc 43 21 113 3.35 1.19 0.99
Blenniidae – combtooth blennies
Hypsoblennius gentilis (Girard, 1854) bcd 20 44 109 3.99 1.15 0.99
Clinidae – kelp blennies
Gibbonsia elegans (Cooper, 1864) bd 10 23 106 20.37 1.15 . 0.99
Heterostichus rostratus Girard, 1854 abcd 329 23 315 1.23 1.13 . 0.99
Gobiidae - gobies
Clevelandia ios (Jordan & Gilbert, 1882) bc 310 10 56 0.37 1.16 0.99
Quietula y-cauda (Jenkins & Evermann,
1889) bc 64 22 64 1.17 1.16 0.97
Pleuronectiformes
Paralichthyidae – sand flounders
Citharichthys stigmaeus Jordan &
Gilbert, 1882 abc 263 22 101 0.65 1.17 0.99
Paralichthys californicus (Ayers, 1859) abc 62 57 430 8.26 1.16 0.98
Pleuronectidae – right-eyed flounders
Pleuronicthys guttulatus Girard, 1854 abc 48 17 204 2.31 1.21 . 0.99
Pleuronichthys ritteri Starks & Morris, 1907 abd 24 78 156 0.17 1.25 0.97
Cynoglossidae - tonguefishes
Symphurus atricaudus (Jordan & Gilbert,
1880) ab 18 64 138 20.88 1.08 . 0.99
a, White Seabass Gill Net Survey; b, San Diego Bay Fisheries Inventory and Utilization Survey; c,
Morro Bay Fish Survey; d, Cryptic reef fish collections from King Harbor and Agua Hedionda; e, Heat
Treatments from Encina Generating Station, Cabrillo Power Plant, and Huntington Beach Generation
Station; f,Opportunistic non-scientific hook and line and spear catches; g, Data provided by Michael
Domeier
Table 1. Continued.
MORPHOMETRIC RELATIONSHIPS OF CAIFORNIA MARINE FISHES 219
Tab
le2
.L
eng
tha
nd
wei
gh
tp
ara
met
ers,
sam
ple
size
(N),
inp
ut
len
gth
typ
e,a
nd
len
gth
cha
ract
eris
tics
of
the
sam
ple
for
14
cart
ilag
ino
us
fish
spec
ies
(Su
bcl
ass
Ela
smo
bra
nch
ii)
an
d57
ray-f
inn
edfi
shsp
ecie
s(C
lass
Act
ino
pte
rygii
)co
mm
on
toso
uth
ern
an
dce
ntr
al
Cali
forn
ia.
length
chara
cter
isti
cspara
met
ers
of
the
rela
tionsh
ip
Sci
enti
fic
Na
me
Su
rvey
NT
yp
eM
in.
(m
m)
Ma
x.
(m
m)
ab
R2
Het
ero
do
nti
form
es
Het
ero
do
nti
da
e-
bu
llh
ead
shark
s
Het
ero
do
ntu
sfr
an
cisc
i(G
irard
,1
85
5)
ab
65
1T
L1
70
87
01
.18
E-0
52
.94
0.9
8
La
mn
ifo
rmes
Alo
pii
da
e-
thre
sher
shark
s
Alo
pia
svu
lpin
us
(Bo
nn
ate
rre,
17
88)
a1
6T
L8
70
25
60
3.1
1E
-04
2.3
60
.90
Ca
rch
arh
inif
orm
es
Scy
lio
rhin
ida
e-
cat
shark
s
Cep
ha
losc
yll
ium
ven
trio
sum
(Ga
rma
n,
18
80)
a3
07
TL
33
59
50
3.0
0E
-06
3.1
30
.92
Tri
ak
ida
e-
ho
un
dsh
ark
s
Gale
orh
inus
gale
us
(Lin
na
eus,
17
58)
a1
02
TL
25
01
90
07
.78
E-0
62
.93
0.9
5
Mu
stel
us
cali
forn
icu
sG
ill,
18
64
ab
44
1T
L3
45
12
00
7.2
7E
-07
3.2
20
.95
Mu
stel
us
hen
lei
(Gil
l,1
86
3)
a3
87
TL
34
01
10
08
.07
E-0
73
.21
0.9
5
Tri
ak
isse
mif
asc
iata
Gir
ard
,1
85
5a
73
6T
L1
60
15
45
5.9
5E
-06
2.9
50
.97
Hex
an
chif
orm
es
Hex
an
chid
ae
-co
wsh
ark
s
No
tory
nch
us
cep
edia
nu
s(B
on
na
terr
e,1
78
8)
a3
0T
L1
18
51
90
08
.61
E-0
73
.22
0.8
8
Sq
ua
lifo
rmes
Sq
ua
lid
ae
-d
og
fish
shark
s
Squ
alu
sa
can
thia
sL
inn
aeu
s,1
75
8a
19
1T
L3
20
12
00
8.2
9E
-08
3.5
70
.94
Sq
uati
nif
orm
es
Sq
ua
tin
ida
e-
an
gel
shark
s
Squati
na
cali
forn
ica
Ay
ers,
18
59
ab
20
6T
L3
20
12
00
7.8
1E
-06
3.0
20
.94
220 SOUTHERN CALIFORNIA ACADEMY OF SCIENCES
length
chara
cter
isti
cspara
met
ers
of
the
rela
tionsh
ip
Sci
enti
fic
Na
me
Su
rvey
NT
yp
eM
in.
(m
m)
Ma
x.
(m
m)
ab
R2
Ra
jifo
rmes
Rh
ino
bati
da
e–
gu
ita
rfis
hes
Rh
ino
ba
tos
pro
du
ctus
Ay
res,
18
54
ab
11
1T
L2
25
13
40
3.4
3E
-06
3.0
10
.95
My
lio
ba
tifo
rmes
Uro
try
go
nid
ae
-A
mer
ica
nro
un
dst
ingra
ys
Uro
ba
tis
ha
ller
i(C
oo
per
,1
86
3)
ab
c5
56
DW
74
27
55
.73
E-0
53
.02
0.9
6
Gy
mn
uri
da
e-
bu
tter
fly
ray
s
Gy
mn
ura
ma
rmo
rata
(Co
op
er,
18
64)
ab
26
DW
25
01
01
02
.74
E-0
63
.20
0.9
8
My
lio
bati
da
e-
eagle
ray
sa
nd
ma
nta
s
My
lio
ba
tis
cali
forn
ica
Gil
l,1
86
5a
bc
27
0D
W1
80
10
00
2.3
2E
-05
2.9
40
.95
Alb
uli
form
es
Alb
uli
da
e-
bo
nef
ish
es
Alb
ula
gil
ber
tiP
feil
er&
Va
nd
erH
eid
en,
20
11
ab
42
SL
55
36
02
.05
E-0
73
.77
0.9
8
Clu
pei
form
es
En
gra
uli
da
e-
an
cho
vie
s
An
cho
aco
mp
ress
a(G
irard
,1
85
8)
b6
3S
L4
51
14
1.1
1E
-05
3.0
50
.94
An
cho
ad
elic
ati
ssim
a(G
ira
rd,
18
54)
b6
37
SL
19
67
7.5
5E
-06
3.0
90
.90
Clu
pei
da
e-
her
rin
gs
Sard
ino
ps
sag
ax
(Jen
yn
s,1
84
2)
ab
c2
47
SL
75
28
01
.19
E-0
53
.02
0.9
1
Au
lop
ifo
rmes
Sy
no
do
nti
da
e–
liza
rdfi
shes
Syn
od
us
luci
oce
ps
(Ay
res,
18
55)
ab
c8
0S
L6
13
60
2.1
2E
-06
3.2
60
.98
Batr
ach
oid
ifo
rmes
Ba
tra
cho
idid
ae
-to
ad
fish
es
Pori
chth
ys
my
ria
ster
Hu
bb
s&
Sch
ult
z,1
93
9a
b3
28
SL
16
49
01
.38
E-0
52
.98
0.9
9
Pori
chth
ys
no
tatu
sG
irard
,1
85
4a
b7
7S
L2
34
60
2.2
1E
-05
2.9
20
.92
Tab
le2
.C
on
tin
ued
.MORPHOMETRIC RELATIONSHIPS OF CAIFORNIA MARINE FISHES 221
Tab
le2
.C
on
tin
ued
.
length
chara
cter
isti
cspara
met
ers
of
the
rela
tionsh
ip
Sci
enti
fic
Na
me
Surv
eyN
Typ
eM
in.
(m
m)
Ma
x.
(m
m)
ab
R2
Mu
gil
ifo
rmes
Mu
gil
ida
e-
mu
llet
s
Mu
gil
cep
ha
lus
Lin
na
eus,
17
58
ab
c4
0S
L5
35
20
2.9
6E
-05
2.9
30
.99
Ath
erin
ifo
rmes
Ath
erin
op
sid
ae
-N
ewW
orl
dsi
lver
sid
es
Ath
erin
op
sa
ffin
is(A
yer
s,1
86
0)
ab
c1
75
6S
L1
11
93
3.8
3E
-06
3.2
40
.99
Ath
erin
op
sis
cali
forn
ien
sis
(Gir
ard
,1
85
4)
ab
c4
20
SL
31
36
01
.03
E-0
53
.01
0.9
4
Leu
rest
hes
ten
uis
(Ay
ers,
18
60)
bc
17
SL
32
16
68
.25
E-0
73
.52
0.9
8
Bel
on
ifo
rmes
Hem
iram
ph
idae
-h
alf
bea
ks
Hy
po
rha
mp
hu
sro
sae
(Jo
rdan
&G
ilb
ert,
1880)
b18
SL
24
127
6.2
3E
-07
3.3
20.9
4
Bel
on
ida
e-
nee
dle
fish
es
Str
on
gylu
raex
ilis
(Gir
ard
,1
85
4)
ab
10
SL
51
76
01
.36
E-0
73
.39
.0
.99
Cy
pri
no
do
nti
form
es
Fu
nd
uli
da
e-
top
min
no
ws
Fundulu
sparv
ipin
nis
Gir
ard
,1854
bc
198
SL
14
78
1.4
9E
-05
3.0
80.9
9
Ga
ster
ost
eifo
rmes
Sy
ng
na
thid
ae
-p
ipef
ish
es
Sy
ng
na
thu
sle
pto
rhy
nch
us
Gir
ard
,1854
bc
814
SL
33
248
8.4
5E
-08
3.3
50.9
5
Sco
rpaen
ifo
rmes
Sco
rpa
enid
ae
-sc
orp
ion
fish
es
Sco
rpa
ena
gu
tta
taG
irard
,1
85
4a
bd
61
8S
L3
03
05
3.6
6E
-05
2.9
80
.94
Seb
ast
esatr
ovi
rens
(Jo
rdan
&G
ilb
ert,
1880)
a102
SL
125
270
9.4
4E
-06
3.2
20.9
2
Seb
ast
esca
rna
tus
(Jo
rdan
&G
ilb
ert,
1880)
a36
SL
110
240
1.7
2E
-04
2.6
80.9
1
Seb
ast
esra
stre
llig
er(J
ord
an
&G
ilb
ert,
1880)
ab
225
SL
52
350
2.9
3E
-05
3.0
10.9
1
222 SOUTHERN CALIFORNIA ACADEMY OF SCIENCES
Tab
le2
.C
on
tin
ued
.
length
chara
cter
isti
cspara
met
ers
of
the
rela
tionsh
ip
Sci
enti
fic
Na
me
Surv
eyN
Typ
eM
in.
(m
m)
Ma
x.
(m
m)
ab
R2
Hex
agra
mm
ida
e-
gre
enli
ng
s
Ox
yle
biu
sp
ictu
sG
ill,
18
62
ad
17
SL
27
15
01
.52
E-0
53
.11
0.9
8
Co
ttid
ae
-sc
ulp
ins
Art
ediu
sco
rall
inus
(Hu
bb
s,1
92
6)
d2
0S
L1
25
93
.19
E-0
52
.88
0.9
9
Lep
toco
ttu
sa
rmatu
sG
irard
,1
85
4a
bc
82
7S
L1
11
90
2.3
2E
-05
2.9
80
.99
Ru
sca
riu
scr
ease
ri(H
ub
bs,
19
26)
d1
74
SL
10
44
1.4
9E
-05
3.1
70
.99
Sco
rpa
enic
hth
ys
ma
rmo
ratu
s(A
yre
s,1
85
4)
ad
13
6S
L4
43
90
2.0
6E
-05
3.0
60
.95
Per
cifo
rmes
Po
lyp
rio
nid
ae
-w
reck
fish
es
Ste
reole
pis
gig
as
Ay
ers,
18
59
ag
96
SL
12
52
00
31
.07
E-0
42
.80
0.9
9
Ser
ran
ida
e-
sea
ba
sses
Pa
rala
bra
xcl
ath
ratu
s(G
irard
,1
85
4)
ab
cd6
36
SL
19
57
52
.09
E-0
53
.01
0.9
8
Pa
rala
bra
xm
acu
lato
fasc
iatu
s
(Ste
ind
ach
ner
,1
86
8)
ab
d4
30
SL
32
35
02
.16
E-0
53
.03
0.9
9
Pa
rala
bra
xn
ebu
life
r(G
irard
,1
85
4)
ab
d6
35
SL
27
41
02
.89
E-0
52
.95
0.9
9
Ha
emu
lid
ae
-g
run
ts
Anis
otr
emus
davi
dso
nii
(Ste
ind
ach
ner
,1
87
6)
ab
de
75
0S
L2
03
30
1.6
4E
-05
3.1
30
.98
Sci
aen
ida
e-
dru
ms
an
dcr
oa
ker
s
Atr
act
osc
ion
no
bil
is(A
yre
s,1
86
0)
ab
f6
54
8S
L7
11
22
02
.97
E-0
52
.87
0.9
6
Chei
lotr
ema
satu
rnum
(Gir
ard
,1
85
8)
ab
34
4S
L3
53
65
3.8
3E
-05
2.9
10
.92
Cy
no
scio
np
arv
ipin
nis
Ay
res,
18
61
b1
0S
L1
79
46
56
.52
E-0
52
.74
0.9
9
Men
tici
rrhus
undula
tus
(Gir
ard
,1
85
4)
ab
43
2S
L1
65
52
02
.58
E-0
52
.91
0.9
4
Ro
nca
dor
stea
rnsi
i(S
tein
da
chn
er,
18
76
)a
be
57
7S
L1
38
54
23
.96
E-0
52
.91
0.9
6
Ky
ph
osi
da
e-
sea
chu
bs
Her
mosi
lla
azu
rea
Jen
kin
s&
Ever
ma
nn
,1
88
9a
53
SL
16
53
10
1.1
4E
-05
3.2
10
.85
MORPHOMETRIC RELATIONSHIPS OF CAIFORNIA MARINE FISHES 223
Tab
le2
.C
on
tin
ued
.
len
gth
chara
cter
isti
csp
ara
met
ers
of
the
rela
tio
nsh
ip
Sci
enti
fic
Nam
eS
urv
eyN
Typ
eM
in.
(m
m)
Max
.(m
m)
ab
R2
Em
bio
toci
da
e-
surf
per
ches
Cy
ma
tog
ast
era
gg
reg
ata
Gib
bo
ns,
18
54
ab
c8
79
SL
26
14
02
.08
E-0
53
.07
0.9
7
Mic
rom
etru
sm
inim
us
(Gib
bo
ns,
18
54
)a
bc
88
SL
21
13
52
.91
E-0
53
.02
0.9
6
Po
ma
cen
trid
ae
–d
am
self
ish
es
Ch
rom
isp
un
ctip
inn
is(C
oo
per
,1
86
3)
ad
14
8S
L2
42
10
2.6
9E
-05
3.0
20
.98
Hy
psy
po
ps
rub
icu
ndu
s(G
irard
,1
85
4)
a1
39
SL
90
22
03
.07
E-0
53
.11
0.8
8
La
bri
da
e-
wra
sses
an
dp
arr
otf
ish
es
Sem
icoss
yphus
pulc
her
(Ay
res,
18
54)
a1
94
SL
45
45
58
.45
E-0
52
.80
0.9
2
Ble
nn
iid
ae
-co
mb
too
thb
len
nie
s
Hy
pso
ble
nn
ius
gen
tili
s(G
irard
,1
85
4)
bcd
30
SL
34
10
91
.58
E-0
53
.10
0.9
5
Hy
pso
ble
nn
ius
jen
kin
si(J
ord
an
&E
ver
ma
nn
,
18
96)
d7
3S
L1
38
08
.64
E-0
63
.20
0.9
8
Cli
nid
ae
-k
elp
ble
nn
ies
Gib
bo
nsi
ael
ega
ns
(Co
op
er,
18
64)
bd
11
5S
L1
61
07
4.8
0E
-06
3.2
40
.99
Het
erost
ichus
rost
ratu
sG
irard
,1
85
4a
bcd
52
1S
L2
34
00
3.9
7E
-06
3.1
70
.99
Lab
riso
mid
ae
-la
bri
som
idb
len
nie
s
Para
clin
us
inte
gri
pin
nis
(Sm
ith
,1
88
0)
bcd
48
9S
L9
58
1.1
1E
-05
3.1
00
.98
Go
bie
soci
da
e-
clin
gfi
shes
Go
bie
sox
rhes
sod
on
Sm
ith
,1
88
1d
42
SL
72
52
.11
E-0
53
.04
0.9
6
Go
bii
dae
-go
bie
s
Cle
vela
ndia
ios
(Jo
rdan
&G
ilb
ert,
1882)
bc
329
SL
65
62.5
5E
-06
3.4
20.9
5
Qu
ietu
lay
-ca
ud
a(J
enk
ins
&E
ver
ma
nn
,1
88
9)
bc
64
SL
22
64
1.2
2E
-05
3.0
50
.95
Rh
ino
go
bio
ps
nic
ho
lsii
(Bea
n,
18
82)
d5
8S
L2
48
66
.28
E-0
63
.26
0.9
9
Sp
hyra
enid
ae
-b
arr
acu
da
s
Sp
hy
raen
aa
rgen
tea
Gir
ard
,1854
ab
436
SL
300
965
2.4
6E
-05
2.7
60.9
4
Sco
mb
rid
ae
-m
ack
erel
s
Sco
mb
erja
po
nic
us
Ho
ttu
yn
,1
78
2a
b2
74
SL
16
03
95
1.9
3E
-05
2.9
30
.92
224 SOUTHERN CALIFORNIA ACADEMY OF SCIENCES
length
chara
cter
isti
cspara
met
ers
of
the
rela
tionsh
ip
Sci
enti
fic
Na
me
Surv
eyN
Typ
eM
in.
(m
m)
Ma
x.
(m
m)
ab
R2
Ple
uro
nec
tifo
rmes
Pa
rali
chth
yid
ae
-sa
nd
flo
un
der
s
Cit
ha
rich
thy
sst
igm
aeu
sJo
rdan
&G
ilb
ert,
1882
ab
c285
SL
22
135
8.3
3E
-06
3.2
00.9
8
Para
lich
thys
cali
forn
icus
(Ay
ers,
18
59)
ab
c6
23
SL
57
81
02
.55
E-0
52
.91
0.9
5
Xy
stre
ury
sli
ole
pis
Jord
an
&G
ilb
ert,
1880
ab
36
SL
105
400
3.0
7E
-06
3.3
60.9
4
Ple
uro
nec
tid
ae
-ri
gh
t-ey
efl
ou
nd
ers
Ple
uro
nic
thy
sg
utt
ula
tus
Gir
ard
,1856
ab
c118
SL
17
340
5.8
0E
-05
2.8
50.9
9
Ple
uro
nic
hth
ys
ritt
eri
Sta
rks
&M
orr
is,
19
07
ab
d1
07
SL
17
23
02
.95
E-0
52
.98
0.9
6
Cy
no
glo
ssid
ae
-to
ng
uef
ish
es
Sy
mp
hu
rus
atr
ica
ud
us
(Jo
rdan
&G
ilb
ert,
1880)
ab
32
SL
34
185
1.0
7E
-05
3.0
00.9
4
a,
Wh
ite
Sea
bass
Gil
lN
etS
urv
ey;
b,
San
Die
go
Bay
Fis
her
ies
Inven
tory
an
dU
tili
zati
on
Su
rvey
;c,
Mo
rro
Bay
Fis
hS
urv
ey;
d,
Cry
pti
cre
effi
shco
llec
tio
ns
fro
mK
ing
Ha
rbo
ra
nd
Ag
ua
Hed
ion
da
;e,
Hea
tT
rea
tmen
tsfr
om
En
cin
aG
ener
ati
ng
Sta
tio
n,
Ca
bri
llo
Po
wer
Pla
nt,
an
dH
un
tin
gto
nB
each
Gen
era
tio
nS
tati
on
;f,
Op
po
rtu
nis
tic
no
n-
scie
nti
fic
ho
ok
an
dli
ne
an
dsp
ear
catc
hes
;g
,D
ata
pro
vid
edb
yM
ich
ael
Do
mei
er
Tab
le2
.C
on
tin
ued
.
MORPHOMETRIC RELATIONSHIPS OF CAIFORNIA MARINE FISHES 225
nearest gram (g) either in the field or from frozen specimens that were brought back to
the laboratory.
All statistical analyses were performed using R (R Core Development Team 2012).
Standard length to total length conversion equations were established using linear
regression analyses. Length-length models were fitted to the equation TL 5 a+bSL where
SL is standard length (mm) and TL is total length (mm) (Table 1). Length-weight models
were fitted to the equation W 5 aLb, where W is the wet body weight (g) and L is the total
length (mm) or disc width (mm) (Table 2) by log-transforming both the length and weight
data, performing linear regression analyses. Estimated parameters were then back-
transformed to the original scale for reporting. Obvious outliers were removed prior to
model fitting. While some species had a low sample size (N , 30), we report parameters
here for those where 1) the naturally occurring size range was adequately represented in
the sample, 2) the models fit the data well (Tables 1, 2), and 3) the lack of published
information on the species made the parameter estimates of high value (Froese 2006).
Parameters for some species described here have been previously published (e.g. Miller
et al. 2008, Love 2011). However, there is value in including parameters for all species
that we had sufficient data for where sampling locations differ across studies and/or
larger sample sizes were available, permitting future users of the parameters more options
depending on their intended use.
Acknowledgements
Funding for data collection was provided by the following agencies: California
Department of Fish and Wildlife’s Ocean Resources Enhancement Hatchery Program
(ORHEP), Port of San Diego, San Luis Obispo Science and Ecosystem Alliance
(SLOSEA), and NRG Cabrillo Power Operations Inc.
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