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_____________Mun. Ent. Zool. Vol. 1, No. 2, June 2006__________ I

MUNIS

ENTOMOLOGY & ZOOLOGY

Ankara / Turkey

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Scope: Munis Entomology & Zoology publishes a wide variety of papers on all aspects of Entomology and Zoology from all over the world, including studies mainly on systematics, taxonomy, nomenclature, fauna, biogeography, biodiversity, ecology, morphology, behavior, conservation, paleobiology and other aspects. Submission of Manuscripts: Works published or under consideration elsewhere (including on the Internet) will not be accepted. At first submission, one double spaced hard copy (text and tables) with figures (not original) must be sent to the Editors, Dr. Hüseyin Özdikmen for publication in MEZ. All manuscripts should be submitted as a Word file or PDF file in an e-mail attachment. If electronic submission is not possible due to limitations of electronic space at the sending or receiving end, unavailability of e-mail, etc., we will accept “hard” versions, in triplicate, accompanied by an electronic version stored on floppy disk or CD-ROM. Review Process: When submitting manuscripts, all authors must provide the name of at least three qualified experts (along with provide their address, subject fields and e-mails). Then, the editors send to experts for review. The review process should normally be completed within 45-60 days. After reviewing papers rejected papers are discarded. For accepted papers, authors are asked to modify their papers according to suggestions of the reviewers and editors. Final versions of manuscripts and figures are needed in a digital format.

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_____________Mun. Ent. Zool. Vol. 1, No. 2, June 2006__________ 173

FAUNISTICAL AND SYSTEMATICAL STUDIES ON THE TRYPETINAE (DIPTERA: TEPHRITIDAE) IN THE

TURKEY ALONG WITH A NEW RECORD TO TURKISH FAUNA

Murat Kütük* and Mehmet Özaslan**

* University of İnönü, Faculty of Science and Art, Department of Biology, 44280 Malatya, TURKEY, E-mail: [email protected] ** University of Gaziantep, Faculty of Science and Art, Department of Biology, 27310 Malatya, TURKEY, E-mail: [email protected] [Kütük, M. & Özaslan, M. 2006. Faunistical and systematical studies on the Trypetinae (Diptera: Tephritidae) in Turkey along with a new record to Turkish fauna. Munis Entomology & Zoology 1 (2): 173-178] ABSTRACT: This study is based on material collected in Turkey in of 1999 - 2005. Four genera and five species of Trypetinae were found to occur in Turkey. Rhagoletis berberidis Jermy is a new record for fauna for Turkey. Key to genera and species of this subfamily occurring in Turkey are provided. The locality and host plants of specimens examined and figures of aculei and wings of species are given. Distribution of the species in Turkey and rest of world stated. KEY WORDS: Tephritidae, New Record, Key, Systematics, Fauna, Turkey.

The subfamily Trypeinae is widespread in the world. Twenty nine

genera and forty three species have been listed from Europe (Merz, 1994). Trypetinae differ from other Tephritidae by the following combination of characters [only the major characters are listed; more complete lists of characters can be found in Freidberg & Kugler (1989) and Merz (1994)]: Postoculer setae acuminate, usually black or brown rarely paler. Frontal stripe usually hairy; arista bare, hairy or plumose; proboscis almost always capitate, with broad labella; scapular setae usually distinct; dorsocentral setae usually situated at about level of anterior supra-alar setae; scutellum flat or convex; wing pattern often with transverse or oblique bands, rarely reticulate; abdomen without distinct pollinosity or with distinct pattern of pollinosity; sixth tergum of female usually much shorter than fifth.

Larvae infest fleshy plants that have fruits and some develop in flower buds, stems or leaves (Freidberg & Kugler, 1989).

MATERIAL AND METHODS

The specimens were collected in Turkey in 1999 - 2005. Flies were

killed in a killing jar with potassium cyanide. The material is deposited in the museum of İnönü University, Department of Biology, Malatya.

Species were identified using the keys by Freidberg & Kugler (1989) and Merz (1994). Terminology follows White et al. (2000). Extensive synonymy and bibliography is provided by Norrbom et al. (1999). Host


plants were determined by Prof. Şemsettin Civelek (University of Fırat, Department of Biology, Elazığ, Turkey).

RESULTS

Four genera (Ceratitis Mac Leay Euleia Walker, Myopardalis Bigot

and Rhagoletis Loew) and their five species (Ceratitis capitata Wiedemann, Euleia heraclei Linnaeus, Carpomya pardalina Bigot, Rhagoletis berberidis Jermy and Rhagoletis cerasi Linnaeus) were collected from Turkey between the years 1999 - 2005. Rhagoletis berberidis is a new record for fauna of Turkey. Distribution of the species in Turkey and rest of world was stated.

KEY TO GENERA OF TRYPETINAE

1. Wing pattern ceratitis type, with 3 bands as well as several small

black spots at basal 0.4 (Fig. 2) …..……………….. Ceratitis Mac Leay - Wing pattern different, lacking small black spots at basal (Figs. 1, 3,

4, 5) ………………………………………………………………………………….……2 2. Wing with 3-4 transverse bands (Figs. 3, 4, 5); antenna with distinct

apicodorsal point; head not distinctly oval; fronto-facial angle about 90 0 and rather projecting……………………………………….3

- Wing not with transverse bands, and different (Fig. 3); antenna rounded at apex; head oval; fronto-facial angle broader and rounded………………………………………… ……...…………. Euleia Walker

3. Hind femur with strong anteroventral subapical setae; vein R4+5 usually without dorsal setulae ………………………… Rhagoletis Loew

- Hind femur without strong anteroventral subapical setae; vein R4+5 with dorsal setulae …………………………….……..…… Carpomya Costa

Carpomya pardalina Bigot, 1891

(Figs. 1, 6) Specimens examined: 12 ♂♂, 15 ♀♀, Adana, Balcalı, 37.02 N, 35.22 E, 60 m, 17.X.2000; 2 ♂♂, 2 ♀♀, Hatay, Kırıkhan, Çiloğlan, 36.30 N, 36.26 E, 110 m, 08.V.2002 (M. Kütük). Host plants: in Freidberg & Kugler, 1989. Distribution: Egypt, Israel, Turkey, Kenya, and Caucasus to W. Indian (Giray, 1979; Freidberg & Kugler, 1989; Norrbom et al. 1999). Measurements (length in mm): Male Body 5.0-6.4, wing 4.0-4.6. Female body: 6.3-7.3, wing 4.5-5.3.

Ceratitis capitata (Wiedemann, 1824) (Figs. 2, 7)

Specimens examined: 27 ♂♂, 35 ♀♀, Adana, Balcalı, 37.02 N, 35.22 E, 60 m, 17.X.2000; 24 ♂♂, 20 ♀♀, Adana, Balcalı, 37.01 N, 35.21 E, 70 m, 16.X.2001; 4 ♂♂, 7 ♀♀, Antalya, Finike, 36.23 N, 30.24 E, 30 m,


20.VI.1999; 5 ♂♂, 3 ♀♀, Antalya, Finike, 36.27 N, 30.12 E, 55 m, 24.VI.2000; 2 ♂♂, 1 ♀, Antalya, Alanya, 36.33 N, 30.12 E, 100 m, 11.VII.2000; 5 ♂♂, 5 ♀♀, Hatay, Kırıkhan, Çiloğlan, 36.30 N, 36.26 E, 110 m, 08.V.2002 (M. Kütük). Host plants: in De Meyer et al. (2002) Distrubiton: Tropical and subtropical regions of Old and New Worlds. The species has been introduced to many countries. Its origin is apparently the Afrotropical region and Turkey (Giray, 1979; Norrbom et al. 1999; Özgür & Kütük, 2003). Measurements (length in mm): Male Body 4.0-5.0, wing 3.8-4.5. Female body: 4.5-6.0, wing 3.5-4.7.

Euleia heraclei (Linnaeus, 1758)

(Figs. 3, 8) Specimens examined: 2 ♂♂, 1 ♀, Adana, Karataş, Beyköy, 36.48 N, 35.21 E, 20 m, 03.V.2001; 2 ♂♂, 2 ♀♀, Adana, Balcalı, 37.02 N, 35.21 E, 100 m, 25.V.2001; 2 ♂♂, 2 ♀♀, Burdur, Yeşilova, Eşeler mountain, 37.30 N, 29.33 E, 1340 m, 23.VI.2000; 1 ♂, 2 ♀♀, Isparta, Yenişarbademli, 37.41 N, 31.21 E, 1230 m, 13.VII.2000; 3 ♂♂, 4 ♀♀, Isparta, Yenişarbademli, 37.41 N, 31.21 E, 1230 m, 12.VI.2001 (M. Kütük). Host plants: in Freidberg & Kugler, 1989; Merz, 1994. Distribution: Afghanistan, Algeria, Armenia, Azerbaijan, England, Estonia, Georgia, Israel, Japan, Kazakhstan, Kyrghyzstan, Latvia, Letonia, Middle East, Moldovia, Morocco, Russia, Switzerland, Tadjikistan, Turkey, Turkmenistan Ukraine, and Uzbekistan (Foote, 1984; White, 1988; Freidberg & Kugler, 1989; Merz, 1994; Norrbom et al. 1999). Measurements (length in mm): Male Body 4.1-5.5, wing 3.8-5.2. Female body: 5.5-6.0, wing 4.5-5.7.

KEY TO SPECIES OF RHAGOLETIS

1. Vein M reaches costa in the apical band of wing (Fig. 5); hyaline gap

between third and fourth bands reaches about vein R4+5; oligofagus species, host plants are Prunus and Lonicera …………………………………………………………..………….. cerasi (Linnaeus)

- Vein M reaches costa at the margin of apical band of wing (Fig. 4); hyaline gap between third and fourth bands reaches about vein R2+3; monofagus species, host plant only is Berberis vulgaris .......................…………………………….…………….......... berberidis Jermy

Rhagoletis berberidis Jermy, 1961 (Figs. 4, 9)

Specimens examined: 2 ♂♂, 1 ♀, Isparta, Yenişarbademli, 37.41 N, 31.21 E, 1230 m, 13.VII.2000; 2 ♀♀, Isparta, Aksu, Yakaköy, 37.43 N, 31.14 E, 1820 m, 14.VII.2000; 3 ♂♂, 3 ♀♀, Isparta, Aksu, Çayır plateau, 37.47 N, 31.14 E, 1925 m, 14.VII.2000; 2 ♂♂, Burdur, Soğanlı, 37.34 N, 30.15 E, 1515 m, 15.VII.2000; 3 ♂♂, 2 ♀♀ Isparta, Yenişarbademli, 37.41


N, 31.21 E, 1230 m, 12.VI.2001; 3 ♂♂, 3 ♀♀, Sivas Kangal, Sipahikoyağı, 38.13 N, 36.12 E, 1450 m, 04.VII.2005; 4 ♂♂, 6 ♀♀, Kayseri, Yahyalı, 38.01 N, 35.25 E, 1570 m, 06.VII.2005 (M. Kütük). Host plants: in Merz, 1994. Distribution: Armenia, Austria, Azerbaijan, Georgia, Hungary, Moldova, Russia, Switzerland and Ukraine (Foote, 1984; Merz, 1994; Norrbom et al. 1999).

This species is new record for Turkis fauna. Measurements (length in mm): Male Body 4.0-5.1, wing 3.2-4.0. Female body: 4.5-6.2, wing 3.5-4.5.

Rhagoletis cerasi Linnaeus, 1826 (Figs. 5,10)

Specimens examined: 3 ♂♂, 5 ♀♀, Adana, Pozantı, Alpu, 37.28 N, 34.54 E, 1120 m, 23.V.2001; 8 ♂♂, 3 ♀♀, Adana, Pozantı, 37.38 N, 34.59 E, 1220 m, 17.V.2002; 1 ♂, 1 ♀, Isparta, Senirkent, Gençali, 38.13 N, 31.03 E, 925 m, 16.VI.1999; 3 ♂♂, Isparta, Senirkent, Aşağıkaşıkara, 38.15 N, 30.48 E, 940 m, 22.VI.2000; 5 ♂♂, Isparta, Yenişarbademli, 37.41 N, 3121 E, 1230 m, 12.VI.2001 (M. Kütük). Host plants: in Giray, 1979; White, 1988; Merz, 1994. Distrubition: Central and North Europe, Russia, Estonia, Latvia, Ukraine, Letonia, Azerbaijan, Georgia, Armenia, Kazakhstan, Uzbekistan, Tadjikistan, Kyrghyzstan, Turkmenistan, England, Switzerland, Turkey and West Siberia (Giray, 1979; Foote, 1984; White, 1988; Merz, 1994; Norrbom et al. 1999). Measurements (length in mm): Male Body 4.3-5.5, wing 3.3-4.5. Female body: 5.3-6.2, wing 3.5-5.0.

DISCUSSION

Rhagoletis berberidis is a new record for the fauna of Turkey. This study contributed to Turkish Tephritidae Fauna and identification of genera and their species of Trypetinae in Turkey. The species of Trypetinae is not widespread in Turkey.

ACKNOWLEDGEMENTS We thank Prof. Şemsettin Civelek (University of Fırat, Department of Biology, Elazığ, Turkey) for determinining the host plants of Trypetinae. We are grateful to the İnönü Üniversity project of Research Foundation (Project number: 2004-3) for the support provided for this project.

LITERATURES CITED

De Meyer, M., Copeland, R. S., Lux, S. A., Mansel, M., Quilici, S., Wharton, R., White, I. M. & Zenz, N. J. 2002. Annotated check list of host plants for Afrotropical fruit flies (Diptera: Tephritidae) of the genus Ceratitis. Koninklijk Museumvoor Midden-Afrika, Tervuren, Belgie. Zoologische Dokumetatie Vol. 27, 91 pp.


Foote, R. H. 1984. Family Tephritidae, In: A. Soos and L. Papp eds., Catalogue of Palaearctic Diptera (ed., R. C. Foote). Budapest and Elsevier Science Publishers, Amsterdam, Vol. 9, pp. 66-149. Freidberg, A. & Kugler, J. 1989. Fauna Palaestina. Insecta IV. Diptera: Tephritidae. Israel Academy of Sciences and Humanities, Jerusalem, 212 pp. Giray, H. 1979. Türkiye Trypetidae (Diptera) Faunasına Ait İlk Liste. Türkiye Bitki Koruma Dergisi 3 : 35 – 46. Merz, B. 1994. Diptera, Tephritidae. Insecta Helvetica Fauna, Hge press, Vol: 10, Geneve, 198 pp. Özgür, A. F. & Kütük, M. 2003. Adana İli Meyve Sinekleri (Tephritidae: Diptera) Faunasının Tespiti. Ç.Ü.Z.F. Dergisi 18 (2): 35-44. White, I. M. 1988. Tephritid Flies (Diptera: Tephritidae). Handbooks for the Identification of British Insects 10 (5a), 134 pp. White, I. M., Headrick, D. H., Norrbom A. L. & Carroll, L. E. 2000. 33 Glossary. In: Fruit Flies (Tephritidae): Phylogeny and Evolution of Behavior (eds., Aluja, M. and Norrbom, A. L.). CRC, Boca Raton, Washington, pp. 881-924.

[Received October 2005. Accepted January 2006]

Figures 1-5. The wings pattern of Trypetinae: 1- Carpomya pardalina, 1- Ceratitis capitata, 3- Euleia heraclei, 4- Rhagoletis berberidis, 5- Rhagoletis cerasi.

1

3

2

4

5


Figures 6-10. The aculeus of Trypetinae: 6- Carpomya pardalina, 7- Ceratitis capitata, 8- Euleia heraclei, 9- Rhagoletis berberidis, 10- Rhagoletis cerasi.

10

6

7

8

9


NOMENCLATURAL CHANGES FOR FOURTEEN TRILOBITES GENERA

Hüseyin Özdikmen*

* Gazi Üniversitesi, Fen-Edebiyat Fakültesi, Biyoloji Bölümü, 06500 Ankara / TÜRKİYE, e-mail: [email protected] [Özdikmen, H. 2006. Nomenclatural changes for fourteen Trilobites genera. Munis Entomology & Zoology, 1 (2): 179-190]

ABSTRACT: Fourteen junior homonyms were detected amongst the Trilobites genera and the following replacement names are proposed: Limbadiscus Korobov, 1980 for Natalina Romanenko, 1978; Kazakhius nom. nov. for Elegantaspis Ivshin, 1962; Neoiranella nom. nov. for Iranella Hupé, 1953; Suluderella nom. nov. for Mareda Kobayashi, 1942; Aldanianus nom. nov. for Comptocephalus Repina, 1964; Neodrepanura nom. nov. for Drepanura Bergeron, 1899; Kiyakius nom. nov. for Pionaspis Zhang, 1983; Galbertianus nom. nov. for Hollardia Alberti, 1964; Neograciella nom. nov. for Graciella Rozova, 1963; Samgonus nom. nov. for Lampropeltis Öpik, 1967; Atilayus nom. nov. for Deltocephalus Ogienko, 1969; Wolfartius nom. nov. for Farsia Wolfart, 1974; Neoblairella nom. nov. for Blairella Rasetti, 1965 and Neoregina nom. nov. for Regina Egorova, 1967. Accordingly, new combinations are herein proposed for the species currently included in these genera respectively: Limbadiscus incitus (Romanenko, 1978) comb. nov.; Kazakhius elegantulus (Ivshin, 1962) comb. nov.; Neoiranella latefrons (King, 1937) comb. nov.; Suluderella mukazegata (Kobayashi, 1942) comb. nov.; Aldanianus mitis (Repina, 1964) comb. nov.; Neodrepanura premesnili (Bergeron, 1899) comb. nov.; Kiyakius ichthyura (Zhang, 1983) comb. nov.; Galbertianus hollardi (Alberti, 1964) comb. nov.; Neograciella graciensis (Rozova, 1963) comb. nov.; Samgonus nitens (Öpik, 1967) comb. nov.; Atilayus orientalis (Ogienko, 1969) comb. nov.; Wolfartius abundans (Wolfart, 1974) comb. nov.; Neoblairella crassimarginata (Rasetti, 1965) comb. nov. and Neoregina opipara (Egorova, 1967) comb. nov. KEY WORDS: nomenclatural changes, homonymy, replacement names, Trilobites.

The purpose of the present paper is to bring the taxonomy of trilobites into accordance with the requirements of the International Code of Zoological Nomenclature (1999). It considers homonymous genus group names of trilobites introduced from 1758 to 2005.

In an effort to reduce the number of homonyms in Trilobites, I systematically checked all generic names published. I found fourteen trilobites genera and one species whose names had been previously published for other taxa, making them junior homonyms. In accordance with Article 60 of the International Code of Zoological Nomenclature, I propose substitute names for these genus and species group names.


TAXONOMY

Order AGNOSTIDA Family HEBEDISCIDAE

Genus LIMBADISCUS Korobov, 1980 Natalina Romanenko, in Repina & Romanenko, 1978. Trudy Inst. Geol. Geofiz. sib. Otd. 382: 128. (Trilobita: Agnostida: Eodiscina: Eodiscoidea: Hebediscidae). Preoccupied by Natalina Pilsbry, in Tryon & Pilsbry, 1893. Man. Conch., (2) 8, 135. (Mollusca: Gastropoda: Pulmonata: Stylommatophora: Acavoidea: Rhytididae).

Remarks: The name Natalina was initially introduced by Pilsbry, 1893 as a replacement name for the preoccupied genus name Aerope Martens, 1860 of the gastropod family Rhytididae (with the type species Helix cafra Férussac, 1821). Subsequently, Romanenko, 1978 described a trilobite genus of the family Hebediscidae (with the type species Natalina incita Romanenko, 1978 from Sanashtykgol Horizon, Gorny Altay, S. Siberia, Russia) under the same generic name. Thus, the genus Natalina Romanenko, 1978 is a junior homonym of the genus Natalina Pilsbry, 1893. I propose for the genus Natalina Romanenko, 1978 the junior subjective synonym Limbadiscus Korobov, 1980 that is the type species Limbadiscus dilatum Korobov, 1980 from Egyngolskaya Fm, NW Mongolia (Jell, 1997). Summary of nomenclatural changes: Limbadiscus Korobov, 1980 = Natalina Romanenko, 1978 (non Pilsbry, 1893). Limbadiscus incitus (Romanenko, 1978) comb. nov. = Natalina incita Romanenko, 1978 = Limbadiscus dilatum Korobov, 1980.

Order ASAPHIDA

Family APHELASPIDIDAE Genus KAZAKHIUS nom. nov.

Elegantaspis Ivshin, 1962. Upper Cambrian trilobites of Kazakhstan; Part 2. Akad. Nauk Kazakh. SSR, Inst. Geol. Nauk, Alma-Ata: 80. (Trilobita: Asaphida: Asaphina: ?Anomocaroidea: Aphelaspididae). Preoccupied by Elegantaspis Heintz, 1929. Skrift. Svalbard Ishavet, Oslo, No. 22, 61. (Pisces: Placodermi: Phlyctaenioidei: Phlyctaeniina: Phlyctaeniidae).

Remarks: Ivshin (1962) proposed the genus name Elegantaspis with the type species Elegantaspis elegantula Ivshin, 1962 from Seletinsky Horizon, Kazakhstan. Unfortunately, the generic name was already preoccupied by Heintz (1929), who had described the genus Elegantaspis in the fossil fish family Phlyctaeniidae. Thus, the genus Elegantaspis Ivshin, 1962 is a junior homonym of the generic name Elegantaspis Heintz, 1929. I propose a new replacement name Kazakhius nom. nov. for Elegantaspis Ivshin, 1962. The name is dedicated to the country Kazakhstan. The name is masculine in gender.


Summary of nomenclatural changes: Kazakhius nom. nov. = Elegantaspis Ivshin, 1962 (non Heintz, 1929). Kazakhius elegantulus (Ivshin, 1962) comb. nov. = Elegantaspis elegantula Ivshin, 1962.

Family DIKELOCEPHALIDAE

Genus NEOIRANELLA nom. nov. Iranella Hupé, 1953. in Piveteau, Traité de Paléont., 3, 189 ; 1955, Ann. Paléont., 41, 97 (117). (Trilobita: Asaphida: Asaphina: Dikelocephaloidea: Dikelocephalidae). Preoccupied by Iranella Uvarov, 1922. J. Bombay nat. Hist. Soc., 28, 729. (Insecta: Orthoptera, Acrididae, Catantopinae).

Remarks: Hupé (1953) proposed the genus name Iranella with the type species Saratogia latefrons King, 1937 from Sayad Hassan, Iran. Unfortunately, the generic name was already preoccupied by Uvarov (1922), who had described the genus Iranella in the orthopteran family Acrididae. Thus, the genus Iranella Hupé, 1953 is a junior homonym of the generic name Iranella Uvarov, 1922. I propose a new replacement name Neoiranella nom. nov. for Iranella Hupé, 1953. The name dedicated to the current genus name.

I have reason in principle to rename the homonym Iranella, but should also know that such a new name may be unnecessary because Saratogia latefrons King, 1937 has already been referred to Maladioidella abdita (Salter, 1866) by Rushton & Hughes (1996); this was, of course, a subjective decision. “Rushton & Hughes (1996) discussed Maladioidella and its synonyms Cedarellus and Iranella, and redescribed the widespread species M. abdita (Salter, 1866)” (Rushton et al., 2002). But Jell & Adrain (2003) given the trilobite genera Kuruktagella T. Zhang, 1981 and Cedarellus Lazarenko, 1966 as junior subjective synonyms of Maladioidella Endo, 1937 in the family Idahoiidae on pages 355, 394 and 472. Also the genus Iranella is given by Jell & Adrain (2003) as a junior subjective synonyms of Maladioidella Endo, 1937 on page 388. But on page 469 the genus name Iranella recorded as a available generic name without synonym in the family Dikelocephalidae. Summary of nomenclatural changes: Neoiranella nom. nov. = Iranella Hupé, 1953 (non Uvarov, 1922). Neoiranella latefrons (King, 1937) comb. nov. = Saratogia latefrons King, 1937.

Family SAUKIIDAE

Genus SULUDERELLA nom. nov. Mareda Kobayashi, 1942. Jap. J. geol. Geogr., 18, 297. (Trilobita: Asaphida: Asaphina: Dikelocephaloidea: Saukiidae). Preoccupied by Mareda Walker, 1855. List Specimens Lep. Ins. Brit. Mus., 5, 1157. (Lepidoptera: Zygaenoidea: Limacodidae: subfamily unassigned).

Remarks: The generic name Mareda Walker, 1855 was proposed for a genus of moths family Limacodidae (with the type species Mareda


ferruginea Walker, 1855 by monotypy). Subsequently, the generic name Mareda Kobayashi, 1942 was introduced for a new trilobite genus (with the type species Mareda mukazegata Kobayashi, 1942 from Fengshan Fm, Shanxi, China) of the family Saukiidae. Thus, the genus Mareda Kobayashi, 1942 is a junior homonym of the generic name Mareda Walker, 1855. I propose for the genus Mareda Kobayashi, 1942 the new replacement name Suluderella nom. nov. The name is given in honour to Prof. Dr. Zekiye Suludere. The name is feminine in gender. Summary of nomenclatural changes: Suluderella nom. nov. = Mareda Kobayashi, 1942 (non Walker, 1855). Suluderella mukazegata (Kobayashi, 1942) comb. nov. = Mareda mukazegata Kobayashi, 1942.

Order CORYNEXOCHIDA Family DINESIDAE

Genus ALDANIANUS nom. nov. Compsocephalus Repina, 1964 in Repina et al. [Lower Cambrian biostratigraphy of the Sayan-Altay folded region]. (Akademiya Nauk SSSR, Sibirskoe Otdelenie, Institut Geologii i Geofiziki, Izdatelstvo: Moscow). 308 p. (Trilobita: Corynexochida: Corynexochina: Corynexochoidea: Dinesidae). Preoccupied by Compsocephalus White, 1845. Ann. Mag. nat. Hist., 15, 39. (Insecta: Coleoptera: Scarabaeoidea: Scarabaeidae: Cetoniinae).

Remarks: The genus Compsocephalus was erected by White, 1845 with the type species Compsocephalus horsfieldianus White, 1845. Later, the genus Compsocephalus was described by Repina, 1964 in Repina et al., 1964 with the type species Compsocephalus mitis Repina, 1964 from Aldanian, Sayan Altay Fold Belt, Russia. However, the name Compsocephalus Repina, 1964 is invalid under the law of homonymy, being a junior homonym of Compsocephalus White, 1845.

In Repina’s paper Compsocephalus appears on page 148 without a species name being assigned. On page 307 Lepidocephaloides is introduced with the type species L. mitis, and Lepidocephaloides mitis is the name given in the explanation of plate 33, figures 12-14. Compsocephalus is re-introduced without description on page 308, in a continuation of the account of L. mitis. The fact that Repina made a hand-written change on a copy of her work is irrelevant to the present nomenclatural problem. Also, Jell & Adrain (2003) mentioned that Lepidocephaloides Repina, 1964 is typographical error for Compsocephalus on p. 307 - hand corrected by author in copy held by A. R. Palmer – borne out by assignment of type species on p. 308.

Under the circumtances, Lepidocephaloides was not made available by Repina within the term of the ICZN. Namely, Lepidocephaloides is not a synonym of Compsocephalus. So I propose to substitute the junior homonym name Compsocephalus Repina, 1964 for the name Aldanianus nom. nov.

http://www.trilobites.info/ordcorynexochida.htm


Summary of nomenclatural changes: Aldanianus nom.nov. = Compsocephalus Repina, 1964 (non White, 1845). Aldanianus mitis (Repina, 1964) comb. nov. = Compsocephalus mitis Repina, 1964.

Order LICHIDA

Family DAMESELLIDAE Genus NEODREPANURA nom. nov.

Drepanura Bergeron, 1889. Bull. Soc. géol. France, 27 (3): 509. (Trilobita: Lichida: Lichina: Dameselloidea: Damesellidae). Preoccupied by Drepanura Schoett, 1891. Bih. svenska VetensAkad., 17 (4), no. 8, 19. (Collembola: Entomobryidae).

Remarks: The name Drepanura was initially introduced by Schoett, 1891 for a genus of the collembolan family Entomobryidae (with the type species Drepanura californica Schoett, 1891). Subsequently, Bergeron, 1899 described a trilobite genus of the family Damesellidae (with the type species Drepanura premesnili Bergeron, 1899 from Kushan Fm, Shandong, China) under the same generic name. Thus, the genus Drepanura Bergeron, 1899 is a junior homonym of the genus Drepanura Schoett, 1891. I propose for the genus Drepanura Bergeron, 1899 the new replacement name Neodrepanura nom. nov. The name is dedicated to the current generic name. Summary of nomenclatural changes: Neodrepanura nom. nov. = Drepanura Bergeron, 1899 (non Schoett, 1891). Neodrepanura premesnili (Bergeron, 1899) comb. nov. = Drepanura premesnili Bergeron, 1899.

Family DAMESELLIDAE

Genus KIYAKIUS nom. nov. Pionaspis Zhang, in Qiu et al., 1983. [Paleontological atlas of east China. 1. Volume of Early Paleozoic.]. Geological Publishing House, Peking: 176. (Trilobita: Lichida: Lichina: Dameselloidea: Damesellidae). Preoccupied by Pionaspis Denison, 1964. Fieldiana, Geol. 13: 386. (Chordata: Agnatha: †Cynthaspidiformes: Cynthaspida: †Cyathaspididae).

Remarks: Zhang (1983) proposed the genus Pionaspis with the type species Pionaspis ichthyura Zhang, 1983 from Kushan Fm, Zhejiang, China. Unfortunately, the generic name was already preoccupied by Denison (1964), who had described the genus Pionaspis in †Cynthaspidiformes. Thus, the genus Pionaspis Zhang, 1983 is a junior homonym of the generic name Pionaspis Denison, 1964. I propose a new replacement name Kiyakius nom. nov. for Pionaspis Zhang, 1983. The name is given in honour of my colleague Prof. Dr. Suat Kıyak. The name is masculine in gender. Summary of nomenclatural changes: Kiyakius nom. nov. = Pionaspis Zhang, 1983 (non Denison, 1964). Kiyakius ichthyura (Zhang, 1983) comb. nov. = Pionaspis ichthyura Zhang, 1983.


Order PROETIDA Family TROPIDOCORYPHIDAE

Genus GALBERTIANUS nom. nov. Hollardia Alberti, 1964. Senckenberg. leth. 45: 123. (Trilobita: Proetida: Proetina: Proetoidea: Tropidocoryphidae). Preoccupied by Hollardia Poey, 1861. Mem. Cuba, 2, 348. (Pisces: Actinopterygii: Tetraodontiformes: Triacanthodidae).

Remarks: The generic name Hollardia Poey, 1861 was proposed for a genus of fish family Triacanthodidae (with the type species Hollardia hollardi Poey, 1861). Subsequently, the generic name Hollardia Alberti, 1964 was introduced for a new trilobite genus (with the type species Hollardia hollardi Alberti, 1964 from Ain Tagh Sh, An Targa, W Morocco) of the family Tropidocoryphidae. Thus, the genus name Hollardia Alberti, 1964 is a junior homonym of the generic name Hollardia Poey, 1861 and also the species name Hollardia hollardi Alberti, 1964 that is the type species of Hollardia Alberti, 1964 is invalid under the law of homonymy, being a primary junior homonym of Hollardia hollardi Poey, 1861 that is the type species of Hollardia Poey, 1861. Under the Zoological Code (ICZN) they must be rejected and replaced. I propose to substitute the junior homonym genus group name Hollardia Alberti, 1964 for the nomen novum Galbertianus and the junior primary homonym species group name Hollardia hollardi Alberti, 1964 for the comb. nov. Galbertianus hollardi. The genus group name Galbertianus is given in honour to G. K. B. Alberti who is the author of the genus name. The name is masculine in gender. Summary of nomenclatural changes: Galbertianus nom. nov. = Hollardia Alberti, 1964 (non Poey, 1861). Galbertianus hollardi (Alberti, 1964) comb. nov. = Hollardia hollardi Alberti, 1964 (non Poey, 1861).

Order PTYCHOPARIIDA

Family LONCHOCEPHALIDAE Genus NEOGRACIELLA nom. nov.

Graciella Rozova, 1963. Geologiya Geofiz. Novosibirsk 1963 (9): 14. (Trilobita: Ptychopariida: Ptychopariina: Ptychoparioidea: Lonchocephalidae). Preoccupied by Graciella Jordan, 1894. Novit. zool., 1, 215. (Insecta: Coleoptera: Chrysomeloidea: Cerambycidae: Lamiinae: Tragocephalini).

Remarks: Rozova (1963) proposed the genus name Graciella with the type species Graciella graciensis Rozova, 1963 from Yurakisky Horizon, NW Siberia, Russia. Unfortunately, the generic name was already preoccupied by Jordan (1894), who had described the genus Graciella in the coleopteran family Cerambycidae (Neave, 1939). Thus, the genus Graciella Rozova, 1963 is a junior homonym of the generic name Graciella Jordan, 1894. I propose a new replacement name Neograciella

http://www.trilobites.info/ordptychopariida.htm

http://www.funet.fi/pub/sci/bio/life/pisces/actinopterygii/index.html

http://www.funet.fi/pub/sci/bio/life/pisces/actinopterygii/tetraodontiformes/index.html

http://www.funet.fi/pub/sci/bio/life/pisces/actinopterygii/tetraodontiformes/triacanthodidae/index.html

http://www.funet.fi/pub/sci/bio/life/pisces/actinopterygii/tetraodontiformes/triacanthodidae/index.html



nom. nov. for Graciella Rozova, 1963. The name dedicated to the current generic name. Summary of nomenclatural changes: Neograciella nom. nov. = Graciella Rozova, 1963 (non Jordan, 1894). Neograciella graciensis (Rozova, 1963) comb. nov. = Graciella graciensis Rozova, 1963.

Family PLETHOPELTIDAE

Genus SAMGONUS nom. nov. Lampropeltis Öpik, 1967. Bull.Bur.Miner.Resour.Geol.Geophys.Aust. 74 (1):182. (Trilobita: Ptychopariida: Uncertain suborder: Uncertain superfamily: Plethopeltidae). Preoccupied by Lampropeltis Fitzinger, 1843. Syst. Rept., 25. (Reptilia: Squamata: Serpentes: Colubridae: Colubrinae: Lampropeltini).

Remarks: Firstly, the genus Lampropeltis was established by Fitzinger, 1843 for snakes family Colubridae with the type species Coluber getulus Linnaeus, 1766. Later, the genus Lampropeltis was proposed by Öpik, 1967 for trilobites family Plethopeltidae with the type species Lampropeltis nitens Öpik, 1967 from Steamboat Sst, Queensland, Australia. However, the name Lampropeltis Öpik, 1967 is invalid under the law of homonymy, being a junior homonym of Lampropeltis Fitzinger, 1843. I propose to substitute the junior homonym name Lampropeltis Öpik, 1967 for the nomen novum Samgonus. The name is dedicated to trilobitologist Dr. Samuel M. Gon III. The name is masculine in gender. Summary of nomenclatural changes: Samgonus nom. nov. = Lampropeltis Öpik, 1967 (non Fitzinger, 1843). Samgonus nitens (Öpik, 1967) comb. nov. = Lampropeltis nitens Öpik, 1967.

Family PROASAPHISCIDAE Genus ATILAYUS nom. nov.

Deltocephalus Ogienko, 1969. Geologiya Geofiz.Novosibirsk 1969 (8): 56 (Trilobita: Ptychopariida: Ptychopariina: Ptychoparioidea: Proasaphiscidae). Preoccupied by Deltocephalus Burmeister, 1838. Gen. Ins., 2 (Jassus), [5] (Homoptera: Cicadomorpha: Membracoidea: Cicadellidae: Deltocephalinae: Deltocephalini).

Remarks: The generic name Deltocephalus Burmeister, 1838 was proposed for a genus of homopterous family Cicadellidae (with the type species Cicada pulicaris Fallèn, 1806 by subsequent designation, by Kirschbaum, 1858). The genus Deltocephalus Burmeister, 1838 first introduced as a subgenus of Jassus Fabricius, 1803, now Iassus Fabricius, 1803. Subsequently, the generic name Deltocephalus Ogienko, 1969 was introduced for a new trilobite genus (with the type species Deltocephalus orientalis Ogienko, 1969 from chersk Fm, S Siberia, Russia) of the family Proasaphiscidae. Thus, the genus Deltocephalus Ogienko, 1969 is a junior homonym of the generic name Deltocephalus


Burmeister, 1838. I propose for the genus Deltocephalus Ogienko, 1969 the new replacement name Atilayus nom. nov. The name is given in honour to my colleague Atılay Yağmur Okutaner. The name is masculine in gender. Summary of nomenclatural changes: Atilayus nom. nov. = Deltocephalus Ogienko, 1969 (non Burmeister, 1838). Atilayus orientalis (Ogienko, 1969) comb. nov. = Deltocephalus orientalis Ogienko, 1969.

Family PROASAPHISCIDAE

Genus WOLFARTIUS nom. nov. Farsia Wolfart, 1974. Geol. Jb. (B) 8: 116. (Trilobita: Ptychopariida: Ptychopariina: Ptychoparioidea: Proasaphiscidae). Preoccupied by Farsia Amsel, 1961. Ark. Zool. (N.S.) 13: 375. (Insecta: Lepidoptera: Pyraloidea: Pyralidae: Phycitinae).

Remarks: The genus Farsia was erected by Amsel, 1961 with the type species Farsia pallorella Amsel, 1961. Later, the genus Farsia was described by Wolfart, 1974 with the type species Farcia abundans Wolfart, 1974 from Kushanian, SE Iran. However, the name Farsia Wolfart, 1974 is invalid under the law of homonymy, being a junior homonym of Farsia Amsel, 1961. I propose to substitute the junior homonym name Farsia Wolfart, 1974 for the nomen novum Wolfartius. The name is given in honour to R. Wolfart who is the current author of the generic name. The name is masculine in gender. Summary of nomenclatural changes: Wolfartius nom. nov. = Farsia Wolfart, 1974 (non Amsel, 1961). Wolfartius abundans (Wolfart, 1974) comb. nov. = Farsia abundans Wolfart, 1974.

Family PTYCHOPARIIDAE

Genus NEOBLAIRELLA nom. nov. Blairella Rasetti, 1965. J. Paleont. 39: 1012. (Trilobita: Ptychopariida: Ptychopariina: Ptychoparioidea: Ptychopariidae). Preoccupied by Blairella Miller & Gurley, 1896. Bull. Illinois Mus. nat. Hist., 11: 6. (Mollusca: Bivalvia: Heterodonta: Veneroida: Carditacea: family uncertain).

Remarks: Rasetti (1965) proposed the genus Blairella with the type species Blairella crassimarginata Rasetti, 1965 from Pleasant Hills Fm, Pennsylvania, USA. Unfortunately, the generic name was already preoccupied by Miller & Gurley (1896), who had described the genus Blairella in Bivalvia. Thus, the genus Blairella Rasetti, 1965 is a junior homonym of the generic name Blairella Miller & Gurley, 1896. I propose a new replacement name Neoblairella nom. nov. for Blairella Rasetti, 1965. The name is dedicated to the current generic name. Summary of nomenclatural changes: Neoblairella nom. nov. = Blairella Rasetti, 1965 (non Miller & Gurley, 1896). Neoblairella crassimarginata (Rasetti, 1965) comb. nov. = Blairella crassimarginata Rasetti, 1965.


Family PTYCHOPARIIDAE Genus NEOREGINA nom. nov.

Regina Egorova, 1967. Paleont.Zh. 1967 (1): 77. (Trilobita: Ptychopariida: Ptychopariina: Ptychoparioidea: Ptychopariidae). Preoccupied by Regina Baird & Girard, 1853. Smithson. misc. Coll., 2 (5), 45. (Reptilia: Squamata: Serpentes: Colubridae).

Remarks: Firstly, the genus Regina was described by Baird & Girard, 1853 for snakes family Colubridae with the type species Regina grahamii Baird & Girard, 1853. Later, the genus Regina was proposed by Egorova, 1967 for trilobites family Ptychopariidae with the type species Regina opipara Egorova, 1967 from Khara-tass Fm, N Siberia, Russia. However, the name Regina Egorova, 1967 is invalid under the law of homonymy, being a junior homonym of Regina Baird & Girard, 1853. In accordance with article 60 of the International Code of Zoological Nomenclature, I propose to substitute the junior homonym name Regina Egorova, 1967 for the nomen novum Neoregina. The name is dedicated to the current genus name. Summary of nomenclatural changes: Neoregina nom. nov. = Regina Egorova, 1967 (non Baird & Girard, 1853). Neoregina opipara (Egorova, 1967) comb. nov. = Regina opipara Egorova, 1967.

LITERATURE CITED

Alberti, G. K. B. 1964. Neue Trilobiten as dem marokkanischen und deutschen Unter-und Mitteldevon. Senckenbergiana lethaea 45: 115-133. Amsel, H. G. 1961. Die Microlepidopteren der Brandt'schen Iran-Ausbeute 5. Teil. Arkiv för Zoologi 13 (17): 323-445. Baird, S. F. & Girard, C. 1853. Catalogue of North American reptiles in the museum of the Smithsonian Institution. Part I. Serpentes. 172 pp. Bergeron, J. N. 1889. Etude de quelques trilobites de Chine. Bulletin de la Societé Géologique de France, 3rd series 27: 499-519. Burmeister, H. C. C. 1838. Rhynchota. No. 2. Genera Insectorum 1, pls 6, 12, 14, 15. Denison, R. H. 1964. The Cyathaspididae: A family of Silurian and Devonian jawless vertebrates. Fieldiana: Geology 13: 307-473. Egorova, L. I. 1967. [Some trilobites from the Lower and Middle Cambrian of the Siberian Platform]. Paleontologicheskij Zhurnal, 1967 (1): 68-78. Fitzinger, L. J. F. J. 1843. Systema reptilium. Fasciculus primus, Amblyglossae. indobonae, Braumuller et Seidel, Vienna, 106 pp. Heintz, A. 1929. Die downtonischen and devonischen Vertebraten von Spitsbergen. 2. Acanthaspida. Skr. Svalb. Ish. 22: 1-81. Hupé, P. 1953. Classification des Trilobites. Annales de Paléontologie, 39: 61-168.


International Comission of Zoological Nomenclature. 1999. International Code of Zoological Nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature, London. Ivshin, N. K. 1962. [Upper Cambrian trilobites of Kazakhstan. Part 2]. (Akademii Nauk Kazakhskoe SSR: Alma-Ata) 412 pp. Jell, P. A. 1997. Introduction to the Suborder Eodiscina. pp. 384-405. In Kaesler, R.L. (ed.) Treatise on invertebrate paleontology, Part O, Arthropoda 1. Trilobita, Revised. Volume 1: Introduction, Order Agnostida, Order Redlichiida. (Geological Society of America and University of Kansas: Boulder, Colorado and Lawrence, Kansas). 530p. Jell, P. A. & Adrain, J. M. 2003. Available generic names for trilobites. Memoirs of the Queensland Museum, 48 (2): 331-553. Jordan, K. 1894. On some new genera and species of Coleoptera in the Tring Museum. Novitates Zoologicae 1: 488-503. Kirschbaum, C. L. 1858. Der Gegend von Wiesbaden gesammelten Arten der Gattung Jasus. Jahrbücher des Nassauischer Verein für Naturkunde 13: 355-358. Kobayashi, T. 1942. The Rakuroan Complex of the Shansi Basin and its surroundings. Japanese Journal of Geology and Geography 18 (4): 283-306. Korobov, M. N. 1980. [Biostratigraphy and miomerid trilobites from the Lower Cambrian of Mongolia.]. Joint Soviet-Mongolian Scientific Research Geological Expedition, Transactions 26: 5-108. Miller, S. A. & Gurley, W. F. E. 1896. New species of Paleozoic invertebrates from Illinois and other states. Illinois State Museum of Natural History, Bulletin 11: 1–50. Neave, S. A. 1939. Nomenclator Zoologicus. The Zoological Society of London, 2: 509. Ogienko, L. V. 1969. [On the problem of Middle Cambrian in the basin of the Up-stream of the River Lena]. Geologiya i Geofizika 1969 (8): 55-62. Öpik, A. A. 1967. The Mindyallan fauna of north-western Queensland. Bureau of Mineral Resources Geology & Geophysics Australia Bulletin 74: 1-404 / 1-167 (2 vols.). Pilsbry, H. A. 1893. in Tryon, G. W. & Pilsbry, H. A. Manual of Conchology. Philadelphia: Conchological Section, Academy of Natural Sciences Ser. 2 (8):1-314 pp. Poey, F. 1861. Memorias sobra la historia natural de la Isla de Cuba, acompañadas de sumarios Latinos y extractos en Francés. Tomo 2. La Habana. Memorias sobra la historia natural de la Isla de Cuba, 2 (1861): ,337-442. Qiu, H., Lu, Y., Zhu, Z., Bi, D., Lin, T., Zhou, Z., Zhang, Q., Qian, Y., Ju, T., Han, N. & Wei, X. 1983. [Trilobita]. In Paleontological Atlas of East China. Part 1: Early Paleozoic. (Nanjing Institute of Geology and Mineral Resources. Geological Publishing House: Beijing). 657 pp. Rasetti, F. 1965. Middle Cambrian trilobites of the Pleasant Hill Formation in central Pennsylvania. Journal of Paleontology 39: 1007-1014. Repina, L. N., Khomentovsky, V. V., Zhuravleva, I. T. & Rozanov, A. Yu. 1964. [Lower Cambrian biostratigraphy of the Sayan-Altay folded region]. (Akademiya Nauk SSSR, Sibirskoe Otdelenie, Institut Geologii i Geofiziki, Izdatelstvo: Moscow). 313 pp.


Repina, L. N. & Romanenko, E. 1978. [Trilobites and stratigraphy of the Lower Cambrian of Altay]. Akademiya Nauk SSSR, Sibirskoe Otdelenie, Instituta Geologii i Geofiziki, Trudy 382: 1-304. Rozova, A. V. 1963. [A biostratigraphic scheme for the Upper and upper Middle Cambrian and new Upper Cambrian trilobites]. Geologiya i Geofizika 1963 (9): 3-19. Rushton, A. W. A. & Hughes, N. C. 1996. Biometry, systematics and biogeography of the late cambrian trilobite Maladioidella abdita. Transactions of the royal Society of edinburgh: earth sciences 86: 247–56. Rushton, A. W. A., Cocks, L. R. M. & Fortey, R. A. 2002. Upper Cambrian trilobites and brachiopods from Severnaya Zemlya, Arctic Russia, and their implications for correlation and biogeography. Geol. Mag. 139 (3): 281-290. Schoett, H. 1891. Beiträge zur Kenntniss kalifornischer Collembola. Bihang Till K. Svenska Vet.-Akad. Handlingar, Band 17, Afd. IV, N:o 8, Stockholm, p.1-25. Uvarov, B. P. 1922. Records and descriptions of Orthoptera from S. W. Asia. Journal of Bombay Natural History Society 28: 719-738. Walker, F. 1855. List of specimens Lepidopterous Insects in the collection of the British Museum, London, 5: 1157. White, A. 1845. Description of two apparently new species of lamellicorn beetles. Annals and Magazine of Natural History 15: 38-41. Wolfart, R. 1974. Die Fauna (Brachiopoda, Mollusca, Trilobita) des älteren Ober-Kambriums (Ober-Kushanian) von Dorah Shah Dad, Südost-Iran, und Surkh Bum, Zentral-Afghanistan. Geologisches Jahrbuch, Reihe B, 8: 71-184.

[Received October 2005. Accepted February 2006]


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SEASONAL OCCURRENCE AND BIOLOGY OF GLOBE THISTLE CAPITULUM WEEVIL LARINUS ONOPORDI

(F.) (COLEOPTERA: CURCULIONIDAE) IN NORTHEASTERN TURKEY

Levent Gültekin*

* Atatürk University, Faculty of Agriculture, Department of Plant Protection, 25240 Erzurum, TURKEY. e-mail: [email protected] [Gültekin, L. 2006. Seasonal occurrence and biology of globe thistle capitulum weevil Larinus onopordi (F.) (Coleoptera: Curculionidae) in Northeastern Turkey. Munis Entomology & Zoology 1 (2): 191-198] ABSTRACT: The biology of weevil species, Larinus onopordi (F.) living on globe thistle Echinops sphaerocephalus L. was described for the first time. The species is univoltine, adults overwinter under hiding object as aggregated. Eggs are inserted into the flowerhead; L. onopordi deposited eggs superficially, 1-23 (3.76) eggs each capitulum may be occur in one flowerhead, larvae develop in capitulum eating and destroying the seeds. In late summer, larvae complete their growth; prepare pupal case 1-4 (0.84) case in each flowerhead and pupation occurs in here. New generation adults emerge from flowerhead after waiting several days. Thus, this weevil has one generation per year in northeatern Turkey. A braconid wasp, Bracon facialis Thom. (Hymenoptera: Braconidae) reared from the pupal stage of L. onopordi. KEY WORDS: Larinus onopordi, Echinops sphaerocephalus, biology, Curculionidae, Lixinae, northeastern Turkey.

Echinops L. are perennial or biennial spiny herbs with erect ridged

stems. Leaves simple to three-pinnatisect, petiolate or sessile. Capitula single-flowered, congested into globose heads subtended by small reflexed bracts. The genus Echinops has sixteen species in all Turkey. Of these, Echinops sphaerocephalus L. is distributed mainly in northeastern Anatolia (Davis, 1975). This plant is on the weed list in Turkey in common pastures, meadows, environs of airport, highways and railways (Uluğ et al., 1993), maize, cotton fields, field margins and other crops (citrus, okra, grape, soybean, wheat, tomato, sesame, groundnut and bean) in Çukurova Region (Okşar & Uygur, 2000). Also, E. sphaerocephalus is on the invasive weed list of North America (Hartman & Nelson, 2000) and it is commonly found in Nebraska (McCarty et al., 1984).

Even though informed about the weed characteristics of E. sphaerocephalus in current literature, this plant also has medicinal uses (Kucherov, 1978). E. sphaerocephalus contains the non-poisonous alkaloid, echinopsin, which gives sufficiently recovery in chronic radiation sickness (Gubina & Omelyanenko, 1959). Polysaccharides isolated from E. sphaerocephalus have been used in traditional Chinese medicine as drugs with anti-inflammatory effect and anti-tumor promoting action in the osseous system (Horvath et al., 1998).

Except for its useful peculiarities discussed above and though E. sphaerocephalus may become a weed in some areas, it is used as a honey


plant: E. sphaerocephalus and E. ritro were planted in Michigan, USA (Wrobleswka et al., 1993). Values of honey efficiency and attraction for bees were found highest among those recommended for gardens as investigated in ten plants in Poland (Jabonski, 1990). In addition, Burzynski et al. (1981) indicated that several species of shrubs and herbaceous plants have been introduced into young Polish Scots pine stands as nectar plants, the most effective being Peucedanum oreoselinum L., Daucus carota L., Echinops sphaerocephalus and Asclepias syriaca L.

The genus Larinus Dejean comprising ca. 180 species is represented by approximately 140 species in the Palaearctic (Csiki, 1934). Larinus is species-richest in the Mediterranean where about 100 species occur (Ter-Minassian, 1967). Host plant range is confined to the tribe Cardueae of the family Asteraceae (Zwölfer et al., 1971).

According to Ter-Minassian (1967), Larinus onopordi (F.) host plant is E. sphaerocephalus. Though this weevil has a large distribution area and is common, there is no biological record published in current literature. In the present paper, seasonal occurrence and biology of L. onopordi were described on the widely perceived plant E. sphaerocephalus, for the first time.

MATERIAL & METHODS

The study was conducted in Northeastern Anatolia in 2003; collections were also made in Eastern Anatolia and Mediterranean Turkey in 2004 and 2005. Biological observations were made weekly (sometimes with intervals of 10–15 days) in the research areas throughout the season. Different biological stages of L. onopordi feeding on the both Echinops species were observed mostly in natural habitats; observations were made also in the laboratory and in the rearing cage in the field. In the provinces of Northeastern Anatolia where the weevil is common, it was collected from 20 flowerheads randomly at every visit and dissected under microscope in the laboratory. Seasonal occurrence of biological stages and duration of stages were described in the field.

RESULTS

This species main host plant is E. sphaerocephalus L. in Northeast Anatolia, but rarely prefers E. orientalis Truatv. to complete their generation in this plant. Overwintered adults associated with host plants from mid May. After feeding several days on leaves (Fig. 1), they mated (Fig. 2) and laid their eggs in flowerheads using rostrum by excavation of the oviposition hole. In addition, the adults damage the stem below flowerhead opening vertical holes. Adult activity period lasted approximately 44 days (Fig. 9). Adult population rise to a peak level in the last week of May and adult occurrence per flowerhead is 0.05-0.9 (0.59) (Table, 1-2). Mating activity duration lasted approximately the


adult activity period which is 37 days. After depositing eggs, females closed on eggs a pale green secretion then this secretion dried, hardened and turned black (Fig. 3). The eggs were laid superficially and it was easy to see the eggs when opening this dry secretion (Fig. 4). There were 1-23 (3.76) eggs in one flowerhead and each flowerhead had 0.35-8.2 (3.76) eggs, on average which oviposit on upper and lateral part of flowerhead (Table 2). Eggs occurrence span was 33 days (Fig. 9). Hatching and feeding in flowerhead larvae continue to grow (Fig. 5.) untill mid September. Larval period duration was 79 days and each flowerhead had 0.2-1.55 (0.84). From the third week of July, pupae were seen in flowerhead (Fig. 6). In one flowerhead, four larvae can grow, also mature larva made up the pupal cell by using mouth secretion and plant remnant where 4 pupal cells occur. If there are more than one larva or pupa in one flowerhead, this flowerhead abnormally shaped. Pupa duration was 51 days and there were 0.2-0.7 (0.5) pupa on each flowerhead, on average. Upon reaching the new generation adult stage, they waited in the flowerhead 7-10 days (Fig. 7) and emerged from here for find a suitable hibernating site such as under stones. This weevil showed aggregation behavior which is possibly seen in 1-3 specimens together hiding under a stone for hibernation. Thus, L. onopordi produced one generation per year.

A braconid wasp, Bracon facialis Thom. (Hymenoptera: Braconidae) reared from the pupal stage of L. onopordi.

DISCUSSION

As field investigation results show, L. onopordi main host plant is E. sphaerocephalus (Fig. 8), however, it can complete their generation on E. orientalis in NE Anatolia. In addition, this weevil species used two different Echinops species and E. sphaerocephalus in SE Anatolia, Central Anatolia and Mediterranean Turkey as a host plant. Conducted in 2004-2005 field expedition covered approximately half of Turkey (eastern part) examining 106 different localities, L. onopordi host range was confined to only the Echinops genus. According to current literature, L. onopordi host plants were E. sphaerocephalus and E. ritro (Ter-Minassian, 1967; Campobasso et al., 1999). This weevil is distributed in Algeria, Tunisia, Syria, Lebanon, Iran, Egypt, Caucasus, Cyprus, Greece, Turkey, Italy, southeastern Ukraine, Lower Volga region, Transcaucasus, southern Kazakhstan, Turkmenistan, Tajikistan, southern Europe (Petri, 1907; Csiki, 1934; Ter-Minassian, 1967; Fremuth, 1982). In the eastern part of Turkey, this weevil is distributed in Adana, Adıyaman Artvin, Bingöl, Bitlis, Diyarbakır, Elazığ, Erzurum, Erzincan, Gaziantep, Hatay, Iğdır, Kars, Kırıkkale, Kilis, Malatya, Nevşehir, Osmaniye, Sivas and Şanlıurfa provinces.

Kasparyan & Gültekin, (2002) informed that an ichneumonid wasp, Exeristes roborator F. (Hymenoptera: Ichneumonidae) is parasitoid of this species. By this present paper an additional parasitoid is recorded


which is including Braconidae. Even if this plant is on the weed list of Turkey (Uluğ et. al., 1993;

Okşar & Uygur, 2000), E. sphaerocephalus is being used as both medicinal plant (Gubina & Omelyanenko, 1959; Kucherov, 1978; Horvath et al., 1998) and cultivated as a honey garden plant (Jabonski, 1990; Wrobleswka et al., 1993). There is not sufficient data about the economical importance of this plant to come to final conclusions regarding this is invasive weed. In investigated region this plant occur roadsides, field margins, abandoned fields and non-cultivated natural habitats commonly. Here, it is not suitable to declare E. sphaerocephalus as an invasive plant and L. onopordi a potential biological control agent. It is possible to discuss, here, the main feeder of E. sphaerocephalus is L. onopordi which is consuming seed of this plant. Accordingly its biological peculiarities and the relative population level, this weevil can balance this plant population level and prevent for jumping an invasive weed plant group.

ACKNOWLEDGEMENTS

I am sincerely grateful to Dr. Vladimir I. Dorofeyev (Botanical Institute, Russian Academy of Sciences, St. Petersburg, Russia) for identification of plants, Dr. Boris A. Korotyaev (Zoological Institute, Russian Academy of Sciences, St. Petersburg, Russia) for giving an opportunity to visit his laboratory, and Dr. S. Belokobyl'skii (Zoological Institute, Russian Academy of Sciences, St. Petersburg for identification of braconid wasps. The study was supported by the Collaborative Linkage Grants No. 978845 and, NR-CLG-981318 of the NATO Life Science and Technology Programme and TUBITAK–TOVAG–1050038.

LITERATURE CITED

Burzynski, J., Rodziewicz, A.& Kolk, A. 1981. Attractiveness of nectar plants to parasites of forest insects, especially those of the European pine shoot moth (Rhyacionia buoliana). Prace Instytutu Badawczego Lesnictwa Poland. 1981 (584/589): 75-78. Campobasso, G., Colonnelli, E., Knutson, L., Terragitti, G. & Cristofaro, M. 1999. Wild Plants and Their Associated Insects in the Palearctic Region, Primarily Europe and the Middle East. United States Department of Agriculture, ARS-147, IV +, Agricultural Research Service, Washington, USA., 243 pp. Csiki, E. 1934. Coleopterorum Catalogus auspiciis et auxilio W. Junk editus a S. Schenkling. Pars 134. Curculionidae: subfam. Cleoninae. Junk, Berlin, 152 pp. Davis, P. H. 1975. Flora of Turkey and the East Aegean Islands. Edinburg University Press, Edinburgh, 890 pp. Fremuth, J. 1982. Cleoninae aus der Türkei und den angrenzenden Gebieten. Fragmenta Entomologica 16 (2): 239-258.


Gubina, G. P. & Omelyanenko, L. M. 1959. Some data on Echinopsin alkaloid treatment of patients for chronic radiation sickness. Vestnik Rentgenologii i Radiologii 34(5): 29-34. Hartman, R. L. & Nelson, B. E. 2000. Working List of Invasive Vascular Plants of Wyoming, With Vernacular Names from Major Works. http://www.rmh.uwyo.edu/ wyinvasives/wyweeds.pdf Horvath, Z., Gyemant, G., Danos, B. & Nanasi, P. 1998. Investigation of polysaccharides of Echinops species. Medicinal plant polysaccharides I. Acta Pharmaceutica Hungarica 68(4):214-219. Jabonski, B. 1990. Nectar secretion and honey-yielding efficiency of important honey plants grown in Poland. Pszczelnicze Zeszyty Naukowe 34: 43-49. Kasparyan, D. R. & Gültekin, L. 2002. First records of two ichneumonid wasps from Northeastern Turkey (Hymenoptera: Ichneumonidae). Zoosystematica Rossica 11(1): 218. Kucherov, E. V. 1978. The medicinal flora of Bashkiria and its rational utilization. Herba Hungarica 17 (1): 5-10. McCarty, M. K., Scifres, C. J. & Robinson, L. R. 1984. A descriptive guide for major Nebraska thistles. Nebraska Agricultural Experiment Station; (SB 493): 27. Okşar, M. & Uygur, S. 2000. Weeds and their biological control possibilities in the Çukurova Region. Türkiye Herboloji Dergisi 3 (1): 27-36. Petri, K. 1907. Bestimmungs-Tabellen der europäischen Coleopteren. LX. Heft (60.), Curculionidae II, Genus Larinus und Verwandte. E. Reitter, Brünn, pp. 51-146. Ter-Minassian, M. E. 1967. Zhuki-dolgonosiki podsemejstva Cleoninae fauny SSSR. Tsvetozhily i stebleedy (triba Lixini). Nauka, Leningrad. (English translation: Weevils of the Subfamily Cleoninae in the Fauna of the USSR. Tribe Lixini. USDA Agricultural Research Service, Washington, D. C. by Amerind Publishing Co. Pvt. Ltd., New Delhi, 1978, 166 pp.). Uluğ, E., Kadıoğlu, İ.& Üremiş, İ. 1993. Weeds of Turkey and Some Characteristic. Tarım ve Köyişleri Bakanlğı, Zirai Mücadele Araştırma Enstitisü Müdürlüğü, Adana, No: 78. 513 pp. Wrobleswka, A., Ayers, G. S. & Hoopingarner, R. A. 1993. Nectar production dynamics and bee reward: a comparison between Chapman's honey plant (Echinops sphaerocephalus L.) and blue globe thistle (Echinops ritro L.). American Bee Journal 133 (11): 789-796. Zwölfer, H., Frick, K. E. & Andres, L. A. 1971. A study of the host plant relationships of European members of the genus Larinus (Col: Curculionidae). Technical Bulletin of the Commonwealth Institute for Biological Control 14: 97–143.

[Received September 2005. Accepted January 2006]


Figure 1-8. Biological stages of Larinus onopordi (F.): 1- adult and feeding damage on leaves. 2- mating. 3- oviposition mark. 4- eggs. 5- larva and pupa. 6- pupa. 7- new generation adult. 8- host plant and habitat.


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Table 1. Presence and seasonal activity of Larinus onopordi (F.) in NE Anatolia in 2003

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4.5.2003 Kars 0 0 0 0 0 0 0 0 (not bud)

6.5.2003 Bingöl 0 0 0 0 0 0 0 0 (not bud)

19.5.2003 Bingöl 1 0 0 0 0 0 0 20

26.52003 Bingöl 28 4 0 0 0 0 0 20

1.6.2003 Kars 18 5 27 0 0 0 0 20

8.6.2003 Bingöl 17 3 164 0 0 0 0 20

11.6.2003 Kars 8 2 90 12 0 0 0 20

17.6.2003 Erzincan 5 1 88 20 0 0- 0 20

3.7.2003 Bingöl 6 2 7 31 0 0 0 20

22.7.2003 Kars 0 0 0 22 4 0 0 20

3.8.2003 Kars 0 0 0 16 9 1 0 20

30.8.2003 Erzincan 0 0 0 13 14 4 2 20

14.9.2003 Erzincan 0 0 0 4 13 12 7 20

Table 2. Relative population density of Larinus onopordi (F.) in NE Anatolia in 2003

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19.52003 Bingöl 0 0.05 0 0 0 0 0

26.52003 Bingöl 0 1.4 0 0 0 0 0

1.6.2003 Kars 45 0.9 1.35 0 0 0 0

8.6.2003 Bingöl 75 0.85 8.2 0 0 0 0

11.6.2003 Kars 75 0.4 4.5 0.6 0 0 0

17.6.2003 Erzincan 70 0.25 4.4 1.0 0 0 0

3.7.2003 Bingöl 80 0.30 0.35 1.55 0 0 0

22.7.2003 Kars 65 0 0 1.1 0.2 0 0

3.8.2003 Kars 75 0 0 0.8 0.45 0.1 0

30.8.2003 Erzincan 85 0 0 0.65 0.7 0.2 0.1

14.9.2003 Erzincan 80 0 0 0.2 0.65 0.6 0.35

Total average

72.2 0.59 3.76 0.84 0.5 0.3 0.225


AN EVALUATION ON THE KNOWN TAXA OF NECYDALINAE AND APATOPHYSEINAE

FROM TURKEY WITH A SYN. NOV. (COLEOPTERA: CERAMBYCIDAE)

Hüseyin Özdikmen* and Semra Turgut* * Gazi Üniversitesi, Fen-Edebiyat Fakültesi, Biyoloji Bölümü, 06500 Ankara, TÜRKİYE, e-mails: [email protected] and [email protected] [Özdikmen, H. & Turgut, S. 2006. An evaluation on the known taxa of Necydalinae and Apatophyseinae from Turkey (Coleoptera: Cerambycidae). Munis Entomology & Zoology 1 (2): 199-204]

ABSTRACT: As a summary of the present paper, the taxa of Turkish Necydalinae and Apatophyseinae fauna that were recorded by various authors from different localities of Turkey are evaluated zoogeographically. For this reason, the distribution patterns in Turkey and rest of the world for each taxon of related subfamilies are determined. Also, chorotypes for each taxon are given in the part of zoogeographical remarks. As a result of the present paper, Necydalis hadullai Szallies, 1994 is a syn. nov. of Necydalis ulmi Chevrolat, 1838. KEY WORDS: Necydalinae, Apatophyseinae, Cerambycidae, Coleoptera, Zoogeography, Fauna, Turkey.

The longhorn beetles or Cerambycidae are classified together with Chrysomelidae and Bruchidae in the superfamily Chrysomeloidea. But, some authors recognized Cerambycidae as a separate superfamily Cerambycoidea (Svacha & Danilevsky, 1986). Cerambycidae sensu stricto is divided into several subfamilies. These are Parandrinae, Prioninae, Lepturinae, Necydalinae, Spondylidinae, Apatophyseinae, Cerambycinae and Lamiinae. Parandrinae are not represented in Turkey. Moreover, as seen below, the subfamilies Necydalinae and Apatophyseinae are represented only a few species in Turkey. Also, the recorded data of both subfamilies Necydalinae and Apatophyseinae from Turkey have not been adequate since only a few works have been presented on Turkish fauna of Necydalinae and Apatophyseinae until now.

Subfamily Necydalinae Tribe Necydalini

Genus Necydalis Linnaeus, 1758 Necydalis hadullai Szallies, 1994

Current records from Turkey: The species is known only from type locality in Turkey. Type loc.: Isparta province: Kovada, S. Eğridir. Remarks: This name is available but invalid as being a junior subjective synonym of Necydalis ulmi Chevrolat, 1838. The species is currently placed in the genus Necydalis.

As seen below, Szallies (1994) stated that “Necydalis hadullai n. sp. seems to be substitute for Necydalis ulmi in the South Turkey; the differences seem to me significant enough to describe Necydalis hadullai


n. sp. as own species.” However, Necydalis ulmi is recorded for the first time for Turkey in 2005. The real status of distribution patterns in Turkey of N. ulmi need to be clarified. Szallies (1994) mentioned “the last sternite only behind the middle most broadly, stretched. The anal plate (between tergite and sternite) is notched. The points of the paramers are rounded” as the most diagnostic characters. But the original description of Necydalis hadullai is based on only one specimen. These characters are very likely individual variations. Moreover, the other mentioned differences between N. hadullai and N. ulmi by Szallies (1994) does not seem distinguishable enough. Finally we think the original description of N. hadullai does not seem significant enough to describe a new species.

The original description of N. hadullai was given by Szallies (1994) as follows: “Beschreibung: Flügeldecken kastanienbraun, hintere Seitenränder geschwärzt. Fühler einfarbig kastanienbraun, die vier ersten Glieder glänzender, die restlichen matt. Die Beine ebenfalls kastanienbraun, Hinterschenkel und -tibien an den Spitzen schwarz, die Spitze der Mittelschenkel auf jeder Seite mit schwarzem Fleck. Hintertarsen gelb. Das 1. Sternit bis auf den Hinterrand, das 4. und 5. ganz schwarz, ebenso das letzte Tergit, sonst ist der Hinterleib rotbraun.

Schildchen schwarz, gerunzelt und fast flach, hinten eine kleine Längsvertiefung aufweisend. Zueinander gelegene Seitenränder der Flügeldecken zum Apex hin deutlich verbreitert. - Halsschild mit zwei glatten Beulen und einem Wulst vor dem Hinterrand. Um diese Erhebungen herum lang goldgelb behaart (wie bei Necydalis ulmi Chevrolat).

Letztes Sternit erst hinter der Mitte am breitesten, gestreckt. Die Analplatte (schaut hinten zwischen Tergit und Sternit heraus) ist eingekerbt (Abb. 2). Die Spitzen der Parameren sind abgerundet (Abb. 4).

Grösse 31 mm. Anmerkung: Necydalis hadullai n. sp. steht der bekannten Art

Necydalis ulmi nahe, so dass die Beschreibung ihn in erster Linie von diesem abgrenzt; ergänzende Zeichnungen der wesentlichen Merkmale beider Arten sollen dabei helfen (Abb. 2 - 5). Necydalis hadullai n. sp. scheint Necydalis ulmi in der Südtürkei zu ersetzen; die Unterschiede scheinen mir bedeutend genug, Necydalis hadullai n. sp. als eigene Art zu beschreiben.

Das Tier wurde von Quercus geklopft.”

Necydalis major major Linnaeus, 1758 Current records from Turkey: The species has been reported by Acatay (1948, 1961, 1968) and Lodos (1998) from Turkey without exact localities. Range: Europe, Siberia (Winkler, 1924-1932); European Russia, Ciscaucasia, the Black Sea coast of Caucasus, all Siberia, Sakhalin, Europe, Japan (Danilevsky & Miroshnikov, 1985); temperate Palaearctics


from Europe to Far East (Svacha & Danilevsky, 1986); Europe (Bense, 1995); Europe, European Russia, European Kazakhstan (Althoff & Danilevsky, 1997); Europe, Caucasus, Transcaucasia, Siberia, In Asia: from Northern Kazakhstan, Altai Mts., Tuva, Amur basin and Primorie region to China, Korean peninsula and Sakhalin (Sama, 2002); Europe, Caucasus, Balkan peninsula, Crimea peninsula, Siberia, Far East of the Russia, China and Korea (Danilevsky, 2005). Remarks: The species Necydalis major Linnaeus, 1758 has two subspecies in the world. These are the nominate Necydalis major major Linnaeus, 1758 and the other Necydalis major aino Kusama, 1975. This species is represented by nominative subspecies in Turkey. Unfortunately any record with exact locality in Turkey has not been given according to the literature. For this reason the real status in Turkey of this taxon need to be clarified. A general distribution of this taxon is given above. The other subspecies Necydalis major aino Kusama, 1975 occurs only in Japan, Sakhalin and Kurile Islands and ? Mongolia according to Kusama & Takakuwa (1984) and Danilevsky (2005). Also accordnig to Danilevsky (2005) Necydalis major major Linnaeus, 1758 and Necydalis major aino Kusama, 1975 are represented in Russia by subspecies.

Necydalis sabatinelli Sama, 1994 Current records from Turkey: There are four records from Turkey. These are: Type loc.: Bolu province: Abant; Bolu province: Abant as Necydalis sp. (Sama, 1982); Bolu province: Abant (Tauzin, 2000); Bolu province: Yedigöller National Park (Malmusi & Saltini, 2005). Range: Endemic to Turkey. Apparently distributed only in a local area of North Turkey.

Necydalis ulmi Chevrolat, 1838 = Necydalis hadullai Szallies, 1994 syn. nov.

Current records from Turkey: There is only one record from European Turkey as Kırklareli province: Demirköy (Malmusi & Saltini, 2005) and also there is only one record from South-West Anatolia as Isparta province: Kovada, S. Eğridir (as Necydalis hadullai Szallies, 1994). Range: Europe (Winkler, 1924-1932); Caucasus, Transcaucasus, Europe, Southwest Ukraine (Danilevsky & Miroshnikov, 1985); Central and Southern Europe, Caucasus, Transcaucasia (Svacha & Danilevsky, 1986); Europe (Bense, 1995); (Althoff & Danilevsky, 1997); Europe, Caucasus, Transcaucasia (Sama, 2002); Europe, Balkan Peninsula, Caucasus (Danilevsky, 2005). Remarks: The species has been known only from European Turkey (=Thracia) of Turkey until this study. At present, it is also known in South-West Anatolia due to the record of Isparta province of Necydalis hadullai Szallies, 1994 syn. nov. The real status of distribution patterns in Turkey of this taxon need to be clarified.


Subfamily Apatophyseinae Tribe Apatophyseini

Genus Apatophysis Chevrolat, 1860 Apatophysis anatolica Heyrovsky, 1938

Current records from Turkey: There are three records from Turkey. These are: Type loc.: Konya province: Akşehir; Aksaray province: Tuz Lake, Eskil (Adlbauer, 1992); Konya province: Karapınar (Özdikmen & Demir, 2006). Range: Endemic to Turkey. Apparently distributed only in Central Turkey.

Apatophysis caspica Semenov, 1901 Current records from Turkey: There are two records from Turkey. Konya province: Tuz Lake and Akşehir (Demelt, 1967); Iğdır province: Reynhanli and Kars province: S. E. slopes of Ararat (Villiers, 1967). Range: Transcaspia (Winkler, 1924-1932); Turkmenia, the western coast of the Caspian sea to the south from Derbenta, Caucasus, Armenia, Northern Iran (Danilevsky & Miroshnikov, 1985); Jordan (Sama et al., 2002); Caucasus, Iran, Central Asia, Near East (Danilevsky, 2005). Remarks: Although two records are known from Turkey, both records must be corrected. Primarily, the records in Villiers (1967) are from Iğdır province which is near Armenian and Iranian borders too. Because S. E. slopes of Ararat is in Iğdır province of Turkey in the present day, we think that these records are unsuspicious according to known distributional patterns in the world. Unfortunately the record in Demelt (1967) seems to be suspicious because of the known distributional patterns of this species in the world. So it probably belongs to the other species Apatophysis anatolica Heyrovsky, 1938 known only from Central Anatolia such as above. Moreover apparently even these two species may be synonyms. Without study the type materials, the recent record from Jordan in Sama et al. (2002) indicate possibility of this status. For this reason the real systematical and taxonomical status of these species need to be clarified.

ZOOGEOGRAPHICAL REMARKS The present zoogeographical characterization is based on the chorotype classification of Anatolian fauna, recently proposed by Vigna Taglianti et al. (1999). Necydalis major major Linnaeus, 1758 has the Sibero-European chorotype. Necydalis ulmi Chevrolat, 1838 has the European chorotype. Apatophysis caspica Semenov, 1901 has the Turanian chorotype. Necydalis sabatinelli Sama, 1994 and Apatophysis anatolica Heyrovsky, 1938 are Anatolian endemic species only known with a few specimens.


ACKNOWLEDGEMENTS We thank my friend Dr. Mikhail L. Danilevsky (Russia) for his very valuable comments and contribution, also for supplying us with a photo of the holotype N. hadullai. Also thanks to Dr. Roberto Fabri (Italy) for his help to reach the literature Malmusi & Saltini (2005).

LITERATURE CITED

Acatay, A. 1948. Zararlı orman böcekleri teşhis anahtarı. T. C. Tarım Bakanlığı Orman Genel Müdürlüğü yayınları, Özel Sayı: 76, Ocak Matbaası, İstanbul, Türkiye, 113 pp. Acatay, A. 1961. Zararlı orman böcekleri teşhis anahtarı. İstanbul Üniversitesi Orman Fakültesi yayınları No: 76, Fakülteler Matbaası, İstanbul, Türkiye, 152 pp. Acatay, A. 1968. Zararlı orman böcekleri teşhis anahtarı. İstanbul Üniversitesi Orman Fakültesi yayınları No: 132, Kutulmuş Matbaası, İstanbul, Türkiye, 154 pp. Adlbauer, K. 1992. Zur Faunistik und Taxonomie der Bockkäferfauna der Türkei II (Coleoptera, Cerambycidae). Entomofauna, 13 (30): 485-509. Althoff, J. & Danilevsky, M. L. 1997. A Check-List of Longicorn Beetles (Coleoptera, Cerambycoidea) of Europe. Slovensko Entomološko Društvo Štefana Michielija. Ljubljana, 64 pp. Bense, U. 1995. Illustrated key to the Cerambycidae (excl. Dorcadionini) and Vesperidae of Europe. Margraf Verlag, Germany, 512 pp. Danilevsky, M. L. 2005. Systematic list of Longicorn Beetles (Cerambycoidea) of the territory of the former USSR (on the base of "Systematic list of Longicorn Beetles of the USSR" by A.L. Lobanov, M.L. Danilevsky, S.V. Murzin, 1981, and computer databases by A.L. Lobanov, 1979-1990). Available from: http://www..zin.ru/Animalia/ Coleoptera/eng/dbase30. htm Danilevsky, M. L. & Miroshnikov, A. I. 1985. The longicorn beetles of the Caucasus (Coleoptera, Cerambycidae) A key book. Krasnodar, USSR, 419 pp. Demelt, C. V. 1967. Nachtrag zur Kenntnis der Cerambyciden-Fauna Kleinasiens. Entomologische Blatter, 63 (2): 106-109. Kusama, K. & Takakuwa, M. 1984. Parandrinae. Prioninae. Spondylinae. Aseminae. Lepturinae (part.). Cerambycinae. Lamiinae (part.). The Longicorn-beetles of Japan in Color. Kodansha, Tokyo: 131-172, 201-493, 511-549. Lodos, N. 1998. Entomology of Turkey VI (General, Aplied and Faunistic). Ege Ü. Ziraat Fak. Yayınları No: 529, E. Ü. Faculty of Agriculture Press, İzmir, 300 pp. Malmusi, M. & Saltini, L. 2005. Cerambycidae raccolti dai componenti del Gruppo Modenese Scienze Naturali durante escursioni in Turchia tra il 1987-2003 (Contributo alla Fauna dei Cerambycidae di Turchia). Quaderno di studi e notizie di storia naturale della Romagna, n. 21, 28 pp. (unpublished). Özdikmen, H. & Demir, H. 2006. Notes on longicorn beetles fauna of Turkey (Coleoptera: Cerambycidae). Munis Entomology & Zoology, 1 (1): 157-166. Sama, G. 1982. Contributo allo studio dei coleotteri Cerambycidae di Grecia e Asia Minore. Fragmenta Entomologica, Roma, 16 (2): 205-227.


Sama, G. 1994. Cerambycidae nuovi o poco noti del Mediterraneo Orientale (Coleoptera, Cerambycidae). Lambillionea 94 (1): 9-13. Sama, G. 2002. Atlas of the Cerambycidae of Europe and the Mediterranean Area, Volume I, Kabourek, Zlin, 173 pp. Svacha, P. & Danilevsky, M. L. 1986. Cerambycoid larvae of Europe and Soviet Union (Coleoptera, Cerambycoidea). Part I. Acta Universitatis Carolinae – Biologica 30 (1986): 1-176. Szallies, A. 1994. Drei neue Coleopteren-Arten aus der Türkei (Coleoptera: Prostomidae, Cerambycidae). Entomol. Z. 104 (13): 259-263. Taglianti, A. V., Audisio, P. A., Biondi, M., Bologna, M. A., Carpaneto, G. M., De Biase, A., Fattorini, S., Piattella, E., Sindaco, R., Venchi, A. & Zapparoli, M. 1999. A proposal for a chorotype classification of the Near East fauna, in the framework of the Western Palaearctic Region. Biogeographia 20: 31-59. Tauzin, P. 2000. Complement a l’inventaire des Coleopteres Cerambycidae de Turquie. L’Entomologiste, 56 (4): 151-153. Villiers, A. 1967. Coléoptéres Cérambycides de Turquie (1. Partie) - L’ Entomologiste, 23 (1): 18-22. Winkler, A. 1924-1932. Catalogus Coleopterorum regionis palaearcticae. Verlag von Albert Winkler, 1135-1226.

[Received October 2005. Accepted January 2006]

Fig. 1. The provinces of Turkey.


TWO NEW SUBSPECIES OF CARABUS (LIMNOCARABUS) CLATHRATUS LINNÉ, 1761 (COLEOPTERA, CARABIDAE) FROM SIBERIA.

Dmitry Obydov*

*Timiryazev’s State Museum of Biology, 123242, Malaya Gruzinskaya str. 15, Moscow, RUSSIA. e-mail: [email protected] [Obydov, D. 2006. Two new subspecies of Carabus (Limnocarabus) clathratus Linné, 1761 (Coleoptera: Carabidae) from Siberia. Munis Entomology & Zoology, 1 (2): 205-210] ABSTRACT: Two new subspecies of Carabus (Limnocarabus) clathratus Linné, 1761: Carabus (Limnocarabus) clathratus jacutensis ssp. n. and Carabus (Limnocarabus) clathratus narymicus ssp. n. are described from Siberia. The distinguishing characters are discussed. KEY WORDS: Coleoptera, Carabidae, Carabus (Limnocarabus) clathratus, new subspecies, Siberia.

Carabus (Limnocarabus) clathratus has been described without indication of locality. The species is widely distributed in Eurasia: from southern France and northern British Islands to Central Yakutia (Russia) to the east and to Azerbajan and northern Iran to the south. Closely related species Carabus (Limnocarabus) maacki Morawitz, 1862 is distributed in Russia, from south-eastern Transbaikalie to South Ussuri Land. The wide area of Carabus clathratus is inhabited by many unequally differing populations. Some of the groups of populations, undoubtedly, must be considered as subspecies. Carabus clathratus clathratus is distributed in northern, central and eastern Europe. Widely distributed subspecies Carabus clathratus auraniensis Müller, 1902 inhabit Balkans and Turkey; Carabus clathratus stygius Ganglbauer, 1890 – Azerbajan and northern Iran; Carabus clathratus antonelli Luigioni, 1921 – central Italy; Carabus clathratus arelatensis Lapouge, 1903 – southern France; Carabus clathratus jansoni Kraatz, 1890 inhabit Scotland. Thus, the majority of populations constitutes a continuous sequence of forms slightly different in size, coloration and ratios, but it is hardly possible to consider these forms as a subspecies. Now a considerable number of subspecies of Carabus clathratus are described. The majority of subspecific names must be regarded as synonyms. Earlier it was considered that nominotypical subspecies of Carabus clathratus is distributed in Russia from western borders up to central Yakutia, though widely distributed species often is represented by the extremely detached subspecies at the edge of the area. The description of two new subspecies of Carabus clathratus from Siberia is given below. Carabus clathratus is the typical of the wetlands, on the banks of canals, under moss or cut canes, in fallen tree trunks as well as in the


water habitat. Indeed, it can stay for long periods under water and is a good swimmer and underwater hunter.

DESCRIPTION Carabus (Limnocarabus) clathratus jacutensis ssp. n. (Fig. 1).

Holotype: male with label: “Yakutsk, 27.VI.1994” (the collector is not known). Papatypes: 2 males, female, same data and same locality; 4 males, 3 females: “Yakutia, Yakutsk env., Berdigestyakh, 1-5.VII.1994” (the collector is not known). The holotype is preserved in the collection of the State Museum of Biology (Moscow, Russia). The paratypes are preserved in the collection of the State Museum of Biology (Moscow, Russia) and in the collection of Mr E. Tarasov (Moscow, Russia).

Body length in males is 23.5 - 24.0 mm (including mandibles), width

8.3 - 8.8 mm; body length in females is 23.8 - 24.2 mm, width 8.3 - 9.0 mm.

Head not thickened, ratio width of pronotum/width of head 2.21; eyes strongly convex; mandibles short, broad, evenly curved; surface of mandibles smooth; terebral tooth of the right and left mandibles bi-dentate, strongly prominent; retinaculum of the right mandible strongly prominent, retinaculum of the left mandible less prominent. Frontal furrows short and shallow, inside smooth. Frons, vertex and neck with dense fine punctures, laterally frons with few coarse wrinkles. Labrum wider than clypeus, moderately notched, without lateral setae. Antenna protruding beyond the base of pronotum by three-four apical segments; palpi slightly dilated; penultimate segment of the maxillary palpi equal to the last segment; penultimate segment of the labial palpi with two setae. Mentum tooth narrow, triangular, equal or slightly longer than lateral lobes; submentum with two setae.

Prothorax very broad, broadest at about middle; ratio width/length 1.47. Pronotum convex with sparse fine punctuation and sparse fine wrinkles. Median longitudinal line distinct; basal foveae very deep and large, inside finely-punctured. Sides of pronotum broadly margined; lateral margin slightly bent upwards; lobes of hind angles short, evenly rounded, slightly bent downwards. Lateral margin with two lateral setae. Elytra oval, relatively convex, widest at about middle; ratio length/width 1.60; ratio width of elytra/width of pronotum 1.33; shoulders prominent; sides of elytra broadly margined, elytral margin coarsely-granular. Elytral sculpture heterodynamous; secondary elytral interspaces strongly convex, more convex than in Carabus clathratus


clathratus; striae coarsely-punctured; primary elytral foveoles large, well-marked. Wings are reduced or partially reduced.

Ventral body surface smooth, metepisternum finely-punctured, slightly longer than its width; sides of abdomen slightly wrinkled; sternal sulci absent.

Shape of aedeagus and endophallic structure are characteristic for the species.

Legs of normal length; male fore tarsi with four dilated segments bearing hairy pads.

Coloration very bright, green with metallic lustre; often margins of pronotum and elytrae bronze or reddish bronze; primary elytral foveoles bronze, reddish bronze or goldish bronze; mandibles, antenna, palpi, legs and ventral body surface black. Differential diagnosis. The new subspecies differs from nominotypical subspecies by more dense punctuation of frons, vertex and neck; more convex pronotum and elytra, more convex secondary elytral interspaces. Coloration of the new subspecies brighter, green with metallic lustre and bronze or reddish bronze elytral margins (in nominotypical subspecies coloration dim, black with green lustre, pronotum and elytra without colour margins). Body of the new subspecies shorter (23.5 - 24.2 mm, in C. clathratus clathratus body length is 27.0 - 34.0 mm). Distribution. Russia, East Siberia, Central Yakutia. Habitat. The beetles are connected to the coasts of the reservoirs of various types and usually do not occur far from water. Carabus (Limnocarabus) clathratus narymicus ssp. n. (Fig. 2). Holotype: male with label: “West Siberia, Narym Distr., Piakurur River Vall., 6.VII.1990, T. Buchatzkaya leg.” Papatypes: male, 2 females, same data and same locality. The holotype and the paratypes are preserved in the collection of the State Museum of Biology (Moscow, Russia).

Body length in males is 23.0 - 24.3 mm (including mandibles), width 8.4 - 8.6 mm; body length in females is 24.6 - 27.2 mm, width 8.5 - 10.2 mm.

Head not thickened, ratio width of pronotum/width of head 2.0; eyes strongly convex; mandibles short, broad, slightly curved; surface of mandibles smooth; terebral tooth of the right and left mandibles bi-dentate, slightly prominent; retinaculum of right and left mandibles moderately prominent. Frontal furrows deep and long, inside smooth or


with few fine punctures. Frons nearly smooth; vertex and neck with sparse fine punctures, laterally frons and vertex with few coarse wrinkles. Labrum wider than clypeus, moderately notched, without lateral setae. Antenna protruding beyond the base of pronotum by three-four apical segments; palpi moderately dilated; penultimate segment of the maxillary palpi equal to the last segment; penultimate segment of the labial palpi with two setae. Mentum tooth narrow, triangular, slightly shorter than lateral lobes; submentum with two setae.

Prothorax broad, broadest at about middle; ratio width/length 1.50. Pronotum weakly convex with sparse fine punctuation and sparse fine wrinkles, laterally with more rough sculpture. Median longitudinal line distinct; basal foveae deep, inside coarsely-punctured. Sides of pronotum relatively broadly margined; lobes of hind angles short, evenly rounded, slightly bent downwards. Lateral margin with two lateral setae.

Elytra oval, relatively convex, widest at about middle; ratio length/width 1.50; ratio width of elytra/width of pronotum 1.37; shoulders prominent; sides of elytra relatively broadly margined, elytral margin coarsely-granular. Elytral sculpture heterodynamous; primary elytral interspaces strongly convex (nearly like secondary) represented by rows of short links and tubercles; striae coarsely-punctured; primary elytral foveoles shallow, indistinct. Wings are reduced.

Ventral body surface smooth, metepisternum nearly smooth, slightly longer than its width; sides of abdomen slightly wrinkled; sternal sulci absent.

Shape of aedeagus and endophallic structure are characteristic for the species.

Legs of normal length; male fore tarsi with four dilated segments bearing hairy pads.

Coloration of pronotum and elytra one-colored black; primary elytral foveoles black, sometimes with weak reddish lustre; mandibles, antenna, palpi, legs and ventral body surface black. Differential diagnosis. The new subspecies differs from nominotypical subspecies by less dense punctuation of frons, less convex pronotum and elytra, strongly convex primary elytral interspaces, which are represented by rows of short links and tubercles and shallow and indistinct primary elytral foveoles. Coloration of pronotum and elytra of the new subspecies one-colored black; primary elytral foveoles black, sometimes with weak reddish lustre (in nominotypical subspecies coloration brighter with green lustre, primary elytral foveoles bronze, reddish-bronze or gold, well-marked). Body of the new subspecies shorter (23.0 - 27.2 mm, in C. clathratus clathratus body length is 27.0 - 34.0 mm). Remark. The new subspecies resembles Carabus (Limnocarabus) maacki Morawitz, 1862 by the character of the elytral sculpture. Distribution. Russia, West Siberia, the Ob River basin, Narym District.


Habitat. The beetles were collected on the wet meadow in the valley of Piakurur River.

ACKNOWLEDGEMENTS

I wish to express my heartfelt gratitude to Mr Evgeny Tarasov (Moscow) and Mrs Tatiana Buchatzkaya (Moscow) who provided me with the materials for study.

LITERATURE CITED

Ganglbauer L. 1890. Carabus (Limnocarabus) stygius n. sp. Wien. ent. Ztg. 9: 117. Kraatz G. 1890. Ueber Varietäten des Carabus (Limnocarabus) clathratus. Wien. ent. Ztg. 9: 187-188. Lapouge G. 1903. Tableaux de détermination des formes du genre “Carabus”. L’Echange. Lyon 25: 164-166, 181-182. Linné C. 1761. Fauna Suecica sistens Animalia Sueciae. Regni: Mammalia, Aves, Amphibia, Pisces, Insecta, Vermes. Distributa per Classes et Ordines, Genera et Species, cum Differentiis Specierum, Synonymis, Austorum, Nominibus, Incolarum, Locis Natalium, Descriptionibus Insectorum. Editio Altera. Auctior.– Stockholmiae: 578 pp. Luigioni P. 1921. Coleotteri d’Italia. Mem. Pont. Acc. Nuovi Lincei, s. II, XVIII: 277 pp. Morawitz A. 1862. Vorläufige Diagnosen neuer Coleopteren aus Südost-Sibirien. Mélanges Biologiques, Bull. Acad. Sci. St. Pétersbourg, T. IV (2): 180-228. Müller J. 1902. Beschreibungen neuer dalmatinischer Koleopteren. Von Josef Müller, Supplement an der Staats-Realschule in Triest. Münch. Koleopt. Zs.: 192-194.

[Received February 2006. Accepted February 2006]


Fig. 1. Carabus (Limnocarabus) clathratus jacutensis ssp. n. (holotype).

Fig. 2. Carabus (Limnocarabus) clathratus narymicus ssp. n. (holotype).


NEOMRAZEKIELLA NOM. NOV., A REPLACEMENT NAME FOR THE GENERIC NAME MRAZEKIELLA BREHM, 1949

(CRUSTACEA: COPEPODA: CANTHOCAMPTIDAE)

Hüseyin Özdikmen* and Giuseppe L. Pesce**

* Gazi Üniversitesi, Fen-Edebiyat Fakültesi, Biyoloji Bölümü, 06500 Ankara / TÜRKİYE, e-mail: [email protected] ** Dipartimento di Scienze Ambientali, University of L’ Aquila, ITALY, e-mail: [email protected] [Özdikmen, H. & Pesce, G. L. 2006. Neomrazekiella nom. nov., a replacement name for the generic name Mrazekiella Brehm, 1949 (Crustacea: Copepoda: Canthocamptidae). Munis Entomology & Zoology, 1 (2): 211-214] ABSTRACT: A junior homonym was detected amongst the Crustacea (Copepoda, Canthocamptidae). The following replacement name is proposed: Neomrazekiella for Mrazekiella Brehm, 1949. KEY WORDS: Mrazekiella, Neomrazekiella, Crustacea, Copepoda, Canthocamptidae, homonymy, replacement name.

Whilst recently researching the “Nomenclator Zoologicus” (Neave 1939-1950) a homonymous copepod generic name was noticed. The opportunity is here taken to provide a replacement name for it in accordance with the International Code of Zoological Nomenclature (2000): Neomrazekiella nom.nov. for Mrazekiella Brehm, 1949.

Crustacea Copepoda Canthocamptidae

Neomrazekiella (nom. nov.)

Remarks. The genus-group name Brehmiella was described by Chappuis, 1929 as a subgenus belonging to the genus Attheyella Brady, 1880 in Copepoda (Crustacea). However, the genus name Brehmiella was proposed by Pascher, 1928 in Chrysomonadida (Protozoa). Therefore, Brehmiella Chappuis, 1929 was a junior homonym of Brehmiella Pascher, 1928. The junior homonym was duly changed by Brehm, 1949, who proposed the genus-group name Mrazekiella nomen novum pro Brehmiella Chappuis, 1929 in Arch. Hydrobiol., Stuttgart, 42: 515 in the family Canthocamptidae (Crustacea, Copepoda, Harpacticoidea).

However, prior to Brehm's action, the genus Mrazekiella Kijenskii, 1926 had been described in the family Radiophryidae (Protozoa, Ciliophora, Oligohymenophorea, Astomatida). The name Mrazekiella Brehm, 1949, proposed for a subgenus of the genus Attheyella Brady, 1880 (Crustacea, Copepoda, Harpacticoidea, Canthocamptidae), is preoccupied by Mrazekiella Kijenskii, 1926 (Protozoa, Ciliophora, Oligohymenophorea, Astomatida, Radiophryidae). It cannot be used as a valid genus name in Crustacea by homonymy.


Consequently, the name Neomrazekiella nom. nov. is here proposed as a replacement name. As a result of this, Mrazekiella Brehm, 1949 is a syn. nov. of Neomrazekiella nom. nov.

Mrazekiella Kijenskii, 1926 (Protozoa) is recorded in Nomenclator Zoologicus vol. 3 and Mrazekiella Brehm, 1949 (Crustacea) is recorded in Nomenclator Zoologicus vol. 6 by Neave and Edwards & Hopwood. Ethimology: from preexisting generic name Mrazekiella. Summary of nomenclatural changes: Attheyella (Neomrazekiella) nom. nov. = Attheyella (Mrazekiella) Brehm, 1949 (non Kijenskii, 1926) Attheyella (Neomrazekiella) alta Shen & Sung 1965 comb. nov. [China; fresh waters] Attheyella (Neomrazekiella) americana (Herrick, 1884) comb. nov. [USA; marsh] Attheyella (Neomrazekiella) amurensis (Borutzy, 1936) comb. nov. [URSS, China; brackish waters] Attheyella (Neomrazekiella) borutzkyi (Smirnov, 1930) comb. nov. [URSS; fresh waters] Attheyella (Neomrazekiella) byblis (Chang & Kim, 1992) comb. nov. [Korea, Japan; springs] Attheyella (Neomrazekiella) dentata dentata (Poggenpol, 1874) comb. nov. [Europe, Japan, China; fresh, brackish waters] Attheyella (Neomrazekiella) dentata americana (Herrick, 1884) comb. nov. [North America; fresh waters] Attheyella (Neomrazekiella) dentata coronata (Daday, 1913) comb. nov. [Asia, Mongolia;?] Attheyella (Neomrazekiella) dogieli (Rylov, 1923) comb. nov. [Asia, USA; fresh waters] Attheyella (Neomrazekiella) illinoisensis illinoisensis (Forbes, 1876) comb. nov. [USA, Europe, Asia; continental waters] Attheyella (Neomrazekiella) illinoisensis hyperborea (Willey, 1931) comb. nov. [North America; continental waters] Attheyella (Neomrazekiella) meridionalis (Dussart, 1982) comb. nov. [Madagascar; fresh waters, lakes] Attheyella (Neomrazekiella) mongoliana (Shen & Chang, 1966) comb. nov. [China; fresh waters] Attheyella (Neomrazekiella) naphtalica (Por, 1983) comb. nov. [Israel; springs] Attheyella (Neomrazekiella) nordenskioldii nordenskioldii (Lilljeborg, 1902) comb. nov. [URSS, Japan, USA; fresh waters] Attheyella (Neomrazekiella) nordenskioldii volgensis (Borutzky, 1952) [URSS; brackish waters]


Attheyella (Neomrazekiella) northumbrica trisaetosa (Chappuis, 1929) comb. nov. [Europe, USA; fresh waters] Attheyella (Neomrazekiella) northumbricoides (Brehm, 1923) comb. nov. [China; brackish waters] Attheyella (Neomrazekiella) osmana (Kiefer, 1955) comb. nov. [Europe, Israel; brackish waters] Attheyella (Neomrazekiella) f. otmanli (Apostolov,1969) comb. nov. [Bulgaria; brackish waters] Attheyella (Neomrazekiella) paranaphtalica (Pesce & Galassi, 1988) comb. nov. [Italy, Sicily included; phreatic] Attheyella (Neomrazekiella) pilosa (Chappuis, 1929) comb. nov. [central America; cenotes] Attheyella (Neomrazekiella) quinquespinosa (Shen & Tai, 1964) comb. nov. [China; fresh waters] Attheyella (Neomrazekiella) spinipes (Reid, 1987) comb. nov. [U.S.A.; springs] Attheyella (Neomrazekiella) stachanovi (Borutzky, 1930) comb. nov. [Sakhaline islands; brackish waters?] Attheyella (Neomrazekiella) tetraspinosa (Chang, 1993) comb. nov. [Korea; springs] Attheyella (Neomrazekiella) trispinosa (Brady, 1880) comb. nov. [Europe, Africa, Israel, fresh, brackish waters] Attheyella (Neomrazekiella) ussuriensis (Rylov, 1933) comb. nov. [URSS: brackish waters] Attheyella (Neomrazekiella) yunnanensis (Shen & Tai, 1979) comb. nov. [China; fresh waters] Attheyella (Neomrazekiella) weigoldi (Brehm, 1923) comb. nov. [China; fresh waters] Attheyella (Neomrazekiella) wulmeri wulmeri (Kerhervé, 1914) comb. nov. [Europe; fresh, interstitial waters] Attheyella (Neomrazekiella) wulmeri occidentalis (Roy, 1932) comb. nov. [France; fresh waters] Attheyella (Neomrazekiella) wulmeroides (Borutzky, 1930) comb. nov. [URSS; continental waters]

AKCNOWLEDGEMENT

Sincere thanks to Janet W. Reid (USA) for reviewing the MS.


LITERATURE CITED

Brehm, V. 1949. Prioritatsbedingte Namensanderung. Archiv fur Hydrobiologie 42: 515. Chappuis, P. A. 1929. Die Unterfamilie der Canthocamptinae. Archiv fur Hydrobiologie 20: 471-516. Edwards, M. A. & Hopwood, A. T. 1966. Nomenclator Zoologicus. The Zoological Society of London 6 (1946-1955): 177. Edwards, M. A. & Vevers, H. G. 1975. Nomenclator Zoologicus. The Zoological Society of London 7 (1956-1965). Edwards, M. A. & Tobias, M. A. 1993. Nomenclator Zoologicus. The Zoological Society of London 8 (1966-1977). Edwards, M. A., Mainly, P. & Tobias M. A. 1996. Nomenclator Zoologicus. The Zoological Society of London 9 (1978-1994). International Comission Zoological Nomenclature. 1999. The International Trust for Zoological Nomenclature, London. Neave, S. A. 1939. Nomenclator Zoologicus. The Zoological Society of London, 1. Neave, S. A. 1939. Nomenclator Zoologicus. The Zoological Society of London, 2. Neave, S. A. 1939. Nomenclator Zoologicus. The Zoological Society of London, 3: 223. Neave, S. A. 1940. Nomenclator Zoologicus. The Zoological Society of London, 4. Neave, S. A. 1950. Nomenclator Zoologicus. The Zoological Society of London, 5. Pascher A. 1928. Eine eigenartige rhizopodiale Flagellate. Archiv fur Protistenkunde 63: 239 (pp. 227-240).

[Received October 2005. Accepted February 2006]


CONTRIBUTIONS TO THE KNOWLEDGE OF TURKISH AUCHENORRHYNCHA WITH TWELVE NEW RECORDS

(HOMOPTERA: CICADELLIDAE)

Emine Demir* * Gazi University, Faculty of Science & Arts, Department of Biology, 06500 Teknikokullar Ankara, TURKEY. E-mail: [email protected]. [Demir, E. 2006. Contributions to the knowledge of Turkish Auchenorrhyncha with twelve new records (Homoptera: Cicadellidae). Munis Entomology & Zoology 1 (2): 215-236] ABSTRACT: In this study, 5225 samples of Cicadellidae specimen collected from Antalya region in south-west Turkey during field studies in the years 1999 and 2001-2003 are examined. 111 species are found to be distributed in this region. Distribution of these species in Turkey and their host plants are given along with their locality records. 79 of these examined species are new records for Antalya and Macropsis albae Wagner, 1950, Iassus pseudocerris Abdul-Nour, 1998, Anoscopus assimilis (Signoret, 1879), Ribautiana cruciata (Ribaut, 1931), Eupteryx insulana (Ribaut, 1948), Euscelis alsioides Remane, 1967, Fieberiella chioscola Meyer-Arndt, 1988, Paralimnellus cingulatus (Dlabola, 1960), Paralimnus phragmitis (Boheman, 1847), Laminacutus libanensis Abdul-Nour, 2001, Platymetopius enatus Dlabola, 1974 and Psammotettix nodosus (Ribaut, 1925) are new records for Turkey’s fauna. Key words: Homoptera, Auchenorrhyncha, Cicadellidae, fauna, new record, Turkey.

Turkey’s first faunistic records of Cicadellidae family are given by Fahringer (1922), Haupt (1930), Kerville (1939), Zachvatkin (1937, 1946), Dlabola (1957, 1963, 1971a, 1971b, 1974, 1979, 1981, 1985), Linnavuori (1965), Dworakowska (1969, 1970, 1971, 1972, 1976, 1982) Kalkandelen (1972, 1974, 1980, 1985) and Kartal (1981, 1983). Lodos and Kalkandelen (1981-1988) compiled the faunistic records of their studies up to 1988 and stated that 423 Cicadellidae species were distributed in Turkey, in the checklist they prepared. Later, the studies of Abdul-Nour (1988), Meyer-Arndt (1988), Kartal (1988), Kartal and Zeybekoğlu (1991, 1992, 1994a, 1994b, 1997), Zeybekoğlu (1996, 1998), Zeybekoğlu and Bulut (2000), Kartal, Zeybekoğlu and Dursun (2001), Başpınar and Uygun (1991a, 1991b, 1991c, 1992a, 1992b), Güçlü and Özbek (1994, 1995), Demir (2004a, 2004b, 2005a, 2005b) added 38 records of species belonging to Cicadellidae. According to the literature, 461 species of Cicadellidae were known to be distributed in Turkey.

The province of Antalya, which is selected as the study area, is located

on the Mediterrenean coastline in southwest Turkey, between 29 15 -

32 36 eastern longtitudes and 36 06 - 37 27 nothern lattitudes. The elevation ranges between 0 and 3070 m. This province is important because of its biological diversity with four national parks and three nature protection areas; namely, Olympos Beydağları Coastal National Park, Termessos Güllük Mountain National Park, Köprülü Kanyon National Park, Altınbeşik National Park, Çığlıkara Nature Protection Area, Dibek Nature Protection Area and Alacadağ Nature Protection Area.


28 Cicadellidae species were found to be distributed in the study area, as a result of compilation of the records in the studies of the authors mentioned above. Climate of the Study Area: The region is under the influence of Mediterrenean climate that is an outer tropical climate, in which photoperiodism is daily and seasonal, precipitation occurs mostly in cold or relatively colder seasons, summer is the dry period and summer drought coincides with maximum summer temperature. All the stations except Gazipaşa (796.3 mm), which is the easternmost station, receive precipitation over 1000mm, according to Akman (1995). On the coastline, 60-65% of precipitation is received in winter and 0.5-2% in summer. Vegetation of the Study Area: There are communities dominated by Pinus brutia forest, garrigue and maquis along the coastline at approximately up to 1000 m above sea level, although they are damaged by humans in many places. Pinus brutia can sometimes be seen above 1000 meters and on the coast it is mixed with Pinus pinea. Ceratonia siliqua, Pistacia lentiscus, Euphorbia dendroides, Myrthus communis, Clematis cirrhosa, Rubia tinctoria, Teucrium creticum and Capparis spinosa are examples of common characteristic species of the coastline. Examples of common characteristic species of the zone just above the coastline are Quercus coccifera, Pistacia terebinthus ssp. palaestina, Crataegus aronia ssp. aronia, Fontanesia philliraeoides, Rhamnus oleoides ssp. graecus, Arbutus andrachne, Melica eligulata, Eryngium falcatum and Rubia tenuifolia. On the eastern half of the region at elevations 1500m and higher, there are forests comprising of Cedrus libani, Abies cilicica, Pinus nigra ssp. pallasiana, Quercus infectoria ssp. boissieri, Quercus libani, Juniperus excelsa, and Junuperus drupacea. The western parts of the region is covered with Cedrus libani forests at elevations 1500m and higher. Although cedars here show floristic structure, blackpine forests are extremely poor. Lonicera nummulariifolia ssp. glandulifera, Digitalis cariensis and Acer hyrcanum ssp. sphaerocaryum are examples of characteristic species in this part, Akman (1995).

MATERIALS AND METHODS

Field studies in 143 different localities were executed in the months

March-October in the years 1997-1999 and 2001-2003. In this study, 5225 adult Cicadellidae samples were collected and analyzed. Samples were collected by sweeping the plants with a wooden shaft net and the host plants of samples collected from only one species of plant are determined. Homopterans, found in the net after sweeping were vacuumed with an aspirator. The samples in the aspirator were then killed in ethyl acetate in a jar and put in standard envelopes and prepared according to the standard methods to produce museum material. The studied samples are preserved in the collection of the author.


RESULTS

It is found that there are 111 species of Cicadellidae family in the study area. Along with their distribution in Turkey and the plants found to be their host, the examined materials of these species are given as a list, below. 79 of these examined species are new records for Antalya and 12 are new records for Turkey (see Table 1).

Family: Cicadellidae Subfamily: Ulopinae

Utecha trivia (Germar, 1821) Material: Gazipaşa, Çalıpınar, 25 m, 12.05.2001 1♀; Güllük Dağı, Araştırma Ormanı, 750 m, 21.05.2002 2♂1♀. It has been collected from weeds in Pinus brutia forest. Distribution in Turkey: Ağrı, Ankara, Bilecik, Çanakkale, Çorum, Denizli, Edirne, Elazığ, Gümüşhane, İzmir, Kastamonu, Malatya, Manisa, Mardin, Muğla, Nevşehir, Rize, Siirt, Sivas, Urfa (Dlabola, 1957, 1981; Lodos & Kalkandelen, 1981; Demir, 2006). Remarks: New for Antalya.

Subfamily: Megophthalminae Megophthalmus scabripennis Edwards, 1915

Material: Akseki, Geriş, 700 m, 17.04.2001 1♂; Manavgat, Demirciler, 105 m, 22.04.2002 2♂1♀. It has been collected from weeds in Pinus brutia forest. Distribution in Turkey: Adana, Ankara, Bursa, İzmir, Manisa, Mersin, Muğla (Dlabola, 1957, 1981; Lodos & Kalkandelen, 1981; Başpınar & Uygun, 1991a). Remarks: New for Antalya.

Subfamily: Macropsinae Macropsis albae Wagner, 1950

Material: Elmalı, Çobanisa, 1240 m, 27.07.1999 6♀; Akçay, 1210 m, 13.07.2002 1♀; Gümüşyaka, 1220 m, 13.07.2002 4♀; Korkuteli, Koru Dağı, Kartal Yaylası, 1600 m, 23.06.2002 1♂2♀; Güllük Dağı, Araştırma Ormanı, 550 m, 19.07.2002 1♀. On Salix. Distribution in Turkey: There is no record from Turkey. Remarks: New for Turkey.

Macropsis graminea (Fabricius, 1798) Material: Elmalı, Akçay, 1100 m, 28.07.1999 1♀. On Populus. Distribution in Turkey: Adıyaman, Ağrı, Ankara, Bitlis, Çanakkale, Diyarbakır, Edirne, Elazığ, Eskişehir, Gümüşhane, Kahramanmaraş, Kırklareli, Konya, Kütahya, Manisa, Tokat, Trabzon, Tunceli, Van (Dlabola, 1957, 1981; Lodos & Kalkandelen, 1981; Demir, 2006). Remarks: New for Antalya.

Subfamily: Agalliinae Anaceratagallia laevis (Ribaut, 1935)

Material: Gündoğmuş, Serinyaka, 700 m, 25.07.1999 4♂; Manavgat, Bucakşeyhler, 18-28.06.1997 1♂4♀; Çayyazı, 20.07.1999 2♂2♀; Manavgat, 10 m, 20.07.1999 1♂1♀; Demirciler, 65 m, 21.07.1999 2♀; Karaöz, 20 m, 22.07.1999 1♀; Bucakşeyhler, 30 m, 23.07.1999 2♀; Bucakşeyhler, 24.07.1999 1♂1♀; Bucakşeyhler, 13.08.1999 5♂10♀; Demirciler, 10 m, 17.08.1999 2♂; Elmalı, 1080 m, 28.07.1999 2♂1♀; Manavgat, 10.08.1999 1♀; Alanya, Obaköy, 25 m, 12.08.1999 1♀; Manavgat, Belenobası, 130 m, 15.04.2001 1♂; Manavgat-Çeltikçi, 14.04.2001 1♀; Alanya, Okurcalar, 12.05.2001 3♂4♀; Gazipaşa, Çalıpınar, 25 m, 12.05.2001 1♂1♀; Manavgat, Demirciler, 105 m, 11.05.2001 1♀; Demirciler, 40 m, 11.05.2001 1♀; Alanya, Keşefli, 12.05.2001 1♂; Manavgat, Beşkonak, 145 m, 07.06.2001 4♂; Demirciler, 65 m, 05.06.2001 1♂; Kemer, 06.06.2001 2♀; Manavgat, Bucakşeyhler, 45 m, 19.07.2001 2♀; Alanya, Kargıcak, 21.07.2001 1♀; Gündoğmuş, Çiçekoluk, 960 m, 25.07.2001 1♂1♀; Manavgat, Bucakşeyhler, 70 m, 23.04.2002 3♂; Demirciler, 20 m, 18.05.2002 1♀; Güllük Dağı, Araştırma Ormanı, 750 m, 21.05.2002 1♂; Manavgat, Demirciler, 40 m, 20.06.2002 1♂2♀; Demirciler, 65 m, 15.07.2002 3♂2♀; Alanya, Kargıcak, 22.08.2002 1♀; Kemer, Ovacık, 1150 m, 20.08.2002 1♂; Finike, Yeşilyurt,


18.04.2001 1♂3♀; Kemer, Çamyuva, 18.04.2001 2♂1♀; Gazipaşa, Macarköy, 24.04.2002 3♀; Kale, Köşkerler, 240 m, 24.06.2002 2♂; Kumluca, Kavakköy, 06.09.2002 2♂3♀; Finike, Turunçova, 06.09.2002 1♂1♀; Gündoğmuş, Güneycik, 240 m, 10.09.2002 3♀; Manavgat, Taşağıl, Kilimli, 40 m, 08.09.2002 1♂; Serik, Aspendos, 20.08.2001 2♂. It has been collected from weeds in Pinus brutia forest. Distribution in Turkey: Adana, Ağrı, Ankara, Bilecik, Bitlis, Çanakkale, Diyarbakır, Elazığ, Erzurum, Hatay, İzmir, İstanbul, Kırklareli, Konya, Malatya, Manisa, Mardin, Mersin, Muğla, Nevşehir, Niğde, Ordu, Samsun, Siirt, Urfa, Yozgat, Zonguldak (Zachvatkin 1946; Dlabola, 1957, 1981; Lodos & Kalkandelen, 1981; Başpınar & Uygun, 1991a). Remarks: New for Antalya.

Austroagallia sinuata (Mulsant et Rey, 1855) Material: Gündoğmuş, Serinyaka, 700 m, 25.07.1999 2♂3♀; Manavgat, Bucakşeyhler, 15-26.08.1997 2♂2♀; Alanya, Obaköy, 25 m, 12.08.1999 1♀; Korkuteli, Kargalık, 1250 m, 27.07.1999 1♀; Manavgat, Karaöz, 90 m, 22.07.1999 1♂; Akseki, Güçlüköy, 610 m, 18.08.1999 1♀; Manavgat, Bucakşeyhler, 13.08.1999 1♂; Hacıobası, 26.08.2003 1♂¸ Kemer, Böğürtlenözü, 21.08.2001 1♂1♀. It has been collected from weeds in Pinus brutia forest and Sesamum indicum. Distribution in Turkey: Adana, Ankara, Diyarbakır, Erzincan, Hakkari, İzmir, Iğdır, Kırşehir, Mardin, Mersin, Muğla, Nevşehir, Samsun, Şırnak (Dlabola, 1957, 1981; Lodos & Kalkandelen, 1981; Başpınar & Uygun, 1991a). Remarks: New for Antalya.

Dryodurgades anatolicus Dlabola, 1957 Material: Termessos, Güllük Dağı, 900-1000 m, 22.07.2001 3♂; Gündoğmuş, Güneycik, 210 m, 21.04.2002 1♂3♀; Elmalı, Çığlıkara, 1600 m, 28.08.2003 4♀. It has been collected from Juniperus and shrubs. Distribution in Turkey: Adana, Ankara (Dlabola, 1957; Demir, 2004b, 2006). Remarks: New for Antalya.

Subfamily: Idiocerinae Rhytidodus boluicus Dlabola, 1970

Material: Manavgat, Bucakşeyhler, 24.07.1999 5♂6♀; Elmalı, Gümüşyaka, 1220 m, 13.07.2002 1♀; Serik, Aspendos, Köprüçayı, 16.10.2001 2♀; Manavgat, Boztepe, Alara, 15.10.2001 1♂; Serik, Aspendos, 20.08.2001 1♂1♀; Finike, Turunçova, 06.09.2002 2♀. On Populus and Salix. Distribution in Turkey: Adana, Ankara, Çankırı, Diyarbakır, Erzurum, İzmir, Kayseri, Manisa (Lodos & Kalkandelen, 1982a; Güçlü & Özbek, 1994; Demir, 2006). Remarks: New for Antalya and endemic to Turkey.

Tremulicerus mesopyrrhus (Kirschbaum, 1868 ) Material: Manavgat, Bucakşeyhler, 24.07.1999 2♂3♀; Antalya: Elmalı, Çobanisa, 1240 m, 27.07.1999 1♂1♀; Manavgat, Hacıobası, 26.08.2003 5♂9♀; Serik, Aspendos, 20.08.2001 1♂. On Populus. Distribution in Turkey: Adana, Ankara, Bitlis, Çankırı, Erzurum, Gümüşhane, İzmir, Konya, Manisa, Van (Lodos & Kalkandelen, 1982a; Güçlü & Özbek, 1994; Demir, 2006). Remarks: New for Antalya.

Sulamicerus ancorarius (Dlabola, 1964) Material: Manavgat, Demirciler, 10 m, 17.08.1999 9♂4♀; Demirciler, 40 m, 24.07.2001 2♂. On Pistacia. Distribution in Turkey: Diyarbakır, Konya, Siirt (Lodos & Kalkandelen, 1982a). Remarks: New for Antalya.

Subfamily: Iassinae Batracomorphus signatus Lindberg, 1923

Material: Alanya, Okurcalar, 12.05.2001 1♀. It has been collected from weeds in fields. Distribution in Turkey: Adana, Iğdır, Mardin (Lodos & Kalkandelen, 1982b). Remarks: New for Antalya.

Iassus pseudocerris Abdul-Nour, 1998 Material: Gündoğmuş, Serinyaka, 625 m, 22.06.2002 1♂. On Quercus. Distribution in Turkey: There is no record from Turkey. Remarks: New for Turkey.

Subfamily: Dorycephalinae Paradorydium mustafai Demir, 2005

Distribution in Turkey: Antalya (Demir, 2005b).


Paradorydium paradoxum (Herrich-Schaeffer, 1837) Distribution in Turkey: Ankara, Antalya, Diyarbakır (Lodos & Kalkandelen, 1982b; Demir, 2004b, 2005b, 2006).

Paradorydium occidentale Lindberg, 1954 Distribution in Turkey: Antalya (Demir, 2005b).

Eupelix cuspidata (Fabricius, 1775) Material: Akseki, Geriş, 700 m, 17.04.2001 1♂; Manavgat, Yayaalan, 540 m, 14.05.2001 1♂; Tilkiler, 470 m, 14.05.2001 1♂; Gazipaşa, Çalıpınar, 25 m, 12.05.2001 1♂; Manavgat, Bucakşeyhler, Seleukeia, 250 m, 23.04.2002 8♂1♀; Demirciler, 105, 22.04.2002 2♂; Demirciler, 105 m, 18.05.2002 1♂; Gündoğmuş, Çiçekoluk, 960 m, 19.05.2002 2♂; Termessos, Güllük Dağı, 750 m, 21.05.2002 2♂; Alanya, Okurcalar, 20.05.2002 1♂; Korkuteli, Koru dağı, Kartal Yaylası, 1600 m, 23.06.2002 2♂. It has been collected from weeds in Pinus brutia forest. Distribution in Turkey: Adana, Adıyaman, Afyon, Ankara, Artvin, Bitlis, Çanakkale, Diyarbakır, Erzurum, Kahramanmaraş, Konya, Malatya, Mardin, Mersin, Nevşehir, Niğde, Urfa, Van (Dlabola, 1957, 1981; Lodos & Kalkandelen, 1982b; G Başpınar & Uygun, 1991a; Güçlü & Özbek, 1994; Demir, 2006). Remarks: New for Antalya.

Subfamily: Hecalinae Hecalus glaucescens (Fieber, 1866)

Material: Manavgat, Bucakşeyhler, 13.08.1999 1♀; Manavgat, 10.08.1999 1♀; Elmalı, 1080 m, 28.07.1999 3♂; Manavgat, 18.09.1998 1♂; Bucakşeyhler, 24.07.1999 1♂; Belenobası, 130 m, 15.04.2001 2♀; Akseki, Güçlüköy, 680 m, 17.04.2001 1♂; Manavgat, Taşağıl, Sağırin, 40 m, 03.06.2001 1♀; Beşkonak, 145 m, 07.06.2001 2♂; Düzlerçamı, 280 m, 22.07.2001 1♀; Alanya, Kargıcak, 21.07.2001 1♂; Gündoğmuş, Güneycik, 240 m, 21.04.2002 1♀; Manavgat, Bucakşeyhler, 70 m, 23.04.2002 1♀; Demirciler, 40 m, 20.06.2002 3♀; Güllük Dağı, Araştırma Ormanı, 550 m, 19.07.2002 1♀; Manavgat, Hacıobası, 26.08.2003 1♂; Elmalı, Tekkeköy, 1100 m, 28.08.2003 1♂. It has been collected from weeds in Pinus brutia forest. Distribution in Turkey: Adana, Adıyaman, Ankara, Hatay, Malatya, Manisa, Mersin, Muğla, Siirt (Lodos & Kalkandelen, 1982b; Başpınar & Uygun, 1991b). Remarks: New for Antalya.

Subfamily: Aphrodinae Anoscopus albifrons (Linnaeus, 1758)

Material: Manavgat, Bucakşeyhler, 70 m, 23.04.2002 1♂. It has been collected from weeds in Pinus brutia forest. Distribution in Turkey: İzmir (Lodos & Kalkandelen, 1982b). Remarks: New for Antalya.

Anoscopus assimilis (Signoret, 1879) Material: Termessos, Güllük Dağı Araştırma Ormanı, 750 m, 21.05.2002 2♂1♀. It has been collected from weeds in Pinus brutia-Quercus forest. Distribution in Turkey: There is no record from Turkey. Remarks: New for Turkey.

Aphrodes makarovi Zachvatkin, 1948 Material: Manavgat, Demirciler, Külcüler, 170 m, 15.04.2001 1♀; Yukarıışıklar-Tilkiler, 460 m, 14.05.2001 2♂1♀; Demirciler, 105 m, 11.05.2001 1♂; Yaylaalan, 540 m, 14.05.2001 1♂; Tilkiler, 220 m, 14.05.2001 1♀; Tilkiler, 470 m, 14.05.2001 1♂; Alanya, Okurcalar, 12.05.2001 1♂; Manavgat, Bucakşeyhler, 70 m, 23.04.2002 3♂2♀; Demirciler, 20 m, 18.05.2002 1♂; Termessos, Güllük Dağı, 750 m, 21.05.2002 4♂1♀; Alanya, Okurcalar, 20.05.2002 1♀; Gündoğmuş, Güneycik, 240 m, 19.05.2002 1♂. It has been collected from weeds in Pinus brutia forest. Distribution in Turkey: Adana, Ankara (Başpınar & Uygun, 1991b; Demir, 2006). Remarks: New for Antalya.

Subfamily: Typhlocybinae Alebra albostriella (Fallén, 1826)

Material: Manavgat, Bucakşeyhler, 24.07.1999 1♀; Demirciler, 40 m, 17.07.2002 1♀. It has been collected from Quercus and maquis. Distribution in Turkey: Adana, Ankara, Artvin, Bitlis, Bolu, Çorum, Hatay, İzmir, Kahramanmaraş, Kayseri, Kocaeli, Kütahya, Ordu


(Dlabola, 1957, 1981; Linnavuori, 1965; Lodos & Kalkandelen, 1983b; Demir, 2006). Remarks: New for Antalya.

Micantulina stigmatipennis (Mulsant et Rey, 1855) Material: Kaş, Gömbe, Uçarsu, 1700 m, 28.07.1999 2♀; Beldibi, 1400 m, 29.07.1999 5♂1♀; Alanya, Okurcalar, 12.05.2001 1♂1♀; Keşefli, 12.05.2001 1♀; Manavgat, Demirciler, 40 m, 11.05.2001 2♀; Elmalı, Gölova-Bozhöyük, 1200 m, 06.06.2001 2♂19♀; Manavgat, Taşağıl, Kilimli, 40 m 03.06.2001 1♂1♀; Taşağıl, Sağırin, 40 m, 03.06.2001 1♂; Korkuteli, Beğiş-Karabayır, 1310 m, 06.06.2001 1♂; Korkuteli-Kargalık, 1060 m, 06.06.2001 7♂12♀; Finike, Gökcük, 310 m, 06.06.2001 4♂7♀; Manavgat, Ilıca, 16.10.2001 1♂3♀; Demirciler, Külcüler, 150 m, 14.10.2001 7♀; Örenşehir, 15.10.2001 1♂2♀; Bucakşeyhler, 70 m, 23.04.2002 2♀; Kaş, Beldibi, 1450 m, 13.07.2002 1♀; Hisarçandır, 770 m, 20.08.2002 1♂; Elmalı, Tekkeköy, 1100 m, 28.08.2003 2♂5♀. On Verbascum. Distribution in Turkey: Ankara, Burdur, Isparta, İstanbul, İzmir, Manisa, Van (Dlabola, 1957, 1981; Lodos & Kalkandelen, 1983b; Demir, 2006). Remarks: New for Antalya.

Kybos virgator (Ribaut, 1933) Material: Elmalı, Çobanisa, 1240 m, 27.07.1999 7♀; Akçay, 1100 m, 28.07.1999 3♂15♀; Akçay, 1100 m, 29.07.1999 5♀; Manavgat, Ilıca, 16.10.2001 1♀; Korkuteli, Koru Dağı, Kartal Yaylası, 1600 m, 23.06.2002 1♂7♀; Elmalı, Gümüşyaka, 1220 m, 13.07.2002 1♀; Tekkeköy, 1100 m, 28.08.2003 3♂20♀. On Salix. Distribution in Turkey: Ankara, Bilecik, İzmir, Kırşehir, Konya, Van (Lodos & Kalkandelen, 1983c; 2004a). Remarks: New for Antalya.

Empoasca decipiens Paoli, 1930 Material: Manavgat, Bucakşeyhler, 13.08.1999 1♀; Akseki, Güçlüköy, 610 m, 18.08.1999 2♂5♀; Elmalı, 1080 m, 28.07.1999 1♂2♀; Manavgat, 10.08.1999 1♀; Bucakşeyhler, 30 m, 23.07.1999 2♀; Bucakşeyhler, 24.07.1999 2♂; Bucakşeyhler, 15-26.08.1997 3♀; Bucakşeyhler, 18-28.06.1997 2♂; Demirciler, 40 m, 14.04.2001 1♀; Demirciler, 65 m, 14.04.2001 1♀; Yukarıışıklar-Tilkiler, 460 m, 14.05.2001 1♀; Beşkomak, 145 m, 07.06.2001 1♂2♀; Demirciler, 40 m, 05.06.2001 1♂; Demirciler, 40 m, 17.07.2002 1♂1♀; Elmalı, Gümüşyaka, 1220 m, 13.07.2002 2♀; Kemer, Çamyuva, 18.04.2001 7♂11♀; Kumluca, Kavakköy, 18.04.2001 7♂9♀; Finike, Yeşilyurt, 18.04.2001 17♂39♀; Kaş, İslamlar, 220 m, 15.05.2001 38♂56♀; Kale, Köşkerler, 240 m, 15.05.2001 51♂47♀; Alanya, Konaklı, 25 m, 24.04.2002 16♂9♀; Gazipaşa, Macarköy, 24.04.2002 23♂16♀; Kemer, Karabucak, 85 m, 25.05.2002 11♂19♀; Finike, Erenkavak, 65 m, 24.06.2002 29♂12♀; Kumluca, Kavakdibi, 130 m, 24.06.2002 8♂9♀; Serik, Camili, 26.07.2001 2♂5♀; Manavgat, Karaöz, 70 m, 18.08.2001 4♂6♀; Serik, Aspendos, 20.08.2001 8♂14♀; Kemer, Çamyuva, 21.08.2001 7♂2♀; Manavgat, Ilıca, 07.09.2001 6♂5♀; Alanya, Kargıcak, 09.09.2001 3♂10♀; Finike, Turunçova, 06.09.2002 2♂9♀. It has been collected from Salix, Sesamum indicum, Ulmus and weeds in Pinus brutia forest. Distribution in Turkey: Adana, Mersin (Dlabola, 1957; Başpınar & Uygun, 1991b). Remarks: New for Antalya.

Asymmetrasca decedens (Paoli, 1932) Material: Manavgat, 10.08.1999 2♀; Bucakşeyhler, 30 m, 23.07.1999 2♂1♀; Bucakşeyhler, 15-26.08.1997 5♀; Bucakşeyhler, 18-28.06.1997 1♂; Bucakşeyhler, 45 m, 19.07.2001 1♂; Kumluca, Kavakköy, 18.04.2001 11♂12♀; Finike, Yeşilyurt, 18.04.2001 7♂15♀; Kaş, İslamlar, 220 m, 15.05.2001 32♂53♀; Kale, Köşkerler, 240 m, 15.05.2001 32♂46♀; Alanya, Konaklı, 25 m, 24.04.2002 7♂11♀; Gazipaşa, Macarköy, 24.04.2002 7♂13♀; Kemer, Karabucak, 85 m, 25.05.2002 3♂13♀; Kumluca, Kavakdibi, 130 m, 24.06.2002 6♂11♀; Serik, Camili, 26.07.2001 1♂7♀; Manavgat, Karaöz, 70 m, 18.08.2001 1♂6♀; Serik, Aspendos, 20.08.2001 2♂3♀; Kemer, Çamyuva, 21.08.2001 1♂8♀; Alanya, Kargıcak, 09.09.2001 1♂13♀; Finike, Turunçova, 06.09.2002 2♂5♀. It has been collected from weeds in Pinus brutia forest. Distribution in Turkey: Adana, Konya, Mersin, (Dlabola, 1957 Lodos & Kalkandelen, 1983c; Başpınar & Uygun, 1991b). Remarks: New for Antalya.

Edwardsiana tshinari Zachvatkin, 1947 Material: Kemer, Ovacık, 1150 m, 20.08.2002 1♂1♀. On Platanus orientalis. Distribution in Turkey: İstanbul, Kocaeli (Dlabola, 1971). Remarks: New for Antalya.

Ribautiana cruciata (Ribaut, 1931) Material: Manavgat, Demirciler, 40 m, 17.07.2002 1♂3♀. On Ulmus. Distribution in Turkey: There is no record from Turkey. Remarks: New for Turkey.


Eupteryx cypria (Ribaut, 1948) Material: Kaş, Kalkan, Yeşilköy, 29.07.1999 5♂8♀; Alanya, Obaköy, 25 m, 12.08.1999 1♀; Manavgat, Karaöz, 20 m, 22.07.1999 4♀; Demirciler, 20 m, 18.05.2002 7♂9♀; Finike, Erenkavak, 65 m, 24.06.2002 2♂1♀; Kale, Eseler, 55 m, 24.06.2002 1♂1♀. It has been collected from weeds in fields and Pinus brutia forest. Distribution in Turkey: Adana (Dlabola, 1957). Remarks: New for Antalya.

Eupteryx fahringeri Melichar, 1911 Material: Kaş, Gömbe, Uçarsu, 1700 m, 28.07.1999 4♂8♀. It has been collected from Verbascum in alpine zone. Distribution in Turkey: Bursa, İzmir (Lodos & Kalkandelen, 1984c). Remarks: New for Antalya and endemic to Turkey.

Eupteryx gyaurdagica Dlabola, 1957 Material: Korkuteli, Koru Dağı, Kartal Yaylası, 1600 m, 23.06.2002 1♂1♀. It has been collected from weeds in fields. Distribution in Turkey: Adana, Afyon, Ankara, Bitlis, Bolu, Isparta, İzmir, Manisa, Nevşehir, Sinop (Dlabola, 1957; Lodos & Kalkandelen, 1984b; Demir, 2006). Remarks: New for Antalya.

Eupteryx insulana (Ribaut, 1948) Material: Manavgat, Demirciler, 20 m, 18.05.2002 1♀; Demirciler, 40 m, 20.06.2002 1♂. It has been collected from weeds in fields. Distribution in Turkey: There is no record from Turkey. Remarks: New for Turkey.

Zyginella pulchra Löw, 1885 Material: Kemer, Ovacık, 1150 m, 20.08.2002 59♂34♀; Kemer, Çamyuva, 21.08.2001 9♂24♀. On Platanus orientalis. Distribution in Turkey: Adana, Aksaray, Diyarbakır, Hatay, İzmir, Konya, Samsun (Lodos & Kalkandelen, 1984a). Remarks: New for Antalya.

Zyginidia sohrab Zachvatkin, 1947 Material: Alanya, Okurcalar, 12.05.2001 1♂; Kemer, Çamyuva, 18.04.2001 4♂6♀; Finike, Yeşilyurt, 18.04.2001 1♂5♀; Alanya, Konaklı, 25 m, 24.04.2002 3♂; Kale, Eseler, 55 m, 24.06.2002 7♂2♀; Serik, Camili, 20.08.2001 6♀; Kumluca, Kavakköy, 06.09.2002 4♂7♀; Gazipaşa, Macarköy, 24.04.2002 4♂2♀; Aksu, Topallar, 26.07.2001 3♂4♀; Finike, Erenkavak, 130 m, 24.06.2002 5♂1♀; Kemer, Böğürtlenözü, 18.04.2001 7♂4♀; Kemer, Karabucak, 25.05.2002 3♂10♀. It has been collected from weeds in fields and Pinus brutia forest. Distribution in Turkey: Adıyaman, Bingöl, Bitlis, Diyarbakır, Elazığ, Gaziantep, Hakkari, Kars, Kocaeli, Iğdır, İstanbul, Malatya, Siirt, Urfa, Van (Dlabola, 1971; Lodos & Kalkandelen, 1984d; Kalkandelen, 1985). Remarks: New for Antalya.

Zygina discolor Horváth, 1897 Material: Kemer, Çamyuva, 25.05.2002 5♀. On Quercus. Distribution in Turkey: Ankara, İzmir, Malatya (Dlabola, 1981; Lodos & Kalkandelen, 1984d). Remarks: New for Antalya.

Arboridia adanae (Dlabola, 1957) Material: Manavgat, Demirciler, 40 m, 17.07.2002 2♀; Kumluca, Kavakdibi, 130 m, 24.06.2002 1♂3♀; Kemer, Karabucak, 85 m, 25.05.2002 1♂2♀. It has been collected from Ulmus and maquis vegetation. Distribution in Turkey: Adana, Adıyaman, Ankara, Çankırı, Denizli, Diyarbakır, Eskişehir, Gaziantep, Kayseri, Kırıkkale, Kırşehir, Konya, Mardin, Nevşehir, Niğde, Sivas, Urfa, Yozgat, (Dlabola, 1957; Lodos & Kalkandelen, 1984d). Remarks: New for Antalya.

Frutioidia sanguinosa (Rey, 1891) Material: Kemer, Çamyuva, 21.08.2001 2♂4♀. On Amygdalus. Distribution in Turkey: Adana (Dlabola, 1957). Remarks: New for Antalya.

Tamaricella cypria (Ribaut, 1948) Material: Manavgat, Taşağıl, 60 m, 27.08.2003 2♂; Kemer, Böğürtlenözü, 21.08.2001 1♂1♀. On Tamarix. Distribution in Turkey: Ankara, Kırıkkale, Malatya (Dlabola, 1981; Lodos & Kalkandelen, 1984c). Remarks: New for Antalya.


Subfamily: Deltocephalinae Grypotes staurus Ivanoff, 1885

Material: Manavgat, Bucakşeyhler, 18-28.06.1997 1♂1♀; Bucakşeyhler, 15-26.08.1997 2♂1♀; Demirciler, 10 m, 17.08.1999 1♀; Bucakşeyhler, 13.08.1999 1♀; Alanya, Keşefli, 21.07.2001 2♀; Manavgat, Demirciler, 105 m, 18.05.2002 1♂; Demirciler, 105 m, 21.06.2002 4♂1♀; Demirciler, 40 m, 20.06.2002 2♂2♀; Bucakşeyhler, 22.07.2003 1♂; Kemer, Çamyuva, 18.04.2001 3♂4♀; Kaş, İslamlar, 220 m, 15.05. 2001 6♂2♀; Alanya, Kestel, 24.04.2002 8♂17♀; Serik, Sarıabalı, 20.08.2001 1♂3♀; Manavgat, Taşağıl, Kilimli, 40 m, 08.09.2002 1♂5♀; Manavgat, Bucakşeyhler, 30 m, 08.09.2001 2♀; Kemer, Böğürtlenözü, 21.08.2001 5♂1♀; Kemer, Karabucak, 85 m, 25.05.2002 3♂1♀. On Pinus brutia. Distribution in Turkey: Adana, Hatay, İzmir, Konya, Muğla, Samsun, Sinop (Linnavuori, 1965; Lodos & Kalkandelen, 1985a; Zeybekoğlu, 1994, 1998). Remarks: New for Antalya.

Goniagnathus brevis (Herrich-Schäffer, 1835) Distribution in Turkey: Adana, Amasya, Ankara, Antalya, Burdur, Diyarbakır, Edirne, Manisa, Mardin, Niğde, Samsun, Sinop, Tokat (Dlabola, 1957; Kalkandelen, 1974; Lodos & Kalkandelen, 1985a; Başpınar & Uygun, 1991b; Kartal & Zeybekoğlu, 1991, Zeybekoğlu, 1998; Demir, 2004a, 2004b, 2006).

Goniagnathus guttulinervis (Kirschbaum, 1868) Distribution in Turkey: Antalya (Demir, 2004a).

Opsius cypriacus Lindberg, 1958 Material: Manavgat, Taşağıl, Sağırin, 40 m, 03.06.2001 1♂1♀; Hacıobası, 14.10.2001 13♂17♀; Örenşehir, 15.10.2001 7♂6♀; Ilıca, 16.10.2001 1♂; Hacıobası, 22.06.2002 1♀; Hacıobası, 26.08.2003 2♂5♀; Taşağıl, 60 m, 27.08.2003 5♂10♀. On Tamarix. Distribution in Turkey: Adana, Amasya, Ankara, Artvin, Aydın, Diyarbakır, Edirne, Erzincan, Erzurum, İzmir, Kırıkkale, Kütahya, Malatya, Manisa, Nevşehir, Samsun, Siirt, Van, Yozgat (Linnavuori, 1965; Lodos & Kalkandelen, 1985a). Remarks: New for Antalya.

Opsius smaragdinus Emeljanov, 1964 Material: Kemer, Çamyuva, 21.08.2001 1♂1♀. On Tamarix. Distribution in Turkey: Nevşehir (Lodos & Kalkandelen, 1985a). Remarks: New for Antalya.

Opsius stactogalus Fieber, 1866 Material: Manavgat, 10.08.1999 2♂2♀; Serik, Aspendos, Köprüçayı, 16.10.2001 1♂2♀; Aspendos, 18.07.2002 31♂36♀; Kale, Beymelek, 20.08.2002 3♂4♀. On Tamarix. Distribution in Turkey: Ankara, İzmir, Kırıkkale, Malatya Nevşehir, Samsun, Sinop (Kalkandelen, 1974; Lodos & Kalkandelen, 1985a; Zeybekoğlu, 1998). Remarks: New for Antalya.

Orosius albicinctus Distant, 1918 Material: Manavgat, Bucakşeyhler, 45 m, 19.07.2001 4♂6♀; Demirciler, 40 m, 24.07.2001 2♀; Örenşehir, 15.10.2001 1♂; Hacıobası, 26.08.2003 2♂; Kumluca, Kavakköy, 06.09.2002 2♀. It has been collected from Sesamum indicum and weeds in fields. Distribution in Turkey: Adana, Mersin (Dlabola, 1981; Başpınar & Uygun, 1991c). Remarks: New for Antalya.

Neoaliturus fenestratus (Herrich-Schäffer, 1834) Material: Gündoğmuş, Serinyaka, 700 m, 25.07.1999 2♂4♀; Manavgat, Çayyazı, 20.07.1999 1♂2♀; Beşkonak, 145 m, 07.06.2001 1♂; Gündoğmuş, Çiçekoluk, 960 m, 25.07.2001 1♂2♀; Manavgat, Bucakşeyhler, Seleukeia, 250 m, 23.04.2002 3♂; Demirciler, 105 m, 22.04.2002 2♂; Termessos, Güllük Dağı, Araştırma Ormanı, 750 m, 21.05.2002 2♂2♀; Kumluca, Kavakköy, 18.04.2001 1♂1♀; Serik, Camili, 26.07.2001 1♀. It has been collected from weeds in Pinus brutia forest. Distribution in Turkey: Adana, Ağrı, Amasya, Ankara, Bolu, Çorum, Diyarbakır, Edirne, Erzurum, Giresun, İzmir, Iğdır, Kocaeli, Mardin, Mersin, Nevşehir, Ordu, Samsun, Sinop, Sivas, Tokat, Urfa, Van (Kalkandelen, 1974; Lodos & Kalkandelen, 1985a; Başpınar & Uygun, 1991c; Kartal & Zeybekoğlu, 1991; Zeybekoğlo, 1998; Demir, 2006). Remarks: New for Antalya.

Neoaliturus guttulatus (Kirschbaum, 1868) Material: Kemer, Çamyuva, 18.04.2001 1♂. It has been collected from weeds in Pinus brutia forest. Distribution in Turkey: Adana, Ankara, Bitlis, Diyarbakır, Iğdır, İzmir,


Konya, Nevşehir, Tokat, Urfa (Kalkandelen, 1974; Lodos & Kalkandelen, 1985a; Demir, 2006). Remarks: New for Antalya.

Neoaliturus pulcher (Haupt, 1927) Material: Manavgat, Demirciler, 105 m, 22.04.2002 1♀. It has been collected from weeds in Pinus brutia forest. Distribution in Turkey: Diyarbakır, Muğla (Lodos & Kalkandelen, 1985a). Remarks: New for Antalya.

Circulifer haematoceps (Mulsant et Rey, 1855) Material: Akseki, Güçlüköy, 610 m, 18.08.1999 5♂15♀; Manavgat, Bucakşeyhler, 15-26.08.1997 1♂; Bucakleyhler, 24.07.1999 1♀; Gündoğmuş, Serinyaka, 700 m, 25.07.1999 1♂; Manavgat, Demirciler, 105 m, 22.04.2002 1♂; Demirciler, 20 m, 18.05.2002 1♀; Alanya, Okurcalar, 20.05.2002 1♂; Manavgat, Hacıobası, 26.08.2003 1♂3♀; Kale, Köşkerler, 240 m, 24.06.2002 2♂; Manavgat, Beşkonak, 145 m, 08.09.2002 1♂2♀. It has been collected from weeds in fields and Pinus brutia forest. Distribution in Turkey: Adana, Ağrı, Ankara, Antalya, Bitlis, Diyarbakır, Erzurum, Gaziantep, Hakkari, Hatay, Kahramanmaraş, Kocaeli, Konya, Mardin, Mersin, Muğla, Nevşehir, Siirt, Sivas, Tunceli, Urfa, Van (Kalkandelen, 1974; Başpınar & Uygun, 1991c; Kartal, Zeybekoğlu & Dursun, 2001; Demir, 2004b). Remarks: Known to Antalya.

Nesoclutha erythrocephala (Ferrari, 1882) Material: Antalya: Kaş, Kalkan, Yeşilköy, 29.07.1999 1♂; Manavgat, Bucakşeyhler, 30 m, 23.07.1999 1♀; Bucakşeyhler, 13.08.1999 1♀. It has been collected from weeds in maquis vegetation. Distribution in Turkey: Adana, Amasya, Tokat (Dlabola, 1957; Zeybekoğlu, 1998). Remarks: New for Antalya.

Cicadulina bipunctata (Melichar, 1904) Material: Manavgat, Bucakşeyhler, 24.07.1999 1♂5♀; Manavgat, 18.09.1998 1♂2♀; Kaş, Kalkan, Yeşilköy, 29.07.1999 2♂2♀; Manavgat, Bucakşeyhler, 13.08.1999 20♂11♀; ; Bucakşeyhler, 15-26.08.1997 2♂3♀; Bucakşeyhler, 18-28.06.1997 1♀; Demirciler, 10 m, 17.08.1999 1♂; Alanya, Obaköy, 25 m, 12.08.1999 2♂10♀; Okurcalar, 12.05.2001 1♀; Finike, Turunçova, 50 m, 06.06.2001 1♀; Manavgat, Örenşehir, 15.10.2001 1♀; Çakış, 16.10.2001 1♀; Gündoğmuş, Güneycik, 240 m, 21.04.2002 1♀; Kemer, Ovacık, 1150 m, 20.08.2002 1♀; Kemer, Çamyuva, 18.04.2001 11♂18♀; Kumluca, Kavakköy, 18.04.2001 2♂6♀; Kale, Eseler, 55 m, 15.05.2001 3♂1♀; Serik, Camili, 26.07.2001 21♂13♀; Aksu, Topallar, 26.07.2001 1♂5♀; Alanya, Kestel, 24.04.2002 17♂28♀; Gazipaşa, Macarköy, 24.04.2002 7♂16♀; Finike, Erenkavak, 65 m, 24.06.2002 23♂12♀; Manavgat, Karaöz, 70 m, 18.08.2001 8♂3♀; Kemer, Böğürtlenözü, 21.08.2001 26♂17♀; Manavgat, Ilıca, 07.09.2001 3♂7♀; Finike, Yeşilyurt, 06.09.2002 1♂9♀; Manavgat, Taşağıl, 55 m, 08.09.2002 2♂6♀; Gündoğmuş, Güneycik, 210 m, 10.09.2002 1♂; Kaş, İslamlar, 220 m, 15.05.2001 14♂29♀; Alanya, Konaklı, 25 m, 24.04.2002 5♂2♀; Kemer, Karabucak 85 m, 25.05.2002 11♂2♀; Kumluca, Kavakdibi, 130 m, 24.06.2002 12♂3♀; Serik, Sarıabalı, 20.08.2001 1♂5♀. It has been collected from weeds in fields and Pinus brutia forest. Distribution in Turkey: Adana, İzmir, Mersin (Lodos & Kalkandelen, 1985b; Başpınar & Uygun, 1991c). Remarks: New for Antalya.

Balclutha hebe (Kirkaldy, 1906) Material: Manavgat, Bucakşeyhler, 13.08.1999 4♂9♀; Bucakşeyhler, 24.07.1999 2♂5♀; Alanya, Obaköy, 25 m, 12.08.1999 4♂9♀; Manavgat, Demirciler, 40 m, 11.05.2001 1♀; Gündoğmuş, Güneycik, 210 m, 08.06.2001 1♀; Manavgat, Boztepe, Alara, 15.10.2001 9♂14♀; Çakış, 16.10.2001 1♂10♀; Hacıobası, 14.10.2001 1♀, Gündoğmuş, Güneycik, 210 m, 21.04.2002 1♂; Manavgat, Demirciler, 20 m, 18.05.2002 1♀; Kemer, Ovacık, 1150 m, 20.08.2002 1♂; Gazipaşa, Asarcık, 24.04.2002 7♂2♀; Gündoğmuş, Güneycik, 210 m, 10.09.2002 3♀; Manavgat, Taşağıl, 55 m, 08.09.2002 3♂1♀; Kemer, Böğürtlenözü, 21.08.2001 3♂5♀; Serik, Dikmen, 26.07.2001 4♂. It has been collected from weeds in fields. Distribution in Turkey: Adana, Adıyaman, Ankara, Bitlis, Burdur, Diyarbakır, Elazığ, Gaziantep, Konya, Malatya, Mardin, Mersin, Muğla, Nevşehir, Urfa (Lodos & Kalkandelen, 1985b; Başpınar & Uygun, 1991c). Remarks: New for Antalya.

Balclutha rhenana Wagner, 1939 Material: Kemer, Çamyuva, 18.04.2001 2♂5♀; Kaş, İslamlar, 220 m, 15.05.2001 1♂3♀; Kumluca, Kavakdibi, 130 m, 24.06.2002 1♂1♀; Kemer, Böğürtlenözü, 21.08.2001 1♂3♀; Finike, Turunçova, 06.09.2002 2♂; Yeşilyurt, 18.04.2001 2♂1♀; Kemer, Çamyuva,


25.05.2002 2♂2♀. It has been collected from weeds in fields. Distribution in Turkey: Erzurum, Malatya, Mersin (Lodos & Kalkandelen, 1985b). Remarks: New for Antalya.

Balclutha rosea (Scott, 1896) Material: Manavgat, 10.08.1999 2♂2♀; Manavgat, 18.09.1998 1♀; Karaöz, 90 m, 22.07.1999 1♂; Bucakşeyhler, 15-26.08.1997 3♀; Bucakşeyhler, 18-28.06.1997 2♀; Bucakşeyhler, 24.07.1999 1♂3♀; Kaş, Kalkan, Yeşilköy, 29.07.1999 1♂1♀; Manavgat, Demirciler, 65 m, 14.04.2001 2♀; Demirciler, Külcüler, 170 m, 15.04.2001 1♀; Gündoğmuş, Güneycik, 210 m, 17.04.2001 1♀; Manavgat, Demirciler, 105 m, 11.05.2001 6♀; Demirciler, 40 m, 11.05.2001 1♀; Gazipaşa, Çalıpınar, 25 m, 12.05.2001 45♂30♀; Alanya, Okurcalar, 12.05.2001 4♂10♀; Manavgat, Beşkonak, 145 m, 07.06.2001 3♂1♀; Taşağıl, Kilimli, 60 m, 03.06.2001 2♀; Taşağıl, Sağırin, 40 m, 03.06.2001 1♂1♀; Taşağıl, 55 m, 07.06.2001 2♀; Finike, Turunçova, 50 m, 06.06.2001 2♂3♀; Kemer, 06.06.2001 1♂3♀; Alanya, Kargıcak, 21.07.2001 1♂; Manavgat, Örenşehir, 15.10.2001 1♂1♀; Gazipaşa, Çalıpınar-Kahyalar, 20 m, 20.05.2002 5♂3♀; Alanya, Yeşilköy, 20.05.2002 1♀; Manavgat, demirciler, 40 m, 20.06.2002 1♂. It has been collected from weeds in fields. Distribution in Turkey: Adana, Diyarbakır, Hatay, İzmir, Konya, Mersin, Nevşehir (Linnavuori, 1965; Lodos & Kalkandelen, 1985b). Remarks: New for Antalya.

Macrosteles fieberi (Edwards, 1889) Material: Manavgat, Bucakşeyhler, 18-28.06.1997 1♂1♀; Elmalı, 1080 m, 28.07.1999 1♂. It has been collected from weeds in fields. Distribution in Turkey: Adana, Ağrı, Ankara, Bitlis, Bolu, Çankırı, Hakkari, Konya, Nevşehir, Osmaniye, Van (Dlabola, 1957, 1981; Kalkandelen, 1974; Lodos & Kalkandelen, 1985b; Kartal, Zeybekoğlu & Dursun, 2001). Remarks: New for Antalya.

Macrosteles forficula (Ribaut, 1927) Material: Kemer, Çamyuva, 25.05.2002 2♂4♀; Finike, Erenkavak, 65 m, 24.06.2002 1♂3♀; Kemer, Karabucak, 85 m, 25.05.2002 1♂3♀; Kumluca, Kavakköy, 18.04.2001 3♂. It has been collected from weeds in fields. Distribution in Turkey: Adana, Amasya, Ankara, Bolu, Erzincan, Erzurum, Konya, Samsun, Sinop, Van (Kalkandelen, 1974; Lodos & Kalkandelen, 1985b; Zeybekoğlu, 1998). Remarks: New for Antalya.

Macrosteles quadripunctulatus (Kirschbaum, 1868) Material: Finike, Turunçova, 50 m, 06.06.2001 1♀; Korkuteli, Datköy, 960 m, 13.07.2002 1♂; Kemer, Böğürtlenözü, 18.04.2001 3♂1♀; Finike, Yeşilyurt, 18.04.2001 1♂7♀; Kale, Eseler, 55 m, 24.06.2002 5♂2♀; Kemer, Karabucak, 85 m, 25.05.2002 1♂6♀; Aksu, Topallar, 26.07.2001 2♂5♀; Serik, Camili, 26.07.2001 2♂1♀; Finike, Erenkavak, 65 m, 24.06.2002 2♂2♀. It has been collected from weeds in fields. Distribution in Turkey: Adana, Adıyaman, Ağrı, Amasya, Ankara, Bilecik, Bitlis, Diyarbakır, Erzincan, Hatay, Isparta, İzmir, Konya, Malatya, Manisa, Mersin, Nevşehir, Ordu, Samsun, Uşak, Van, Zonguldak (Kalkandelen, 1974; Lodos & Kalkandelen, 1985b; Başpınar & Uygun, 1991c; Zeybekoğlu, 1998). Remarks: New for Antalya.

Recilia schmidtgeni (Wagner, 1939) Material: Manavgat, Bucakşeyhler, 18-28.06.1997 2♂4♀; Manavgat, 18.09.1998 3♀; Bucakşeyhler, 13.08.1999 1♂; Demirciler, 10 m, 17.08.1999 2♂2♀; Kaş, Kalkan, Yeşilköy, 29.07.1999 1♂2♀; Elmalı, 1080 m, 28.07.1999 1♂; Manavgat, Taşağıl, Sağırin, 40 m, 03.06.2001 1♂; Bucakşeyhler, 70 m, 23.04.2002 1♂; Demirciler, 20 m, 18.05.2002 1♀; Alanya, Yeşilköy, 20.05.2002 1♂; Kaş, İslamlar, 220 m, 15.05.2001 3♂1♀; Kale, Köşkerler, 240 m, 15.05.2001 2♂3♀; Alanya, Konaklı, 25 m, 24.04.2002 3♂5♀; Serik, Sarıabalı, 20.08.2001 4♀; Manavgat, Karaöz, 70 m, 18.08.2001 1♂1♀; Kemer, Çamyuva, 21.08.2001 5♂4♀; Böğürtlenözü, 06.09.2002 2♂4♀; Alanya, Okurcalar, 09.09.2001 5♂; Manavgat, Beşkonak, 145 m, 08.09.2002 3♂1♀; Gazipaşa, Asarcık, 24.04.2002 3♂2♀; Kemer, Karabucak, 85 m, 25.05.2002 4♂1♀; Kaş, Kalkan, Üzümlü, 285 m, 15.05.2001 1♂1♀; Kumluca, Kavakdibi, 130 m, 24.06.2002 5♀. It has been collected from weeds in fields and Pinus brutia forest. Distribution in Turkey: Adana, Ankara, Diyarbakır, Erzincan, İzmir, Mersin, Nevşehir, Samsun, Sinop, Tokat, Urfa (Kalkandelen, 1974; Lodos & Kalkandelen, 1985c; Başpınar & Uygun, 1991c; Kartal & Zeybekoğlu, 1991; Zeybekoğlu, 1998). Remarks: New for Antalya.


Stymphalus rubrostriatus (Horvath, 1907) Material: Demirciler, 40 m, 20.06.2002 1♂3♀. It has been collected from Olea europea and subtropic vegetation. Distribution in Turkey: Diyarbakır, İstanbul, Mardin, Mersin (Dlabola, 1981; Lodos & Kalkandelen, 1985c; Başpınar & Uygun, 1992a). Remarks: New for Antalya.

Chiasmus conspurcatus (Perris, 1857) Material: Manavgat, 21.07.1999 3♂2♀; Manavgat, 10 m, 20.07.1999 1♀; Beşkonak, 145 m, 07.06.2001 2♀; Finike, Turunçova, 50 m, 06.06.2001 1♂; Alanya, Yeşilköy, 20.05.2002 1♂1♀; Elmalı, Gökpınar, 1200 m, 13.07.2002 1♀; Kemer, Çamyuva, 25.05.2002 1♂1♀. It has been collected from weeds in fields and Pinus brutia forest. Distribution in Turkey: Adana, Amasya, Ankara, Antalya, Artvin, Çankırı, Diyarbakır, Konya, Rize, Sinop, Tokat, Urfa (Kalkandelen, 1974; Lodos & Kalkandelen, 1985c; Başpınar & Uygun, 1992a; Zeybekoğlu, 1998). Remarks: Known to Antalya.

Doratura homophyla (Flor, 1861) Material: Elmalı, 1080 m, 28.07.1999 1♂2♀; Gökpınar, 1200 m, 13.07.2002 1♂; Korkuteli, Ulucak, 1165 m, 13.07.2002 2♂3♀; Güllük Dağı, Araştırma Ormanı, 550 m, 19.07.2002 11♂3♀; Elmalı, Tekkeköy, 1100 m, 28.08.2003 1♀. It has been collected from weeds in Garden and Pinus brutia forest. Distribution in Turkey: Adana, Amasya, Ankara, Bolu, Diyarbakır, Eskişehir, Gaziantep, Hatay, Sinop, Van (Linnavuori, 1965; Kalkandelen, 1974; Lodos & Kalkandelen, 1985c; Başpınar & Uygun, 1992a; Zeybekoğlu, 1998). Remarks: New for Antalya.

Aconurella prolixa (Lethierry, 1885) Material: Manavgat, Bucakşeyhler, 13.08.1999 3♂1♀; Demirciler, 10 m, 17.08.1999 2♂10♀; Manavgat, 10.08.1999 1♂13♀; Manavgat, 21.07.1999 3♂19♀; Manavgat, 18.09.1998 1♂10♀; Çayyazı, 20.07.1999 3♀; Manavgat, 10 m, 20.07.1999 4♀; Korkuteli, Yazır-İmrahor, 950 m, 27.07.1999 1♀; Manavgat, 26.07.1999 1♂1♀; Bucakşeyhler, 30 m, 23.07.1999 1♀; Bucakşeyhler, 24.07.1999 2♂5♀; Elmalı, Akçay, 1100 m, 28.07.1999 1♀; Elmalı, 1080 m, 28.07.1999 6♂5♀; Manavgat, Belenobası, 130 m, 15.04.2001 3♀; Beşkonak, 145 m, 07.06.2001 1♂2♀; Taşağıl, 55 m, 07.06.2001 1♀; Demirciler, 40 m, 20.06.2002 1♀; Korkuteli, Ulucak, 1165 m, 13.07.2002 1♀; Kaş, Kalkan, Üzümlü, 285 m, 15.05.2001 1♂1♀. It has been collected from weeds in fields and Salix. Distribution in Turkey: Adana, Ankara, Antalya, Artvin, Erzincan, Erzurum, Gaziantep, Hakkari, Kahramanmaraş, Mersin, Nevşehir, Rize (Linnavuori, 1965; Lodos & Kalkandelen, 1985c; Başpınar & Uygun, 1992a). Remarks: Known to Antalya.

Fieberiella aschei Meyer-Arndt, 1988 Material: Elmalı, Çığlıkara, 1600 m, 28.08.2003 1♂1♀; Korkuteli, 30 km W, 670 m, 28.08.2003 2♀. It has been collected from maquis vegetation. Distribution in Turkey: Antalya (Meyer-Arndt, 1988). Remarks: Endemic to Turkey and known to Antalya.

Fieberiella chioscola Meyer-Arndt, 1988 Material: Düzlerçamı, 280 m, 22.07.2001 2♂. On Crateagus. Distribution in Turkey: There is no record from Turkey. Remarks: New for Turkey.

Fieberiella macchiae Linnavuori, 1962 Material: Manavgat, Evrenleryavşı, Söylerkale, 250 m, 16.03.2002 1♂2♀. On Myrthus communis. Distribution in Turkey: Adana, Antalya, Denizli, Kütahya, Muğla, Nevşehir, Zonguldak (Lodos & Kalkandelen, 1986a). Remarks: Known to Antalya.

Fieberiella oenderi Dlabola, 1985 Material: Manavgat, Bucakşeyhler, 45 m 24.07.1999 1♂1♀; İbradı, Üzümdere, 470 m, 18.08.1999 1♀; Gündoğmuş, Çiçekoluk, 960 m, 25.07.1999 1♂; Manavgat, Demirciler, 10 m, 17.08.1999 1♀; Demirciler, 65 m, 14.04.2001 1♀; Demirciler, 105 m, 11.05.2001 1♀; Kemer, 06.06.2001 1♀; Termessos, Güllük Dağı, 900-1000 m, 22.07.2001 3♀; Akseki, Murtiçi, 540 m, 25.07.2001 1♂2♀; Manavgat, Demirciler, 40 m, 24.07.2001 4♂2♀; Demirciler, Külcüler, 150 m, 14.10.2001 1♂; Akseki, Güçlüköy, 550 m, 21.04.2002 1♂; Manavgat, Demirciler, 105 m, 22.04.2002 1♀; Gündoğmuş, Çiçekoluk, 960 m, 19.05.2002 1♂; Manavgat, Demirciler, 20 m, 21.06.2002 1♀; Demirciler, 40 m, 20.06.2002 2♀; Demirciler, 40 m, 17.07.2002 1♂1♀; Çığlık, 330 m, 19.07.2002 1♀; Beşkonak, 180 m, 27.08.2003 1♀. It has been collected from Pistacia terebinthus, Myrthus communis, Fontanesia philliraeoides and maquis


vegetation. Distribution in Turkey: Antalya, Isparta (Dlabola, 1985; Lodos & Kalkandelen, 1986a). Remarks: Endemic to Turkey and known to Antalya.

Synophropsis lauri (Horváth, 1897) Material: Güllük Dağı, Araştırma Ormanı, 550 m, 19.07.2002 1♀; Kemer, Çamyuva, 21.08.2001 1♂. On Laurus nobilis. Distribution in Turkey: Amasya, Ankara, Hatay, İstanbul, Mersin, Rize, Samsun, Sinop, Tokat, Trabzon, Yalova (Lodos & Kalkandelen, 1986a). Remarks: New for Antalya.

Docotettix cornutus Ribaut, 1948 Material: Manavgat, Bucakşeyhler, 18-28.06.1997 1♂; İbradı, Üzümdere, 470 m, 18.08.1999 1♂; Gündoğmuş, Çiçekoluk, 960 m, 25.07.1999 1♂; Manavgat, Bucakşeyhler, 45 m, 24.07.1999 7♂5♀; Demirciler, 10 m, 17.08.1999 5♂3♀; Demirciler, 65 m, 14.04.2001 1♂; Yukarıışıklar-Tilkiler, 460 m, 14.05.2001 1♀; Akseki, Murtiçi, 540 m, 25.07.2001 1♀; Gündoğmuş, Çiçekoluk, 960 m, 25.07.2001 1♀; Manavgat, Demirciler, 40 m, 24.07.2001 1♂; Bucakşeyhler, 45 m, 19.07.2001 1♂; Gündoğmuş, Kalecik, 500 m, 25.07.2001 4♂3♀; Manavgat, Demirciler, Külcüler, 150 m, 14.10.2001 4♀; Evrenleryavşı, Söylerkale, 250 m, 16.03.2002 2♂9♀; Gündoğmuş, Güneycik, 210 m, 17.03.2002 1♀; Manavgat, Demirciler, Külcüler, 230 m, 22.04.2002 1♂; Demirciler, 105 m, 22.04.2002 1♂2♀; Gündoğmuş, Güneycik, 210 m, 21.04.2002 2♂1♀; Serinyaka, 640 m, 19.05.2002 1♂; Manavgat, Demirciler, 105 m, 18.05.2002 2♀; Demirciler, 105 m, 21.06.2002 4♂6♀; Demirciler, 20 m, 21.06.2002 2♂; Demirciler, 40 m, 20.06.2002 1♂; Güllük dağı, Araştırma Ormanı, 750 m, 19.07.2002 1♂2♀; Serik, Sarıabalı, 18.07.2002 1♂; Kaş, Dirgenler, 265 m, 13.07.2002 1♂1♀; Manavgat, Demirciler, 40 m, 17.07.2002 1♂1♀; Beşkonak, 180 m, 27.08.2003 4♀; Köprülü Kanyon, 495 m, 27.08.2003 1♀; Korkuteli, 30 km W, 670 m, 28.08.2003 1♂2♀. On Myrthus communis, Olea europaea and Fontanesia philliraeoides. Distribution in Turkey: Antalya, Aydın, İzmir (Lodos & Kalkandelen, 1986a). Remarks: Known to Antalya.

Exitianus capicola (Stål, 1855) Material: Manavgat, 21.07.1999 33♂17♀; Alanya, Obaköy, 25 m, 12.08.1999 3♂12♀; Manavgat, Değirmenli, 04.08.1999 1♂; Manavgat, 10.08.1999 3♂6♀; Bucakşeyhler, 24.07.1999 9♂18♀; Bucakşeyhler, 30 m, 23.07.1999 3♂1♀; Manavgat, 26.07.1999 1♂2♀; Manavgat, 18.09.1998 13♂8♀; Bucakşeyhler, 13.08.1999 18♂17♀; Demirciler, 40 m, 21.07.1999 4♂3♀; Demirciler, 10 m, 17.08.1999 6♂2♀; Manavgat, 10 m, 20.07.1999 1♂9♀; Çayyazı, 20.07.1999 1♂2♀; Karaöz, 20 m, 22.07.1999 2♂3♀; Kaş, Kalkan, Yeşilköy, 29.07.1999 1♀; Manavgat, Belenobası, 130 m, 15.04.2001 4♂1♀; Demirciler, 65 m, 14.04.2001 1♀; Demirciler, 65 m, 11.05.2001 2♂3♀; Alanya, Okurcalar, 12.05.2001 2♀; Keşefli, 12.05.2001 1♂; Manavgat, Demirciler, 65 m, 05.06.2001 2♂1♀; Taşağıl, 55 m, 07.06.2001 07.06.2001 1♂; Demirciler, 40 m, 05.06.2001 1♀; Beşkonak, 145 m, 07.06.2001 3♂1♀; Finike, Turunçova, 50 m, 06.06.2001 1♀; Alanya, Kargıcak, 21.07.2001 2♂1♀; Manavgat, Ilıca, 16.10.2001 1♂1♀; Çakış, 16.10.2001 1♂1♀; Bucakşeyhler, Seleukeia, 250 m, 23.04.2002 2♂; Demirciler, 105 m, 22.04.2002 1♂; Demirciler, 105 m, 18.05.2002 1♀; Alanya, Kargıcak, 22.07.2002 2♂3♀; Manavgat, Hacıobası, 26.08.2003 1♀; Taşağıl, 55 m, 08.09.2002 5♂11♀; Kemer, Böğürtlenözü, 06.09.2002 9♂1♀; Manavgat, Ilıca, 07.09.2001 3♂; Serik, Dikmen, 26.07.2001 8♂3♀; Gazipaşa, Asarcık, 24.04.2002 2♂3♀; Macarköy, 24.04.2002 2♂; Kumluca, Kavakköy, 18.04.2001 2♂1♀; Finike, Yeşilyurt, 18.04.2001 3♀; Kemer, Çamyuva, 21.08.2001 4♂2♀; Gazipaşa, Çalıpınar, 25 m, 09.09.2001 2♂; Kale, Eseler, 55 m, 15.05.2001 2♀. It has been collected from weeds in fields and subtropic vegetation. Distribution in Turkey: Adana, Antalya, İzmir, Mersin, Muğla (Lodos & Kalkandelen, 1986a; Başpınar & Uygun, 1992a). Remarks: Known to Antalya.

Exitianus fasciolatus (Melichar, 1911) Material: Alanya, Okurcalar, 12.05.2001 1♀; Manvagat, Demirciler, 65 m, 11.05.2001 1♀; Demirciler, 105 m, 11.05.2001 1♂1♀; Yukarıışıklar-Tilkiler, 460 m, 14.05.2001 1♂; Tilkiler, 470 m, 14.05.2001 1♂; Yaylaalan, 540 m, 14.05.2001 1♀; Gündoğmuş, Çiçekoluk, 960 m, 19.05.2002 1♀. It has been collected from weeds in fields and Pinus brutia forest. Distribution in Turkey: Adana, Bitlis, Diyarbakır, Gaziantep, İzmir, Kahramanmaraş, Konya, Mardin, Mersin, Nevşehir, Niğde, Sakarya, Urfa (Lodos & Kalkandelen, 1986a; Başpınar & Uygun, 1992a). Remarks: New for Antalya.


Tetartostylus illyricus (Kirschbaum, 1868) Material: Manavgat, Bucakşeyhler, 18-28.06.1997 1♂; Demirciler, 105, 18.05.2002 1♀. It has been collected from weeds in fields and Pinus brutia forest. Distribution in Turkey: Ankara, Edirne, İstanbul (Kalkandelen, 1974; Lodos & Kalkandelen, 1986a). Remarks: New for Antalya.

Laminacutus libanensis Abdul-Nour, 2002 Material: Manavgat, Bucakşeyhler, 15-26.08.1997 2♀; Kemer, Çamyuva, 21.08.2001 2♂3♀. It has been collected from weeds in Pinus brutia forest and light traps. Distribution in Turkey: There is no record from Turkey. Remarks: New for Turkey.

Platymetopius enatus Dlabola, 1974 Material: Termessos, Güllük Dağı, 900-1000 m, 19.07.2002 1♂. It has been collected from weeds in Pinus brutia-Quercus forest. Distribution in Turkey: There is no record from Turkey. Remarks: New for Turkey.

Platymetopius guttatus Fieber, 1869 Material: Manavgat, Demirciler, 10 m, 17.08.1999 1♀. On Myrthus communis. Distribution in Turkey: Adana, Amasya, Bursa, İzmir, Ordu, Samsun, Sinop, Tokat, Trabzon (Lodos & Kalkandelen, 1986b; Kartal & Zeybekoğlu, 1991; Zeybekoğlu, 1998). Remarks: New for Antalya.

Platymetopius henribauti Dlabola, 1961 Material: Manavgat, Bucakşeyhler, 24.07.1999 1♀; Demirciler, 20 m, 18.05.2002 1♂. It has been collected from weeds and Populus. Distribution in Turkey: Ankara, Konya, Sivas (Kalkandelen, 1974; Lodos & Kalkandelen, 1986b; Demir, 2006). Remarks: New for Antalya.

Platymetopius quercinus Dlabola, 1974 Material: Akseki, Murtiçi, 540 m, 25.07.2001 1♀. On Quercus. Distribution in Turkey: Adıyaman, Kırşehir, Mardin (Lodos & Kalkandelen, 1986b). Remarks: New for Antalya.

Proceps acicularius Mulsant et Rey, 1855 Material: Manavgat, Demirciler, 40 m, 14.04.2001 1♀; Demirciler, 40 m, 11.05.2001 1♂; Kemer, 06.06.2001 1♀; Kemer, Çamyuva, 18.04.2001 1♀. On Sorghum halepense. Distribution in Turkey: Ankara (Dlabola, 1957). Remarks: New for Antalya.

Anoplotettix eckerleini Dlabola, 1965 Material: Manavgat, Beşkonak, Köprülü Kanyon, 495 m, 07.06.2001 2♀; Demirciler, 20 m, 18.05.2002 1♀; Gündoğmuş, Güneycik, 210 m, 22.06.2002 5♀. On Arbutus andrachne and Pistacia terebinthus. Distribution in Turkey: Antalya (Lodos & Kalkandelen, 1986b). Remarks: Known to Antalya.

Anoplotettix kalkandeleni Dlabola, 1971 Material: Gündoğmuş, Güneycik, 210 m, 22.06.2002 2♀. On Arbutus andrachne. Distribution in Turkey: Antalya (Dlabola, 1971). Remarks: Endemic to Turkey and known to Antalya.

Anoplotettix rodosicus Dlabola, 1974 Material: Gazipaşa, Çalıpınar, 25 m, 12.05.2001 2♀; Güllük Dağı, 750 m, 21.05.2002 8♂1♀; Gazipaşa, Çalıpınar-Kahyalar, 20 m, 20.05.2002 1♀. It has been collected from weeds Pinus brutia forest. Distribution in Turkey: Antalya, Muğla (Lodos & Kalkandelen, 1986b). Remarks: Known to Antalya.

Phlepsius intricatus (Herrich-Schaffer, 1838) Material: Manavgat, 10.08.1999 2♂2♀; Bucakşeyhler, 13.08.1999 3♂1♀; Bucakşeyhler, 24.07.1999 1♂1♀; Çayyazı, 20.07.1999 2♂2♀; Demirciler, 40 m, 05.06.2001 1♂; Beşkonak, 145 m, 07.06.2001 1♂; Elmalı, Akçay, 1210 m, 13.07.2002 1♀. It has been collected from weeds in fields and Pinus brutia forest. Distribution in Turkey: Ankara, Amasya, Bolu, Çanakkale, Diyarbakır, Isparta, İzmir, Kars, Kırşehir, Malatya, Mardin, Muğla, Nevşehir, Samsun, Trabzon (Linnavuori, 1965; Kalkandelen, 1974; Lodos & Kalkandelen, 1986c; Kartal & Zeybekoğlu, 1991; Zeybekoğlu, 1998; Demir, 2006). Remarks: New for Antalya.

Selenocephalus ankarae Dlabola, 1957 Material: Termessos, Güllük Dağı, 900-1000 m, 22.07.2001 8♀; Güllük Dağı, 540 m, 22.07.2001 2♀; Manavgat, Bucakşeyhler, 45 m, 19.07.2001 1♀; Akseki, Murtiçi, 540 m,


25.07.2001 1♂2♀; Termessos, Güllük Dağı, 900-1000 m, 19.07.2002 5♂4♀; Kumluca, Sarıcasu, 575 m, 13.07.2002 2♂; İbradı, Akşahap-İbradı, 530 m, 21.08.2002 2♀. On Phlomis, Fontanesia philliraeoides and shrubs. Distribution in Turkey: Amasya, Ankara, Antalya, Muğla, Tokat (Dlabola, 1957; Kalkandelen, 1974; Lodos & Kalkandelen, 1986c; Zeybekoğlu, 1998; Demir, 2004b). Remarks: Endemic to Turkey and known to Antalya.

Selenocephalus pallidus Kirschbaum, 1868 Material: Manavgat, Bucakşeyhler, 45 m, 24.07.1999 1♀; Bucakşeyhler 30 m 23.07.1999 1♂; Gündoğmuş, Çiçekoluk, 960 m, 25.07.1999 1♂; Akseki, Murtiçi, 540 m, 25.07.2001 1♂1♀; Gündoğmuş, Kalecik, 500 m, 25.07.2001 2♀; Güneycik, 210 m, 22.06.2002 5♂1♀; Manavgat, Demirciler, 40 m, 20.06.2002 1♂; Serik, Sarıabalı, 18.07.2002 1♂1♀; Kaş, Kemerköy, 260 m, 13.07.2002 3♂1♀; Gündoğmuş, Güneycik, 210 m, 14.07.2002 4♂3♀; Kemer, Olimpos Milli Parkı, 20.08.2002 4♀; Hisarçandır, 770 m, 20.08.2002 3♀; Kumluca, Altınyaka-Karacağaç, 910 m, 20.08.2002 2♂5♀; İbradı, Ormana, 1000 m, 21.08.2002 1♀; Manavgat, Beşkonak, 180 m, 27.08.2003 1♂8♀. On Cistus creticus, Olea europaea and shrubs. Distribution in Turkey: Adana, Ankara, Antalya, Aydın, Bolu, Burdur, Bursa, Çanakkale, Çankırı, Çorum, Denizli, Isparta, İzmir, Kayseri, Kırşehir, Konya, Manisa, Samsun, Sinop, Tokat, Yozgat (Dlabola, 1957; Linnavuori, 1965; Kalkandelen, 1974; Lodos & Kalkandelen, 1986c; Kartal & Zeybekoğlu, 1991). Remarks: Known to Antalya.

Hardya anatolica Zachvatkin, 1946 Material: Korkuteli, Koru Dağı, Kartal Yaylası, 1600 m, 23.06.2002 1♀. It has been collected from weeds in fields. Distribution in Turkey: Adıyaman, Ankara, Antalya, Bolu, Elazığ, Erzincan, Isparta, İzmir, Kahramanmaraş, Karaman, Kayseri, Kırşehir, Malatya, Nevşehir, Ordu (Zachvatkin, 1946; Dlabola, 1957; Linnavuori, 1965; Kalkandelen, 1974; Lodos & Kalkandelen, 1987a; Demir, 2004b, 2006). Remarks: Known to Antalya.

Eohardya fraudulenta (Horváth, 1903) Material: Manavgat, Bucakşeyhler, Seleukeia, 250 m, 23.04.2002 1♀; Güllük Dağı, Araştırma Ormanı, 750 m, 21.05.2002 3♂; Kemer, Çamyuva, 18.04.2001 1♂. It has been collected from weeds in fields and Pinus brutia forest. Distribution in Turkey: Ankara, İstanbul (Zachvatkin, 1964; Kalkandelen, 1974; Lodos & Kalkandelen, 1987a; Demir, 2006). Remarks: New for Antalya.

Cicadula lineatopunctata (Matsumura, 1908) Material: Kemer, Çamyuva, 18.04.2001 2♂2♀. It has been collected from weeds in fields. Distribution in Turkey: Adana, Ankara, Aydın, Bolu, Bursa, Çankırı, Diyarbakır, Düzce, Erzincan, İzmir, Malatya, Mardin, Sakarya, Sinop, Urfa (Dlabola, 1957; Kalkandelen, 1974; Lodos & Kalkandelen, 1987a). Remarks: New for Antalya.

Mocydiopsis monticola Remane, 1961 Material: Akseki, Güçlüköy, 680 m, 17.04.2001 1♂; Manavgat, Demirciler, 65 m, 05.06.2001 1♀; Taşağıl, 55 m, 07.06.2001 1♀; Beşkonak, 145 m, 07.06.2001 4♀; Demirciler, 40 m, 05.06.2001 8♂16♀. It has been collected from weeds in Pinus brutia forest. Distribution in Turkey: Adana, Diyarbakır, Mersin, Samsun, Uşak (Lodos & Kalkandelen, 1987a, Başpınar & Uygun, 1992b). Remarks: New for Antalya.

Thamnotettix creticus Dlabola, 1974 Material: Akseki, Güçlüköy, 680 m, 17.04.2001 3♂1♀; Geriş, 700 m, 17.04.2001 8♂1♀; Gündoğmuş, Güneycik, 210 m, 17.04.2001 1♂; Manavgat, Bucakşeyher, 70 m, 23.04.2002 1♂; Gündoğmuş, Serinyaka, 640 m, 19.05.2002 2♂1♀; İbradı, Başlar, 1440 m, 21.08.2002 1♂. It has been collected from weeds in fields and Pinus brutia forest.Distribution in Turkey: Adana, Amasya, Hatay, Mersin, Samsun, Sinop, Tokat, Osmaniye (Kartal, 1983; Lodos & Kalkandelen, 1987a; Kartal & Zeybekoğlu, 1991). Remarks: New for Antalya.

Thamnotettix vittmeri Dlabola, 1971 Material: Gündoğmuş, Güneycik, 210 m, 17.04.2001 1♀; Manavgat, Demirciler, 65 m, 22.04.2002 1♂; Bucakşeyhler, Seleukeia, 250 m, 23.04.2002 2♀. It has been collected from weeds in fiel ds and Pinus brutia forest. Distribution in Turkey: İzmir, Muğla (Dlabola, 1971). Remarks: New for Antalya and endemic to Turkey.


Thamnotettix zelleri (Kirschbaum, 1868) Material: Manavgat, Hocalı, 15.04.2001 1♀; Demirciler, 65 m, 14.04.2001 37♂19♀; Demirciler, Külcüler, 170 m, 15.04.2001 17♂23♀; Demirciler, 40 m, 14.04.2001 19♂17♀; Belenobası, 130 m, 15.04.2001 8♂5♀; Saraçlı, 15.04.2001 5♂7♀; Yaylaalan, 540 m, 14.05.2001 1♂5♀; Tilkiler, 220 m, 14.05.2001 1♂1♀; Demirciler, 65 m, 11.05.2001 6♂3♀; Yukarıışıklar-Tilkiler, 460 m, 14.05.2001 1♂18♀; Demirciler, 40 m, 11.05.2001 3♂12♀; Demirciler, 65 m, 05.06.2001 1♂1♀; Beşkonak, 145 m, 07.06.2001 2♀; Demirciler, 40 m, 22.04.2002 10♂14♀; Bucakşeyhler, 70 m, 23.04.2002 5♂1♀; Bucakşeyhler, Seleukeia, 250 m, 23.04.2002 2♂4♀; Demirciler, 105 m, 22.04.2002 4♂; Demirciler, 65 m, 22.04.2002 13♂14♀; Akseki, Geriş-Cemerler, 650 m, 21.04.2002 2♂1♀; Gündoğmuş, Güneycik, 240 m, 21.04.2002 11♂7♀; Güneycik, 240 m, 19.05.2002 1♂3♀; Çiçekoluk, 960 m, 22.06.2002 1♂; Kemer, Çamyuva, 18.04.2001 1♀. It has been collected from weeds in fields and Pinus brutia forest.Distribution in Turkey: Adana, Antalya, Aydın, Bursa, Çanakkale, Çorum, Edirne, Hatay, İzmir, Kırklareli, Manisa, Mersin, Muğla, Tekirdağ (Lodos & Kalkandelen, 1987a; Başpınar & Uygun, 1992b). Remarks: Known to Antalya.

Limotettix striola (Fallén, 1806) Material: Manavgat, Bucakşeyhler, 18-28.06.1997 1♂; Akseki, Geriş, 700 m, 17.04.2001 2♀; Kaş, İslamlar, 220 m, 15.05.2001 1♂. It has been collected from weeds in Pinus brutia forest. Distribution in Turkey: Ağrı, Ankara, Artvin, Bolu, Bursa, Çankırı, Diyarbakır, Erzincan, Erzurum, Iğdır, İstanbul, Kahramanmaraş, Konya, Malatya, Nevşehir, Niğde, Sakarya, Urfa, Van, Zonguldak (Dlabola, 1957; Kalkandelen, 1974; Lodos & Kalkandelen, 1987b). Remarks: New for Antalya.

Euscelidius mundus (Haupt, 1927) Material: Manavgat, Bucakşeyhler, 18-28.06.1997 15♂18♀; Kemer, Çamyuva, 18.04.2001 3♂6♀; Karabucak, 25.05.2002 5♂11♀. It has been collected from weeds in fields and Pinus brutia forest. Distribution in Turkey: Adana, Ankara, Diyarbakır, Erzurum, Gaziantep, Hatay, Isparta, İzmir, Mardin, Mersin, Urfa, Van (Linnavuori, 1965; Kalkandelen, 1974; Lodos & Kalkandelen, 1987b; Başpınar & Uygun, 1992b). Remarks: New for Antalya.

Euscelidius schencki (Kirschbaum, 1868) Material: Manavgat, Demirciler, 40 m, 05.06.2001 1♂3♀. It has been collected from weeds in fields and Pinus brutia forest. Distribution in Turkey: Adana, Ankara, Bursa, Diyarbakır, İzmir, Mersin, Siirt (Kalkandelen, 1974; Lodos & Kalkandelen, 1987b; Başpınar & Uygun, 1992b). Remarks: New for Antalya.

Euscelidius variegatus (Kirschbaum, 1858) Material: Manavgat, Evrenleryavşı, Söylerkale, 250 m, 16.03.2002 1♂2♀. On Cistus creticus. Distribution in Turkey: Bursa, İstanbul (Lodos & Kalkandelen, 1987b). Remarks: New for Antalya.

Euscelis alsioides Remane, 1967 Material: Manavgat, Demirciler, 65 m, 14.04.2001 1♂; Alanya, Okurcalar, 12.05.2001 7♂6♀; Manavgat, Yukarıışıklar, 200 m, 14.05.2001 2♀; Tilkiler, 220 m, 14.05.2001 1♂; Demirciler, 105 m, 11.05.2001 1♀; Demirciler, 40 m, 05.06.2001 3♂9♀; Alanya, Okurcalar, 20.05.2002 1♂. It has been collected from weeds in fields and Pinus brutia forest. Distribution in Turkey: There is no record from Turkey. Remarks: New for Turkey.

Euscelis alsius Ribaut, 1952 Material: Manavgat, Bucakşeyhler, 15-26.08.1997 2♀; Gündoğmuş, Çiçekoluk, 960 m, 25.07.1999 1♂7♀; Serinyaka, 700 m, 25.07.1999 1♂3♀; Manavgat, Demirciler, 65 m, 21.07.1999 1♂; Korkuteli, Yazır-İmrahor, 950 m, 27.07.1999 1♀; Gazipaşa, Çalıpınar, 25 m, 12.05. 2001 8♂10♀; Alanya, Okurcalar, 12.05.2001 7♂5♀; Keşefli, 12.05.2001 1♀; Manavgat, Yukarıışıklar-Tilkiler, 460 m, 14.05.2001 1♂1♀; Demirciler, 40 m, 11.05.2001 1♂; Demirciler, 65 m, 11.05.2001 1♂; Beşkonak, 145 m, 07.06.2001 4♂4♀; Demirciler, 65 m, 05.06.2001 3♂2♀; Taşağıl, Kilimli, 40 m, 03.06.2001 1♀; Taşağıl, 55 m, 07.06.2001 1♂1♀; Demirciler, 40 m, 05.06.2001 4♂; Gündoğmuş, Güneycik, 210 m, 08.06.2001 7♂2♀; Çiçekoluk, 960 m, 25.07.2001 2♂1♀; Güllük Dağı, Araştırma Ormanı, 750 m, 21.05.2002 1♀; Gazipaşa, Çalıpınar-Kahyalar, 20 m, 20.05.2002 21♂6♀; Alanya, Yeşilköy, 20.05.2002 1♂3♀; Gündoğmuş, Serinyaka, 625 m, 22.06.2002 1♀; Korkuteli, Söğütcük, 1010 m, 13.07.2002 2♂. It has been collected from weeds in fields and Pinus brutia forest. Distribution in Turkey: Adana, Ankara, Bolu, Mersin, Nevşehir, Niğde, Sakarya, Van


(Linnavuori, 1965; Kalkandelen, 1974; Başpınar & Uygun, 1992b). Remarks: New for Antalya.

Euscelis incisus (Kirschbaum, 1858) Material: Manavgat, Bucakşeyhler, 18-28.06.1997 2♀; Bucakşeyhler, 45 m, 24.07.1999 1♂; Bucakşeyhler, 30 m, 23.07.1999 1♂; Çayyazı, 20.07.1999 1♂. It has been collected from weeds in fields and Pinus brutia forest. Distribution in Turkey: Adana, Ankara, Burdur, Edirne, Eskişehir, Isparta, Kayseri, Kocaeli, Mersin (Dlabola, 1957; Kalkandelen, 1974; Başpınar & Uygun, 1992b ; Demir, 2004b, 2006). Remarks: New for Antalya.

Euscelis lineolatus Brullé, 1832 Material: Manavgat, Belenobası, 130 m, 15.04.2001 2♂1♀; Demirciler, 65 m, 14.04.2001 1♀; Hocalı, 215 m, 15.04.2001 1♂3♀; Akseki, Geriş, 700 m, 17.04.2001 3♀; Alanya, Okurcalar, 12.05.2001 1♀; Manavgat, Yukarıışıklar-Tilkiler, 460 m, 14.05.2001 1♀; Yaylaalan, 540 m, 14.05.2001 1♀. It has been collected from weeds in fields and Pinus brutia forest. Distribution in Turkey: Ankara, Artvin, Balıkesir, İzmir, Kırıkkale, Kırklareli, Kırşehir, Konya, Malatya, Manisa, Niğde, Samsun, Trabzon, Uşak (Kerville, 1939; Lodos & Kalkandelen, 1987b; Kartal & Zeybekoğlu, 1991). Remarks: New for Antalya.

Artianus manderstjernii (Kirschbaum, 1868) Material: Erenkaya, 680 m, 25.07.1999 1♀; Elmalı, 1080 m, 28.07.1999 2♂3♀; Korkuteli, Beğiş-Karabayır, 1310 m, 06.06.2001 1♂; Gündoğmuş, Çiçekoluk, 960 m, 25.07.2001 1♀; Korkuteli, Koru Dağı, Kartal Yaylası, 1600 m, 23.06.2002 2♂1♀; Akseki, Erenkaya, 630 m, 22.06.2002 1♀; Güllük Dağı, Araştırma Ormanı, 550 m, 19.07.2002 22♂19♀; Elmalı, Gökpınar, 1200 m, 13.07.2002 6♂3♀; Korkuteli, Söğütcük, 1010 m, 13.07.2002 8♂17♀; Elmalı, Çığlıkara, 1600 m, 28.08.2003 2♂3♀; Çığlıkara, 1820 m, 28.08.2003 1♀. It has been collected from weeds in fields and Pinus brutia forest. Distribution in Turkey: Ağrı, Ankara, Balıkesir, Çankırı, Diyarbakır, Edirne, Gaziantep, İzmir, Kırklareli, Urfa, Van, Zonguldak (Dlabola, 1957; Kalkandelen, 1974; Lodos & Kalkandelen, 1987b; Demir, 2004b, 2006). Remarks: New for Antalya.

Zercanus rubroocellatus Dlabola, 1965 Material: Manavgat, Bucakşeyhler, 18-28.06.1997 1♀. It has been collected from weeds in Pinus brutia forest. Distribution in Turkey: Adana (Lodos & Kalkandelen, 1987b). Remarks: New for Antalya.

Paralimnus phragmitis (Boheman, 1847) Material: Manavgat, 10.08.1999 1♀. On Arundo donax. Distribution in Turkey: There is no record from Turkey. Remarks: New for Turkey.

Paralimnellus cingulatus (Dlabola, 1960) Material: Manavgat, Örenşehir, 15.10.2001 1♂. It has been collected from weeds in subtropic vegetation. Distribution in Turkey: There is no record from Turkey. Remarks: New for Turkey.

Arocephalus languidus (Flor, 1861) Material: Manavgat, Bucakşeyhler, 13.08.1999 1♀. It has been collected from weeds in Pinus brutia forest. Distribution in Turkey: Ankara, Kars (Lodos & Kalkandelen, 1987c). Remarks: New for Antalya.

Psammotettix alienus (Dahlbom, 1850) Material: Elmalı, 1080 m, 28.07.1999 18♂14♀; Akçay, 1100 m, 29.07.1999 1♂; Finike, Turunçova, 50 m, 06.06.2001 2♂3♀; Kemer, 06.06.2001 14♂14♀; Akseki, Erenkaya, 630 m, 22.06.2002 10♂2♀; Korkuteli, Koru dağı, Kartal Yaylası, 1600 m, 23.06.2002 13♂13♀; Datköy, 960 m, 13.07.2002 35♂35♀; Ulucak, 1165 m, 13.07.2002 6♂5♀; Söğütcük, 1010 m, 13.07.2002 2♂; Elmalı, Çığlıkara, 1820 m, 28.08.2003 1♀; Çığlıkara, 1600 m, 28.08.2003 2♂2♀; Kemer, Çamyuva, 18.04.2001 13♂5♀; Kale, Köşkerler, 240 m, 15.05.2001 27♂19♀; Kumluca, Kavakdibi, 130 m, 24.06.2002 31♂48♀; Finike, Turunçova, 06.09.2002 3♂8♀; Kemer, 06.09.2002 1♂4♀ Düzlerçamı, 280 m, 22.07.2001 1♀; Finike, 06.06.2001 1♀; Güllük Dağı, Araştırma Ormanı, 750 m, 21.05.2002 2♀; Elmalı, Akçay, 1210 m, 13.07.2002 1♂1♀; Gökpınar, 1200 m, 13.07.2002 14♂17♀. It has been collected from weeds in fields and Pinus brutia forest. Distribution in Turkey: Adana, Amasya, Ankara, Balıkesir, Bolu, Bursa, Çorum, Diyarbakır, Erzincan, Erzurum, Hakkari, Iğdır, Kastamonu, Konya, Malatya, Mardin, Mersin, Muş, Nevşehir, Sakarya, Samsun, Siirt, Sinop, Uşak, Van, Yozgat


(Kalkandelen, 1974; Lodos & Kalkandelen, 1987c; Kartal et. al, 1991, 2001; Demir, 2004b, 2006). Remarks: New for Antalya.

Psammotettix cerinus (Lindberg, 1948) Material: Manavgat, 21.07.1999 5♂2♀; Manavgat, 10 m, 20.07.1999 5♀; Bucakşeyhler, 30m, 23.07.1999 1♂3♀; Çayyazı, 20.07.1999 7♂11♀; Karaöz, 20 m, 22.07.1999 1♂; Değirmenli, 04.08.1999 1♂; Demirciler, 10 m, 17.08.1999 1♂1♀; Bucakşeyhler, 13.08.1999 5♂4♀; Bucakşeyhler, 24.07.1999 1♂3♀; Manavgat, 18.09.1998 2♀; Manavgat, 26.07.1999 1♂; Manavgat-Çeltikçi, 14.04.2001 1♂; Demirciler, 65 m, 14.04.2001 1♀; Demirciler, Külcüler, 170 m, 15.04.2001 1♂; Belenobası, 130 m, 15.04.2001 13♂6♀; Yaylaalan, 540 m, 14.05.2001 2♂; Alanya, Okurcalar, 12.05.2001 1♂3♀; Keşefli, 12.05.2001 1♂; Manavgat, Beşkonak, 145 m, 07.06.2001 1♂; Demirciler, 40 m, 05.06.2001 1♂; Bucakşeyhler, 70 m, 23.04.2002 2♂6♀; Alanya, Yeşilköy, 20.05.2002 3♀; Okurcalar, 20.05.2002 1♀; Gazipaşa, Çalıpınar-Kahyalar, 20 m, 20.05.2002 1♀; Manavgat, Demirciler, 40 m, 20.06.2002 1♂; Alanya, Kargıcak, 22.08.2002 1♀. It has been collected from weeds in fields and subtropic vegetation. Distribution in Turkey: Adana, Hatay, Mersin (Dlabola, 1957; Linnavuori, 1965; Lodos & Kalkandelen, 1987c; Başpınar & Uygun, 1992b). Remarks: New for Antalya.

Psammotettix confinis (Dahlbom, 1850) Material: Elmalı, Tekkeköy, 1100 m, 28.08.2003 1♂. It has been collected from weeds in fields. Distribution in Turkey: Adana, Ağrı, Balıkesir, Bitlis, Edirne, Erzincan, Erzurum, Giresun, Hakkari, Iğdır, Isparta, Konya, Ordu, Samsun, Sivas, Van (Dlabola, 1957; Kalkandelen, 1974; Lodos & Kalkandelen, 1987c; Kartal et. al., 1991, 2001). Remarks: New for Antalya.

Psammotettix nodosus (Ribaut, 1925) Material: Antalya: Manavgat, 10 m, 20.07.1999 5♂5♀; Alanya, Okurcalar, 20.05.2002 6♂4♀ Alanya, Okurcalar, 12.05.2001 1♂. It has been collected from weeds in fields. Distribution in Turkey: There is no record from Turkey. Remarks: New for Turkey.

Psammotettix pictipennis (Kirschbaum, 1868) Material: Manavgat, Bucakşeyhler, 18-28.06.1997 1♂1♀; Akseki, Erenkaya, 680 m, 25.07.1999 3♀. It has been collected from weeds in fields and Pinus brutia forest. Distribution in Turkey: Adana, Ankara, Bartın, Bolu, Çankırı, Çorum, Erzincan, Hakkari, Iğdır, Kahramanmaraş, Kayseri, Karaman, Kocaeli, Malatya, Niğde, Urfa, Van (Dlabola, 1957; Kalkandelen, 1974; Lodos & Kalkandelen, 1987c; Kartal et. al., 2001). Remarks: New for Antalya.

Psammotettix provincialis (Ribaut, 1925) Material: Elmalı, Çobanisa, 1240m, 27.07.1999 3♂1♀; Gündoğmuş, Serinyaka, 700 m, 25.07.1999 1♀; Manavgat, Çayyazı, 20.07.1999 1♂; Akseki, Geriş, 700 m, 17.04.2001 17♂18♀; Güçlüköy, 680 m, 17.04.2001 2♂5♀; Gündoğmuş, Çiçekoluk, 960 m, 25.07.2001 2♂; Alanya, Yeşilköy, 20.05.2002 2♂2♀; Gündoğmuş, Güneycik, 240 m, 10.09.2002 2♂3♀; Kemer, Böğürtlenözü, 18.04.2001 7♂19♀; Kaş, Kalkan, Üzümlü, 285 m, 15.05.2001 2♂5♀; Gündoğmuş, Serinyaka, 640 m, 19.05.2002 1♀; İbradı, Akşahap-İbradı, 530 m, 21.08.2002 1♀. It has been collected from weeds in fields and Pinus brutia forest. Distribution in Turkey: Adana, Ankara, Antalya, Bolu, Çankırı, Diyarbakır, Edirne, Erzurum, İstanbul, İzmir, Kayseri, Konya, Mersin, Nevşehir, Sakarya, Samsun, Van (Dlabola, 1957; Kalkandelen, 1974; Lodos & Kalkandelen, 1987c; Kartal & Zeybekoğlu, 1991; Başpınar & Uygun, 1992b; Demir, 2006). Remarks: Known to Antalya.

Psammotettix striatus (Linnaeus, 1758) Material: Manavgat, Bucakşeyhler, 18-28.06.1997 26♂26♀; Bucakşeyhler, 15-26.08.1997 2♂1♀; Manavgat, 10.08.1999 1♂7♀; Manavgat, 21.07.1999 5♂11♀; Manavgat, 26.07.1999 1♀; Bucakşeyhler, 30 m, 23.07.1999 1♂1♀; Değirmenli, 4.08.1999 2♀; Bucakşeyhler, 13.08.1999 5♂8♀; Bucakşeyhler, 45 m, 24.07.1999 1♂; Demirciler, 10 m, 17.08.1999 1♂1♀; Demirciler, 40 m, 21.07.1999 1♂; Demirciler, 65 m, 21.07.1999 1♀; Bucakşeyhler, 24.07.1999 3♂4♀; Manavgat, 18.09.1998 5♂4♀; Demirciler, 65 m, 14.04.2001 2♀; Belenobası, 130 m, 15.04.2001 1♂2♀; Manavgat-Çeltikçi, 14.04.2001 1♂; Taşağıl, Sağırin, 40 m, 03.06.2001 2♂; Demirciler, 65 m, 05.06.2001 6♂7♀; Beşkonak, 145 m, 07.06.2001 43♂51♀; Bucakşeyhler, 22.07.2003 2♂10♀; Hacıobası, 26.08.2003 1♂1♀ Manavgat, Yaylaalan, 540 m, 14.05.2001 1♀; Taşağıl, Kilimli, 40 m, 03.06.2001 3♀; Taşağıl, 55 m, 07.06.2001 5♀; Alanya, Kargıcak, 21.07.2001 1♀; Manavgat, Demirciler, 65 m, 22.04.2002


1♂; Bucakşeyhler, 70 m, 23.04.2002 4♀; Demirciler, 105 m, 18.05.2002 1♀; Demirciler, 20 m, 18.05.2002 1♀; Manavgat, Demirciler, 40 m, 20.06.2002 4♂3♀; Demirciler, 20 m, 21.06.2002 1♂; Demirciler, 40 m, 17.07.2002 1♂; Serik, Dikmen, 18.07.2002 1♀. It has been collected from weeds in fields and Pinus brutia forest. Distribution in Turkey: Adana, Ankara, Kayseri, Kocaeli, Konya, Mersin, Nevşehir, Samsun (Dlabola, 1957; Kalkandelen, 1974; Kartal & Zeybekoğlu, 1991; Başpınar & Uygun, 1992b). Remarks: New for Antalya.

Ebarrius cognatus (Fieber, 1869) Material: Akseki, Güçlüköy, 680 m, 17.04.2001 1♂. It has been collected from weeds in Pinus brutia forest.. Distribution in Turkey: Ankara, Bolu, Çankırı, Erzincan, Kars, Kayseri, Niğde, Van (Dlabola, 1957; Kalkandelen, 1974; Lodos & Kalkandelen, 1987d; Demir, 2004b, 2006). Remarks: New for Antalya.

Jassargus kurdicus Remane and Schulz, 1976 Material: Düzlerçamı, 280 m, 22.07.2001 1♂4♀. It has been collected from weeds in Pinus brutia forest. Distribution in Turkey: Kırıkkale (Lodos & Kalkandelen, 1987d). Remarks: New for Antalya.

* This study is prepared from the Ph. D. Thesis approved by the Institute of Natural Sciences of Gazi University on August 2005.

ACKNOWLEDGEMENTS

I thank Dr. Yasemin Özdemir, Dr. Mustafa Özdemir, Dr. Işıl Özdemir, Dr. Hani Abdul-Nour and my family for their help during different phases of this project.

LITERATURE CITED Abdul-Nour, H. 1988. Studies on the genus Mocydia Edwards, 1922, with the description of a new species: M aegea n. sp. (Homoptera Auchenorrhyncha Cicadellidae). Marburger Entomologische Publikation. 2(4): 131-148. Akman, Y. 1995. Türkiye Orman Vejetasyonu. Ankara Üniv. Fen Fak. Yayınları, Ankara. 326 pp. Başpınar, H. & Uygun, N. 1991-1992. Doğu Akdeniz Bölgesi turunçgil bahçelerindeki Cicadellidae türleri üzerinde faunistik ve sistematik çalışmalar I-V. Türkiye entomoloji dergisi 15(2-3-4): 89-106; 157-172; 203-222, 16 (1-2): 47-64; 99-114. Demir, E. 2004a. Goniagnathus guttulinervis (Kirschbaum, 1868) new to Turkey, with data on ditribution of the Genus in Antalya (Homoptera: Auchenorryncha: Cicadellidae). Acta entomologica Slovenica 12 (2): 255-257. Demir, E. 2004b. Karyağdı Dağı (Ankara) Auchenorrhyncha Faunası Hakkında (Homoptera). Priamus 11(1/2): 17-27. Demir, E. 2005a. Two new records of Cicadellidae fauna in Turkey (Homoptera, Auchenorrhyncha). Zoology in the Middle East 35: 111. Demir, E. 2005b. Review of Paradorydium Kirkaldy (Homoptera, Auchenorrhyncha, Cicadellidae) from Turkey, with the description of a new species. Entomological News 116(2): 75-82. Demir, E. 2006. Contributions to the knowledge of Turkish Auchenorrhyncha (Homoptera) with a new record, Pentastiridius nanus (Ivanov, 1885). Munis entomology and zoology 1(1): 97-122.


Dlabola, J. 1957. Results of the Zoological Expedition of the National Museum in Prague to Turkey 20. Homoptera, Auchenorrhyncha. Acta entomologica Musei Nationale Pragae 31(469): 19-68. Dlabola, J. 1963. A Revision of the Leafhopper Genus Macropsidius Rib. (Homoptera, Auchenorrhyncha). Casopis Ceskoslovenske Scolecnosti Entomologicke 60(1-2): 115-124. Dlabola, J. 1971. Taxonomische und Chorologische Erganzungen der Zikadenfauna von Anatolien, Iran, Afghanistan und Pakistan (Homoptera, Auchenorrhyncha). Acta entomologica bohemoslovaca 68: 377-396. Dlabola, J. 1971. Taxonomische und chorologische Erzangungen zur türkischen und iranischen Zikadenfauna (Homoptera, Auchenorrhyncha). Acta faunustica entomologica Musei Nationale Pragae 14(163): 115-138. Dlabola, J. 1974. Zur Taxonomie und Chorologie einiger Mediterraner Zikaden (Homoptera, Auchenorrhyncha). Acta Zoologica Academia Scientiarum Hungaricae 20(3-4): 289-308. Dlabola, J. 1979. Neue Zikaden aus Anatolien, Iran und aus Süeeuropaischen Landern (Homoptera, Auchenorrhyncha). Acta Zoologica Academia Scientiarum Hungaricae 25(3-4): 235-257. Dlabola, J. 1981. Ergebnisse der Tschechoslowakisch-Iranischen Entomologischen Expeditionen nach dem Iran (1970 und 1973). Acta entomologica Musei Nationale Pragae 40: 127-311. Dlabola, J. 1985. Zwei neue Fieberiella-Arten aus der Türkei und Spanien (Homoptera,Cicadellidae). Türkiye Bitki Koruma Dergisi 9: 75-78. Dworakowska, I. 1969. On the Genera Zyginella Löw and Limasolla Dlab. (Cicadellidae, Typhlocybinae). Bulletin de L'Academie Polonaise des Sciences 17(7): 433-438. Dworakowska, I. 1970. On the Genus Arboridia Zachv. (Auchenorrhyncha, Cicadellidae, Typhlocybinae). Bulletin de L'Academie Polonaise des Sciences 18(10): 607-615. Dworakowska, I. 1971. On the genera related to Tamaricella Zachv. and some other Erythroneurini (Hom., Cicadellidae, Typhlocybinae). Annales Entomologica Fennica 37(2): 99-121. Dworakowska, I. 1972. On some Species of the Genus Eupteryx Curt. (Auchenorrhyncha, Cicadellidae, Typhlocybinae). Bulletin de L'Academie Polonaise des Sciences 20(10): 727-734. Dworakowska, I. 1976. Kybos Fieb., subgenus of Empoasca Walsh (Auchenorrhyncha, Cicadellidae, Typhlocybinae) in Palaearctic. Polska Akademia Nauk 21(13): 387-463. Dworakowska, I. 1982. Typhlocybini of Asia. Entomologische Abhandlungen 45(6): 99-181. Fahringer, J. 1922. Eine Rhynchotenausbeute aus der Türkei, Kleinasien und den benachbarten Gebieten. Konowia 1: 296-307. Gadeau de Kerville, H. 1939. Recit Sommaire du Voyage en Liste Methodique des Invertebres et des Vertebres Recoltes en Asie-Mineure. Voyage Zoologique D’Henri Gadeau de Kerville en Asie-Mineure (Avril-Mai, 1912). Paul Le Chevalier, Paris. 148 pp. Güçlü, Ş. & Özbek, H. 1994. Erzurum Yöresinde Cicadellidae (Homoptera, Auchenorrhyncha) Türleri Üzerinde Faunistik ve Sistematik Çalışmalar II. Idiocerinae,


Iassinae, Dorycephalinae, Aphrodinae, Cicadellinae. Türkiye entomoloji dergisi 18(2): 103-117. Güçlü, Ş. & Özbek, H. 1995. Türkiye Cicadellidae (Homoptera) Faunası İçin Yeni Kayıtlar. Türkiye entomoloji dergisi 19(4): 305-308. Haupt, H. 1930. Ein neuer Paralimnus Mats. Aus Kleinasien (Homopt. Cicad.). Deutsche Entomologische Zeitschrifft : 207-208. Kalkandelen, A. 1972. Orta Anadolu Cicadellidae: Euscelinae Faunasından Dört Yeni Tür ve Bir Alttür. İstanbul Üniversitesi Fen Fakültesi Mecmuası 37(3-4): 153-163. Kalkandelen, A. 1974. Orta Anadolu'da Homoptera: Cicadellidae familyası türlerinin taksonomileri üzerinde araştırmalar. Zirai Mücadele ve Karantina Genel Müdürlüğü, Ankara. 220 pp. Kalkandelen, A. 1980. Contributions to the families Delphacidae and Cicadellidae (Homoptera) from Turkey. Türkiye Bitki Koruma Dergisi 4(3): 147-154. Kalkandelen, A. 1985. Four new species of genus Zyginidia (Zyginidia) Haut (Homoptera: Cicadellidae) and with notes on the taxonomy and distributions of the species of this genus in Turkey. Türkiye Bitki Koruma Dergisi 9: 13-25. Kartal, V. & Zeybekoğlu, Ü. 1991. Samsun Ondokuz Mayıs Üniversitesi Kurupelit Kampüsü Çevresinde Deltocephalinae (Hom, Auch., Cicadellidae) Alt Familyası Türlerinin Faunistik Yönden İncelenmesi. Ondokuz Mayıs Üniversitesi Fen Dergisi 3(1): 232-257. Kartal, V. & Zeybekoğlu, Ü. 1992. Türkiye Cicadellidae (Hom, Auchenorrhyncha) Faunası İçin Yeni Türler. Turkish Journal of Zoology 16: 349-352. Kartal, V. & Zeybekoğlu, Ü. 1994a. A New Species Allygidius nihati n. sp. (Hom, Auch., Cicadellidae) from Turkey. Turkish Journal of Zoology 18: 259-261. Kartal, V. & Zeybekoğlu, Ü. 1994b. Türkiye Cicadellidae (Hom, Auchenorrhyncha) Faunası İçin Yeni Bir Tür Kaydı. Turkish Journal of Zoology 18: 37-39. Kartal, V. & Zeybekoğlu, Ü. 1997. Türkiye Faunası İçin Yeni Üç Diplocolenus Rib. (Hom, Auch., Cicadellidae, Deltocephalinae) Türü Üzerine Taksonomik Bir Araştırma. Turkish Journal of Zoology 21: 291-294. Kartal, V. 1083. Neue Homopteren aus der Türkei II (Homoptera Auchenorrhyncha). Marburger Entomologische Publikation 1(8): 235-248. Kartal, V. 1981. Neue Homopteren aus der Türkei-I. Priamus 1(1): 24-30. Kartal, V. 1988. Türkiye Cicadellid Faunası İçin Yeni Bir Cins Kazachtanicus Dlab.,1961 (Homoptera, Auchenorrhyncha, Cicadellidae. Doğa Zooloji D. : 68-69. Kartal, V., Zeybekoğlu, Ü. & Dursun, A. 2001. A faunistic study on Deltocephalinae (Homoptera: Auchenorrhyncha: Cicadellidae) in Yüksekova (Hakkari). Türkiye entomoloji dergisi 25(2): 83-92. Lodos, N. & Kalkandelen, A. 1981. Preliminary list of Auchenorrhyncha with notes on distribution and importance of species in Turkey VII. Family Cicadellidae: Ulopinae, Megophthalminae, Ledrinae, Macropsinae and Agallinae. Türkiye Bitki Koruma Dergisi 5(4): 215-230.


Lodos, N. & Kalkandelen, A. 1982a. Preliminary list of Auchenorrhyncha with notes on distribution and importance of species in Turkey VIII. Family Cicadellidae: Idiocerinae. Türkiye Bitki Koruma Dergisi 6(1): 15-28. Lodos, N. & Kalkandelen, A. 1982b. Preliminary list of Auchenorrhyncha with notes on distribution and importance of species in Turkey IX. Family Cicadellidae: Iassinae, Penthiminae, Dorycephalinae, Hecalinae and Aphrodinae. Türkiye Bitki Koruma Dergisi 6: 147-159. Lodos, N. & Kalkandelen, A. 1983a. Preliminary list of Auchenorrhyncha with notes on distribution and importance of species in Turkey X. Family Cicadellidae: Xestocephalinae, Stegelytrinae and Cicadellinae. Türkiye Bitki Koruma Dergisi 7: 23-28. Lodos, N. & Kalkandelen, A. 1983b. Preliminary list of Auchenorrhyncha with notes on distribution and importance of species in Turkey XI. Family Cicadellidae, Typhlocybinae: Alebrini and Dikraneurini. Türkiye Bitki Koruma Dergisi 7: 107-115. Lodos, N. & Kalkandelen, A. 1983c. Preliminary list of Auchenorrhyncha with notes on distribution and importance of species in Turkey XII. Family Cicadellidae, Typhlocybinae: Empoascini. Türkiye Bitki Koruma Dergisi 7: 153-165. Lodos, N. & Kalkandelen, A. 1984a. Preliminary list of Auchenorrhyncha with notes on distribution and importance of species in Turkey XIII. Family Cicadellidae, Typhlocybinae: Typhlocybini (Part I-II). Türkiye Bitki Koruma Dergisi 8: 33-44; 87-97. Lodos, N. & Kalkandelen, A. 1984c. Preliminary list of Auchenorrhyncha with notes on distribution and importance of species in Turkey XV. Family Cicadellidae, Typhlocybinae: Erythroneurini (Part I-II). Türkiye Bitki Koruma Dergisi 8: 159-168; 201-210. Lodos, N. & Kalkandelen, A. 1985a. Preliminary list of Auchenorrhyncha with notes on distribution and importance of species in Turkey XVII. Family Cicadellidae, Deltocephalinae: Grypotini, Goniagnathini and Opsiini (Part I). Türkiye Bitki Koruma Dergisi 9: 79-90. Lodos, N. & Kalkandelen, A. 1985b. Preliminary list of Auchenorrhyncha with notes on distribution and importance of species in Turkey XVIII. Family Cicadellidae, Deltocephalinae: Macrostelini (Part II). Türkiye Bitki Koruma Dergisi 9(3): 147-161. Lodos, N. & Kalkandelen, A. 1985c. Preliminary list of Auchenorrhyncha with notes on distribution and importance of species in Turkey XIX. Family Cicadellidae, Deltocephalinae: Deltocephalini, Scaphytopiini, Doraturini. Türkiye Bitki Koruma Dergisi 9: 207-215. Lodos, N. & Kalkandelen, A. 1986a. Preliminary list of Auchenorrhyncha with notes on distribution and importance of species in Turkey XX. Family Cicadellidae, Deltocephalinae: Fieberiellini, Stirellini and Tetartostylini. Türkiye Bitki Koruma Dergisi 10(1): 25-32. Lodos, N. & Kalkandelen, A. 1986b. Preliminary list of Auchenorrhyncha with notes on distribution and importance of species in Turkey XXI. Family Cicadellidae, Deltocephalinae: Athysanini (Part I-IV). Türkiye Bitki Koruma Dergisi 10 (3-4): 131-139; 203-211, 11 (1-2): 29-40.; 97-109. Lodos, N. & Kalkandelen, A. 1987c. Preliminary list of Auchenorrhyncha with notes on distribution and importance of species in Turkey XXV. Family Cicadellidae, Deltocephalinae: Paralimnini Distans (Part I-II). Türkiye Entomoloji Dergisi 11 (3-4): 151-162; 195-202. Lodos, N. & Kalkandelen, A. 1988. Preliminary list of Auchenorrhyncha with notes on distribution and importance of Turkey XXVII. (Addenda and Corrigenda). Türkiye Entomoloji Dergisi 12(1): 11-22.


Meyer-Arndt, S. 1988. Vier neue Arten der Gattung Fieberiella Signoret, 1879 aus Griechenland und der Türkei (Homoptera Auchenorrhyncha Cicadellidae). Marburger Entomologische Publikation 2(4): 179-208. Zachvatkin, A. A. 1937. Deux Zyginidia nouvellus des Turquie (Homoptera, Eupterygidae). Soc. Nat. Moscou (Sect. Biol.) Bull. 46: 317-322. Zachvatkin, A. A. 1946. Studies on the Homoptera of Turkey. Trans. R. Ent. Soc. London 97(6): 149-176. Zeybekoğlu, Ü. & Bulut, N. 2000. Türkiye Cicadellidae (Homoptera, Auchenorrhyncha) Faunası İçin Yeni İki Tür Kaydı ve Taksonomik Özellikleri. OMÜ Fen-Edebiyat Fakültesi Fen Dergisi 11(1): 85-90. Zeybekoğlu, Ü. 1996. Türkiye’de saptanan Ebarrius Ribaut, 1946 (Homoptera, Auchenorryncha, Cicadellidae, Deltocephalinae) cinsine bağlı türler üzerine sistematik bir araştırma. Turkish Journal of Zoology 20: 187-189. Zeybekoğlu, Ü. 1998. The species of Deltocephalinae (Homoptera: Auchenorrhyncha: Cicadellidae) found in the Middle and East Black Sea Regions. Türkiye Entomoloji Dergisi 22(1): 37-45.

[Received January 2006. Accepted March 2006]

Table 1. New records for Turkey that collected from Antalya province in South Turkey according to the present paper.

Subfamily

New record for Turkey

Macropsinae Macropsis albae Wagner, 1950 Iassinae Iassus pseudocerris Abdul-Nour, 1998

Aphrodinae Anoscopus assimilis (Signoret, 1879) Typhlocybinae Ribautiana cruciata (Ribaut, 1931) Typhlocybinae Eupteryx insulana (Ribaut, 1948)

Deltocephalinae Fieberiella chioscola Meyer-Arndt, 1988 Deltocephalinae Laminacutus libanensis Abdul-Nour, 2001 Deltocephalinae Platymetopius enatus Dlabola, 1974 Deltocephalinae Euscelis alsioides Remane, 1967 Deltocephalinae Paralimnus phragmitis (Boheman, 1847) Deltocephalinae Paralimnellus cingulatus (Dlabola, 1960) Deltocephalinae Psammotettix nodosus (Ribaut, 1925)


A NEW SUBSTITUTE NAME FOR THE GENUS RHABDOTUS TOWNES, 1946 (HYMENOPTERA: ICHNEUMONIDAE)

NON CHAUDOIR, 1865

Hüseyin Özdikmen* and Semra Turgut*

* Gazi Üniversitesi, Fen-Edebiyat Fakültesi, Biyoloji Bölümü, 06500 Ankara / TÜRKİYE, e-mail: [email protected] and [email protected] [Özdikmen, H. & Turgut, S. 2006. A new substitute name for the genus Rhabdotus Townes, 1946 (Hymenoptera: Ichneumonidae) non Chaudoir, 1865. Munis Entomology & Zoology, 1 (2): 237-238] ABSTRACT: A junior homonym was detected amongst the Hymenoptera and the following replacement name is proposed: Oezdemirus for Rhabdotus Townes, 1946 (Ichneumonidae: Ichneumoninae). Accordingly, seven new combinations are herein proposed for all seven valid species currently included in Rhabdotus Townes, 1946 (Ichneumonidae). KEY WORDS: Oezdemirus, Rhabdotus, homonymy, Hymenoptera, Ichneumonidae, replacement name

TAXONOMY

Oezdemirus nom. nov., substitute name

Rhabdotus Townes, 1946. Bol. Ent. venezol., 5, 55. (Insecta: Hymenoptera: Ichneumonoidea: Ichneumonidae: Ichneumoninae: Ichneumonini). Preoccupied by Rhabdotus Chaudoir, 1865. Bull. Soc. imp. Nat. Moscou, 38 (2), 94. (Insecta: Coleoptera: Carabidae).

The genus Rhabdotus was erected by Chaudoir (1865) with the type

species Rhabdotus reflexus Chaudoir, 1865 in Coleoptera (Carabidae). It is still used as a valid generic name in Coleoptera. Also the Neotropical genus Rhabdotus was described by Townes (1946) with the type species Ichneumon intentus Cresson, 1873 in Hymenoptera (Ichneumonoidea: Ichneumonidae: Ichneumoninae: Ichneumonini) from Mexico. It is still a valid generic name in Hymenoptera. However, the name Rhabdotus Townes, 1946 is invalid under the law of homonymy, being a junior homonym of Rhabdotus Chaudoir, 1865. Under the Zoological Code (ICZN, 1999) it must be rejected and replaced. In accordance with article 60 of the International Code of Zoological Nomenclature, fourth edition (1999), we propose to substitute the junior homonym Rhabdotus Townes, 1946 with the nomen novum Oezdemirus. As a result of this, Rhabdotus Townes, 1946 is replaced with Oezdemirus nom. nov. The following new combination is Oezdemirus intentus (Cresson, 1873), comb. nov. along with six other new combinations for all seven valid species currently included in Rhabdotus Townes, 1946 (Hymenoptera: Ichneumonidae).


SYSTEMATIC ACCOUNTS Order Hymenoptera Superfamily Ichneumonoidea Family Ichneumonidae Subfamily Ichneumoninae Genus Oezdemirus new replacement name

For Rhabdotus Townes, 1946, junior homonym of Rhabdotus Chaudoir, 1865. Type species: Ichneumon intentus Cresson, 1873, from Mexico.

Gender: Masculine Etymology: Oezdemirus dedicated to the last names of our colleagues Dr. Yasemin Özdemir, Dr. Işıl Özdemir and Dr. Mustafa Özdemir. Species account and distribution: seven species; known from Mexico and Guatemala in the Neotropical Region (Yu and Horstmann (1997). Mandatory new combinations: Oezdemirus alternans (Brullé, 1846) new comb. from Rhabdotus. Oezdemirus famelicus (Cresson, 1868) new comb. from Rhabdotus. Oezdemirus gracilentissimus (Dalla Torre, 1902) new comb. from Rhabdotus. This name was given as new name for Ichneumon gracilentus Cresson, 1868 preoccupied by Ichneumon gracilentus Wesmael, 1845. Oezdemirus intentus (Cresson, 1873) new comb. from Rhabdotus. Oezdemirus salvini (Cameron, 1885) new comb. from Rhabdotus. Oezdemirus teres (Cresson, 1868) new comb. from Rhabdotus. Oezdemirus trunculentus (Cameron, 1885) new comb. from Rhabdotus.

ACKNOWLEDGEMENTS

Thanks to Dr. Yasemin Özdemir (Nazife Tuatay Plant Protection

Museum, Ankara, Turkey) for references support.

LITERATURE CITED

Chaudoir, M. de. 1865. Essai sur les Féronides de l'Australie et de la Nouvelle-Zélande. Bull. Soc. Imp. Nat. Moscou, 38 (2): 65-112 [94]. International Comission of Zoological Nomenclature. 1999. International Code of Zoological Nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature, London. Townes, H. K. 1946. The generic position of the Neotropic Ichneumonidae with types in the Philadelphia and Quebec museums described by Cresson, Hooker, Norton, Provancher and Viereck. Boletin de Entomologia Venezolana. 5: 29-63 [55]. Yu, D. S. and K. Horstmann. 1997. A catalogue of world Ichneumonidae (Hymenoptera), Part 1: Subfamilies Acaenitinae to Ophioninae, Memoirs of the American Entomological Institute, 58 (1): 1-763 [642]. Yu, D. S. and K. Horstmann. 1997. A catalogue of world Ichneumonidae (Hymenoptera), Part 2: Subfamilies Orthocentrinae to Xoridinae, Memoirs of the American Entomological Institute, 58 (2): 764 -1558.

[Received December 2005. Accepted March 2006]


ADDITIONAL RECORDS FOR THE ODONATA FAUNA OF EAST MEDITERRANEAN REGION OF TURKEY

Ali Salur* and Suat Kıyak**

* Hitit University, Sciences and Arts Faculty, Department of Biology, 19030, Çorum / TURKEY. e-mail: [email protected] ** Gazi Üniversitesi, Fen-Edebiyat Fakültesi, Biyoloji Bölümü, 06500 Ankara/TÜRKİYE, e-mail: [email protected] [Salur, A. & Kıyak, S. 2006. Additional records for the Odonata fauna of East Mediterranean Region of Turkey. Munis Entomology & Zoology 1 (2): 239-252] ABSTRACT: In this study, odonatan specimens collected from the provinces Mersin, Adana, Hatay, Kahramanmaraş and Osmaniye in East Mediterranean region of Turkey, July-September in 2002 and April-August in 2003 have been established faunistically. It has been determined that they belong to 52 species and subspecies of 25 genera of 9 families. It discovered that Ischnura intermedia is a new record for fauna of the research area. KEY WORDS: Odonata, Additional records, East Mediterranean region, Turkey

There have been many studies reported about the Odonata fauna of

Turkey until now. Among these Demirsoy (1982) and Demirsoy (1995) (unpublished project report) were the detailed ones. The research area was studied by Ardıç & Uygun at 1996 as a presentation called “Doğu Akdeniz Bölgesi Odonata faunasının saptanması”. They reported that 43 species belonging to Odonata had been determined. Materials belonging to Ardıç and Uygun are stored at the museum of Çukurova University Faculty of Agriculture, Plant Protection Department. As a result of a careful investigation on the samples of the museum, it mistakes were determined in the identification of some species. Wrong identifications and the correct ones were given at the end of the present study (Table 1).

All samples were collected from the East Mediterranean region of Turkey (Fig. 1) in the years of 2002-2003 and they were collected by using a net trap and having been killed in the killing jars vapour of ethyl acetate. These samples are deposited in Zoological Museum of Gazi University (=ZMGU).

Family CALOPTERYGIDAE

Calopteryx splendens splendens (Harris, 1782) Materials: Kahramanmaraş: 1 male, Tanır (Hurman stream), 38º25’N 36º55’E, 1248 m, 26.07.2003.

Calopteryx splendens intermedia Sélys, 1887 Materials: Adana: 1 female, Karaisalı (Salbaş village, Üçürge brook), 37º08’N 35º08’E, 98 m, 18.07.2002; 1 male, Karaisalı (Topaktaş, Körkün stream), 37º09’N 35º09’E, 104 m, 18.07.2002; 2 males, 1 female, Feke (Tufanbeyli road, Göksu river), 37º51’N 35º58’E, 661 m, 19.08.2002; 3 males, Saimbeyli (Tırtat, Göksu river), 37º52’N 36º00’E, 635 m, 19.08.2002; 1 male, Aladağ (Eyner, Small stream), 37º24’N 35º27’E, 250 m, 18.05.2003; 1 male, 2 females, Ceyhan (Karaçay), 37º03’N 35º57’E, 44 m, 16.05.2003; 1 male, Karaisalı (Kuzgun village), 37º09’N 35º05’E, 162 m, 18.05.2003; 4 males, 2 females, Karaisalı (Güvenç village), 37º12’N 35º05’E, 170 m, 18.05.2003; 1 male Karaisalı (Hacılı, Körkün stream), 37º17’N 35º19’E, 278 m, 18.05.2003; 1 male, Aladağ (Kahyalar), 37º17’N 35º23’E,


208 m, 18.05.2003; 1 male Aladağ (Eyner, Small stream), 37º24’N 35º27’E, 250 m, 18.05.2003; 1 male, Aladağ (Boztahta, Simit bridge), 37º23’N 35º26’E, 150 m, 05.06.2003; 1 male, Aladağ (Seyhan), 37º22’N 35º28’E, 158 m, 05.06.2003; 1 male, İmamoğlu-Kozan road (Tırmıl brook), 37º20’N 35º45’E, 85 m, 05.06.2003; 1 male, Kozan (Horzum, Tersakan-Pınargözü brook), 37º39’N 35º51’E, 600 m, 05.06.2003; 1 male, Feke (Göksun bridge), 37º42’N 35º52’E, 05.06.2003; 1 male, Kozan (Horzum, Tersakan (Pınargözü) brook), 37º39’N 35º51’E, 600 m, 05.06.2003; 1 male, Kozan (Kayhan), 37º22’N 35º50’E, 82 m, 06.06.2003; 1 male, Tufanbeyli (Güzelim village, Güzelim brook), 38º07’N 36º10’E, 1430 m, 10.06.2003; 1 female, Adana, Aladağ (Büyüksofulu), 37º32’N 35º09’E, 940 m, 29.07.2003; Hatay: 1 male, 1 female, İskenderun (Arsuz), 36º09’N 36º01’E, 520 m, 07.06.2003; 1 male, Samandağ (Karaçay, Büyükdere), 36º08’N 36º01’E, 500 m, 07.06.2003; 1 male, Narlıca, 36º15’N 36º15’E, 93 m, 08.06.2003; 3 males, Kırıkhan’ın kuzeyi (Kamışlar village), 36º38’N 36º26’E, 105 m, 08.06.2003; 1 male, Hassa (Akbez), 36º50’N 36º32’E, 100 m, 08.06.2003; 2 males, Hassa (Akbez), 36º50’N 36º37’E, 105 m, 08.06.2003; 4 males, 1 female, Yayladağ (Çabala-Kızılçad road), 35º52’N 36º06’E, 440 m, 09.06.2003; 4 males, Yayladağ (Dağardı road), 35º52’N 36º06’E, 440 m, 09.06.2003; 4 males, Yayladağ (Dervent brook kolu), 35º53’N 36º01’E, 400 m, 09.06.2003; 4 males, Yayladağ,, 35º53’N 36º00’E, 450 m, 09.06.2003; Kahramanmaraş: 4 males, 2 females, Andırın (Çatak mevkii, Andırın brook), 37º30’N 36º21’E, 545 m, 06.06.2003; 2 males, 1 female, Andırın (Çınar geçidi), 37º36’N 36º20’E, 1080 m, 06.06.2003; 3 females, Afşin (Yazdere, Hurman stream), 38º09’N 37º00’E, 1320 m, 10.06.2003; Mersin: 2 males, Tarsus (Keşbükü, Tarsus stream), 37º01’N 34º45’E, 139 m, 17.07.2002; 1 female, Silifke (Kapızlı), 36º23’N 34º02’E, 22 m, 19.05.2003; 1 male, eastern of Bozyazı, 36º06’N 33º00’E, 22 m, 03.06.2003; 1 male Bozyazı (Gözce), 36º09’N 33º10’E, 40 m, 03.06.2003; 2 males, 1 female, Bozyazı (between Gözce-Gözsüzce), 36º11’N 33º12’E, 115 m, 03.06.2003; 1 male, 2 females, Mut (Kışlaköy, Göksu river), 36º29’N 33º22’E, 200 m, 03.06.2003; 4 males, 1 female, Silifke (Kapızlı), 36º23’N 34º02’E, 24 m, 20.04.2003; 1 female, Tarsus (Keşbükü, Tarsus stream), 37º01’N 34º45’E, 139 m, 31.07.2003; Osmaniye: 1 male, Kadirli (Akova), 37º20’N 35º57’E, 47 m, 06.06.2003.

Calopteryx virgo festiva (Brullé, 1832) Materials: Adana: 2 females, Saimbeyli (Saimbeyli brook, Ayçukuru), 37º57’N 36º04’E, 850 m, 19.08.2002; 1 female, Kozan (Horzum, Tersakan-Pınargözü brook), 37º39’N 35º51’E, 600 m, 05.06.2003; 2 males, Aladağ (Büyüksofulu), 37º32’N 35º09’E, 940 m, 29.07.2003; Hatay: 1 male, Erzin (Başlamış village, Erzin ılıcası), 36º57’N 36º15’E, 452 m, 19.07.2002; Kahramanmaraş: 1 female, Andırın (Çatak, Andırın brook), 37º30’N 36º21’E, 545 m, 06.06.2003; 1 male, Andırın (Kesik), 37º25’N 36º20’E, 1127 m, 06.06.2003; 1 male, 2 females, Andırın (Pakmaksızlar village), 37º39’N 36º21’E, 1183 m, 06.06.2003; 1 male, Suçatı, 37º45’N 36º41’E, 656 m, 10.06.2003; Mersin: 1 male, Anamur (Ormancık village, Alaköprü brook), 36º10’N 32º53’E, 107 m, 27.06.2002; Osmaniye: 2 males, Yarpuz (Haraz plateau), 29.07.2003; 4 males, Yarpuz Plateau, 29.07.2003.

Family EUPHAEIDAE Epallage fatime (Charpentier, 1840)

Materials: Adana: 1 male, 2 females, Pozantı, Gerdibi village civarı, 37º29’N 35º08’E, 954 m, 21.06.2002; 2 males, 1 female, Karaisalı (Güvenç village), 37º12’N 35º05’E, 170 m, 18.05.2003; 1 male, Karaisalı (Ömerli Village), 37º17’N 35º15’E, 191 m, 18.05.2003; 1 female, Aladağ (Kahyalar), 37º17’N 35º23’E, 208 m, 18.05.2003; 1 female, Aladağ (Topallı village, Çatalan dam), 37º19’N 35º23’E, 124 m, 18.05.2003; 1 male, Aladağ (Eyner, Small stream), 37º24’N 35º27’E, 250 m, 18.05.2003; 1 male, Aladağ (Boztahta, Simit bridge), 37º23’N 35º26’E, 150 m, 05.06.2003; 1 male, 1 female, Kozan (Kozan dam), 37º33’N 35º49’E, 340 m, 05.06.2003; 1 male, 1 female, Kozan (Horzum, Tersakan -Pınargözü brook), 37º39’N 35º51’E, 600 m, 05.06.2003; 1 male, 1 female, Feke (Göksun bridge), 37º42’N 35º52’E, 05.06.2003; 1 male, Aladağ (Kıcak), 37º35’N 35º14’E, 1075 m, 29.07.2003; Hatay: 1 female, Yayladağ (Çalı boğazı), 35º52’N 36º05’E, 496 m, 09.06.2003; 3 males, 1 female, Yayladağ (Çabala-Kızılçad road), 35º52’N 36º06’E, 440 m, 09.06.2003; 3 males, Hatay, Yayladağ (Dağardı road), 35º52’N 36º06’E, 440 m, 09.06.2003; 3 males, Yayladağ (race of Dervent brook), 35º53’N 36º01’E, 400 m, 09.06.2003; 3 males, Yayladağ, 35º53’N 36º00’E, 450 m, 09.06.2003; 1 female, Samandağ (between Tomruk-Çevlik),


36º15’N 35º49’E, 35ºm, 16.04.2003; 1 female, Samandağ (between Tomruk-Çevlik) 36º14’N 35º50’E, 34 m, 16.04.2003; Kahramanmaraş: 1 female, Andırın (Kesim), 37º25’N 36º14’E, 200 m, 19.07.2002; 3 males, 1 female, Andırın (Çatak, Andırın brook), 37º30’N 36º21’E, 545 m, 06.06.2003; 1 female, Andırın (Çınar geçidi), 37º36’N 36º20’E, 1080 m, 06.06.2003; 3 females, Bulutoğlu, 37º38’N 36º41’E, 520 m,10.06.2003; Mersin: 1 male, 1 female, Tarsus (Sandal village, Söğütlü plateau), 565 m, 37º09’N 34º48’E, 17.07.2002; 1 male, Bozyazı (Gözce), 36º09’N 33º10’E, 40 m, 03.06.2003; 6 males, 2 females, Bozyazı (Gözsüzce), 36º11’N 33º12’E, 115 m, 03.06.2003; 2 males, Aydıncık (Gülnar road), 36º10’N 33º21’E, 30 m, 03.06.2003; 1 female, Aydıncık (Bozağaç-Gülnar road, Menekşe brook), 36º16’N 33º23’E 680 m, 03.06.2003; 1 male, Mut (Kışlaköy, Göksu river), 36º29’N 33º22’E, 200 m, 03.06.2003; 2 males, Çamlıyayla (Kurtçukuru), 37º10’N 34º45’E, 581 m, 30.07.2003; Osmaniye: 2 males, Kadirli (Akova), 37º20’N 35º57’E, 47 m, 06.06.2003.

Family LESTIDAE

Lestes macrostigma (Eversmann, 1836) Materials: Adana: 5 females, Karataş (Bahçeköy), 36º35’N 35º25’E, 2 m, 17.05.2003; 3 females, Karataş (Bahçeköy), 36º35’N 35º27’E, 2 m, 17.05.2003; Hatay: 1 male, 1 female, İskenderun (Arsuz), 36º29’N 35º59’E, 5 m, 15.05.2003; 1 male, 3 females, Antakya (Campus of Mustafa Kemal University), 36º19’N 36º12’E, 120 m, 16.05.2003; Mersin: 1 male, 1 female, Mut (Kargıcak), 36º25’N 33º40’E, 320 m, 04.06.2003.

Lestes viridis parvidens Artobolevski, 1929 Materials: Adana: 1 male, 1 female, Aladağ (Seyhan), 37º22’N 35º28’E, 158 m, 05.06.2003; Mersin: 1 male, 3 females, Bozyazı (Gözce), 36º09’N 33º10’E, 40 m, 03.06.2003.

Sympecma fusca (Vander Linden, 1823) Materials: Adana: 1 male, Pozantı (Çamlıbel village), 37º28’N 35º06’E, 987 m, 21.06.2002.

Family PLATYCNEMIDIDAE Platycnemis dealbata Klug, 1863

Materials: Adana: 4 males, Seyhan river, 36º59’N 35º10’E, 34 m, 18.07.2002; 2 males, 2 females, Karaisalı road (Seyhan river-canal), 37º00’N 35º10’E, 47 m, 18.07.2002; 4 males, Karaisalı (Salbaş village, Çakıt brook), 37º06’N 35º06’E, 84 m, 18.07.2002; 3 males, Karaisalı (Salbaş village, Üçürge brook), 37º08’N 35º08’E, 98 m, 18.07.2002; 7 males, 5 females, Karaisalı (Topaktaş, Körkün stream), 37º09’N 35º09’E, 104 m, 18.07.2002; 3 males, İmamoğlu road, 37º03’N 35º26’E, 115 m, 18.07.2002; 4 males, 2 females, İmamoğlu (Hakkıbeyli, Han brook), 37º10’N 35º33’E, 187 m, 18.07.2002; 1 male, 1 female, İmamoğlu (İncedere), 37º14’N 35º38’E, 95 m, 18.07.2002; 1 male, 1 female, İmamoğlu-Kozan road (Kurukulak brook), 37º17’N 35º42’E, 47 m, 18.07.2002; 2 males, Kozan (Aslanlı, Tırmıl brook), 37º20’N 35º45’E, 74 m, 18.07.2002; 1 male, Kozan (Deliçay), 37º34’N 35º47’E, 119 m, 18.07.2002; 1 male, Kozan (Kayhan), 37º22’N 35º49’E, 84 m, 19.07.2002; 2 males, 4 females, Ceyhan road (Karaçay), 37º03’N 37º57’E, 35ºm, 19.07.2002; 1 male, 2 females, Ceyhan (Ceyhan river), 37º03’N 35º48’E, 34 m, 19.07.2002; 5 males, 3 females, Karataş (Yassıören, Seyhan canal), 36º43’N 35º11’E, 12 m, 19.04.2003; 1 female, Ceyhan (Karaçay), 37º03’N 35º57’E, 44 m, 16.05.2003; 1 male, Karataş (Topraklı), 36º39’N 35º20’E, 10 m, 17.05.2003; 5 males, 5 females, Karaisalı (Güvenç village), 37º12’N 35º05’E, 170 m, 18.05.2003; 1 female, Karaisalı (Ömerli village, Çatalan dam), 37º17’N 35º21’E, 190 m, 18.05.2003; 1 female, İmamoğlu (İncedere), 37º14’N 35º37’E, 135 m, 05.06.2003; 1 male, Kozan (Kayhan), 37º22’N 35º50’E, 82 m, 06.06.2003; 1 male, 2 females, İmamoğlu-Kozan road (Kurukulak brook), 37º17’N 35º42’E, 66 m, 05.06.2003; 2 males, İmamoğlu-Kozan road (Tırmıl brook), 37º20’N 35º45’E, 85 m, 05.06.2003;1 male, Aladağ (Gedikli), 37º29’N 35º09’E, 920 m, 29.07.2003; Hatay: 1 male, 1 female, İskenderun (Sarıseki, Sarıseki brook), 36º40’N 36º13’E, 9 m, 20.07.2002; 5 females, Reyhanlı (Antakya road), 36º14’N 36º23’E, 96 m, 20.07.2002; 1 male, Antakya (Nallıca, Asi river), 36º15’N 36º14’E, 100 m, 20.07.2002; 2 females, Kırıkhan (Hassa road 4.km), 36º32’N 36º23’E, 113 m, 21.07.2002; 3 males, Kırıkhan (Hassa road, Bulgur spring), 36º37’N 36º26’E, 165 m, 21.07.2002; 4 males, 1 female, Kırıkhan (Gölbaşı), 36º30’N 36º28’E, 95 m, 16.05.2003; 4 males, 1 female,


Kırıkhan’ın kuzeyi, 36º34’N 36º24’E, 113 m, 16.05.2003; 2 males, 2 females, İskenderun (Büyükdere), 36º33’N 36º07’E, 10 m, 15.05.2003; 6 males, 1 female, İskenderun (Sarıseki), 36º40’N 36º13’E, 26 m, 07.06.2003; 3 males, İskenderun (Arsuz), 36º09’N 36º01’E, 520 m, 07.06.2003; 1 male, Samandağ (Karaçay, Büyükdere), 36º08’N 36º01’E, 500 m, 07.06.2003; 1 male, Antakya (Demirköprü, Asi river), 36º14’N 36º21’E, 102 m, 08.06.2003; 2 males, Reyhanlı (Müşrüfe, Afrin river), 36º19’N 36º32’E, 112 m, 08.06.2003; 2 males, Kırıkhan (Büyükkarasu brook), 36º30’N 36º24’E, 100 m, 08.06.2003; 1 male, 1 female, Kırıkhan’ın kuzeyi 36º34’N 36º23’E, 105 m, 08.06.2003; 1 male, northern of Kırıkhan (Kamışlar village), 36º38’N 36º26’E, 105 m, 08.06.2003; 1 male, 1 female, Hassa (Akbez), 36º50’N 36º32’E, 100 m, 08.06.2003; 1 male, Hassa (Akbez), 36º50’N 36º37’E, 105 m, 08.06.2003; Kahramanmaraş: 1 male, Andırın (Gökahmetler, Aslantaş dam), 37º24’N 36º16’E, 250 m, 06.06.2003; 1 male, 1 female, Menzelet Dam, 37º45’N 36º42’E, 675 m, 10.06.2003; 1 male, Ilıca (Menzelet dam), 37º49’N 36º48’E, 690 m, 26.07.2003; 1 male, Elbistan (Afşin road, DSİ regulator), 38º11’N 37º05’E, 1138 m, 26.07.2003; Osmaniye: 1 male, Çukurköprü, 37º20’N 35º55’E, 52 m, 19.07.2002; 1 male, Kadirli (between Çukurköprü-Aydınlar), 37º20’N 35º56’E, 51 m, 19.07.2002; 1 male, Kadirli (Yalnızdut), 37º17’N 36º02’E, 62 m, 19.07.2002; 2 males, 2 females, Kadirli (Karabacak), 37º14’N 36º02’E, 53 m, 19.07.2002; 1 male, Kadirli (Orhaniye, Seyhan brook), 37º12’N 36º04’E, 48 m, 19.07.2002; 3 males, 2 females, Toprakkale (Yolçatı brook), 37º03’N 36º06’E, 55 m, 19.07.2002.

Platycnemis kervillei (Martin, 1909) Materials: Adana: 1 female, Karataş cape (dalyan), 36º35’N 35º19’E, 10 m, 17.05.2003; 1 male, 1 female, Aladağ (Kahyalar), 37º17’N 35º23’E, 208 m, 18.05.2003; 2 males, 1 female, Aladağ (Boztahta, Simit bridge), 37º23’N 35º26’E, 150 m, 05.06.2003; 2 males, İmamoğlu-Kozan road (Kurukulak brook), 37º17’N 35º42’E, 66 m, 05.06.2003; 1 female, Kozan (Horzum, Tersakan -Pınargözü brook), 37º39’N 35º51’E, 600 m, 05.06.2003; HATAY: 1 male, 1 female, Yayladağ (Çabala-Kızılçad road), 35º52’N 36º06’E, 394 m, 15.05.2003; 1 female, Yayladağ (Kureyşi brook), 35º54’N 36º03’E, 394 m, 15.05.2003; 1 male, 2 females, Yayladağ (race of Dervent brook), 35º53’N 36º01’E, 400 m, 15.05.2003; 1 male, Yayladağ (Çalı boğazı), 35º52’N 36º05’E, 496 m, 09.06.2003; 5 males, 2 females, Yayladağ (Çabala-Kızılçad road), 35º52’N 36º06’E, 440 m, 09.06.2003; 3 males, 2 females, Yayladağ (Dağardı road), 35º52’N 36º06’E, 440 m, 09.06.2003; 1 male, 1 female, Yayladağ (race of Dervent brook), 35º53’N 36º01’E, 400 m, 09.06.2003; 2 males, 1 female, Yayladağ, 35º53’N 36º00’E, 450 m, 09.06.2003; Kahramanmaraş 1 male, 1 female, Andırın (Çatak, Andırın brook), 37º30’N 36º21’E, 545 m, 06.06.2003; 1 female, Andırın (Kesik), 37º25’N 36º20’E, 1127 m, 06.06.2003.

Platycnemis pennipes (Pallas, 1771) Materials: Adana: 1 male, 1 female, Pozantı (Gerdibi village), 37º29’N 35º08’E, 954 m, 21.06.2002; 12 males, 3 females, Tufanbeyli (between Karsavuran-Pınarlar, İncedere), 38º13’N 36º13’E, 1400 m, 19.08.2002; 1 female, İmamoğlu (İncedere), 37º14’N 35º37’E, 135 m, 05.06.2003; 4 females, Tufanbeyli (Güzelim village, Güzelim brook), 38º07’N 36º10’E, 1430 m, 10.06.2003; Kahramanmaraş: 1 female, Suçatı (Menzelet dam), 37º45’N 36º41’E, 656 m, 10.06.2003; 2 males, 2 females, Menzelet dam, 675 m, 37º45’N 36º42’E, 10.06.2003; 1 female, Göksun (Çardak), 36º06’N 36º51’E, 1329 m, 10.06.2003; 1 female, Afşin (Yazdere, Hurman stream), 38º09’N 37º00’E, 1320 m, 10.06.2003; 1 male, Çağlayancerit (Başdevrişli), 37º42’N 37º10’E, 1260 m, 27.07.2003; Mersin: 1 female, Silifke (Kapızlı), 36º23’N 34º02’E, 22 m, 19.05.2003; 4 males, 1 female, Bozyazı, 36º06’N 32º57’E, 10 m, 03.06.2003; 1 female, eastern of Bozyazı, 36º06’N 33º00’E, 22 m, 03.06.2003; 1 male, Bozyazı (between Gözce-Gözsüzce), 36º11’N 33º12’E, 115 m, 03.06.2003; 1 male, Aydıncık (Yenikaş, Soğuksu brook), 36º08’N 33º17’E, 2 m, 03.06.2003; 1 male, 1 female, Aydıncık (Gülnar road), 36º10’N 33º21’E, 30 m, 03.06.2003; 1 male, Tarsus (Akçakocalı, Berdan dam), 36º57’N 34º50’E, 75 m, 04.06.2003.

Family COENAGRIONIDAE Ischnura elegans ebneri Schmidt, 1938

Materials: Adana: 4 males, 1 female, Karaisalı (Seyhan river), 36º59’N 35º10’E, 34 m, 18.07.2002; 2 males, 2 females, Karaisalı road (Seyhan river-canal), 37º00’N 35º10’E, 47 m, 18.07.2002; 1 male, İmamoğlu (İncedere), 37º14’N 35º38’E, 95 m, 18.07.2002; 1 male,


Kozan (Kayhan), 37º22’N 35º49’E, 84 m, 19.07.2002; 1 female, Tufanbeyli (between Karsavuran-Pınarlar, İncedere), 38º13’N 36º13’E, 1400 m, 19.08.2002; 2 males, Zeytinli (canal), 36º59’N 35º07’E, 25 m, 18.07.2002; 3 males, Ceyhan (Karaçay), 37º03’N 35º57’E, 44 m, 16.05.2003; 2 males, 2 females, Ceyhan (Ceyhan river), 37º03’N 35º48’E, 35ºm, 16.05.2003; 1 male, 1 female, Karataş (Topraklı, Dalyan-Ilgınlık), 36º39’N 35º20’E, 10 m, 17.05.2003; 1 male, 1 female, Karataş (Bahçeköy), 36º35’N 35º25’E, 2 m, 17.05.2003; 1 male, 1 female, Karataş (Bahçeköy), 36º35’N 35º27’E, 2 m, 17.05.2003; 1 male, 2 females, Karataş cape (dalyan), 36º35’N 35º19’E, 10 m, 17.05.2003; 1 male, 1 female, Karaisalı (Abacı, Seyhan dam), 37º09’N 35º11’E, 150 m, 18.05.2003; 1 male, 1 female, Karaisalı (Güvenç village), 37º12’N 35º05’E, 170 m, 18.05.2003; 1 male, 1 female, Karaisalı (Ömerli village, Çatalan dam), 37º17’N 35º21’E, 190 m, 18.05.2003; 1 female, Aladağ (Topallı village, Çatalan dam), 37º19’N 35º23’E, 124 m, 18.05.2003; 1 male, Kozan (Kayhan), 37º22’N 35º50’E, 82 m, 06.06.2003; 17 males, 9 females, Karataş (Yassıören, Seyhan canal), 36º43’N 35º11’E, 12 m, 19.04.2003; 8 males, 6 females, Karataş (Karagöçer), 36º42’N 35º05’E, 13 m, 19.04.2003; 1 male, 1 female, Karataş (Tuz lake), 36º43’N 35º02’E, 6 m, 19.04.2003; 10 males, 8 females, Karataş (Tabaklar village), 36º44’N 35º00’E, 10 m, 19.04.2003; 2 males, 1 female, Karataş (Konaklı village), 36º39’N 35º09’E, 12 m, 19.04.2003; 9 males, Karataş (Gölkaya village), 36º36’N 35º23’E, 17 m, 19.04.2003; 6 males, 2 females, Karataş (Ömer lake), 36º42’N 35º33’E, 19 m, 19.04.2003; HATAY: 1 male, 2 females,İskenderun, Sarıseki (Sarıseki brook), 36º40’N 36º13’E, 17 m, 20.07.2002; 2 males, İskenderun (Güzin brook), 36º34’N 36º09’E, 13 m, 20.07.2002; 2 males, 1 female, Kırıkhan, 36º28’N 36º21’E, 112 m, 20.07.2002; 1 male, Kırıkhan (Kumlu road), 36º27’N 36º22’E, 152 m, 20.07.2002; 1 male, Reyhanlı (Antakya road), 36º14’N 36º23’E, 94 m, 20.07.2002; 2 males, Samandağ (Asi river), 36º04’N 36º00’E, 100 m, 21.07.2002; 3 males, Hassa (Akbez, Kilis il sınırı), 36º50’N 36º37’E, 384 m, 21.07.2002; 1 male, İskenderun (Sarıseki), 36º40’N 36º13’E, 26 m, 15.05.2003; 3 males, İskenderun (Arsuz), 36º29’N 35º59’E, 5 m, 15.05.2003; 1 male, İskenderun (Arsuz, Arsuz stream), 36º24’N 35º53’E, 10 m, 15.05.2003; 2 males, 1 female, Antakya (Campus of Mustafa Kemal University), 36º19’N 36º12’E, 120 m, 16.05.2003; 2 males, 4 females, Kırıkhan (Gölbaşı), 36º30’N 36º28’E, 95 m, 16.05.2003; 1 male, 1 female, northern of Kırıkhan, 36º34’N 36º24’E, 113 m, 16.05.2003; 1 male, Samandağ (Karaçay, Küçükdere), 36º09’N 36º01’E, 520 m, 07.06.2003; 1 male, Antakya (Demirköprü, Asi river), 36º14’N 36º21’E, 102 m, 08.06.2003; 2 males, 1 female, Reyhanlı, 36º14’N 36º34’E, 187 m, 08.06.2003; 1 male, Reyhanlı (Müşrüfe, Afrin river), 36º19’N 36º32’E, 112 m, 08.06.2003; 1 male, Kırıkhan (Gölbaşı), 36º30’N 36º28’E, 100 m, 08.06.2003; 2 males, 2 females, Antakya (Campus of Mustafa Kemal University), 36º19’N 36º12’E, 120 m, 17.04.2003; 2 males, 2 females, Kumlu (Karasüleymanlı, canal III), 36º23’N 36º24’E, 91 m, 18.04.2003; 3 males, 2 females, Kırıkhan (Gölbaşı village), 36º30’N 36º28’E, 95 m, 18.04.2003; Kahramanmaraş: 2 females, (Hasancıklı, stream of between Menzelet dam and Sır dam), 37º36’N 36º47’E, 470 m, 10.06.2003; 1 male, Menzelet dam, 675 m, 37º45’N 36º42’E, 10.06.2003; 1 male, Elbistan (Afşin road, DSİ Regulator), 38º11’N 37º05’E, 1138 m, 26.07.2003; 2 males, Menzelet Dam, 37º45’N 36º45’E, 645 m, 26.07.2003; Mersin: 2 males, 2 females, Tarsus, 36º55’N 34º55’E, 15 m, 20.04.2003; 1 male, 1 female, Tarsus (Berdan stream), 36º54’N 34º55’E, 23 m, 20.04.2003; 1 male, 1 female, Deliçay, 36º50’N 34º42’E, 28 m, 20.04.2003; 1 male, 1 female, Yenişehir (near harbour), 36º47’N 34º36’E, 10 m, 20.04.2003; 2 males, Erdemli (Arpaçbahşiş), 36º38’N 34º21’E, 18 m, 20.04.2003; 1 male, 1 female, Silifke (Kapızlı), 36º23’N 34º02’E, 24 m, 20.04.2003; 1 male, 1 female, Erdemli (Limonlu stream), 36º32’N 34º13’E, 7 m, 19.05.2003; 1 female, Bozyazı, 36º06’N 32º57’E, 10 m, 03.06.2003; 2 males, Silifke (Kapızlı), 36º23’N 34º02’E, 22 m, 31.07.2003; 1 male, Centrum, 36º46’N 34º36’E, 10 m, 31.07.2003; Osmaniye: 1 male, Kadirli (Yalnızdut), 37º17’N 36º02’E, 62 m, 19.07.2002; 1 male, 1 female, Kadirli (Koçyurdu), 37º13’N 36º02’E, 50 m, 19.07.2002; 1 male, Kadirli (Orhaniye, Seyhan brook), 37º12’N 36º04’E, 48 m, 19.07.2002; 1 male, Toprakkale (Karaçay), 37º03’N 36º12’E, 114 m, 19.07.2002; 1 male, 2 females, near Centrum, 37º06’N 36º18’E, 140 m, 16.05.2003; 1 male, Kadirli (Akova), 37º20’N 35º57’E, 47 m, 06.06.2003.

Ischnura intermedia Dumont, 1974 Materials: Adana: 1 male, Aladağ (Büyüksofulu), 37º33’N 35º11’E, 1020 m, 29.07.2003. Remark: This species is new for east Mediterranean region of Turkey.


Erythromma viridulum Charpentier, 1840 Materials: Hatay: 1 male, Kırıkhan (Gölbaşı village), 36º30’N 36º28’E, 95 m, 16.05.2003.

Coenagrion puella (Linnaeus, 1758) Materials: Kahramanmaraş: 4 males, Göksun (Çardak), 36º06’N 36º51’E, 1329 m, 10.06.2003; 3 males, Göksun (Terbüzek I), 38º04’N 36º27’E, 1390 m, 10.06.2003.

Coenagrion syriaca (Morton, 1924) Materials: Adana: 1 male, Kozan (Kozan dam), 37º31’N 35º50’E, 300 m, 05.06.2003; Hatay: 1 male, Kırıkhan (Gölbaşı village), 36º30’N 36º28’E, 95 m, 18.04.2003; 5 males, 2 females, İskenderun (Sarıseki), 36º40’N 36º13’E, 26 m, 15.05.2003; Kahramanmaraş: 2 males, Afşin (Büyükkızılcık village, Kızılcık brook), 38º09’N 36º42’E, 1400 m, 10.06.2003; Mersin: 1 male, 1 female, Silifke (Kapızlı), 36º23’N 34º02’E, 24 m, 20.04.2003; Osmaniye: 1 male, Kadirli (Akova), 37º20’N 35º57’E, 47 m, 06.06.2003.

Cercion lindeni (Sélys, 1840) Materials: Hatay: 1 male, Kırıkhan (Gölbaşı), 36º30’N 36º28’E, 95 m, 16.05.2003; 1 male, Kırıkhan (Gölbaşı), 36º30’N 36º28’E, 100 m, 08.06.2003; 1 male, Kırıkhan (Gölbaşı village), 36º30’N 36º28’E, 95 m, 18.04.2003.

Ceriagrion georgifreyi (Schmidt, 1953) Materials: Hatay: 1 male, İskenderun (Sarıseki), 36º40’N 36º13’E, 26 m, 15.05.2003; Mersin: 1 male, 1 female, Bozyazı, 36º06’N 32º57’E, 10 m, 03.06.2003; 1 female, Aydıncık (Gülnar road), 36º10’N 33º21’E, 30 m, 03.06.2003.

Family AESHNIDAE Anax immaculifrons Rambur, 1842

Materials: Adana: 1 male , Kozan (Kozan dam), 37º31’N 35º50’E, 300 m, 05.06.2003; 2 males, Aladağ (Simit bridge), 37º23’N 35º26’E, 150 m, 29.07.2003.

Anax imperator Leach, 1815 Materials: Adana: 1 male, Karaisalı (Çatalan dam), 37º11’N 35º16’E, 72 m,18.07.2002; 1 male, Tufanbeyli (Güzelim, race of Sarız stream), 38º08’N 36º10’E, 1390 m, 19.08.2002; 1 female Karataş cape (dalyan), 36º35’N 35º19’E, 10 m, 17.05.2003 ; 1 female, Karaisalı (Abacı, Seyhan dam), 37º09’N 35º11’E, 150 m, 18.05.2003; 1 female, Kozan (Kozan dam), 37º31’N 35º50’E, 300 m, 05.06.2003; Hatay: 1 male, Antakya (Campus of Mustafa Kemal University), 36º19’N 36º12’E, 120 m, 16.05.2003; 1 male, Kırıkhan (Gölbaşı), 36º30’N 36º28’E, 95 m, 16.05.2003; 1 male, Kırıkhan’ın kuzeyi, 36º34’N 36º24’E, 113 m, 16.05.2003; 1 male, Reyhanlı, 36º14’N 36º34’E, 187 m, 08.06.2003; Kahramanmaraş: 1 male, Andırın (between Yeniköy-Kesim), 37º25’N 36º14’E, 200 m, 19.07.2002; 1 male, Afşin (Alimpınar), 38º12’N 36º53’E, 1285 m, 10.06.2003; Mersin: 1 male, Tece (Tece Stream), 36º41’N 34º27’E, 24 m, 31.07.2003; Osmaniye: 1 male , near Centrum, 37º06’N 36º18’E, 140 m, 16.05.2003.

Anax parthenope (Selys, 1839) Materials: Hatay: 1 male , Kırıkhan (Gölbaşı), 36º30’N 36º28’E, 100 m, 08.06.2003; 1 male, Hassa (Akbez), 36º50’N 36º37’E, 105 m, 08.06.2003; 2 females, Samandağ (between Tomruk-Çevlik), 36º16’N 35º48’E, 25 m, 16.04.2003; 1 male, Yayladağ (race of Dervent brook), 400 m, 17.04.2003; 1 male, Kırıkhan (Gölbaşı), 36º30’N 36º28’E, 100 m, 08.06.2003; Kahramanmaraş: 1 male, Andırın (Gökahmetler, Aslantaş dam), 37º24’N 36º16’E, 250 m, 06.06.2003; Mersin: 1 male, Tarsus (Akçakocalı, Berdan dam), 36º57’N 34º50’E, 75 m, 04.06.2003; Osmaniye: 1 male , Kadirli (Yalnızdut), 37º17’N 36º02’E, 62 m, 19.07.2002.

Hemianax ephippiger (Burmeister, 1839) Materials: Hatay: 1 male, Kırıkhan (Gölbaşı village), 36º30’N 36º28’E, 95 m, 18.04.2003.

Caliaeschna microstigma (Schneider, 1845) Materials: Adana: 1 male, Pozantı, Dağdibi village, 37º39’N 34º59’E, 1239 m, 20.06.2002; 1 male, Adana, Aladağ (Eyner), 37º25’N 35º26’E, 200 m, 05.06.2003; 1 male, Aladağ (Seyhan), 37º22’N 35º28’E, 158 m, 05.06.2003; 1 male, Kozan (Horzum, Tersakan -Pınargözü brook), 37º39’N 35º51’E, 600 m, 05.06.2003; 1 male, Aladağ (Gedikli), 37º29’N 35º09’E, 920 m, 29.07.2003; 1 male, Aladağ (Büyüksofulu), 37º32’N 35º09’E, 940 m, 29.07.2003; Hatay: 1 male, Erzin (Başlamış village, Erzin ılıcası), 36º57’N 36º15’E, 452 m,


19.07.2002; 1 male, 1 female, Yayladağ (Çabala-Kızılçad road), 35º52’N 36º06’E, 440 m, 09.06.2003; Kahramanmaraş: 1 male, Andırın (Pakmaksızlar village), 37º39’N 36º21’E, 1183 m, 06.06.2003; 1 male, Çağlayancerit (Yenipınar), 37º42’N 37º09’E, 1278 m, 27.07.2003; Mersin: 3 males, Mut (Şarlak), 36º27’N 33º22’E, 243 m, 03.06.2003; 1 female, Sarıkavak, 36º36’N 33º38’E, 460 m, 03.06.2003; 1 male, Çamlıyayla (Papazın Bahçesi), 37º14’N 34º37’E, 761 m, 30.07.2003; 1 male, Çamlıyayla (Sebil, Tarsus stream), 37º07’N 34º32’E, 643 m, 30.07.2003; Osmaniye: 1 male, Hasanbeyli, 16.05.2003.

Family GOMPHIDAE Gomphus davidi Sélys, 1887

Materials: Adana: 5 males, Karaisalı (Ömerli village, Çatalan dam), 37º17’N 35º21’E, 190 m, 18.05.2003; 2 males, 1 female, Aladağ (Boztahta, Simit bridge), 37º23’N 35º26’E, 150 m, 05.06.2003; Hatay 1 male, İskenderun (Sarıseki), 36º40’N 36º13’E, 26 m, 07.06.2003; 1 female, İskenderun (Arsuz), 36º09’N 36º01’E, 520 m, 07.06.2003; 1 male, Reyhanlı (Müşrüfe, Afrin river), 36º19’N 36º32’E, 112 m, 08.06.2003; 1 female, northern of Kırıkhan (Kamışlar village), 36º38’N 36º26’E, 105 m, 08.06.2003; 1 female, Hassa (Akbez), 36º50’N 36º32’E, 100 m, 08.06.2003; 1 female, Yayladağ (Dağardı road), 35º52’N 36º06’E, 440 m, 09.06.2003; 1 male, Yayladağ (Görentaş village), 35º54’N 36º09’E, 568 m, 09.06.2003; MERSIN: 1 male, Mezitli stream, 36º43’N 34º30’E, 10 m, 19.05.2003.

Gomphus flavipes lineatus Bartenev, 1929 Materials: Adana: 1 female, Ceyhan road (Karaçay), 37º03’N 37º57’E, 35ºm, 19.07.2002.

Onychogomphus assimilis (Schneider, 1845) Materials: Adana: 1 male, 1 female, Pozantı (Kamışlı, between Gedikli-Hamidiye), 37º30’N 35º03’E, 917 m, 29.07.2003; 1 male, 1 female, Aladağ (Büyüksofulu), 37º32’N 35º09’E, 940 m, 29.07.2003; Mersin: 1 male, Tarsus (Keşbükü, Tarsus stream), 37º01’N 34º45’E, 139 m, 17.07.2002.

Onychogomphus forcipatus albotibialis Schmidt, 1954 Materials: Adana: 1 male, Feke (Göksu river), 37º48’N 35º55’E, 963 m, 19.08.2002; Mersin: 2 males, 1 female, Bozyazı (Gözce), 36º09’N 33º10’E, 40 m, 03.06.2003; Osmaniye: 1 female, Yarpuz (Haraz Plateau), 29.07.2003.

Onychogomphus lefebvrei (Rambur, 1842) Materials: Adana: 1 male, Aladağ (Eyner), 37º25’N 35º26’E, 200 m, 29.07.2003; 1 male, Aladağ (Simit bridge), 37º23’N 35º26’E, 150 m, 29.07.2003; Hatay: 2 males, İskenderun (Arsuz), 36º09’N 36º01’E, 520 m, 07.06.2003; 2 males, northern of Kırıkhan (Kamışlar village), 36º38’N 36º26’E, 105 m, 08.06.2003; 2 males, 1 female, Hassa (Akbez), 36º50’N 36º32’E, 100 m, 08.06.2003; 1 male, Yayladağ (Dağardı road), 35º52’N 36º06’E, 440 m, 09.06.2003; 1 female, İskenderun (Sarıseki), 36º40’N 36º13’E, 26 m, 28.07.2003; Kahramanmaraş: 1 male, Andırın (Kesim village, Keşiş brook), 37º25’N 36º15’E, 170 m, 19.07.2002; 1 male, Ilıca (Menzelet Dam), 37º49’N 36º48’E, 690 m, 26.07.2003; 3 males, Çağlayancerit (Bozlar, Aksu stream), 37º39’N 37º27’E, 890 m, 27.07.2003; Mersin: 5 males, Tarsus (Sandal village, Söğütlü plateau), 565 m, 37º09’N 34º48’E, 17.07.2002; 1 male, Aydıncık (Gülnar road), 36º10’N 33º21’E, 30 m, 03.06.2003; 1 male, Aydıncık (Gülnar road), 36º12’N 33º21’E, 186 m, 03.06.2003; 1 male, Tece (Tece stream), 36º41’N 34º27’E, 24 m, 04.06.2003; 1 male, Silifke (Kargıcak, Kargıcak brook), 36º26’N 33º38’E, 195 m, 31.07.2003; Osmaniye: 5 males, Kadirli (Akova), 37º20’N 35º57’E, 47 m, 06.06.2003.

Onychogomphus macrodon (Selys, 1887) Materials: Hatay: 1 male, Northern of Kırıkhan (Kamışlar village), 36º38’N 36º26’E, 105 m, 08.06.2003.

Paragomphus lineatus (Sélys, Hagen, 1850) Materials: Adana: 1 male, Aladağ (Eyner), 37º25’N 35º26’E, 200 m, 29.07.2003; Hatay: 1 female, between Belen-Kırıkhan, 36º27’N 36º15’E, 195 m, 20.07.2002; 1 female, between Kırıkhan-Kumlu, 36º25’N 36º23’E, 88 m, 20.07.2002; Kahramanmaraş: 1 male, Andırın (between Yeniköy-Kesim), 37º25’N 36º14’E, 200 m, 19.07.2002.


Family CORDULEGASTRIDAE Cordulegaster insignis insignis Schneider, 1845

Materials: Adana: 1 male, Aladağ (Kahyalar), 37º17’N 35º23’E, 208 m, 18.05.2003; 2 males, Aladağ (Eyner), 37º25’N 35º26’E, 200 m, 05.06.2003; Hatay: 1 female, Hassa (Akbez), 36º50’N 36º37’E, 105 m, 08.06.2003; 1 male, Antakya (Campus of Mustafa Kemal University), 36º19’N 36º12’E, 120 m, 16.05.2003; Kahramanmaraş: 1 male, Andırın (Gökahmetler, Aslantaş dam), 37º24’N 36º16’E, 250 m, 06.06.2003; Mersin: 1 female, Çamlıyayla (Papazın bahçesi), 37º14’N 34º37’E, 761 m, 30.07.2003; Osmaniye: 1 male, Yarpuz (Haraz Plateau), 29.07.2003.

Family LIBELLULIDAE Libellula depressa Linnaeus, 1758

Materials: Adana: 1 male, Tufanbeyli (Pınarlar), 38º13’N 36º13’E, 1400 m, 10.06.2003; 1 male, Tufanbeyli (Güzelim village, Güzelim brook), 38º07’N 36º10’E, 1430 m, 10.06.2003; Hatay: 1 male, Narlıca, 36º15’N 36º15’E, 93 m, 08.06.2003; Kahramanmaraş: 1 male, Andırın (Kesik), 37º25’N 36º20’E, 1127 m, 06.06.2003; 1 male, Afşin (Alimpınar), 38º12’N 36º53’E, 1285 m, 10.06.2003; 1 male, Afşin (Büyükkızılcık village, Kızılcık brook), 38º09’N 36º42’E, 1400 m, 10.06.2003; 1 male, Göksun (Salyan), 1340 m, 38º02’N 36º38’E, 10.06.2003; 2 males, Afşin (Yazdere, Hurman stream), 38º09’N 37º00’E, 1320 m, 10.06.2003.

Libellula fulva Müller, 1764 Materials: Mersin: 1 male, Bozyazı (Gözce), 36º09’N 33 10’E, 40 m, 03.06.2003.

Libellula pontica (Sélys, 1887) Materials: Hatay: 1 male, northern of Kırıkhan, 36º34’N 36º24’E, 113 m, 16.05.2003; Mersin: 1 male, Silifke (Kapızlı), 36º23’N 34 02’E, 22 m, 19.05.2003.

Orthetrum brunneum (Fonscolombe, 1837) Materials: Adana: 1 male, Karaisalı (Topaktaş, Körkün stream), 37º09’N 35º09’E, 104 m, 18.07.2002; 1 male, Tufanbeyli (Güzelim, race of Sarız stream), 38º08’N 36º10’E, 1390 m, 19.08.2002; 1 male, Tufanbeyli (İncedere), 38º13’N 36º13’E, 1400 m, 19.08.2002; 1 female, Kozan (Kozan dam), 37º31’N 35º50’E, 300 m, 05.06.2003; 1 male, Feke (Tenkerli village), 37º44’N 35º52’E, 05.06.2003; 1 male, Aladağ, 37º33’N 35º22’E, 730 m, 29.07.2003; Kahramanmaraş: 1 male, Andırın (Kesik), 37º25’N 36º20’E, 1127 m, 06.06.2003; 1 male, Afşin (Büyükkızılcık village, Kızılcık brook), 38º09’N 36º42’E, 1400 m, 10.06.2003; 2 males, 2 females, Pazarcık (Kartalkaya Dam), 37º29’N 37º17’E, 730 m, 27.07.2003; 1 male, Ekinözü (Gaziler), 37º59’N 37º06’E, 1084 m, 26.07.2003; 1 male, Çağlayancerit (Aksu village, Aksu stream), 37º44’N 37º22’E, 987 m, 27.07.2003; 1 male, Çağlayancerit (Başdevrişli), 37º42’N 37º10’E, 1260 m, 27.07.2003; 1 female, Elbistan (Afşin Road, DSİ Regulator), 38º11’N 37º05’E, 1138 m, 26.07.2003.

Orthetrum cancellatum (Linnaeus, 1758) Materials: Hatay: 1 male, Antakya (Campus of Mustafa Kemal University), 36º19’N 36º12’E, 120 m, 16.05.2003; Mersin: 1 male, Tarsus (Kurtçukuru, Kadıncık-I dam), 37º08’N 34 44’E, 420 m, 17.07.2002.

Orthetrum chrysostigma (Burmeister, 1839) Materials: Adana: 1 male, Kozan (Kozan dam), 37º31’N 35º50’E, 300 m, 05.06.2003; 1 male, Aladağ (Göksu'ya Akan Dere, Simit Bridge), 37º23’N 35º26’E, 150 m, 29.07.2003; Hatay: 8 males, 1 female, İskenderun, Sarıseki (Sarıseki brook), 36º40’N 36º13’E, 17 m, 20.07.2002; 1 male, Kırıkhan (Hassa road, Bulgur spring), 36º37’N 36º26’E, 165 m, 21.07.2002; 1 female, İskenderun (Arsuz), 36º29’N 35º59’E, 5 m, 15.05.2003; 1 male, İskenderun (Arsuz), 36º09’N 36º01’E, 520 m, 07.06.2003, 1 male, Antakya (Demirköprü, Asi River), 36º14’N 36º21’E, 102 m, 08.06.2003; Kahramanmaraş: 2 males, Andırın (between Yeniköy-Kesim), 37º25’N 36º14’E, 200 m, 19.07.2002; 5 males, Andırın (Kesim village, Keşiş brook), 37º25’N 36º15’E, 170 m, 19.07.2002; Mersin: 1 male, eastern of Bozyazı, 36º06’N 33 00’E, 22 m, 03.06.2003; 1 male, Bozyazı (Gözce), 36º09’N 33º10’E, 40 m, 03.06.2003.


Orthetrum coerulescens anceps (Schneider, 1845) Materials: Adana: 1 male, Aladağ (Büyüksofulu village), 37º33’N 35º10’E, 1024 m, 21.06.2002; 1 female, Karaisalı (Salbaş village, Çakıt brook), 37º06’N 35º06’E, 84 m, 18.07.2002; 1 male, Karaisalı (Topaktaş, Körkün stream), 37º09’N 35º09’E, 104 m, 18.07.2002; 1 male, Karaisalı (Güvenç village), 37º12’N 35º05’E, 170 m, 18.05.2003; Mersin: 3 males, Bozyazı, 36º06’N 32º57’E, 10 m, 03.06.2003; 1 male, Erdemli (Arpaçbahşiş), 36º38’N 34º22’E, 54 m, 04.06.2003; 1 male, Erdemli (Arpaçbahşiş), 36º38’N 34º22’E, 54 m, 31.07.2003.

Orthetrum sabina (Drury, 1770) Materials: Adana: 2 males, Karaisalı (Çatalan dam), 37º11’N 35º16’E, 72 m,18.07.2002; 3 females, Karaisalı (Çiçekli village, Seyhan dam), 37º08’N 35º18’E, 73 m, 18.07.2002; 1 male, Karataş (Topraklı, Dalyan-Ilgınlık), 36º39’N 35º20’E, 10 m, 17.05.2003; 1 male, , Karataş (Bahçeköy), 36º35’N 35º25’E, 2 m, 17.05.2003; 2 males, 2 females, Karaisalı (Abacı, Seyhan dam), 37º09’N 35º11’E, 150 m, 18.05.2003; 1 male, 2 females, Aladağ (Topallı village, Çatalan dam), 37º19’N 35º23’E, 124 m, 18.05.2003; 1 male, Karaisalı (Eğlence, Çatalan dam), 37º17’N 35º13’E, 135 m, 18.05.2003; 1 female, Ceyhan (Ceyhan river), 37º03’N 35º45’E, 200 m, 29.07.2003; Kahramanmaraş: 1 male, Andırın (Between Yeniköy-Kesim), 37º25’N 36º14’E, 200 m, 19.07.2002; 1 male, , Andırın (Gökahmetler, Aslantaş dam), 37º24’N 36º16’E, 250 m, 06.06.2003; MERSIN: 1 male, Tece (Tece Stream), 36º41’N 34º27’E, 24 m, 31.07.2003; Hatay: 1 male , İskenderun (Sarıseki), 36º40’N 36º13’E, 17 m, 20.07.2002; 1 female, İskenderun (Sarıseki, Sarıseki brook), 36º40’N 36º13’E, 9 m, 20.07.2002; 1 male, between Kırıkhan-Kumlu, 36º25’N 36º23’E, 88 m, 20.07.2002; 1 male, 1 female, Kumlu, 36º23’N 36º24’E, 95 m, 20.07.2002; 1 female , Reyhanlı (Antakya road 10.km, canal), 36º14’N 36º23’E, 94 m, 20.07.2002; 1 male, İskenderun (Arsuz), 36º29’N 35º59’E, 5 m, 15.05.2003; 1 male, Antakya (Campus of Mustafa Kemal University), 36º19’N 36º12’E, 120 m, 16.05.2003; 1 female , Kırıkhan (Gölbaşı), 36º30’N 36º28’E, 95 m, 16.05.2003; 1 male, Kırıkhan (Gölbaşı), 36º30’N 36º28’E, 100 m, 08.06.2003; 1 male, İskenderun (Arsuz), 36º09’N 36º01’E, 520 m, 07.06.2003; 1 male, Reyhanlı, 36º14’N 36º34’E, 187 m, 08.06.2003; 2 males, Kırıkhan (Başpınar, Özkızılkaya bridge, canal), 36º27’N 36º28’E, 95 m, 08.06.2003; 2 males, Kırıkhan (Gölbaşı), 36º30’N 36º28’E, 100 m, 08.06.2003; 1 male, Northern of Kırıkhan 36º34’N 36º23’E, 105 m, 08.06.2003; Osmaniye: 1 male, Centrum, 37º06’N 36º18’E, 140 m, 16.05.2003; 1 male, Kadirli (Akova), 37º20’N 35º57’E, 47 m, 06.06.2003.

Orthetrum taeniolatum (Schneider, 1845) Materials: Adana: 1 female, Kozan (Kozan Dam, Kılgen), 37º31’N 35º49’E, 300 m, 18.07.2002; 1 male, 1 female, Kozan (Deliçay), 37º34’N 35º47’E, 119 m, 18.07.2002; 1 male, Karaisalı (Kuzgun village), 37º09’N 35º05’E, 162 m, 18.05.2003; 2 males, Karaisalı (Güvenç village), 37º12’N 35º05’E, 170 m, 18.05.2003; 2 females, Aladağ (Kahyalar), 37º17’N 35º23’E, 208 m, 18.05.2003; 1 female, İmamoğlu-Kozan road (Kurukulak brook), 37º17’N 35º42’E, 66 m, 05.06.2003; Hatay: 1 male, Antakya (Paşa village, Bedirge brook), 36º26’N 36º13’E, 100 m, 15.05.2003; 1 male, İskenderun (Arsuz), 36º24’N 35º53’E, 15 m, 15.05.2003; 1 female, between Belen-Kırıkhan, 36º27’N 36º15’E, 195 m, 20.07.2002; 1 female, Kırıkhan (Hassa road, Bulgur spring), 36º37’N 36º26’E, 165 m, 21.07.2002; Mersin:1 female, Mezitli stream, 36º43’N 34º30’E, 10 m, 19.05.2003; 1 male, 1 female, Tece (Tece stream), 36º41’N 34º27’E, 24 m, 19.05.2003; 1 female, Erdemli (Tömük), 36º28’N 34º21’E, 20 m, 19.05.2003; 1 female, Erdemli (Arpaçbahşiş, Kargıcak brook), 36º36’N 34º19’E, 14 m, 19.05.2003; 1 male, Erdemli (Tömük), 36º28’N 34º21’E, 20 m, 31.07.2003; 2 males, Çamlıyayla (Kadıncık-I dam), 37º09’N 34º43’E, 429 m, 30.07.2003.

Crocothemis erythraea (Brullé, 1832) Materials: Adana: 1 male, Aladağ (Kahyalar), 37º17’N 35º23’E, 208 m, 18.05.2003; Hatay: 1 female, İskenderun (Güzin Brook), 36º34’N 36º09’E, 13 m, 20.07.2002; 1 female, İskenderun (Sarıseki), 36º40’N 36º13’E, 9 m, 20.07.2002; 1 male, Hassa (Akbez), 36º50’N 36º37’E, 105 m, 08.06.2003; 2 males, Kırıkhan (Gölbaşı), 36º30’N 36º28’E, 100 m, 08.06.2003; 1 male, İskenderun (Sarıseki, Sarıseki brook), 36º40’N 36º13’E, 9 m, 20.07.2002; 1 male, İskenderun (Güzin brook), 36º34’N 36º09’E, 13 m, 20.07.2002; 1 male, Kumlu, 36º23’N 36º24’E, 95 m, 20.07.2002; 1 male, Kumlu (Müşrüfe, Afrin river), 36º18’N 36º32’E, 104 m, 20.07.2002; 1 female, Reyhanlı (Antakya road), 36º14’N 36º23’E, 94 m, 20.07.2002; Kahramanmaraş: 1 female, Pazarcık (Kartalkaya Dam), 37º29’N 37º17’E, 730


m, 27.07.2003; Mersin: 1 male, Bozyazı, 36º06’N 32º57’E, 10 m, 03.06.2003; 1 male, Silifke (Göksu River), 36º24’N 33º48’E, 70 m, 31.07.2003; 1 male, Silifke (Kapızlı), 36º23’N 34º02’E, 22 m, 31.07.2003; 1 male, Merkez (Akarsu), 36º46’N 34º36’E, 10 m, 31.07.2003; 1 male, Tece (Tece Stream), 36º41’N 34º27’E, 24 m, 31.07.2003; Osmaniye: 1 male, Toprakkale crossroads (Karaçay), 37º03’N 36º12’E, 114 m, 19.07.2002; 1 male, Kadirli (Akova, marsh), 37º20’N 35º57’E, 47 m, 06.06.2003.

Crocothemis servilia (Drury, 1770) Materials: Adana: 1 male, Karaisalı (Çiçekli village, Seyhan dam), 37º08’N 35º18’E, 73 m, 18.07.2002; 1 male, 1 female, Karataş (Topraklı, Dalyan-Ilgınlık), 36º39’N 35º20’E, 10 m, 17.05.2003; Hatay: 1 male, İskenderun (Güzin brook), 36º34’N 36º09’E, 13 m, 20.07.2002; 2 males, Kırıkhan (Gölbaşı), 36º30’N 36º28’E, 95 m, 16.05.2003; 1 female, Reyhanlı (Başpınar, Özkızılkaya Brook), 36º27’N 36º28’E, 95 m, 08.06.2003; 1 male, İskenderun (Sarıseki), 36º40’N 36º13’E, 26 m, 28.07.2003; Mersin: 1 female, Erdemli (Limonlu stream), 36º32’N 34º13’E, 7 m, 19.05.2003; 1 male, Silifke (Kapızlı), 36º23’N 34º02’E, 22 m, 19.05.2003; 1 male, Bozyazı (Gözce), 36º09’N 33º10’E, 40 m, 03.06.2003; 1 male, Aydıncık (Gülnar Road), 36º10’N 33º21’E, 30 m, 03.06.2003; 1 male, Tarsus (Akçakocalı, Berdan Dam), 36º57’N 34º50’E, 75 m, 04.06.2003; 1 male, Çamlıyayla (Kadıncık-I dam), 37º09’N 34º43’E, 429 m, 30.07.2003; 1 male, 1 female, Silifke (Göksu River), 36º24’N 33º48’E, 70 m, 31.07.2003; 1 female, Silifke (Kapızlı), 36º23’N 34º02’E, 22 m, 31.07.2003; Osmaniye: 1 male, Kadirli (Akova), 37º20’N 35º57’E, 47 m, 06.06.2003.

Sympetrum fonscolombei (Sélys, 1840) Materials: Adana: 3 males, 3 females, Karaisalı (Çiçekli village, Seyhan dam), 37º08’N 35º18’E, 73 m, 18.07.2002; 2 males, 1 female, Karaisalı (Salbaş village, Çakıt brook), 37º06’N 35º06’E, 84 m, 18.07.2002; 2 males, 1 female, Aladağ (Topallı village, Çatalan dam), 37º19’N 35º23’E, 124 m, 18.05.2003; 1 male, Karaisalı (Ömerli village, Çatalan dam), 37º17’N 35º21’E, 190 m, 18.05.2003; 1 male, Aladağ, 37º33’N 35º22’E, 730 m, 29.07.2003; Hatay: 1 male, 1 female, Belen (Kırıkhan kavşağı), 36º27’N 36º15’E, 195 m, 20.07.2002; 1 male, Reyhanlı (Antakya road), 36º14’N 36º31’E, 103 m, 20.07.2002; 1 male, Altınözü (Büyükburç, Yarseli dam), 36º11’N 36º19’E, 148 m, 21.07.2002; 2 males, Hassa (Akbez), 36º50’N 36º37’E, 384 m, 21.07.2002; 2 females, 1 male, Yayladağ (Görentaş village), 35º54’N 36º09’E, 568 m, 09.06.2003; Kahramanmaraş: 2 males, Andırın (between Yeniköy-Kesim), 37º25’N 36º14’E, 200 m, 19.07.2002; 1 male, 2 females, Türkoğlu (Minehüyük, Büyükgöl), 37º14’N 36º49’E, 483 m, 22.07.2002; 2 males, Türkoğlu (Maraş road, Aksu stream), 37º27’N 36º53’E, 461 m, 22.07.2002; 2 males, 2 females, Hasancıklı (Ceyhan river), 456 m, 22.07.2002; 1 male, Menzelet dam, 37º45’N 36º45’E, 645 m, 26.07.2003; 1 female, Çağlayancerit (Aksu village, Aksu stream), 37º44’N 37º22’E, 987 m, 27.07.2003; 2 males, Pazarcık-Narlı road (Kırni), 37º25’N 37º09’E, 622 m, 27.07.2003; Mersin: 2 males, 2 females, Tarsus (Kurtçukuru, Kadıncık-I dam), 37º08’N 34º44’E, 420 m, 17.07.2002; 1 female, Tarsus (Keşbükü, Tarsus Stream), 37º01’N 34º45’E, 139 m, 17.07.2002; 1 female, Aydıncık (Gülnar road), 36º10’N 33º21’E, 30 m, 03.06.2003; 1 male, 1 female, Tarsus (Akçakocalı, Berdan dam), 36º57’N 34º50’E, 75 m, 04.06.2003; Osmaniye: 1 male, Kaypak (Kalecik dam), 37º09’N 36º27’E, 600 m, 16.05.2003.

Sympetrum meridionale (Sélys, 1841) Materials: Adana: 1 female, Aladağ (Kıcak), 37º35’N 35º14’E, 1075 m, 29.07.2003; 1 male, Aladağ (Büyüksofulu), 37º32’N 35º09’E, 940 m, 29.07.2003; Hatay: 1 female, Yayladağ (Görentaş village), 35º54’N 36º09’E, 568 m, 09.06.2003; Kahramanmaraş: 1 female, Türkoğlu (Maraş road, Aksu stream), 37º27’N 36º53’E, 461 m, 22.07.2002; Mersin: 1 female, Tarsus (Kurtçukuru, Kadıncık-I dam), 37º08’N 34º44’E, 420 m, 17.07.2002; 1 male, Çamlıyayla (Papazın bahçesi), 37º14’N 34º37’E, 761 m, 30.07.2003; Osmaniye: 1 male, 1 female, Yarpuz (Haraz plateau), 29.07.2003.

Sympetrum sanguineum (Müller, 1764) Materials: Adana: 1 female, Saimbeyli (Saimbeyli brook, Ayçukuru), 37º57’N 36º04’E, 850 m, 19.08.2002; Hatay: 1 male, Reyhanlı (Antakya road), 36º14’N 36º23’E, 94 m, 20.07.2002; 1 male, Yayladağ (Görentaş village), 35º54’N 36º09’E, 568 m, 09.06.2003; Kahramanamaraş: 2 males, 1 female, Türkoğlu (Minehüyük, Büyükgöl), 37º14’N 36º49’E, 483 m, 22.07.2002; 1 male, Çağlayancerit (Bozlar, Aksu stream), 37º39’N 37º27’E, 890 m, 27.07.2003; Mersin: 1 male, Tarsus (Akçakocalı, Berdan dam), 36º57’N 34º50’E, 75 m, 04.06.2003.


Sympetrum striolatum (Charpentier, 1840) Materials: Adana: 1 male, Karaisalı (Çiçekli village, Seyhan dam), 37º08’N 35º18’E, 73 m, 18.07.2002; 1 female, Aladağ (Gedikli), 37º29’N 35º09’E, 920 m, 29.07.2003; 1 female, Aladağ (Kıcak), 37º35’N 35º14’E, 1075 m, 29.07.2003; Hatay: 2 females, İskenderun (Arsuz), 36º29’N 35º59’E, 5 m, 15.05.2003; 1 male, Hassa (Akbez), 36º50’N 36º37’E, 105 m, 08.06.2003; 1 male, Antakya (Campus of Mustafa Kemal University), 36º19’N 36º12’E, 120 m, 16.05.2003; Mersin: 1 male, 1 female, Bozyazı (Gözce), 36º09’N 33º10’E, 40 m, 03.06.2003; 1 male, Çamlıyayla (Okkoyağı village, Tarsus stream), 37º13’N 34º38’E, 770 m, 30.07.2003; 2 males, Çamlıyayla (Papazın bahçesi), 37º14’N 34º37’E, 761 m, 30.07.2003; 1 male, Çamlıyayla (Okkoyağı village, Tarsus stream), 37º13’N 34º38’E, 770 m, 30.07.2003; Osmaniye: 1 male, Yarpuz (Haraz plateau), 29.07.2003.

Brachythemis fuscopalliata (Sélys, 1887) Materials: Adana: 3 males, Karaisalı (Karaömerli village, Seyhan dam), 37º06’N 35º20’E, 79 m, 18.07.2002; 1 female, Karaisalı (Çiçekli village, Seyhan dam), 37º08’N 35º18’E, 73 m, 18.07.2002; 1 male, Karataş (Topraklı), 1 female,36º39’N 35º20’E, 10 m, 17.05.2003; Karataş (Bahçeköy), 36º35’N 35º25’E, 2 m, 17.05.2003; Mersin: 1 female, Erdemli (Tömük), 36º28’N 34º21’E, 20 m, 19.05.2003; Kahramanmaraş: 1 male, 2 females, Andırın (Gökahmetler, Aslantaş dam), 37º24’N 36º16’E, 250 m, 06.06.2003; Hatay: 1 male, Reyhanlı (Müşrüfe, Afrin river), 36º19’N 36º32’E, 112 m, 08.06.2003; 1 female, Kırıkhan (Başpınar, Özkızılkaya bridge, stream), 36º27’N 36º28’E; 1 male, 1 female, Yayladağ (Görentaş village), 35º54’N 36º09’E, 568 m, 09.06.2003.

Brachythemis leucosticta (Burmeister, 1839) Materials: Hatay: 1 female Kırıkhan (Başpınar, Özkızılkaya bridge, stream), 36º27’N 36º28’E.

Diplacodes lefebvrei (Rambur, 1842) Materials: Adana: 5 females, Karataş (Topraklı, Dalyan-Ilgınlık), 36º39’N 35º20’E, 10 m, 17.05.2003; Hatay: 3 females, İskenderun, Sarıseki (Sarıseki brook), 36º40’N 36º13’E, 17 m, 20.07.2002; 5 males, 1 female, İskenderun (Sarıseki, Sarıseki brook), 36º40’N 36º13’E, 9 m, 20.07.2002; 1 male, İskenderun (Sarıseki), 36º40’N 36º13’E, 26 m, 07.06.2003; Mersin: 1 male, Bozyazı (Dikilitaş beach), 36º05’N 32º55’E, 3 m, 03.06.2003; 1 male, Aydıncık (Gülnar road), 36º10’N 33º21’E, 30 m, 03.06.2003; 1 female, Aydıncık (Gülnar road), 36º12’N 33º21’E, 186 m, 03.06.2003; 2 males, Tarsus (Akçakocalı, Berdan dam), 36º57’N 34º50’E, 75 m, 04.06.2003; 1 male, Centrum (small stream), 36º46’N 34º36’E, 10 m, 31.07.2003; 1 male, Silifke (Kapızlı), 36º23’N 34º02’E, 22 m, 31.07.2003.

Trithemis annulata (Beauvois, 1807) Materials: Adana: 1 male, Karaisalı road (Seyhan river-canal), 37º00’N 35º10’E, 47 m, 18.07.2002; 2 males, 4 females, Karaisalı, 37º02’N 35º10’E, 69 m, 18.07.2002; 1 male, Karaisalı (Salbaş village, Çakıt brook), 37º06’N 35º06’E, 84 m, 18.07.2002; 2 males, Karaisalı (Salbaş village, Üçürge brook), 37º08’N 35º08’E, 98 m, 18.07.2002; 1 female, between Adana-Karaisalı, 37º02’N 35º10’E, 69 m, 18.07.2002; 3 males, Karaisalı (Çatalan dam), 37º11’N 35º16’E, 72 m,18.07.2002; 1 male, İmamoğlu (Hakkıbeyli, Han brook), 37º10’N 35º33’E, 187 m, 18.07.2002; 1 male, İmamoğlu-Kozan road (Tanrı brook), 37º18’N 35º43’E, 63 m, 18.07.2002; 4 females, Kozan (Kozan dam, Kılgen), 37º31’N 35º49’E, 300 m, 18.07.2002; 1 male, Zeytinli (canal I), 36º59’N 35º07’E, 25 m, 18.07.2002; 1 female, Karaisalı (Abacı, Seyhan dam), 37º09’N 35º11’E, 150 m, 18.05.2003; 1 male, Kozan (Deliçay), 37º34’N 35º47’E, 119 m, 18.07.2002; 1 male, Karaisalı (Ömerli village, Çatalan dam), 37º17’N 35º21’E, 190 m, 18.05.2003; 1 female, Karaisalı (Ömerli village), 37º17’N 35º15’E, 191 m, 18.05.2003; 1 female, Aladağ (Boztahta, Simit bridge), 37º23’N 35º26’E, 150 m, 05.06.2003; 1 male, İmamoğlu-Kozan road (Tırmıl brook), 37º20’N 35º45’E, 85 m, 05.06.2003; 1 female, Kozan (Deliçay), 37º24’N 35º47’E, 110 m, 05.06.2003; 1 male, Kozan (Kozan dam), 37º31’N 35º50’E, 300 m, 05.06.2003; 1 male, Kozan (Kayhan), 37º22’N 35º50’E, 82 m, 06.06.2003; 1 male, Ceyhan (Ceyhan river), 37º03’N 35º45’E, 200 m, 29.07.2003; 1 male, Aladağ (Simit bridge), 37º23’N 35º26’E, 150 m, 29.07.2003; HATAY: 1 female, İskenderun, Sarıseki (Sarıseki brook), 36º40’N 36º13’E, 17 m, 20.07.2002; 1 male, İskenderun (Güzin brook), 36º34’N 36º09’E, 13 m, 20.07.2002; 1 male, 2 females, between Belen-Kırıkhan, 36º27’N 36º15’E, 195 m, 20.07.2002; 1 male, Kırıkhan -Kumlu road, 36º27’N 36º22’E, 152 m, 20.07.2002; 1 male, between Kırıkhan-Kumlu, 36º25’N 36º23’E, 88 m, 20.07.2002; 4 males, 1 female, Kumlu (Müşrüfe, Afrin river), 36º18’N 36º32’E, 104


m, 20.07.2002; 1 male, Antakya (Demirköprü, Asi river), 36º14’N 36º21’E, 101 m, 20.07.2002; 3 males, Antakya (Nallıca, Asi river), 36º15’N 36º14’E, 100 m, 20.07.2002; 1 female, Altınözü (Büyükburç, Yarseli dam), 36º11’N 36º19’E, 148 m, 21.07.2002; 5 males, 1 female, Kırıkhan (Hassa road, Bulgur spring), 36º37’N 36º26’E, 165 m, 21.07.2002; 1 female, Hassa (Akbez), 36º50’N 36º37’E, 384 m, 21.07.2002; 1 female, İskenderun (Sarıseki), 36º40’N 36º13’E, 26 m, 15.05.2003; 1 female, İskenderun (Arsuz), 36º29’N 35º59’E, 5 m, 15.05.2003; 1 male, İskenderun (Arsuz, Arsuz stream), 36º24’N 35º53’E, 10 m, 15.05.2003; 1 male, Belen (Çakallı), 36º26’N 36º16’E, 170 m, 15.05.2003; 2 males, Antakya (Campus of Mustafa Kemal University), 36º19’N 36º12’E, 120 m, 16.05.2003; 1 female, Kırıkhan (Gölbaşı), 36º30’N 36º28’E, 95 m, 16.05.2003; 1 male, northern of Kırıkhan, 36º34’N 36º24’E, 113 m, 16.05.2003; 1 male, 1 female, İskenderun (Sarıseki), 36º40’N 36º13’E, 26 m, 07.06.2003; 1 female, İskenderun (Arsuz), 36º09’N 36º01’E, 520 m, 07.06.2003; 1 male, Samandağ (Karaçay, Küçükdere), 36º09’N 36º01’E, 520 m, 07.06.2003; 1 male, Samandağ (Karaçay, Büyükdere), 36º08’N 36º01’E, 500 m, 07.06.2003; 1 male, Narlıca, 36º15’N 36º15’E, 93 m, 08.06.2003; 1 female, Antakya (Demirköprü, Asi river), 36º14’N 36º21’E, 102 m, 08.06.2003; 3 males, Reyhanlı, 36º14’N 36º34’E, 187 m, 08.06.2003; 2 males, 1 female, Kırıkhan (Başpınar, Özkızılkaya bridge, stream), 36º27’N 36º28’E; 1 male, Kırıkhan (Gölbaşı), 36º30’N 36º28’E, 100 m, 08.06.2003; 1 male, Kırıkhan (Büyükkarasu brook), 36º30’N 36º24’E, 100 m, 08.06.2003; 3 males, northern of Kırıkhan 36º34’N 36º23’E, 105 m, 08.06.2003; 1 male, Reyhanlı (Müşrüfe, Afrin river), 36º19’N 36º32’E, 112 m, 08.06.2003; 1 male, 1 female, Yayladağ (Görentaş village), 35º54’N 36º09’E, 568 m, 09.06.2003; Kahramanmaraş: 1 male, 1 female, Andırın (between Yeniköy-Kesim), 37º25’N 36º14’E, 200 m, 19.07.2002; 1 male, Andırın (Anacık village, Aslantaş dam), 37º23’N 36º19’E, 185 m, 19.07.2002; 1 female, Türkoğlu (Maraş road, Aksu stream), 37º27’N 36º53’E, 461 m, 22.07.2002; 1 male, Türkoğlu-Maraş road (Aksu stream), 37º29’N 36º53’E, 457 m, 22.07.2002; 2 males, Andırın (Gökahmetler, Aslantaş dam), 37º24’N 36º16’E, 250 m, 06.06.2003; Mersin: 1 male, Tarsus (Sandal village, Söğütlü plateau), 565 m, 37º09’N 34º48’E, 17.07.2002; 1 male, 1 female, Tarsus (Kurtçukuru, Kadıncık-I dam), 37º08’N 34º44’E, 420 m, 17.07.2002; 2 males, Tarsus (Tarsus stream, Berdan-I canal), 36º55’N 34º55’E, 14 m, 18.07.2002; 1 male, Bozyazı (Gözce), 36º09’N 33º10’E, 40 m, 03.06.2003; 1 male, Erdemli (Arpaçbahşiş), 36º38’N 34º22’E, 54 m, 04.06.2003; 1 male, 1 female, Tarsus (Akçakocalı, Berdan dam), 36º57’N 34º50’E, 75 m, 04.06.2003; 3 males, Silifke (Kapızlı), 36º23’N 34º02’E, 22 m, 19.05.2003; 1 male, Silifke (Kapızlı), 36º23’N 34º02’E, 22 m, 31.07.2003; 1 female, Çamlıyayla (Kurtçukuru, Kurtçukuru brook), 37º10’N 34º45’E, 581 m, 30.07.2003; 1 male, Erdemli (Arpaçbahşiş), 36º38’N 34º22’E, 54 m, 31.07.2003; 1 male, Erdemli (Limonlu stream), 36º32’N 34º13’E, 7 m, 31.07.2003; 1 male, Merkez, 36º46’N 34º36’E, 10 m, 31.07.2003; 1 male, Tece (Tece stream), 36º41’N 34º27’E, 24 m, 31.07.2003; OSMANIYE: 2 males, Çukurköprü, 37º20’N 35º55’E, 52 m, 19.07.2002; 1 female, Kadirli (between Çukurköprü-Aydınlar), 37º20’N 35º56’E, 51 m, 19.07.2002; 1 male, Kadirli (Aydınlar), 37º21’N 36º00’E, 56 m, 19.07.2002; 1 male, Kadirli (Yalnızdut), 37º17’N 36º02’E, 62 m, 19.07.2002; 1 female, Kadirli (Orhaniye, Seyhan brook), 37º12’N 36º04’E, 48 m, 19.07.2002; 1 male, Toprakkale (Yolçatı brook), 37º03’N 36º06’E, 55 m, 19.07.2002; 1 male, Kaypak (Kalecik dam), 37º09’N 36º27’E, 600 m, 16.05.2003; 1 male, 1 female, near Centrum, 37º06’N 36º18’E, 140 m, 16.05.2003; 1 male, 2 females, Kadirli (Mehmetli, Mehmetli dam), 06.06.2003.

Trithemis arteriosa (Burmeister, 1839) Materials: Adana: 2 males, Kozan (Kozan dam), 37º31’N 35º50’E, 300 m, 05.06.2003; Hatay: 1 male, İskenderun, Sarıseki (Sarıseki brook), 36º40’N 36º13’E, 17 m, 20.07.2002; 1 male, Yayladağ (Çabala-Kızılçad road), 35º52’N 36º06’E, 394 m, 15.05.2003; 2 males, Yayladağ (Çabala-Kızılçad road), 35º52’N 36º06’E, 440 m, 09.06.2003; Mersin: 1 male, Aydıncık (Gülnar road), 36º12’N 33º21’E, 186 m, 03.06.2003; 1 male, Aydıncık (Bozağaç-Gülnar road, Menekşe brook), 36º16’N 33º23’E 680 m, 03.06.2003; Osmaniye: 1 male, Yarpuz plateau, 29.07.2003.

Trithemis festiva (Rambur, 1842) Materials: Adana: 1 male, Aladağ (Simit bridge), 37º23’N 35º26’E, 150 m, 29.07.2003; Mersin: 1 male, Bozyazı (Gözce), 36º09’N 33º10’E, 40 m, 03.06.2003; 1 male, 1 female,


Bozyazı (between Gözce-Gözsüzce), 36º11’N 33º12’E, 115 m, 03.06.2003; 2 males, Aydıncık (Gülnar road), 36º10’N 33º21’E, 30 m, 03.06.2003.

Selysiothemis nigra (Vander Linden, 1825) Materials: Hatay: 2 females, Hassa (Akbez), 36º50’N 36º37’E, 384 m, 21.07.2002; Osmaniye: 2 males, Kadirli (Orhaniye, Seyhan brook), 37º12’N 36º04’E, 48 m, 19.07.2002.

ACKNOWLEDGEMENTS

We are most grateful for the financial support given by Gazi University, Ministry of BAP (Project no: 05/2002-34) and we express our sincere thanks to Prof.Dr. Nedim Uygun (Çukurova University Faculty of Agriculture, Plant Protection Department), whon enabled us to check the collection of Odonata. The present study is a part of dissertation which name is “Odonata Fauna of Mediterranean Region of Turkey”.

LITERATURE CITED

Demirsoy, A. 1982. Türkiye Faunası, Odonata. TÜBİTAK, 4, 8, 154 pp. Demirsoy, A. 1995. Türkiye Faunası, Odonata. TÜBİTAK, 4, 8, 446 pp. (Unpublished project report) Ardıç, A. & Uygun, N. 1996. Doğu Akdeniz Bölgesi Odonata faunasının saptanması. Türkiye 3. Entomoloji Kongresi, 24-28 Eylül, Ankara.


Fig. 1 Map of the East Mediterranean region of Turkey.


Table 1. Wrong identifications and the true ones of Ardıç&Uygun (1996).

In Ardıç ve Uygun (1996) Correct

Calopteryx splendens (Harris, 1782) Calopteryx splendens intermedia (Selys 1887)

Calopteryx virgo (Linnaeus, 1758) Calopteryx virgo festiva (Brullé 1832)

Platycnemis latipes (Rambur, 1842) Platycnemis dealbata Selys & Hagen 1850

Lestes viridis (Vander Linden, 1825) Lestes viridis parvidens (Artobolevski 1929)

Lestes virens (Charpentier, 1845) Lestes virens vestalis (Rambur 1842)

Ischnura elegans (Vander Linden, 1823) Ischnura elegans ebneri Schmidt, 1938

Onychogomphus forcipatus (Linnaeus, 1758)

Onychogomphus forcipatus albotibialis Schmidt, 1954

Paragomphus genei (Selys, 1841) Paragomphus lineatus (Sélys & Hagen, 1850)

Gomphus vulgatissimus (Linnaeus, 1841) Gomphus davidi Selys, 1887

Orthetrum ramburi (Schneider, 1848) Orthetrum coerulescens anceps (Schneider, 1845)

Orthetrum coerulescens (Fabricius, 1789) Orthetrum chrysostigma (Burmeister, 1839)

Sympetrum vulgatum (Linnaeus, 1758) Doubtful, specimens was crushed.


NEW SUBSTITUTE NAMES FOR TWO ENNOMINAE GENERA (LEPIDOPTERA: GEOMETRIDAE)

Hüseyin Özdikmen* and Selma Seven**

* Gazi Üniversitesi, Fen-Edebiyat Fakültesi, Biyoloji Bölümü, 06500 Ankara / TÜRKİYE, e-mail: [email protected] ** Gazi Üniversitesi, Fen-Edebiyat Fakültesi, Biyoloji Bölümü, 06500 Ankara / TÜRKİYE, e-mail: [email protected] [Özdikmen, H. & Seven, S. 2006. New substitute names for two Ennominae genera (Lepidoptera: Geometridae). Munis Entomology & Zoology, 1 (2): 253-256] ABSTRACT: Two junior homonyms were detected amongst the Lepidoptera and the following replacement names are proposed: Begumia for Lobus Fletcher, 1979 and Eliphia for Moneta Warren, 1906 (Geometridae: Ennominae). Accordingly, new combinations are herein proposed for all type species currently included in monotypic genera Lobus Fletcher, 1979 and Moneta Warren, 1906. KEY WORDS: Begumia, Eliphia, Lobus, Moneta, homonymy, replacement names

BEGUMIA NOM. NOV., SUBSTITUTE NAME

Lobus Fletcher, 1979. In Nye [Ed.] The generic names of moths of the world. Vol. 3. Geometroidea. Publications Br. Mus. nat. Hist. No. 812: 118. (Insecta: Lepidoptera: Geometroidea: Geometridae: Ennominae). Preoccupied by Lobus Martin, 1972. J.Kans.ent.Soc. 45: 8. (Insecta: Diptera: Asiloidea: Asilidae).

The Afrotropical and Oriental genus Lobus was erected by Martin

(1972) in Diptera (Asiloidea: Asilidae). It is still used as a valid generic name in Diptera (Joseph & Pauri 1983, 1984, 1991, 1992, 1998). Also the Australian genus Lobus was proposed by Fletcher (1979) as an objective replacement name for Scioglyptis Meyrick, 1892 with the type species Scioglyptis lithinopa Meyrick, 1892 by subsequent designation by Turner, 1917 in Lepidoptera (Geometroidea: Geometridae: Ennominae) from Australia. Scioglyptis Meyrick, 1892 is a junior homonym of Scioglyptis Guest, 1887 (Lepidoptera: Geometridae: Ennominae). Scioglyptis Guest, 1887 and Lobus Fletcher, 1979 are still valid generic names in Lepidoptera. Scoble (1999) included Scioglyptis Meyrick, 1892 as a junior synonym of Lobus Fletcher, 1979 that had only one species. However, the name Lobus Fletcher, 1979 is invalid under the law of homonymy, being a junior homonym of Lobus Martin, 1972. Under the Zoological Code (ICZN, 1999) it must be rejected and replaced. In accordance with article 60 of the International Code of Zoological Nomenclature, fourth edition (1999), we propose to substitute the junior homonym Lobus Fletcher, 1979 for the nomen novum Begumia. As a result of this, Lobus Fletcher, 1979 is replaced with Begumia nom. nov. The following new combination is established: Begumia lithinopa (Meyrick, 1892), comb. nov.


SYSTEMATIC ACCOUNTS

Order Lepidoptera Superfamily Geometroidea Family Geometridae Subfamily Ennominae Genus Begumia new replacement name

For Lobus Fletcher, 1979, junior homonym of Lobus Martin, 1972. Type species: Scioglyptis lithinopa Meyrick, 1892, from Australia.

Gender: Feminine Etymology: Begumia dedicated to Begüm Seven Çalışkan. Species account and distribution: Monotypic; known from Australia: Queensland, Brisbane. Mandatory new combination: Begumia lithinopa (Meyrick, 1892) new comb. from Lobus.

Lobus Martin, 1972 (Diptera) is recorded in Nomenclator Zoologicus

vol. 8: 269 p. by Edwards & Tobias (1993) and Lobus Fletcher, 1979 (Lepidoptera) is recorded in Nomenclator Zoologicus vol. 9: 354 p. by Edwards et al. (1996).

ELIPHIA NOM. NOV., SUBSTITUTE NAME

Moneta Warren, 1906. Novit. zool., 13, 159. (Insecta: Lepidoptera: Geometroidea: Geometridae: Ennominae). Preoccupied by Moneta Cambridge, 1870. Proc. zool. Soc. London, 1870, 737. (Arachnida: Araneae: Labidognatha: Theridiidae).

The genus Moneta was erected by Cambridge (1870) with the type

species Moneta spinigera Cambridge, 1870 in Araneae (Labidognatha: Theridiidae). It is still used as a valid generic name in Araneae (Okuma 1994, Zhu 1998, Song et al. 1999, Yoshida 2001, 2003, Namkung 2003). Also the genus Moneta was described by Warren (1906) with the type species Moneta plenicolor Warren, 1906 by original designation in Lepidoptera (Geometroidea: Geometridae: Ennominae) from Angabunga River, British New Guinea [Papua New Guinea]. It is still a valid generic name in Lepidoptera. Scoble (1999) included only one species in the genus Moneta Warren, 1906. However, the name Moneta Warren, 1906 is invalid under the law of homonymy, being a junior homonym of Moneta Cambridge, 1870. Under the Zoological Code (ICZN, 1999) it must be rejected and replaced. In accordance with article 60 of the International Code of Zoological Nomenclature, fourth edition (1999), we propose to substitute the junior homonym Moneta Warren, 1906 for the nomen novum Eliphia. As a result of this, Moneta Warren, 1906 is replaced with Eliphia nom. nov. The following new combination is established: Eliphia plenicolor (Warren, 1906), comb. nov.


SYSTEMATIC ACOUNTS

Order Lepidoptera Superfamily Geometroidea Family Geometridae Subfamily Ennominae Genus Eliphia new replacement name

For Moneta Warren, 1906, junior homonym of Moneta Cambridge, 1870. Type species: Moneta plenicolor Warren, 1906, from Angabunga River, British New Guinea [Papua New Guinea].

Gender: Feminine Etymology: Eliphia dedicated to Elif Gül Özdikmen. Species account and distribution: Monotypic; known from British New Guinea [Papua New Guinea]: Angabunga River. Mandatory new combination: Eliphia plenicolor (Warren, 1906) new comb. from Moneta.

Moneta Cambridge, 1870 (Arachnida) and Moneta Warren, 1906

(Lepidoptera) are recorded in Nomenclator Zoologicus vol. 3: 204 p. by Neave (1940).

LITERATURE CITED

Cambridge, O. P. 1870. On some new genera and species of Araneida. Proceedings of the Zoological Society of London 1870: 728-747. Edwards, M. A. & Tobias, M. A. 1993. Nomenclator Zoologicus. The Zoological Society of London, 8 (1966-1977): 1-627 [269]. Edwards, M. A., Manly, P. & Tobias, M. A. 1996. Nomenclator Zoologicus. The Zoological Society of London, 9 (1978-1994): 1-747 [354]. Fletcher, D. S. 1979. In Nye, IWB. [ed.]. The Generic Names of Moths of the World, Vol. 3. Geometroidea. Publications British Museum of natural History. No. 812: 118. International Comission of Zoological Nomenclature. 1999. International Code of Zoological Nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature, London. Joseph, A. N. T. & Parui, P. 1983. A new species of Lobus Martin (Diptera: Asilidae) from S. India. Bulletin of the Zoological Survey of India, Calcutta 5 (1): 67-69. Joseph, A. N. T. & Parui, P. 1984. Three small, little-known genera of Asilidae (Diptera) from the Oriental Region. Bulletin of the Zoological Survey of India, Calcutta 6 (1-3): 247-252. Joseph, A. N. T. & Parui, P. 1991. On some Asilidae (Diptera) present in the B. P. Bishop Museum, Hololulu III. Bulletin of the Zoological Survey of India, Calcutta 88 (3-4): 241-253. Joseph, A. N. T. & Parui, P. 1992. Asilidae (Diptera) from Goa, Karnataka and Tamil. Records of the Zoological Survey of India 92 (1-4): 99-106.


Joseph, A. N. T. & Parui, P. 1998. The Fauna of India and adjacent Countries - Diptera (Asilidae) Part I. xvi + 278 pp. Calcutta. Martin, C. H. 1972. Genital morphology and species of the eastern hemisphere genus Lobus. Journal of Kansas Entomology Society 45: 7-17. Namkung, J. 2003. The Spiders of Korea, 2nd. ed. Kyo-Hak Publ. Co., Seoul, 648 pp. Neave, S. A. 1940. Nomenclator Zoologicus. The Zoological Society of London 3: 1-1065 [204]. Okuma, C. 1994. Spiders of the genera Episinus and Moneta from Japan and Taiwan, with descriptions of two new species of Episinus (Araneae: Theridiidae). Acta arachnologica Tokyo 43: 5-25. Scoble, M. J. (editor), Pitkin, L. M., Parsons, M., Honey, M. R. & Pitkin, B. R. 1999. Geometrid Moths of the World: A Catalogue. Apollo Books, Stenstrup. 1312 pp., 2 Volumes. Song, D. X., Zhu, M. S. & Chen, J. 1999. The Spiders of China. Hebei Sci. Technol. Publ. House, Shijiazhuang, 640 pp. Warren, W. 1906. New Drepanidea, Thyrididae, Uraniidae, and Geometridae from British New Guinea. Novitates Zoologicae 13: 61-161. Yoshida, H. 2001. A revision of the Japanese genera and species of the subfamily Theridiinae (Araneae: Theridiidae). Acta arachnologica Tokyo 50: 157-181. Yoshida, H. 2003. The spider family Theridiidae (Arachnida: Araneae) from Japan. Arachnological Society of Japan, 224 pp. Zhu, M. S. 1998. Fauna Sinica: Arachnida: Araneae: Theridiidae. Science Press, Beijing, xi + 436 pp.



HABITAT SELECTION AND DIVERSITY OF GROUND BEETLES (CARABIDAE) ON AHIR MOUNTAIN

(K.MARAŞ, TURKEY), IN MEDITERRANEAN REGION

Serap Avgın*

* Çukurova Üniversitesi, Fen-Edebiyat Fakültesi, Biyoloji Bölümü, Balcalı, Adana, TÜRKİYE. e-mail: [email protected] [Avgın, S. 2006. Habitat selection and diversity of ground beetles (Carabidae) in Ahır Mountain (Kahramanmaraş, Turkey), in Mediterranean Region. Munis Entomology & Zoology 1 (2): 257-266] ABSTRACT: This study was carried out on Ahır Mountain (Kahraman Maraş, Turkey), in the Mediterranean Region between April and October of 2004 and 2005. In this region, there is not any work about ground beetles (Coleoptera, Carabidae) so far. A total of 24 ground beetles species were recorded in the pitfall traps. Among them, Calathus (Calathus) libanensis pluriseriatus Putzeys, 1873 and Calathus (Calathus) syriacus Chaudoir, 1863 were the most abundant species. Separately, habitat selection (grassland, forest edge and forest interior) of all species, diversity, monthly distribution and chorotypes, ecology of these species are given in this study. It was recorded that the diversity of ground beetles was significantly higher in the forest edge and grassland than in the forest interior. There was no significant difference in the diversity of ground beetles of the grassland and the forest edge in Ahır Mountain. KEY WORDS : Carabidae, ground beetle, Ahır mountain.

Ahır mountain is the chain of the Southeast Toros mountains and this

area is located within the boundaries of Kahraman Maraş province, Turkey, in the Mediterranean. The altitude of Ahır mountain changes between 600 metre and 2301 metres. This area has a Mediterranean climate, the main characteristics of which are : dry summers and warm and rainy winters. The seasonal precipitation regime during the year is winter, spring, autumn and summer. This is a typical first variant of the East Mediterranean climate (Akman, 1990; Varol, 2003).

The study area lies within the Mediterranean phytogeographical region. This region is dominated by the forest species Pinus brutia Ten., Pinus nigra J.F. 7411 hectare forest field is present in Ahır mountain. 4532 hectare of this forest is fertile but 2879 hectare of it is infertile. This region is important for its forest in Turkey. The grass and bushes dominated in Mediterranean Region are found at the forest edge and unforest regions. In this region, research about insects is rare and there is not any work about ground beetles (Coleoptera, Carabidae) so far.

Ground beetles are very important on the biological agenda. They are generalist predators which can greatly reduce insect pest (Varchola & Dunn, 1999). The ground beetles have been an important study group in ecological research. Because they are diverse and abundant, their ecology and systematics are relatively well known, and they seem to be highly sensitive to habitat changes and, thus, good indicators of environmental change (Thiele, 1977; Ings & Hartley, 1999 ; Magura et al. 2000 ;


Melnychuk et al., 2003). There are many quantitative studies on the distribution of ground beetles in various habitats in the world. For example, some research has reported distribution of ground beetles in jack pine, oak and maple, and on aspen stands with various histories of defoliation and different soil types (Epstein & Kulman, 1990). It was discovered in a work on the effects of habitat structure on the ground beetle community composition of a regenerating native forest in the Deeside region of northeast Scotland (Ings & Hartley, 1999). Similar findings have been reported by Magura et al., 2000 and Gutierrez et al., 2004. In addition, much of the previous work using carabids as enviromental indicators has concentrated on moorland habitats (Gardner et al., 1997; Ingls & Hartley, 1999). Most studies on the carabid communities of woodlands have concentrated on the effects of management practices such as afforestation of uplands (Butterfield et al., 1995), soil scarificaiton (Parry & Rodger, 1986) and pest control (Walsh et. Al., 1993). However, only a few studies have been carried out in pine forest (Young & Armstrong, 1995; Ingls & Hartley, 1999).

In the literature much of the distribution of ground beetles in various habitats in the world were recorded (Epstein & Kulman, 1990; Gandhi et al., 2001; Armstrong & McKinlay, 1997; Allegro & Sciaky, 2003; Döring & Kromp, 2003).

Although many studies were carried out about the effect of different habitats, different climate, different altitude and different season on ground beetles in different countries, in Turkey such studies are very rare. The main aim of the present study is to determine habitat selection (grassland, forest edge and forest interior), diversity, monthly distribution and ecology of ground beetles in Ahır Mountain (Kahraman Maraş, Turkey), in the Mediterranean Region. The study focusses on the use of carabids as biological agenda forming the base. It also aimed to open the way to the works on the ecology of ground beetles in Turkey. This is the first study with this aim done about ground beetles in Turkey.

MATERIAL AND METHODS

The material of this investigation was collected from the Ahır mountain (Kahraman Maraş, Turkey) between April and October of 2004 and 2005. This region is mostly dominated by pine forest (P. brutia and P.nigra). Three habitats, forming a transect, were studied: 1). Forest interior, 2). Forest edge and 3). Grassland. Samples were collected using pitfall traps that were establihed in each of the 3 habitats. The pitfall traps consisted of plastic cups filled with 25 % ethylene glycol set into the ground. The traps were emptied monthly (Epstein & Kulman 1990, Armstrong & Mckinlay 1997, Magura et al. 2000). Specimens were dissected in the laboratory. Materials were then identified to species using keys in Müller (1926), Jeannel (1941), Lindroth (1974), Lindroth (1985), Trautner and Geigenmüller (1987), Hurka (1996). The identificaiton of those which weren’t identified were made with Prof. Augusto Vigna Taglianti and Dr.


Paolo Bonavita in Italy. Those which were already identified were checked in Italy with Prof. Augusto Vigna Taglianti and Dr. Paolo Bonavita. The specimens are deposited in the Biology Department of Çukurova University.

RESULTS

In this study many ground beetles collected are a new record for Ahır mountain. A total of 24 ground beetles species were recorded in the pitfall traps, between April and October of 2004 and 2005. The traps were emptied monthly. The species collected in pitfall traps every month are given in table 1. Among them, Calathus (Calathus) libanensis pluriseriatus Putzeys, 1873 and Calathus (Calathus) syriacus Chaudoir, 1863 were the most abundant species. The information collected about the ecology of ground beetles and chorotypes is given below.

Genus Brachinus Weber, 1801 Ecology: In this research, four species belong to this genus were collected in the traps forest edge and mostly in the region where grassland dominated.

“Bombardier Beetles”. These species are found often in open country on clayey soils, mostly in higher numbers. The larvae of all Brachinidae are probably parasitic (Trautner & Geigenmüller, 1987). Brachinus (Brachinus) ejaculans Fischer von Waldheim, 1828 Chorotype : Turano-European (Casale & Vigna Taglianti, 1999). Brachinus (Brachynidius) bodemeyeri Apfelbeck, 1904 Chorotype : Turano-European (Casale & Vigna Taglianti, 1999). Brachinus (Brachynidius) brevicollis Motschulsky, 1844 Chorotype : Turano- Mediterranean (Casale & Vigna Taglianti, 1999). Brachinus (Brachynidius) explodens Duftschmid, 1812 Chorotype : Asiatic-European (Casale & Vigna Taglianti, 1999).

Genus Calosoma Weber, 1801

Ecology: In this study, only one species belonging to this genus were collected in the traps put in the forest interior.

This genus is know from lowlands to mountains, more commonly only at the time of mass occurrence of lepidoptera forest pests (Hurka, 1996). They mostly prey on caterpillars. Larval development is very short, only 2-3 weeks. Larvae are predatory, too (Trautner & Geigenmüller, 1987). Calosoma sycophanta (Linné, 1758) Chorotype : Palearctic (Holarctic) (Casale & Vigna Taglianti, 1999).

Genus Carabus Linné, 1758 Ecology: In this study, three species belong to this genus were collected in the traps put at the edge and in the interior of the forest but the forest


interior traps, yielded. They preferably feed on earth-worms, snails, slugs, and numerous

insects, however, they do also feed on carrion and sometimes even on vegetable matter. Digestion is extra-intestinal. Usually single eggs are laid in the ground. The larval development takes several weeks. Larvae are predators. Pupation takes place in the ground. Almost all species are mainly active at night. For their defense they can spray acid fluids from their anal glands and salivate digestive ferment. Wings reduced in nearly all species (Lindroth, 1985; Trautner & Geigenmüller, 1987). Carabus (Mimocarabus) maurus paphius L. Redtenbacher, 1843 Chorotype : South-West Asiatic (Casale & Vigna Taglianti, 1999). Carabus (Chaetomelas) morawitzi montesamanus Mandl, 1967 Chorotype : South-East Anatolian (Casale & Vigna Taglianti, 1999). Carabus (Lamprostus) mulsantianus nurdagensis Blumenthal, 1967 Chorotype : South-East Anatolian (Casale & Vigna Taglianti, 1999).

Genus Leistus Fröhlich, 1799

Ecology: In this study, only one species belong to this genus was collected in the traps put both in the forest interior and the forest edge.

These species are found in different habitats, some species montane to subalpine. The species of this genus occur among debris in more or less shady places, where they prey upon mites, Collembola, etc. They are nocturnal (Lindroth, 1985; Trautner & Geigenmüller, 1987). Leistus (Pogonophorus) spinibarbis rufipes Chaudoir, 1843 Chorotype :Cyprus and Anatolian (Casale & Vigna Taglianti, 1999).

Genus Notiophilus Duméril, 1806

Ecology: In this study, only one species belonging to this genus was collected and it was collected in the trap which was put in the forest interior.

They live on the litter-layer of forests, heaths, or meadows, some at shores. They are active during the day and hunt by sight. The species are diurnal, sun-loving insects, very rapid in their movement. They are visually hunting beetles, preying upon mites, Collembola and other arthropods. The larvae have specialized on Collembola to a greater extent than have the adults (Lindroth, 1985; Trautner & Geigenmüller, 1987; Hurka, 1996). Notiophilus danieli Reitter, 1897 Chorotype : East Mediterranean (Casale & Vigna Taglianti, 1999).

Genus Metallina Motschulsky, 1850

Ecology: In this study, only one species belonging to this genus was collected and it was collected in the trap which was put in the region where grassland dominated.


They are Brachypterous, rarely macropterous. These live in drier to semi-moist and unshaded Calluna heaths, mostly on peaty soil and hills (Trautner & Geigenmüller, 1987; Hurka, 1996). Metallina (Neja) sporadica (J. Sahlberg, 1903) Chorotype : East Mediterranean (Casale & Vigna Taglianti, 1999).

Genus Amara Bonelli, 1810

Ecology: In this research, two species belonging to this genus were mostly collected in the traps at the forest edge and mostly in the region where the grassland dominated.

They are macropterous, observed in flight (Hurka, 1996). Many species are active at daytime. Imagines are partly or completely phytophagous, they preferably feed on flowers and seeds (e. g. of Cruciferes). Larvae are mostly predatory and live in burrows (Trautner & Geigenmüller, 1987). Very distributed and very common. A xerophilous species living in open country on dry, sandy or clayey soil with rather dense but short vegetation. Notably on heaths and grasssland; also on cultivated land, often on lawns in parks and in gardens. The beetle is pronouncedly diurnal, often seen running on the surface in bright sunshine. It feeds on plant seeds and has been reported to be harmful to winter cereals. Breeding takes places in spring (Lindroth, 1985). Amara (Amara) aenea (De Geer, 1774) Chorotype : Palearctic (Casale & Vigna Taglianti, 1999). Amara (Amara) proxima Putzeys, 1866 Chorotype : South European (Casale & Vigna Taglianti, 1999).

Genus Ophonus Dejean, 1821

Ecology: In this study, only one species belonging to this genus was collected in the traps at the forest ledge and mostly at the region where grassland dominated.

These species is brachypterous, rarely macropterous. It occurs in dry, open habitats, usually on gravelly, clay-mixed chalky soil with short, sparse vegetation, e.g. in dry meadows and grassland, often on southern hill-sides. The species is regularly found in company with Harpalus, Brachinus, etc. It is usually considered a typical spring breeder, but finds of newly emerged beetles in mid and late June suggest that autumn breeding may also occur (Lindroth, 1985; Trautner & Geigenmüller, 1987; Hurka, 1996). Ophonus (Hesperophonus) subquadratus (Dejean, 1829) Chorotype : Mediterranean (Casale & Vigna Taglianti, 1999).

Genus Harpalus Latreille, 1802

Ecology: In this study, only one species belonging to this genus were collected in the traps at the forest edge and mostly at the region where grassland dominated.


Many species are partly or preferably phytophagous, at least as imago (Trautner & Geigenmüller, 1987). These species is found in unshaded habitats; steppe, vineyards, fields, fallows; lowlands to hills (Hurka, 1996). Harpalus (Harpalophonus) metallinus Ménétriés, 1836 Chorotype : South-West Asiatic (Casale & Vigna Taglianti, 1999).

Genus Odotoncarus Solier, 1835

Ecology: In this study, only one species belonging to this genus was collected in the traps at the forest edge and mostly at the region where grassland dominated. Odotoncarus asiaticus (Chaudoir, 1852) Chorotype : East Mediterranean (Casale & Vigna Taglianti, 1999).

Genus Ditomus Bonelli, 1810

Ecology: In this research, only one species belonging to this genus was collected in the traps at the forest edge and mostly in the region where the grassland vegetation dominated.

These species are mostly phytophagous. They live mostly on sandy and partly saline soils near the coast and for aestivation in the dry summer plantago-seeds are collected and stored in the subterranean burrows (Trautner & Geigenmüller, 1987). Ditomus calydonius calydonius (P. Rossi, 1790) Chorotype : Cyprus and Anatolian (Casale & Vigna Taglianti, 1999).

Genus Dixus Billberg, 1820

Ecology: In this research, two species belonging to this genus were collected in the traps at the forest edge and mostly in the region where the grassland dominated.

These species are macropterous. They live on steppe, lowlands (Hurka, 1996). Dixus eremita (Dejean, 1825) Chorotype : East Mediterranean (Casale & Vigna Taglianti, 1999). Dixus obscurus (Dejean, 1825) Chorotype : East Mediterranean (Casale & Vigna Taglianti, 1999).

Genus Calathus Bonelli, 1810

Ecology: In this study, two species belonging to this genus were collected. These species were collected in the traps which were put mostly in the forest interior.

This commonly distributed species usually lives in open country on different kinds of moderately dry ground with sparse vegetation, achieving its greatest abundance on sandy soil. It is a common inhabitant of dry meadows, grassland, dunes and heaths; also on agricultural land and in thin forests, mainly of Pinus. It is frequent in the fields up to the lower


alpine region. It is probably chiefly carnivorous (Lindroth, 1985; Hurka, 1996). Calathus (Calathus) libanensis pluriseriatus Putzeys, 1873 Chorotype : Anatolian (Casale & Vigna Taglianti, 1999). Calathus (Calathus) syriacus Chaudoir, 1863 Chorotype : South-West Asiatic (Casale & Vigna Taglianti, 1999).

Genus Anchomenus Bonelli, 1810

Ecology: In this research, only one species belonging to this genus was collected in the traps at the forest edge and mostly in the region where the grassland dominated.

They are macropterous, observed in flight. They are in unshaded, dry to moderately moist habitats ; fields, steppe, pastures, edges of small woods ; from lowlands to mountains, often gregariously (Lindroth, 1985; Hurka, 1996). Anchomenus (Anchomenus) dorsalis (Pontoppidan, 1763) Chorotype : Palearctic (Casale & Vigna Taglianti, 1999).

Genus Microlestes Schmidt-Goebel, 1846

Ecology: In this research, only one species belonging to this genus was collected mostly in the region where grassland dominated.

This species is brachypterous, rarely macropterous. They live in rather dry habitats, indifferent to shade: steppe, pastures, brick-yards; lowlands to mountains, often in hills (Hurka, 1996). They are found preferably on sandy or clayey soils which are exposed to sunlight and at least partly poor in vegetation (Lindroth, 1985; Trautner & Geigenmüller, 1987). Microlestes maurus (Sturm, 1827) Chorotype : Turano European (Casale & Vigna Taglianti, 1999).

Genus Lamprias Bonelli, 1810

Ecology: In this research, only one species belonging to this genus was mostly collected in the region where grassland dominated.

Probably all species are parasites or macropterous, observed in flight. They rarely live in dry to moderatly moist, unshaded habitats : steppe, pastures ; lowlands to foothills. On low vegetation in open country, mostly in spring (Trautner & Geigenmüller, 1987; Hurka, 1996). Lamprias cyanocephalus (Linné, 1758) Chorotype : Palearctic (Casale & Vigna Taglianti, 1999).

DISCUSSION

In the recent studies, generally the ecology of carabids which have a

very important biological agenda has been more important rather than the systematic study of ground beetles. As in this research, much research on


the diverstiy of ground beetles and on their effect on habitat structure in the forest region of different countries has been investigated (Debinski & Brussard, 1994; Molnar et al., 2001; Gutierrez et al., 2004).

On Ahır mountain it was observed that the diversity of ground beetles was significantly higher in the forest edge and grassland than in the forest interior. There was no significant difference in the diversity of ground beetles of the grassland and the forest edge. Similar results were obtained from research carried out in oak- hornbeam forest in Hungary (Molnar et al., 2001). This also emphasized the edge effect on ground beetles, because forested habitats were usually significantly less diverse than the open areas (Magura & Tothmeresz, 1998; Molnar et al., 2001).

Also, the monthly distribution of carabids collected in pitfall traps was given in this research (table 1) and this information shows seasonal distribution of ground beetles on Ahır mountain. In addition, similar studies have been carried out in many countries, but the results showed some differences according the climate of that country (Epstein & Kulman, 1990; Armstrong & McKinlay, 1997).

ACKNOWLEDGEMENTS

I would like to thank Prof Augusto Vigna Taglianti (Italy) and Dr. Paolo Bonavita (Italy), for their valuable help and advice and The Scientific and Technical Research Council of Turkey (TUBITAK) for financial support with this research.

LITERATURE CITED

Akman, Y. 1990. İklim ve biyoiklim. Ankara : Palme Yayınları. Allegro, G. & Sciaky, R. 2003. Assessing the potential role of ground beetles (Coleoptera, Carabidae) as bioindicators in poplar stands, with a newly proposed ecological index (FAI). Forest Ecology and Management 175: 275-284. Armstrong, G. & McKinlay, R. G. 1997. Vegetation management in organic cabbages and pitfall catches of carabid ceetles. Agriculture, Ecosystems and Environment 64: 267-276. Butterfield, J., Luff, M. L., Baines, M. & Eyre, M. D. 1995. Carabid beetle communities as indicators of conservation potential in upland forest. For. Ecol. Manage 79: 63-77. Casale, A. & Vigna Taglianti, A. 1999. Caraboid Beetles (excl. Cicindelidae) of Anatolia, and their biogeographical significance (Coleoptera, Caraboidea). Biogeographia, Lavori della Società Italiana di Biogeografia 20: 277-406. Debinski, D. & Brussard, P. F. 1994. Using biodiversity data to assess species- habitat relationships in Glacier National Park, Montana. Ecological Applications 4: 833-843. Döring, T. F. & Kromp, B. 2003. Which carabid species benefit from organic agriculture?- a review of comparative studies in winter cereals from Germany and Switzerland. Agriculture, Ecosystems and Environment 98: 153-161.


Epstein, M. E. & Kulman, H. M. 1990. Habitat distribution and seasonal occurrence of carabid beetles in East- Central Minnesota. The American Midland Naturalist 123 : 209-225. Gandhi, K. J. K., Spence, J. R., Langor, D. W. & Morgantini, L. E. 2001. Fire residuals as habitat reserves for epigaeic beetles (Coleoptera : Carabidae and Staphylinidae). Biological Conservation 102 : 131-141. Gardner, S. M., Hartley, S. E., Davies, A. & Palmer, S. C. F. 1997. Carabid communities on heather moorlands in northeast Scotland: the consequences of grazing pressure for community diversity. Biol. Cons. 81: 275-286. Gutierrez, D., Menendez, R. & Mendez, M. 2004. Habitat-based conservation priorities for carabid neetles within whe Picos De Europa National Park, Northern Spain. Biological Conservation 115: 379-393. Hurka, K. 1996. Carabidae of The Czech and Slovak Republics. Kabourek Publishing, Zin: 1-565. Ings, T. C. & Hartley, S. E. 1999. The effect of habitat structure on carabid communities during the regeneration of a native scottish forest. Forest Ecology And Management 119 : 123-136. Jeannel, R. 1941. Faune De France, coleopteres carabiques. Premiere partie, Paris: 39: 1-571. Lindroth, C. H. 1974. Handbooks for the identification of British insect. Royal Entomological Society of London: 4 (2): 1-149. Lindroth, C. H. 1985. The carabidae (Coleoptera) of Fennoscandia and Denmark. fauna Entomologica Scandinavica. E.J. Brill/Scandinavian Science Press Ltd.: 15 (1): 1-225. Magura, T. & Tothmeresz, B. 1998. Edge effect on carabids in an oak-hornbeam forest at the Aggtelek National Park (Hungary), Acta Phytopathol. Entomology Hungary 33: 379-387. Magura, T., Tothmeresz, B. & Bordan, Zs. 2000. Effects of Nature Management Practice on Carabid Assemblages (Coleoptera : Carabidae) in a Non-Native Plantation. Biological Conservation 93: 95-102. Melnychuk, N. A., Olfert, O., Youngs, B. & Gillott, C. 2003. Abundance and diversity of carabidae (Coleoptera) in different farming systems. Agriculture, Ecosystems and Environment 95 : 69-72. Molnar, T., Magura, T., Tothmeresz, B. & Elek, Z. 2001. Ground beetles (Carabidae) and edge effect in oak-hornbeam forest and grassland transects. Eur. J. Soil. Biol. 37 : 297-300. Müler, D. G. 1926. I coleotteri della venezia giulia. catalogo ragionato. parte I. : Adephaga. Trieste. Tipografia Fratelli Mosettig: 1-304. Pary, W. H. & Rodger, D. 1986. The effect of soil scarification on the ground beetle fauna of a Caledonian pine forest. Scot. For. 40: 1-9. Thiele, H. U. 1977. Carabid beetles in their environments. Springer, Berlin. 369. Trautner, J. & Geigenmüller, K. 1987. Tiger beetles groud beetles, illustrated key to the cicindelidae and carabidae of Europe. Josef Margraf Publisher, Germany, 488.


Varchola, J. M. & Dunn, J. P. 1999. Changes in ground beetle (Coleoptera: Carabidae) assemblages in farming systems bordered by complex or simple roadside vegetation. Agriculture, Ecosystems and Environment 73: 41-49. Varol, Ö. 2003. Flora of Başkonuş Mountain (Kahramanmaraş). Turk Journal of Botanical 27: 117-139. Walsh, P. J., Day, K. R., Leather, S. R. & Smith, A. 1993. The influence of soil type and pine species on the carabid community of a plantation forest with a history of pine beauty moth infestation. Forestry 66: 135-146. Young, M. R. & Armstrong, G. 1995. The effect of age, stand density and variability on insect communities in native woodlands. In: Aldhous, J. R. (Ed.), Our Pinewood Heritage. Forestry Commission, the Royal Society for the Protection of Birds, Scottish Natural Heritage, Inverness, 206-221.

[Received December 2005. Accepted March 2006] Table 1. The carabids collected from Ahır Mountain (Kahramanmaraş, Turkey) between April and October of 2004 and 2005.


NOMENCLATURAL CHANGES IN CERAMBYCIDAE (COLEOPTERA)

Hüseyin Özdikmen*

* Gazi Üniversitesi, Fen-Edebiyat Fakültesi, Biyoloji Bölümü, 06500 Ankara / TÜRKİYE, e-mail: [email protected] [Özdikmen, H. 2006. Nomenclatural changes in Cerambycidae (Coleoptera). Munis Entomology & Zoology, 1 (2): 267-269] ABSTRACT: A replacement name, Fehmii nom. nov. is proposed for the preoccupied genus name Damiria Fairmaire, 1900 (Cerambycidae: Cerambycinae: Cleomenini). Also, a clarification on the validity of the generic name Zotalemimon Pic, 1925 is presented. KEY WORDS: Fehmii, Damiria, Zotalemimon, Diboma, homonymy, replacement name, Coleoptera, Cerambycidae.

Fehmii nom. nov., substitute name

Damiria Fairmaire, 1900. Bull. Soc. ent. France, 1900, 89. (Coleoptera: Chrysomeloidea: Cerambycidae: Cerambycinae: Cleomenini). Preoccupied by Damiria Keller, 1891. Z. wiss. Zool., 52, 308. (Porifera: Cellularia: Demospongiae: Ceractinomorpha: Poecilosclerida: Microcionina: Acarnidae).

Remarks: The name Damiria was initially introduced by Keller (1891) for a genus of the sponges family Acarnidae (with the type species Damiria simplex Keller, 1891). For the present, Anisotylacanthaea Vacelet, 1969; [Damira] de Laubenfels (lapsus) and Xytopsene de Laubenfels, 1936 are the junior names of Damiria Keller, 1891 (Hooper, 2000). Subsequently, Fairmaire (1900) described a beetle genus of the family Cerambycidae (with the type species Damiria perrieri Fairmaire, 1900 by monotypy) under the same generic name. Thus, the genus Damiria Fairmaire, 1900 is a junior homonym of the genus Damiria Keller, 1891. According to Article 60 of the International Code of Zoological Nomenclature, I propose for the genus Damiria Fairmaire, 1900 the new replacement name Fehmii nom. nov. The name dedicated to Fehmi Özdikmen. The name is masculine in gender. Distribution: Known from Madagascar. Summary of nomenclatural changes: Fehmii new replacement name = Damiria Fairmaire, 1900 (non Keller, 1891). Fehmii perrieri (Fairmaire, 1900) new comb. from Damiria.

Also, Damiria Keller, 1891 (Porifera) and Damiria Fairmaire,

1900 (Coleoptera) are recorded in Nomenclator Zoologicus vol. 2: 10 p. by Neave (1939).


A clarification on the validity of the generic name Zotalemimon Pic, 1925

Diboma Thomson, 1864. Syst. Cerambycid., 46. (Coleoptera: Chrysomeloidea: Cerambycidae: Lamiinae: Desmiphorini). Preoccupied by Diboma Walker, 1863. List Specimens Lep. Ins. Brit. Mus., 27, 8. (Lepidoptera: Pyraloidea: Pyralidae: Pyralinae).

Remarks: Thomson (1864) proposed the genus Diboma with the type species Diboma tranquilla Thomson, 1864 in the beetles family Cerambycidae. Unfortunately, the generic name was already preoccupied by Walker (1863), who had described the genus Diboma with the type species Diboma abscissalis Walker, 1863 by monotypy in the moth family Pyralidae. Thus, the genus Diboma Thomson, 1864 is a junior homonym of the generic name Diboma Walker, 1863. According to the International Code of Zoological Nomenclature, it must be rejected and replaced. However, any replacement name for Diboma Thomson, 1864 is unnecessary because there are apparently several synonyms. The generic names Zotalemimon Pic, 1925; Donysia Gressitt, 1940 and Sybrocentrura Breuning, 1947 are the junior synonyms of Diboma Thomson, 1864 (Kusama & Takakuwa, 1984; Heffern, 2005). So I propose to substitute name Zotalemimon Pic, 1925 that is available and the senior synonym of the generic name Diboma Thomson, 1864 for Diboma Thomson, 1864. Distribution: Known from Madagascar, Comores and SE Asia: beginning from southern Japan and Taiwan and extending as far south as Malaysia. Summary of nomenclatural changes: Zotalemimon Pic, 1925 = Diboma Thomson, 1864 (non Walker, 1863) = Donysia Gressitt, 1940 = Sybrocentrura Breuning, 1947. Zotalemimon tranquilla (Thomson, 1864) new comb. from Diboma. Zotalemimon procera (Pascoe, 1859) new comb. from Diboma. Zotalemimon costata (Matsushita, 1933) new comb. from Diboma. Zotalemimon posticata (Gahan, 1890) new comb. from Diboma. Zotalemimon borneotica (Breuning, 1969) new comb. from Diboma.

Also, Diboma Walker, 1863 (Lepidoptera) and Diboma Thomson,

1864 (Coleoptera) are recorded in Nomenclator Zoologicus vol. 2: 66 p. by Neave (1939).


LITERATURE CITED Breuning, S. 1947. Nouvelles formes de Longicornes du Musée de Stockholm. Arkiv för Zoologi. Stockholm. 39A (6): 1-68. Fairmaire, L. M. H. 1900. Descriptions de Coléoptères recueillis par M. H. Perrier de la Bâthie à Madagascar [Siagona, Tetragonoderus, Pygora, Acmaeodera, Myiodola, Damiria et Cassiodopsis]. Bulletin de la Société Entomologique de France 1900: 85-90. Gressitt, J. L. 1940. The longicorn beetles of Hainan Island. Philippine Journal of Science. Manila. 72 (1/2): 1-239. Heffern, D. J. 2005. Catalog and bibliography of Longhorned beetles from Borneo (Coleoptera: Cerambycidae). Available from: http://www.zin.ru/animalia/Coleoptera/ pdf/borneo_catalog_electronic_version_2005-1.pdf; Version January 2005. Hooper, J. N. A. 2000. “Sponguide”. Guide to Sponge collection and identification. Available from: http://www.qmuseum.qld.gov.au/organisation/sections/SessileMarine Invertebrates/spong.pdf ; Version August 2000). International Comission of Zoological Nomenclature. 1999. International Code of Zoological Nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature, London. Keller, C. 1891. Die Spongienfauna des rothen Meeres (2. Hälfte). Zeitschrift für Wissenschaftliche Zoologie 52: 294-368. Kusama, K. & Takakuwa, M. 1984. Parandrinae. Prioninae. Spondylinae. Aseminae. Lepturinae (part.). Cerambycinae. Lamiinae (part.). The Longicorn-beetles of Japan in Color. Kodansha, Tokyo: 131-172, 201-493, 511-549. Neave, S. A. 1939. Nomenclator Zoologicus. The Zoological Society of London 2: 1-1025 [10, 66]. Pic, M. 1925. Nouveautés diverses. Mélanges Exotico-Entomologiques. Moulins. 44: 1-32. Thomson, J. 1864. Systema cerambycidarum ou expose de tous les genres compris dans la famille des cerambycides et familles limitrophes. Mémoire de la Société royale des sciences de Liège 19: 1-540. Walker, F. 1863. List of the specimens of lepidopterous insects in the collection of the British Museum. Part XXVII. Crambites & Tortricites. British Museum (Natural History), London, 1-286 pp.

[Received January 2006. Accepted February 2006]


NEW SUBSTITUTE NAMES FOR TWO PREOCCUPIED TACHINID GENERA PARAGONIA MESNIL, 1950 AND

MENETUS ALDRICH, 1926 (DIPTERA)

Hüseyin Özdikmen*

* Department of Biology, Faculty of Science and Arts, University of Gazi, 06500 Ankara / Turkey, e-mail: [email protected] [Özdikmen, H. 2006. New substitute names for two preoccupied Tachinid genera Paragonia Mesnil, 1950 and Menetus Aldrich, 1926 (Diptera). Munis Entomology & Zoology, 1 (2): 270-272] ABSTRACT: Two junior homonyms were detected amongst the Tachinidae and the following replacement names are proposed: Mesnilius for Paragonia Mesnil, 1950 and Aldrichomyia for Menetus Aldrich, 1926. Accordingly, new combinations are herein proposed for the type species currently included in monotypic genera Paragonia Mesnil, 1950 and Menetus Aldrich, 1926. Keywords: Mesnilius, Paragonia, Aldrichomyia, Menetus, homonymy, replacement names

TAXONOMY Order Diptera

Family Tachinidae Mesnilius nom. nov., substitute name

Paragonia Mesnil, 1950. In Lindner, Flieg. palaearkt. Reg. Stuttgart, 64g, 106 (1950). (Diptera: Tachinidae: Goniinae: Sturmiini). Preoccupied by Paragonia Huebner, [1823]. Verz. bekannt. Schmett., (19) 292. (Lepidoptera: Geometroidea: Geometridae: Ennominae).

Remarks: Mesnil (1950) proposed the genus Paragonia with the type species Paragonia portentosa Mesnil, 1950 by original designation in the fly family Tachinidae (Diptera) (Cantrell & Crosskey, 1989; Evenhuis, 1996; O’Hara & Wood, 2004; O’Hara, 2005). Unfortunately, the generic name was already preoccupied by Huebner (1823), who had described the genus Paragonia with the type species Phalaena tasima Cramer, 1779 by monotypy in the moth family Geometridae (Lepidoptera). Scoble et al. (1999) included 9 species and 2 subspecies (inc. nominates) in the genus Paragonia Huebner, 1823 and also Clysia Guenée, 1858 as a junior synonym of the genus Paragonia. Thus, Paragonia Mesnil, 1950 is invalid under the law of homonymy, being a junior homonym of Paragonia Huebner [1823]. In accordance with article 60 of the International Code of Zoological Nomenclature, fourth edition (1999), I propose to substitute the junior homonym Paragonia Mesnil, 1950 for the nomen novum Mesnilius. Etymology: from L. P. Mesnil who current author name of preexisting genus Paragonia. The name is masculine in gender.


Distribution: Known from Australia (WA). Summary of nomenclatural changes; Mesnilius new replacement name = Paragonia Mesnil, 1950 (non Huebner, [1823]) Mesnilius portentosa (Mesnil, 1950) new comb. from Paragonia Mesnil, 1950.

Aldrichomyia nom. nov., substitute name

Menetus Aldrich, 1926. Trans. Amer. ent. Soc., 52, 23. (Diptera: Brachycera: Tachinidae: Dexiinae: Voriini). Preoccupied by Menetus Adams & Adams, 1855. Gen. Moll., 2, 262. (Mollusca: Gastropoda: Pulmonata: Hygrophila: Planorboidea: Planorbidae).

Remarks: The name Menetus was initially introduced by Adams & Adams (1855) for a genus of the gastropod family Planorbidae (with the type species Menetus opercularis (Gould, 1847)). At present, the genus Menetus Adams & Adams, 1855 includes 5 species and 2 subspecies as Menetus alabamensis Pilsbry, 1895; Menetus brogniartianus (I. Lea, 1842); Menetus dilatatus dilatatus (Gould, 1841); Menetus dilatatus avus Pilsbry, 1905; Menetus floridensis F. C. Baker, 1945; Menetus opercularis (Gould, 1847) and Menetus sampsoni Ancey, 1885 (Baker, 1945; Thompson, 1983; Burch, 1989; Turgeon et al., 1998). Subsequently, Aldrich (1926) described a new nearctic fly genus of the family Tachinidae (with the type species Brachicoma macropogon Bigot, 1889 by monotypy) under the same generic name (O’Hara & Wood, 2004). Thus, Menetus Aldrich, 1926 is invalid under the law of homonymy, being a junior homonym of Menetus Adams & Adams, 1855. In accordance with article 60 of the International Code of Zoological Nomenclature, fourth edition (1999), I propose for the genus Menetus Aldrich, 1926 the new replacement name Aldrichomyia nom. nov. Etymology: from Aldrich who current author name of preexisting genus Menetus. The name is masculine in gender. Distribution: Known from nearctic region. Summary of nomenclatural changes: Aldrichomyia new replacement name = Menetus Aldrich, 1926 (non Adams & Adams, 1855). Aldrichomyia macropogon (Bigot, 1889) new comb. from Menetus Aldrich, 1926.

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LITERATURE CITED Adams, H. & Adams, A. 1855. The Genera of Recent Mollusca; Arranged According to their organization. London : John Van Voorst, 2: 93-284. Aldrich, J. M. 1926. Notes on muscoid flies with retracted hind crossvein, with key and several new genera and species. Trans. Am. Entomol. Soc. 52: 7-28. Baker, F. C. 1945. The Molluscan Family Planorbidae. The University of Illinois Press, Urbana, Illinois, USA, 530 + xxxvi. Burch, J. B. 1989. North American Freshwater Snails. Malacological Publications, Hamburg, Michigan, USA, p. 365. Cantrell, B. K. & Crosskey, R. W. 1989. Family Tachinidae. Pp. 733–784. In Evenhuis, N.L., ed., Catalog of the Diptera of the Australasian and Oceanian Regions. Bishop Museum Special Publication 86. Bishop Museum Press and E.J. Brill. 1155 pp. Evenhuis, N. L. 1996. Revised online version of Cantrell and Crosskey (1989). Website address: http://hbs.bishopmuseum.org/aocat/tachinidae.html. (Web page dated 11 November 1996, accessed 19 January 2005.) Huebner, J. 1816-1826. Verzeichniss bekannter Schmetterlinge. Augsberg, J. Hübner. International comission of zoological nomenclature. 1999. International Code of Zoological Nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature, London. 306 pp. Mesnil, L. P. 1950. Larvaevorinae (Tachininae). Fliegen Palaearkt. Reg. 64g: 105-60. Scoble, M. J. (editor), Pitkin, L. M., Parsons, M., Honey, M. R. & Pitkin, B. R. 1999. Geometrid Moths of the World: A Catalogue. Apollo Books, Stenstrup. 1312 pp., 2 Volumes. O'Hara, J. E. 2005. World genera of the Tachinidae (Diptera) and their regional occurrence. PDF document, 69 pp. Available from: http://www.nadsdiptera.org/Tach/Genera/Gentach.pdf O'Hara, J. E. & Wood, D. M. 2004. Catalogue of the Tachinidae (Diptera) of America north of Mexico. Memoirs on Entomology, International 18. 410 pp. O'Hara, J. E. & Wood, D. M. 2004. Checklist of the Tachinidae (Diptera) of America north of Mexico. PDF document, 42 pp. Available from: http://www.nadsdiptera.org/Tach/Cklist/Cktach.pdf (Web page dated 28 January 2004) Thompson, F. G. 1983. The planorbid snail Miromenetus dilatatus avus (Pilsbry) in the West Indies and Central America Nautilus, 97 (2): 68-69. Turgeon, D. D., Quinn, J. F. Jr., Bogan, A. E., Coan, E. V., Hochberg, F. G., Lyons, W. G., Mikkelsen, P. M., Neves, R. J., Roper, C. F. E., Rosenberg, G., Roth, B., Scheltema, A., Thompson, F. G., Vecchione, M. & Williams, J. D. 1998. Common and scientific names of aquatic invertebrates from the United States and Canada: Mollusks. 2nd Edition. American Fisheries Society, Special Publication 26:ix-526.

[Received January 2006. Accepted February 2006]

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ADDITIONAL NOTES ON AQUATIC COLEOPTERA FAUNA OF TURKEY WITH A NEW RECORD

(HELOPHORIDAE, HYDROPHILIDAE)

Suat Kıyak*, Savaş Canbulat**, Ali Salur*** and Mustafa Darılmaz*

* Gazi University, Sciences and Arts Faculty, Department of Biology, 06500 Ankara / TURKEY. e-mail: [email protected], [email protected] ** Sakarya University, Arts and Sciences Faculty, Department of Biology, Sakarya / TURKEY. e-mail: [email protected] *** Hitit University, Sciences and Arts Faculty, Department of Biology, 19030, Çorum / TURKEY. e-mail: [email protected] [Kıyak, S., Canbulat, S., Salur, A. & Darılmaz, M. 2006. Additional notes on aquatic Coleoptera fauna of Turkey with a new record (Helophoridae, Hydrophilidae). Munis Entomology & Zoology, 1 (2): 273-278]

ABSTRACT: In this study, 31 aquatic beetle specimens collected from the provinces Denizli, Aydın, Isparta and Antalya in south-west Mediterranean region of Turkey, between 2000–2002 have been established faunistically. It has been determined that they belong to 13 species of 7 genera of 2 families. 7 of these examined species are new records for south-west Mediterranean region and Helophorus (s.str.) grandis Illiger, 1798 is a new record for Turkey. KEY WORDS: Coleoptera, Helophoridae, Hydrophilidae, fauna, new record, Turkey.

A total of 34 genera of aquatic Hydrophilidae are known from the

Palaearctic and Oriental regions. So far, 15 genera and 77 species of Hydrophilidae have been recorded from Turkey (Gentili & Chiesa, 1975; Lenistea, 1978; Gentili, 1979, 2000; Henegouwen, 1986; Hansen, 1987; Hebauer, 1993, 1994; Ribera et al., 1997; Schödl, 1997, 1998; Cuppen & Van Maanen, 1998; İncekara et al., 2003a, 2003b, 2004a, 2005a, 2005b; Mart et al., 2003; Hansen, 2004; Hebauer & Ryndevich, 2005; Darılmaz & Kıyak, 2006).

The Helophoridae is also a large family comprising of a single subfamily containing a single genus, Helophorus Fabricius. This genus includes about 190 species (Angus, 1984, 1985, 1988, 1992; Smetana, 1985). 51 species have been recorded belonging to the six subgenera in Turkey, previously (Lenistea, 1978; Angus, 1984, 1985, 1988, 1992; Hebauer, 1994; Mart & Erman, 2001; Hansen, 2004; İncekara et. al. 2004b, 2005b, 2005c, 2005d; Darılmaz & Kıyak, 2006). With this study, one new record was added to the Turkish Helophoridae fauna.

METHODS

All samples were collected from south-west Mediterranean region in the years 2000-2002. The samples were collected from spring water areas, with either a sieve, ladle or water-net having a 1 mm mesh size. The beetles were killed with 70% alcohol and in the laboratory cleaned of


debris with a small paintbrush. The aedeagophore was dissected under a stereo-microscope and left in 10% KOH solution for about 1–2 hours. Materials have been deposited in the Zoological Museum of Gazi University (=ZMGU), Ankara, Turkey.

Family Helophoridae

Helophorus (Helophorus) grandis Illiger 1798

Materials: Antalya: 1 male, Korkuteli, the stream flow to Korkuteli dam, 37º05’N 30º08’E, 1074 m, 22.04.2001. Remarks: Helophorus grandis Illiger 1798 is here recorded from Turkey for the first time. H. grandis is a west and central European species, occurring throughout Britain and Ireland, France except the Mediterranean part, the northern half of Spain, Germany, Switzerland, Denmark, southern Sweden, south west Finland, Norway, Latvia, Estonia, Lithuania, Poland and in Russia north east to Syktivkar and south to about Moscow. Outside Europe H. grandis occurs in the Atlas Mountains of Morocco and it has also been introduced to eastern North America (Smetana, 1985; Angus, 1992).

Helophorus (Helophorus) syriacus Kuwert, 1885

Materials: Denizli: 1 female, Buldan, Süleymanlı lake, 38º03’N 28º46’E, 1175 m, 19.05.2001. Distribution in Turkey: Adana, Antakya, Diyarbakır, Mardin, Amanos Mountains (Gavur Mountains) (Angus, 1988, 1992; Mart & Erman, 2001). Remarks: New for south-west Anatolia.

Helophorus (Rhopalohelophorus) griseus Herbst, 1793

Materials: Aydın: 2 females, Söke, Sarıkemer, Büyük Menderes, 37º32’N 27º22’E, 21 m, 20.05.2001. Distribution in Turkey: Bolu, Bursa, Edirne, Erzincan, Erzurum, Gümüşhane, İstanbul, Kayseri (Angus 1988; Mart & Erman, 2001; İncekara et al., 2004b). Remarks: New for south-west Anatolia.

Helophorus (Rhopalohelophorus) discrepans Rey, 1885

Materials: Antalya: 1 male, Kaş, Gömbe, Girdev lake, 36º40’N 29º40’E, 1815 m, 24.06.2000. Distribution in Turkey: Ankara, Ardahan, Artvin, Bolu, Erzincan, Erzurum, Hakkâri, Kayseri, Van (Angus, 1988, 1992; Mart & Erman, 2001; İncekara et al., 2004b). Remarks: New for south-west Anatolia.


Helophorus (Rhopalohelophorus) longitarsis Wollaston 1864 Materials: Aydın: 2 males, 4 females, Akeniköy, 37º30’N 27º19’E, 60 m, 16.04.2000. Distribution in Turkey: Aksaray, Ankara, Burdur, Erzincan (Angus, 1988; Mart & Erman, 2001; İncekara et al., 2004a; Darılmaz & Kıyak, 2006). Remarks: New for south-west Anatolia.

Family Hydrophilidae

Berosus (Enoplurus) spinosus (Steven, 1808) Materials: Denizli: 1 female, Çivril, Bucak, the west of Işıklı lake, 38º14’N 29º51’E, 838 m, 26.06.2001. Distribution in Turkey: Adana, Afyon, Aksaray, Ankara, Antalya, Aydın, Edirne, Elazığ, İçel, Kars, Kırşehir, Malatya, Van (Cuppen & Van Maanen, 1998; İncekara et al., 2003a; Darılmaz & Kıyak, 2006). Remarks: New for Denizli.

Laccobius (Dimorpholaccobius) simulatrix d'Orchymont, 1932 Materials: Aydın: 1 male, 2 females, Söke, Sarıkemer, Büyük Menderes, 37º32’N 27º22’E, 21 m, 20.05.2001. Distribution in Turkey: Ağrı, Aksaray, Ankara, Antalya, Artvin, Aydın, Balıkesir, Bayburt, Bitlis, Bolu, Bursa, Çanakkale, Çorum, Denizli, Edirne, Erzincan, Erzurum, Hakkâri, Isparta, İçel, İstanbul, İzmir, Kahramanmaraş, Kars, Kayseri, Kırklareli, Kırşehir, Manisa, Muğla, Niğde, Osmaniye, Samsun, Sivas, Trabzon, Van, (Gentili & Chiesa, 1975; Gentili, 2000; İncekara et al., 2003a; Mart et al., 2003; Darılmaz & Kıyak, 2006). Remarks: Known to Aydın.

Laccobius (Dimorpholaccobius) sulcatulus Reitter, 1909

Materials: Antalya: 1 male, Korkuteli, Öküzgözü I Dam, 36º53’N 30º02’E, 1200 m, 22.04.2001. Distribution in Turkey: Amasya, Ankara, Antalya, Ardahan, Bayburt, Bitlis, Burdur, Diyarbakır, Erzincan, Erzurum, Gümüşhane, Kars, Konya, Manisa, Toros Mountains, Van (Gentili & Chiesa, 1975; Gentili, 1979; Gentili, 2000; İncekara et al., 2003a; Mart et al., 2003; Hansen, 2004). Remarks: Known to Antalya.

Enochrus (Lumetus) bicolor (Fabricius, 1792)

Materials: Denizli; 1 male, 1 female, Çivril, near of the Beydilli village, Işıklı Lake, 38º15’N 29º55’E, 840 m, 20.06.2002. Distribution in Turkey: Aksaray, Ankara, Antalya, Erzincan, İçel, Kars, Kırşehir, Van (Lenistea, 1978; Hansen, 1987; Schödl, 1998; İncekara et al., 2003a, 2005a; Darılmaz & Kıyak, 2006). Remarks: New for Denizli.


Enochrus (Lumetus) fuscipennis (Thomson, 1884) Materials: Denizli; 4 females, Çivril, near of the Beydilli village, Işıklı Lake, 38º15’N 29º56’E, 850 m, 18.05.2001; Aydın: 4 females, Söke, Avşar, Azap Lake, 37º35’N 27º26’E, 37 m, 23.06.2001. Distribution in Turkey: Aksaray, Artvin, Erzincan, Erzurum, Rize (İncekara et al., 2005a, Darılmaz & Kıyak, 2006). Remarks: New for south-west Anatolia.

Limnoxenus niger (Gmelin, 1790)

Materials: Aydın: 1 male, 1 female, Söke, Yeşilköy, Azap Lake, 37º34’N 27º26’E, 17 m, 23.06.2002. Distribution in Turkey: Without detailed locality (İncekara et al., 2003a). Remarks: Probably with a new record for south-west Anatolia of Turkey.

Hydrochara caraboides (Linnaeus, 1758)

Materials: Isparta: 1 male, Senirkent, Gençali village, Eğirdir Lake, 38º14’N 30º45’E, 950 m, 21.06.2000. Distribution in Turkey: İzmir (İncekara et al., 2003a). Remarks: New for south-west Anatolia.

Hydrophilus (Hydrophilus) piceus (Linnaeus, 1758)

Materials: Denizli; 1 male, 1 female, Çivril, near of the Beydilli village, Işıklı Lake, 38º11’N 30º03’E, 920 m, 22.06.2000. Distribution in Turkey: Erzurum (İncekara et al., 2003a). Remarks: New for south-west Anatolia.

LITERATURE CITED

Angus, R. B. 1984. Towards a Revision of the Palearctic Species of Helophorus F. (Coleoptera, Hydrophilidae) I, Entomological Review 63 (3): 89-119. Angus, R. B. 1985. Towards a Revision of the Palearctic Species of Helophorus F. (Coleoptera, Hydrophilidae) II, Entomological Review 64 (4): 128-162. Angus, R. B. 1988. Notes on the Helophorus (Coleoptera, Hydrophilidae) occurring in Turkey, Iran and neighbouring territories, Revue suisse de Zoologie 95 (1): 209-248. Angus, R. B. 1992. Insecta, Coleoptera, Hydrophilidae, Helophorinae. Süsswasserfauna von Mitteleuropa 20/10-2, Gustav Fischer Verlag, Jena, 144 pp. Cuppen, J. G. M. & Van Maanen, B. 1998. Distribution and habitats of Berosus in The Netherlands (Coleoptera: Hydrophilidae), Entomologische Berichten (Amsterdam) 58 (11): 213-223. Darılmaz, M. & Kıyak, S. 2006. A contribution to the knowledge of the Turkish water beetles fauna (Coleoptera), Munis Entomology & Zoology 1 (1): 129-144.


Gentili, E. & Chiesa, A. 1975. A Revisione dei Laccobius paleartici (Coleoptera: Hydrophilidae), Estratto dalle Memorie della Società Entomologica Italiana 54: 1-187. Gentili, E. 1979. Aggiunte alla revisione dei Laccobius paleartici (Coleoptera, Hydrophilidae), Estratto dal Bollettino della Societa' Entomologica Italiana 111 (1-3): 43-50. Gentili, E. 2000. Distribuzione Del Genere Laccobius (Coleoptera, Hydrophilidae), In Anatolia E Problemi Relativi. Biogeographia XXI: 173-214. Hansen, M. 1987. The Hydrophiloidea (Coleoptera) of Fennoscandia and Denmark Fauna. Entomologica Scandinavica 18. E. J. Brill. Leiden, The Netherlands 254. pp. Hansen, M. 2004. Hydrophilidae. In Löbl, I. & Smetana, A. 2004. Catalogue of Palaearctic Coleoptera. Volume 2. Hydrophiloidea – Histeroidea – Staphylinoidea. Apollo Books, Stenstrup. 942 pp. Hebauer, F. & Ryndevich, S. K. 2005. New data on the Distribution of Old World Hydrophilidae (Coleoptera), Acta Coleopterologica XXI (1): 43-51. Hebauer, F. 1993. European Chaetarthria, Latissimus 3: 1-3. Hebauer, F. 1994. The Hydrophiloidea of Israel and the Sinai (Coleoptera, Hydrophiloidea), Zoology in the Middle East 10: 73-137. Henegouwen, A. B. 1986. Revision of the European species of Anacaena Thomson (Coleptera, Hydrophilidae), Entomologica Scandinavica 17: 393-407. İncekara, Ü., Mart, A. & Erman, O. 2003a. Check list of the Hydrophilidae (Coleoptera) Species of Turkey and a New Record for the Turkish Fauna, Turkish Journal of Zoology 27: 47-53. İncekara, Ü., Mart, A. & Hayat, R. 2003b. A new record for the Turkish fauna: Cercyon ustulatus (Preyssler, 1790) (Coleoptera, Hydrophilidae), Türkiye Entomoloji Dergisi 27 (2): 91-94. İncekara, Ü., Mart, A. & Erman, O. 2004a. First records of Anacaena lutescens (Stephens, 1829) and Cercyon littoralis (Gyllenhal, 1808) from Turkey (Coleoptera, Hydrophilidae), Zoology in the Middle East 31: 103-106. İncekara, Ü., Mart, A. & Erman, O. 2004b. Distribution of Turkish Helophorus Fabricius, 1775 (Coleoptera, Helophoridae) I. Subgenus Rhopalhelophorus, with Two New Records, Journal of the Entomological Research Society 6 (2): 51-62. İncekara, Ü., Mart, A. & Erman, O. 2005a. Studies on Turkish Hydrophilidae (Coleoptera) I. Genus Enochrus Thomson, 1859, Turkish Journal of Zoology 29: 155-158. İncekara, Ü., Mart, A. & Erman, O. 2005b. Some Notes on Two Newly Recorded Aquatic Coleoptera (Hydrophilidae, Helophoridae) Species from Turkey, Fırat Üniversitesi Fen ve Mühendislik Bilimleri Dergisi 17 (2): 449-454. İncekara, Ü., Mart, A. & Erman, O. 2005c. Distribution of Turkish Helophorus Fabricius, 1775 (Coleoptera, Helophoridae) II. Subgenus Atracthelophorus, with Two New Records and Some Ecological Notes, Turkish Journal of Zoology 29: 237-242. İncekara, Ü., Mart, A. & Erman, O. 2005d. Helophorus erzurumicus sp. n., a new species from Turkey and some notes on H. ponticus Angus, 1988 (Coleoptera: Hydrophilidae), Aquatic Insects 27 (1): 43–46.


Lenistea, M. A. 1978. Hydradephaga und Palpicornia. pp. 291-314. In, Illies, J. (Editor). Limnofauna Europea. Gustav Fischer Verlag, The Netherlands. 521 pp. Mart, A. & Erman, O. 2001. A Study on Helophorus Fabricius, 1775 (Coleoptera, Hydrophilidae) Species, Turkish Journal of Zoology 25: 35-40. Mart, A., Incekara, Ü. & Erman, O. 2003. A New Species of Laccobius Erichson, 1837 (Coleoptera: Hydrophilidae: Hydrophilinae) from Turkey, Aquatic Insects 25 (1): 75-80. Ribera, I., Schodl, S. & Hernando, C. 1997. Enochrus ater (Kuwert) and E. salomonis (Sahlberg) (Coleoptera: Hydrophilidae), two widespread but overlooked species new to the European fauna, Hydrobiologia 354: 183-188. Schödl, S. 1997. Taxonomic studies on the genus Enochrus (Coleoptera: Hydrophilidae), Entomological Problems 28 (1): 61-66. Schödl, S. 1998. Taxonomic revision of Enochrus (Coleoptera: Hydrophilidae) I. The E. Bicolor species complex, Entomological Problems 29 (2): 111-127. Smetana, A. 1985. Revision of the subfamily Helophorinae of the Nearctic region (Coleoptera: Hydrophilidae), Memoirs of the Entomological Society of Canada 131: 1-154.

[Received January 2006. Accepted April 2006]


A ZOOGEOGRAPHICAL REVIEW OF SPONDYLIDINAE IN TURKEY (COLEOPTERA: CERAMBYCIDAE)

Hüseyin Özdikmen* and Semra Turgut* * Gazi Üniversitesi, Fen-Edebiyat Fakültesi, Biyoloji Bölümü, 06500 Ankara, TÜRKİYE, e-mails: [email protected] and [email protected] [Özdikmen, H. & Turgut, S. 2006. A zoogeographical review of Spondylidinae in Turkey (Coleoptera: Cerambycidae). Munis Entomology & Zoology 1 (2): 279-288] ABSTRACT: The taxa of Turkish Spondylidinae fauna that were recorded by various authors from different localities of Turkey are evaluated zoogeographically with the present paper. For this reason, the distribution patterns in Turkey and rest of the world for each taxon of related subfamily are determined. Also, chorotypes for each taxon are given in the parts of zoogeographical remarks. Key Words: Coleoptera, Cerambycidae, Spondylidinae, Turkey, Zoogeography, Fauna

The longhorn beetles or Cerambycidae are classified together with Chrysomelidae and Bruchidae in the superfamily Chrysomeloidea. But, some authors recognized Cerambycidae as a separate superfamily Cerambycoidea (Svacha & Danilevsky, 1986). Cerambycidae sensu stricto is divided into several subfamilies. These are Parandrinae, Prioninae, Lepturinae, Necydalinae, Spondylidinae, Apatophyseinae, Cerambycinae and Lamiinae. Parandrinae are not represented in Turkey. Moreover, the subfamilies Necydalinae and Apatophyseinae are represented only a few species in Turkey. The Turkish Spondylidinae fauna has not been evaluated as detailed by any authors yet. The works on the fauna of Spondylidinae in Turkey have been carried out especially towards late second half of the 20. century mostly faunistically. These are: Fairmaire (1884), Schimitschek (1944), Alkan (1946), Erdem (1947), Acatay (1948, 1961, 1963 and 1968), Defne (1954), Çanakçıoğlu (1956 and 1983), Demelt & Alkan (1962), Demelt (1963 and 1967), Villiers (1967), Erdem (1968), Besçeli (1969), Gfeller (1972), Gül-Zümreoğlu (1972 and 1975), Tosun (1975), Erdem & Çanakçıoğlu (1977), Lobanov et al. (1981), Sekendiz (1981), Sama (1982 and 2002), Danilevsky & Miroshnikov (1985), Svacha & Danilevsky (1986), Öymen (1987), Önder et al. (1987), Adlbauer (1988 and 1992), Sama & Rapuzzi (1993), Yüksel (1996), Lodos (1998), Tauzin (2000), Alkan (2000), Tozlu (2001), Tozlu et al. (2002), Danilevsky (2005), Özdikmen & Çağlar (2004), Özdikmen & Şahin (2005), Malmusi & Saltini (2005), Özdikmen (2006). But, any work included the subfamily Spondylidinae completely is presently related to Turkish fauna. The known taxa of Spondylidinae in the studies above have been given together with other examined Cerambycidae taxa by various authors step by step. So nowadays, such data about the Turkish Spondylidinae fauna have reached a considerable level. The main aim of the present paper is to


evaulate the Turkish Spondylidinae fauna zoogeographically and faunistically by using known data. In this paper classification and nomenclature of the longhorn beetles suggested by Danilevsky, 2005 and Althoff & Danilevsky, 1997 are followed. All genera are listed in the same order as in Danilevsky (2005), Althoff & Danilevsky (1997) and Sama (2002). Within the genera the species are listed alphabetically. Each name of a species or subspecies is accompained by the author’s name and description date. The data, “Current records from Turkey”, “Range” and “Zoogeographical Remarks” under the title for each taxon is given in this present text. The present zoogeographical characterization is based on the chorotype classification of Anatolian fauna, recently proposed by Vigna Taglianti et al. (1999).

Subfamily Spondylidinae

Tribe Saphanini Genus Drymochares Mulsant, 1847

Drymochares starcki Ganglbauer, 1888 The species has three subspecies in the World. These are nominate Drymochares starcki starcki Ganglbauer, 1888; Drymochares starcki cavazzutii Sama & Rapuzzi, 1993 and Drymochares starcki ivani Sama & Rapuzzi, 1993. Drymochares starcki starcki Ganglbauer, 1888 occurs only in Caucasus. Drymochares starcki cavazzutii is known from Abchazia, Armenia and north-eastern Turkey. Drymochares starcki ivani is only known from northern part of Turkey (Sama & Rapuzzi, 1993). These taxa may be distinguished by the different pubescence of elytra. Tauzin (2000) mentioned that Ilgaz Mountains in Kastamonu province was the hybridization area for the two subspecies occurring in Turkey.

Drymochares starcki cavazzutii Sama & Rapuzzi, 1993 Current records from Turkey: Artvin prov.: pass between from Borçka and Hopa (type locality); Turkey (Lobanov et al., 1981); Artvin prov.: from Borçka to Hopa (Sama, 1982); Northern Turkey (Danilevsky & Miroshnikov, 1985); Eastern Black Sea Region (Svacha & Danilevsky, 1986); Artvin prov.: Çifteköprüler and Murgul (Petek forest); Gümüşhane prov.: Zigana Mts. (Adlbauer, 1988; Sama & Rapuzzi, 1993); Giresun prov.: Dereli; Ordu prov.: Harçbeli; Hasancık; Gölköy; Gürgentepe; Mesudiye; Tokat prov.: Niksar; Samsun prov.: Yenigün, Alaçam; Sinop prov.: Dranaz Mt., Yenikonak; Kastamonu prov.: Çatalzeytin, between Azdavay and Pınarbaşı, Ilgaz Mts. (Sama & Rapuzzi, 1993); Samsun prov.: Ladik and ?Layvikent (=Yakakent); Gümüşhane prov.: Şiran; Artvin prov.: Murgul; Ordu prov.: Akkuş; Kastamonu prov.: Ilgaz Mts. (Tauzin, 2000); Samsun prov.: Kavak, Hacılar pass; Artvin prov.: Cankurtaran pass (Malmusi & Saltini, 2005). Range: Turkey, Caucasus (Sama & Rapuzzi, 1993). Zoogeographical remarks: The subspecies is distributed in Turkey from Artvin to Kastamonu provinces in North Turkey. All old records before 1993 from Turkey should belong to this subspecies.

Drymochares starcki ivani Sama & Rapuzzi, 1993 Current records from Turkey: Bolu province: Abant (type locality) ; Bolu province (Tauzin, 2000); Bolu province: Abant; Zonguldak province: from Karadere to Eğerci (Malmusi & Saltini, 2005). Range: Turkey. Zoogeographical remarks: The subspecies is distributed only towards West of Kastamonu province in North Turkey. Up to now, it has been known from a rather local area in Turkey.

Genus Saphanus Serville, 1834 Saphanus piceus (Laicharting, 1784)

Current records from Turkey: There is no published record from Turkey. However, this species collected in Turkey is preserved in the collection of Stanislav Kadlec (Czechia) according to Danilevsky (2004). Range: Europe (Winkler, 1924-1932); Central and North of South Europe (Svacha & Danilevsky, 1986); Europe (Bense, 1985); Europe (Althoff & Danilevsky, 1997); Europe: from French Alps to Bulgaria and Greece (Sama, 2002); Europe


and Balkan Peninsula (Danilevsky, 2005). Zoogeographical remarks: The species is distributed only in Europe and probably European Turkey. It has the S-European chorotype.

Genus Oxypleurus Mulsant, 1839 Oxypleurus nodieri Mulsant, 1839

Current records from Turkey: Antalya province: Bey Mts. (Demelt, 1963); Southern Turkey (Sama, 2002). Range: Mediterranean Region (Winkler, 1924-1932); Crimea, Europe, North Africa (Danilevsky & Miroshnikov, 1985); Mediterranean area (inc. Canary Islands), as far eastwards as the Black Sea, South Africa (Svacha & Danilevsky, 1986); Europe, North Africa (Bense, 1985); Europe (Althoff & Danilevsky, 1997); Southern Europe, Southern Turkey, Canary Islands, Madeira, North Africa, imported to South Africa (Sama, 2002); Crimea Peninsula, Caucasus, North Africa, South Europe, Balkan Peninsula (Danilevsky, 2005). Zoogeographical remarks: The species is distributed only in South part of Turkey. It has the Mediterranean chorotype.

Tribe Anisarthronini Genus Anisarthron Redtenbacher, 1845 Anisarthron barbipes (Schrank, 1781)

Current records from Turkey: The species has been reported only by Lodos (1998) from Turkey without exact locality. Range: Europe (Winkler, 1924-1932); Central Europe, up to Northern Italy and North of the Balkan Peninsula (Svacha & Danilevsky, 1986); Europe (Bense, 1985); Europe: from Italy to Ukraine (Althoff & Danilevsky, 1997); Western, Central and Eastern Europe (Sama, 2002); Europe and Balkan Peninsula (Danilevsky, 2005). Zoogeographical remarks: Unfortunately any record with exact locality in Turkey has not been given according to the literature yet. For this reason the real status in Turkey of this taxon needs to be clarified. It has the S-European chorotype.

Genus Alocerus Mulsant, 1862 Alocerus moesiacus (Frivaldsky, 1838)

Current records from Turkey: Turkey (Svacha & Danilevsky, 1986); Isparta province: Eğirdir, Kovada (Adlbauer, 1992); Turkey (Lodos, 1998); Antalya province: Kumluca (Tozlu et al., 2002); Kırklareli province: İğneada, Mert lake (Özdikmen & Çağlar, 2004). Range: Mediterranean Region (Winkler, 1924-1932); Mediterranean Region (Svacha & Danilevsky, 1986); South Europe, North Africa (Bense, 1985); South Europe: from Portugal to Bulgaria and Greece (Althoff & Danilevsky, 1997). Zoogeographical remarks: The species is distributed in European Turkey and South Anatolia in Turkey. It has the Mediterranean chorotype.

Tribe Asemini Genus Nothorhina Redtenbacher, 1845 Nothorhina muricata (Dalman, 1817)

Current records from Turkey: The species has been reported as N. punctata by Lobanov et al., 1981 and Svacha & Danilevsky, 1986 from Turkey without exact locality. Range: Central and North Europe (Winkler, 1924-1932); Japan (Kusama & Takakuwa, 1984); Europe, South-West Siberia, Asia Minor, Japan (Svacha & Danilevsky, 1986); Europe, European Russia (Bense, 1985); Europe, European Russia, European Kazakhstan (Althoff & Danilevsky, 1997); Northern Palaearctic Region from Western Europe to Japan (Sama, 2002); Europe, Balkan Peninsula, Kazakhstan, West Siberia, Near East, Japan (Danilevsky, 2005). Zoogeographical remarks: Unfortunately any record with exact locality in Turkey has not been given according to the literatures yet. For this reason the real status in Turkey of this taxon needs to be clarified. It has the Sibero-European chorotype.

Genus Asemum Eschscholtz, 1830 Asemum striatum (Linnaeus, 1758)

Current records from Turkey: Turkey (Acatay, 1948, 1961 and 1968); Sinop province: Hassan, 45 km SE Sinop (Villiers, 1967); Gümüşhane province: Torul Mts. (Gfeller, 1972); Gümüşhane province (Öymen, 1987); Tokat province: Topçam Mt. and Yıldız Mt., Almus, Teknecik (Adlbauer, 1992); Turkey (Lodos, 1998); Kars province: Sarıkamış (Tozlu, 2001); Kars province: Sarıkamış (Tozlu et al., 2002); Northern Turkey (Sama, 2002). Range: Palaearctic Region (Winkler, 1924-1932); European Russia, Caucasus, Siberia, Europe, Mongolia, North-East China, Japan, Korea, Northern America (Danilevsky & Miroshnikov,


1985); Palaearctic Region, North America (Svacha & Danilevsky, 1986); Europe, European Russia (Bense, 1985); Europe, European Russia, European Kazakhstan (Althoff & Danilevsky, 1997); Europe, Northern Turkey, Caucasus, Siberia, Manchuria, Korea, Sakhalin, Japan, North America (Sama, 2002); Europe, Balkan Peninsula, Caucasus, Republics of Central Asia, Kazakhstan, West and East Siberia, Far East Russia, Sakhalin, Mongolia, Korea, Japan (Danilevsky, 2005). Zoogeographical remarks: Apparently the species is distributed only in North parts of Turkey. It has the Holarctic chorotype.

Asemum tenuicorne Kraatz, 1879 Current records from Turkey: Ankara province: Kızılcahamam (Demelt, 1967); Turkey (Lobanov et al., 1981); Turkey (Danilevsky & Miroshnikov, 1985); Turkey (Svacha & Danilevsky, 1986); Turkey (Lodos, 1998); Asia Minor (Central and South-Eastern Turkey) (Sama, 2002). Range: Crimea, Caucasus, Syria (Winkler, 1924-1932); Crimea, Caucasus, Transcaucasia, Near East, Turkey (Danilevsky & Miroshnikov, 1985); Crimea, Caucasus, Transcaucasia, Turkey, Syria, Greece, ? Sweden (Svacha & Danilevsky, 1986); Italy, Greece, Romania, Sweden, Crimea (Althoff & Danilevsky, 1997); Europe (incl. Sweden, Italy, Greece, Rumania, Crimea, Central and South-Eastern Turkey, West Caucasus, Northern Spain) (Sama, 2002); Crimea, Caucasus, Europe, Near East (Danilevsky, 2005). Zoogeographical remarks: Apparently the species is distributed in Central and South parts of Turkey. It has probably the Turano-European chorotype.

Genus Arhopalus Serville, 1834 Arhopalus ferus (Mulsant, 1839)

Current records from Turkey: Hatay province: Akbez (Fairmaire, 1884); Bolu province: Abant as C. polonicus (Alkan, 1946); İzmir province: Central; Aydın province: Kuşadası (Demelt, 1963); Antalya province: Gebiz, Bozburan Mt. as Arhopalus tristis (Villiers, 1967); İzmir province: Bornova; Tokat province: Turhal (Gül-Zümreoğlu, 1972 and 1975); Turkey as Arhopalus tristis Lobanov et al., 1981); Eskişehir province: Bokuyak (=?Bozüyük) as Arhopalus tristis (Sama, 1982); Çanakkale province: Gelibolu; Ankara province: Botanic Garden as Arhopalus tristis (Öymen, 1987); Çanakkale province: İntepe as Arhopalus tristis (Adlbauer, 1988); Tokat province: Turhal; İzmir province; Marmara Region, Aegean Region (Lodos, 1998); Adana province: Balcalı and Kozan; Antalya province: Central, Serik, Alanya, Topçam and Akseki; Bilecik province: Central; Kütahya province: Tavşanlı (Tozlu et al., 2002); Aydın province : Kuşadası ; Kırklareli province : İslambeyli ; Muğla province : Marmaris (Malmusi & Saltini, 2005) ; Turkey (Özdikmen & Şahin, 2006). Range: Europe, Siberia (Winkler, 1924-1932); European Russia, Caucasus, Transcaucasia, Siberia, Europe, Near East, Northern China, North Africa (Danilevsky & Miroshnikov, 1985); Most of Palaearctic Region (Svacha & Danilevsky, 1986); Europe, European Russia, North Africa (Bense, 1995); Europe, European Russia, European Kazakhstan (Althoff & Danilevsky, 1997); North Africa, Europe, Caucasus, Transcaucasia, Syria, Israel, Siberia, China (Sama, 2002); Europe, Balkan Peninsula, Crimea, Caucasus, Kazakhstan, West Siberia, Far East Russia, North Africa, Near East, China (Danilevsky, 2005). Zoogeographical remarks: Apparently the species is rather widely distributed in Turkey (especially West half of Turkey). It has the Sibero-European and the Turano-Europeo-Mediterranean chorotypes.

Arhopalus rusticus (Linnaeus, 1758) Current records from Turkey: Between Ordu province and Bolu Mts. (Alkan, 1946); Sinop province: Ayancık (Schimitschek, 1944); Turkey (Acatay, 1948, 1961 and 1968); Tokat province: Arguslu (Villiers, 1967); Bolu province: Bolu Mts., Mengen (Demelt, 1967); Karabük province: Büyükdüz Research Forest (Besçeli, 1969); Denizli province: Buldan (Gül-Zümreoğlu, 1975); Antalya province: Manavgat (Karavca place), Korkuteli (Yazır forests), Serik (Belek forest) (Tosun, 1975); Bolu province, Sinop province: Ayancık, Gümüşhane province, Trabzon province: Akçaabat, Antalya province: Manavgat, Korkuteli, Serik, Denizli province (Erdem & Çanakçıoğlu, 1977); Gümüşhane province: Torul, Örümcek Serisi forests, Trabzon province: Akçaabat, around Düzköy (Sekendiz, 1981); Bolu province, Sinop province: Ayancık, Gümüşhane province, Trabzon province: Akçaabat, Antalya province: Manavgat, Korkuteli and Serik, Denizli province (Çanakçıoğlu, 1983); Northern Turkey (Danilevsky & Miroshnikov, 1985); Artvin province: Atilla Forest, Muğla province: Central, Balıkesir province: Dursunbey, Alaçam, İstanbul province: Büyükada and Belgrad Forest (Öymen, 1987); Turkey (Önder et al., 1987); Kütahya province, Kastamonu province: Yaralıgöz pass, Sinop province: Boyabat, Dıranaz pass, Rize province: İkizdere (Adlbauer,


1992); Artvin province: Şavşat (Yayla, Kocabey place) and (Veliköy, Karagöl Forests), Gümüşhane province: Karanlıkdere Forests (Yüksel, 1996); Gümüşhane province, Artvin province, Kars province, East Anatolian Region, Aegean Region, Western Black Sea Region, Mediterranean Region (Lodos, 1998); Artvin province: Ardanuç, Ovacık (Alkan, 2000); Kars province: Sarıkamış (Tozlu, 2001); Ankara province: Çamlıdere, Bayburt province: Central, Gümüşhane province: Central and Kürtün, Kars province: Sarıkamış (Tozlu et al., 2002); Turkey (Sama, 2002); Ordu province: Perşembe, Çaytepe Özdikmen, 2006); Karabük province: Büyükdüz Research Forest (Özdikmen & Şahin, 2006). Range: Europe (Winkler, 1924-1932); European Russia, Caucasus, Transcaucasia, Siberia, Europe, North Iran, North Turkey, Mongolia, Korea, Northern China, Japan, North Africa, North Amerika (Danilevsky & Miroshnikov, 1985); Most of Palaearctic Region, North America (Svacha & Danilevsky, 1986); Europe, European Russia, North Africa (Bense, 1995); Europe, European Russia, European Kazakhstan (Althoff & Danilevsky, 1997); North Africa, Europe, Turkey, Caucasus, Iran, Middle Asia, Siberia, Japan, North America, European Russia, European Kazakhstan (Sama, 2002); Europe, Balkan Peninsula, Crimea, Caucasus, Kazakhstan, West and East Siberia, Far East Russia, Sakhalin, Near East, Iran, Mongolia, Korea, Japan (Danilevsky, 2005). Zoogeographical remarks: Apparently the species is widely distributed in Turkey. It has the Holarctic chorotype.

Arhopalus syriacus (Reitter, 1895) Current records from Turkey: İzmir province: Kemalpaşa as Criocephalus syriacus (Demelt & Alkan, 1962); İzmir province: Kemalpaşa, Antalya province: Bey Mt. as Criocephalus syriacus (Demelt, 1963); Antalya province: Kepez forests (Tosun, 1975); Artvin province: Ardanuç Forests as Criocephalus syriacus (Sekendiz, 1981); Muğla province: Yeşilyurt (Adlbauer, 1992); Gaziantep province, Şanlıurfa province, İzmir province: Kemalpaşa as Criocephalus syriacus (Lodos, 1998); Adana province: Balcalı, Bağtepe and Kozan, Antalya province: Central (Tozlu et al., 2002); Nevşehir province: Cappadocia (Malmusi & Saltini, 2005); İçel province: Mut, Alahandüzü (Özdikmen & Şahin, 2006). Range: Syria (Winkler, 1924-1932); Mediterranean area, Canary Islands (Svacha & Danilevsky, 1986); South Europe (Bense, 1995); South Europe (Althoff & Danilevsky, 1997); Mediterranean Region (Sama, 2002). Zoogeographical remarks: Apparently it distributed in South parts of Turkey. For this reason, the Artvin record in Sekendiz (57) is very doubtful as a possible wrong identification. The status of this record must be revised. It has the Mediterranean chorotype.

Genus Tetropium Kirby, 1837 Tetropium castaneum (Linnaeus, 1758)

Current records from Turkey: Turkey (Acatay, 1948, 1961 and 1968); Düzce province, Zonguldak province: Devrek, Western Black Sea Region (Erdem & Çanakçıoğlu, 1977, 1983); Düzce province: Çiçekli forest, Zonguldak province: Devrek, Beldibi, Davulgu forest, Zonguldak province: Devrek, Karadere as Tetropium luridum (Defne, 1954); North Anatolia as Tetropium luridum (Erdem, 1968); Bolu province: Seben (Öymen, 1987); Turkey (Önder et al., 1987); Artvin province: Atila Forests (Yüksel, 1996); Zonguldak province, Bolu province, Western Black Sea Region (Lodos, 1998); Artvin province: Central (Alkan, 2000). Range: Central and North Europe, Siberia (Winkler, 1924-1932); European Russia, Northern Caucasus, Siberia, Europe, Mongolia, Northern China, Korea, Northern Japan (Danilevsky & Miroshnikov, 1985); Most of Palaearctic Region, from Europe to Japan (Svacha & Danilevsky, 1986); Europe, European Russia (Bense, 1995); Europe, European Russia, European Kazakhstan (Althoff & Danilevsky, 1997); Northern Palearctic region from Europe to Japan (Sama, 2002); Europe, Balkan Peninsula, Crimea, Caucasus, Kazakhstan, West and East Siberia, Far East Russia, Sakhalin, Mongolia, China, Korea, Japan (Danilevsky, 2005). Zoogeographical remarks: Apparently it is distributed only in North parts of Turkey. It has the Sibero-European chorotype.

Tetropium fuscum (Fabricius, 1787) Current records from Turkey: Turkey (Acatay, 1948, 1961 and 1968); North Anatolia (Erdem, 1968); Artvin province: Şavşat (Veliköy, Karagöl Forests), (Karagöl, Meşeli, Mukali district), Ardanuç (Tepedüzü, Şahinkaya) (Yüksel, 1996); Turkey (Lodos, 1998); Artvin province: Şavşat, Karagöl (Alkan, 2000). Range: Central and North Europe, Siberia (Winkler, 1924-1932); Japan (Kusama & Takakuwa, 1984); European Russia, Georgia, West Siberia, Europe (Danilevsky & Miroshnikov, 1985); Europe, West Siberia (Svacha &


Danilevsky, 1986); Europe, European Russia (Bense, 1995); Europe, European Russia, European Kazakhstan (Althoff & Danilevsky, 1997); Europe, Western Siberia, European Kazakhstan (Sama, 2002); Europe, Balkan Peninsula, Caucasus, Kazakhstan, West Siberia, Japan (Danilevsky, 2005). Zoogeographical remarks: Apparently it is distributed only in North-East Turkey. It has the Sibero-European chorotype.

Tetropium gabrieli Weise, 1905 Current records from Turkey: The species has only been reported by Acatay, 1948, 1961 and 1968 from Turkey without exact locality. Range: Central Europe (Winkler, 1924-1932); Central Europe and some adjacent regions, reaching to Danmark, Germany, France, Great Brtain (Svacha & Danilevsky, 1986); Central and North Europe (Bense, 1995); Europe, ? North parts of European Russia (Althoff & Danilevsky, 1997); Europe from British Isles to Russia (Sama, 2002); Europe (Danilevsky, 2005). Zoogeographical remarks: Any record with exact locality in Turkey has not been given according to the literature yet. Unfortunately the records in Acatay (1948, 1961 and 1968) seem to be very doubtful. Because the species must not occur in Turkey according to the known distributional patterns in the world. Therefore Acatay’s records are not valid. It has the Centraleuropean chorotype.

Tribe Spondylidini Genus Spondylis Fabricius, 1775

Spondylis buprestoides (Linnaeus, 1758) Current records from Turkey: Sinop province: Ayancık (Schimitschek, 1944); Kars province: Sarıkamış (Erdem, 1947); Turkey (Acatay, 1948, 1961, 1963 and 1968); Bursa province: Keles, Kocayayla place (Çanakçıoğlu, 1956); Trabzon province: Zigana Mts. (Villiers, 1967); Ankara province: Kızılcahamam (Demelt, 1967); Karabük province: Büyükdüz Research Forest (Besçeli, 1969); Turkey (Lobanov et al., 1981); Turkey (Danilevsky & Miroshnikov, 1985); Trabzon province: Maçka (Öymen, 1987); Artvin province: Ardanuç (Yüksel, 1996); Turkey (Lodos, 1998); Artvin province: Ardanuç, Tosunlu (Alkan, 2000); Kars province: Sarıkamış (Tozlu, 2001); Kars province: Sarıkamış (Tozlu et al., 2002); Northern and South-Western Turkey (Sama, 2002); İstanbul province: Şile (Malmusi & Saltini, 2005); Ankara province: Kızılcahamam, Çamkoru (Özdikmen & Şahin, 2006). Range: Palaearctic Region (Winkler, 1924-1932); European Russia, Caucasus, Transcaucasia, Siberia, Europe, Turkey, Northern Iran, Mongolia, Northern China, Korea, Japan (Danilevsky & Miroshnikov, 1985); Most of Palaearctic Region, from Europe to Japan (Svacha & Danilevsky, 1986); Europe, European Russia (Bense, 1995); Europe, European Russia, European Kazakhstan (Althoff & Danilevsky, 1997); Europe, Northern and South-Western Turkey, Siberia, Sakhalin, Northern Mongolia, Northern China, Korea, Japan, ? North Africa (Sama, 2002); Europe, Balkan Peninsula, Crimea, Caucasus, Kazakhstan, West and East Siberia, Far East Russia, Sakhalin, Mongolia, China, Korea, Japan (Danilevsky, 2005). Zoogeographical remarks: Apparently it is distributed only in North parts of Turkey. It has the Sibero-European chorotype.

DISCUSSION

The Turkish Spondylidinae subfamily is represented by 4 tribes as Saphanini, Anisarthronini, Asemini and Spondylidini. The tribe Saphanini includes 3 genera as Drymochares, Saphanus and Oxypleurus. The tribe Anisarthronini includes 2 genera as Anisarthron and Alocerus. The tribe Asemini includes 4 genera as Nothorhina, Asemum, Arhopalus and Tetropium. The tribe Spondylidini includes only one genus as Spondylis normally. In the tribe Saphanini, the genus Drymochares is represented by 2 subspecies as D. s. cavazzutii and D. s. ivani of the species Drymochares starcki. The other two genera, Saphanus and Oxypleurus are represented


by one each species as Saphanus piceus (Laicharting, 1784) and Oxypleurus nodieri Mulsant, 1839 respectively. In the tribe Anisarthronini, both genera Anisarthron and Alocerus are represented by one species each as Anisarthron barbipes (Schrank, 1781) and Alocerus moesiacus (Frivaldsky, 1838) respectively. In the tribe Spondylidini, the genus Spondylis is represented by the species Spondylis buprestoides (Linnaeus, 1758). In the tribe Asemini, the genus Nothorhina is represented by the species Nothorhina muricata (Dalman, 1817). The genus Asemum is represented by 2 species as Asemum striatum (Linnaeus, 1758) and Asemum tenuicorne Kraatz, 1879. The genus Arhopalus is represented by 3 species as Arhopalus ferus (Mulsant, 1839), Arhopalus rusticus (Linnaeus, 1758) and Arhopalus syriacus (Reitter, 1895). According to literatures, the genus Tetropium has been represented by 3 species as Tetropium castaneum (Linnaeus, 1758), Tetropium fuscum (Fabricius, 1787) and Tetropium gabrieli Weise, 1905. But, as seen in the text, Tetropium gabrieli has only been known by Acatay’s records from Turkey. However, Acatay’s records are wrong. We think the species must not occur in Turkey according to its known distribution patterns in the world. Consequently, the Turkish Spondylidinae subfamily includes 4 tribes, 10 genera, 14 species and 2 subspecies until now according to the present paper. Taking into consideration the status of the species Tetropium gabrieli, the known real taxa number of Turkish Spondylidinae is 15 as species and subspecies. Moreover, Sama (pers. com.) mentioned that Anisarthron barbipes, Saphanus piceus and all species of the genus Tetropium are very likely absent in Turkey. Under these circumtances, Turkish Spondylidinae includes 4 tribes, 7 genera, 9 species and 2 subspecies. Zoogeographically, Asemum striatum (Linnaeus, 1758) and Arhopalus rusticus (Linnaeus, 1758) have the Holarctic chorotype. Oxypleurus nodieri Mulsant, 1839, Alocerus moesiacus (Frivaldsky, 1838) and Arhopalus syriacus (Reitter, 1895) have the Mediterranean chorotype. Saphanus piceus (Laicharting, 1784) and Anisarthron barbipes (Schrank, 1781) have the S-European chorotype. Nothorhina muricata (Dalman, 1817), Tetropium castaneum (Linnaeus, 1758), Tetropium fuscum (Fabricius, 1787) and Spondylis buprestoides (Linnaeus, 1758) have the Sibero-European chorotype. Arhopalus ferus (Mulsant, 1839) has the Sibero-European and the Turano-Europeo-Mediterranean chorotypes. Asemum tenuicorne Kraatz, 1879 has probably the Turano-European chorotype. Tetropium gabrieli Weise, 1905 has the Centraleuropean chorotype. The subspecies Drymochares starcki cavazzutii Sama & Rapuzzi, 1993 has been known from Turkey and Caucasus. The other subspecies Drymochares starcki ivani Sama & Rapuzzi, 1993 are known only from Turkey yet.


ACKNOWLEDGEMENTS Thank to Dr. Gianfranco Sama (Italy) for his review and comments.

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Defne, M. Ö. 1954. Batı Karadeniz Bölgesindeki Göknarların Zararlı Böcekleri ve Mücadele Metodları. T. C. Tarım Bakanlığı, Orman Genel Müdürlüğü Yay. No. 105, 60-72. Demelt, C. V. & Alkan, B. 1962. Short information of Cerambycidae Fauna of Turkey. Bitki Koruma Bülteni, 2 (10): 49-56. Demelt, C. V. 1963. Beitrag zur Kenntnis der Cerambycidenfauna Kleinasiens und 13. Beitrag zur Biologie palaearkt. Cerambyciden, sowie Beschreibung einer neuen Oberea-Art. Entomologische Blatter, 59 (3) : 132-151. Demelt, C. V. 1967. Nachtrag zur Kenntnis der Cerambyciden-Fauna Kleinasiens. Entomologische Blatter, 63 (2): 106-109. Erdem, R. 1947. Sarıkamış Ormanlarında Entomolojik Gözlemler, T.C. Tarım Bakanlığı, Orman Genel Müdürlüğü Yayınları, Özel Sayı: 52, Ankara. Erdem, R. 1968. Useful and Harmful Insects of Forest, İstanbul Univ. Publication of Forest Faculty, 43-58, İstanbul, Turkey (Turkish with English summary). Erdem, R. & Çanakçıoğlu, H. 1977. Pests of Wood in Turkey, İstanbul Univ. Publication of Forest Faculty, 113-134, İstanbul, Turkey (Turkish with English summary). Fairmaire, M. L. 1884. Liste des Coleopteres recueillis par M. l’abbe David a Akbes (Asie-Mineure) et Descriptions des Especes Nouvelles. Annales de la Societe entomologique de France, Octobre (1884): 165-180. Gfeller, W. 1972. Cerambycidae (Coleoptera) der Türkei-Persienexpedition 1970 der Herren Dr. H. c. W. Wittmer und U. v. Botmer. Mitteilungen der Entomologischen Geselschaft Basel, 22 (1): 1-8. Gül-Zümreoğlu, S. 1972. Catalogue of Insect and common pests (1928-1969). T. C. Publications of Agriculture Ministry, Bornova, İzmir, 119 pp. Gül-Zümreoğlu, S. 1975. Investigations on taxonomy, host plants and distribution of the Longhorned Beetles (Cerambycidae-Coleoptera) in Aegean Region. T. C. Ministry of Food, Agriculture and Stockbreeding, No : 28, , İstiklal Press, İzmir, 208 pp. Kusama, K. & Takakuwa, M. 1984. Parandrinae. Prioninae. Spondylinae. Aseminae. Lepturinae (part.). Cerambycinae. Lamiinae (part.). The Longicorn-beetles of Japan in Color. Kodansha, Tokyo: 131-172, 201-493, 511-549. Lobanov, A. L., Danilevsky, M. L. & Murzin, S. V. 1981. [Systematic list of Longicorn beetles (Coleoptera. Cerambycidae) of the USSR. 1]. Rev. d’Entom. 60 (4): 784-803 [in Russian]. Lodos, N. 1998. Entomology of Turkey VI (General, Aplied and Faunistic). Ege Ü. Ziraat Fak. Yayınları No: 529, E. Ü. Faculty of Agriculture Press, İzmir, 300 pp. Malmusi, M. & Saltini, L. 2005. Cerambycidae raccolti dai componenti del Gruppo Modenese Scienze Naturali durante escursioni in Turchia tra il 1987-2003 (Contributo alla Fauna dei Cerambycidae di Turchia). Quaderno di studi e notizie di storia naturale della Romagna, n. 21, 28 pp. (unpublished). Önder, F., Karsavuran, Y., Tezcan, S. & Önder, P. 1987. Scientific and Turkish names of some useful and harmful species of Agricultural, Forestic and Domestic Animals in Turkey, T.C. Ministry of Agriculture, Ankara, Turkey (Turkish with English summary). Öymen, T. 1987. The Forest Cerambycidae of Turkey. İ. Ü. Forest Faculty, İstanbul, 146 pp.


Özdikmen, H. 2006. Contribution to the knowledge of Turkish longicorn beetles fauna (Coleoptera: Cerambycidae). Munis Entomology & Zoology, 1 (1): 71-90. Özdikmen, H. & Çağlar, Ü. 2004. Contribution to the Knowledge of Longhorned Beetles (Coleoptera, Cerambycidae) from Turkey, Subfamilies Prioninaei Lepturinae, Spondylidinae and Cerambycinae. Journal of the Entomological Research Society, 6 (1): 39-69. Özdikmen, H. & Şahin, Ö. 2006. Longhorned beetles collection of the entomology museum of Central Anatolia forestry research directorship, Ankara, Turkey (Coleoptera, Cerambycidae). G. U. Journal of Science 19 (1): 1-8. Sama, G. 1982. Contributo allo studio dei coleotteri Cerambycidae di Grecia e Asia Minore. Fragmenta Entomologica, Roma, 16 (2): 205-227. Sama, G. 2002. Atlas of the Cerambycidae of Europe and the Mediterranean Area, Volume I, Kabourek, Zlin, 173 pp. Sama, G. & Rapuzzi, P. 1993. Revisione dei generi Saphanus Serville, 1834 e Drymochares Mulsant, 1847. Lambillionea, 93: 278–294. Schmitschek, E. 1944. Forstinsekten der Türkei und Ihre Umwelt Grundlagen der türkischen Forstentomologie, Volk und Reich Verlag Prag, 125-141 pp. Sekendiz, O. A. 1981. Researches on important damaging animal species of Eastern Black Sea Region. K. T. Ü. Publication of Forest Faculty, No : 12, Trabzon, 114 pp. Svacha, P. & Danilevsky, M. L. 1986. Cerambycoid larvae of Europe and Soviet Union (Coleoptera, Cerambycoidea). Part I. Acta Universitatis Carolinae – Biologica, 30 : 1-176. Taglianti, A. V., Audisio, P. A., Biondi, M., Bologna, M. A., Carpaneto, G. M., De Biase, A., Fattorini, S., Piattella, E., Sindaco, R., Venchi, A. & Zapparoli, M. 1999. A proposal for a chorotype classification of the Near East fauna, in the framework of the Western Palaearctic Region. Biogeographia 20: 31-59. Tauzin, P. 2000. Complement a l’inventaire des Coleopteres Cerambycidae de Turquie. L’Entomologiste, 56 (4): 151-153. Tosun, İ. 1975. The damaging insect species at conifer forests in Mediterranean Region and researches on their predators and parasites. İ. Ü. Orman Fakültesi Dergisi, 26 (2): 218-254. Tozlu, G. 2001. Sarıkamış (Kars) Ormanlarında Sarıçam (Pinus sylvestris L.)’da zarar yapan Elateridae, Buprestidae, Cerambycidae, Curculionidae (Coleoptera) ve Diprionidae (Hymenoptera) familyalarına bağlı türler üzerinde çalışmalar. Türkiye Entomoloji Dergisi 25(2): 133-146. Tozlu, G., Rejzek, M. & Özbek, H. 2002. A contribution to the knowledge of Cerambycidae (Coleoptera) fauna of Turkey. Part I: Subfamilies Prioninae to Cerambycinae. Biocosme Mèsogèen, Nice, 19 (1-2): 55-94. Villiers, A. 1967. Coléoptéres Cérambycides de Turquie (1. Partie) - L’ Entomologiste, 23 (1): 18-22. Winkler, A. 1924-1932. Catalogus Coleopterorum regionis palaearcticae. Verlag von Albert Winkler, 1135-1226. Yüksel, B. 1996. The damaging insect species at oriental spruce forests and their predators and parasite species –I (Pest Insects), K.T.Ü. Institute of Science, Engineering of Forest, Trabzon, 222 pp. (Turkish with English summary).



THE OCCURRENCE OF CHORISTONEURA MURINANA IN ABIES CILICICA FORESTS IN ASIA MINOR

Oğuzhan Sarıkaya* and Mustafa Avcı*

* Süleyman Demirel University, Forestry Faculty, Forest Engineering Department, 32260 Isparta, TURKEY, e-mail: [email protected] [Sarıkaya, O. & Avcı, M. 2006. The occurrence of Choristoneura murinana in Abies cilicica forests in Asia Minor. Munis Entomology & Zoology 1 (2): 289-296] ABSTRACT: Although widespread in European fir forests, Choristoneura murinana (Hübner) was not known in Asia Minor until 2000. Owing to the serious damage caused by this budworm in the Lakes District to stands of Abies cilicica Carr., this study was launched on its distribution, morphology, biology and injury. Forewings of the C. murinana adults are yellowish-beige colored. The mean wingspan between forewings was 22.8±1.7 mm (mean ± standard deviation) in female and 18.5±1.6 mm (mean±SD) in male adults. The main flying period of C. murinana lasts from the mid of June to the end of July. The moth is univoltine and overwinters in larval stage. C. murinana has a total of 6 larval instars, with greyish-light green mature larvae. Pupation takes place among the last needles they feed on. C. murinana causes serious damage to fresh shoots, buds and needles, especially when the larvae molt to third stage in the second half of April. Spring feeding of the larvae causes more damage than summer and autumn feeding. Key words: Choristoneura murinana, fir forest, the Lakes District, budworm, Turkey

The European fir budworm (=EFB) Choristoneura murinana (Hübner) (Lep.: Tortricidae) feeds on the needles, buds and shoots of fir trees (Du Merle & Géraud, 1988). It was recorded for the first time in Asia Minor in 2000, in Abies cilicica Carr. stands in Yenişarbademli, county of Isparta, Turkey (Sarıkaya & Avcı, 2002). EFB is a pest of the fir, and after heavy defoliation the tree weakens and is attacked by bark beetles (especially Pityokteines curvidens Germ. and Cryphalus piceae Ratz.) which may cause the death of the tree.

EFB is widespread in the northern hemisphere especially in Central and Eastern Europe. Damage caused by C. murinana was first recorded in 1805 in Thüringerwald forests in Germany (Bucher, 1953). Outbreaks of the insect in different regions of Germany were recorded by Schimitschek (1936) and Franz (1940). Outbreaks of EFB were reported from Central and Eastern Europe (Patocka, 1960; Birova, 1966; Du Merle et al., 1992; Géraud, 1988; Du Merle & Brunet, 1991; Du Merle et al., 1990; Markalas & Bogenschütz, 1995). Beside these studies, the moth was recorded in central Russia and Siberia (Ivliev et al., 1970; Galkin, 1980).

Du Merle et al. (1992) and Bogenschütz (1997) stated that EFB caused serious damage throughout A. alba regions in Europe. Their studies showed that C. murinana, which was first known as monophagus on A. alba, caused damage on other species of trees in various regions of Europe and Siberia also. The occurrence of the moth in Yenişarbademli fir forests suggests that Asia Minor is included in the range of EFB and A. cilicica is a new host plant (Sarıkaya & Avcı, 2002). The study on the


biology and ecology of the moth was launched owing to the severe damages inflicted to the local fir stands.

MATERIALS & METHODS

Field studies were conducted between 2001 and 2003 in May-August in a mixed forest in Yenişarbademli (Fig. 1). The stand includes mainly A. cilicica, a few Pinus nigra Arnold and Juniperus excelsa Bieb. The morphology and biology of EFB was studied on laboratory reared individuals. Larvae were removed from the damaged fir trees at different altitudes (between 1060 and 1400 m) of the study area. A survey was conducted by Delta pheromone traps in neighbor fir stands. The larvae were put into plastic jars (25x25x30 cm) and fed with fresh fir shoots. In addition to the laboratory reared pupae, additional pupae were taken from the forest and put into jars. The development was monitored until adult emergence. Measurements of the wingspan of 30 ♂ and 30 ♀ were taken. The sex ratio and the life span of moths were assessed. The mating took place in reproduction jars. The eggs were collected from the shoots and maintained in the laboratory at 21-24 °C.

RESULTS Distribution

After the finding of the budworm in A. cilicica forests of Yenişarbademli, a county of Isparta, in 2000, investigations were conducted also in A. cilicica forests of Lakes District, southwest Turkey. However, EFB was found only in Yenişarbademli Taurus fir (A. cilicia) forests. Biometrics

The wings of adult are yellowish-beige colored with light stripes and brown patterns. Forewings bear had light stripes and brown patterns. Hind wings are brownish grey (Fig.2). Male are smaller than female moths, and the mean wingspan was 22.8±1.7 mm (18-24 mm) in female and 18.5±1.6 mm (16-21 mm) in male moths.

The eggs are laid in two rows and attached on the needles. The eggs are oval, the upper surface is convex and lower surface is disc-shaped, with a width of 0.8-1 mm (0.9±0.06 mm). Mean number of eggs in an egg mass was 26±3.4 (12-42) (n=76).

The newly hatched larvae are greenish-yellow, with reddish-brown head capsules. The first instar larvae are 2 mm long on average. The mature larvae are grayish-light green with blackish-brown head capsules. Abdomen segments are black and clearly visible. The thoracic legs are black. The mean length of mature larvae is 1.6±0.25 cm (1.2-2.1 cm) (Fig. 3). The mean width of the head capsules of last (sixth) instar larvae is 1.60±0.32 mm (1.43-1.80 mm, n=32).

The prepupa is greyish green like last instar larvae. 1-2 days after pupating, the pupae turn to brown (Fig. 4). The pupa mean length is 9.8±1.3 mm (8-12 mm, n=75). Its mean width is 3.6±0.22 mm (3-4 mm,


n=75). Male pupae were smaller (0.33±0.0035 g) and lighter than female pupae (0.48±0.012 g). Feeding and damage symptoms

The larvae feed on young shoots, buds and needles. They seem to prefer the current year needles. The heavily defoliation is caused by the advance instar larvae (3-6) soon after hibernation. They move between shoots by spinning webs, surrounding two or more neighbouring needles used for shelter. Larvae caused heavy damage on the shoots (Fig. 5). Basic Biology C. murinana is a univoltine species. The main flight period lasts from the mid of June to the end of July. Lifespan of adults was 6 days in laboratory. The ratio of male/female was determined as 1/1.

Fresh egg masses are green; they turn to light green in 2nd day and to yellow-green 4-5 days after deposition. Incubation lasted 7-9 days in the laboratory. Hatching began on June 17 in 2002 and on June 15 in 2003. In the forest hatching occurred one week later.

The larvae terminated hibernation in the second half of April. It was observed that larvae were carried by the wind between shoots and trees. The 6th instar larvae were observed at the end of May.

Pupation takes place inside webs surrounded by half consumed needles. In laboratory (21-24 °C) pupation occurred on June 3 and adult emergence occurred on June 12. Hence, pupa stage lasted about 9 days. In the forest the pupation extended for about 12-14 days.

DISCUSSION

The first record of C. murinana in Turkey was in 2000 in the Abies cilicica stands in Yenişarbademli district. Similar to other species of Abies, A. cilicica is also a suitable host for EFB. In Europe, Priesner et al. (1980), Du Merle & Cornic (1991), Du Merle et al. (1992) and Bogenschütz (1997) described the biology and the damage caused by the budworm on A. alba. Kailidis & Georgevits (1971) found the budworm on A. cephalonica and A. borisi regis. In addition to these results, other hosts were identified by Du Merle et al. (1992) (A. grandis), Du Merle & Cornic (1989; 1991) (Cedrus atlantica Endl.) and Du Merle et al. (1990) (Picea abies L.).

Du Merle & Cornic (1991) conducted research in three different regions of France (Combe Noire, Mont-Ventoux and Luberon) and reported the wingspan between forewings of mature males and females. In their research in Combe Noire in 1985, they determined the wingspan between forewings of mature males to be between 13.0-21.2 mm and 18 mm on average. In the study in Mont-Ventoux, they observed that this distance in mature males is between 12.0-20.6 mm and is 17.6 mm on average. In the same study in Luberon region in 1988, they reported that the wingspan between forewings of male matures was between 14.2-18.6 mm and 16.6 mm on average whereas in female matures this distance was between 14.8-21.8 mm and 18.8 on average. Du Merle et al. (1992) found


the wingspan between forewings of matures to be 22.6 mm in females and 18.2 mm in males. It can be seen that the results of these investigations and our measures do not have great difference, except the fact that the values that are found in Luberon region are smaller than what we have found.

Du Merle & Brunet (1991) counted the number of C. murinana eggs in A. alba forests in Combre-Noire region of France and they determined the average number of eggs in an egg packet as 17.6. Du Merle et al. (1992) determined the average number of eggs as 23.3 in their research in Massif Central region of France. Our results are similar to the ones that were reported for the Massif Central region, whereas they were different from the results obtained from Combre Noire forests.

Du Merle & Brunet (1991) reported that eggs changed yellow green in color about 5-6 days after laying and larvae hatched in 10-12 days. We determined that hatching occurred 10-12 days after depositing.

Bogenschütz (1991) reported that the lengths of mature larvae were usually between 1.2-1.8 cm and this value may be as high as 2.2 cm. In our study this value was determined to be between 1.2 and 2.1 cm (average 1.6±0.25 cm). It can be seen that values about the length of mature larva do not have any significant difference.

Du Merle et al. (1992) determined the width of pupa to be between 2.5-3.4 mm in their study which was conducted in France. Also in same study they published that the average weight of male pupa was 0.035 gr whereas in female, it was 0.052 gr. Bogenschütz (1991) reported that the average weight of male pupa was 0.040 gr in female it was 0.052 gr. and the length of pupae were between 8-14 mm. When our work is compared with the two works mentioned above it can be seen that our results are similar to those of Du Merle et al. (1992).

Bogenschütz (1991) reported that the flight period of the moth is from the end of June until the end of July. Du Merle & Brunet (1991) determined this period to be from the beginning of July until the beginning of August in natural A. alba forest and afforestation field which is constituted by C. atlantica trees in France. Markalas & Bogenschütz (1995) reported the flying period to continue from mid June until mid July. When our work is compared with the other papers, it can be seen that our results are similar to the results of Markalas & Bogenschütz (1995) in Greece. According to the results of Du Merle & Brunet (1991)’s study, which was conducted in France, the flying period starts later than in Greece and in Turkey.

It was observed that the females deposit their eggs in a double row on the needles in the middle and upper crowns of fir trees. Also Géraud et al. (1987), Du Merle & Brunet (1991) and Bucher (1953) reported that females lay their eggs on the upper shoots of trees.

In our study the larvae started to hatch on June 17 in 2002 and on June 15 in 2003 in laboratory conditions. Bogenschütz (1997) reported that the hatching of larvae began at the end of June. When our results are compared with the results of Bogenschütz (1997), it is obvious that larvae


hatched earlier in Yenişarbademli. Pupa duration was between 12-14 days in our study. Also Bogenschütz (1997) reported this period to continue for about 14 days. In both studies the results concerning pupa duration are similar.

In conclusion, it appears that the outbreak of C. murinana on Abies cilicica forests in Turkey is similar to other outbreaks observed in Europe. Further work is needed in order to better characterize this apparently isolated population of the moth.

ACKNOWLEDGEMENTS

We thank Kevin R. TUCK (Entomology Department, The British

Natural Museum) for the identification of the moth, Dr. Zvi Mendel and Dr. Asko Lehtijarvi (Forestry Faculty, Suleyman Demirel University) for the revision of the manuscript and for the useful suggestions.

LITERATURE CITED

Birova, H. 1966. Studium der Parasitierung der Eier des Tannentriebwicklers (Choristoneura murinana Hbn.) und deren Eignung für Arten der Gattung Trichogramma Westw. Biologica 21 (5): 329–338. Bogenschütz, H. 1991. Eurasian species in forestry. In World Crop Pests, Tortricid Pests, vol 5. Elsevier, Amsterdam, pp. 673–677. Bogenschütz, H. 1997. The fir lepidoptera of Slovakia and studies on fir pest insects in the Black Forest (Southwest Germany). Biologia, 52 (2): 141–145. Bucher, G. E. 1953. Biotic factors of control of the European fir budworm, Choristoneura murinana (Hbn.) (N. Comb), in Europe. Can. J. Agric. Sci., 33:448–469. Du Merle, P., Avolio, S. & Chambon, J. P. 1990. Sulla presenza di Choristoneura

murinana (Hb.) e di due altri lepidottei tortricidi nelle abetine di Calabria e di Lucania, LItalia Forestale e montana. 40: 197-212. Du Merle, P. & Brunet, S. 1991. From green to yellow or yellowish white: egg-colour changes in relation to oviposition rank in the fir budworm Choristoneura murinana (Hb.). (Lep., Tortricidae). J. Appl. Ent., 111: 342-348. Du Merle, P., Brunet, S. & Cornic, J. F. 1992. Polyphagous potentalities of Choristoneura murinana (Hb.) (Lep.: Tortricidae): A “monophagous” folivore extending its host range, J. Appl. Ent., 113: 18-40. Du Merle, P. & Cornic, J. F. 1989. Répartition, niveaux de population et risques de pullulation de la tordeuse du Sapin, Choristoneura murinana (Lepidoptera: Tortricidae), en

France. Résultats d une enquete par piégeage sexuel. Ann. Soc. Ent. Fr., 25: 265–288. Du Merle, P. & Cornic, J. F. 1991. Monitoring the reproductive capacity of Choristoneura murinana (Lepidoptera: Tortricidae) populations by measuring the size of male moths caught in sex pheromone traps. Acta Ecol., 12: 368–383. Du Merle, P. & Géraud, D. 1988. Estimation des niveaux d’infestation passes de la tordeuse du sapin Choristoneura murinana (Hb.) (Lép., Tortricidae) par analyse retrospective des pertes de feuillage et de croissance des arbres. Ann. Sci. For., 45: 357-381.


Franz, J. 1940. Der Tannentriebwickler Cacoecia murinana Hb. Beitrage zur Bionomie und Ökologie. Z. Ang. Ent., 27: 585–620. Galkin, G. 1980. Pikhtovaya listovertka-tolstushka-massovyi vreditel’ pikhtovo-elovykh lesov v Krasnoyarskom krae. Lesnoe Khozyajstovo. 3: 65-67. Géraud, D., Cornic, J. F. & Du Merle, P. 1987. Observations sur la distribution spatiale et sur la mesure des populations pré-imaginales de la tordeuse du sapin, Choristoneura murinana Hb. (Lepidoptera., Tortricidae). J. Appl. Ent., 103: 258-278. Ivliev, L. A., Sinchilina, E. M., Kashcheev, M. A. &. Kononov, D. G. 1970. O vspyshke massovogo razmnozheniya pikhtovoj listovertki (Choristoneura murinana Hb.) v. Primorskom krae. Entomol. Issledov. Na Dal’nem Vostoke (Vladivostok) 1: 13-39. Kailidis, D. S. & Georgevits, R. 1971. Ta entoma tis elatis (biologia, simasia, katapolemisis). Deltion Ereunon, 38, Thessaloniki, pp. 1-82. Markalas, S. & Bogenschütz, H. 1995. Preminary results in the biology of Choristoneura murinana (Lepidoptera, Tortricidae) in Greece with sex pheromones. Proceedings of the 5th Hellenic Entomology Meeting, November 1993, Athens, pp. 213–219. Patocka, J. 1960. Die Tannenschmetterlinge der Slowakei, mit Berücksichtigung der Fauna Mitteleuropas. Slovensk Akad., Bratislava. Priesner, E., Bogenschütz, H. & Arn, H. 1980. A sex attractant for the European Fir Budworm Moth, Choristoneura murinana. Z. Naturforsch., 35: 390-398. Sarıkaya, O. & Avcı, M. 2002. Batı Akdeniz Toros Göknarı (Abies cilicica Carr.) Ormanlarında Ağaç Ölümleri. Orman Mühendisliği Dergisi, Sayı: 9-10: 20-24. Schimitschek, E. 1936. Das Massenauftreten des Tannentriebwicklers Cacoecia murinana Hb. in Niederösterreich 1929-1934. Z. Ang. Ent., 22: 565-602.



Fig. 1. Research area

Fig. 2. The female and male moth of C. murinana

♀ ♂


Fig. 3. The mature larva of C. murinana

Fig. 4. The pupa of C. murinana

Fig. 5. Heavy damage of larvae of C. murinana

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