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BRISTOL BAY DATA REPORT NO. 92
Nushagak Sonar Enumeration Project, 1982
·~·
By
R. Eric ~1inard ResearchBiologfst
Division of Commercial Fisheries
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TABLE OF CONTENTS
· Page(s)
List of Figures ............................................... . i
List of Tab 1 es . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . i i
Introduction .................................................. .
Methods and ~1ateri a 1 s ......................................... .
Sonaor Units ..........................•.................... Ca 1 i brati on .............................................. . Sector Distribution ...................................... . Species Apportionment .................................... . AWL Sampling .........................•.....•.............. Climatological Data ...................................... .
Results ....................................................... .
Sonar Counts ............................................. . Sector Di stri but ion ...................................... . Species Apportionment ................... , .........•....... Daily Escapement Estimates ............................... . AWL Sampling ............................................. . Climatological Data ...................................... .
Discussion .....................•................•..............
Literature Cited ........................••.....................
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LIST 0 F FIGURES
J J Figure 1. Location of Nushagak Sonar enumeration project
Nushagak River, Bristol Bay, Alaska.................... 2
J Figure 2. Nushagak River bottom profile at sonar site, 1982...... 3
Figure 3. Nushagak River sonar count distribution, June 14 -
J Figure 4.
August 18, 1982.... .. .. .. . . . . .. .. . .. . ... .. .... .. . . . . . . . 8
Nushagak River sonar count distribution over time and
0 Figure 5.
sector in three day time periods , 1982. . . . . . . . . . . . . . . . . 9
Nushagak River daily escapement estimates by species, 1982.. ... . . . .. . . . . . . . .. ... . .. ....... ... ..... .. .. . ... .. . 17
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LIST OF TABLES
Table 1. Nushagak River escapement by species, 1982 ............... .
Table 2. Number of spawners per index point by species, 1982 ...... .
Table 3. Nushagak sonar drift net CPUE (fish per drift) by day and species, 1982 .............................•...........
Table 4. Nushagak River daily escapement, by species, estimated from gill net CPUE data and sonar counts collected ·at Nushagak Sonar, 1982 .................................. .
Table 5. Nushagak River run timing, by species, estimated by in-season sonar estimates, post-season estimates and commercial catch data, 1982 ........... : .............. .
Appendix Table 1. 1982 north bank of Nushagak River side scan sonar counts by sector .............. ; ........... .
Appendix Table 2. 1982 south bank of Nushagak River side scan sonar counts by sector .......................... .
Appendix Table 3. 1982 north bank of Nushagak River side scan sonar counts by hour ............................ .
Appendix Table 4. 1982 s'outh bank of Nushagak River side scan sonar counts by hour .....•.......................
Appendix Table 5. Nushagak River sockeye salmon escapement.age, weight, length and sex summaries, 1982 ...•.......
Appendix Table 6. Nushagak River chum salmon escapement.age, weight, length and sex summaries, 1982 .......... .
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INTRODUCTION
This report presents results of the 1982 side-scan sonar enumeration project conducted on the Nushagak River in Bristol Bay .. The objective
of this research is to obtain accurate estimates of salmon escapement
into the Nushagak River system soon after the fish have passed through
the commercial fishery. The feasibility of enumerating the five species
of adult salmon returning to the Nushagak was first examined using a single
side scan sonar counter in 1979. Because of .the programs apparent .success
it was expanded to include a second counter in 1980 (McBride eta] 1980).
Since then this program has been run annually with only slight variation in
site location and methodology between years.
The sonar gear and field camp is located near the village of Portage Creek
some 40.2 km (25 miles) from the terminus of the Nushagak commercial fishing
district (Figure 1). The single river channel, approximately 335m in width,
flows at between 1.22 m/sec. and 2.l3in/sec. Maximum depth is approximately
4.6 m in mid-channel. Water depth at 60 ft. offshore of .. the north and south
banks is about 2 meters and 3 meters, respectively (Figure 2).
METHODS AND MATERIALS
Sonar Units
The north and south bank sonar units were installed. on 9 June 1982. The
north bank counter (Serial #7820008) was installed approximately 300 meters
up-stream from the site used in 1981 to take advantage of swifter river
velocity. The south bank counter (Serial #7820002) was installed in the
same location as it has been since 1980. Both units monitored the salmon
1
J Lake ,J
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i Location of Nushagak sonar enumeration project Nushagak River, Bristol Bay, Alaska.
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Wood River
N
0 5 10 15
Miles
Nuyakuk Tower
~
Bristol Bay
2
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w ..., 5
"' "' '1-~
.s:: ..., c. 10
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15
North Bank
r...- r·· -, '-------' c-r L.:.:J ~ L.J L.J L.J L.J l.......:J L.:..J L.J [..__.J L.J :,_____]
-.------------~------- 335 meters ,......._,
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. ' .'· .... ~ . .... I:,
. ' :< ~· •• ·~·lio·~·, ... ' . . . .. . .... ~ . .. .. .... \ . .. .. ,, . . . . ..
• .. II ' <t. I ..
... * ''" '01 . . . .. . . 4. . ' . .. ·• . ,, . .. .• .• . . ..
·.: ... • .;i·' ...... . . . . ... ' ...... ~. 40
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• '.. ,.It'' • . .... ' . 0 0
0 0
0 0 .. 80
Figure 2. Nushagak River· bottpm profile at sonar s1te,:l982.
' .. ·- .... . . . . 240
:#.··-·" . . . . . . . .:: 280
•'· .. ' 320
., __; ~
South Bank
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run, recording counts on an hourly basis, until their removal on August 18.
Seasonal ADF&G personnel continually monitored the counters throughout the
season enabling rapid detection of problems (i.e. debris caught on sub
strate, mis-aimed transducers, under/over counting). The transducers in
both systems were aimed using an oscilloscope under the direction of Al
Menin (Bendix hydroacoustics) immediately following their installation and
then periodically during the season.
Calibration
Each counter was calibrated hourly, with the exception of the early morning
hours, for each day of operation. Calibration consisted of a ten minute
interval where counts produced by the machine were compared with those made
by trained technicians observing an oscilloscope pattern and hand tallying
observed counts. At the end of the interval the machine count was divided
into the oscilloscope·count to yield a percent agreement between the two.
If the percent agreement was consistently less than 80% or greater than 120%
indicating poor accurac~, an adjustment to the fish swimming velocity setting
on the machine was made. The calculated percent agreement was applied to the
corresponding hourly count for all sectors producing an adjusted hourly count.
Adjusted hourly counts were summed for all hours and apportioned by species to
provide rough in-season estimates of fish passage on a daily basis. This
calibration technique proved to be an effective method of measuring machine
accuracy·and adjusting counts.
Sector Distribution
Distribution of counts across the substrate is a good indication of the
performance of the sonar gear. Sector distribution for the north and south
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bank counters was calculated post-seasonally. This was accomplished by
determining the proportional contribution made by each sector for hours
having a high percent agreement (90% or better) between the machine and
oscilloscope and applying that number to the adjusted daily total. More
specifically the working formula was:
Where:
E sJ. X Nb = nb i = 1
s
= sum of i raw counts in sector j with percent agreement of 90% or better.
S = sum of all unadjusted totals with percent of 90% or better.
Nb= adjusted daily total for day b.
nb= estimated sector total for day b.
This procedure insured that for hours in which many false counts occurred,
causing a low percent agreement, sector distribution was not artificially
distorted.
Species Apportionment
In-season, adjusted sonar counts were apportioned by species based on 10
[] fathom drift gill net catches collected several times each day. King (8! in.),
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red (5-3/8 in.) and pink (4! in.) gear were generally fished with equal
effort. The total catch for all gear types was combined at the end of each
day and the percent contribution by species was calculated. Percentages were
applied to the adjusted daily count to yield an estimate of daily escapement
by species.
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Post-seasonally the total adjusted sonar count was apportioned based upon
percent contribution of each species to the combined aerial survey and tower
estimates (Bucher 1982). To arrive at estimates of daily escapement by
species, drift net data collected at the sonar site was examined in terms
of daily catch per unit of effort (CPUE). A daily CPUE (fish per drift)
for each species was calculated by combining catches for all gear types
fished on a given day, and dividing by the total number of drifts made that
day. The result was a daily abundance index, in fish per drift, by species.
An estimate of spawners per gill net index point was derived by dividing the
total number of sonar counts apportioned to a specie by that spectes
accumulative gill net index. Daily escapement estimates for each species
were calculated by multiplying the daily gill net index by the appropriate
number of spawners per gill net index point.
AWL Sampling
Sockeye and chum salmon were captured using a beach seine and sampled for
standard age, sex, and length data. Age was determined from scale samples,
length (mid-eye to fork tail) was recorded to the nearest millimeter, and
sex was determined visually.
Climatological Data
Weather data were collected twice daily at Portage Creek. Precipitation was
measured using aTay'lor Clear View rain gauge, wind direction and velocity
were estimated, air andwater.temperatures were measured to the nearest
degree. Water level was measured at the sonar site by means of a staff
. gauge to the nearest inch.
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RESULTS
Sonar Counts
A total of 2,539,002 adjusted counts were recorded by the two sonar units
(Appendix Tables 1 and 2) located at Portage Creek. The north bank contributed
1,709,837 counts, or 67 percent, while the south bank contributed 829,165
counts for the remaining 33 percent of the total. The distribution of.
counts between the north and south banks is opposite from prior years
(McBride et al 1980; McBride et al 1981) but not unprecedented. Nelson
(personal communications) pointed out that bank preference on the Wood
River is known to change from year to year depending on.the timing of peak
migration and the influence high tides have upon fi.sh passage.
Sector Distribution
The distribution of counts along the north bank substrate was, for the fourth ..
year, markedly bimodal (Figure 3). With 27 percent of the north bank counts
occurring in the last three .sectors, it is presumed that some fish were missed
going around the substrate. On the south bank however, the distribution was
more inshore. Only 8 percent of .the south bank counts occurred over the last
three sectors. It is suspected that the slower river velocities and gentle
slope of the north bank attributes to the offshore distribution. North bank
sector dis.tribution showed a marked shift inward on aoout the 9th of July
(Figure 4), which was coincident with the decline of.the red salmon run and
the start of the pink run. Presumably the reds are stronger swimmers and can
maintain themselves in the $Wift currents fo.und offshore. The south bank
demonstrated fairly consistent inshore distribution throughout the season.
Sonar counts by hour are presented for each sonar unit in Appendix Tables 3
and 4. The percent of sonar counts by hour for the entire season are relatively
consistent between units and show a significant decrease d-uring the hours of
darkness (Figure 3, lower half).
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\ \
NUSHAGAK RIVER SONAR COUNT DISTRIBUTION JUNE 1~ - AUGUST 18, 1982
' ' \ ' ' \
\
' \ ' \
\
' \
---- NORTH BANK ------------ SOUTH BANK
'
-~---=-----~---c-----~~-;,--------~------------~-------------04-~~~~~~~~~~~--~
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p E R c 5 E N T
0
0 1 2 3 ~ 5 6 7 8 9 10 11 12 S E C T 0 R
--- NORTH BANK ------------ SOUTH BANK
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.. ---.......... ' . ;
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... ----- .. ,
0 3 6 9 12' 15 18 21 2~ H 0 U R .
' Figure 3. Nushagak River sonar count distribution, June 14-August 18, 1982. I . .
8.
,-- ' L....J
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NORTH BANK
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SOUTH BANK
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Figure 4 .. Nushagak River sonar count distribution over time and sector in three day time periods, 1982.
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Species Apportionment
In-season estimates, summarized in Table 1, of species composition were
provided daily to fishery managers. Although these estimates were preliminary
they provided indispensable data with which to manage the fishery.
Post-seasonal estimates of species abundance (Table 1) tallied 124,939 king
salmon, 537,682 red salmon, 230,139 chum salmon, 227,237 coho salmon and
1,424,732 pink salmon. A comparison between the post-season estimates with
those made from combined aerial survey and tower counts (Table 1) shows them
to be very similar, indicating that they are probably more representative of
the actual escapement totals than were the in-season estimates.
Daily Escapement Estimates
The number of spawners per gill net index- point was calculated for each species
by dividing the total sonar counts apportioned to a species by the accumulative
gill net index for that species (Table 2): The calculations yielded 5,142 king, •
17,983 red, 6,538 chum, 15,149 coho and 31,661 pink spawners per gill net index
point. Daily escapement estimates by species were derived from the spawner
per gill net index point relationships and daily drift gill net CPUE data
(Table 3). Table 4 presents daily escapement estimates for the five species
returning to the Nushagak River in 1982.
AWL Sampling
Of the·l,360 samples collected at Nushagak Sonar, 633 were sockeye and 727
were chum. Preliminary analysis indicated that the sockeye were composed
of primarily age 52 fish, 60 percent of which were males and 40percent.
females. Average length of the males and females was·;587.1 mm· and 562.2 mm,
respectively (Appendix Table 5). Chum salmon statistics are presented in
Appendix Table 6.
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Table 1. Nushagak River Escapement by Species, 1982.
Estimated Escaeement King Red Chum Coho Pink Total
Inseason Estimate 355,460 507,800 634,750 228,509 812,483 2,539,002
Post-Season Estimate 124,939 537,682 230,139 227,237 1 ,424,732 2,544,733
Combined Aerial Survey Tower Estimate 147,000 601 ,000 256,000 229,676 1 ,611 ,230 2,844,906
Percent Agreement Between Combined Aerial Survey Tower Estimate .85 .90 .90 .99 .88 .89 and Post-Season Sonar Estimates
NOTE: The difference between totals for both sonar estimates is due to rounding error and post-seasonal extrapolation of sonar counts based on drift net CPUE data.
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Table 2. Number of spawners per index point by species, 1982.
Accumulative Spawners Per SEecies EscaEement Index Index Point
King 124,939 24.3 5,142
Red 537,682 29.9 17,983
Chum 230,139 35.2 6,538
Coho 227,237 15.0 15 '149
Pink 1,424,732 45.0 31 ,661
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J Table 3. Nushagak Sonar Drift Net CPUE (Fish Per Drift) by Date and Species, 1982.
J King Red Chum Coho Pink Date Daily Accum. Daily Ace urn. Daily Accum. Daily Accum. Daily Accum.
June 21 1.5 1.5 0 0 .6 .6 22 1.1 2.6 .4 .4 .6 1.2
J 23 1.3 4.·0 1.3 1.6 .3 1.4 24 3.3 7.3 2.2 3.8 1.2 2.6 25 0.8 9.8 .4 4.2 .8 3.4 26 0.5 10.3 0 4.2 2.2 5.6
J 27 .3 10.6 .5 4.7 1.9 7.5 28 .3 10.9 1.2 5.9 1.6 9.2 29 .6 11.5 .8 6.7 .3 9.4 30 1.0 12.5 .9 7.6 1.2 10.6
J July 1 1.1 13.6 2.3 9.8 1.5 12.1 2 1.1 14.6 2.1 11.9 1.1 13.2 3 1.0 15.6 2.0 13.9 3.3 16.5 4 .2 15.8 1.8 15.7 1.0 17.5 5 .8 16.6 1.7 17.4 .8 18.3
J 6 .8 17.4 2.1 19.5 .4 18.7 7 .7 18.1 1.3 20.7 .6 19.3 8 .3 18.4 1.4 22.2 1.4 20.7 9 .4 18.9 .3 22.5 .6 21.3
J 10 .6 19.5 .4 22.9 1.1 22.5 11 .3 19.8 .8 23.7 1.3 23.8 12 .5 20.3 .5 24.2 1.3 25.0 13 .4 20.7 .7 24.9 1.4 26.5 ' . 1 • 1
J 14 .4 21.1 .3 25.2 .4 26.9 . 1 .2 15 .9 22.0 .3 25.5 .4 27.4 . 1 .3 16 .3 22.2 .4 26.0 .4 27.7 . 1 .3 17 .4 22.7 .6 26.6 .6 28.4 . 1 • 1 .4 .7
] 18 .4 23.0 .4 27.0 1.1 29.4 • 1 .2 .3 1.0 19 . 1 23.1 .9 27.9 .9 30.3 0 .2 .4 1.4 20 .2 23.3 .3 28.2 1.3 31.6 0 .2 .6 1.9 21 . 1 23.4 .4 28.6 .4 32.0 • 1 .3 .6 2.5
J 22 .3 23.7 .3 28.9 .3 32.3 .2 .4 1.1 3.6 23 • 1 23.7 .4 29.3 .2 32.5 .3 . 7 .5 4.1 24 .2 23.9 .4 29.7 .4 32.9 .8 1.4 1.9 6.0 25 0 23.9 . 1 29.8 .4 33.2 .4 1.8 .8 6.8
J 26 .4 24.3 . 1 29.9 .4 33.6 .5 2.3 6.1 12.g 27 0 24.3 0 29.9 .4 34.0 0 2.3 2.9 15.9 28 0 24.3 0 29.9 .3 34.3 .7 3.0 3.4 19.3 29 0 24.3 0 29.9 . 1 34.4 .6 3.5 3.4 22.7
J 30 • 1 34.5 .4 4.0 4.6 27.2
31 • 1 34.7 .3 4.2 5.4 32.6 Aug. 1 0 34.7 .2 4.4 3.7 36.3
2 .5 35.2 .5 4.9 2. 1 38.4 3 0 35.2 0 4.9 1. 7 40.1 'l 4 0 35.2 0 4.9 1. 2 41.3
~ 5 0 35.2 • 1 5.0 .3 41.5 6 .4 5.4 .9 42.5 7 . 1 5.5 • 1 42.5 ,-1 8 . 1 5.5 .3 42.8 9 .4 6.0 .4 43.2 ,_
10 .a 6.7 1.1 44.3 11 3.8 10.5 .6 45.0
J 12 1.5 12.0 0 45.0 13 1.0 13.0 0 45.0 14 0 13.0 0 45.0 15 2.0 15.0
J 16 17 18 19
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J Table 4. Nushagak River daily escapement, by species, estimated from gill net CPUE data and sonar counts
cc~lected at Nushagak Sonar, 1982
J Estimated Esca ement King Red Chum Coho Pink Total
Date Daily Accum. Daily Accum. Daily Accum. Daily Accurn. Daily Accum. Dally Accum.
J June 14 281 281 120 120 100 100 501 501 15 589 870 252 372 210 310 1 ,051 1 ,552 16 557 1 ,427 239 611 199 509 995 2,547 17 1 .432 2,859 614 1,225 512 1 ,021 2,558 5,105 18 1,583 4,442 678 1 ,903 565 1,586 2,826 7,931
J 19 1.123 5,565 481 2,384 401 1 ,987 2,005 9,936 20 790 6,355 338 2~722 282 2,269 1 glQ ] ] Jg6 21 7,836 14,191 0 2,722 3,895 6,164 11.731 23,077 22 5,746 19,937 7,133 9,855 3,895 10,059 16,774 39,851 23 6,791 26,728 23,182 33,037 1 ,948 12,007 31,921 71 ,772 24 17,239 43,967 39,230 72,267 7,790 19,797 64,259 136,031
J 25 4,179 48,146 7,133 79,400 5,194 24,991 16 ~Qfi ] 52 53Z 26 2,612 50,758 0 79,400 14,282 39,273 16,894 169,431 27 1 ,567 52,325 8,916 88,316 12,335 51 ,608 22,818 192,249 28 1,567 53,892 21,398 109,714 10,387 61 ,996 33,352 225,601 29 3,134 57,027 14,266 123,980 1 ,948 63,944 19,348 244,949
J 30 5,224 62 ,251 16,049 140,029 7,790 71,734 29 023 2H Ql2
July I 5,746 67,997 41,014 181 ,042 9,738 81,472 56,498 330,510 2 5,746 73,744 37,447 218,490 7,141 88,613 50,334 380,844 3 5,224 78,968 35,664 254,154 21 ,424 110,037 62,312 443,156 4 1 ,045 80,012 32,098 286,251 6,492 116,529 39,635 482,791
D 5 4,179 84,192 30,314 316,566 5 '194 121 ,722 39 68Z 522 gzs 6 4,179 88,371 37,447 354,013 2,597 124,319 44,223 566,701 7 3,657 92,028 23,182 377,194 3,246 127 ,565 30,085 596,786 8 1 ,567 93,595 24,965 402,159 9,089 136,654 35,621 632,407 9 2,090 95,684 5,350 407,509 3,895 140,549 11 ,335 643,742
10 3,134 98,819 7,133 414,642 7,141 147,620 lZ :108 661 ] so
J 11 1,567 100,386 14,266 428,907 8,440 156,130 24,273 685,423 12 2,612 102,998 8,916 437,823 8,440 164,570 19,968 705,391 13 2,090 105,088 12,482 450,306 9,089 173,658 23,661 729,052 14 2,090 107,177 5,350 455,655 2,597 176,255 3,216 3,216 13,253 742,305 15 4,702 111 ,879 5,350 461,005 2,597 178,852 3,216 6 !132 ]5,865 Z58 1ZO
D 16 1,567 113,446 7,133 468,138 2,597 181 ,449 3,216 9,648 14,513 772,683 17 2,090 115,536 10,699 478,837 3,895 185,344 1,354 1,354 3,216 12,864 21,254 793,937 18 2,090 117,625 7,133 485,970 7 '141 192,485 1 ,354 2,708 12,864 25,729 30,582 842,519 19 522 118,148 16,049 502,018 5,843 198,328 0 2,708 9,648 35,377 32,062 856,581 20 1 ,045 119,192 5,350 507,368 8,440 206,768 0 2,708 12,864 48,241 27,699 884,280 21 522 119,715 7,133 514,501 2,597 209,364 1,354 4,062 19,297 67,538 30,903 915,183
J 22 1 ,567 121,282 5,350 519,850 1,948 211,312 2,708 6,771 19,297 86,835 30,870 946,053 23 522 121,804 7,133 526,983 1,298 212,610 4,062 10,833 35,377 122,212 48,392 994,445 24 1,045 122,849 7,133 534,116 2,597 215,207 10,833 21 ,666 16,081 138,292 37,689 1,032,134 25 0 122,849 1,783 535,899 2,597 217,804 5,416 21 ,Qa~ !21 ,lQfi ]22 J26 ZQ 202 ] 103 036 26 2,090 124,939 1,783 537,682 2,597 220,401 6,771 33,853 25,729 225,127 38,970 1,142,006
J 27 0 2,597 222,998 0 33,853 196,182 421,309 192,779 1,340,785 28 1,948 224,945 9,479 43,331 93,267 514,576 104,694 1 ,445,479 29 649 225,594 8,125 51 ,456 109,347 623,923 118,121 1 ,563,600 30 649 226,244 5,416 2!2,8Z2 1Q2 ~4Z ZJ3.ZZ1 lHi !112 1 fiZ2 012 31 649 226,893 4,062 60,935 147,941 881,211 152,652 1 ,831 ,664
J Aug. 1 0 226,893 2,708 63,643 173,669 1 ,054,881 176,377 2,008,041
2 3,246 230,139 6,771 70,413 118,996 1,173,876 129,013 2,137,054 3 0 230,139 0 70,413 67,538 1,241 ,415 67,538 2,204,592 4 0 230,139 0 70,413 54,674 1,296,088 54,674 2,259,266 5 1 ,354 71,767 38,593 1,334,681 39,947 2,299,213 6 5,416 77,184 9,648 1 ,344,330 15,064 2,314,277
J 7 1 ,354 78,538 3,216 1 ,347 ,546 4,570 2,318,847 8 1,354 79,892 9,648 1,357,194 11 .002 2,329,849 9 5,416 85,308 12,864 1 ,370,059 ·18,280 2,348,129
lo 10,833 96,141 35,377 1,405,436 46,210 2,394",339 11 51 ,456 147,597 19,297 1 ,424 ,732 70,753 2,465,092
J 12 20,312 167,908 0 1 ,424,732 20,312. 2,485,404 13 13,541 181 ,449 0 13,541 2,498,945 14 0 181,449 0 0 2,498,9~5 15 27,082 208,531 27,082 2,526,027 16 8,180 216,711 a, 120 2,534,207 17 7,873 224,584 7,873 2,542,080
J 18 2,653 227,237 2,653 2,544,733
J J
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Climatological Data
Climatological data were collected twice daily throughout the season. An
estimated 5.69 inches of rain fell from June 6 to August 18, 1982 for an
average precipitation of .08 inches per day. ~1ean ambient temperature was
11.5° C (52.7° F) while the mean surface water temperature was estimated
to be 10.9° C (51.7° F). Water level dropped an estimated 51 inches over
the course of the season.
Discussion
The operation of .side scan sonar counters at Portage Creek has, for the
past four years, proved exceedingly useful for in-season fishery management.
The array of problems associated with counting fish with sonar at this site
appears to be narrowing. Still, the over-riding problem is how to accurately
apportion sonar counts into individual species. Current in-season methods are
imprecise and tend to suggest artificially altered run timing and magnitude of
species less dominant in numbers. Following the end of the 1982 field season
and during the subsequent data analysis, the problem of species apportionment
was addressed and the contributing factors identified (Minard 1982). In-season
apportionment appears to be skewed by two factors: (1) differing catchability
coefficients for each species and (2) the different time and magnitude with
which each speCies return. An example of how these two factors work in concert
to influence the apportionment process is easily seen in.the case of the king
and red salmon counts. From historical commercial catch data it is clear.that
the king salmon run in the Nushagak typically peaks in. the third week of June
while the sockeye run has traditionally peaked around the fo.urth of July. In
1982 in-season apportionment suggested the peak of escapement for both these
species at about the 5th of July. The reason for this apparent misapportion-
15
J J J ]
J J D J J J J J J J J ]
]
J J
ment of king counts is simply as the sockeye run begins to build, so does
the total number of sonar counts to be apportioned. A single king salmon
early in the run means less, in terms of total escapement, than does a
single king caught during the peak of the sockeye run because of the
different magnitude of the sonar counts at those two times. To compound
the problem, king salmon are readily captured in both king and sockeye drift
nets, used to apportion counts, while sockeye are caught primarily in the
smaller red gear.
To get around the apportionment problem, the 1982 data was evaluated in terms
of daily gill net CPUE by species. The results were encouraging and when
daily escapement estimates using this methodology were plotted and compared
to other indicators of run timing, it was found that the post-season estimates
mimicked more closely the expected trends in run timing. Figure 5 shows
comparative graphs of run timing and magnitude for the in-season and post
season estimates. Table 5 is a comparison of peak catches in the commercial
district with in-season and post-season estimates of peak escapement by
species in 1982. The methods used post-seasonally are probably more
indicative of actual escapement. Application of these methods to in-season
counts has yet to be tested. Problems arise since the calculation of spawners
per index point is dependent upon having the total escapement by species
already estimated.
Offshore distribution appeared to be less of a problem than reported in past
years. This was probably due to at least two factors. First, a new site
approximately 300 meters upstream on the north bank was used for the operation
16
J Fi"gure 5 .. Nushagak daily escapement estimates by speci es, 1.982. 90
J 80
J 70
~ Pink
J "' -g 60 INSEASON ESTIMATE "' "' :::l. 0
l ;:: 50 Chum ~ .., <:::
J ~ 40 QJ c.
"' u
"' 30 UJ
D Red 20
0 10
D 1 25 30 5 10 20 25 30 4 9 14 19 June July Pink August
J 100 DATE
0 9.0
80 POST SEASON ESJI~~TE
0 ~ 70 "'
0 "0 <:::
"' "' :::l 60 0 .<=
Coho J f-~ ..., 50 <::: QJ
.E QJ
0 c. 40 "' u
"' UJ
D 30
0 20
10
: I LJ
15 20 25 30 5 10 15 20 25 30 4 9 14 .19 Q June 17 July August
·----- -- ·-- - ····-- . --
.]
J J J J J 0 J 0 J 0 0
D I [] ! ..
0 0
J J '
I
Table 5. Nushagak River run timing by species estimated by in-season sonar estimates, post-season sonar estimates and commercia 1 catch data, 1982. ·
Run Timing Peak Da
In-Season Post-Season Commercial seecies Sonar Estimate Sonar Estimate Catch
King 7/ 6 6/24 6/23 - 6/26
Red 7/ 6 7/ 1 7/ 5 - 7/ 6
Chum 7/10 7/ 3 7/ 5- 71 7
Coho 7/25 8/ll 8/ 9 - 8/10
Pink 7/26 7/26 7/25 - 7/26
18
J ]
D D 0 D 0 0 0 0 J J 8 0 0 []
J D 0
.of the sonar gear. This site took advantage of a 3% higher river velocity
which presumably forces the fish shoreward. Secondly, 1982 was a pink
year. The pinks being smaller than are sockeye, chum and coho, tend to be
weaker swimmers. Their preference for the slower velocities found toward
the inner sectors was clearly demonstrated when sector distribution was
evaluated over time (Figure 4). Because of the overall magnitude of the
pink run, the seasonal plot of sector distribution was probably skewed
inward. Had the plot concerned only the sockeye run, then I believe there
would have been a higher percentage of counts occurring in the offshore
sectors.
1982 was the first year that the program was extended to enumerate coho
salmon. In-season and post-season analysis indicated that there were no
special problems with enumerating this species with sonar and.that aspect
of this year's operation is considered to be highly successful.
. 19
J J J ]
J J 0 0 0 J 0 0 g
0 0 [J
J 0 D
LITERATURE CITED
Bucher, W. A. Spawning Ground Surveys in the Nushagak and Togiak Districts of Bristol Bay, 1982. Bristol Bay Data Report No. 93. 29 pp.
McBride, D. H. and Mesiar, D. 1980. Nushagak Sonar Enumeration Project, 1980. ADF&G Bristol Bay Data Report No. 83. 45 pp.
McBride, D. H. and Mesiar, D. 1981. Nushagak Sonar Enumeration Project, 1981. ADF&G Bristol Bay Data Report No. 88. 24 pp.
Minard, R. E. 1982. Nushagak Sonar Species Apportionment Results/Methods, 1982. ADF&G Division of Commercial Fisheries Memorandum. 10 pp. ·
20
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J 0 J
0 0 0
J J •
J
Appendix Table 1.
1982 NORTH BANK OF HUSHAGAK RIVER SIDE SCAN SONAR COUNTS BY SECTOR
DATE
6 14 6 IS 616 6 17 6 1B
619 620 621 622 623
624 625 626 6 27 628
629 630 7 1 7 2 7 J
7 4 7 5 7 6 7 7 7 •
7 9 710 711 7 12 713
7 14.
71S 7 16 7 17 7 18
719 720 721 722 723
SECTOR I 2
2 4
21 1J 19
54 58 58 62 58
BO 356 218 140 631
157 177
2879 7262 8273
105 351 295
2J
•• ISO
93 1960
916 1665·
244 741 929
1682 3515
6586 5729 2792 6166 4769
4 6
IS 18 19
17 27 84
174 101
156 332 704
1117 6019
623 910
8748 5014 2816
277 785 855 133 583
366 347 737
1672 522
581 3181 .2227 2523 2279
4791 5997 3429
12305 13877
7 24 16080 1-4202 7 25 19308 12007 7 26 6476 15083 7 27 18035 10229 7 28 11081 10183
729 730 731 8 I • 2
• J • 4 • 5 • 6 • 7
• • • 9 810 8 II • 12
• 1J 814 B IS 816 a 17
• 18
15750 17660
6995 11334
5240
1962 1238 3393
209 659
156 290 623 527 408
593 266 228 380
77
82
8340 11809 24168 15017 14636
6083 2898 4829
461 275
326 ... 1639 3371
314
438 1348
381 2255 2552
153
3
7 9
25 273 410
413 603 951
1060 555
405 482
2272 2338 6191
650 892
5981 6376 1672
292 472 940 790
1246
801 581
1552 2886 U40
2463 3518 2312 1736 1391
2448 2436 1601 6555
10916
3514 2474 7746 2590 2086
1490 1637 5169 2126 3840
4196 1338 1799
708 474
533 1243 1148 4457 565
670 1637
533 1399 151-4
435
14 II IS
6 25
13 25
105 71
115
215 277
1756 825
2102
322 398
4039 2334
704
107 623 ... 622 396
304 396
1631 1731 628
599 3255 1251 686 927
5
6 II 4 9
12
5 18
152 74
127
233 239 486 288
1212
308 225
1451 1837 1232
264 1837 437
2287 430
199 21581
925 438 264
253 522
1459 457 773
1203 --- 760·. 709 405 318 640
1820 699 3811 980
1296 1067 3148
754 655
673 338
1305 444 610
1049 355 687 480 290
467 343 279
1225 251
825 460 381. 268 410
169
532 407
1859 580 284 ... 280 375 105 102
195 134 410 228 168
.118 126
66 356 126
206 39
247
" 96
128
6
9 24 II
2 9
5 16 ... 86
352
417 478 684 252 710
229 163
1741 1254 1584
2904 1911 12-44 2820 2089
3087 SOB
158-4 6109 202
197 471
118"1 276 502
7
54 45 II 22 37
23 21
!66 155 240
584 578
1240 749
1184
602 234
2611 2615 1760
4792 2993 8484 4405 7995
21-\3 1783 6491 2328 661
356 1565 2331 969
2743
• 65 81 78
137 153
60 30
217 160 376
461 1293 972 873
1937
1173 1904 3057 6203 2024
13516 9851
16196 11710 20524
8210 19671 5927 3542 585
1142 2171 2217 1184 2820
. 887 -2026 ---1689 754 1"17"1 -43-43
2245 7852 14977 633 1473 2171
1033 1195 1507
590 295
1478 625 353
270 444
8293 1460
165
41 205 229 240 214
81 227 lBO 198 439
387 249 114 93 45
112
1124 817
4198 1679 723
1060 1536 1114 1444
534
469 636 458
1120 917
311 310 672 540 816
980 304 152 140
51
266
2104 6018 6734 2740 2113
1304 2096 1663 1592 1246
1025 1050 931
5229 2079
1052 1011
525 ... 534
1341 366 171 344 "167
148
9
36
" 189 400 470
242 93
292 236 387
686 946
1458 516 940
1957 2810 1294 2226 1320
8189 4759 9725
13149 14202
20532 14468 6225 1393 1556
2397 1784 1421 4427 1236
10
51 167 210
"' 625
481 102 364 306 689
724 1888 2073
817 1133
1492 4162 2232 1578 2024
6263 8809 -4943
13704 8221
10080 10737 3700 1433 684
995 1229
777 11275
1275
1689 ... 1Z24-4945 1896 7216 2069 1301 1301 3144 1809
8756 11550 2276 1526 6490 6876 4808 15271 7660 11125
14083 3019 1266 1296 1030
846 941 679
3753 1284
437 363 115 421 565
644 195 343 192 148
128
4374 2096 1898 1234 1475
1293 1150 600
2412 673
·JIB 305 262 448
·1098
851 132 381 93 71
107
II
25 167 174 673 582
347 152 330 664 595
413 4383 4950 3716 1334
1577 4149 1763 1059 1760
1036 1015 1400 2531 2028
1695 2423
11086 2010 476
690 1229
692 1823 1777
971 3544 2230 1314 1862
1836 13066 5450 8103 3500
1426 2178 1512 1277 1170
1822 5307 931
2-467 .,. 407 455 279 290 502
799 257 495 23 96
82
CUMULATIVE 12 TOTAL TOTAL
66 160 137 415 376
239 178
1739 673 698
874 3904 9225
10554 11026
9532 11951
1383 669
1408
2281 1699 2086 2513
21472
683 4043 216-4 3562" 991
431 1803
218 639
2666
844 2914
563 939 872
2176 668
5887 1661 1743
2031 1661 2721 919
1958
1813 3148 836
-4029 703
378 400
81 84 31
335 187 418 64 19
41
339-' 784 890
2467 2737
1899 1323 4606 3721 4293
5248 15156 26038 22185 34419
18622 27975 37179 38427 26577
40026 35105 47045
. 54687 79274
48250 76631 43982 28020
9374
10348 21469 17018 27677 21904
339 1123 2013 4480 7217
9116 10439 15045 18766 23059
28307 -43-463 69501 91686
126105
1-44727 172702 209881 248308 274885
314911 350016 397061 451748 531022
579272 655903 699885 727905 737279
747627 769096 786114 813791 835695
25118 860813 35146 895959 45932 941891 36677 978568 45775 1024343
63760 1088103 59929 1148032 71425 1219457 67075 1286532 51506 1338038
512t9 1389337 44754 1434091 56479 1490570 382t8 .1528818 320 6 "1560824 I .
~~~!: ~!~~~~~: 15782 1615800 21336 1637136 8164 1645300
45~4 1649884 5939 1655823 5869! 1661692
12766 1674458 5649 1680107
8019 1688176 5440 1693616 3844 1697460 5280 1702740 5246 1707986
1851 i709837
198371 248234 132962 .53058 5123.3 55841 99361 209639 202093 181660 124803 152582 1709837
21
J J ~]
J J J J
0 J J J J J 0
J '
~]
·Appendix Table 2.
1982 SOUTH BANK OF NUSHAGAK RIVER SIDE SCAN SONAR COUNTS BY SECTOR
SECTOR DATE
6 14 41 6 15 18 6 16 26 6 17 42 6 18 1
6 19 7 6 20 23 6 21 24 6 22 234 6 23 74
6 24 127 6 25 81 6 26 68 6 27 407 6 28 362
6 29 547 6 30 2255 7 1 4218 7 2 13736 7 3 1686
2
a • 0
10 5
11 32 34
1278 329
121 16 79
222 225
1013 1298
11872 15698
5528
7 4 2492 7603 7 s 5618 11301 7 6 3922 . 2350 7 7 132 2576 7 8 130 1868
7 9 142 379 7 10 20 183 711 52 99 7 12 24 196 713 5 128
7 14 104 404 7 15 95 143 7 16 309 970 7 17 341 2846 7 18 1199 2172
7 19 2299 2227 7 20 1172 5336 7 21 6269 4011 7 22 12134 1-4275 7 23 2317 6780
7 24 22504 8223 7 25 12896 2466 7 26 28402 11895 7 27 54041 10665 7 28 41288 14338
3
5
• 1 1 1
6 1
27 579 439
236 B!
141 505 496
3039 2101 1379 1369 1511
1608 2635 2403 1521 1319
450 346 303 149 101
236 155
. 266 662 .,. !55
1060 754
1777 8368
2060 1582 1272
426 435
7 29 730 7 31 a 1 a 2
24682 20051 30074 25940
8946
5483 162
a 3
• • a 5 ~ 6 a 7
a· a a •
. 8 10 911 a 12
a 13 914 a 15 916 a 17
a 1a
3288 921 692
. 423 623
372 142
1950 . 669
757·
371 461 36
282 62
600
9851 747 8526 620 5859 413 9497 603
2975 193 575 347 145 640 307 581 160 212
136 471 58 94
506 1481 2338 3272
69 581
429 869 214 873 173 . 932 179 1217 452 BOO
57 29
• 16 30 2 6 5
7 2
325 1776
433
644 564 507 912 557
965 677 292 605 365
366 530 911 334 517
490 202 303 245 B!
271 511 339 649 469
129 972 307 954
3671
73~ 672 290 270 211
122 262 439 210 450
169 252 939 771 255
453 346 572 909 573
417 389 545 384 297
23
5
3 14 3 5 6
3 2
492 2082
433
704 1434 1312 1867 392
••• 539 460 576 319
199 563
"541 163 354
492 255 577 652 531
254 1224 U7 519 290
126 939 123 954 496
192 196 120
B! 119
10 149 191 70
459
47 110 434 954 260
136 163 144 220 229
52 63
110 205
21
2
6
0 16 5 1
• 3 5
507 1332
336
427 935 912
1190 57
241 440 309 257 152
390 431 211 23
196
309 235 309 425 397
173 630 244 416 135
194 564
22 353 397
57 169 95 22 13
0 37 70 7
170
47 39 .
269 392 146
27 37
" <5· 139
29 32 55
269 62
7
9 22 16
1 3
12 7
1345 2187
946
703 1901 1430 2035 779
531 914 931 776 729
329 1060
634 404 415
••• 503 <31 651 464
346 1058
661 629 !99
403 966 302
2209 1058
217 293 350
65 132
25 296 177 161 409
56 252 495 979 674
227 262 235 !55 267
226 63
255 219 164
a
21 27 9 3 6
13 3
1141 1887
591
427 1096 1413 1507 517
396 952 399 736 691
299 962 945 504 475
m 463 443 379 760
334 975 603 573 194
229 972 134 776 959
192 415 435
•• 204
96 174 131
. 245 .
510
25 260 640 639 644
309 599 102 115. 313
104 111 !59 102
92
13
9
7
• 926 1181
490
253 790 692
1129 541
643 272 221 565 792
140 663 555 179 429
993 405 309 301 471
299 630 395 199 247
239 650
95 1296
992
164 645 316 !03 257
122 299 131 203 229
45 165 579 340 676
507. 357 530 !55 176
214 32
221 205 246
15
343126 199205 57564 31682 25308 15467 35411 29802 24875
22
10
11 46 6
10 17
5 4
912 1113
470
364 693 597 946 713
396 670 362 559 639
!59 979 356 310 601
1035 392 361 317 330
306 226 259 232 135
171 939 269
1473 529
103 495 479 124 509
56 237 219 301 212
22 315 423 431 600
599 829 289 225 245
336 119 255 436
92
13
25547
11
14 34 10
1 13
17 2
569 749 267
326 467 299 569 470
177 469 362 656 699
73 979 304 210 509
269 379 251 293 249
249 511 lOB 307 111
216 59! 194
1797 1058
50 293 659 130 224
51 !00 119 105 297
42 236
"516 464 397
272 913 433
75 176
249 !51
97 129 297
32
CUHULATIVE 12 TOTAL TOTAL
13 21 17
2 14
16 2
737 739 374
269 966 492 664 333
224 321 557 519 660
164 910
·224 210 331
341 259 145 139 B!
362 202 129 290
73
113 309 313
2601 2315
•• 247 453 101 257
65 50
!SO ., 25
19 119 206 306 254
54 334 163 110 121
110 16 62 64 92
11
162 267 104
91
•• 107 97
6939 15134
5071
4601 9024 7931
11852 5432
8650 10806 21252 36052 13760
162 429 533 624 713
920 907
7846 22980 28051
32652 41676 49607 61459 66891
75541 86347
107599 143651 157411
13810 171221 26429 197650 13256 210906 6565 217471 7133 224604
6551 231155 3642 234797 3583 238380 3770 242150 3598 245748
3326 249074 6360 255434 4698 260132 7650 267782 5687 273469
6491 279960 14057 294017 12782 306799 40399 347198 28940 376138
34575 410713 20338 431051 44754 475805 66077 541882 57986 599868
30864 630732 32243 662975 40866 703841 33!;i91 737432 21806 759238
6927 766165 3490 769655 5966 775621 6577 782198 4891 787089
3561 790650 4133 794783 6501 801284 8288 809572 3645 813217
3406 816623 2524 919147 2900 822047 3689 825736 2627 828363
802 929165
21276 19902 829165
J ]
J J
]
~-] L
0 J J J J
J
[J
0
0
Appendix Table 3.
1982 NORTH BANK OF NUSHAGAK RIVER SIDE SCAN SONAR COUNTS BY HOUR
H 0 U R DATE 2 3 4 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 TOTAL
6 14 4 6 15 13 6 16 2 6 17 62 6 18 102
6 19 6 20 6 21 622 6 23
624 6 25 6 ::!6 6 ::!7 6 28
139 41 53
146 119
95 207 289 147 468
4 8 7
84 90
186 33 54 97
136
61 125 145 125 295
16 2
58 BB
142 42 49 65
101
39 139 93
154 226
2 18 s
78 81
75 37 42 72
123
26 118
48 170 205
11 1 17 33 9 12
73 72 89 100
6 29 5 17 44 79
121
4 22 5B 99
128
6 21 61
134 83
lS 34 26
130 58
7 B 13 BB 3B 57 44
58 23 56 55 59 110 81 85 29 53
13 47 25 24 6 22 28 28 41 49
103 102 199 246
44 18 40 34 42 32 44 25 32 24 96 105 63 86
74 0 0 155 149 190 95 198 203
49 61 80 158 126 56 105 162 147
92 51 77
116 121
70 57 61 89
115
99 57 so
161 115
146 54 82 86 63
82 79 30 13 41 49 57 223 93 65
65 67 60 69
311 200 347 127 23 105
57 68
243 102 92
146 114 as 96 211 221 212 us BJ 353 709 2579 2044 86 154 34 505 398 248 73 67 665 801 14 509 1575 2147 304 271 42 1067 2383 705 20 819 111 478 1894 914 477 431 2447 1947 96 BOO
57 52
403 163 145
72 76
351 146 489
33 110 289 193 230
92
" 197 231 199
38 42
228 293 176
31 52
299 185 260
51 so
189 188 384
59
" 410 185 279
95 41
854 150 368
32 44
210 169 169
290 331 316 331 299 459 480 521 374 157 252 203 99 954 565 1110 1347 1018 1163 765 793 410 3786 3498 847 1456 4030 3205 170 481 314 232 881 919 4684 3187 1041 1071 1626 957 80 1489 4695 2618 1610 4793 2543 1883 1626 1831
5 51 47
135 167
40 so
134 126 323
114 229 688 955 573
339 784 890
2467 2737
1899 1323 4606 3721 4293
5248 15156 26038 22185 34419
6 29 268 6 30 173 7 1 .ns 7 2 203 7 3 534
m~semmm==1m~mrna1mmm--~--m•~ 90 30 54 144 317 4635 1978 2896 1499 2392 2949 1424 2269 453 798 1452 778 1182 633 293 248 736 552 27975
589 127 94 1207 2730 3126 4609 1712 4973 3887 2048 1723 328 254 346 813 1062 1142 2241 1533 434 334 1152 37179 1aO 207 1015 2044 459 4213 6687 4309 4828 954 604 498 1278 1020 1042 1411 1605 2333 1783 942 2aO 310 222 38427 266 99 940 1965 374 1192 1963 3031 1317 333 1275 409 610 513 739 990 2399 2251 2174 1402 779 792 200 26577
7 4 137 885 215 431 3941 3408 2993 2963 S757 6148 1464 332 1283 1335 820 516 1276 1127 830 1737 480 651 894 403 7 5 132 529 1066 1360 3295 987 882 1060 968 1467 2612 3696 1093 2068 4749 1376 1608 1249 720 2918 445 171 344 310 7 6 460 489 722 1305 1198 769 2648 4963 2522 549 1011 1794 902 786 2991 4001 1322 709 1091 3047 972 1713 6510 4571 7 7 2833 2392 1561 1387 2343 1617 1534 956 2649 2i31 3955 1380 1783 1859 2470 5379 3396 5616 1391 2009 1583 825 1654 1984 7 8 3501 2629 1384 817 1506 937 281 571 2186 3968 5851 3729 5593 6729 8574 9621 3835 1977 776 1820 844 1515 4822 5808
7 9 2307 577 427 7 10 2048 1a69 895 7 11 2839 2385 613 7 12 340 299 262 7 13 209 242 162
174 442 1595 4985 1a19 1024 2.<132 1156 22a7 144a 2693 2823 2112 2418 1740 2124 a46 2097 2368 3411 4945 73 171 236 1344 43.<13 1642 1559 4042 4043 5.<167 5017 5755 5609 7554 5694 3115 2328 23S7 2902 <1728 38<10
618 479 487 709 1303 782 2004 2073 3310 2285 3387 2948 2356 1407 1280 1575 1039 968 4144 4717 274 857 2406 3991 139.<1 328 201 248 899 1876 259 2892 2741 2917 1939 <120 434· 895 896 427 770 329 11469mrn~mm273116m~~===~~millmm
7 14 203 264 63 54 90 125 127 96 90 142 1 1s 195 143 zn 1a6 333 265 -269 313 455 27B 7 16 256a 1742 149 155 379 500 412 374 303 270 7 17 1013 1462 1351 1155 996 770 1469 1366 1411 a16 7 18 321 102 172 67 555 563 1884 1080 1191 1005
246 677 324 510 755 1101 864 547 570 776 653 698 sse 14so 2593 43o 95 630 526 278 765 a61 521 447
Sa2 2a5 655 502 355 2361 998 456 654 739 479 1030 2153 866 796 297
563 525 862 642 n3 7os 747 1282 1634 2312 3796 926 a43 687 1182 1539 945 847 278 974 1226 2124 2796 1976 434 339 764 2702 2357 1354
7 19 435 375 319 789 334 1963 892 1248 1018 753 342 1816 1393 1540 1714 809 709 1207 1915 847 1460 365 714 2161 7 20 670 807 765 972 569 868 3642 946 1a06 1255 927 1250 1420 733 2186 1117 4070 979 1837 1163 1670 1200 1190 3104 7 21 1006 543 614 587 769 228 766 1148 4820 1497 699 1172 2496 1347 1724 1536 3740 5159 3411 4154 4299 1691 1208 1318 7 22 1185 952 831 1650 -1397 2437 995 1360 2633 1462 1237 1434 712 1520 1308 1517 2206 1420 1616 1488 2099 1830 2810 57a. 7 23 148 166 410 666 2393 3280 1320 393 557 1284 1355 2126 1811 3935 1164 1454 4624 2239 2479 1836 3333 3173 3999 1630
7 24.1358 408 354 278 1299 1700 1444 1165 319 1671 841 2108 13S7 3780 1600 2102 5245 4706 2851 3505 9663 5841 5421 4744 7 25 1569 611 207 261 1872 2401 5753 5381 3026 1799 724 1336 2110 615 1046 1597 3895 1462 3436 5878 5091 2a52 2519 44a8 7 26 3324 2054 967 614 1401 2557 7608 8341 3483 3244 3269 2891 1245 2634 1167 3061 3561 3711 2764 2501 3299 2280 1590 3a59 7 27 3694 1455 723 675 960 2254 4740 4908 6584 7561 4332 2857 1112 1327 1784 3007 2666 844 2336 2422 2569 3859 2441 1965 7 28 1669 3a6 390 sao 831 2413 4139 3433 5438 6654 2194 787 1680 689 1765 2673 2218 2495 1558 1483 2239 2002 2291 1499
40026 35105 47045 54687 79274
4a250 76631 43982 28020
9374
10348 21469 17018 27677 21904
25118 35146 45932 36677 45775
63760 59929 71425 67075 51506-
7 29 775 704 7 30 1945 1409 7 31 1859 1467 a 1 1475 930 8 2 1212 839
417 448 532 307__ 405
270 75a 2000 3283 2705 4938 5880 2093 1925 1460 2238 1961 2521 2110 3514 1916 2296 1992 592 2569 2382 51299 571 1501 5496 2006 2181 1554 2033 1397 1413 1396 1006 1435 2190 2892 1989 2543 1785 1395 1865 2064 2240 44754 374 921 2908 5251 3885 4092 4504 1240 1920 2041 1618 1484 5342 1981 3235 2751 1268 1507 1905 3095 1299 56479
a 3 399 8 4 1200 8 5 329 a 6 473 s 7 267
9 8 113 a 9 43 a 10 169 a 11 60 8 12 8
a 13 20 e 14 35 9 15 30 s 16 23 s 17 24
254 _386_17_97 __ 3261 3075_2601_1578 1761_1066_1524 _1368 __ 922 __ 876 ___ 924 ___ 949 ___ 911__1156 _2675_2999__ 3193 __ 2260_ 38248_ 207 898 2315 3550 4542 2578 2042 2044 1177 696 479 1022 594 1170 794 673 831 1243 1406 806 483 32006
428 404 393 964 739 474 176 303 394 513 546 751 174 223 205
563 1284 1920 2109 1757 647. 351 272 473 744 1070 372 400 a32 1212 1775 1290 839 775 475 342 393 326 341 692 765 594 841 498 "s23 405 340 684 621 1311 702 65a 1693 1096 775 652 618 2623 1078 885 419 1321 1481 282 485 410 813 286 285 487 466 .<131 124 176 279
747 610 487 384 2241 1060 1190 899 333 464 601 538 756 2185 761 385 618 540 789 928 834 1073 11.<19 . 673 799 623 959 783 1262 752 353 223 620 558 399 329 296 295 181 93
20794 18400 15782 21336 a164
so 50
104 32 31
116 52 64 45 31
34 11 32 262 29 29 33 20 23 38
76 128 345 326 292 290 94 105 234 273 332 146 67 131 398 ·197 335 216 so 73 820 1163 706 700 24 31 191 791 792 385
221 261 102- 119 57 99
767 719 261 114
269 182 93 275 114 154 154 126 177 2Sa 246 281" 615 467 1n 90 111 152 206 256 292 829 . 896 1149 869 660 492 464 86 93 183 665 399 371 172
140 481 528 630 132
173 400 629 493 126
167 457 701 662 260
324 454 SOB 379 199
212 334 316
76 246
~u=~mmmmmmm-mmm~==~~ 3739ffimNmm=mmm5==•-m~1m~w 26 22 119 197 416 331 281 215 140 218 168 222 130 178 142 137 186 194 207· 191 uum=mwmm•mm1~rnmww~mm• 44~1urn~mrumrum=~a1143mm~~m~
79. 45a4 93 5939 66 5a69 32 12766 56 5649
53 24 37 43 42
8069 5440 3844 5280 5246
' 18 11 29 30 18 36 63 92 71 133 120 79 77 83 4a 54 180 159 125 131 94 110 51 42 25 ·1851 ---- --- ---- ---- ---- ---- ---- ---- --- ___ ..:. --- ---- ~-- ---- --- ---- ---- -- -- --- --- --- ---- ----
48416346302174923457455526706597880994379242391082712806787460447691037663592294935687899975439796138546276559887?0721031709837
' 23
J
]
J J
J J J J J
J J J J
Appendix Table 4.
1982 SOUTH BANK OF NUSHAGAK RIVER SIDE SCAN SONAR COUNTS BY HOUR
H 0 U R DATE 1 2 3 5 6 7 B 9 10 11 12 13 14 15 16 17 !B 19 20 21 22 23 24 TOTAL
6 14 6 6 15 14 6 16 12 6 17 12 6 18 1
6 19 5 6 20 3 6 21 s 6 22 561 6 23 210
6 24 49 6 25 374 6 26 427 6 27 538 6 28 598
6 29 152 6 30 107 7 1 430 7 2 751 7 3 773
7 4 318 7 5 239 7 6 252 7 7 128 7 a 157
7 9 127 7 10 103 7 11 124 7 12 144 i 13 167
7 14 7 15 7 16 7 17 7 18
167 us 235
60 12
7 19 266 7 20 519 7 21 814 7 22 139 7 23 121
7 24 .172 7 25 80 7 26 498 7 27 520 7 28 286
3 2 12 7 1 7 3 10 0 0
3 3 1
393 117
0 0 0
213 101
1 B 7
5 B 0 4 9 20 .23 13
416918 915229
0 16 0 1 5
2 26 15 12 8 10
2 2 6 14 9 7 9 1 1 0 3 1 3 3 8 7 14 0 0 1 3 2 • 2 3 5 4 0 0 9 2 7 1 5 2
7 2 ' 4 7 9
11 5 6 298 617 749 94 400 511
0 5 11, 7 0 3 5 4 8 9 30 9
903 1076 1246 632 220 140 319 232
B 2
76
~· m
6 3 5 3 2 2
133 248 408 813 1261 722 292 225 265
0 1
m 902 1<5
,. 0
511 ro 153
• 2 3 5 6 1 3 0 1 17
6 17
<21 211 110
2 2
407 60< 123
• 9 2 2 B
1 0
526
"' 233
1 16 B 0 3
7 7
526 <06 180
7 6 2 2
' • B
610 537
95
0 11 12 3 8 10 2 1 1 3 ,_ 2
1 5 5
3 0
567 <86 239
5 2
"' 533 3<0
0 3
957 377 156
~~oo~mmmu1~m141•mm~rum~mM~mwm m~-~~-~~mmm~G-m•mlli~mm-mwrn=,•m~~~~mm~mmmillmmmmmw mrn==rnmm~rnmm~m~~w~mmwrn-= 299 421 262 198 382 252 201 213 236 119 6 141 22 26 16 83 94 225 182 289 646 333 188
m~millW!rn--~m~~m~wmmmmmrnmm '''"~~m~u!•mmmm~m=•~mmrn~m~ mmromm~~=-~~~mrn~m=-m=~-~~~m 651 1951 329 671 899 1115 2168 2246 2557 2205 2045 3239 3004 1851 1470 1415 1859 1377 1041 968 11!57 946 137 m~~m•ro~mro!rummmmmm-916mm-mm
m~m~mm~m~mG~ill-mlmmill--lm~~ 258 330 202 569 415 523 1130 1025 1128 4668 3378 1938 2675 936 1004 854 849 1090 729 807 328 916 438 amm~lamrnrnm~mrnmrnmmm<1Smm~~91
rn1U371~B~mm-m219mm~mmmmrum~=m 113~nmmmmmm~m3u~mwmm~wrumm113
106 184 65 136 189 89 66 100 87 70 56 144
122 110 143 187 124 99 55 81
251 277 199 168 333 185 136 172
96 122
120 376 360 185 458 113 158 128 119 362 228 228 119 143 145 255 357 262 227 181 192 99 87 101 215 229 205 101 92 114 121 276 192 203 142
128 236 2347 167 90 122 119 195
148 69 52 63 94 97 90 150 86 86 160 206
72
" 78 29 51
~
39 38 31 17
19 150 156 128 135 153 105 213 214 134 249 141 161 71 266 253 194 173 85 254 244 404 345 354 502 192 35 121 129 93 115 58 151 146 127 233 237 372 254 74 270 476 335 163 382 253 410 233 435 595 129 822 34 389 361 245 165 193 354 246 361 458 580 353 173 - -- ·-··· . --------·------------------------------·-
140 226 112 <B2 3<3
!59 2<9
" 511 157
162 144 184 141 71 108
142 336 216 212
142 130 115 86 122 243
103 110 112 346 195 153
99 316 227
115 336 367
"' 15<
98 141 369 815 284 7-48 146 334 96 102
193 177
"' 307 66
155 m
75 555 336
64 99
129 116 106 66
201 163 161 .. ,
nooM~~rnm~rnrnmmw~m~mwrnnwm~ ~115131•mwwmm~rn~-~•rnmmm~~mlrn m~w~rul<!mmm~m•~mmm-mm~mm~ 288 587 1646 1723 3316 4828 1153 1026 3424 3735 2332 1091 1184 1440 1752 2185 2128 1500 547 1227 1804 1072 272 131 158 85 461 1137 2075 1109 461 574 1333 1302 1641 1916 1459 937 908 2733 3258 2973 1076 2196 644 252
109 139 81 49
427 109 194 133
as 185
155 489 1123 2729 5368 1541 2406 3723 1838 2148 1294 1082 2926 1642 1143 1286 1442 825 399 403 193 %~~=~-m-~-•m=mlrn=wmm~~
125 565 1393 3470 2931 3861 2170 948 1513 2111 1541 4046 3482 1640 1683 2634 3070 2488 2033 1245 771 rnm~m6=-~=~~=--~~~3113~~~= 133 567 1873 2714 2591 4099 5976 5112 4680 4292 5300 4043 2248 2560 1358 1998 1999 2675 1125 1693 394
162 267 104
91 89
107 87
6939 15134 5071
- 4601 9024 7931
11852 5432
8650 10806 21252 36052 13760
13810 26429 13256 6565 7133
6551 3642 3583 3770 3598
3326 6360 4698 7650 5687
6491 14057 12782 40399 28940
34575 20338 44754 66077 57986
729 7 30
-, "31 B 1 B 2
241 103 75 152 91 48
-tea -19a- -9o-
61 486 1350 1526 2441 5090 3921 3113 1968 2282 670 1675 1851 572 768 807 810 312 579 66 97 30864 39 1314 2600 107 52 184 1146 2200 3239 4381 5657 2418 1782 2386 1105 1043 588 254 425 270 762 32243 89 ·1345-1920 3137 2109 ·1149 ·t649 1466 ·1302_··1754· 2172 4507-5526· 3368 ·3128 2753-1324· 1137- 457· ·140 - 58 · 40866-45 359 2987 4203 4233 2996 2198 1711 910 1102 1269 1321 1811 1611 2096 1553 737 952 569 432 242 33591 d•-==~m~~~-mmm~w~mw~~~
65 121 68 173 87 50
a 3 67 e .., 151 a s 133 8 6 118 B 7 105
B B B 9 B 10 B 11 B 12
B 13 B 14 B 15 816 B 17
B !B
.. 82 50
lOB 27
32 25
6 9
21
"
35 17 90 68 88 195 70 104 74 33
35 21 30 41 36 35
124 51 24 22
13 . 15 12 29 14 34 12 19 7 ,.
3 6
46 127 71 205
193 366 115 265 71 226
534 307
"' 392 270
20
•• 26 39 28
13 27 2< 10 71
•
76 189 sa 157 26 94 59 236 26 133.
22 97 10 94 23 149 19 112 16 72
77 46
796 2130 274 432 376 195 322 251 533 743
920 428 382 156 313 155 415 410 438 267
.,. 106 79 ...
233
132 171
•• 304 141
"' .,. 210 277 155
302 219 192 215
"' 28
400 175 72 . 169 64 606 223. 112 158 76 331 359 191. 474 209 180 396 248 448 600 179 417 166 319 383
167 30 145 63 188 140 t3o · 15S. 15~ 121 178 68 69 57 108 119 148 168 157 193 87 .65 263 356 . 466
85 101 66 36 20
139 368 120 197 60 892 91 62
217 453
222 79 so
1<5 20
67 75 148 104 106 102 229 318 57 326 1214 7S1 352 291 . 312
225 186 276 273 222 134 229 166 150 271 199 229 155 173 174
11 12 26
123 101 139 89 114 680 197 329 48 107
191 295 94 159
154 97 569. 549 218 217
433 540 116 67 328 299 438 161 70 109
16 31
" 7 48 44 41 843
482 1238 157 2.28
329 552 339 691 99
135 89
298 <95 76
11
185 <90 722 463 66
132 57 ,.,
190 •• 15
60 35 35 <16 74 228
258 98 176 94
128 187 340 2<0 100
5< 48
100 240
60
22
140 157 660 295
" 66
146 ., 131
10
51
22 ., ,., 106 169
, .. Bl
891 203 27
158 138 131 126 56
.,
78 75 99 134 94 112 77 80 34 54
62 38 36 30
432 \ 221 166 25 30 20
83 31 S1 33 66 30 17 11 52 19
51 23
6927 3490 5966 6577 4891
3561 4133 6501 8288 3645
3406 2524 2900 3689 2627
802
12614 8493 9496 75751982139893501474953547352504845299042572438374723342226443493925238110389983305732703295522713021746 829165
• 24
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]
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[J
AH;:endix 'll:lble 5 • N.Jshagak River s:xXeye salmn escapement age, length am sex stmraries, 1SS2.
1'GE GlUJP
42 52 53 62 63
MIILES
Percent 4.80 48.80 o.oo 2.10 6.40
Av Lergth 444.67 597.83 o.oo 621.50 600.56
std Error 8.43 1.17 0.00 7.15 3.04
Sanp Size 27 272 0 12 36
FEM\LES
Percent .80 31.90 .10 1.60 3.50
Av Lergth 469.60 562.23 537.00 591.44 570.40
std Error 26.ll 1.38 o.oo 8.67 3.57
Sanp Size 5 177 1 9 20
SEXES a:M3INED
Percent 5.60 80.70 .10 3.70 9.90
Av Lergth 448.23 583.76 537.00 608.50 589.90
std Error ll.l9 1.25 o.oo 7.80 3.23
Sanp Size 32 449 1 21 56
-25-'
'lbtal
62.10
587.07
2.13
347
37.90
562.20
2.47
212
100.00
577.64
2.26
559
J J J
Appendix Table 6. Nusha~ak n J r1ALES
c, ERCEIIT
JV LENGTH TD ERROR
SAMP SIZE
]EHALES
lERCENT
l.Jv LENGTH STD ERROR
DAHP SIZE
SEXES COMBINED
JERCEIIT
c;t! I.EIIGTH 'lTD ERROR :JAHP SEE
J j
']
0 J J J J
31
.40
566.00 25.24
3
.30
531.00 27.00
2
.70
~51 • 00 25.94
5
river esc~peMent of chuM
AGE 41
20.40
603.09 1. 98 134
20.10
568.40 2.20
131
40.50
585.87 2.09
265
GROUP 51
36.80
613.17 1.95 240
18. 10
581.80 2.43
119
54.90
602.83 2.11 ~eo .::; ,J !
-26-
sa.l11on, a·3e and len'3th
61
3.00
638.85 6.01
20
.90
592.00 11.70
6
3,90
628.04 7.33
26
TOTAL
.;o. 60
610.74 2.34 397
39.40
574.81 2.72
258
100.00
596.58 2.49
65'S
(MM) by Se):, 1982