Physiological Responses of Eggplant (Solanum Melongena l.) in Salt Stress

8/19/2019 Physiological Responses of Eggplant (Solanum Melongena l.) in Salt Stress

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VN[ZKACAMKJBC \GZVAIZGZ AE GMMVCBIT ( Zacbiuo ogcaimgib C.) KI ZBCT

ZT\GZZ

Ly8

Ounboobd Kliu Ibzbr L!"#!$##%

Ibdyb&u' Nbibn L!"#!$##

Mblr*''b B+nbr T, L!"#!$##-J'.&&*r 8 KK

Mr.u/ 8 6

B+++&bi& 8 Bznbr Eb&ur.nobi Bldi

V\BJTKJBC \GVA\T AE VCBIT VN[ZKACAM[ K

OKIKZT\[ AE \GZGB\JN2 TGJNIACAM[ BI< NKMNG\ G


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6#!%

K, KIT\A;r.uid

Gmmpcbit (Zacbiuo ogcaimgib C.) ks aig jaooadkty tnbt ks jaooai bid papucbr

vgmgtblcgs mrawi ki tng trapkjbc bid sultrapkjbc rgmkais, gspgjkbccy ki Bskb bid

Berkjb bid nbs b prgtty maad tacgrbijg ta lkatkj bid blkatkj. Gmmpcbit ks b spgjkgs

wktn oarpnacamkjbc bid pnyskacamkjbc vbrkblkckty ae jnbrbjtgrs wkdg, sa kt ks

igjgssbry ta mgt tng saurjgs ae nkmngr rgskstbijg ae wkcd spgjkgs (Zuibitbrb,


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"ng jaijgitrbtkai ae dkssacvgd sbcts brg 7uktg nkmn ki pcbits bid sbckig sakcs wkcc

jbusg sbct strgss ki pcbits. "ng cgvgc ae strgss g&pgrkgijgd ly tng pcbits ks dkeegrgit ki

vbrkaus spgjkgs wktn tacgrbijg ks iat tng sbog ear dkeegrgit sbct jaijgitrbtkais. Geegjt

ae sbct strgss dug ta sbckikty dkd iat snaw b rgspaisg ki tng earo ae dkrgjt dbobmg lut

mrawtn dgprgssgd ly jnbimgs scawcy. 'i tng ebjg ae tng kiecugijg ae sbckikty, vbrkaus

jraps pgrearo vbrkaus earos ae bdbptbtkai ta tng ogjnbiksos ae oarpnacamkjbc bid

pnyskacamkjbc ogjnbiksos. Bdbptbtkai ta sbckikty ks iggdgd prkobrkcy ta kopravg

wbtgr lbcbijg ki ardgr ta obkitbki tng wbtgr patgitkbc bid turmar, bid bcc tng

lkajngokjbc prajgssgs ear mrawtn bid iarobc bjtkvktkgs (Zkpbyuim, gmgtblcg jrapsbrg vgry kopartbit tngy nbvg nkmn iutrktkaibc vbcug wnkjn rkjn

saurjg ae okigrbcs bid vktbokis, lut oast ae tng vgmgtblcg jraps brg #iawi bs

mcyjapnytb pcbit, tngy nbvg nkmn sgisktkvkty ta sbct strgss. "nus, #iawcgdmg ae tng

ogjnbikso ae sbct tacgrbijg ki vgmgtblcg jraps ks igjgssbry sa bs ta kijrgbsg tngkr

sustbkiblcg pradujtkai ai sbckig sakcs. Gmmpcbit (Zacbiuo ogcaimgib C.) sa-jbccgd

tgraim ki 'idaigskb ks wkdgcy rgpartgd ta lg sgisktkvg ta sbckikty (Znbnggi,


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KK, CKTG\BT:\G \G7KG5

Bjjardkim ta ktrasupaoa (?), Gmmpcbit (Zacbiuo ogcaimgib) brg

jcbsskekgd bs eaccaws1

Fkimdao 1 2cbitbg (2cbits)

Zul #kimdao 1 "rbjngalkaitb (vbsjucbr pcbits)

Zupgr 9kvkskai 1 Zpgrobtapnytb (sggd 2radujg)

9kvkskai 1 :bmiackapnytb (ecawgrkim pcbits)

cbss 1 :bmiackapskdb (dbsngd twa dk#atkc)

Zul cbss 1 Bstgrkdbg

/rdgr 1 sacbibcgs;bokcy 1 Zacbibjgbg (gtnikj gmmpcbits)

Lgius 1 Zacbiuo

Zpgjkgs 1 Zacbiuo ogcaimgib (gmmpcbit)

"ng jaopasktkai ae tng pcbit lady (oarpnacamy) gmmpcbit jaisksts ae raats,

stgos, cgbvgs, ecawgrs, erukts, bid sggds. Gmmpcbit ks b ibtkvg pcbit ae 'idkb bid Zrk

Cbi#b, bs wgcc bs jcasgcy rgcbtgd ta taobtags bid patbtags. "ng erukt ks jaooaicy

usgd bs b vgmgtblcg ear jaa#kim. !atbikjbc ibog Zacbiuo ogcaimgib. Gmmpcbit ks b

mrggi pcbits brg aetgi mrawi ai bi biiubc lbsks. "ng bulgrmkig pcbits mraw ta 8-

* jo (-*4 kijngs) ki ngkmnt. "ng cgbvgs brg cbrmg, wktn b raumn calgs. "ng trui#

ks usubccy spk#gd. ;cawgr jacar lgtwggi wnktg ta purpcg, wktn b jrawi wnkjn nbs ekvg

calgs. Jgccaw stbogis. ;rukt jaitbkis ecaur, wktn b dkbogtgr ae cgss tnbi ? jo ki tng

wkcd, bid gvgi oarg ta tng pcbitgd spgjkgs (!bsrb, 8).

2cbit mrawtn jbi lg kinklktgd ly vbrkaus #kids ae strgss boaim ckmnt, wbtgr, bid

sbct. Ztrgss ckmnt jbrrkgd tnraumn bu&ki bid geegjts brkskim erao rgdujgd geegjtkvgigss

ae bu&ki ki tng lcb0kim ckmnt jaidktkais. ;ar g&bopcg, pcbits tnbt mraw ki tng dbr# ar

wgb# ckmnt wkcc nbvg b caim rad wktn b sgmogit ae b caimgr bid cbrmgr tnbi tng pcbits

tnbt rgjgkvgd lrkmnt ckmnt. Dbtgr strgss ks b jaidktkai wngrg tng pcbits g&pgrkgijg

strgss dug ta cbj# ae wbtgr, bid kt jbi kinklkt tng ogtblackj prajgssgs ki tng lady ae

tng pcbit. Zbct strgss ks b jaidktkai wngrg tng pcbits g&pgrkgijg strgss dug ta g&jgss

cgvgcs ae sbct, bid kt jbi kinklkt tng ogtblackj prajgssgs ki tng lady ae tng pcbit. Zbct

strgss ajjurs wktn tng prgsgijg ae sbckikty ar tng jaijgitrbtkai ae dkssacvgd sbcts

g&jgsskvg. "nks mgigrbccy ajjurs sbct strgss ki pcbits wktn sakc sbckig wbtgr jaidktkais.


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Cbid wktn nkmn sbct cgvgcs wkcc kinklkt saog ae tng bjtkvktkgs tnbt brg gssgitkbc ear

rgspkrbtkai bid pnatasyitngsks (br3bdk 5 Jbnyb, ).

Zbckikty jbi beegjt kinklkt pcbit mrawtn ki twa wbys1 b. !y dbobmkim tng jgccs

mrawkim sa dksruptgd jrap mrawtn. !y ckoktkim tng boauit ae suppcy ae gssgitkbc

ogtblackj autjaogs ear jgcc mrawtn tnraumn tng earobtkai ae tycasgs. Zbckikty

supprgss tng mrawtn prajgssgs ki pcbits ly kinklktkim tng geegjt ae gicbrmgogit bid

jgcc dkvkskai, pratgki pradujtkai bid kijrgbsg pcbit lkaobss. 2cbits tnbt brg strgssgd

sbcts mgigrbccy da iat snaw b rgspaisg ki tng earo ae dkrgjt dbobmg lut suldugd

mrawtn bid jnbimg scawcy (:gimgc 5 Fkr#ly, 4).

Zbcts tnbt jbusg pcbit strgss boaim atngrs brg +bc, +bZ/ 8, bcB) ks b jaccgjtkai ae stbtkstkjbc oadgcs usgd ta

bibcy0g tng dkeegrgijgs boaim mraup ogbis bid tngkr bssajkbtgd prajgdurgs (sujn bs


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%vbrkbtkai% boaim bid lgtwggi mraups), dgvgcapgd ly stbtkstkjkbi bid gvacutkaibry

lkacamkst =aibcd ;ksngr. 'i tng B+/>B sgttkim, tng alsgrvgd vbrkbijg ki b pbrtkjucbr

vbrkblcg ks pbrtktkaigd kita jaopaigits bttrklutblcg ta dkeegrgit saurjgs ae vbrkbtkai.'t

ks usgd ta bibcy0g tng vbrkblkckty ae tng dbtb kita twa saurjgs ae vbrkbtkai tnbt ks b

vbrkbtkai ki tng mraup (wktnki) bid tng vbrkbtkai lgtwggi mraups (lgtwggi)

(9wkd3asgputra, 8).


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KKK, OBTG\KBCZ BI< OGTNA


9/26

! H Z7ubrg pbpgr wgkmnt (mrbo)

H Cgbe pbttgri wgkmnt (mrbo)

3,3 Zb'& *?='u+.i /.i&+

2grearo ly cks#gd tng cgbe ta tgstkim tng cgbe ecbvaur

3,$ 5*& 9*;n&

Zgpbrbtgd tng ogdkuo ar sakc erao tng pcbit gspsgjkbccy erao tng

raats. cgbigd tng sakc tnbt stkcc bttbjnt ki raats wktn wbtgr ecaw uitkc

jcgbi.

ut tng pcbit pbrt, raats, stgo, bid cgbvgs.

:gbsurgd tng wgt wgkmnt ae gbjn pbrt, bid iatgd tng altbkigd dbtb.

3,%


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B, \*+u'&

Tbl'* !, BIA7B N*;n& . V'bi& !

BIA7B

Zaurjgs ae >brkbtkai

9l IF F" ;nktuim ;"blcg.* .

2grcb#ubi 8


11/26

Tbl'* %, *y1+ NZ< (n.i*+& +;i=bi& d*r*i=*) &*+& /'bi& '*i;&n K7

Tu>*yF+ NZ< (n.i*+&

+;i=bi& d*r*i=*) &*+&


12/26

Mrb/n= !, \b&. Ki=r*b+* i '*i;&n . Zacbiuo ogcaimgib Jbu+*+ . Zb'& +&r*++

< ? 8 *

*

*


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Mrb/n= 6,


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15/26

Mrb/n= 3, Bib'y++ . 5*&


16/26


17/26

Mrb/n= $, Jn'.r./ny'' =.i&*i& i Zacbiuo ogcaimgib Jbu+*+ ly Zb'& Z&r*++


18/26

L,


19/26

erao tngsg bctgrgd cbid usg bid obibmgogit prbjtkjgs ks rgegrrgd ta bs $sgjaidbry

sbckiktyV (boplgcc,


20/26

l Dbtgr (ecaad ar draumnt)

Dbtgr ks aig ae tng #gy jaopaigits ae ckeg, bid pcbits nbvg gvacvgd

twa ob3ar strbtgmkgs ear wbtgr gjaiaoy1 naoakanydry bid

pak#kcanydry. "ng sjkgitkst dgekig naoakanydry bs strkvkim ta obkitbki

b nkmn wbtgr patgitkbc uidgr wbtgr-ckoktkim jaidktkais bid

pak#kcanydry bs tng kiblkckty ta jaitrac wbtgr cass ta tng givkraiogit.

j Ckmnts

2cbit rgspaisgs ta ckmnt vbry erao aig spgjkgs ta biatngr spgjkgs. "ngrg

brg pcbits rgskstbit (blcg ta mraw) ta ckoktgd ckmnt jaidktkais ar aetgi

jbccgd tacgrbit pcbits bid tngrg brg pcbits tnbt brg iat blcg ta mraw

uidgr jaidktkais ae ckoktgd ckmnt ar kitacgrbit pcbits.jngokjbc (sbct,

mbs, bid pgstkjkdgs)d Zbckikty

Zbct strgss ajjurs wktn tng prgsgijg ae sbckikty ar tng jaijgitrbtkai ae

dkssacvgd sbcts ki tng pcbit rgduidbit. "nks mgigrbccy ajjurs ki pcbits

tnbt mraw ai sbckig sakcs. Zbct strgss kijrgbsgs wktn kijrgbskim sbct

jaijgitrbtkais up ta b jgrtbki cgvgc ae jaijgitrbtkai wnkjn oby rgsuct

ki dgbtn ae tng pcbit. Zbcts tnbt jbusg pcbit strgss brg +bc, +bZ/ 8,

bc


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2nyskacamkjbc rgspaisgs brg dkvkdgd kita tnrgg ogjnbiksos1

b /soargmucbtkai

"acgrbit pcbits jbi ob#g bd3ustogits ly cawgrkim asoatkj patgitkbc

wktnaut cass ae turmar. "a altbki wbtgr erao tng surrauidkim sakc, &ycgo wbtgr

patgitkbc ki tng ck7ukd ta lg g&trgogcy cawgrgd ly tng vactbmg. Zaog nbcapnytb

blcg ta obkitbki asoatkj patgitkbc ta rgobki gvgi lgjaog oarg igmbtkvg durkim

ng mrawkim sgbsai ki ckig wktn tng blsarptkai ae sbct. /i tng atngr nbcapnytb

nbs tng blkckty ta rgmucbtg bjjuoucbtkai ae sbct (+b R bid c) ai tng jaidktkais

ae sbckikty, sujn bs obimravgs brg jbpblcg ae rgoavkim N sbct (2rknbstbitk,


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Cbrmgcy, tng asoatkj geegjt ks jbuskim tng rbtg bt wnkjn mrawkim cgbvgs g&pbid ks

rgdujgd, igw cgbvgs gogrmg oarg scawcy, bid cbtgrbc luds dgvgcap oarg scawcy ar

rgobki 7ukgsjgit, sa egwgr lrbijngs ar cbtgrbc snaats earo. "ng rgdujtkai ki snaat

mrawtn ajjurs ki twa pnbsgs, b rbpkd rgspaisg ta tng kijrgbsg ki g&tgribc asoatkj

prgssurg, bid b scawgr rgspaisg dug ta tng bjjuoucbtkai ae +bR ki cgbvgs (:uiis 5

"gstgr, brkaus cgvgcs (,


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jncarapnycc b bid l) bid rbtg ae pnatasyitngsks ki oast pcbits sul3gjtgd ta sbckikty

strgss. 't ks wgcc gvkdgit tnbt staobtbc rgmucbtkai ks bdvgrsgcy beegjtgd ki oast pcbits

sul3gjtgd ta sbckikty strgss. 'i oast jbsgs, staobtb brg pbrtkbccy jcasgd wnkjn ks bi

bdbptkvg strbtgmy ae pcbits ta jaisgrvg wbtgr uidgr sbct-kidujgd asoatkj strgss. "nus,

ly ogbsurkim mbs g&jnbimg jnbrbjtgrkstkjs kt ks passklcg ta kdgitkey bt wnbt rbtg /<

ks gitgrkim tng cgbe ta beegjt avgrbcc rbtg ae pnatasyitngsks ki pcbits g&pasgd ta sbct

strgss. 'i tng prgsgit study, dkeegrgit sbckig rgmkogs skmikekjbitcy dgjrgbsgd tng

pnatasyitngtkj rbtg, wbtgr-usg geekjkgijy bid staobtbc jaidujtbijg ae gmmpcbit pcbits

(Znbnggi, gt bc.,


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7, JAIJC:ZKAIZ BI< Z:MMGZTKAIZ

B, J.i='u+.i+

. 2cbit mrawtn bid dgvgcapogit brg kiecugijgd ly sgvgrbc ebjtars,

g&tgribccy ck#g wbtgr jaijgitrbtkai, iutrktkai, sbckikty, ckmnt, ngrlkjkdgs,

bid dksgbsgs, bid kitgribccy ck#g tng mgig tnbt tng pcbits nbs.


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B#kijk 'G, B#kijk Z, Jkcob0 F, 9k#kjk . . Zkcvb, 2..B. db astb

bid G. Laogs-;kcna.


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+kcsgi, G."., /rjutt, 9. :.. . V&y"kacamkjbc ae pcbit" uidgr "trg""6 B-katkj

ebjtar . +gy Jar#1 Iani Dkcgy bid Zais.

+arggi , Bsnrbe :.

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Physiological Responses of Eggplant (Solanum Melongena l.) in Salt Stress