Reza Hakimimofrad- fish populations Abundance Estimates

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    Estimating Abundance

    Reading: Chapter 10

    Survey design Visual censuses

    Acoustic methods

    Trawl surveys

    Depletion estimates

    Mark-recapture estimates

    Egg Production Methods

    Fishery-dependent CPUE

    Estimating Abundance

    Why do we need to estimate abundance?

    To estimate:

    1. Stock size

    2. Recruitment

    3. Mortality

    4. Spatial distribution

    Estimating Abundance

    Survey design

    A central problem is obtaining an abundanceindex that is proportional to stock size

    Well-designed survey should provide estimates of: average fish abundance or density and

    Spatial distribution (survey boundaries?)

    Accuracy vs. Precision

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    Accuracy Precision

    Estimating Abundance

    Survey design

    A central problem is obtaining an abundanceindex that is proportional to stock size

    Well-designed survey should provide estimates of:

    average fish abundance or density and

    Spatial distribution (survey boundaries?)

    Accuracy vs. Precision

    Bias vs. Variance

    precision ( error) = $

    Sample size (n)

    5 10 15 20 25 30 35

    SampleE

    rror(%)

    40

    50

    60

    70

    80

    90

    100

    110

    Sample error vs. sample size

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    Estimating Abundance

    Survey design

    Stratification by habitat type or depth Combine abundance estimates across strata

    Increases precision

    Systematic vs. Random sampling

    Systematic can be more precise and generallyreduces costs

    Estimating Abundance

    Visual censuses

    Require clear, shallow waters

    Best with non-cryptic fish that dont avoid divers

    Can see fish and habitat Transects most common

    Point counts (timed or instantaneous)

    Behavior

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    Estimating Abundance

    Acoustics

    Use of sound waves to detect fish (swim bladder)

    Best for pelagic fishes

    Target strength is species-specific and must bedetermined experimentally

    Simultaneous trawling to ground-truth catch

    Problems with acoustic shadows and avoidance

    Very promising for well understood pelagic stocks

    Estimating Abundance

    Depletion (or Removal) estimates

    Relation between abundance and catch rate

    Requires:

    Closed population

    Short fishing period (no recruitment) Catchability proportional to abundance

    CPUE (C/f) = qNtNt = N0 KtCPUE (C/f) = qN0 qKt

    Plot CPUE vs. cumulative catch (K) (known as Leslie method)

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    Kt

    CPUE

    Leslie Method

    Estimate of N0Slope = -q

    Consecutive sweeps with 100ft. seine (Fall 2003)

    1st haul 2nd haul 3rd haul

    Numbercaptured

    0

    5

    10

    15

    20

    25

    30

    35

    40

    45

    Pinfish

    Cumulative Catch

    40 50 60 70 80 90 100

    CPUE

    0

    10

    20

    30

    40

    50

    Mullet

    Cumulative Catch

    20 25 30 35 40

    CPUE

    0

    5

    10

    15

    20

    25

    Spot

    Cumulative Catch

    5 10 15 20 25

    CPUE

    0

    2

    4

    6

    8

    10

    12

    14

    Shrimp

    Cumulative Catch

    5 10 15 20 25

    CPUE

    0

    2

    4

    6

    8

    10

    Depletion estimates of abundance (Fall 2003)

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    60ft seine pulled inside 100ft seine (Fall 2003)

    1st haul 2nd haul 3rd haul 4th haul

    Numbercapture

    d

    0

    5

    10

    15

    20

    25

    Mullet

    Spot

    Pinfish

    Blue crab

    Fort Fisher Field trip 2004Depletion estimation using 100ft. seine

    Seine haul

    1st haul 2nd haul 3rd haul

    Numberscaptured

    0

    20

    40

    60

    80

    100

    120

    140

    Pinfish

    Mojarra

    Atl silverside

    Ladyfish

    Total fish

    Pinfish

    K70 80 90 100 110 120 130

    CPUE

    0

    20

    40

    60

    80

    100

    All species

    K75 100 125 150 175 200 225 250 275 300

    CPUE

    0

    20

    40

    60

    80

    100

    120

    140

    160

    180

    200

    Mojarra

    K0 50 100 150 200

    CPUE

    0

    10

    20

    30

    40

    Atl. silverside

    K0 10 20 30 40 50

    CPUE

    0

    5

    10

    15

    20

    Depletion estimates of abundance (Fall 2004)

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    Pinfish

    Cumulative catch (K)

    500 520 540 560 580

    CPUE(#perhaul)

    -100

    0

    100

    200

    300

    400

    500

    600

    Atl. silverside

    Cumulative catch (K)

    70 80 90 100

    CPUE(#perhaul)

    0

    10

    20

    30

    40

    50

    60

    70

    80

    Atl. croaker

    Cumulative catch (K)

    7.8 8.0 8.2 8.4 8.6 8.8 9.0 9.2 9.4

    CPUE(#perhaul)

    0

    2

    4

    6

    8

    10

    All species

    Cumulative catch (K)

    850 900 950 1000 1050

    CPUE(#perhaul)

    -200

    0

    200

    400

    600

    800

    1000

    Depletion estimates of abundance (Fall 2005)

    Total Length (mm)

    0 20 40 60 80 100 120 140

    Relativefrequency(%)

    0

    10

    20

    30

    40

    50

    60

    20 ft seine

    Total Length (mm)0 20 40 60 80 100 120 140

    Relativefrequency(%)

    0

    2

    4

    6

    8

    10

    12

    14

    16

    60 ft seine

    n = 71

    n = 210

    Size-selectivity of beach seines (Fall 2005)

    Estimating Abundance

    Depletion (or Removal) estimates DeLury Method

    CPUE (C/f) = qNtCPUE (C/f) = qN0(Nt/N0)

    ln CPUE = ln qN0 + ln (Nt/N0)

    Substitute Nt/N0 = e-qE

    ln CPUE = ln qN0qE

    Plot ln CPUE vs. cumulative effort (E)

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    Cumulative effort (E)

    lnCPUE

    DeLury Method

    slope = -q

    y-int. = ln qN0

    Estimating Abundance

    Trawl surveys

    Very widely used, most common

    Mesh size regulates fish size

    Constant catchability (q) essential; lack ofstandardization is major problem

    Consistent gear design, tow speed, duration helpto maintain q

    C = qfN

    CPUE = qD

    Stock biomass = D x area

    Estimating Abundance

    Trawl surveys

    Many factors affect catchability (q)

    Tow speed

    Depth

    Time of day Vessel noise

    Mostly, q is unknown, but..

    If q is constant, then estimated stock biomass willbe proportional to actual stock size

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    Surveys

    Annual method

    Daily method

    Estimating Abundance

    Whats wrong with using CPUE from fishery?

    It provides catch and effort data from large areasover long time scales, so why not use it?

    Often times it is used, only data available

    Landings data omits discards (bycatch, undersize)

    Catch/effort data hard to get for every boat

    CPUE (LPUE) rarely proportional to abundance

    No gear standardization

    Capture efficiency increases with time

    Fishers dont fish randomly

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    Fig. 10.15. Spatialdistribution

    of commercialtrawling effort(hours per year)in the North Sea

    Fig. 4.16. Distributionof Atlantic cod in theGulf of St. Lawrence,showing range expansionand contraction overtwenty years

    Fig. 4.17. Occurrence oflow, medium, and highcatches of Atlantic cod

    in research vesselsurveys as thefishery collapsed

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    Fig. 4.18. How the

    calculation of meancatch rate can affectthe interpretation offishery trends, examplefrom northern cod

    0

    1

    2

    3

    4

    5

    6

    7

    8

    9

    0 1000 2000 3000 4000

    Biomass (Tons)

    CatchabilityCoefficien

    Abundance

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    Hypoaggregation

    Hyperaggregation

    Abundance

    Density

    Abundance

    CatchperEffort(CPUE)

    CPUE remains high due to aggregation of fish