93

Systematic List of the Noctuoidea of Europe(Notodontidae, Nolidae, Arctiidae, Lymantriidae, Erebidae,

Micronoctuidae, and Noctuidae)

Michael FIBIGER and Hermann H. HACKER

Superfamily NOCTUOIDEA LATREILLE, 1809

Remarks to Classification

In the European List of Noctuidae (FIBIGER and HACKER, 1991) we refrained from writing a justification for subdividingthe family into subfamilies and tribes. Our understanding of the classification of the family has progressed sincethen, but only to some extent therefore several paraphyletic or even polyphyletic groupings still remain in theNoctuidae (s.l.) and in the other families dealt with here. Most of the tribes listed here are monophyletic and webelieve that most of them will stand the test of time. In groups where research is lasting, we refer to the latestknowledge available. For example the family Noctuidae has now been divided into three families, the Noctuidae,Micronoctuidae and Erebidae, these corresponding roughly to the groups previously called “quadrifid“ and “trifid“noctuids. Ongoing research, however, suggests that the arctiid clade might be derived from within the Erebidae(WELLER and MITCHELL, pers. com.). Other important results have been published in the last 13 years, some ofwhich are: KITCHING and RAWLINS (in KRISTENSEN, 1998); KITCHING and YELA (1999); SPEIDEL, FÄNGER and NAUMANN(1996); SPEIDEL and NAUMANN (1996); POOLE (1995, and his catalogue from 1989); the North American Moths ofNorth America (MONA) book series: LAFONTAINE and POOLE (1991), POOLE (1995), LAFONTAINE (1998, 2004); BECK(1996, 1999, 2000); the volumes of the book series Noctuidae Europaeae (1990-2003); and many papers inEsperiana.

Systematists agree that the Noctuoidea form a monophyletic clade, which is supported by both structural andmolecular research. Also the Notodontidae, Arctiidae, Lymantriidae, Nolidae (probably), and Noctuidae (sensustricto – the “trifids“), are believed to be monophyletic groups. However, it appears probable that the Erebidae(the “quadrifids“) are probably paraphyletic in that the arctiid/lymantriid clade is probably derived from within theErebidae. Since 1991 some of the European subfamilies of the former “Noctuidae“ have been removed andplaced in the Nolidae, which was upgraded to family status, and some tribes have changed in classificationamong the Notodontidae, Arctiidae, Lymantriidae, Erebidae, Micronoctuidae, and Noctuidae. Finally, the Arctiidaehave been shown to be a monophyletic family and are closely related to the Erebidae (MITCHELL et al., 2000;JACOBSEN and WELLER, 2001), and the tiny genus Micronoctua has been shown to belong to a new family, theMicronoctuidae, a family of many undescribed genera. The many changes in the classification within thesuperfamily resulted in the need to expand this new European list to include all of the Noctuoidea.

For the additional families included in the check list since 1991, we generally follow KITCHING and RAWLINS (1998)for the Lymantriidae, MILLER (1991) for the Notodontidae, and JACOBSEN and WELLER (2001) for the Arctiidae.Finally for the Erebidae and Noctuidae we refer to the list from 1991 with supplements (FIBIGER and HACKER,1992, 1998), KITCHING and RAWLINS (1998), FIBIGER and SKULE (2004), and especially to the accompanying paperby FIBIGER and LAFONTAINE (2005) in this volume of Esperiana. This latter paper is the result of intensive workwhere lepidopterological traditions from both sides of the Atlantic – particularly intiated and encouraged by J. D.LAFONTAINE (Ottawa) – called for the necessity to evaluate and share knowledge, so that through collaboration,we could built the foundation for this new European list from a holarctic perspective.

Esperiana Buchreihe zur Entomologie Bd. 11: 93-205 Schwanfeld, 29. Juni 2005 ISBN 3-938249-01-3

Authors‘ addresses:

Michael FIBIGER, Molbechs Allé 49, DK-4180, Sorø, Denmark E-mail: fibiger@mail1.stofanet.dkHermann H. HACKER, Kilianstraße 10, 96231 Bad Staffelstein, Germany E-mail: hermann-heinrich.hacker@t-online.de

94

Systematic order of the Families and Subfamilies of the European Noctuoidea

Name number of species page

Family NOTODONTIDAE STEPHENS, 1829

Subfamily THAUMETOPOEINAE AURIVILLIUS, 1889 5 97

Subfamily PYGAERINAE DUPONCHEL, [1845] 6 97

Subfamily NOTODONTINAE STEPHENS, 1829Tribus NOTODONTINI STEPHENS, 1829 22 97Tribus DICRANURINI DUPONCHEL, [1845] 13 98

Subfamily PHALAERINAE BUTLER, 1886 5 99

Subfamily HETEROCAMPINAE NEUMOGEN & DYAR, 1894 3 99

Family NOLIDAE BRUAND, 1846

Subfamily NOLINAE BRUAND, 1846 21 99

Subfamily CHLOEPHORINAE STAINTON, 1859Tribus CHLOEPHORINI STAINTON, 1859 2 100Tribus SARROTHRIPINI HAMPSON, 1894 7 100

Subfamily EARIADINAE HAMPSON, 1912 5 101

Family ARCTIIDAE LEACH, [1815]

Subfamily LITHOSIINAE BILLBERG, 1820Tribus LITHOSIINI BILLBERG, 1820 39 101

Subfamily SYNTOMINAE HERRICH-SCHÄFFER, [1846]Tribus SYNTOMINI HERRICH-SCHÄFFER, [1846] 10 103

Subfamily ARCTIINAE LEACH, [1815]Tribus ARCTIINI LEACH, [1815] 47 103Tribus CALLIMORPHINI WALKER, [1865] 12 106

Family LYMANTRIIDAE HAMPSON, [1893]

Subfamily LYMANTRIINAE HAMPSON, [1893] 15 106Tribus LYMANTRIINI HAMPSON, [1893]Tribus LEUCOMINI GROTE, 1895 1 107Tribus ORGYIINI WALLENGREN, 1861 14 107

Family EREBIDAE LEACH, [1815]1

Subfamily RIVULINAE GROTE, 1895 3 108

Subfamily BOLETOBIINAE GROTE, 1895 4 108

Subfamily HYPENODINAE FORBES, 1954 8 108

Subfamily ARAEOPTERONINAE FIBIGER, 20052 1 109

Subfamily EUBLEMMINAE FORBES, 1954Tribus EUBLEMMINI FORBES, 1954 38 109

Subfamily HERMINIINAE LEACH, [1815] 19 110

Subfamily HYPENINAE HERRICH-SCHÄFFER, [1851] 12 112

1 MITCHELL et al. (1997; 2000); FIBIGER & LAFONTAINE (2005).2 In: FIBIGER & LAFONTAINE, 2005, - in this volume of Esperiana.

95

Subfamily PHYTOMETRINAE HAMPSON, 1913 7 112

Subfamily AVENTIINAE TUTT, 1896 1 113

[Subfamily EREBINAE LEACH, [1815] 3

Subfamily CALPINAE BOISDUVAL, 1840Tribus CALPINI BOISDUVAL, 1840 1 113Tribus SCOLIOPTERYGINI HERRICH-SCHÄFFER, [1852]4 1 113

Subfamily CATOCALINAE BOISDUVAL, [1828]Tribus TOXOCAMPINI GUENÉE, 1852 29 113Tribus ACANTHOLIPINI FIBIGER & LAFONTAINE, 20055 1 114Tribus ARYTRURINI FIBIGER & LAFONTAINE, 20056 1 114Tribus MELIPOTINI GROTE, 1895 (= SYNEDINI FORBES, 1954)7 8 115Tribus EUCLIDIINI GUENÉE, 18528 5 115Tribus OPHIUSINI GUENÉE, 1837 18 115Tribus CATOCALINI BOISDUVAL, [1828] 29 116

Subfamily EUTELIINAE GROTE, 18829 2 117

Family MICRONOCTUINAE FIBIGER, 200510 1 117

Family NOCTUIDAE LATREILLE, 180911

Subfamily PLUSIINAE BOISDUVAL, [1828]Tribus ABROSTOLINI EICHLIN & CUNNINGHAM, 1978 5 118Tribus ARGYROGRAMMATINI EICHLIN & CUNNINGHAM, 1978 9 118Tribus PLUSIINI BOISDUVAL, [1828]

Subtribus AUTOPLUSIINA KITCHING, 1987 7 118Subtribus EUCHACIINA CHOU and LU, 1979 17 119Subtribus PLUSIINA BOISDUVAL, [1828] 21 120

Subfamily EUSTROTIINAE GROTE, 1882 9 120

Subfamily BAGISARINAE CRUMB, 1956 2 121

Subfamily ACONTIINAE GUENÉE, 184112

Tribus ACONTIINI GUENÉE, 1841 6 121Tribus ARMADINI WILTSHIRE, 1961 2 122Tribus AEDIINI BECK, 1960 2 122

Subfamily PANTHEINAE SMITH, 1898 3 122

Subfamily DILOBINAE AURIVILLIUS, 1889 1 122

Subfamily RAPHIINAE BECK, 1996 1 122

Subfamily ACRONICTINAE HEINEMANN, 1859 24 123

Subfamily METOPONIINAE HERRICH-SCHÄFFER, [1851]13 17 124

Subfamily CUCULLIINAE HERRICH-SCHÄFFER, [1850] 60 125

3 See FIBIGER & LAFONTAINE, 2005, - in this volume of Esperiana. No European species have yet been recognised yet.4 See FIBIGER & LAFONTAINE, 2005.5 See FIBIGER & LAFONTAINE, 2005, - in this volume of Esperiana6 See FIBIGER & LAFONTAINE, 2005, - in this volume of Esperiana.7 See FIBIGER & LAFONTAINE, 2005, - in this volume of Esperiana.8 See FIBIGER & LAFONTAINE, 2005, - in this volume of Esperiana.9 No members of Stictopterinae in Europe, - although Zebeeba falsalis is placed sometimes in Stictopterinae erronously.10 In FIBIGER & LAFONTAINE, 2005, - in this volume of Esperiana.11 See FIBIGER & LAFONTAINE, 2005, - in this volume of Esperiana.12 See FIBIGER & LAFONTAINE, 2005, - in this volume of Esperiana.13 See FIBIGER & LAFONTAINE, 2005, - in this volume of Esperiana.

Name number of species page

96

Subfamily ONCOCNEMINAE FORBES & FRANCLEMONT, 1954 59 126

Subfamily AMPHIPYRINAE GUENÉE, 1837 13 129

Subfamily PSAPHIDINAE GROTE, 1896Tribus PSAPHIDINI GROTE, 1896 12 130Tribus FERALIINI POOLE, 1995 1 131

Subfamily HELIOTHINAE BOISDUVAL, [1828] 23 131

Subfamily CONDICINAE POOLE, 1995 6 132

Subfamily ERIOPINAE HERRICH-SCHÄFFER, [1851] 2 132

Subfamily BRYOPHILINAE GUENÉE, 1852 29 132

Subfamily XYLENINAE GUENÉE, 183714

Tribus PSEUDEUSTROTIINI BECK, 1996 1 134Tribus PRODENIINI FORBES, 1954 3 134Tribus ELAPHRIINI BECK, 1996 2 134Tribus CARADRININI BOISDUVAL, 1840

Subtribus CARADRININA BOISDUVAL, 1840 48 134Subtribus ATHETINA FIBIGER & LAFONTAINE, 200515 5 136

Tribus COSMIINI GUENÉE, 1852 16 136Tribus DYPTERYGIINI FORBES, 1954 12 137Tribus ACTINOTIINI BECK, 1996 5 138Tribus PHLOGOPHORINI HAMPSON, 1918 8 138Tribus APAMEINI GUENÉE, 1841

Subtribus OXYTRYPIINA GOZMANY, 1970 1 139Subtribus APAMEINA GUENÉE, 1841 107 139Subtribus SESAMIINA FIBIGER & GOLDSTEIN, 2005 2 143

Tribus EPISEMINI GUENÉE, 1852 16 143Tribus XYLENINI GUENÉE, 1837

Subtribus XYLENINA GUENÉE, 1837 76 144Subtribus ANTITYPINA FORBES & FRANCLEMONT, 1954 69 148

Subfamily HADENINAE GUENÉE, 1837Tribus ORTHOSIINI GUENÉE, 1837 21 151Tribus THOLERINI BECK, 1996 4Tribus HADENINI GUENÉE, 1837 119 153Tribus LEUCANIINI GUENÉE, 1837 36 158Tribus ERIOPYGINI FIBIGER & LAFONTAINE, 200515 7 159Tribus GLOTTULINI GUENÉE, 1852 1 160

Subfamily NOCTUINAE LATREILLE, 1809Tribus AGROTINI RAMBUR, 1848

Subtribus AUSTRANDESIINA ANGULO & OLIVARES, 1990 1 160Subtribus AGROTINA RAMBUR, 1848 135 160

Tribus NOCTUINI LATREILLE, 1809Subtribus AXYLIINA FIBIGER & LAFONTAINE, 2005 3 165Subtribus NOCTUINA LATREILLE, 1809 131 165

Total 1576

Name number of species page

14 See FIBIGER & LAFONTAINE, 2005, - in this volume of Esperiana.15 See FIBIGER & LAFONTAINE, 2005, - in this volume of Esperiana

97

Family NOTODONTIDAE STEPHENS, 1829

Subfamily THAUMETOPOEINAE AURIVILLIUS, 1889

Genus Thaumetopoea HÜBNER, [1820]

processionea (LINNAEUS, 1758)solitaria (FREYER, 1838)pityocampa ([DENIS & SCHIFFERMÜLLER], 1775)pinivora (TREITSCHKE, 1834)herculeana RAMBUR, 1866

Subfamily PYGAERINAE DUPONCHEL, [1845]

Genus Pygaera OCHSENHEIMER, 1810

timon (HÜBNER, [1803])

Genus Clostera SAMOUELLE, 1819

curtula (LINNAEUS, 1758)albosigma (FITCH, 1855)16

pigra (HUFNAGEL, 1766)anachoreta ([DENIS & SCHIFFERMÜLLER], 1775)anastomosis (LINNAEUS, 1758)

Subfamily NOTODONTINAE STEPHENS, 1829

Tribus NOTODONTINI STEPHENS, 1829

Genus Notodonta OCHSENHEIMER, 1810

dromedarius (LINNAEUS, 1758)torva (HÜBNER, [1803])tritophus ([DENIS & SCHIFFERMÜLLER], 1775)ziczac (LINNAEUS, 1758)

Genus Drymonia HÜBNER, [1819]

dodonaea ([DENIS & SCHIFFERMÜLLER], 1775)ruficornis (HUFNAGEL, 1766)obliterata (ESPER, [1785])17 (= melagona BORKHAUSEN, 1790)querna ([DENIS & SCHIFFERMÜLLER], 1775)velitaris (HUFNAGEL, 1766)

Genus Pheosia HÜBNER, 1819

tremula (CLERCK, 1759)gnoma (FABRICIUS, 1776)

.16 New to Europe from the Ural Mountains (K. NUPPONEN & FIBIGER, 2002). This Eastern Palaearctic species was previously

known westward to the Sayan and Altai Mountains (SCHINTLMEISTER, 1984; K. NUPPONEN & FIBIGER, 2002).17 The lectotype of obliterata ESP. was designated by HACKER (1999).

98

Genus Paradrymonia KIRIAKOFF, 1967

vittata (STAUDINGER, 1892)

Genus Pterostoma GERMAR, 1812

palpina (CLERCK, 1759)

Genus Ptilophora STEPHENS, 1828

plumigera ([DENIS & SCHIFFERMÜLLER], 1775)

Genus Leucodonta STAUDINGER, 1892

bicoloria ([DENIS & SCHIFFERMÜLLER], 1775)

Genus Ptilodon HÜBNER, 1822

capucina (LINNAEUS, 1758)cucullina ([DENIS & SCHIFFERMÜLLER], 1775)

Genus Odontosia HÜBNER, 1819

carmelita (ESPER, [1798])18

sieversii (MÉNÉTRIÉS, 1856)

Genus Rhegmatophila STANDFUSS, 1888

alpina (BELLIER, 1881)richelloi HARTIG, 1939

Genus Gluphisia BOISDUVAL, 1829

crenata (ESPER, [1785])

Tribus DICRANURINI DUPONCHEL, [1845]

Genus Cerura SCHRANK, 1802

vinula (LINNAEUS, 1758)iberica (TEMPLADO & ORTIZ, 1966)erminea (ESPER, [1783])intermedia (TEICH, 1896)19

przewalskyi (ALPHÉRAKY, 1882)20

Genus Furcula LAMARCK, 1816

furcula (CLERCK, 1759)bicuspis (BORKHAUSEN, 1790)aeruginosa (CHRISTOPH, 1873)21

18 Year of description cf. HEPPNER (1981) and HACKER (1999).19 South Russia: Astrakhan district (SCHINTLMEISTER, 1982; ANIKIN et al., 2000a).20 South Russia: Uralsk district (SCHINTLMEISTER, 1982; ANIKIN et al., 2000a).21 South Russia: Astrakhan, Volgograd, Samara and Saratov districts (SCHINTLMEISTER, 1982; ANIKIN et al., 2000a).

99

bifida (BRAHM, 1787)interrupta (CHRISTOPH, 1867)22

Genus Euhampsonia DYAR, 1897

cristata (BUTLER, 1877)23

Genus Neoharpyia DANIEL, 1965

verbasci (FABRICIUS, 1798)

Genus Dicranura REICHENBACH, 1817

ulmi ([DENIS & SCHIFFERMÜLLER], 1775)

Subfamily PHALAERINAE BUTLER, 1886

Genus Phalera HÜBNER, 1819

bucephala (LINNAEUS, 1758)bucephalina STAUDINGER, 1901bucephaloides (OCHSENHEIMER, 1810)

Genus Peridea STEPHENS, 1828

anceps (GOEZE, 1781)korbi (REBEL, 1918)24

Subfamily HETEROCAMPINAE NEUMOGEN & DYAR, 1894

Genus Stauropus GERMAR, 1812

fagi (LINNAEUS, 1758)

Genus Harpyia OCHSENHEIMER, 1810

milhauseri (FABRICIUS, 1775)

Genus Spatalia HÜBNER, 1819

argentina ([DENIS & SCHIFFERMÜLLER], 1775)

Family NOLIDAE BRUAND, 184625

Subfamily NOLINAE BRUAND, 1846

22 South Russia: Astrakhan, Volgograd and Saratov districts (SCHINTLMEISTER, 1982; ANIKIN et al., 2000a).23 This species with Pacifiac distribution was collected once in the Samara district of S Russia: Zhiguli (ANIKIN et al., 2000a). It

is doubtful if it is resident in the western Palaearctic region.24 Romania (ssp. herculeana POPESCU-GORJ & CAPUSE, 1963) (SCHINTLMEISTER, 1982; 1989).25 The arrangement follows FIBIGER & SKULE (in press).

100

Genus Antennola de FREINA & WITT, 1984

impura (MANN, 1862)

Genus Meganola DYAR, 1898

togatulalis (HÜBNER, 1796)strigula ([DENIS & SCHIFFERMÜLLER], 1775)kolbi DANIEL, 193526

gigantula (STAUDINGER, 1879)albula ([DENIS & SCHIFFERMÜLLER], 1775)

Genus Nola LEACH, [1815]

cucullatella (LINNAEUS, 1758)tutulella ZERNY, 1927confusalis (HERRICH-SCHÄFFER, [1847])cicatricalis (TREITSCHKE, 1835)harouni WILTSHIRE, 195127

aerugula (HÜBNER, 1793)

crambiformis REBEL, 1902)27a

centonalis HÜBNER, 1796; subsp. holsatica SAUBER, 1916)28

karelica (TENGSTRÖM, 1869)dresnayi (WARNECKE, 1946)squalida STAUDINGER, 1871cristatula (HÜBNER, 1793)subchlamydula STAUDINGER, 1871chlamitulalis (HÜBNER, [1813])thymula MILLIÈRE, 1867

Subfamily CHLOEPHORINAE STAINTON, 1859

Tribus CHLOEPHORINI STAINTON, 1859

Genus Bena BILLBERG, 1820

bicolorana (FUESSLY, 1775) (=quercana [DENIS & SCHIFFERMÜLLER],1775; = prasinana auctt. nec LINNAEUS)29

Genus Pseudoips HÜBNER, 1822

prasinana (LINNAEUS, 1758) (=fagana FABRICIUS, 1781)30

Tribus SARROTHRIPINI HAMPSON, 1894

Genus Nycteola HÜBNER, 1822

revayana (SCOPOLI, 1772)columbana (TURNER, 1925)31

26 The question whether kolbi DANIEL, 1935 deserves full species or being just a synonym or a subspecies of strigula ([DENIS &SCHIFFERMÜLLER], 1775) is unsolved. We follow the opinions of BESHKOV (2000) and RAKOSY & SZEKELY (1995) and treat it atspecies rank.

27 This species occurs in the south and south-eastern islands of Greece (FIBIGER & KARSHOLT, 1998).27a Type locality of crambiformis REBEL, 1902 is S Ural, Orenburg (ANIKIN et al., 2000) (pers. comm. K. NUPPONEN).28 FIBIGER & KARSHOLT (1998).29 MIKKOLA & HONEY (1993); FIBIGER & HACKER (1998).30 Stat. rev. (MIKKOLA & HONEY, 1993): lectotype designation by LEMPKE. Therefore the Linnean name has been interpreted

incorrectly (FIBIGER & HACKER, 1998).31 Bracket.

101

degenerana (HÜBNER, [1799])asiatica (KRULIKOVSKY, 1904)siculana (FUCHS, 1899)kuldzhana OBRAZTSOV, 195332

Genus Characoma WALKER, 1863

nilotica (ROGENHOFER, 1882)33

Subfamily EARIADINAE HAMPSON, 1912

Genus Earias HÜBNER, [1825]34

clorana (LINNAEUS, 1761)albovenosana OBERTHÜR, 191735

syriacana BARTEL, 190336

vernana (FABRICIUS, 1787)insulana (BOISDUVAL, 1833)

Family ARCTIIDAE LEACH, [1815]37

Subfamily LITHOSIINAE BILLBERG, 1820

Tribus LITHOSIINI BILLBERG, 1820

Genus Nudaria HAWORTH, 1809

mundana (LINNAEUS, 1761)

Genus Thumatha WALKER, 1866

senex (HÜBNER, 1808)

Genus Paidia HÜBNER, 1819

rica (FREYER, 1858)griseola (ROTHSCHILD, 1933)cinerascens HERRICH-SCHÄFFER, 1847

Genus Apaidia HAMPSON, 1900

mesogona (GODART, 1824)rufeola (RAMBUR, 1832)

32 DUFAY (1958) stated records of this species from Russia, Sarepta. Futher specimens were taken in the Urals (K. NUPPONEN &FIBIGER, in press).

33 Pardasena virgulana (MABILLE, 1880): The statement about records from Malta is due to a misunderstanding. The referenceconcerns a specimen taken in England, most probably imported.

34 The African taxon E. biplaga WALKER, a pest on cotton, has been recorded four times in Europe: three times in England - oneof which could be a migrant (Martin HONEY, pers. comm.) - and one in Denmark, 2004, also taken on open land, and it couldbe a migrant.

35 A revision of the taxonomic status of albovenosana is needed.36 A revision of the taxonomic status of syriacana is needed.37 See FIBIGER & LAFONTAINE (2005).

102

Genus Miltochrista HÜBNER, 1819

miniata (FORSTER, 1771)

Genus Cybosia HÜBNER, 1819

mesomella (LINNAEUS, 1758)

Genus Pelosia HÜBNER, 1819

muscerda (HUFNAGEL, 1766)obtusa (HERRICH-SCHÄFFER, 1847)plumosa (MABILLE, 1900)

Genus Atolmis HÜBNER, 1819

rubricollis (LINNAEUS, 1758)

Genus Lithosia FABRICIUS, 1798

quadra (LINNAEUS, 1758)

Genus Eilema HÜBNER, 1819

depressa (ESPER, [1787])griseola (HÜBNER, 1803)lurideola (ZINCKEN, 1817)complana (LINNAEUS, 1758)pseudocomplana (DANIEL, 1938)morosina (HERRICH-SCHÄFFER, 1847)caniola (HÜBNER, 1808)palliatella (SCOPOLI, 1763)pygmaeola (DOUBLEDAY, 1847)predotae (SCHAWERDA, 1927)lutarella (LINNAEUS, 1758)muscula (STAUDINGER, 1899)sororcula (HUFNAGEL, 1766)uniola (RAMBUR, 1866)interpositella STRAND, 1920albicosta (ROGENHOFER, 1894)rungsi (TOULGOET, 1960)

Genus Zobida BIRKET-SMITH, 1965

bipuncta (HÜBNER, 1824)

Genus Setema de FREINA & WITT, 1984

cereola (HÜBNER, 1803)

Genus Setina SCHRANK, 1802

irrorella (LINNAEUS, 1758)flavicans (GEYER, 1836)alpestris ZELLER, 1865aurita (ESPER, [1787])cantabrica (de FREINA & WITT, 1985)

103

roscida ([DENIS & SCHIFFERMÜLLER],1775)

Subfamily SYNTOMINAE HERRICH-SCHÄFFER, [1846]

Genus Amata FABRICIUS, 1807

phegea (LINNAEUS, 1758)ragazzii (TURATI, 1917)kruegeri (RAGUSA, 1904)nigricornis (ALPHÉRAKY, 1883)38

caspia (STAUDINGER, 1877)39

transcaspica OBRATZSOV, 196640

turgaica OBRATZSOV, 196641

Genus Dysauxes HÜBNER, 1819

ancilla (LINNAEUS, 1758)famula (FREYER, 1836)punctata (FABRICIUS, 1781)

Subfamily ARCTIINAE LEACH, [1815]

Tribus ARCTIINI LEACH, [1815]

Genus Ocnogyna LEDERER, 1853

corsica (RAMBUR, 1832)baetica (RAMBUR, 1836)loewii (ZELLER, 1846)zoraida (de GRASLIN, 1837)parasita (HÜBNER, 1790)

Genus Artimelia RAMBUR, 1866

latreillei (GODART, 1823)

Genus Chelis RAMBUR, 1866

maculosa (GERNING, 1780)simplonica (BOISDUVAL, 1840)dahurica (BOISDUVAL, 1843)42

caecilia (KINDERMANN, 1853)42a

Genus Sibirarctia DUBATOLOV, 198743

kindermanni (STAUDINGER, 1867)44

38 S. Russia (ANIKIN et al., 2000a).39 S. Russia (ANIKIN et al., 2000a).40 Only known from the type series from South Russia: Uralsk region (ANIKIN et al., 2000a).41 Only known from the type series from South Russia: Uralsk region (ANIKIN et al., 2000a).42 S Russia: Bashkiria reserve; S. Ural (DUBATOLOV, 1988; [1994]; 1996; ANIKIN et al., 2000a).42a S. Ural: Moskovo, steppe hills 2 km to the west from the main flow of Ural River, 5.VII.2003 (leg. K. NUPPONEN) (K. NUPPONEN

pers. comm.).43 Combination cf. DUBATOLOV (1987).44 Described from S Russia: Orenburg district.

104

Genus Watsonarctia de FREINA & WITT, 1984

deserta (BARTEL, 1902) (=casta ESPER, [1785])

Genus Phragmatobia STEPHENS, 1828

fuliginosa (LINNAEUS, 1758)placida (FRIVALDSKY, 1835)luctifera ([DENIS & SCHIFFERMÜLLER],1775)

Genus Cymbalophora RAMBUR, 1866

rivularis (MÉNÉTRIÉS, 1832)pudica (ESPER, [1785])45

Genus Parasemia HÜBNER, 1820

plantaginis (LINNAEUS, 1758)

Genus Grammia RAMBUR, 1866

qvenseli (PAYKULL, 1793)45a

Genus Holoarctia FERGUSON, 1984

cervini (FALLOU, 1864)puengeleri (O. BANG-HAAS, 1927)46

Genus Spilosoma CURTIS, 1825

lutea (HUFNAGEL, 1766)lubricipeda (LINNAEUS, 1758)urticae (ESPER, [1789])

Genus Hyphantria HARRIS, 1841

cunea (DRURY, 1773)

Genus Diaphora STEPHENS, 1827

mendica (CLERCK, 1759)

Genus Cycnia HÜBNER, 1818

sordida (HÜBNER, 1803)luctuosa (GEYER, 1833)

Genus Lacydes WALKER, 1855

spectabilis (TAUSCHER, 1806)

45 Year of description cf. HEPPNER (1981) and HACKER (1999).45a Spelling: L. AARVIK, pers. comm.46 S.Russia: Ural (ANIKIN et al., 2000b); subsp. puengeleri occurs in the Urals; subsp. fridolini (TORSTENIUS, 1971) occurs in

Swedish and Norwegian Lapland.

105

Genus Rhyparia HÜBNER, 1820

purpurata (LINNAEUS, 1758)

Genus Rhyparioides BUTLER, 1877

metelkana (LEDERER, 1861)

Genus Diacrisia HÜBNER, 1819

sannio (LINNAEUS, 1758)

Genus Hyphoraia HÜBNER, 1820

aulica (LINNAEUS, 1758)testudinaria (FOURCROY, 1785)dejeani (GODART, 1822)

Genus Pararctia SOTAVALTA, 1965

lapponica (THUNBERG, 1791)tundrana (TSHISTJAKOV, 1990)47

Genus Sibirarctia DUBATOLOV, 1987

kindermanni (HAMPSON, 1901)47a

Genus Acerbia SOTAVALTA, 1965

alpina (QUENSEL, 1802)

Genus Borearctia DUBATOLOV, 1984

menetriesi (EVERSMANN, 1846)

Genus Pericallia HÜBNER, 1820

matronula (LINNAEUS, 1758)

Genus Arctia SCHRANK, 1802

tigrina (VILLERS, 1789)caja (LINNAEUS, 1758)olschwangi DUBATOLOV, 199048

flavia (FUESSLY, 1779)villica (LINNAEUS, 1758)festiva (HUFNAGEL, 1766)

47 Pararctia subnebulosa (DYAR, 1899) is a nearctic species, which was mixed with P. tundrana (see eg. TSHISTJAKOV 1990 andDUBATOLOV’s internet pages) in former times. Thus, P. subnebulosa is not an European species. P. tundrana is known from arcticregions of Russia, westward to Polar Ural (pers. comm. K. NUPPONEN). According to DUBATOLOV ([1994]; 1996) it was recordedalso from North Russia: Murmansk, Arkhangelsk regions; Komi republic.

47a The species occurs in the Ural Mountains.48Arctia olschwangi is known from Polar Ural, and the Southern part of Yamal Peninsula and Eastern Yakutia, altogether 6 exx.

(pers. comm. K. NUPPONEN).

106

Tribus CALLIMORPHINI WALKER, [1865](= UTETHEISINI FORBES, 1960; = NYCTEMERINI WALKER, [1865]) (JACOBSON & WELLER, 2001)

Genus Callimorpha LATREILLE, 1809

dominula (LINNAEUS, 1758)

Genus Euplagia HÜBNER, 1820

quadripunctaria (PODA, 1761)

Genus Tyria HÜBNER, 1819

jacobaeae (LINNAEUS, 1758)

Genus Dodia DYAR, 1901

albertae DYAR, 190149

Genus Spiris HÜBNER, 1819

striata (LINNAEUS, 1758)

Genus Coscinia HÜBNER, 1819

cribraria (LINNAEUS, 1758)bifasciata (RAMBUR, 1832)mariarosae EXPÓSITO, 1991benderi (MARTEN, 1957)romeii (SAGARRA, 1924)libyssa (PÜNGELER, 1907)

Genus Utetheisa HÜBNER, 1819

pulchella (LINNAEUS, 1758)

Family LYMANTRIIDAE HAMPSON, [1893]

Subfamily LYMANTRIINAE HAMPSON, [1893]

Tribus LYMANTRIINI HAMPSON, [1893]

Genus Lymantria HÜBNER, [1819]

monacha (LINNAEUS, 1758)dispar (LINNAEUS, 1758)atlantica (RAMBUR, 1837)lapidocola (HERRICH-SCHÄFFER, 1851)

Genus Ocneria HÜBNER, [1819]

rubea ([DENIS & SCHIFFERMÜLLER], 1775)eos REISSER, 1962ledereri (MILLIÈRE, 1868)

49 North Russia: Murmansk, Arkhangelsk regions; Komi Republic; N. Ural (DUBATOLOV, [1994]; 1996).

107

Genus Albarracina STAUDINGER, 1883

warionis (OBERTHÜR, 1881)

Genus Parocneria DYAR, 1897

detrita (ESPER, [1785])terebinthi (FREYER, 1838)

Genus Euproctis HÜBNER, [1819]

Subgenus Euproctis HÜBNER, [1819]

chrysorrhoea (LINNAEUS, 1758)

Subgenus Sphrageidus MAES, 1984

similis (FUESSLY, 1775)

Genus Penthophera GERMAR, 1812

morio (LINNAEUS, 1767)

Genus Casama WALKER, 1865

innotata (WALKER, 1855)50

Genus Arctornis GERMAR, 1810

l-nigrum (MÜLLER, 1764)

Tribus LEUCOMINI GROTE, 1895

Genus Leucoma HÜBNER, [1822]

salicis (LINNAEUS, 1758)

Tribus ORGYIINI WALLENGREN, 1861

Genus Dicallomera BUTLER, 1881

fascelina (LINNAEUS, 1758)

Genus Calliteara BUTLER, 1881

pudibunda (LINNAEUS, 1758)abietis ([DENIS & SCHIFFERMÜLLER], 1775)

Genus Laelia STEPHENS, 182851

coenosa (HÜBNER, [1808])

50 SAMMUT (2000): Malta, Benghisa, 23.X.1987 (leg. A. SEGUNA).51 This genus is often erroneously placed in the tribe Lymantriini.

108

Genus Gynaephora HÜBNER, [1822]

selenitica (ESPER, [1789])groenlandica (WOCKE, 1874) (= G. lugens KOZHANCHIKOV, 1948)51a

Genus Orgyia OCHSENHEIMER, 1810

recens (HÜBNER, [1819])antiqua (LINNAEUS, 1758)antiquiodes (HÜBNER, [1822])aurolimbata GUENÉE, 1835trigotephras BOISDUVAL, 1828corsica (BOISDUVAL, 1834)rupestris RAMBUR, 1832dubia (TAUSCHER, 1806)

Family EREBIDAE LEACH, [1815]52

Subfamily RIVULINAE GROTE, 189553

Genus Rivula GUENÉE, [1845]

tanitalis REBEL, 191253a

sericealis (SCOPOLI, 1763)

Genus Zebeeba KIRBY, 1892, comb. n.54

falsalis (HERRICH-SCHÄFFER, 1839)

Subfamily BOLETOBIINAE GROTE, 189555

Genus Parascotia HÜBNER, [1825]

detersa (STAUDINGER, 1892)nisseni TURATI, 190556

lorai AGENJO, 1967fuliginaria (LINNAEUS, 1761)

Subfamily HYPENODINAE FORBES, 195457

Genus Hypenodes DOUBLEDAY, 1850

humidalis DOUBLEDAY, 1850 (= turfosalis WOCKE, 1850)

51a G. groenlandica WOCKE, 1874) (= G. lugens KOZHANCHIKOV, 1948), was recorded in 1994 in the Polar Ural (leg. J. KULLBERG & J.

JALAVA). The taxon is probably not conspecific with Gynaephora rossii (CURTIS, 1835) (pers. comm. K. NUPPONEN).52 The systematic order and sequence of tribes, genera, and species follow mainly FIBIGER & HACKER (1991, 1992, and 1998);

KITCHING & RAWLINS (1998); FIBIGER & SKULE (2004), GOATER et al. (2003); and especially FIBIGER & LAFONTAINAE (2005), - in thisvolume of Esperiana.

53 We maintain this taxon at subfamily level because of character states in the larvae and in the genitalia of both sexes; seeFIBIGER & LAFONTAINAE (2005), - in this volume of Esperiana.

53a Recorded in Europe on Crete.54 This genus and species, formerly placed in the subfamily Stictopterinae, was transferred to the subfamily Rivulinae by

SPEIDEL et al. (1996) (see FIBIGER & LAFONTAINE, 2005, - in this volume of Esperiana).55 FIBIGER & LAFONTAINAE (2005), - in this volume of Esperiana.56 No brackets (FIBIGER & HACKER, 1998).57 See FIBIGER and LAFONTAINE, 2005.

109

anatolica SCHWINGENSCHUSS, 1938 (= nigritalis RONKAY, 1984)58

orientalis STAUDINGER, 1901 (= nesiota REBEL, 1916)59

kalchbergi STAUDINGER, 1876

Genus Schrankia HÜBNER, [1825]

costaestrigalis (STEPHENS, 1834) (= hartigi BERIO, 1991)60

intermedialis REID, 197261

taenialis (HÜBNER, [1809])balneorum (ALPHÉRAKY, 1880)62

Subfamily ARAEOPTERONINAE FIBIGER, 200563

Genus Araeopteron HAMPSON, 189364

ecphaea (HAMPSON, 1914)65

Subfamily EUBLEMMINAE FORBES, 195466

Tribus EUBLEMMINI FORBES, 1954

Genus Calymma HÜBNER, [1823]

communimacula ([DENIS & SCHIFFERMÜLLER], 1775)

Genus Odice HÜBNER, [1823]

arcuinna (HÜBNER, 1790)blandula (RAMBUR, 1858)pergrata (RAMBUR, 1858)suava (HÜBNER, [1813])jucunda (HÜBNER, [1813])

Genus Eublemma HÜBNER, [1821] (=Coccidiphaga SPULER, 1907)67

minutata (FABRICIUS, 1794) (= noctualis HÜBNER, 1796)viridula (GUENÉE, 1841)caprearum (DRAUDT, 1933)68

elychrysi (RAMBUR, 1833)

58 Synonymy cf. NOWACKI & FIBIGER (1996); FIBIGER & HACKER (1998).59 With reference to the list from Cyprus in Esperiana (FIBIGER, NILSSON & SVENDSEN, 1998).60 Synonymy cf. FIBIGER & HACKER (1998).61 This taxon is probably a hybrid, - under revision (FIBIGER, in prep.).62 Ukraine, Crimea and Ural (NUPPONEN & FIBIGER, 2002).63 The tribe is under revision. The character states of the tribe are described by FIBIGER & LAFONTAINE (2005), and in detail by

FIBGER & HACKER (2001). The tribe includes approximately 150 species distributed worldwide, most of which are undescribed(FIBIGER, in prep.).

64 New genus for the European fauna. Its exact position close to Eublemminae and Acontiinae is yet uncertain and underrevision (FIBIGER & HACKER, 2001).

65 FIBIGER & AGASSIZ (2001). Greece and Spain: Mallorca. Now also found in the mainland of Spain, and in Italy, incl.Sicily (leg.S. BELLA, M. FIBIGER, O. KARSHOLT & G. JEPPESEN; and new to France from Corsica, Sept. 2003 (leg. D. NILSSON & P. SKOU).Either the species is spreading rapidly, or it has recieved intensified attention recently.

66 Transfered from the ‘trifine‘ (Noctuidae, s.s.) to the ‘quadrifine‘ (Erebinae) (FIBIGER & LAFONTAINE, 2005); and partly revised,upgraded to subfamily rank, with autapomorphies listed in FIBIGER & HACKER (2002).

67 Synonym of Eublemma HÜBNER, [1821] (FIBIGER & HACKER, 2002).68 Bona species. (FIBIGER, in press).

110

pulchralis (VILLERS, 1789) (=candidana FABRICIUS, 1794; = rectifascia de JOANNIS, 1909)69

ochreola (STAUDINGER, 1900) (=thurneri ZERNY, 1936)70

ostrina (HÜBNER, [1808])porphyrinia (FREYER, 1845)parva (HÜBNER, [1808])debilis (CHRISTOPH, 1884)71

panonica (FREYER, 1840)amasina (EVERSMANN, 1842)rosea (HÜBNER, 1790)72

amoena (HÜBNER, [1803]) (= E. respersa sensu auct. nec [D & S.],1775)73

purpurina ([DENIS & SCHIFFERMÜLLER], 1775)pura (HÜBNER, [1813])himmighoffeni (MILLIÈRE, 1867)pallidula (HERRICH-SCHÄFFER, 1856)straminea (STAUDINGER, 1892)hansa (HERRICH-SCHÄFFER, 1851)74

gratiosa (EVERSMANN, 1854)candicans (RAMBUR, 1858)75

albida (DUPONCHEL, 1843) (=albicans GUENÉE, 1852; = gueneei SPULER, 1907)76

parallela (FREYER, 1842)pusilla (EVERSMANN, 1837) (= concinnula HERRICH-SCHÄFFER, 1851)polygramma (DUPONCHEL, [1842])ragusana (FREYER, 1844)scitula (RAMBUR, 1833) (= turatii BERIO, 2002)77

Genus Rhypagla NYE, 1975

lacernaria (HÜBNER, [1813])

Genus Metachrostis HÜBNER, [1820]

velox (HÜBNER, [1813]) (= velocissima TURATI, 1926)velocior (STAUDINGER, 1892)dardouini (BOISDUVAL, 1840)

Subfamily HERMINIINAE LEACH, [1815]

Genus Orectis LEDERER, 1857

massiliensis (MILLIÈRE, [1864]) (=euprepiata DANNEHL, 1933)78

proboscidata (HERRICH-SCHÄFFER, [1851]) (=barteli TURATI, 1907)79

69 Synonym (LERAUT, 1997).70 Synonym cf. HACKER (1999e).71 SE. Russia: one � (ZI St. Petersburg); 1 � (Zool. Mus. Novosibirsk) (FIBIGER & HACKER, 1998).72 Phalaena rosea HÜBNER, 1790 is not a homonym of Phalaena rosea sensu FOURCROY, 1785, as the latter is a misidentification

of Bombyx rosea FABRICIUS, 1775, a junior synonym of Miltochrista miniata (FORSTER, 1771) (HONEY, in litt.) (cf. KARSHOLT, in:KARSHOLT & RAZOWSKI, 1996) (FIBIGER & HACKER, 1998).

73 Misidentification, originated from HÜBNER, who himself corrected it (HONEY, in litt.) (cf. KARSHOLT, in: KARSHOLT & RAZOWSKI,1996; FIBIGER & HACKER, 1998).

74 Bona species (HACKER, 1999e; FIBIGER & HACKER, 2004).75 Name of species (cf. FIBIGER & HACKER, 1998). Rediscovered in southern Spain, after more than 100 years of virtual absence

(Penny HALE, pers. comm.).76 Synonyms (KARSHOLT, in: KARSHOLT & RAZOWSKI, 1996; FIBIGER & HACKER, 1998).77 FIBGER & HACKER (2002) syn.78 RAINERI & ZILLI (1995); FIBIGER & HACKER (1998).79 RAINERI & ZILLI (1995); FIBIGER & HACKER (1998).

111

Genus Idia HÜBNER, [1813]

calvaria ([DENIS & SCHIFFERMÜLLER], 1775)

Genus Simplicia GUENÉE, 1854

rectalis (EVERSMANN, 1842)

Genus Paracolax HÜBNER, [1825]

tristalis (FABRICIUS, 1794) (= glaucinalis auct., nec LINNAEUS, 1758, nec [DENIS & SCHIFFERMÜLLER], 1775; =derivalis HÜBNER, 1796)

Genus Nodaria GUENÉE, 1854

nodosalis (HERRICH-SCHÄFFER, [1851])

Genus Macrochilo HÜBNER, [1825]

cribrumalis (HÜBNER, 1793)

Genus Herminia LATREILLE, 1802

tarsipennalis TREITSCHKE, 1835tarsicrinalis (KNOCH, 1782)flavicrinalis ANDREAS, 1910 (= flavicrinaloides POOLE, 1989; = gigantea TURATI, 1911)grisealis ([DENIS & SCHIFFERMÜLLER], 1775)tenuialis (REBEL, 1899)

Genus Polypogon SCHRANK, 1802

tentacularia (LINNAEUS, 1758)gryphalis (HERRICH-SCHÄFFER, 1851)

Genus Pechipogo HÜBNER, 1825

strigilata (LINNAEUS, 1758) (= barbalis CLERCK, 1759)plumigeralis (HÜBNER, [1825]) (=crinalis TREITSCHKE, 1829)80

simplicicornis (ZERNY, 1935)81

Genus Zanclognatha LEDERER, 1857

lunalis (SCOPOLI, 1763)zelleralis (WOCKE, 1850)

80 YELA, HONEY & RONKAY (1997); FIBIGER & HACKER (1998).81 YELA, HONEY & RONKAY (1997); FIBIGER & HACKER (1998).

112

Subfamily HYPENINAE HERRICH-SCHÄFFER, [1851]

Genus Zekelita WALKER, 1863 (=Rhynchodontodes WARREN, 1913)82

Subgenus Rhynchodontodes WARREN, 1913

antiqualis (HÜBNER, [1809])

Subgenus Ravalita LÖDL & MAYERL, 1998

ravulalis (STAUDINGER, 1879)ravalis (HERRICH-SCHÄFFER, 1851) (= revolutalis ZELLER, 1852)83

Genus Hypena SCHRANK, 1802

Subgenus Hypena SCHRANK, 1802

proboscidalis (LINNAEUS, 1758)rostralis (LINNAEUS, 1758)obesalis TREITSCHKE, 1829obsitalis (HÜBNER, [1813])palpalis (HÜBNER, 1796) (= extensalis GUENÉE, 1854)munitalis MANN, 1861opulenta (CHRISTOPH, 1877)lividalis (HÜBNER, 1796)

Subgenus Bomolocha HÜBNER, [1825]

crassalis (FABRICIUS, 1787)

Subfamily PHYTOMETRINAE HAMPSON, 191384

Genus Phytometra HAWORTH, 1809

luna ZERNY, 1927sanctiflorentis (BOISDUVAL, 1834)viridaria (CLERCK, 1759)amata (BUTLER, 1878)85

Genus Raparna MOORE, 1882

conicephala (STAUDINGER, 1870)

Genus Colobochyla HÜBNER, [1825]

salicalis ([DENIS & SCHIFFERMÜLLER], 1775)

Genus Trisateles TAMS, 1939

emortualis ([DENIS & SCHIFFERMÜLLER], 1775)

82 LÖDL & MAYERL (1998), FIBIGER & HACKER (1998).83 LÖDL & MAYERL (1998), FIBIGER & HACKER (1998). Apart from the population of SE European Russia, this species occurs also

in Greece on Samos (leg. P. SVENDSEN), and on the Dodokanesian Islands, Leros, Kalymnos, and Rhodos (leg. FIBIGER,JEPPESEN, MADSEN & NILSSON).

84 FIBIGER & LAFONTAINAE (2005),-in this volume of Esperiana.85 This species of Eastern Palearctic distribution was surprisingly found in Slovenia (3 males). Whether or not its presence in

Europe is autochthonous cannot be decided at present (KÜHNE, 1997; FIBIGER & HACKER, 1998).

113

Subfamily AVENTIINAE TUTT, 189686

Genus Laspeyria GERMAR, 1810

flexula ([DENIS & SCHIFFERMÜLLER], 1775)

Subfamily CALPINAE BOISDUVAL, 184087

Tribus CALPINI BOISDUVAL, 1840

Genus Calyptra OCHSENHEIMER, 1816

thalictri (BORKHAUSEN, 1790)

Tribus SCOLIOPTERYGINI HERRICH-SCHÄFFER, [1852]

Genus Scoliopteryx GERMAR, 1810

libatrix (LINNAEUS, 1758)

Subfamily CATOCALINAE BOISDUVAL, [1828]88

Tribus TOXOCAMPINI GUENÉE, 1852

Genus Chrysorithrum BUTLER, 1878

flavomaculata (BREMER, 1861)

Genus Exophyla GUENÉE, 184189

rectangularis (GEYER, [1828])

Genus Anumeta WALKER, 1858

atrosignata WALKER, 1858spilota (ERSCHOV, 1874)90

henkei STAUDINGER, 1877fractistrigata ALPHÉRAKY, 1882cestis (MÉNÉTRIÉS, 1848)

Genus Lygephila BILLBERG, 1820

lusoria (LINNAEUS, 1758)lubrica (FREYER, 1846)ludicra (HÜBNER, 1790)glycyrrhizae (RAMBUR, 1866)

86 FIBIGER & LAFONTAINAE (2005),-in this volume of Esperiana.87 FIBIGER & LAFONTAINAE (2005),-in this volume of Esperiana.88 General arrangement see GOATER, RONKAY and FIBIGER, (2003); the new arrangement of the tribes is following FIBIGER &

LAFONTAINAE (2005),-in this volume of Esperiana.89 Spelling.90 Bona species (GOATER, RONKAY & FIBIGER, 2003).

114

fonti YELA & CALLE, [1990]91

pastinum (TREITSCHKE, 1826)viciae (HÜBNER, [1822])craccae ([DENIS & SCHIFFERMÜLLER], 1775) (= herrerai YELA, 1990)92

procax (HÜBNER, [1813]) (= limosa TREITSCHKE, 1826)

Genus Tathorhynchus HAMPSON, 189493

exsiccata (LEDERER, 1855)

Genus Autophila HÜBNER, [1823]

Subgenus Autophila HÜBNER, [1823]

hirsuta (STAUDINGER, 1870)libanotica (STAUDINGER, 1901)94

limbata (STAUDINGER, 1871)dilucida (HÜBNER, [1808])asiatica (STAUDINGER, 1888)rosea (STAUDINGER, 1888)95

Subgenus Cheirophanes BOURSIN, 1955

cataphanes (HÜBNER, [1813])anaphanes BOURSIN, 1940chamaephanes BOURSIN, 194096

ligaminosa (EVERSMANN, 1851)maura (STAUDINGER, 1888)97

Genus Apopestes HÜBNER, [1823]

spectrum (ESPER, [1787])

Tribus ACANTHOLIPINI FIBIGER & LAFONTAINE, 200598

Genus Acantholipes LEDERER, 1857

regularis (HÜBNER, [1813])

Tribus ARYTRURINI FIBIGER & LAFONTAINE, 200599

Genus Arytrura JOHN, 1912

musculus (MÉNÉTRIÉS, 1859)

91 Year of publication (Misc. Zool. 12: 205).92 Year of publication; herrerai YELA, 1990 is a synonym of L. craccae ([DENIS & SCHIFFERMÜLLER], 1775) (GOATER, RONKAY &

FIBIGER, 2003).93 Raised again to generic level, see FIBIGER and LAFONTAINE (2005).94 FIBIGER (1992a).95 Now six more specimens from Malta (Paul SAMMUT, Antony SEGUNA, and Aldo CATANIA pers. comm.).96 BOURSIN (1955) and FIBIGER & HACKER (1992; 1998): South Russia: Kamenka near Saratov; also reported from the South Ural

Mountains of Russia (NUPPONEN & FIBIGER, 2002).97 Recorded from Malta (SAMMUT, 1984; FIBIGER & HACKER, 1998).98 See FIBIGER and LAFONTAINE (2005).99 See GOATER, RONKAY and FIBIGER (2003).

115

Tribus MELIPOTINI GROTE, 1895 (= SYNEDINI FORBES, 1954)100

Genus Drasteria HÜBNER, 1818 (=Protomelipotis BERIO, 2002, syn. n.)101

flexuosa (MÉNÉTRIÉS, 1848)tenera (STAUDINGER, 1877)caucasica (KOLENATI, 1846)picta (CHRISTOPH, 1877)cailino (LEFEBVRE, 1827)saisani (STAUDINGER, 1882)sesquistria (EVERSMANN, 1854)rada (BOISDUVAL, 1848)

Tribus EUCLIDIINI GUENÉE, 1852102

Genus Euclidia OCHSENHEIMER, 1816103

Subgenus Euclidia OCHSENHEIMER, 1816

glyphica (LINNAEUS, 1758)

Subgenus Callistege HÜBNER, [1823]

mi (CLERCK, 1759)fortalitium (TAUSCHER, 1809)

Subgenus Gonospileia HÜBNER, [1823]

munita (HÜBNER, [1813])triquetra ([DENIS & SCHIFFERMÜLLER], 1775)

Tribus OPHIUSINI GUENÉE, 1837104

Genus Catephia OCHSENHEIMER, 1816

alchymista ([DENIS & SCHIFFERMÜLLER], 1775)

Genus Zethes RAMBUR, 1833

insularis RAMBUR, 1833

Genus Pericyma HERRICH-SCHÄFFER, [1851]

albidentaria (FREYER, 1842)

Genus Heteropalpia BERIO, 1939

acrosticta (PÜNGELER, 1904)105

100 See GOATER, RONKAY and FIBIGER (2003) and FIBIGER and LAFONTAINE (2005).101 Protomelipotis BERIO, 2002 was described with sesquistria (EVERSMANN, 1854) as its type species.102 See FIBIGER and LAFONTAINE (2005).103 Arrangement of subgenera and species following GOATER, RONKAY & FIBIGER (2003).104 See GOATER, RONKAY and FIBIGER (2003).105 Recorded from Malta (leg. SAMMUT) (GOATER, RONKAY & FIBIGER, 2003).

116

Genus Pandesma GUENÉE, 1852

robusta (WALKER, [1858])

Genus Cerocala BOISDUVAL, 1828

scapulosa (HÜBNER, [1808])algiriae OBERTHÜR, 1876 (= insana HAMPSON, 1913 nec HERRICH-SCHÄFFER, 1858, nec STAUDINGER, 1871)106

Genus Ophiusa OCHSENHEIMER, 1816

tirhaca (CRAMER, 1777)

Genus Minucia MOORE, [1885]

lunaris ([DENIS & SCHIFFERMÜLLER], 1775)

Genus Clytie HÜBNER, [1823]

delunaris (STAUDINGER, 1889)107

illunaris (HÜBNER, [1813])108

gracilis (BANG-HAAS, 1907)109

syriaca (BUGNION, 1837)

Genus Dysgonia HÜBNER, [1823]

rogenhoferi (BOHATSCH, 1880)110

torrida (GUENÉE, 1852)algira (LINNAEUS, 1767)

Genus Grammodes GUENÉE, 1852 (= Genus Prodotis JOHN, 1910)

bifasciata (PETAGNA, 1787)stolida (FABRICIUS, 1775)

Tribus CATOCALINI BOISDUVAL, [1828]111

Genus Catocala SCHRANK, 1802

Subgenus Catocala SCHRANK, 1802

fulminea (SCOPOLI, 1763)nymphaea (ESPER, [1787])eutychea (TREITSCHKE, 1835)mariana RAMBUR, 1858disjuncta (GEYER, [1828])

106 Taxonomy cf. GOATER, RONKAY & FIBIGER ( 2003) or HACKER (2001).107 Within the revision of the genus Clytie HBN. (HACKER, 2001) not mentioned for the European fauna, but reported by LVOVSKI

(1971) from S Russia: Astrakhan. See GOATER, RONKAY and FIBIGER (2003).108 C. sancta (STAUDINGER, 1898) has never been found in Europe (cf. the revision of the genus Clytie HBN.; HACKER, 2001).109 Revision of the genus Clytie HÜBNER, [1823]) (HACKER, 2001); also reported from the South Ural Mountains of Russia

(NUPPONEN & FIBIGER, 2002).110 Reported by LVOVSKI (1971) from S Russia: Astrakhan. See GOATER, RONKAY and FIBIGER (2003).111 See GOATER, RONKAY and FIBIGER (2003).

117

separata (FREYER, 1848)112

conversa (ESPER, [1787])neonympha (ESPER, [1805])nymphagoga (ESPER, [1787])brandti (HACKER & KAUTT, 1999)113

diversa (GEYER, [1828])hymenaea ([DENIS & SCHIFFERMÜLLER], 1775)fraxini (LINNAEUS, 1758)conjuncta (ESPER, [1787])adultera MÉNÉTRIÉS, 1856nupta (LINNAEUS, 1767)electa (VIEWEG, 1790)elocata (ESPER, [1787])oberthueri (AUSTAUT, 1879)114

deducta EVERSMANN, 1843puerpera (GIORNA, 1791)orientalis STAUDINGER, 1877115

dilecta (HÜBNER, [1808])sponsa (LINNAEUS, 1767)promissa ([DENIS & SCHIFFERMÜLLER], 1775)detrita WARREN, 1913lupina HERRICH-SCHÄFFER, [1851]pacta (LINNAEUS, 1758)optata (GODART, 1824)

Subfamily EUTELIINAE GROTE, 1882

Genus Eutelia HÜBNER, [1823]

adulatrix (HÜBNER, [1813])adoratrix (STAUDINGER, 1892)

Family MICRONOCTUIDAE FIBIGER, 2005116

Genus Micronoctua FIBIGER, 1997

karsholti FIBIGER, 1997117

Family NOCTUIDAE LATREILLE, 1809118

Subfamily PLUSIINAE BOISDUVAL, [1828]119

112 C. separata FRR. is a full species and not a infraspecific form of disjuncta (GEYER, [1828]) (HACKER, 1998c; FIBIGER & HACKER,1998).

113 This species was described from Iran and it occurs in a very isolated population in Greece: Epirus (ssp. schaideri HACKER& HABELER, 1999).

114 FIBIGER (1991) and FIBIGER & HACKER (1992; 1998): Spain.115 New for the European fauna from the western and European part of Kazakhstan and the South Ural Mountains of Russia

(HACKER & MIATLEUSKI, 2001; NUPPONEN & FIBIGER, 2002).116 In FIBIGER & LAFONTAINE, 2005,- in this volume of Esperiana.117 Described from southern Turkey and the south-eastern Greek Islands (FIBIGER, 1997a; FIBIGER & HACKER, 1998).118 The systematic order and sequence of species follow mainly FIBIGER & HACKER (1991, 1992, and 1998; FIBIGER & SKULE

(2004); and FIBIGER & LAFONTAINAE, 2005,-in this volume of Esperiana.119 See GOATER, RONKAY and FIBIGER (2003).

118

Tribus ABROSTOLINI EICHLIN & CUNNINGHAM, 1978

Genus Abrostola OCHSENHEIMER, 1816

tripartita (HUFNAGEL, 1766) (= triplasia auctt. nec LINNAEUS, 1758)120

asclepiadis ([DENIS & SCHIFFERMÜLLER], 1775)121

agnorista DUFAY, 1956triplasia (LINNAEUS, 1758) (= trigemina WERNEBURG, 1864)122

clarissa (STAUDINGER, 1900)123

Tribus ARGYROGRAMMATINI EICHLIN & CUNNINGHAM, 1978

Genus Trichoplusia McDUNNOUGH, 1944

ni (HÜBNER, [1803])

Genus Thysanoplusia ICHNOSÉ, 1973

orichalcea (FABRICIUS, 1775)daubei (BOISDUVAL, 1840)circumscripta (FREYER, 1831)

Genus Ctenoplusia DUFAY, 1970

accentifera (LEFEBVRE, 1827)limbirena (GUENÉE, 1852)vittata (WALLENGREN, 1856)124

Genus Chrysodeixis HÜBNER, [1821]

chalcites (ESPER, [1789])125

acuta (WALKER, [1858])

Tribus PLUSIINI BOISDUVAL, [1828]

Subtribus AUTOPLUSIINA KITCHING, 1987126

Genus Macdunnoughia KOSTROWICKI, 1961

confusa (STEPHENS, 1850)

Genus Diachrysia HÜBNER, [1821]

chryson (ESPER, [1789])

120 Stat. rev. (MIKKOLA & HONEY, 1993; FIBIGER & HACKER, 1998).121 Name of species (cf. FIBIGER & HACKER, 1991).122 Stat. rev. (MIKKOLA & HONEY, 1993; FIBIGER & HACKER, 1998).123 Reported from Bulgaria and European Turkey (BESHKOV, 2000; GOATER, L. RONKAY & FIBIGER, 2003).124 England, 1996 (B. GOATER, pers. com.; FIBIGER & HACKER, 1998).125 It should be mentioned here that the closely related (and also superficially very similar) SE Asian sister-species, C. eriosoma

(DOUBLEDAY, 1843) recently has been recorded on several occations in Germany and Sweden (once), imported as pestspecies into green-houses (HEINICKE, 2000; PALMQVIST, 2004 - with genitalia illustrated).

126 In FIBIGER & LAFONTAINE, 2005,- in this volume of Esperiana.

119

chrysitis (LINNAEUS, 1758)stenochrysis (WARREN, 1913) (= tutti KOSTROWICKI, 1961)127

generosa (Staudinger, 1900)128

nadeja (OBERTHÜR, 1880)zosimi (HÜBNER, [1822])

Subtribus EUCHALCIINA CHOU and LU, 1979129

Genus Euchalcia HÜBNER, [1821]

variabilis (PILLER, 1783) (= cuprea ESPER, [1787])130

bellieri (KIRBY, 1900) (=galloi BERIO, 1989)131

siderifera (EVERSMANN, 1846)italica (STAUDINGER, 1882)consona (FABRICIUS, 1787)modestoides POOLE, 1989 (= modesta HÜBNER, 1786 praeocc.)biezankoi (ALBERTI, 1965)chlorocharis (DUFAY, 1961)emichi (ROGENHOFER & MANN, 1873)132

Genus Polychrysia HÜBNER, [1821]

moneta (FABRICIUS, 1787)esmeralda (OBERTHÜR, 1880)

Genus Panchrysia HÜBNER, [1821]

aurea (HÜBNER, [1803]) (=deaurata ESPER, [1787] nec GOEZE, 1781; =chryson BORKHAUSEN, 1792 nec ESPER,[1789]133

v-argenteum (ESPER, [1798])ornata (Bremer, 1864)134

dives (EVERSMANN, 1844)

Genus Lamprotes REICHENBACH (Leipzig), 1817

c-aureum (KNOCH, 1781)

Genus Plusidia BUTLER, 1879

cheiranthi (TAUSCHER, 1809)

127 Taxonomy cf. GOATER, RONKAY & FIBIGER (2003).128 Recorded from S Russia (GOATER, L. RONKAY & FIBIGER, 2003) and Ukraine (KLJUTSCHKO, pers. comm.).129 In FIBIGER & LAFONTAINE, 2005,- in this volume of Esperiana.130 For the nomenclatural questions concerning the taxa cuprea ESPER, [1787] and modestoides POOLE, 1989 cf. KONONENKO et

al. (1998) and especially HACKER (1998b). HACKER designated a lectotype of cuprea and this species is conspecific withvariabilis (PILLER, 1783). Subsp. uralensis (EVERSMANN, 1842) in the South Ural mountains; subsp. fuscolivacea VARGA &RONKAY, 1984 in Bulgaria.

131 Synonym of galloi BERIO, 1989 by FIBIGER & HACKER (1992; 1998).132 Recorded new for Europe on Kalymnos, Greece (M. FIBIGER, G. JEPPESEN, D. NILSSON & A. MADSEN, in prep.).133 Phalaena Noctua deaurata ESPER, [1787] is a homonym of Phalaena deaurata GOEZE, 1781 (HACKER, 1998b; FIBIGER &

HACKER, 1998).134 New for the European fauna from the Ural mountains (K. NUPPONEN & FIBIGER, in press; GOATER, L. RONKAY & FIBIGER, 2003).

120

Subtribus PLUSIINA BOISDUVAL, [1828]135

Genus Autographa HÜBNER, [1821]

gamma (LINNAEUS, 1758) (=messmeri SCHADEWALD, 1992; =voelkeri SCHADEWALD, 1992)136

mandarina (FREYER, 1845)pulchrina (HAWORTH, 1809)buraetica (STAUDINGER, 1892)jota (LINNAEUS, 1758)macrogamma (EVERSMANN, 1842)aemula ([DENIS & SCHIFFERMÜLLER], 1775)excelsa (KRETSCHMAR, 1862)bractea ([DENIS & SCHIFFERMÜLLER], 1775)

Genus Megalographa LAFONTAINE & POOLE, 1991

biloba (STEPHENS, 1830)137.

Genus Cornutiplusia KOSTROWICKI, 1961

circumflexa (LINNAEUS, 1767)

Genus Syngrapha HÜBNER, [1821]

devergens (HÜBNER, [1813])rilaecacuminum VARGA & L. RONKAY, 1982138

parilis (HÜBNER, [1809])hochenwarthi (HOCHENWARTH, 1785)diasema (BOISDUVAL, 1829)microgamma (HÜBNER, [1823])ain (HOCHENWARTH, 1785)interrogationis (LINNAEUS, 1758)

Genus Plusia OCHSENHEIMER, 1816

festucae (LINNAEUS, 1758)putnami (GROTE, 1873)

Subfamily EUSTROTIINAE GROTE, 1882139

Genus Phyllophila GUENÉE, 1852

obliterata (RAMBUR, 1833)

Genus Protodeltote UEDA, 1984

pygarga (HUFNAGEL, 1766)

Genus Deltote REICHENBACH (Leipzig), 1817

deceptoria (SCOPOLI, 1763)

135 In FIBIGER & LAFONTAINE, 2005,- in this volume of Esperiana.136 New synonyms (FIBIGER , 1993b; FIBIGER & HACKER, 1998).137 Several times migrant in England (GOATER, RONKAY & FIBIGER, 2003).138 Bona species (GOATER, RONKAY & FIBIGER, 2003).139 In FIBIGER & LAFONTAINE, 2005,- in this volume of Esperiana.

121

uncula (CLERCK, 1759)bankiana (FABRICIUS, 1775)

Genus Eulocastra BUTLER, 1886

diaphora (STAUDINGER, 1879)

Genus Pseudozarba WARREN, 1914

bipartita (HERRICH-SCHÄFFER, 1850)

Genus Ozarba WALKER, 1865

moldavicula (HERRICH-SCHÄFFER, 1851)lascivalis (LEDERER, 1855)

Subfamily BAGISARINAE CRUMB, 1956140

Genus Xanthodes GUENÉE, 1852

albago (FABRICIUS, 1794)

Genus Pardoxia VIVES & GONZALEZ, 1981141

graellsi (FEISTHAMEL, 1837)

Subfamily ACONTIINAE GUENÉE, 1841142

Tribus ACONTIINI GUENÉE, 1841143

Genus Acontia OCHSENHEIMER, 1816144

Subgenus Acontia OCHSENHEIMER, 1816145

lucida (HUFNAGEL, 1766)melanura (TAUSCHER, 1809) (=titania sensu auctt. nec ESPER, [1798])146

titania (ESPER, [1798]) (=urania FRIVALDSKY, 1835)147

candefacta (HÜBNER, [1831])147a

Subgenus Emmelia HÜBNER, [1821], stat. nov.148

trabealis (SCOPOLI, 1763)viridisquama (GUENÉE, 1852)

140 Arrangement following CRUMB (1956); KITCHING AND RAWLINS (1999).141 Junior author.142 New autapomorphies are found during the revision by LEGRAIN, HACKER & FIBIGER, in prep. The subfamily comprises more

than 250 species worldwide, many of which are not yet described. See FIBIGER & LAFONTAINE, 2005.143 See FIBIGER & LAFONTAINE (2005).144 The tribe Acontiini is under revision by LEGRAIN, HACKER & FIBIGER, in prep, who state new autapomorphies.145 Revised by LEGRAIN, HACKER & FIBIGER, in prep.146 Bona sp. (HACKER, 1998b).147 Synonym (HACKER, 1998b).147a This nearctic species was introduced to Ukraine and is spreading there rapidly (KLJUTSHKO et al., 2004).148 Revised by LEGRAIN, HACKER & FIBIGER, in prep.

122

Tribus ARMADINI WILTSHIRE, 1961149

Genus Armada STAUDINGER, 1884150

panaceorum (MÉNÉTRIÉS, 1848)barrygoateri FIBIGER, L. & G. RONKAY, 2003

Tribus AEDIINI BECK, 1960151

Genus Aedia HÜBNER, [1823]152

funesta (ESPER, [1766])leucomelas (LINNAEUS, 1758)

Subfamily PANTHEINAE SMITH, 1898153

Genus Panthea HÜBNER, [1820]

coenobita (ESPER, 1785)

Genus Trichosea GROTE, 1875

ludifica (LINNAEUS, 1758)

Genus Colocasia OCHSENHEIMER, 1816

coryli (LINNAEUS, 1758)

Subfamily DILOBINAE AURIVILLIUS, 1889154

Genus Diloba BOISDUVAL, 1840

caeruleocephala (LINNAEUS, 1758)155

Subfamily RAPHIINAE BECK, 1996156

Genus Raphia HÜBNER, [1821]

hybris (HÜBNER, [1813])

149 See FIBIGER & LAFONTAINE (2005).150 See FIBIGER & LAFONTAINE (2005).151 See FIBIGER & LAFONTAINE (2005).152 See FIBIGER & LAFONTAINE (2005).153 See FIBIGER & LAFONTAINE (2005).154 See FIBIGER & LAFONTAINE (2005).155 We tentatively place the often debated Dilobinae here, according to the similarity of the genitalia with those of C. coryli L..

(V. KONONENKO, pers. comm.).156 See FIBIGER & LAFONTAINE (2005).

123

Subfamily ACRONICTINAE HEINEMANN, 1859157

Genus Oxicesta HÜBNER, [1819]

geographica (FABRICIUS, 1787)chamoenices HERRICH-SCHÄFFER, 1845 (=nervosa de VILLERS, 1789, nec RETZIUS, 1783)158

serratae ZERNY, 1927

Genus Eogena GUENÉE, 1852

contaminei (EVERSMANN, 1847)

Genus Moma HÜBNER, [1820]

alpium (OSBECK, 1778)

Genus Acronicta OCHSENHEIMER, 1816

Subgenus Acronicta OCHSENHEIMER, 1816

alni (LINNAEUS, 1767)cuspis (HÜBNER, [1813])tridens ([DENIS & SCHIFFERMÜLLER], 1775)psi (LINNAEUS, 1758)aceris (LINNAEUS, 1758)leporina (LINNAEUS, 1758)

Subgenus Subacronicta KOZHANTSHIKOV, 1950

megacephala ([DENIS & SCHIFFERMÜLLER], 1775)

Subgenus Hyboma HÜBNER, [1820]

strigosa ([DENIS & SCHIFFERMÜLLER], 1775)

Subgenus Viminia CHAPMAN,1890

menyanthidis (ESPER, [1789])auricoma ([DENIS & SCHIFFERMÜLLER], 1775)euphorbiae ([DENIS & SCHIFFERMÜLLER], 1775)cinerea (HUFNAGEL, 1766)158a

orientalis MANN, 1862rumicis (LINNAEUS, 1758)

Genus Craniophora SNELLEN, 1867

ligustri ([DENIS & SCHIFFERMÜLLER], 1775)pontica (STAUDINGER, 1879)

Genus Simyra OCHSENHEIMER, 1816

nervosa ([DENIS & SCHIFFERMÜLLER], 1775)albovenosa (GOEZE, 1781)

157 See FIBIGER & LAFONTAINE (2005).158 Homonym (LERAUT, 1997; FIBIGER & HACKER, 1998).158a The status of this taxon is not yet studied.

124

dentinosa FREYER, 1839

Subfamily METOPONIINAE HERRICH-SCHÄFFER, [1851]160

Genus Panemeria HÜBNER, [1823]

tenebrata (SCOPOLI, 1763) (=jocosa ZELLER, 1847)161

tenebromorpha RÁKOSY, HENTSCHOLEK & HUBER, 1996162

Genus Apaustis HÜBNER, [1823]

rupicola ([DENIS & SCHIFFERMÜLLER], 1775) (=theophila STAUDINGER, 1866)163

Genus Alvaradoia AGENJO, 1984164

numerica (BOISDUVAL, 1840)

Genus Aegle HÜBNER, [1823]

semicana (ESPER, [1798]) (=vespertalis HÜBNER, [1813])165

vespertinalis (RAMBUR, 1858)166

agatha (STAUDINGER, 1861)kaekeritziana (HÜBNER, [1799])exquisita BOURSIN, 1969

Genus Metaegle HAMPSON, 1908

pallida (STAUDINGER, 1892)167

Genus Megalodes GUENÉE, 1852

eximia (FREYER, 1845)

Genus Synthymia HÜBNER, [1823]

fixa (FABRICIUS, 1787)

Genus Haemerosia BOISDUVAL, 1840

renalis (HÜBNER, [1813])vassilininei A. BANG-HAAS, 1912

160 Following MATTHEWS (1991) the genera listed here were previously placed in the Nearctic subfamily Stiriinae GROTE, 1882,see FIBIGER & LAFONTAINE, 2005.

161 Synonym (FIBIGER & HACKER, 1998; cf. BERIO, 2002).162 This species was described from N. Greece163 Synonym (FIBIGER & HACKER, 1998).164 Transferred from Eustrotiinae.165 Synonym, see HACKER, 1998b; FIBIGER & HACKER, 1998.166 Year of publication.167 BESHKOV (2000): Bulgaria.

125

Genus Mesotrosta LEDERER, 1857

signalis (TREITSCHKE, 1829)

Genus Mycteroplus HERRICH-SCHÄFFER, [1850]

puniceago (BOISDUVAL, 1840)

Genus Tyta BILLBERG, 1820167a

luctuosa ([DENIS & SCHIFFERMÜLLER], 1775)

Subfamily CUCULLIINAE HERRICH-SCHÄFFER, [1850]

Genus Cucullia SCHRANK, 1802168

Subgenus Cucullia SCHRANK, 1802169

argentina (FABRICIUS, 1787)bubaceki KITT, 1925magnifica FREYER, 1840 (= scopula FISCHER von WALDHEIM, 1839)splendida (STOLL, 1782)scopariae DORFMEISTER, 1853fuchsiana EVERSMANN, 1842fraudatrix EVERSMANN, 1837formosa ROGENHOFER, 1860absinthii (LINNAEUS, 1761)170

argentea (HUFNAGEL, 1766)spectabilisoides POOLE, 1989 (= spectabilis HÜBNER, [1813] nec TAUSCHER, 1806)cineracea FREYER, 1842lindei HEYNE, 1899171

artemisiae (HUFNAGEL, 1766)praecana EVERSMANN, 1843mixta FREYER, 1842cemenelensis BOURSIN, 1923172

lactea (FABRICIUS, 1787)xeranthemi BOISDUVAL, 1840propinqua EVERSMANN, 1842tiefi TSHETVERIKOV, 1956lactucae ([DENIS & SCHIFFERMÜLLER], 1775)fraterna BUTLER, 1878lucifuga ([DENIS & SCHIFFERMÜLLER], 1775)umbratica (LINNAEUS, 1758)biornata FISCHER von WALDHEIM, 1840balsamitae BOISDUVAL, 1840sabulosa STAUDINGER, 1879campanulae FREYER, [1831]santolinae RAMBUR, 1834hartigi RONKAY & RONKAY, 1988calendulae TREITSCHKE, 1835 (= wredowi O. G. COSTA, [1835]; = dellabrunai BERIO, 1980)chamomillae ([DENIS & SCHIFFERMÜLLER], 1775)

167a See FIBIGER & LAFONTAINE, 2005.168 RONKAY & RONKAY (1994).169 See under Shargacucullia RONKAY & RONKAY, 1992.170 Name of the species (cf. RONKAY & RONKAY1994).171 Year of publication.172 Year of publication.

126

inderiensis [HERRICH-SCHÄFFER, 1856]173

santonici (HÜBNER, [1813])naruenensis STAUDINGER, 1879boryphora FISCHER von WALDHEIM, 1840achilleae GUENÉE, 1852improba CHRISTOPH, 1885syrtana (MABILLE, 1888)biskrana (OBERTHÜR, 1918)gnaphalii (HÜBNER, [1813])tanaceti ([DENIS & SCHIFFERMÜLLER], 1775)dracunculi (HÜBNER, [1813])virgaureae BOISDUVAL, 1840amota ALPHÉRAKY, 1887asteris ([DENIS & SCHIFFERMÜLLER], 1775)

Subgenus Shargacucullia RONKAY & RONKAY, 1992174 stat. rev.

scrophulariphaga (RAMBUR, 1833)caninae (RAMBUR, 1833)blattariae (ESPER, [1790]) (= minogenica REBEL, 1916; = eugeniae BECK, 1989)scrophulariphila (STAUDINGER, 1859)erythrocephala (F. WAGNER, 1914) (=scrophularivora GUENÉE, 1852, nomen nudum)gozmanyi RONKAY & RONKAY, 1994 (= blattariae eugeniae auct.)175

scrophulariae ([DENIS & SCHIFFERMÜLLER], 1775)lanceolata (VILLERS, 1789) (=thapsiphaga TREITSCHKE, 1826)176

reisseri (BOURSIN, 1933)lychnitis (RAMBUR, 1833)verbasci (LINNAEUS, 1758)prenanthis (BOISDUVAL, 1840)177

Subgenus Calocucullia RONKAY & RONKAY, 1987, stat. rev.178

celsiae (HERRICH-SCHÄFFER, 1850)

Subfamily ONCOCNEMIDINAE FORBES & FRANCLEMONT, 1954

Genus Brachygalea HAMPSON, 1906179

albolineata (BLACHIER, 1905)

Genus Calophasia STEPHENS, 1829

lunula (HUFNAGEL, 1766)hamifera STAUDINGER, 1863platyptera (ESPER, [1788])180

almoravida de GRASLIN, 1863barthae F. WAGNER, 1929opalina (ESPER, [1794]) (= casta BORKHAUSEN, 1793)

173 Year of publication (HEPPNER, 1982).174 RONKAY & RONKAY (1992; 1994). This taxon was synonymised first by POOLE (1995) and further by KULLBERG et al. (2002), but

we believe that this clade merits subgeneric status, based on the superficial distinctness and by the synapomorphic structuresof their genitalia, especially the vesica

175 RONKAY & RONKAY (1994) sp. n.: Hungary; Slovakia.176 Synonym (LERAUT, 1997; FIBIGER & HACKER, 1998).177 Year of publication.178 RONKAY & RONKAY (1994); stat. rev., see above note 174.179 Synonymy of Criophasia HAMPSON, 1906 cf. RONKAY & RONKAY (1995). The two genera have been described in the same

publication. WARREN (1910) already used Brachygalea, in this case RONKAY & RONKAY, 1995 are right in treating Criophasiaas a synonym.

180 It has not been possible to trace european records of angularis CHRETIÈN, 1911 (= strigmatica ROTHSCHILD, 1913).

127

Genus Pamparama RONKAY & RONKAY, 1995181

acuta (FREYER, 1837)

Genus Behounekia HACKER, 1990

freyeri (FRIVALDSKY, 1835)

Genus Omphalophana HAMPSON, 1906

antirrhinii (HÜBNER, [1803])serrata (TREITSCHKE, 1835)anatolica (LEDERER, 1857)

Genus Sympistis HÜBNER, [1823]

funebris (HÜBNER, [1809]) (= funesta PAYKULL, 1793)lapponica (THUNBERG, 1791)heliophila (PAYKULL, 1793) (= melaleuca THUNBERG, 1791 praeocc.)nigrita (BOISDUVAL, 1840)182 (= subsp. zetterstedti STAUDINGER, 1857)

Genus Oncocnemis LEDERER, 1853

confusa (FREYER, [1839])strioligera LEDERER, 1853campicola LEDERER, 1853nigricula (EVERSMANN, 1847)senica (EVERSMANN, 1856) (= literata BREMER, 1864)

Genus Calliergis HÜBNER, [1821]183

ramosa (ESPER, [1786])

Genus Lophoterges HAMPSON, 1906

millierei (STAUDINGER, 1870)hoerhammeri (F. WAGNER, 1931)

Genus Epimecia GUENÉE, 1839

ustula (FREYER, 1835)

Genus Rhabinopteryx CHRISTOPH, 1889

subtilis (MABILLE, 1888)

Genus Teinoptera CALBERLA, 1891 (=Copiphana HAMPSON, 1906; = Wiltshireria BERIO, 2002 syn. n.)184

olivina (HERRICH-SCHÄFFER, 1852)

181 RONKAY & RONKAY (1995): gen. n.182 RONKAY & RONKAY (1995): zetterstedti (STAUDINGER, 1857) is just a subspecies of nigrita (BOISDUVAL, 1840).183 Name of the genus (cf. FIBIGER & HACKER, 1991).184 Synonym (HACKER, 1998b; FIBIGER & HACKER, 1998). Wiltshireria BERIO, 2002, described for the generotype kraussi REBEL is

also a synonym of Teinoptera CALBERLA, 1891 syn. n.

128

lunaki (BOURSIN, 1940)oliva (STAUDINGER, 1895)185

Genus Cleonymia BERIO, 1966

Subgenus Cleonymia BERIO, 1966 (=Pectinania BERIO, 2002 syn. n.)186

baetica (RAMBUR, 1837)chabordis (OBERTHÜR, 1876)

Subgenus Serryvania BERIO, 1980187

yvanii (DUPONCHEL, 1833)diffluens (STAUDINGER, 1870)opposita (LEDERER, 1870)pectinicornis (STAUDINGER, 1859)

Genus Amephana HAMPSON, 1906

Subgenus Amephana HAMPSON, 1906

anarrhini (DUPONCHEL, [1840])

Subgenus Trigonephra BERIO, 1980

aurita (FABRICIUS, 1787)dalmatica (REBEL, 1919)

Genus Omia HÜBNER, [1821]

cymbalariae (HÜBNER, [1809])banghaasi STAUDER, 1930cyclopea (de GRASLIN, 1837)

Genus Harpagophana HAMPSON, 1906

hilaris (STAUDINGER, 1895)

Genus Recophora NYE, 1975

canteneri (DUPONCHEL, 1833)

Genus Metopoceras GUENÉE, 1850

Subgenus Metopoceras GUENÉE, 1850

felicina (DONZEL, 1844)omar (OBERTHÜR, 1887)

Subgenus Tritomoceras BERIO, 1980

albarracina HAMPSON, 1918khalildja OBERTHÜR, 1884

185 RONKAY & RONKAY (1995): one European specimen recorded from Greece in 1896, coll. ZMUC.186 Pectinania BERIO, 2002, described for the generotype chabordis OBERTHÜR, 1876 is a synonym of Cleonymia BERIO, 1966

syn. n.187 Ronkayiana FIBIGER & HACKER is a nomen nudum.

129

Genus Valerietta DRAUDT, 1938188

bulgarica (DRENOWSKI, 1953)niphopasta (HAMPSON, 1906)189

Genus Lamprosticta HÜBNER, [1820]

culta ([DENIS & SCHIFFERMÜLLER], 1775)

Genus Xylocampa GUENÉE, 1837

areola (ESPER, [1789])mustapha (OBERTHÜR, 1910)

Genus Stilbia STEPHENS, 1829190

anomala (HAWORTH, 1812)andalusiaca STAUDINGER, 1892calberlae (FAILLA-TEDALDI, 1890)faillae PÜNGELER, 1891191

philopalis de GRASLIN, 1852

Genus Praestilbia STAUDINGER, 1892

armeniaca STAUDINGER, 1892

Genus Stilbina STAUDINGER, 1892

Subgenus Lukaschia BECK, 1991192

olympica DIERL & POVOLNY, 1970

Subfamily AMPHIPYRINAE GUENÉE, 1837193

Genus Pyrois HÜBNER, [1820]

cinnamomea (GOEZE, 1781)effusa (BOISDUVAL, [1828])

Genus Amphipyra OCHSENHEIMER, 1816

pyramidea (LINNAEUS, 1758)berbera RUNGS, 1949perflua (FABRICIUS, 1787)livida ([DENIS & SCHIFFERMÜLLER], 1775)tragopoginis (CLERCK, 1759)tetra (FABRICIUS, 1787)

188 HREBLAY (1992); BESHKOV (2000).189 BESHKOV (2000): Bulgaria: Spirka Kresna.190 RONKAY et al. (2001).191 Year of description.192 Subgenus stat. rev. (RONKAY et al., 2001).193 See FIBIGER & LAFONTAINE, 2005.

130

micans LEDERER, 1857molybdea CHRISTOPH, 1867194

submicans KUZNETZOV, 1958stix HERRICH-SCHÄFFER, 1850

Genus Phidrimana KONONENKO, 1989195

amurensis (STAUDINGER, 1892)196

Subfamily PSAPHIDINAE GROTE, 1896197

Tribus PSAPHIDINI GROTE, 1896 198

Genus Asteroscopus BOISDUVAL, 1828199

sphinx (HUFNAGEL, 1766)syriaca (WARREN, 1910) (= decipulae KOVACS, 1966)

Genus Brachionycha HÜBNER, [1819]

nubeculosa (ESPER, 1785)sajana DRAUDT, 1934

Genus Valeria STEPHENS, 1829

oleagina ([DENIS & SCHIFFERMÜLLER], 1775)jaspidea (de VILLERS, 1789)

Genus Meganephria HÜBNER, [1820]

bimaculosa (LINNAEUS, 1767)

Genus Allophyes TAMS, 1942200

oxyacanthae (LINNAEUS, 1758)corsica (SPULER, 1908)201

asiatica (STAUDINGER, 1892)alfaroi AGENJO, 1951cretica PINKER & REISSER, 1978

194 Described from one single female originating from Sarepta. It is unclear, if molybdea CHRISTOPH, 1867 and submicansKUZNETZOV, 1958 are conspecific and why molybdea is said to be a senior synonym of submicans, because according toPOOLE (1989) the name was not preoccupied by an older taxon in the same combination. According to ANIKIN et al. (2000b)the species (given as submicans = molybdea) was recorded in Europe in the districts of Volgograd, Saratov and Uralsk.

195 Not in Apameini, but here.196 This eastern steppe species (E Sibiria; Russian Far East) was reported from the Orenburg region (KONONENKO, 1989),

South Russia (ANIKIN et al., 2000b) and: S Ural, Orenburg (K. NUPPONEN & FIBIGER, 2002).197 See FIBIGER & LAFONTAINE, 2005.198 Systematic position cf. RONKAY & RONKAY (1995).199 Gen. rest. (RONKAY & RONKAY, 1995).200 The genus contains some taxonomic problems concerning the confirmation of species or subspecies status of some taxa

(MAZEL, 1991; FIBIGER & HACKER, 1998; RONKAY et al., 2001).201 With subsp. protai BOURSIN, 1966; and subsp. parenzani de LAEVER, 1977.

131

Tribus FERALIINI POOLE, 1995

Genus Feralia GROTE, 1874

sauberi (GRAESER, 1892)

Subfamily HELIOTHINAE BOISDUVAL, [1828]

Genus Schinia HÜBNER, [1818] (= Protoschinia HARDWICK, 1970)

cardui (HÜBNER, 1790)cognata (FREYER, 1833)purpurascens (TAUSCHER, 1809)scutosa ([DENIS & SCHIFFERMÜLLER], 1775)202

Genus Stenoecia WARREN, 1911

dos (FREYER, 1838)

Genus Pyrocleptria HAMPSON, 1903

cora (EVERSMANN, 1837)

Genus Heliothis OCHSENHEIMER, 1816

viriplaca (HUFNAGEL, 1766)maritima de GRASLIN, 1855ononis ([DENIS & SCHIFFERMÜLLER], 1775)peltigera ([DENIS & SCHIFFERMÜLLER], 1775)nubigera HERRICH-SCHÄFFER, 1851

Genus Helicoverpa HARDWICK, 1965

armigera (HÜBNER, [1808])

Genus Pyrrhia HÜBNER, [1821]

umbra (HUFNAGEL, 1766)exprimens (WALKER, 1857)purpurina (ESPER, [1804])

Genus Periphanes HÜBNER, [1821]

delphinii (LINNAEUS, 1758)victorina (SODOFFSKY, 1849)treitschkei (FRIVALDSKY, 1835)

Genus Chazaria MOORE, 1881 (= Rhodocleptria HAMPSON, 1903)

incarnata (FREYER, 1838)203

202 Combination (MATTHEWS, 1991; FIBIGER & HACKER, 1998).203 POOLE (1989); FIBIGER & HACKER (1998).

132

Genus Aedophron LEDERER, 1857

rhodites (EVERSMANN, 1851)phlebophora LEDERER, 1858

Genus Janthinea GUENÉE, 1852

friwaldskii (DUPONCHEL, 1835)

Genus Hebdomochondra STAUDINGER, 1879

syrticola STAUDINGER, 1879

Subfamily CONDICINAE POOLE, 1995204

Genus Condica WALKER, 1856 (=Platysenta GROTE, 1874)

viscosa (FREYER, 1831)capensis (WALKER, 1857)205

Genus Hadjina STAUDINGER, 1892

wichti (HIRSCHKE, 1904)

Genus Acosmetia STEPHENS, 1829

caliginosa (HÜBNER, [1813])

Genus Eucarta LEDERER, 1857206

amethystina (HÜBNER, [1803])virgo (TREITSCHKE, 1835)

Subfamily ERIOPINAE HERRICH-SCHÄFFER, [1851]207

Genus Callopistria HÜBNER, [1821]

juventina (STOLL, 1782)latreillei (DUPONCHEL, 1827)

Subfamily BRYOPHILINAE GUENÉE, 1852

Genus Cryphia HÜBNER, 1818

Subgenus Cryphia HÜBNER, 1818

204 Arrangement follows KITCHING and RAWLINS (1999).205 Denmark, England; probably migration, but might have been introduced by fruits, vegetables or flowers (FIBIGER & HACKER,

1998). Occurs regularly in southern Spain, Andalusia (G. JEPPESEN, pers. comm.).206 See FIBIGER & LAFONTAINE, 2005.207 The name for the subfamily was already used by several authors (e.g. POOLE, 1995; BECK 1999); it is derived from Eriopidae

HERRICH-SCHÄFFER, [1845], type genus Eriopus TREITSCHKE, 1825 (= Callopistria HÜBNER, [1821]). See FIBIGER & LAFONTAINE,2005.

133

receptricula (HÜBNER, [1803]omalosi SVENDSEN & FIBIGER, 1998208

distincta (CHRISTOPH, 1887)209

fraudatricula (HÜBNER, [1803])amygdalina BOURSIN, 1963simulatricula (GUEÉE, 1852)algae (FABRICIUS, 1775)pallida (BETHUNE-BAKER, 1894)ochsi BOURSIN, 1940210

Subgenus Bryoleuca HAMPSON, 1908

ravula (HÜBNER, [1813])vandalusiae (DUPONCHEL, 1842)ereptricula (TREITSCHKE, 1825)tephrocharis BOURSIN, 1953rectilinea (WARREN, 1909)seladona (CHRISTOPH, 1885)raptricula ([DENIS & SCHIFFERMÜLLER], 1775)gea (SCHAWERDA, 1934)orthogramma BOURSIN, 1954petrea (GUENÉE, 1852)petricolor (LEDERER, 1870)strobinoi DUJARDIN, 1972211

Subgenus Bryophila TREITSCHKE, 1825

domestica (HUFNAGEL, 1766)maeonis (LEDERER, 1865)

Subgenus Bryopsis BOURSIN, 1970

muralis (FORSTER, 1771)amasina (DRAUDT, 1931)

Subgenus Bryonycta BOURSIN, 1955

pineti (STAUDINGER, 1859)

Genus Victrix STAUDINGER, 1879

Subgenus Victrix STAUDINGER, 1879212

agenjoi (FERNÁNDEZ, 1931)

Subgenus Moureia ORFILA & ROSSI, 1956

microglossa (RAMBUR, 1858)

Subgenus Poliobrya HAMPSON, 1908

umovii (EVERSMANN, 1846)

208 Described from Crete (SVENDSEN & FIBIGER, 1998; FIBIGER & HACKER, 1998).209 New for the European fauna from the western and European part of Kazakhstan (HACKER & MIATLEUSKI, 2001).210 This species was described two times (1940/1941) using the same type material (FIBIGER & HACKER, 1998).211 The taxon is under revision.212 Subgenus (VARGA & RONKAY, 1989).

134

Subfamily XYLENINAE GUENÉE, 1837213

Tribus PSEUDEUSTROTIINI BECK, 1996

Genus Pseudeustrotia WARREN, 1913214

candidula ([DENIS & SCHIFFERMÜLLER], 1775)

Tribus PRODENIINI FORBES, 1954

Genus Spodoptera GUENÉE, 1852

exigua (HÜBNER, [1808])cilium (GUENÉE, 1852)littoralis (BOISDUVAL, 1833)

Tribus ELAPHRIINI BECK, 1996

Genus Galgula GUENÉE, 1852

partita GUENÉE, 1852

Genus Elaphria HÜBNER, [1818]215

venustula (HÜBNER, 1790)

Tribus CARADRININI BOISDUVAL, 1840

Subtribus CARADRININA BOISDUVAL, 1840216

Genus Caradrina OCHSENHEIMER, 1816216a

Subgenus Caradrina OCHSENHEIMER, 1816

morpheus (HUFNAGEL, 1766)

Subgenus Platyperigea J. B. SMITH, 1894

terrea FREYER, 1840proxima RAMBUR, [1837] (= infusca RAMBUR, 1858)kadenii FREYER, 1836aspersa RAMBUR, 1834montana BREMER, 1861217

albina EVERSMANN, 1848petraea TENGSTRÖM, 1869 (=grisea EVERSMANN, 1848)

213 See FIBIGER & LAFONTAINE, 2005, in this volume of Esperiana.214 The systematic position of this genus has been disputed for a long time (e.g. ZILLI, 1997). It is surprisingly found to be a

member of this subfamily (D. LAFONTAINE, pers. obs.,-see FIBIGER & LAFONTAINE, 2005,-in this volume of Esperiana).215 See FIBIGER & LAFONTAINE, 2005, in this volume of Esperiana.216 See the revision by HACKER (2004), see also FIBIGER & LAFONTAINE, 2005.216a See the revision by HACKER (2004).217 With subsp. menetriesii KRETSCHMAR, 1863 and rougemonti SPULER, 1908.

135

Subgenus Boursinidrina HACKER, 2004

syriaca STAUDINGER, [1892]germainii (DUPONCHEL, 1835) (=psammopsis BOURSIN, 1967)

Subgenus Kalchbergiana HACKER, 2004

ingrata STAUDINGER, 1897

Subgenus Eremodrina BOURSIN, 1937

vicina STAUDINGER, 1870218

armeniaca (BOURSIN, 1936) (=clara SCHAWERDA, 1928 praeocc.)expansa ALPHÉRAKY, 1887219

inumbrata (STAUDINGER, 1900)220

draudti (BOURSIN, 1936) (=pseudopertinax BOURSIN, 1939)221

zernyi (BOURSIN, 1936)222

ibeasi (FERNÁNDEZ, [1918])fergana STAUDINGER, [1892]223

pertinax (STAUDINGER, 1879)gilva (DONZEL, 1837)flava OBERTHÜR, 1876distigma CHRÈTIEN, 1913

Subgenus Paradrina BOURSIN, 1937

selini BOISDUVAL, 1840fuscicornis RAMBUR, 1832abruzzensis (DRAUDT, 1933)minoica HACKER, 2004suscianja (von MENTZER, 1981)wullschlegeli PÜNGELER, 1903224

fulvafusca HACKER, 2004225

flavirena GUENÉE, 1852 (=muricolor BOURSIN, 1933)noctivaga BELLIER, 1863levantina HACKER, 2004226

clavipalpis (SCOPOLI, 1763)

Genus Hoplodrina BOURSIN, 1937

octogenaria (GOEZE, 1781) (= alsines BRAHM, 1791)blanda ([DENIS & SCHIFFERMÜLLER], 1775)superstes (OCHSENHEIMER, 1816)hesperica DUFAY & BOURSIN, 1960respersa ([DENIS & SCHIFFERMÜLLER], 1775)ambigua ([DENIS & SCHIFFERMÜLLER], 1775)

Genus Chilodes HERRICH-SCHÄFFER, [1849]

maritima (TAUSCHER, 1806)

218 Also from Malta (P. SAMMUT, A. SEGUNA, and A. CATANIA, pers. comm.).219 Recorded from S-Ural.220 Recorded from S-Ural.221 In Europe from Crete (subsp. cretica REISSER, 1958).222 Described from SE European Russia, Sarepta.223 New to the European fauna from S. Russia: 1 �, „S. Rußland“ [Sarepta] (gen.prep. HACKER N 9585�) (Museum für Tierkunde,

Dresden) (FIBIGER & HACKER, 1998).224 With subsp. hispanica MABILLE, 1906, callei YELA, 1987, and scythica HACKER, 2004.225 In Europe known only from Rostov/Don (POLTAVSKI & NEKRASOV, 2002), reported as C. poecila (BOURSIN, 1939).226 It seems likely that C. jacobsi (ROTHSCHILD, 1914) occurs in the southernmost Iberian Peninsula, but the separation from

small specimens of C. flavirena GUENÉE, 1852 is difficult and requires genitalia checking (see HACKER, 2004).

136

distracta (EVERSMANN, 1848)

Genus Scythocentropus SPEISER, 1902 (= Ecbolemia HAMPSON, 1908)227

scripturosa (EVERSMANN, 1854)228

inquinata (MABILLE, 1888)misella (PÜNGELER, 1908)229

Genus Rusina STEPHENS, 1829230

ferruginea (ESPER, [1785])231

Genus Pseudoxestia BOURSIN, 1953232

apfelbecki (REBEL, 1901)

Genus Charanyca BILLBERG, 1820233

trigrammica (HUFNAGEL, 1766) (= erubescens TURATI, 1909)

Subtribus ATHETISINA FIBIGER & LAFONTAINE, 2005234

Genus Athetis HÜBNER, [1821]

gluteosa (TREITSCHKE, 1835) (= kitti REBEL, 1913)235

furvula (HÜBNER, [1808])

Genus Hydrillula TAMS, 1938

pallustris (HÜBNER, [1808])

Genus Proxenus HERRICH-SCHÄFFER, [1850]

lepigone (MÖSCHLER, 1860)hospes (FREYER, 1831)

Tribus COSMIINI GUENÉE, 1852236

Genus Enargia HÜBNER, [1821]

paleacea (ESPER, [1788])abluta (HÜBNER, [1808]) (= imbuta BOISDUVAL, 1840)

227 See HACKER (2001).228 Name of the species (cf. FIBIGER & HACKER, 1991).229 See NUPPONEN and FIBIGER (2002).230 Not in Apameini.231 The nomenclatural valid name of the species - ferruginea (ESPER, [1785]) or tristis (RETZIUS, 1783) - is not yet clear.232 Not in Apameini.233 Not in Apameini.234 In this volume of Esperiana.235 See HUEMER and TARMANN (1993).236 See FIBIGER and LAFONTAINE (2005).

137

Genus Ipimorpha HÜBNER, [1821]

retusa (LINNAEUS, 1761)subtusa ([DENIS & SCHIFFERMÜLLER], 1775)contusa (FREYER, 1849)

Genus Cosmia OCHSENHEIMER, 1816

Subgenus Cosmia OCHSENHEIMER, 1816

diffinis (LINNAEUS, 1767)affinis (LINNAEUS, 1767)

Subgenus Calymnia HÜBNER, [1821]

trapezina (LINNAEUS, 1758)confinis HERRICH-SCHÄFFER, [1849] (= rhodopsis BOURSIN, 1962)pyralina ([DENIS & SCHIFFERMÜLLER], 1775)

Genus Dicycla GUENÉE, 1852

oo (LINNAEUS, 1758)

Genus Atethmia HÜBNER, [1821]

ambusta ([DENIS & SCHIFFERMÜLLER], 1775)algirica (CULOT, 1917)centrago (HAWORTH, 1809)

Genus Mesogona BOISDUVAL, 1840236a

acetosellae ([DENIS & SCHIFFERMÜLLER], 1775)oxalina (HÜBNER, [1803])

Tribus DYPTERYGIINI FORBES, 1954237

Genus Dypterygia STEPHENS, 1829

scabriuscula (LINNAEUS, 1758)

Genus Trachea OCHSENHEIMER, 1816

atriplicis (LINNAEUS, 1758)

Genus Anthracia HÜBNER, [1823]

eriopoda (HERRICH-SCHÄFFER, [1851])ephialtes (HÜBNER, [1822])

Genus Mormo OCHSENHEIMER, 1816

maura (LINNAEUS, 1758)

236a Arrangement of the species and genus according to discussions between LAFONTAINE and FIBIGER.237 See FIBIGER and LAFONTAINE, 2005,-in this volume of Esperiana.

138

Genus Polyphaenis BOISDUVAL, 1840

viridis (VILLERS, 1789) (=sericata ESPER, [1787]238; = albibasis WARREN, 1911)

Genus Olivenebula KISHIDA & YOSHIMOTO, 1977 (= Chlorothalpa BECK, 1996)239

subsericata (HERRICH-SCHÄFFER, [1861])xanthochloris (BOISDUVAL, 1840)

Genus Thalpophila HÜBNER, [1820]

matura (HUFNAGEL, 1766)vitalba (FREYER, 1834)

Genus Phoebophilus STAUDINGER, 1888

veternosus (PÜNGELER, 1908)240

Genus Heterophysa BOURSIN, 1953

dumetorum (GEYER, [1834])

Tribus ACTINOTIINI BECK, 1996241

Genus Hyppa DUPONCHEL, [1845]

rectilinea (ESPER, [1788])

Genus Brachyxanthia BUTLER, 1878

zelotypa (LEDERER, 1853)242

Genus Actinotia HÜBNER, [1821]

polyodon (CLERCK, 1759)radiosa (ESPER, [1804])

Genus Chloantha BOISDUVAL, RAMBUR & GRASLIN, [1836]243

hyperici ([DENIS & SCHIFFERMÜLLER], 1775)

Tribus PHLOGOPHORINI HAMPSON, 1918244

Genus Phlogophora TREITSCHKE, 1825

scita (HÜBNER, 1790)

238 A homonym as stated by several authors (cf. RAINERI & ZILLI, 1995; NOWACKI & FIBIGER, 1996; LERAUT, 1997).239 Combination cf. HACKER (2001).240 FIBIGER & HACKER (1992; 1998): Russia: South Ural Mts.: Guberla.241 See FIBIGER & LAFONTAINE, 2005.242 FIBIGER & HACKER (1992): Russia: Perm („westl. Uralvorland“) (EVERSMANN, 1856).243 FIBIGER (1997b).244 See FIBIGER & LAFONTAINE, 2005. The genus Paranataelia DRAUDT, 1935 and its species tenerifica (HAMPSON, 1906) (= whitei

REBEL, 1906), syn. n., were previously listed as to occur on the Azores (FIBIGER and HACKER, 1991), but it is absent fromthese islands and is an endemic species of the Canary Islands.

139

meticulosa (LINNAEUS, 1758) (= lamii SCHADEWALD, 1992)245

interrupta (HAMPSON, 1908)cabrali PINKER, 1971furnasi PINKER, 1971246

Genus Euplexia STEPHENS, 1829

lucipara (LINNAEUS, 1758)

Genus Auchmis HÜBNER, [1821]

detersa (ESPER, [1787])

Genus Pseudenargia BOURSIN, 1956

ulicis (STAUDINGER, 1859)

Tribus APAMEINI GUENÉE, 1841(= Nonagriini GUENÉE, 1837, a synonym of Apameini)247

Subtribus OXYTRYPINA GOZMANY, 1970248

Genus Oxytrypia STAUDINGER, 1871

orbiculosa (ESPER, [1799])

Subtribus APAMEINA GUENÉE, 1841

Genus Apamea OCHSENHEIMER, 1816

monoglypha (HUFNAGEL, 1766)baischi HACKER, 1989sicula (TURATI, 1909) (=indiges sensu auctt. nec TURATI, 1926)249

syriaca OSTHELDER, 1933 (= tallosi KOVACS & VARGA, 1969)250

lithoxylaea ([DENIS & SCHIFFERMÜLLER], 1775)sublustris (ESPER, [1788])arabs (OBERTHÜR, 1881)crenata (HUFNAGEL, 1766)epomidion (HAWORTH, 1809) (=characterea sensu auctt. nec ([DENIS & SCHIFFERMÜLLER], 1775)251

aquila DONZEL, 1837lateritia (HUFNAGEL, 1766)furva ([DENIS & SCHIFFERMÜLLER], 1775)maillardi (GEYER, [1834])zeta (TREITSCHKE, 1825) (= exulis LEFEBVRE, 1836)michielii VARGA, 1976rubrirena (TREITSCHKE, 1825)platinea (TREITSCHKE, 1825)oblonga (HAWORTH, 1809)remissa (HÜBNER, [1809])

245 FIBIGER (1993b); FIBIGER & HACKER (1998), new synonym.246 Name of the species (cf. FIBIGER & HACKER, 1991); Acores.247 According to the nomenclatural code (2000) we accept the well known name Apameini - see also FIBIGER & LAFONTAINE,

2005,-in this volume of Esperiana. The genera and species, incl. their sequences in this tribe are under revision (ZILLI,RONKAY and FIBIGER, in prep., in Noctuidae Europaeae).

248 See FIBIGER and LAFONTAINE (2005).249 FIBIGER & HACKER (1998).250 FIBIGER & HACKER (1998).251 Valid name (KARSHOLT, in: KARSHOLT & RAZOWSKI, 1996).

140

unanimis (HÜBNER, [1813])illyria FREYER, 1846leucodon (EVERSMANN, 1837)anceps ([DENIS & SCHIFFERMÜLLER], 1775) (= siegeli BERIO, 1985)sordens (HUFNAGEL, 1766)ferrago (EVERSMANN, 1837)pyxina (A. BANG-HAAS, 1910)alpigena (BOISDUVAL, [1837])scolopacina (ESPER, [1788])

Genus Leucapamea SUGI, 1982

ophiogramma (ESPER, [1794])

Genus Eremobina MCDUNNOUGH, 1927 (=Pabulatrix SUGI, 1982)252

pabulatricula (BRAHM, 1791)

Genus Oligia HÜBNER, [1821]

strigilis (LINNAEUS, 1758)versicolor (BORKHAUSEN, 1792)latruncula ([DENIS & SCHIFFERMÜLLER], 1775)dubia (HEYDEMANN, 1942)fasciuncula (HAWORTH, 1809)

Genus Mesoligia BOURSIN, 1965

furuncula ([DENIS & SCHIFFERMÜLLER], 1775)literosa (HAWORTH, 1809) (= faroulti ROTHSCHILD, 1914)

Genus Mesapamea HEINICKE, 1959

Subgenus Mesapamea HEINICKE, 1959

secalis (LINNAEUS, 1758)didyma (ESPER, 1788) (= secalella REMM, 1983)253

. moderata (EVERSMANN, 1843)storai REBEL, 1938 (= acorina PINKER, 1971)

Subgenus Resapamea VARGA & RONKAY, 1992254

hedeni (GRAESER, 1888)

Genus Xylomoia STAUDINGER, 1892255

strix MIKKOLA, 1980graminea (GRAESER, 1889)stangelmaieri MIKKOLA, 1998256

retinax MIKKOLA, 1998257

252 Synonym (FIBIGER & HACKER, 1998).253 M. remmi REZBANYAI-RESER, 1985 (?=insolita REZBANYAI-RESER, 1996): Because the females recorded have strongly deformed

genitalia, we still refrain from accepting these taxa as full species, and hold to the opinion that they are hybrids between M.secalis and M. didyma.

254 S.E. Russia (VARGA & RONKAY, 1992; AHOLA, pers. comm.; KLJUTSCHKO, pers. comm.; LEHMANN, pers. comm.) (FIBIGER &HACKER, 1998).

255 The genus is a true member of Apameini.256 Described from N. Italy/Venezia Giulia, Caorle; figured on plate A: 13.257 One of the paratypes was recorded from S-Ural (Chelyabinsk Distr.) (cf. also K. NUPPONEN & FIBIGER, 2002).

141

Genus Phothedes LEDERER, 1857

captiuncula (TREITSCHKE, 1825)

Genus Eremobia STEPHENS, 1829

ochroleuca ([DENIS & SCHIFFERMÜLLER], 1775) (=asiatica DRAUDT, 1936)258

deccerti HAMPSON, 1908 (=pseudotrachea KRULIKOWSKY, 1909)

Genus Luperina BOISDUVAL, 1829

testacea ([DENIS & SCHIFFERMÜLLER], 1775)kruegeri TURATI, 1912)siegeli (BERIO, 1986)nickerlii (FREYER, 1845)irritaria (A. BANG-HAAS, 1912)259

taurica (KLJUTSCHKO, 1967)rubella (DUPONCHEL, 1835)samnii (SOHN-RETHEL, 1929)dumerilii (DUPONCHEL, 1826)pozzii (CURO, 1883)zollikoferi (FREYER, 1836)

Genus Sidemia STAUDINGER, 1892

spilogramma (RAMBUR, 1871)

Genus Crypsedra WARREN, 1911260

gemmea (TREITSCHKE, 1825)

Genus Amphipoea BILLBERG, 1820

oculea (LINNAEUS, 1761)fucosa (FREYER, 1830)asiatica (BURROWS, 1911)lucens (FREYER, 1845)crinanensis (BURROWS, 1908)

Genus Hydraecia GUENÉE, 1841

micacea (ESPER, [1789]ultima HOLST, 1965nordstroemi HORKE, 1952mongoliensis URBAHN, 1967petasitis DOUBLEDAY, 1847osseola (STAUDINGER, 1882)

Genus Gortyna OCHSENHEIMER, 1816

flavago ([DENIS & SCHIFFERMÜLLER], 1775)

258 Asiatica DRAUDT, 1936 is not a full species but only a subspecies of ochroleuca D. & S. (HACKER, 1998b).259 In Italy subsp. sohnretheli DRAUDT, 1931 (= tiberina (SOHN-RETHEL, 1929).260 It is a true Apameini,-not related to Polymixis (see also HACKER & RONKAY, 1992b).

142

xanthenes (GERMAR, [1842])261 (= franciscae TURATI, 1913)puengeleri (TURATI, 1909)moesiaca HERRICH-SCHÄFFER, 1849borelii (PIERRET, 1837) (= grapholeuca BERIO, 2002, syn. n.)262

cervago EVERSMANN, 1844

Genus Helotropha LEDERER, 1857

leucostigma (HÜBNER, [1808])

Genus Calamia HÜBNER, [1821]

tridens (HUFNAGEL, 1766)

Genus Staurophora REICHENBACH (Leipzig), 1817

celsia (LINNAEUS, 1758)

Genus Argyrospila HERRICH-SCHÄFFER, [1851]

succinea (ESPER, [1798]) (= maculata EVERSMANN, 1842)263

Genus Chortodes TUTT, 1897

minima (HAWORTH, 1809)extrema (HÜBNER, [1809])fluxa (HÜBNER, [1809])pygmina (HAWORTH, 1809)dulcis (OBERTHÜR, 1918)morrisii (MORRIS, 1837) (= subsp. sohn-retheli PÜNGELER, 1907)elymi (TREITSCHKE, 1825)stigmatica (EVERSMANN, 1855)brevilinea (FENN, 1864) (= impudica STAUDINGER, 1892)abrupta (EVERSMANN, 1854) (=contumax PÜNGELER, 1902; = johni PÜNGELER, 1914)

Genus Oria HÜBNER, [1821]

musculosa (HÜBNER, [1808])myodea (RAMBUR, 1858)

Genus Nonagria OCHSENHEIMER, 1816

typhae (THUNBERG, 1784)

Genus Rhizedra WARREN, 1911

lutosa (HÜBNER, [1803])

Genus Celaena STEPHENS, 1829

haworthii (CURTIS, 1829)

261 Year of publication.262 This name was proposed by BERIO (2002) for “G. leucographa BORKH. nec. SCHIFF.“263 The species striata (STAUDINGER, 1897) has never been found authentically in Europe.

143

Genus Phragmatiphila HAMPSON, 1908

nexa (HÜBNER, [1808]) (= insularis TURATI, 1913)264

Genus Archanara WALKER, 1866

geminipuncta (HAWORTH, 1809)neurica (HÜBNER, [1808])dissoluta (TREITSCHKE, 1825)sparganii (ESPER, [1790])algae (ESPER, [1789])

Genus Sedina URBAHN, 1933

buettneri (E. HERING, 1858)

Genus Arenostola HAMPSON, 1910

phragmitidis HÜBNER, [1803]) (= semicana sensu auctt. nec ESPER, [1798])265

Genus Coenobia STEPHENS, 1850

rufa (HAWORTH, 1809)

Subtribus SESAMIINA FIBIGER & GOLDSTEIN, 2005266

Genus Sesamia GUENÉE, 1852

nonagrioides (LEVEBVRE, 1827)cretica LEDERER, 1857

Tribus EPISEMINI GUENÉE, 1852 267 (= CLEOCERINI FORBES & FRANCLEMONT, 1954)

Genus Episema OCHSENHEIMER, 1816

grueneri BOISDUVAL, [1837]glaucina (ESPER, [1789])amasina HAMPSON, 1906268

tersa ([DENIS & SCHIFFERMÜLLER], 1775) 269

gozmanyi RONKAY & HACKER, 1985lederi CHRISTOPH, 1885 (= sareptana ALPHÉRAKY, 1897)korsakovi (CHRISTOPH, 1885)

Genus Cleoceris BOISDUVAL, [1836]

scoriacea (ESPER, [1789])

264 Combination. LERAUT (1997) treats insularis as subspecies of nexa (HÜBNER, [1808]).265 Following KARSHOLT (in: KARSHOLT & RAZOWSKI, 1996) and LERAUT (1997) the valid name of this species should be phragmitidis

(HÜBNER, [1803]. Semicana ESP. is a senior synonym of Aegle vespertalis HBN. (see HACKER, 1998a).266 See FIBIGER and LAFONTAINE, 2005, this volume of Esperiana.267 RONKAY et al. (2001).268 In Europe restricted to Crimea (RONKAY et al., 2001).269 The taxon melanogona (TAUSCHER, 1809) is unsolved; it might be a full species or a synonym of tersa ([DENIS & SCHIFFERMÜLLER],

1775), amasina HAMPSON, 1906, glaucina (ESPER, [1789]), or lederi CHRISTOPH, 1885.

144

Genus Leucochlaena HAMPSON, 1906

Subgenus Leucochlaena HAMPSON, 1906

oditis (HÜBNER, [1822]) (=hispida GEYER, [1832])270

turatii (SCHAWERDA, 1931)seposita TURATI, 1919271

cypraota HAMPSON, 1918 (= muscosa STAUDINGER, 1892)

Subgenus Furcochlaena RONKAY, YELA & HREBLAY, 2001272

fallax (STAUDINGER, 1870)

Genus Ulochlaena LEDERER, 1857

hirta (HÜBNER, [1813])

Genus Eremopola WARREN, 1911

Subgenus Eremopola WARREN, 1911

lenis (STAUDINGER, 1892)

Subgenus Eremochlaena BOURSIN, 1953273

orana (LUCAS, 1894)

Tribus XYLENINI GUENÉE, 1837274

Subtribus XYLENINA GUENÉE, 1837275

Genus Brachylomia HAMPSON, 1906

viminalis