Upload
others
View
8
Download
1
Embed Size (px)
Citation preview
How do ecological and evolutionaryprocesses interact and jointly
influence population dynamics,community structure, and ecosystem
function?
Population,community,ecosystemdynamicsE
Phenotypic expression,ecological interactions
Individualadaptive traits
I
Selective pressureson heritable variation
Eco-evolutionary feedbacks
Looking mathematically at how‘microscopic’ processes operating at
the level of individuals scale up topopulations, communities,
ecosystems
How life history evolution influencespopulation dynamics
Adult / juvenilecompetition Population
dynamicsE
Adult survivaland recruitment
I
Selective pressureson genetic variation
How coevolution affects speciesassociation, coexistence and phenotypes
X YCost ofmutualism/symbiosis,
reward from partner,trait-dependentcompetitionfor rewards
CommunitydynamicsE
Individualinvestment in
mutualism
I
Selective pressureson genetic variation
Individual metabolismAmbienttemperature
Environmentalresources
Ecologicalinteractions
Nichepartitioning
Speciesdiversity
Biochemical Kinetics Hypothesis
Individual metabolismAmbienttemperature
Generationtime
Mutationrate
Speciationrate
Evolutionary extension
‘Universal’ biokineticsprediction fails to capturevariation in large-scale
diversity patterns
• Average organism size is constantacross communities
• Organism size does not evolve
Ambienttemperature
Environmentalresources
Ecologicalinteractions
Morpho-speciesdiversity
Eco-evolutionary dynamics of bodysize along temperature gradients
Geneticmutation
Body size
M
M
M
Evolving trophic networksB
ody
size
Evolutionary time
successful mutation #1 successful mutation #2…
Mechanism: Effect of temperature ontwo-consumer coexistence
Temp.-dep.on resourceproductiononly
Temp.-dep.on consumerrates only
Temp.-dep.on resourceproduction &consumerrates
competitiveexclusion
coexistence by mutual invasibility
coexistence byfacilitation
Global projections: model inputs
• What forms basal resource(evolution not represented)
• Metabolism-mediated temperaturedependence(s): on or off
• Metabolism-mediated temperaturedependence(s): profile
• Communities’ age differential• Environmental geography• Environmental dynamics
Global projections: terrestrial diversity
Also mapping local invasibility how global patterns are affected by dispersal
Stegen, Enquist, Ferriere (Ecol. Lett. 2009)
E
Body Size I Selective pressureson genetic variation
Temperature
OrganismalMetabolism
Trophic structure,size spectrum,
nutrient cycling
Intrinsic mortality,predation risk,
diet &feeding rate
Integrating Temperature- and Size-dependent organismal metabolism into
eco-evolutionary dynamics
Acknowledgments
Mathias Gauduchon, Emily Jones, James Stegen, Dom Carval,Arnaud Pocheville
Judie Bronstein, Don DeAngelis, Fabio Dercole, Brian Enquist,Margaret Evans, Marino Gatto, Richard Law, Hans Metz,David Reznick, Sergio Rinaldi, Joe Travis
Alex Flecker & Lab, Rana El-Sabaawi, Marita DavisonFunding from NSF, ANR, Institut Universitaire de France