7
OBITUARIES Copeia 2011, No. 4, 599–605 Margaret G. Bradbury (1927–2010) Tomio Iwamoto 1 , Gregor M. Cailliet 2 , Daniel M. Cohen 3 , Theodore W. Pietsch 4 , Tom Tucker 5 , Ralph J. Larson 6 , and Marlene L. Martin 7 M ARGARET (‘‘MAGGIE’’) BRADBURY was born in Chicago, Illinois on 15 July 1927 to Margaret and Gerald Bradbury. She had one sibling, a younger brother Gerald, Jr., who had a daughter Jarill Ristine and son Robert. Margaret attended public schools in Chicago and Evanston, Illinois. Her father, an architect, died when she was young, but left her with a great interest in architecture. Jarill Ristine (in litt., 4 Dec. 2010) relates that Margaret’s ‘‘father died when she was probably 11, my dad was 9. He was sent off to boarding school while Maggie lived at home . . . when Aunt Maggie [came] for a visit . . . usually Christmas . . . I was always enchanted. To me she was larger than life, beautiful and exciting to listen to . . . telling wonderful stories about Te Vega and places she had been.’’ In 1947 Margaret entered the field of science as Staff Artist of the Department of Zoology at the Chicago Natural History Museum (now Field Museum of Natural History, FMNH), providing illustrations for (among others) herpetologists Robert F. Inger (1954, 1956) and Karl P. Schmidt, and ichthyologists Robert H. Kanazawa (1952) and Loren P. Woods (Woods and Kanazawa, 1951). The amphibian drawings she did for Inger’s Philippine Amphibia (1954) and the squirrelfish genus Holocentrus for Woods (1955; also Woods and Sonoda, 1973) displayed her early artistic skills. During part of her years at FMNH, she was enrolled at Roosevelt College [now University] in Chicago, at that time a new (opened 1945) college dedicated to equal access to education and to progressive causes. She received her B.S. degree in Zoology in 1955. During the summer of 1955, she collected fishes in the Bahamas on an expedition of the Academy of Natural Sciences, Philadelphia, after which she matriculated as a graduate student at Stanford University (SU). DMC recalls Margaret saying that she entered the SU graduate program as a student of George S. Myers at the urging of FMNH ichthyologists Marion Grey and Loren Woods, and herpetologist Karl P. Schmidt. DMC first met her as a fellow graduate student at SU during one of her Turkish coffee parties—such events perhaps a prelude to a life as a supremely gracious hostess entertaining a constant stream of friends, visitors, and associates. 1 Department of Ichthyology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, California 94118; E-mail: [email protected]. 2 Moss Landing Marine Laboratories, 8272 Moss Landing Road, Moss Landing, California 95039-9647; E-mail: [email protected]. 3 100 Thorndale Drive, Apt. 148, San Rafael, California 94093; E-mail: [email protected]. 4 School of Aquatic and Fishery Sciences, and Burke Museum of Natural History and Culture, University of Washington, Campus Box 355020, Seattle, Washington 98105-5020; E-mail: [email protected]. 5 970 Laurel Avenue, San Mateo, California 94401; E-mail: [email protected]. 6 Department of Biology, 1600 Holloway Avenue, San Francisco State University, San Francisco, California 94132; E-mail: [email protected]. 7 26445 Via Mallorca, Carmel, California 93923; E-mail: [email protected]. F 2011 by the American Society of Ichthyologists and Herpetologists DOI: 10.1643/OT-10-195 Fig. 1. Margaret taking fish photos on board the Te Vega in the Indian Ocean, 1964. (Photograph by Richard Mariscal.)

Margaret G. Bradbury (1927–2010)

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OBITUARIES

Copeia 2011, No. 4, 599–605Margaret G. Bradbury

(1927–2010)

Tomio Iwamoto1, Gregor M. Cailliet2, Daniel M. Cohen3, Theodore W. Pietsch4,Tom Tucker5, Ralph J. Larson6, and Marlene L. Martin7

MARGARET (‘‘MAGGIE’’) BRADBURY was born inChicago, Illinois on 15 July 1927 to Margaret andGerald Bradbury. She had one sibling, a younger

brother Gerald, Jr., who had a daughter Jarill Ristine and sonRobert. Margaret attended public schools in Chicago andEvanston, Illinois. Her father, an architect, died when she wasyoung, but left her with a great interest in architecture. JarillRistine (in litt., 4 Dec. 2010) relates that Margaret’s ‘‘fatherdied when she was probably 11, my dad was 9. He was sent offto boarding school while Maggie lived at home . . . when AuntMaggie [came] for a visit . . . usually Christmas . . . I was alwaysenchanted. To me she was larger than life, beautiful andexciting to listen to . . . telling wonderful stories about Te Vegaand places she had been.’’

In 1947 Margaret entered the field of science as Staff Artist ofthe Department of Zoology at the Chicago Natural HistoryMuseum (now Field Museum of Natural History, FMNH),providing illustrations for (among others) herpetologistsRobert F. Inger (1954, 1956) and Karl P. Schmidt, andichthyologists Robert H. Kanazawa (1952) and Loren P. Woods(Woods and Kanazawa, 1951). The amphibian drawings shedid for Inger’s Philippine Amphibia (1954) and the squirrelfishgenus Holocentrus for Woods (1955; also Woods and Sonoda,1973) displayed her early artistic skills. During part of heryears at FMNH, she was enrolled at Roosevelt College [nowUniversity] in Chicago, at that time a new (opened 1945)college dedicated to equal access to education and toprogressive causes. She received her B.S. degree in Zoology in1955. During the summer of 1955, she collected fishes in theBahamas on an expedition of the Academy of Natural Sciences,Philadelphia, after which she matriculated as a graduatestudent at Stanford University (SU). DMC recalls Margaretsaying that she entered the SU graduate program as a studentof George S. Myers at the urging of FMNH ichthyologistsMarion Grey and Loren Woods, and herpetologist Karl P.Schmidt. DMC first met her as a fellow graduate student at SUduring one of her Turkish coffee parties—such events perhapsa prelude to a life as a supremely gracious hostess entertaininga constant stream of friends, visitors, and associates.

1 Department of Ichthyology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, California 94118; E-mail:[email protected].

2 Moss Landing Marine Laboratories, 8272 Moss Landing Road, Moss Landing, California 95039-9647; E-mail: [email protected] 100 Thorndale Drive, Apt. 148, San Rafael, California 94093; E-mail: [email protected] School of Aquatic and Fishery Sciences, and Burke Museum of Natural History and Culture, University of Washington, Campus Box 355020,

Seattle, Washington 98105-5020; E-mail: [email protected] 970 Laurel Avenue, San Mateo, California 94401; E-mail: [email protected] Department of Biology, 1600 Holloway Avenue, San Francisco State University, San Francisco, California 94132; E-mail: [email protected] 26445 Via Mallorca, Carmel, California 93923; E-mail: [email protected] 2011 by the American Society of Ichthyologists and Herpetologists DOI: 10.1643/OT-10-195

Fig. 1. Margaret taking fish photos on board the Te Vega in the IndianOcean, 1964. (Photograph by Richard Mariscal.)

EARLY YEARS AT STANFORD/HOPKINS MARINE STATION

At Stanford, Margaret confided with DMC that she expectedrather more guidance in her work than Professor Myersgenerally accorded his students. So with some trepidation,she shifted professors (circa 1957) from Myers to Rolf L.Bolin, who held forth at the university’s Hopkins MarineStation in Pacific Grove on the shores of Monterey Bay.Margaret was worried about hurting Myers’ feelings, but herfellow students assured her that Myers had so many studentsthat he really wasn’t bothered about losing one, even a goodone like her. At Hopkins, she faced her new professor mostdays over a noon-hour chess board. Among the faculty atHopkins, Margaret was probably most attached to DonaldP. Abbott, with whom she formed a lifelong friendship,together with his wife Izzie. She built lasting friendshipswith many fellow students who took Bolin’s summer coursein ichthyology and marine ecology, a cohort that becameleaders in marine biology for a generation. ‘‘Stanfordgraduate students in ichthyology and fishery biology spenttime taking courses at the Marine Station . . . and a numberbased their research work at the Station. Large numbers ofStanford undergraduates also attended . . . [as] part of the[then] required curriculum’’ (Brittan, 1997:246).

While on the main campus in Palo Alto between 1955 and1957, Margaret was a teaching assistant in biology and alsoworked as a part-time technical assistant processing SouthPacific fishes at the George Vanderbilt Foundation on the SUcampus. This large collection (about 45,000 lots) was trans-ferred in 1967–68 to the California Academy of Sciences,followed by the SU collections in 1969. During her graduateyears at Hopkins she worked as a technician preparing researchreports for California Cooperative Oceanic Fisheries Investiga-tions at Hopkins (1957); illustrator and illustrating consultant,U.S. Bureau of Commercial Fisheries (now National MarineFisheries Service), Washington, D.C. in 1959 (probably theperiod when she drew the bone figures for the Fishes of theWestern North Atlantic, no. 1, part 3; see Mead and Bradbury,1963); and teaching assistant for marine biology classes atHopkins Marine Station (summers 1961 and 1962). Her Ph.D.in Biological Sciences was received from SU in 1963, but beforethat (1962) she took a position as Assistant Professor of Biologyat MacMurray College, in Jacksonville, Illinois. Her Ph.D.dissertation was a systematic study of the batfish familyOgcocephalidae, a group she continued to study for anotherfour decades, well into her retirement years until her eyesightdeteriorated to the point where she had to give it up.

SAN FRANCISCO STATE UNIVERSITY

In 1963 Margaret returned to California to join the facultyof San Francisco State College (now University, SFSU) asAssistant Professor of Biological Sciences. She becameAssociate Professor in 1967 and full Professor in 1971. Shetaught a wide variety of courses including Ichthyology,Biology of Fishes, Comparative Vertebrate Anatomy, Historyof Biology, Evolution, Introductory Biology for Non-majors,and others. She became a valued member of the biologyfaculty at SFSU, where she had a 31-year career, fully retiringin 1994 after spending eight years in a half-time ‘earlyretirement’ program. RJL, who was a faculty colleague for agood part of Margaret’s career, related that she was active andeffective in service to the department and university at SFSU,serving on the University Promotions Committee for at leastone year and on many departmental committees. One of her

notable contributions came just before her full retirement,when the Biology Department switched from a Zoology/Botany introductory sequence for majors to an integratedone-year, two-semester course. She served on a three-membercommittee that took charge of developing the second-semester course, which concerned evolution, the diversityof life, and ecology. She played a major role in writinglaboratory exercises for this course, where her expertise insystematics and comparative anatomy were very useful.

She mentored and inspired many students, including atleast 13 who received their M.A. degrees under her direction.Margaret’s first Master’s student, Tom Tucker, recalls thatshe had high expectations of her students, took great pridein their academic achievements, and lovingly followed theirchoice of careers. Margaret appointed Tom as her T.A.,which helped support his education; she introduced him tomuseum curation and recommended him to William N.Eschmeyer for a job working to move and integrate theGeorge Vanderbilt Foundation fishes into that of theCalifornia Academy of Sciences. That led to Tom’s subse-quent hire by Earl S. Herald and to his life-time employmentin the Steinhart Aquarium of the Academy. DMC recalls onesummer when he was asked to teach Rolf Bolin’s fish course,with Margaret as T.A. After his first lecture Margaret politelyreminded him on behalf of the students that he had talkednonstop for six hours and the students were desperate for abreak. Students meant so much to Margaret.

Margaret’s concern for students and her sense of socialjustice burst forth during a tumultuous period of studentand faculty protests and strikes at SFSU in 1968–69, whenthe university was headed by an unpopular interimpresident, Dr. S. I. Hayakawa. Margaret was one of thefaculty who went out on strike. When DMC asked her why,she said she didn’t much know about the politics, she justdidn’t want to see any of the kids get hurt. Her support of,and enduring fondness for, her students and her joy atinteracting with them are reflected in many of the tributeswe have received from former students.

TT relates that after he was drafted into the Marines anddeployed overseas, Margaret corresponded with him regu-larly. Stuart G. Poss wrote that he was at the point of

Fig. 2. The consummate hostesses, Margaret Bradbury and PearlSonoda, in their Daly City home, circa 1970. ‘‘Note the teak chest off thedecommissioned Te Vega, the treasured orchids, and the ever-presentSan Francisco Chronicle.’’ (Photograph and quote by Tom Tucker.)

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dropping out of school for lack of money when Margarethelped out by giving him a job as her research assistantstudying batfish feeding. ‘‘I was literally able to get off a dietthat consisted of mostly rice, vegetables and cannedmackerel that I could buy for 35 cents per can.’’ When itcame time to write, revise, and rewrite his thesis, ‘‘she was

always there with suggestions for improvement as she wasthroughout my time at SFSU. She was the source ofencouragement for so many students . . . although shealways had a way of making each feel as though [he or she]was getting all her attention. She really impressed uponme . . . the need for all biologists to learn taxonomy

Fig. 3. Margaret at desk sorting through batfish data, July 2005. (Photograph by Hsuan-Ching Ho.)

Obituaries 601

and demonstrate what she rightfully recognized as amongthe most important contributions that a biologist canmake, providing a better understanding of the biodiversityand richness of the natural world.’’ (Stuart G. Poss, in litt.,30 Nov. 2010)

Margaret obviously loved field work and recognized itsvalue in exposing students to the joys and wonders of ournatural world. She led many class trips collecting up anddown the California coast from Big Sur to Mendocino. Fieldtrips for her ichthyology class were annual rituals and

Fig. 4. Margaret Bradbury and Tom Tucker at Otter Point, August 2009. Lovers Point to the left of Tom’s arm, and Hopkins Marine Station farther inbackground. (Photograph by Tom Tucker.)

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usually included one trip to Half Moon Bay where the entireclass would haul in a large beach seine set in the bay. GMCrecalls Margaret coming to the Moss Landing MarineLaboratories (MLML) area for field trips, including beachseining, otter trawling, and deep-sea trawling. Greg oftentook her ichthyology classes out on the MLML researchvessel and shared data sheets, especially those from MossLanding and Elkhorn Slough field trips, which are now inthe Elkhorn Slough National Estuarine Research Reserve andMonterey Bay National Marine Sanctuary, Sanctuary Inte-grated Monitoring Network. Eric Anderson, who was one ofGreg’s first graduate students, relates that when herichthyology class came to MLML to go out trawling (oncea year), before boarding ‘‘she would invariably crack open abeer to everyone’s surprise, as [she explained] ‘the carbon-ation settles my stomach.’’’ (M. E. Anderson, in litt., 30 Oct.2010). Margaret really liked MLML (which is run by aconsortium of seven CSU campuses, of which SFSU was oneof the founders and home campus of former MLML DirectorJohn Martin), so much so that she served for a long time onthe MLML Governing Board, some of it as Chair of theBoard.

SCIENTIFIC WORK

During the summers of 1963 and 1964, Margaret was able toreturn to her beloved Hopkins Marine Station as an AssistantProfessor of Biology to teach the summer ichthyologycourse. In 1964, she made a memorable 12-week cruise (5October–12 December) across the Indian Ocean on HopkinsMarine Station’s 33-m schooner Te Vega (Fig. 1). The cruisestarted in Mombasa, Kenya, touched on the Amirantes Is.,Seychelles, Maldive Is., Ceylon [Sri Lanka], Nicobar Is.,Phuket, Malaysia, and terminated in Singapore, a distanceof 4,300 nautical miles. The objective of the cruise was to‘‘gather information on the association of organismsoccurring at the depths of the deep scattering layer (DSL)and at levels immediately above and below the DSL.’’(Bradbury et al., 1971:410). The publication resulting fromthis cruise was possibly the most important baselinecontribution to knowledge of the DSL in the Indian Ocean.Margaret made other long cruises on the Te Vega off thecoasts of California and Baja California, and another toEcuador and the Galapagos under the aegis of the StanfordUniversity Oceanographic Program.

TWP summarizes Margaret’s scientific work: Margaret wasbest known among ichthyologists as the world’s authorityon the systematics of the batfishes, family Ogcocephalidae.Her earliest effort relative to this group was a short notepublished in 1962 on the designation of lectotypes ofogcocephalids selected from syntypes in the Museum ofComparative Zoology. But this was soon followed in 1967 by‘‘The Genera of Batfishes,’’ a paper (part of her doctoraldissertation), which, for the first time since the work ofValenciennes (1837), examined the group as a whole,precisely defining the limits of the family and each of thenine genera then recognized, providing a list of species foreach genus (along with information on the whereabouts oftype material), and a phylogenetic analysis. The latterclearly shows that Margaret was on the cutting edge of thelatest developments in biosystematics, applying a ‘cladistic’(Bradbury, 1967:418) approach almost simultaneously withthe 1966 English translation of Willi Hennig’s PhylogeneticSystematics. Following work on batfishes of the WesternCentral Atlantic for FAO Species Identification Sheets

published in 1978, Margaret produced in 1980 an incrediblydetailed and beautifully illustrated revision of the new worldbatfish genus Ogcocephalus in which she recognized 12species, describing five as new to science. Careful attentionwas given to everything, from full annotated synonymiesand statistical analyses of morphometrics, to elaboratedescriptions of external morphology, coloration, and geo-graphic variation and distribution. Equally impressiverevisions of the allied genera Halieutopsis, containingdescriptions of four new species, and Dibranchus, withdescriptions of a new genus and four new species, appearedin 1988 and 1999, respectively. In total, Margaret described16 new species and one new genus. Altogether, Margaret’swork in systematics was exemplary, careful and precise,setting high standards for others to follow. Ted concludes bysaying, those of us who have delved in lophiiformsystematics over the years are especially indebted to her.Grateful acknowledgments have been extended in the formof new species named in her honor: Oneirodes bradburyaeGrey, 1956; Coelophrys bradburyae Endo and Shinohara,1999; and Halieutopsis margaretae Ho and Shao, 2007.

A BEAUTIFUL PERSON, A SUPREME HOSTESS

Most all of Margaret’s close associates can relate stories ofher hospitality, generosity, and warmth. Bill Austin andJohn S. Pearse went to Hopkins in summer 1959 to attendDonald Abbott’s class; there they met Margaret. Bill, whostayed at Hopkins for the rest of his academic career, had anoffice across from Margaret’s in Agassiz Hall, and has ‘‘fondmemories of casual conversations—including her character-istic lilting laugh. We shared an interest in catching a fewrays on the roof of Agassiz when the fog cleared. I rememberhow gracious and kind Margaret was when she offered Dianeand me a place to crash on our visits from Canada. Maggiewas a beautiful person.’’ (Bill Austin, as quoted by J. Pearse,in litt., 28 Oct. 2010). John’s memories are similar: ‘‘WhenVicki and I took the position at UCSC we enjoyed seeingMaggie over the years. We stayed with her and Pearl severaltimes when we visited the Academy . . . she was a closefriend, and as Bill wrote, a beautiful person.’’ (J. Pearse, inlitt., 28 Oct. 2010).

In 1967, Pearl Sonoda, a long-time friend and colleaguefrom the FMNH, left Chicago and the FMNH to becomesenior curatorial assistant at CAS. Pearl moved in withMargaret to a house off Skyline Drive in Daly City, wherethey established themselves as premier hostesses to aconstant stream of visitors (Fig. 2). They later purchased ahouse together near the coast in the Pacific Manor district ofPacifica. GMC recalls Margaret inviting him over in 1972when he arrived at MLML: She introduced me to virtually allthe active ichthyologists in the general area. What a treatthat was! And many long-time friends and colleagues haveresulted from that introduction. James C. Kelley, formerDean of Science and Engineering at SFSU, remarked (in litt.,30 Oct. 2010) that Greg’s story ‘‘was also true for Susan andme. When we first arrived at SFSU, Margaret and Pearl had anice party and invited a group of Marine types for us tomeet.’’ Margaret and Pearl were consummate hostesseswhose warmth and charm made visitors to their home feelcomfortable and welcome—this coupled with good foodand lively conversation left fond memories for everyoneinvolved. Pearl retired in 1996 after 30 years at the Academyand soon after returned to Chicago to live with her sisters.

Obituaries 603

RETIREMENT YEARS

Before and after Margaret fully retired in 1994, she made aconcerted effort to complete a revision of the batfish genusDibranchus (Bradbury, 1999), a paper on two new species ofDibranchus (with John McCosker and Douglas Long, 1999),and an account of the family Ogcocephalidae for the‘‘Annotated Checklist of Fishes’’ series (Bradbury, 2003;Fig 3). RJL noted that Margaret expressed a great deal ofsatisfaction in having completed this work and having beenasked to review other works. During this period she came into the California Academy of Sciences, where she was aFellow and a Research Associate of Ichthyology, on manyoccasions to examine specimens and check literature. Sheinvariably attended the Ichthyology Department’s annualChristmas party and Dave Catania’s fall abalone feast. Afterretirement, Margaret visited often with MLM, wife of JohnMartin, whom Margaret first met in 1959 at Hopkins.Marlene recalls that after John died, Margaret began tocome and stay with her for a few days at each visit. One daywhen MLM came home from teaching at MontereyPeninsula College, Margaret said, ‘‘Today I was walkingaround Pacific Grove, and I suddenly realized that I want tomove down here.’’ (Fig. 4). She had very fond memories ofher student days at Hopkins. Margaret lived across CarmelValley Road from Marlene at Del Mesa Carmel in anapartment that was typical of her—very understated elegant.The elegance, however, did not include the litter boxes forher beloved cats. Margaret became an active member of theBoard of the Friends of Moss Landing Marine Laboratories,serving as President for several years. She was very supportiveof MLML students, both undergrad and graduate, and servedon numerous M.A. and M.S. thesis committees. GMC notesthat at both Del Mesa and later in Pacific Grove, Margarethelped to instruct and counsel other seniors in computer use.She seemed very happy back on the Monterey Peninsuladuring her retirement, meeting new friends and re-establish-ing friendships with old colleagues. After several years at DelMesa, Margaret had experienced a couple of falls and soughtthe longer-term security of a residence that offered nursingcare, so she moved to Canterbury Woods, ‘‘a life careretirement facility in Pacific Grove . . . where other fishytype residents were already living.’’ (David C. Powell, in litt.,7 Dec. 2010). She eventually passed away on 9 October 2010.

Perhaps Margaret’s final contribution to ichthyologycame in the form of a budding graduate student fromTaiwan—Hsuan-Ching (‘‘Hans’’) Ho. From his studentcollecting activities, Hans became interested in studyingdeepsea anglerfishes for his graduate degrees. After a visit toTaiwan by TWP in 2003, Hans decided to focus his doctoralstudies on the batfishes. He first contacted Margaret in 2004and visited her in 2005. During that visit, Hans relatesthat she gave ‘‘many details of each group and reminded methat the depth is an important factor for studying batfishes.She was also very generous to give me all her data sheets[including] . . . data of type specimens . . . and photos (butno drawings).’’ It was his first year of studying batfishes‘‘and these photos and data . . . helped me a lot to befamiliar with most species, and prevent many mistakes.’’Hans revisited her in 2006 and showed her photographs ofTaiwan species and his results from recent visits to museumsaround the world. ‘‘She listened . . . quietly and looked veryexcited with what I had done.’’ The two discussed importanttopics including the taxonomy of Halieutichthys. ‘‘She wasvery humorous . . . [and] . . . also very humble. When I

invited her to review Halieutopsis and other groups together,she declined and said she is no longer working on them andhope[d] [that he could] publish the results.’’ (H.-C. Ho, inlitt. 19 Nov. 2010).

The aura of embracing warmth, comfortable friendliness,and genuine caring interest that emanated from Margaret isabundantly evident from the many stories and anecdotes wehave received from her friends and colleagues. MLMperceptively summarized Margaret’s demeanor as Mid-west-ern warm and yet sophisticated. And, she lived a rich and fulllife on several levels. First and foremost she was a highlysocial person, a trait that carried over into her teaching andher administrative work. She was a talented artist, a careful,dedicated ichthyologist, and a superb educator with greatenthusiasm and concern for students. Her legacy is carried onby the many students who benefited from her guidance,support, encouragement, and mentorship.

ACKNOWLEDGMENTS

We thank the many friends of Margaret who contributed tothis article in various ways; without their input, Margaret’sstory could not have been adequately told, although werealize our shortcomings in writing an obituary of someonewho led such a full and varied life. We thank the followingfor their assistance and words, and apologize to those whosenames have been inadvertently omitted: M. Anderson,K. Andersonnecker, W. Austin, D. Catania, W. Eschmeyer,J. Fong, D. Greenfield, J. Kelley, A. Leviton, J. Ogden, J. andV. Pearse, S. Poss, D. Powell, J. Ristine, M. Rogers, and H. Voris.

LITERATURE CITED

Bradbury, M. G. 1962. Lectotypes of species of Ogcocepha-lidae selected from syntypes in the Museum of Compar-ative Zoology. Breviora 166:1–4.

Bradbury, M. G. 1967. The genera of batfishes (familyOgcocephalidae). Copeia 1967:399–422.

Bradbury, M. G. 1978. Ogcocephalidae, p. 4. unpaged. In:FAO Species Identification Sheets for Fishery Purposes.Western Central Atlantic (Fishing Area 31). Vol. I. Foodand Agriculture Organization of the United Nations,Rome.

Bradbury, M. G. 1980. A revision of the fish genusOgcocephalus, with descriptions of new species from theWestern North Atlantic Ocean (Ogcocephalidae; Lophii-formes). Proceedings of the California Academy ofSciences 42:229–285.

Bradbury, M. G. 1988. Rare fishes of the deep-sea genusHalieutopsis: a review with descriptions of four new species(Lophiiformes: Ogcocephalidae). Fieldiana Zoology (n.s.)44:1–22.

Bradbury, M. G. 1999. A review of the fish genusDibranchus, with descriptions of new species and a newgenus, Solocisquama (Lophiiformes: Ogcocephalidae).Proceedings of the California Academy of Sciences51:259–310.

Bradbury, M. G. 2003. Family Ogcocephalidae Jordan 1895,batfishes. California Academy of Sciences AnnotatedChecklists of Fishes 17:1–17.

Bradbury, M. G., D. P. Abbott, R. V. Bovbjerg, R. N.Mariscal, W. C. Fielding, R. T. Barber, V. B. Pearse, S. J.Proctor, J. C. Ogden, J. P. Wourms, L. R. Taylor, Jr., J. G.Christofferson, J. P. Christofferson, R. M. McPhearson,M. J. Wynne, and P. M. Stromborg, Jr. 1971. Studies on

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the fauna associated with the deep scattering layers in theequatorial Indian Ocean, conducted on R/V Te Vegaduring October and November 1964, p. 409–452. In:Proceedings of an International Symposium on BiologicalSound Scattering in the Ocean, March 31–April 2, 1970.G. B. Farquhar (ed.). Airlie House Conference Center,Warrenton, Virginia.

Bradbury, M. G., J. E. McCosker, and D. J. Long. 1999.Batfishes of the Galapagos Islands with descriptions of twonew species of Dibranchus (Teleostei: Ogcocephalidae).Revue francaise d’aquariologie, herpetologie 25(1998) 3–4:79–88.

Brittan, M. R. 1997. The Stanford school of ichthyology:eighty years (1891–1970) from Jordan (1851–1931) toMyers (1905–1985), p. 233–263. In: Collection Building inIchthyology and Herpetology. T. W. Pietsch and W. D.Anderson, Jr. (eds.). American Society of Ichthyologistsand Herpetologists, Special Publication 3.

Endo, H., and G. Shinohara. 1999. A new batfish,Coelophrys bradburyae (Lophiiformes: Ogcocephalidae)from Japan, with comments on the evolutionary relation-ships of the genus. Ichthyological Research 46:359–365.

Grey, M. 1956. New records of deep sea fishes, including anew species, Oneirodes bradburyae, from the Gulf ofMexico. Copeia 1956:242–246.

Hennig, W. 1966. Phylogenetic Systematics. University ofIllinois Press, Urbana, Illinois.

Ho, H-C., and K-T. Shao. 2007. New species of Halieutopsis(Lophiiformes: Ogcocephalidae), from the western northand eastern central Pacific Ocean. Raffles Bulletin ofZoology, Supplement 4:87–92.

Inger, R. F. 1954. Philippine Zoological Expedition 1946–1947.Systematics and zoogeography of Philippine Amphibia.

Fieldiana: Zoology 33:181–531, 97 figs. [all drawings byBradbury]

Inger, R. F. 1956. Some amphibians from the lowlands of northBorneo. Fieldiana: Zoology 33:389–424. [MGB drawings: fig.85, map; fig. 91 of tadpole of Rhacophorus pardalis pardalis]

Kanazawa, R. H. 1952. More new species and new records offishes from Bermuda. Fieldiana: Zoology 34:71–80. [MGBdrawings: fig. 9, Callechelys bilinearis; fig. 10, Anarchiasyoshiae; fig. 11, Rhegma bermudensis; fig. 12, Gillellus greyae;fig. 13, Microdesmus woodsi]

Mead, G. W., and M. G. Bradbury. 1963. Names of bones.In: Fishes of the Western North Atlantic. H. B. Bigelow (ed.-in-chief). Memoirs of the Sears Foundation for MarineResearch 1(3):20–23.

Valenciennes, A. 1837. Acanthopterygiens a pectoralespediculees, p. 335–507. In: Histoire Naturelle des Poissons12. G. Cuvier and A. Valenciennes (eds.). Levrault, Paris &Strasbourg.

Woods, L. P. 1955. Western Atlantic species of the genusHolocentrus. Fieldiana: Zoology 37:91–119. [MGB draw-ings: fig. 14, heads of Holocentrus coruscus, H. bullisi, and H.vexillarius; fig. 15, Holcentrus marianus; fig. 16, Holocentrusascensionis; fig. 17, Holocentrus rufus; fig. 18, Holocentrusbullisi]

Woods, L. P., and R. H. Kanazawa. 1951. New species andnew records of fishes from Bermuda. Fieldiana: Zoology31:629–644. [MGB drawings of Liopropoma mowbrayi n.sp., Centropyge argi n. sp., and Otophidium schmidti n. sp.]

Woods, L. P., and P. Sonoda. 1973. Order Berycomorphi(Beryciformes), p. 263–396. In: Fishes of the WesternNorth Atlantic. D. M. Cohen (ed.-in-chief). Memoirs of theSears Foundation for Marine Research 1(6):263–396. [MGBdrawings figs. 32, 35, 37, 39; formerly used in Woods, 1955]

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