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Institute of the Biological Problems of the North, Portovaya Str. 18, 685000 Magadan, Russia, e-mail: [email protected]
ABSTRACT: The fine structure of tail appendix of the typical dyplocyst Aploparaksisbulbocirrus Deblock et Rausch, 1968 since first invagination up to stage of maturecysticercoid has been investigated. In composition of exocyst elements of epithelial andmuscle tissues are defined and, besides, few low-differentiated cells of peculiar organiza-tion (“dark cells”) are found. Elements of the connective tissue are not revealed. The exocysthistogenesis is characterized by differences in tegument construction of its outer and innersurfaces, destructive changes of tegument after invagination of scolex, low-differentiallevel of myocytes. The obtained data support the idea of protective and trophic functionsof exocysts.
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KEY WORDS: cysticercoid, diplocyst, ultrastructure, first invagination, scolexogenesis,tail appendix, exocyst.
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Conn D.B., Rocco L.J. 1989. Fine structure of the cellularparenchyma and extracellular matrix of Ophiotaenialoennbergii (Cestoda: Proteocephalidea) // Acta Zoo-logica (Stockholm). Vol.70. P.105–110.
Gabrion C., Gabrion J. 1976. Etude ultrastructurale de lalarvae de Anomotaenia constricta (Cestoda, Cyclo-phyllidea) // Zeitschrift fur Parasitenkunde. Vol.49.P.161–177.
Krasnoshchekov G.P., Moczon T., Pluzhnikov L.T. 1979.Ultrastructure of the cyst of Hymenolepis diminutalarvae // Folia parasitologica. Vol. 6. P.245–251.
Krasnoshchekov G.P., Pluzhnikov L.T. 1980. Differenti-ation and autophagy of tegument of exocyst Parict-erotaenia porosa (Cestoda, Dilepididae) // Helm-inthologia. Vol.17. P. 91–300.
Krasnoshchekov G.P., Pluzhnikov L.T., Gulyaev V.D. 1985.Ultrastructure of the larval organ of metacestode Lat-eriporus geigraphicus Cooper, 1921 (Cestoda: Dilepi-didae) // Folia parasitologica. Vol.32. P.51–59.
Rees G. 1973a. Cysticercoids of three species of Tatria(Cyclophyllidea: Amabiliidae) including T. octacan-tha sp. nov. from the haemocoele of the damsel-flynymphs Pyrrhosoma nimphula, Sulz and Enallagmacyathigerum, Charp. // Parasitology. Vol.66. P.423–446.
Rees G. 1973b. The ultrastructure of the cysticercoid ofTatria octacantha Rees, 1973 (Cyclophyllidea: Am-abiliidae) from the haemocoele of the damsel-fly nymphsPyrrhosoma nimphula, Sulz and Enallagma cyathigerum,Charp. // Parasitology. Vol.67. P.85–103.
Rifkin E., Cheng T.C., Hohl H.R. 1970. The fine structureof the tegument of Tylocephalum metacestodes withemphasis on the new type of microvilli // J. Morph.Vol.130. P.11–24.
Ubelaker J.E., Cooper N.B., Allison V.E. 1970. The finestructure of the cysticercoid of Hymenolepis diminu-ta. I. The outer wall of the capsule // Zeitschrift furParasitenkunde. Vol.34. P.258–270.
Webb R.A. 2005. The organization and fine structure ofthe muscles of the scolex of the cysticercoid of Hyme-nolepis microstoma // Journal of Morphology. Vol.154.P.339–356.