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© ÇÎÎËÎÃÈß ÁÅÑÏÎÇÂÎÍÎ×ÍÛÕ, 2009 Çîîëîãèÿ áåñïîçâîíî÷íûõ, 2009, 6(2): 129145 Ñòðóêòóðà è äèôôåðåíöèðîâêà òêàíåé öèñòèöåðêîèäîâ. 2. Äèôôåðåíöèðîâêà òêàíåé õâîñòîâîãî ïðèäàòêà ó òèïè÷íîé äèïëîöèñòû Aploparaksis bulbocirrus (Cestoda: Hymenolepididae) Â.Ï. Íèêèøèí Èíñòèòóò áèîëîãè÷åñêèõ ïðîáëåì Ñåâåðà ÄÂÎ ÐÀÍ, óë. Ïîðòîâàÿ, 18, Ìàãàäàí 685000 Ðîññèÿ, e-mail: [email protected] ÐÅÇÞÌÅ: Èçó÷åíà òîíêàÿ îðãàíèçàöèÿ ýêçîöèñòû ó ðàçâèâàþùåãîñÿ è çðåëîãî öèñòèöåðêîèäà Aploparaksis bulbocirrus Deblock et Rausch, 1968. Íà ñòàäèè ïåðâîé èíâàãèíàöèè â ñîñòàâå ýêçîöèñòû âûÿâëåíû ýëåìåíòû ïîãðàíè÷íîé è ìûøå÷íîé òêàíåé, à òàêæå íåìíîãî÷èñëåííûå ìàëîäèôôåðåíöèðîâàííûå êëåòêè.  ýêçîöèñòå çðåëîãî öèñòèöåðêîèäà ïîìèìî òåãóìåíòà è ìûøå÷íûõ ýëåìåíòîâ âûÿâëÿþòñÿ «òåìíûå» êëåòêè, ïðåäïîëîæèòåëüíî ðàññìàòðèâàåìûå êàê ìîäèôèêàöèè ìàëîäèô- ôåðåíöèðîâàííûõ êëåòîê. Êëåòêè ñîåäèíèòåëüíûõ òêàíåé íå îáíàðóæåíû. Äëÿ ñôîðìèðîâàííîé ýêçîöèñòû õàðàêòåðíû ðàçëè÷èÿ â ñòðîåíèè òåãóìåíòà åå âíåøíåé è âíóòðåííåé ïîâåðõíîñòåé, äåñòðóêòèâíûå èçìåíåíèÿ òåãóìåíòà ïîñëå èíâàãèíà- öèè ñêîëåêñà, íèçêàÿ ñòåïåíü äèôôåðåíöèðîâêè ìèîáëàñòîâ. Ïîëó÷åííûå äàííûå ñâèäåòåëüñòâóþò î çàùèòíîé è òðîôè÷åñêîé ôóíêöèÿõ ýêçîöèñòû ó èññëåäîâàííûõ òèïè÷íûõ äèïëîöèñò. ÊËÞ×ÅÂÛÅ ÑËÎÂÀ: öèñòèöåðêîèä, òèïè÷íàÿ äèïëîöèñòà, óëüòðàñòðóêòóðà, ïåð- âàÿ èíâàãèíàöèÿ, ñêîëåêñîãåíåç, õâîñòîâîé ïðèäàòîê, ýêçîöèñòà. Structure and differentiation of tissues of cysticercoids. 2. Differentiation of the exocyst in typical diplocyst of Aploparaksis bulbocirrus (Cestoda: Hymenolepididae) V.P. Nikishin Institute of the Biological Problems of the North, Portovaya Str. 18, 685000 Magadan, Russia, e- mail: [email protected] ABSTRACT: The fine structure of tail appendix of the typical dyplocyst Aploparaksis bulbocirrus Deblock et Rausch, 1968 since first invagination up to stage of mature cysticercoid has been investigated. In composition of exocyst elements of epithelial and muscle tissues are defined and, besides, few low-differentiated cells of peculiar organiza- tion (“dark cells”) are found. Elements of the connective tissue are not revealed. The exocyst histogenesis is characterized by differences in tegument construction of its outer and inner surfaces, destructive changes of tegument after invagination of scolex, low-differential level of myocytes. The obtained data support the idea of protective and trophic functions of exocysts.

6 2 129 145 Nikishin - kmkjournals.comkmkjournals.com/upload/PDF/IZ/IZ Vol 06/invert6_2 129_145 Nikishin.pdfmail: [email protected] ABSTRACT: The fine structure of tail appendix of

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Institute of the Biological Problems of the North, Portovaya Str. 18, 685000 Magadan, Russia, e-mail: [email protected]

ABSTRACT: The fine structure of tail appendix of the typical dyplocyst Aploparaksisbulbocirrus Deblock et Rausch, 1968 since first invagination up to stage of maturecysticercoid has been investigated. In composition of exocyst elements of epithelial andmuscle tissues are defined and, besides, few low-differentiated cells of peculiar organiza-tion (“dark cells”) are found. Elements of the connective tissue are not revealed. The exocysthistogenesis is characterized by differences in tegument construction of its outer and innersurfaces, destructive changes of tegument after invagination of scolex, low-differentiallevel of myocytes. The obtained data support the idea of protective and trophic functionsof exocysts.

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KEY WORDS: cysticercoid, diplocyst, ultrastructure, first invagination, scolexogenesis,tail appendix, exocyst.

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