Application of Growth Models in Thinning

Embed Size (px)

Citation preview

  • 7/27/2019 Application of Growth Models in Thinning

    1/60

    APPLICATIONS OF GROWTHMODELS FOR THINNING

    Makrand Gujar

    11-661-003

    I. Ph.D

  • 7/27/2019 Application of Growth Models in Thinning

    2/60

    Thinning

    Thinning is defined as a felling made in an

    immature stand for the purpose of improving

    the growth and form of the trees that remain,after thinning without permanently breaking

    the canopy.

    Thinning consist of series of successive

    felling operations for a number of times

    before crop matures.

    The interval between two successive felling

    may be fixed but it is dependent on time

    required for canopy closure.

    Thinning is carried from sapling stage and is

    continued upto the beginning of the

    regeneration period

    Thinning is applicable to pure and even agedor nearly even aged crop.

  • 7/27/2019 Application of Growth Models in Thinning

    3/60

    Objectives of Thinning

    To improve the hygieniccondition

    To create best condition of

    growth

    Salvage the anticipated lossesof the mercantable volume

    To obtain desirable

    composition of crops

    Improvement in wood quality

    To obtain intermediate yield

    and increase net yield and

    financial out turn

    To reduce risk of pest and

  • 7/27/2019 Application of Growth Models in Thinning

    4/60

    Thinning cycle

    Planned number of years whichelapses between succesive orconsecutive thinning in the samecrop.

    Thinning cycles are not regular

    They are shorter during earlyperiod of growth and longer whenthe crop is middle aged to mature.

    Eg.

    The teak plantation in M.P & Mah.Are thinned at 5th , 10th,20th,30th,45th and 60th year.

    Upto 10 th year, thinning cycle isfive year,from 10th to 30th year it is

    ten year and afterwards it is fifteen

  • 7/27/2019 Application of Growth Models in Thinning

    5/60

    Thinning reduces mortality (or salvages it before it occurs) by reducing the number of trees

    per acre.

    The remaining trees then have more site resources to draw from and typically grow faster

    and healthier.

    By thinning at regular intervals, one can be assured that stress due to overcrowding is

    avoided.

    Thinned trees can then develop stronger root systems and be less prone to windthrow.

    The species composition of a stand can also be influenced by thinning, e.g., depending on

    which tree species are cut and which are retained.

    If sawlogs or veneer logs are sought, thinnings would focus on developing large and high

    quality stems. Thus thinning can improve growing conditions, species composition, tree

    quality, and the economic value of the stand.

    Importantly, poor thinning choices can reduce quality and economic values (e.g., high

    grading or always taking the best trees and leaving the worst).

    However, well planned thinning can provide increases in timber values and economic

    returns. Specific recommendations for thinning are provided in the tree specific guide

    sections.

    ADVANTAGES OF THINNING

  • 7/27/2019 Application of Growth Models in Thinning

    6/60

    Growth Models..

    Models estimating growth and yield of forest stands provideimportant tools for forest management.

    It is useful to distinguish between

    models for prediction, and(intended for management)

    models for understanding(e.g. physiological, process models)

    (Bunnell 1989).

    Typical applications are in

    forecasting for forest planning purposes,

    in the comparison and

    evaluation of silvicultural (pruning, thinning) regimes,and

    in the updating of stand description databases.

    Thinning models comes under models for prediction category

  • 7/27/2019 Application of Growth Models in Thinning

    7/60

    Survival pattern of tree Three distinct survival patterns found during the life of a plantation which must be recognized if the total

    survival pattern is to be modelled.

    The first pattern occurs during the first year after planting.

    Survival during the first year is highly variable and depends on many factors including care of the planting

    stock at the nursery and at the planting site, time of seedling storage, planting crew practices and first year

    climatic factors (such as the amount and distribution of rainfall during the growing season).

    The second easily identifiable survival pattern occurs from year one to some time beyond crown closure.

    Mortality during this period is random and can be attributed to factors other than intra-specific competition

    (although intra-specific competition, i.e. crown closure, may begin during this period). Important factors here

    which may influence survival include levels of inter-specific competition (both woody and herbaceous), stand

    establishment practices and certain stochastic elements such as insect, rodent or disease attacks.

    The third survival pattern occurs from the onset of intra-specific competition induced mortality to rotation.

    During this period, the effects of intra-specific competition are the dominant forces affecting survival (although

    random mortality can still occur). Two important factors which must be considered when modeling survival

    during this period include stand density and site index.

    Amateis et al.,1996

  • 7/27/2019 Application of Growth Models in Thinning

    8/60

    These factors will also affect when competition induced mortality

    begins(obviously, denser stands on better sites will enter this stage of stand

    development sooner). In addition, intermediate silvicultural treatments applied to

    plantations during this period may also affect survival patterns.

    Thinning, in particular, has a direct effect on stand survival because it alters the

    amount and distribution of the growing stock.

    It also changes the overall vigor of the stand by removing smaller, slower growing

    trees (if the thinning is from below) and providing additional growing space for the

    residual stand.

    Therefore, appropriate survival models for this period of intra-specific competition

    should be sensitive to natural changes in stand density due to self-thinning as well

    as artificial changes due to thinnings applied as silvicultural treatments

  • 7/27/2019 Application of Growth Models in Thinning

    9/60

    Growth models used for thinning

    FORECASTis a management-oriented, stand-level, forest-growthand ecosystem-dynamics model. The model was designed to

    accommodate a wide variety ofsilvicultural and harvesting systems

    and natural disturbance events (e.g., fire, wind, insect epidemics) in

    order to compare and contrast their effect on forest productivity,stand dynamics, and a series of biophysical indicators of non-timber

    values.

    COMMIX :A mechanistic model, COMMIX, whichsimulates growth of mixed-species forest stands

    (Bartelink 1998b),

    TADAM

    SILVA

    BWINPRO SORTIE

    http://en.wikipedia.org/w/index.php?title=Stand-level_modelling&action=edit&redlink=1http://en.wikipedia.org/w/index.php?title=Forest_growth_modelling&action=edit&redlink=1http://en.wikipedia.org/wiki/Ecosystemhttp://en.wikipedia.org/wiki/Silviculturehttp://en.wikipedia.org/w/index.php?title=Forest_stand_dynamics&action=edit&redlink=1http://en.wikipedia.org/w/index.php?title=Forest_stand_dynamics&action=edit&redlink=1http://en.wikipedia.org/w/index.php?title=Forest_stand_dynamics&action=edit&redlink=1http://en.wikipedia.org/w/index.php?title=Forest_stand_dynamics&action=edit&redlink=1http://en.wikipedia.org/wiki/Silviculturehttp://en.wikipedia.org/wiki/Ecosystemhttp://en.wikipedia.org/wiki/Ecosystemhttp://en.wikipedia.org/wiki/Ecosystemhttp://en.wikipedia.org/w/index.php?title=Forest_growth_modelling&action=edit&redlink=1http://en.wikipedia.org/w/index.php?title=Forest_growth_modelling&action=edit&redlink=1http://en.wikipedia.org/w/index.php?title=Forest_growth_modelling&action=edit&redlink=1http://en.wikipedia.org/w/index.php?title=Stand-level_modelling&action=edit&redlink=1http://en.wikipedia.org/w/index.php?title=Stand-level_modelling&action=edit&redlink=1http://en.wikipedia.org/w/index.php?title=Stand-level_modelling&action=edit&redlink=1
  • 7/27/2019 Application of Growth Models in Thinning

    10/60

    Thinning effects

    Thinning causes reduction of density and associatedreduction in resource use and competition, whichincrease the growth of the remaining trees and reducestheir mortality rate.

    On the stand scale this effect can be divided into twoeffects.

    First, the total stand production (NPP) is reduced(although very slightly for light thinning) because of thereduced resource capture (Zeide, 2004).

    Second, the self-thinning (density dependent) mortalityis reduced, which is linked to the improved growth of theremaining trees.

  • 7/27/2019 Application of Growth Models in Thinning

    11/60

    3/2 Law of Self-Thinning

    The most famous and most controversial of hypotheses

    is the

    3/2 Law of Self-Thinning(Hutchings 1983, Weller 1987,

    Lonsdale 1990), which says that

    stands of plants reach a limiting density such that the

    average plant mass w =W/N is proportionalto N3/2, i.e.,

  • 7/27/2019 Application of Growth Models in Thinning

    12/60

    CASE STUDIES

  • 7/27/2019 Application of Growth Models in Thinning

    13/60

    Effects of stand composition and thinning in mixed-

    species forests: a modelling approach applied to

    Douglas-fir and beech

    A mechanistic model, COMMIX, which simulates growth of mixed-speciesforest stands (Bartelink 1998b), was used to estimate the response ofmixed-forest growth and development to specific silvicultural treatmentsand stand compositions.

    COMMIX(COMpetition in MIXed stands)

    The model was also used to analyze effects of thinning regimes and stand

    composition on productivity.

    To investigate model performance, simulation estimates were comparedwith field data.(yield tables) (Jansen et al. 1996).

    Thinning regimes included in the yield tables (in terms of basal area to beremoved) were applied in the model runs.

    In the present study, COMMIX was used to investigate the effects of standcomposition and thinning regime on growth and yield.

    In this study, thinning from above (crown thinning or high thinning) wasapplied: to benefit the best trees, the strongest competitors are removed,H. H. BARTELINK,2000

  • 7/27/2019 Application of Growth Models in Thinning

    14/60

    The relative yields of Douglas fir and beech were defined

    as:

    RYD =YDmix / YDmono

    RYB =YBmix / YBmono

    where YDmixand YBmix are the yields of Douglas-fir and beech in

    the mixture (m3 ha1), and

    YDmono and YBmono are the yields in monoculture (m3 ha1),

    The relative total yield was defined as:

    RYT = RYD + RYB.

  • 7/27/2019 Application of Growth Models in Thinning

    15/60

    Two thinning regimes were also defined.

    In Thinning Regime I, thinning intensity (expressed as a fraction of the standing basal area) was

    the same as used in the yield table (the default thinning regime). High thinning was the

    method of thinning applied.

    Beech is generally believed to be at a disadvantage in this mixture type,

    Thinning Regime II was defined so that the thinning intensity in beech was fixed at only 5% of

    the standing basal area, whereas for Douglas-fir the thinning fraction increased from 1.0 times

    the default fraction (i.e., the yield table values) at Age 20, to 1.5 times the default fraction at Age

    70.

    Simulated stands comprised 400 (20 20) trees.

    Initial planting distance was 2.5 m, based on Douglas-fir stand densities.

    The model was set to simulate 50 years of stand growth, from age 20 to 70.

  • 7/27/2019 Application of Growth Models in Thinning

    16/60

    Table 2. Tree components distinguished in the COMMIX

    model and the initial values of the variables. Biomass

    amounts are dry weights.

    Foliage area is one-sided (i.e., projected area). Tree

    diameter (dbh) is measured at breast height (1.30 m

    above the forest floor).

    Main species-specific parameters used COMMIX

    (after Bartelink 1998b). In addition, empiricalparameters are applied to describe the allometry

    between tree diameter at breast height (dbh; 1.30 m

    above the forest floor), dbh and branch biomass,

    dbh and coarse root biomass, dbh and crown

    dimensions, and dbh, height and bole volume.

  • 7/27/2019 Application of Growth Models in Thinning

    17/60

    RESULTS

  • 7/27/2019 Application of Growth Models in Thinning

    18/60

    Figure 1. Simulated LAI development over time for stands with

    different shares of Douglas-fir and beech, for Thinning Regime

    I. Abbreviations: %dg indicates percentage of Douglas-fir.

    Figure 2. Simulated development over time of the total stand

    biomass for stands with different proportions of Douglas-fir and

    beech, for Thinning Regime I. Abbreviations: %dg indicates

    percentage of Douglas-fir.

    Figures 1 to 3 show only the results obtained under Thinning Regime I, because relatively little variation between the two

    thinning regimes were apparent in the stand-level results.

    Figure 3. Simulated development over time of the root/shoot ratio for

    stands with different shares of Douglas-fir and beech, for Thinning

    Regime I. Abbreviations: %dg indicates percentage of Douglas-fir.

    Figure 4. Simulated stand volume increment for stands with different

    proportions of Douglas-fir and beech, for Thinning Regimes I (a) and

    II (b). Abbreviations: % indicates percentage of Douglas-fir.

    Th l ti i ld (RYB RYD d

  • 7/27/2019 Application of Growth Models in Thinning

    19/60

    The relative yields (RYB, RYD, and

    RYT) of beech and Douglas-fir in the

    two scenarios.

    The yield of Douglas-fir in the mixedstand is higher than the yield of a

    monospecific Douglas-fir stand.

    However, when thinning intensities in

    beech are lower than the default-thinning regime (Figure 5b), the RY of

    Douglas-fir in mixtures with a low

    proportion of Douglas-fir is less than 1.0,

    and the RYT exceeds 1.0. Figure 5. Estimated relative yield (RY) for the stands, based onthe stem volume production over a 50-year period, for Thinning

    Regimes I (a) and II (b).

    The lower-left to upper-right solid line represents the yield of the

    Douglas-fir trees in the mixture (RYD), the upper-left to lower-rightsolid line represents the beech yield (RYB),

    the horizontal solid line is the (mixed) stand total yield (RYT).

    The dashed lines show the expected yield if intra- and inter-

    specific interactions were equivalent, i.e., no advantage of mixing

    would exist.

    The position of the individual stands on thex-axis depends on

    their proportion (in terms of basal area) of Douglas-fir.

  • 7/27/2019 Application of Growth Models in Thinning

    20/60

    Figure 7. Relative diameter frequencydistributions of beech (open) and Douglas-

    fir(solid), for Thinning Regime I. Relative

    diameter frequency distributions after 50 years

    of growth (Age 70), for the five initial stand

    conditions (ae). The x-axis shows the dbh

    class(cm), and the y-axis shows the relative

    frequency in the stand.

    Figure 8. Relative diameter frequencydistribution of beech (open) and Douglas-fir

    (solid), for Thinning Regime II. Relative

    diameter frequency distributions after 50 years

    of growth (Age 70), for the five initial

    stand conditions (ae). The x-axis shows the

    dbh class (cm), and the y-axis shows the relative

    frequency in the stand.

  • 7/27/2019 Application of Growth Models in Thinning

    21/60

    The model was validated with field data from yield tables.

    For douglas fir, For Douglas-fir, the yield table data conformed well to the

    results of simulated low thinning in the first decades. However, there were differences between table and model estimates in

    older stands.

    The higher estimates with the model compared with the yield table in older

    stands are in accordance with the findings of Schoonderwoerd and Daamen

    (1995),who reported that the yield table underestimates growth rates instands older than approximately 40 years.

    The simulated beech stand initially grew faster than described by the yield

    table, but slowed down with increasing stand age.

    Simulated stand productivity decreased as stand age increased, which couldbe attributed to the opening up of the canopy.

    The thinning intensity indicated by the yield table is large compared with the

    estimated basal area increment

    Conclusion

  • 7/27/2019 Application of Growth Models in Thinning

    22/60

    Comparing Thinning Regimes I and II reveals that,

    For Douglas-fir, increased thinning intensity of Douglas-fir(Thinning Regime II) results in

    (1) larger diameters in stands with a low share of beech;

    (2) smaller diameters in mixtures with a large share of

    beech, compared with the stand structures arrived at inThinning Regime I.

    For beech, the reduced thinning intensities in both beech and

    Douglas-fir (Thinning Regime II) lead to a higher stem

    number and consequently, a lower mean dbh and asmaller dbh range.

  • 7/27/2019 Application of Growth Models in Thinning

    23/60

    TAUYIELD: A STAND-LEVEL GROWTH AND YIELD

    MODEL FOR THINNED AND UNTHINNED LOBLOLLY

    PINE PLANTATIONS

    TAUYIELD can be used for a variety of purposes including inventory

    updating, evaluating thinning as a silvicultural alternative and as input to

    management decision-making.

    The light-thin and heavy-thin plots used in developing TAUYIELD

    received primarily selection thinnings from below with a few plots first

    receiving a row thinning to provide access followed by a selection

    thinning.

    Trees were selected for removal based on size, vigor, quality and

    spacing. All plots were thinned once and allowed to grow for twelve years.

    Amateis et al.,1996

  • 7/27/2019 Application of Growth Models in Thinning

    24/60

    Basal area and survival for each of four thinning treatments at plot

    establishment, five years, ten years and eighteen years.

  • 7/27/2019 Application of Growth Models in Thinning

    25/60

    The thinning response function developed by Liu et al. (1995)

    was incorporated into each dynamic equation of the system. The

    general form of the function is:

    where: T = thinning response

    Ga, Gb = stand basal area after

    and before thinning, respectively

    A = stand age

    TA = age of thinning

    r, k = parameters to be estimated

  • 7/27/2019 Application of Growth Models in Thinning

    26/60

    Equation (1) has certain desirable biological properties.

    The first is that when no thinning has occurred, the before to after thinning basal arearatio is 1 which means T has no effect on the equation of which it is a part.

    Second, at time of thinning the response is also conditioned to be 1 which means

    there is no immediate response at the time of thinning.

    Third, response to thinning begins at zero and, depending on the magnitudes andsigns of r and k, affects the equation into which it has been incorporated to an

    increasing degree up to some maximum effect and then diminishes over time

    .

    The duration of thinning response (in years) is determined by the value of the duration

    parameter, k.

    The rate parameter, r, is dimensionless and along with G , G , A and TA defines the

    shape of the response function.

  • 7/27/2019 Application of Growth Models in Thinning

    27/60

    In order to project stand basal area, it is necessary to provide TAUYIELD

    with an initial basal area. When one is not available, the following basal area

    prediction equations can be used:

    where: N = number of trees removed in the thinning operation tN = number of trees remaining after thinning a

    G = basal area removed in the thinning operation t

    b - b = parameters to be estimated

    BASAL AREA PREDICTION EQUATIONS

  • 7/27/2019 Application of Growth Models in Thinning

    28/60

  • 7/27/2019 Application of Growth Models in Thinning

    29/60

    Total yield prediction

    In order to predict total cubic foot volume, a multiple linear

    regression equation was formulated:

    where: b - b = parameters to be estimated

  • 7/27/2019 Application of Growth Models in Thinning

    30/60

    Thinned-unthinned

    Figures 11a 11d compare some basic stand development relationships

    for the average unthinned, light thinned and heavy-thinned stand in theregion-wide plantation data set.

    At plot establishment, the average stand conditions were age 15, siteindex 60, 566 trees per acre and 110 square feet per acre of basal area.

    Following thinning, the average light-thinned conditions were 315 treesper acre and 80 square feet per acre of basal area. For the averageheavy-thinned stand, the mean residual number of trees per acre was238 and the mean residual basal area was 65 square feet per acre.

    These figures present projections to age 35 which, for the average stand,is 20 years following thinning.

    (1) Unthinned control plot,

    (2) Lightly thinned plot from

    which approximately

    one-third of the basal

    area was removed, and,

    (3) Heavily thinned plot from

    which approximately

    one-half of the basalarea was removed.

    Comparison of TAUYIELD projected growth and yield data for unthinned

  • 7/27/2019 Application of Growth Models in Thinning

    31/60

    MIC Stand Age

    10 15 20 25 30

    Ft3/ac

    MIC 1:

    traditional309 1,121 2,004 2,716 3,158

    MIC 2:

    genetics396 1,353 2,355 3,135 3,605

    MIC 3:

    MIC2 + F396 1,353 2,637 3,433 3,912

    MIC 4: MIC

    3 + H518 1,670 3,139 4,033 4,502

    MIC 5: MIC

    4 + 2nd

    F&H

    641 2,170 3,645 4,587 5,057

    SRTS-FIA 568 1,138 1,708 2,361 3,013

    1993 RPA 310 1,136 1,892 2,382 2,824

    1997 FIA

    Georgia

    Survey

    420 912 1,540 1,969 2,625

    (Ft3/ac to a 4 in.

    diameter outside

    bark top)

    MIC=management

    intensity class;F=fertilization;

    H=herbicide

    application.

    TAUYIELD assumes

    SI 60 at base age 25

    and planting densityis 600 trees per ac;

    SRTS-FIA, 1993

    RPA, and 1997 FIA

    Georgia survey data

    p p j g y

    MICs with FIA data and modeling assumptions. Merchantable wood

    volume

    Thinning Response and Thinning Bias

  • 7/27/2019 Application of Growth Models in Thinning

    32/60

    Thinning Response and Thinning Bias

    in a Young Scots Pine Stand The study analyses the annual post-thinning response and thinning bias of a young Scots pine

    stand as a function of tree size, competition faced by the tree, and competition that is removed

    around the tree in the thinning treatment.

    The thinning response of a tree was defined as the change of tree growth due to a thinning

    treatment. The thinning bias was defined as the difference between the true growth and model

    prediction.

    A distance-dependent (spatial) and a distance-independent (non-spatial) growth model were

    used in the calculations.

    The distance-dependent growth model was (Miina and Pukkala 2000; Equation 8):

    (id)0.5 = 2.7432 3.0645/(d + 5) + 0.3332d/(age + 5) 0.3115ln(age) 0.1160ln(G) 0.0411CIp

    0.0358CIs

    where id is the future 5-year increment in overbarkdiameter (cm),

    d is the tree diameter (includingbark) at breast height (cm), age is the tree age at breast

    height (years),

    G is the stand basalarea (including bark) (m2ha1),

    CIp and CIs are the competition indices computed from pine and spruce competitors, resp.Timo Pukkala, Jari Miina and Marc Palah,2002

  • 7/27/2019 Application of Growth Models in Thinning

    33/60

    The distance-independent growth model was (Nyyssnen and Mielikinen 1978;

    Equation 4):

    ln(pd) = 5.4625 0.6675ln(T) 0.4758ln(G) + 0.1773ln(D)

    0.9442ln(Hdom) 0.3631ln(d) + 0.7762ln(h) wherepd is the future annual increment in overbark

    diameter growth in the next 5-year period, as a compound interest percentage

    (%),

    T is the stand age (years),

    G is the stand basal area (including bark) (m2ha1), D is the diameter(including bark) of the median basal area tree (cm),

    Hdom is the dominant height (m),

    d is the tree diameter (including bark) at breast height (cm) and

    h is the tree height (m)

    The empirical data were measured from a thinning experiment consisting of ten plots,

    each 40 30 m in size, which were thinned to different stand densities.

    The ten-year post-thinning growth of every remaining tree was measured.

  • 7/27/2019 Application of Growth Models in Thinning

    34/60

    Table . Stand characteristics of plots after thinning in 1986/87 and in 1997.

    N = number of stems per ha,

    G before and G = stand basal area before and after thinning, respectively,

    D = mean diameter at breast height,

    H = mean height, T = mean breast height age, and g = weighted by tree basal area.

  • 7/27/2019 Application of Growth Models in Thinning

    35/60

    Where Cij is competition index for tree j

    nj is the number of neighbour nearer than 6m

    ij is vertical angle defined by the predicted height of the tree j

    The following competition index was computed for every

    tree :

  • 7/27/2019 Application of Growth Models in Thinning

    36/60

    The annual thinning response in

    different classes of (A) diameter, (B)

    retained competition and (C)

    harvested competition Diameter

    growths are predicted with thedistance-dependent model.

    The annual thinning response in

    different classes of (A) diameter, (B)

    retained competition and (C)

    harvested competition. Diameter

    growths are predicted with thedistance-independent model.

  • 7/27/2019 Application of Growth Models in Thinning

    37/60

    Conclusion

    The results indicated that the highest thinning response is among medium-sized

    and co-dominant trees.

    The thinning response is quite small, and even negative for some trees, for two

    years after thinning but it becomes clearly positive from the third year onwards.

    The spatial model underestimated the growth of small trees (which usually face

    high competition) while the non-spatial model overestimated the growth of treesthat are small or face much competition.

    The spatial model used in this study overemphasized the effect of competition while

    the non-spatial model underestimated this effect. Both growth models

    overestimated the growth of trees in heavily thinned places, but this bias

    disappeared in two years.

    The negative bias was more pronounced with a spatial growth model because the

    tendency of the non-spatial model to underestimate the growth of trees facing little

    competition partly compensated for the negative bias.

  • 7/27/2019 Application of Growth Models in Thinning

    38/60

    A generic model of thinning and stand density effects on forest

    growth, mortality and net increment

    The model is based on the relationship between the self-

    thinning limit, i.e. the maximal number of trees that can

    coexist on a fixed area (Nmax; cf. symbols in Tab. I) and

    the mean tree size (b).

    To model the number of trees and the effects of thechanges in forest stand density during the development

    of a stand, a self-thinning equation is used.

    For stands growing at maximum density (closed stands)

    b = k Nmax

    Where, Nmax is number of trees and b is the biomass of

    the average tree

    (Oskar et al.,2009)

  • 7/27/2019 Application of Growth Models in Thinning

    39/60

    Closure (c) after a thinning. The figure shows thinning removal

    ofN0 Nt trees, which are smaller than the average tree (thinning

    from below). This leads to an increase in mean tree size (b)

    from b0 before thinning to bt after the thinning. c after the thinning is

    given by N after thinning (Nt) divided by the Nmax (solid line) corresponding

    to bt (Nmax t). The dashed line shows the development after

    thinning.

  • 7/27/2019 Application of Growth Models in Thinning

    40/60

    The response of growth rate (ug) and mortality (um) tostand closure (c). Dotted straight lines show the responses

    immediatelyafter thinning before the trees have utilized the

    new growing space available. The solid lines show the

    acclimated responses and the dashed lines show the effect

    of a thinning to c = 0.5 followed byacclimation.

  • 7/27/2019 Application of Growth Models in Thinning

    41/60

    All values are means for each species. 1 Number of observations, 2mean stand age during experiment, 3 duration over which mean

    responses were

    estimated, 4 adjusted value, measured value was 1.27

    q is mean tree size,

  • 7/27/2019 Application of Growth Models in Thinning

    42/60

    Examples of the development of un-

    thinned (solid line) and a thinned stand(dashed line) simulated with a forest

    growth model for un-thinned stands

    (Franklin et al., 2009) combined with the

    thinning scenario presented here, including

    the effect of competition

    un-related mortality (Eq. (7)).

    Thinnings that reduce closure (c) to

    c = 0.6 are triggered when c > 0.85. The

    stand is Scots pine planted

    at 3000 trees per ha. Net increment (C) isgross biomass growth

    mortality.

  • 7/27/2019 Application of Growth Models in Thinning

    43/60

    In conclusion, introducing a generic thinning framework

    as presented in this study in large scale scenario

    analyses of forest resource development could

    significantly increase their realism with regard to the

    silvicultural decision space in forest management.

  • 7/27/2019 Application of Growth Models in Thinning

    44/60

    Thinning Impacts on Even-aged

    Stands

    of Eucalyptus in Brazil

    E i t l D i

  • 7/27/2019 Application of Growth Models in Thinning

    45/60

    Experimental Design

    -Replicated randomized complete block with repeated measures

    -6 blocks (two in each installation), each one involving two repetitions;

    - 4 treatments, corresponding to different basal area percentages removed in each thinning :

    Treatment 1: 20% without pruning;

    Treatment 2: 35% without pruning;

    Treatment 3: 50% without pruning;

    Treatment 4: 35% with pruning up to 6.0 meters;

    - Each block contained 8 permanent rectangular plots, with an area of 2,600 m2, totaling 48

    plots (6 blocks x 2 repetitions x 4 treatments)

    Eff t i di i t f t t l h i ht

  • 7/27/2019 Application of Growth Models in Thinning

    46/60

    Effect on periodic increment of average total height

    A B C

    A: only thinning 35% and thinning 35% + pruning were equal

    B and C : Only thinning 20% was different from the other treatments

    There is thinning effect

    Effect on periodic increment of basal area per

  • 7/27/2019 Application of Growth Models in Thinning

    47/60

    ect o pe od c c e e t o basa a ea pe

    hectare

    A B C

    A, B and C: only thinning 35% and thinning 35% + pruning were equal

    There is thinning effect

    Eff t i di i t f l h t

  • 7/27/2019 Application of Growth Models in Thinning

    48/60

    Effect on periodic increment of volume per hectare

    A B C

    A, B and C: only thinning 35% and thinning 35% + pruning were equal

    There is thinning effect

    Eff t i di i t f l t

  • 7/27/2019 Application of Growth Models in Thinning

    49/60

    Effect on periodic increment ofvolume per tree

    A

    A, B and C: only thinning 35% and thinning 35% + pruning were equal

    There is thinning effect

    B C

    Conclusion

  • 7/27/2019 Application of Growth Models in Thinning

    50/60

    Conclusion

    Thinning affected the growth of total height, diameter, basal area per hectare, total

    volume per tree and total volume per hectare, but did not affect the growth of

    dominant height

    Thinning prevented regular tree mortality

    Prunning did not affect the growth trend of the variables analyzed

  • 7/27/2019 Application of Growth Models in Thinning

    51/60

  • 7/27/2019 Application of Growth Models in Thinning

    52/60

    THANK YOU

    Using a Density Management Diagram to Develop

  • 7/27/2019 Application of Growth Models in Thinning

    53/60

    Using a Density-Management Diagram to Develop

    Thinning

    Schedules for Loblolly Pine Plantations

    A density-management diagram is a stocking chart

    based on natural stand development expressed as

    changes in average tree size and trees per acre

    overtime.

    (Dean and Baldwin, 1993)

  • 7/27/2019 Application of Growth Models in Thinning

    54/60

    Thinning s equence for a hypoth etical s tandplot ted on

    the lob lo l lyp ine dens i ty -manugement

    diagram. Dotted l ines denote upp er and low er

    grow ing stock l imi ts for th isexample.

  • 7/27/2019 Application of Growth Models in Thinning

    55/60

    Thinning is an important means to pursue silvicultural

    objectives(e.g. selection of desired tree species,

    promotion of stability and stem quality) that generates

    income opportunities during the long rotation periods in

    temperate and boreal forest ecosystems. As acentrepiece in stand level forest management, thinning

    has received considerable attention in forest research

  • 7/27/2019 Application of Growth Models in Thinning

    56/60

    WHY THINNING?

    Thinning removes surplus trees and concentrates the

    potential wood production of the stand on the selected crop

    trees. By thinning your plantation, you are essentially

    salvaging trees which would eventually die naturally.However, by thinning, you can control which trees survive.

    When the average height of the trees in the forest plantation

    is greater than 10 feet, it is time to ask a professional forester

    about thinning.

    Crown closure, or when the crowns of the trees in the

    plantation begin to crowd each other, is another clue that the

    plantation requires thinning.

    Modeling Diameter Growth and Self-Thinning

  • 7/27/2019 Application of Growth Models in Thinning

    57/60

    g g

    in Planted Sugi

    (Cryptomeria japonica) Stands

    The objectives of this study were to analyze diameter

    growth in relation to natural thinning in high-density

    stands in even-aged, pure plantation forests and to

    develop a growth prediction system based on Japanese

    permanent plot data. the diameter growth rate was formalized as a function of

    DBH, stand density and stand age, using parameters

    derived from full density curves

    Tohru Nakajima*,1, Mitsuo Matsumoto2

    and Norihiko Shiraishi3

    ON STAND DENSITY MANAGEMENT

  • 7/27/2019 Application of Growth Models in Thinning

    58/60

    ON STAND DENSITY MANAGEMENT

    DIAGRAM

  • 7/27/2019 Application of Growth Models in Thinning

    59/60

    Figure 6. Estimated absolute yield of the stands over a 50-

    year period for Thinning Regimes I (a), and II (b).

    The dashed lines show the expected yield if intra- and inter-

    specific interactions were equivalent; i.e., no advantage of

    mixing would exist.

    Solid lines represent the model estimates: deviations from

    the dashed lines indicate that interaction between thespecies occurs.

  • 7/27/2019 Application of Growth Models in Thinning

    60/60

    CROWN COMPETITION FACTOR Foresters base the rules for thinning plantations on

    the crown competition factor (CCF), a measure ofcompetition that integrates tree size and the

    number of trees per acre.

    A plantation with a CCF = 100 indicates that thesum of the tree crown areas equals the area of theplantation.

    A CCF less than 100 indicates an understockedplantation; a CCF greater than 100 indicatescrowding.

    Determine the CCF of plantation by measuring anytwo of the following:

    the average tree size in the plantation, the basalarea of the plantation, and the number of trees per

    acre.

    The number of trees per acre to thin from theplantation will depend on the CCF of the stand andthe upper and lower CCF levels between whichplantation stocking will be maintained.