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12/2/2014 1 Ecology Ecology is the study of life in the environment, as well as how all factors of the living and non-living world interact Earth is an elaborate, interconnected system Because of this, we are concerned with the health of all parts of the world Diversity Much of ecology focuses on Earth’s biodiversity Biodiversity: the sum total of all life on Earth Biodiversity has both intrinsic and extrinsic value and is worth conserving Used to learn more about ourselves and our world Potential source of resources Intrinsic beauty/happiness Global Declines in Biodiversity Currently in sixth mass extinction Wild and domesticated species Ecosystem functions down Organization of Life Habitat: wherever an organism lives To keep systems organized, we describe life on Earth in a string of hierarchies: Species Population Community Ecosystem Biome Biosphere Every organism has a niche, or a role in the environment

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Page 1: ecologylecture2

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Ecology Ecology is the study of life in

the environment, as well as how all factors of the living and non-living world interact

Earth is an elaborate, interconnected system

Because of this, we are concerned with the health of all parts of the world

Diversity Much of ecology focuses on Earth’s biodiversity

Biodiversity: the sum total of all life on Earth

Biodiversity has both intrinsic and extrinsic value and is worth conserving

Used to learn more about ourselves and our world

Potential source of resources

Intrinsic beauty/happiness

Global Declines in Biodiversity Currently in sixth mass extinction

Wild and domesticated species

Ecosystem functions down

Organization of Life Habitat: wherever an organism lives

To keep systems organized, we describe life on Earth in a string of hierarchies:

Species Population Community Ecosystem Biome Biosphere

Every organism has a niche, or a role in the environment

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Biomes Large zones of land or aquatic habitat with similar

climate, organisms, and soils

Caused by climate bands, wind currents, and cycling of the atmosphere

Six major terrestrial biomes: Tundra, taiga, temperate deciduous forest, grassland, tropical rainforest, desert

Some biomes are more fragile than others (tundra, TRF)

Some biomes are more diverse

Energy Flow Energy comes from an initial

source (a producer or autotroph) and flows to organisms that cannot produce their own energy (a consumer or heterotroph)

Most energy on Earth from the process of photosynthesis

Energy supplied to animals by the process of cellular respiration

Energy Flow Energy efficiency depends

on how it is transferred and where it comes from

Energy tiered in “trophic levels” and often arranged in pyramids

“Rule of 10” states than on average, only about 10% of energy transfers from one level to the next

Energy Flow Energy efficiencies

depend also on procurement and transfer:

Herbivores more efficient procurers of food but get less energy from what they consume

Predators get more energy from their food, but have to invest more energy into obtaining it

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Food Chains Show order of energy

transfer Simple link of one

organism to the next Arrows show direction of

energy flow: point from what is being eaten/consumed to what is doing the consuming

All chains start with the sun Each day 1019 kcal of solar

energy = 100 million atomic bombs

Levels of Feeding Producers

Primary consumers

Secondary consumers

Tertiary consumers

Quaternary consumers

Detritivores: feed on decomposing matter, often at all or many levels of the food chain

Food Webs More accurate representation

of energy transfer

As ecosystems are interactive, food chains aren’t always as simple as they seem

Example: Alligator can be a top predator OR consumed by a secondary consumer

Organismal Interactions Organisms in an ecosystem interact in dynamic and

diverse ways

Primarily driven by competition

Competition is what ultimately drives evolution

Resource Partitioning Competition over resources is what leads to

partitioning

Rather than compete, organisms segregate into different niches

Niches are determined based on partitioning

Can lead to adaptive radiation in extreme instancesevolution!!

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Organismal Interactions Symbiosis: organisms working together or closely to

survive

More to come on bees…

Mutualism: both organisms benefit

Commensalism: one benefits other is neither harmed nor helped

Parasitism: one organism benefits, the other is harmed

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Crypsis and Mimicry Crypsis: an organism’s ability to hide or blend in

Highly evolved survival strategy that helps an organism avoid predation

Mimicry: organisms that resemble other, more dangerous organisms or inanimate objects

Typically an example of coevolution

Productivity Solar (or geothermal) energy converted to chemical

energy = Gross Primary Productivity

Energy available for the next trophic level after respiration, etc. is subtracted= Net Primary Productivity

This will tie in to photosynthesis later… don’t forget!

PP varies depending on Amount of CO2

Amount of light

Minerals available

Amount of water

Temperature – roughly doubles each 10oC

All of the above are also photosynthetic limiting factors

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Availability of Resources and Competition Leads to Specific Population Interactions!

Populations Every ecosystem has a limited amount of energy and

resources, so populations cannot grow indefinitely

The maximum amount of organisms in an area is that ecosystem’s carrying capacity

S-shaped curve Carrying Capacity Different organisms have evolved differently to combat

competition and carrying capacity

r and k Strategists r-strategists: Give birth

to many offspring which mostly die early in life

k-strategists: Give birth to a few offspring, care for them, many reach adulthood/sexual maturity

Limiting Factors Ultimately what limits the size of populations are

limiting factors

Either living factors (biotic) or nonliving (abiotic)

Can be dependent on density or not

Can be devastating or regulatory

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Matter Recycling Matter can neither be

created nor destroyed, so all matter on Earth cycles through various phases

Predictable patterns of cycling are what create and dictate Earth’s biogeochemical cycles Biogeochemical cycles

typically have both abiotic and biotic components

Hydrologic Cycle

Carbon Cycle Phosphorus Cycle

Nitrogen Cycle Nitrogen is the primary

component of the atmosphere (~78%)

Atmospheric N cannot be used by plants or animals

It must first be converted through a process called nitrogen fixation

Nitrogen Fixation Natural or synthetic process

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Living N Fixation Done exclusively by prokaryotes

Best-known N fixers are bacteria called Rhizobium Found in most soils

Both free-living and attached to plant roots

Primarily associated with leguminous plants (peanuts, beans, peas, clover, etc.)

Can fix the most bacteria when found in plant root nodules

First step in N-Fixation Bacteria must transform nitrogen gas into ammonium

ions

Catalyzed by the enzyme nitrogenase

Needs a supply of hydrogen, ATP for energy, and NO oxygen (anaerobic)

What we need for N-fixing and where we get it… H+ come from a chemical in

the plant called NADP

ATP (energy) comes from the break-down of sugar in cellular respiration

Anaerobic conditions maintained by plant using a protein called leghaemoglobin

Nitrogen Fixation as a Relationship Considered a type of mutualism:

Rhizobium get a habitat and suitable place to live as well as what they need to fix nitrogen

Plants in turn get a supply of usable nitrogen

Nitrogen Fixing elsewhere Lightning has enough energy to sever the triple bonds

between two N

N then combines with oxygen in nitrogen oxides which are soluble in water

Significant source of nitrogen in areas of heavy thunderstorms (tropics)

The Haber Process Ammonium is an essential

plant nutrient, so it is necessary in many fertilizers

Haber process combines nitrogen and hydrogen gases into ammonia

Takes a lot of energy, though

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How plants use fixed nitrogen If the plant has Rhizobium in its roots: amino acids

synthesized right there in the nodules and then transported to rest of plant Amino acids proteins

If no Rhizobium: plants uptake fixed nitrogen from soil and… Transform nitrates nitrites ammonia amino

acids in the roots, OR

Transport nitrates to leaves where they are transformed eventually into amino acids

Eventual goal is proteins either way

From plants to animals… We can only get nitrogen from organic

molecules like those in plants or other animals

Some amino acids but mostly proteins ingested and broken down into AA’s AA’s absorbed by blood in primarily

the small intestine and transported to all cells

Excess AA’s are broken down via deamination in the liver and excreted in urine.

Animals to soil… Proteins are broken down by decomposers like fungi

and bacteria using protease enzymes

Ammonia is released by these bacteria in the process of ammonification

Ammonia is in turn broken down by different bacteria into first nitrites and then nitrates

Process of nitrification carried out by nitrifying bacteria, especially those in the groups Nitrosomonasand Nitrobacter get energy from nitrification

Requires lots of oxygen

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Denitrification Reverse the process and turn nitrates into nitrogen gas

Common in decomposition sites (compost, wet soils, boggy areas, sewage treatment)

Brings the cycle full circle!

Life is ordered in a hierarchy Kingdom

Phylum

Class

Order

Family

Genus

Species

The Five Kingdoms Prokaryota = Prokaryotes

Sometimes divided into Archaebacteria and Eubacteria

Protoctista = Protists. Called Kingdom “Protista” in the States

Fungi = true mushrooms, slime molds, molds, shelf fungi

Plantae= Plants

Animalia = Animals

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ProkaryotaProtoctista

FungiPlantae

Animalia

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Organizing biology is all about hierarchies!

Already talked about the 5 (6) kingdom system

Each large group is split into smaller groups

(Domain) Kingdom Phylum Class Order Family Genus Species

Carl Linnaeus Nobilized as “Carl Von Linne,”

he was a Swedish botanist who revolutionized the way scientists communicate about organisms

Published the Systema Naturaein 1735

Widely hailed as one of the most important biological texts ever written

Binomial Nomenclature Every organism on Earth that is described by a

scientist has two names (Bi- nomial)

Genus name: Larger group

Species name: specific to ability to inbreed

Why care about binomial nomenclature?

Standardizes scientific discourse

Enables scientists from different cultures to communicate about the same organism

Common names are VERY unreliable!

Dichotomous Keys Series of two-condition questions used to identify

organisms by morphological traits

Based in the logic that genetically (and thus evolutionary hierarchically) similar organism are similar in appearance

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