Hosta Species Update●The Hosta Library●ORG20080329●©W. George Schmid 2010 Original Edition 20080329 1st Revision 20100619

H. kikutii var. polyneuron (F. Maek.) N. Fujita Maekawa in Nakai, T. Iconographia Plantarum Asiae-Orientalis, Tokyo, 4:496–497;

1952 (excluding the synonyms). Acta Phytotax. et Geobot., Vol. 27, 3/4:78 1976. スダレギボウシ = 簾擬宝珠 = Sudare Gibōshi (= rattan blind Hosta)

History, Nomenclature and Habitat: The H. kikutii complex is widespread and shows distinct local phenotypes. Among these are distinct forms with larger leaves having more principal vein pairs than those of H. kikutii var. kikutii forma typica. In addition, the principal veins are spaced distinctly closer. This has given rise to the Japanese name Sudare Gibōshi, derived from the Japanese Kanji 簾, which transliterates to sudare (the name for a familiar Japanese rattan blind, a bamboo blind or screen with closely spaced slats). Thus, the Japanese name translates to “rattan blind hosta.” This varietas is well known in Japan and has been cultivated there for many years under its Japanese name スダレギボウシ = 簾擬宝珠 = Sudare Gibōshi. The scientific species epithet is of Greek extraction, poly meaning many, and neuron standing for veins (nerves), hence the meaning “many-veined.”

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Veining of H. kikutii var. polyneA = H. kikutii var. kikutii (Miyazaki-ken; 宮C = H. kikutii var. polyneuron (Kochi-ken; 高

Comparison of average leaf shape andFrom N. Fujita 1976 (“Veinings of Hosta

uron and H. kikutii var. kikutii 崎県) B = H. pulchella (Ōita-ken; 大分県)知県) D = H. shikokiana (also Kochi-ken)

arrangement of principal vein pairs ”;The genus Hosta (Liliaceae) in Japan)

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Although the name of the typical province of Hyūga-no kuni (日向国) in wkikutii var. kikutii was found, H. kikutii vmountainous regions of eastern Shikoku Widespread populations have been obserIshizuki and Akaishi mountain ranges, in (香川県). Fujita (1976) states that the tySatsunan Islands region (薩南諸島) on Yapparent geographic displacement is tyakusimensis as being synonymous to H. kitype specimen for H. kikutii var. yakusimenFujita’s 1976 placement is not supported bthe two varieties were differentiated with aconsidered distinct entities. Fujita (1976) astating (Fujita 1976, pp. 80-81) that he wakikutii var. polyneuron populations on Yakurecorded) near Ikegawa-chō (池川町 [now

H. kikutii var. polyneuroGrowing along steep bank in Ehime-Court.: © Rojiura-no-Giboshi 路地

n(in situ) スダレギボウシ

ken (愛媛県); Shikoku Island (四国)

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species Hyūga Gibōshi alludes to the old estern Kyūshū (九州), where the type of ar. polyneuron has a main habitat in the Island (四国; meaning “four provinces”). ved in Kochi-ken (高知県), south of the Ehime-ken (愛媛県), and in Kagawa-ken pe was collected by Suzuki in northern akushima (屋久島). The reason for this hat Fujita considered H. kikutii var. kutii var. polyneuron and used Maekawa’s sis to apply to H. kikutii var. polyneuron.

y RAPD/DNA analysis (see Pages 6-7) and single primer OPB-02 and are therefore

lso gave credence to this differentiation by s not able to confirm the presence of H. shima and used a leaf sample collected (as Niyodogawa-chō (仁淀川町)]) in Kochi-

裏のギボウシこと 後ノ上 憲一

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ken for his illustration “Veinings of Hostfeatured in this Species Update were taken var. polyneuron (in situ) exclusively fromprefectures mentioned above. It is possible northern Satsunan Islands, but this has noM.N. Tamura proposed that H. kikutii vataxon.” They suggested two new varieties,Fujita & M. N. Tamura and H. kikutii varH. kikutii var. densinervia differs from H.veins of the leaves more or less close togetscabrinervia differs from H. kikutii var. dethe abaxial surface of the leaves prominenvarieties were formerly assigned to H. kikuH. polyneuron F. Maek.). DNA content (results and that they undoubtedly closelyRynchophorae has been confirmed. RAPDpolyneuron can be differentiated with the s

H. kikutii var. polynAccessions of H. kikutii variants in the gr

In cultivation the scapes are l© W.G. Schm

euron (cult. vouchers) eenhouse of University of Georgia, 1989 eaning but are more upright id 1989.08.29

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a” (shown here on page 1). Photographs of small and large populations of H. kikutii areas in Shikoku Island (四国) in the

that H. kikutii var. polyneuron occurs in the t been confirmed. In 1980, N. Fujita and r. polyneuron be considered a “doubtful

Hosta kikutii F. Maek. var. densinervia N. . scabrinervia N. Fujita & M. N. Tamura.

kikutii var. kikutii in having the principle her and papillose abaxially. H. kikutii var. nsinervia in having the principal veins on tly papillose. Plants belonging to these two tii var. polyneuron (F. Maek.) N. Fujita (= 2C) shows that all H. kikutii show close related (see pages 6-7) and placement in analysis has shown that the H. kikutii var. ingle primer OPB-02 and is therefore con-

sidered a distinct entity and taxon in section Rynchophorae. As to whether it should

red to as 表日本 (t c Ocean coasta cific coastal regions of Shikoku Island (四 G. Murata and H. Koyama

be a doubtful taxon and be replaced by two new varieties on the basis of morphological differentiation is a matter of taxonomic opinion since H. kikutii var. polyneuron shows considerable polymorphism in its various endemic populations.

◄◄◄ H. kikutii

var. polyneuron (in situ)

In situ the scapes are more leaning to be-come subhorizontal at times. Note many and tightly spaced principal veins in Ehi-me-ken ( 愛 媛 県 ) Shikoku Island (四国)©Rojiura-no-Giboshi 路地裏のギボウシ

こと 後ノ上 憲一

Notes on Habitat: The species H. kikutii var. polyneuron and H. kikutii var. caput-avis occupy a habitat on the Pacific side of Japan refer

ransliterated Omote Nihon). In a strict sense, Omote Nihon is applied to Pacifil regions of the main island of Honshū (本州), but the Pa

国) are also included here. (1980) investigated the taxonomic and phytogeographic nature of plants considered rheophytes (rheophyte = a plant that is associated with flowing water). Although the mainland of Japan is located in the warm temperate zone, H. kikutii var. polyneuron and H. kikutii var. caput-avis are by Murata, et. al., considered to occupy habitats similar to those of tropical plants, which grow on rocky shelves near rivers in the bottoms of gorges which are either inundated once to several times a year or receive abnormally high precipitation, which results in heavy run-off. The study further states that these rheophytic hostas have evolved to develop tenacious rhizomes and roots that sprout immediately after a flood and which cling to the crevices of rocks anchoring its slender or down-sloping leaves against rushing water. These plants are resistant to physical damage but grow weakly in shade and for this reason H. kikutii var. polyneuron and H. kikutii var. caput-avis are most frequently found in open valleys or ravines with considerable sun exposure. My own observations indicate that these specific and sometimes unique requirements have led to disjunct, small populations that have undergone longtime adaptation to ecological niches, leading to considerable differentiation and polymorphism. Leaning scapes and slender leaves are an expression of this adaptation. On the other hand, the development of larger leaves to maximise photosynthesis has also been observed and this is the case with H. kikutii

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var. polyneuron. This taxon inhabits deeper ravines and narrow valleys with more shade. The photo-graph shown ◄ ◄ ◄heEhShikoku (

re [in situ in ime-ken (愛媛県);

四 国 )]. Court.: © 路地裏 の

ギボウシ こと後ノ

上 憲 一 The pre-carious habitat mir-rors that of H. hypoleuca in Chichi-iwagawa Gorge ( 乳岩川); Aichi-ken (愛知県).

Plant Morphology: Note: The description below is average for this varietas. H. kikutii var. polyneuron is a polymorphic taxon, which shows considerable macro-morphological differences in its natural populations. Variants have lanceolate to ovate lanceolate leaves with 9-11 (13) pairs of principal veins, which have a narrow spacing. Although other variants of H. kikutii have leaves with relatively tight vein spacing, this varietas has its veins spaced closer than most other Hosta species. The leaf margins can be straight or undulate. In most individuals, the flower morphology is very uniform and matches that of H. kikutii var. kikutii forma typica. In most populations, the scapes are straight but lean considerably and may reach sub-horizontal position, particularly when growing on vertical rock faces. Plant size 60 cm dia., 25 cm high (24 by 12 in.). Petiole 13 by 0.4 cm (5.2 by 0.15 in. wide) ascending in an arch, light green. Leaf 18–25 by 7.5–11 cm (7–10 by –4.50 in.), elliptical, ovate to lanceolate, petiole-leaf transition attenuate to rounded, elongated, drooping tip, with flat surface, slightly convolute at the base, no waves or slight waves margin, not rugose, shiny dark to medium green above, shiny light green below. Venation 9-11 (13), very closely spaced, sunken above, projected below. Scape 50 cm (20 in.), straight, smooth round. Sterile lower bracts small, leaf-like, with exterior bracts enveloping the interior; occasionally ±2 larger sterile leaf-like; fertile bracts navicular, broad, thick, first green, but opening white, withering late, but persisting at anthesis, 2 by 1 cm (0.75 by 0.33 in.), some larger. Raceme short, densely imbricated, 10–20 flowers. Flowers mostly white, shiny, sometimes very

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lightly suffused purple in the center of the tepal (Type A; Schmid 1991 ►); 4.5 by 3 cm across the lobes (1.75 by 1.25 in.), carried

Genome Size: DNA content (2C) in pg (one (10-12) gram) foneuron is 22.9 with a Dl/DAPl ratio of 1.12. Average DNA co(10-12) gram) for the H. kikutii species and botanical varieties and forms has an average 22.8 ± 0.3. (Zonneveld, B.J.M. and F. Van Iren; 2001). As expected, all of the major macromorphological variants (H. kikutii var. kikutii var. polyneuron, and H. kikutii var. caput avis [syn. H. kiku

between 22.4 and 23.0 and t nt in Rynchophorae is confirmed by a close grouping of DNA content values. H. kikutii var. polyneuron and the closely related species H. shikokiana show a DNA content (2C) in pg (10-12 gram) = 22.9 (Zonneveld, B.J.M. and F. Van Iren, 2001), which falls into the range of the species in

horizontally on strong, 2.5 cm long (1.00 in.) pedicels, which sometimes extend beyond the bracts; perianth expanding, funnel-shaped, in the central part dilated bell-shaped, lobes spreading @+angled to the axis of perianth; stamens as long as perianth or extending beyond it. Anthers purple. Late August to early September. Fertile. Karyotype-Chromosomes: Sporophytic Count = 60; 12 large, 48 small; (2n).

r H. kikutii var. poly-ntent (2C) in pg (one

f. leuconota, H. kikutii tii var. tosana]) vary

heir placeme

section Rynchophorae and its placement in the section has been confirmed. Populations of H. kikutii sensu lato, H. kikutii var. polyneuron, and H. shikokiana occur sympatrically on Shikoku Island (四国). ◄ Fig A: DNA Banding Pattern

20 = H. kikutii var. kikutii

22 = H. kikutii var. caput avis (H. kikutii var. tosana)

23 = H. kikutii var. polyneuron

24 = H. kikutii var. yakusimensis (H. kikutii var. kikutii f. leuconota is considered a morph).

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02 = (5’-TGATCCCTGG-3’) and the varietas

ection Rynchophorae.

y: ujita. ol. 27, 3/4:78 1976.

ll. Fujita, near Ikegawa-chō (池川町 [now ochi-ken (高知県); Shikoku, Japan.

ons observed in Kochi-ken (高知県), near the anges, in Ehime-ken (愛媛県 ), and in

in moist, shady valleys and ravines and in with alpine grasses and subshrubs.

. polyneuron Maekawa in Nakai: Iconogr. Plant. Asiae Orient., 5:496–497 1952

ynonymy not accepted; see page 5).

スダレギボウシ = 簾擬宝珠 = SudThe following taxa have beeFujita and M.T. Tamura (19

ザラツキギボウシ = ざらづき擬prominent papilous veins on

アワ キギボウシ = あわ擬宝珠 =Shikoku Island (四国) [pro p

Horticultural Names anH. kikutii var. polyneuron ‘Ishizu’

(石鎚山; Ishizuchisan). Maon this and neighboring mo th the habitat name is superflu s since it could mean any of the many different selections that inhabit these

H. kikutii var. polyneuron ‘Kifukurin’ name is H. ‘Kifukurin Sud ame for a number of different yellow-margined sports but some selections have unique Japanese

DNA Banding: Recent RAPD analysis (Y. Yu, 2002; Sauve, R.J., S. Zhou, Y. Yu, and W.G. Schmid; 2005) has established the banding patterns of four closely related H. kikutii species accessions in section Rynchophorae (See Fig. A). H. kikutii var. polyneuron was examined and its banding pattern is illustrated in Fig. A, which shows comparison using a single primer OPB-H. kikutii var. polyneuron was differentitherefore considered a distinct entity in s

Taxonomic Type and SynonymH. kikutii var. polyneuron (Maekawa) FActa Phytotaxonomica et Geobotanica, VType: Based on the cultivated plant; co

Niyodogawa-chō (仁淀川町)]) in KHab.: Shikoku Island (四国). Populati

Ishizuki and Akaishi mountain rKagawa-ken (香川県); usuallyhigher mountain passes together

Botanical Synonyms:

ated with the single primer OPB-02 and is

H(excluding the synonyms).

. kikutii var. yakusimensis Fujita 1976 (sHH. kikutii F.Maek. var. densinervia N. Fujita & M.T. Tamura, var. nov. 2008 Acta

Phytotax. Gebot. 59 (1) : 31-36 (pro parte tantum). H. kikutii F.Maek. var. scabrinervia N. Fujita & M.T. Tamura, var. nov. 2008 Acta

Phytotax. Gebot. 59 (1) : 31-36 (pro parte tantum).

Japanese Synonyms: are Gibōshi (= rattan blind Hosta). n associated with H. kikutii var. polyneuron by N. 80) (see page 3): 宝珠 = Zaratsuki Gibōshi (= “feels rough” = abaxial principal veins) [pro parte tantum]. Awa Gibōshi (= name of a locality in eastern arte tantum].

d Synonyms: = a selection of the species from Mount Ishizuchi ny populations of H. kikutii var. polyneuron exist untains and to label a selection of the species wiou mountains.

= (name incorrect per ICNCP). The correct are’, This is a collective n

cultivar names. Some yellow-margined sports have been given a uniquecultivar name (see page 12). Some of these plastome mutations may be sportsof a hybrid between H. longipes and H. kikutii. In Japan the yellow-marginedsports are different and collectively known as 黄覆輪スダレギボウシ (= H.‘Kifukurin Sudare’) or have been given a unique cultivar name (see page 12)

H. kikutii ‘Ogon’ = (name incorrect per ICNCP). This name should not be used. Ithas been applied incorrectly to various yellow sports of H. kikutii var. polyneuron. In Japan the yellow sports are collectively known as 黄金スダレ

ギボウシ (= Ogon Sudare Gibōshi; see page 12). Some have been assignedunique cultivar names.

H. kikutii var. polyneuron ‘Shirofukurin’ = (name incorrect per ICNCP). A name forwhite-margined sports of the species. There are two distinct sports incultivation, both named specifically, one from Wakayama-ken (和歌山県),which should be called H. ‘Sudare Shirofukurin Wakayama’ and the otherfrom Shikoku (四国) with the correct name being H. ‘Sudare ShirofukurinShikoku’ (see page 13).

H. kikutii var. polyneuron ‘Shironakafu’ = (name incorrect per ICNCP). This nameshould not be used. It has been applied incorrectly to various white centersports of H. kikutii var. polyneuron. The Japanese collective name is 白中斑

スダレギボウシ= H. ‘Shironakafu Sudare’. In Japan the yellow-marginedsports are different and some have unique cultivar names (see page 12).

.

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H. kikutii var. polyneuron(in situ) スダレギボウシ

Growing along steep bank in Ehime-ken (愛媛県); Shikoku Island (四国)Court.: © Rojiura-no-Giboshi ▪ 路地裏のギボウシこと 後ノ上 憲一

H. kikutii ‘Variegated’ = (name incorrect per ICNCP). This name should not be usedIt has been applied incorrectly to various variegated sports of H. kikutii varpolyneuron.

H. kikutii var. polyneuron in Cultivation: A number of different clonescollected from wild populations represent H. kikutii var. polyneuron in cultivation.

utii (sensu lato) examples date to 1969 from Craig (as ginating with Dr. Miazawa as H. caput-avis and

. r. yakusimensis obtained by Davidson from Suzuki and listed as Summers

kutii var. polyneuron originated with the 1989-1991 Study at UGA, also with W.G. Schmid, and other

ccessions, the polymorphic nature of this lthough uniform in flower morphology,

s. The specimens respond

wild. I Most

Schmidvar. poare cov

◄◄◄ H. kikutii var. polyneuron

.

.

The earliest horticultural H. kikrs [1972] No. 337) oriH-9/Summe

from Epstein as Summers No. 406 (Summers 1972). This was preceded in 1968 by Hkikutii vaNo. 297 (Summers 1972). In 1969, Craig imported another accession named H. tosana and was listed by Summers as No. 358. Purportedly, it originated on Izu Peninsula, which is questionable since H. kikutii (sensu lato) is not native to this part of Japan. Exact identification is no longer possible, although some of these accessions may have been H. kikutii (sensu lato). Unfortunately, these and other non-vouchered imports created some confusion and some of the cultivated clones of these early imports still carry incorrect names in garden. Vouchered accessions of H. ki

Hostaacademic sources. As with other H. kikutii avarietas is reflected in cultivated specimens. Ait shows considerable variation in the leaf sizes and shapewell to good cultivation practice and well prepared soil. Given copious amounts of moisture, cultivated specimens will grow abundantly and become larger than in the

t should also be noted that not all accessions are being propagated.specimens offered in the trade are from Summers, UGA and Hosta Hill R.G. (W.G.

) accession from Shikoku( 四国), with some variegated sports of H. kikutii lyneuron originating with Hideko Gowen. These and other variegated sports ered under the heading Horticultural Progeny, below.

Higashiakaishi-yama (東赤石山) Ehime-ken

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(愛媛県 四国) Court.: © fc2web (四国)

H. kikutii

スダレLeaf mearly s

Shikok

kikutithe trade but not all specimens offeredthis spwere some be explimited

var. phostarIncorr

erica and Europe and consequently the parentage

var. polyn on ►►►(in situ – moist montane forest floor)

ギボウシ (簾擬宝珠) ound showing the colors of pring flush

eur

Kochi-ken (高知県) u Island (四国)

are true representatives of ecies. Since several clones

collected from the habitat, phenotypical differences can ected, but the species has a habitat in which most of the

H. kikutii var. polyneuron ►►► Detail of perianth with tepals スダレギボウシ (簾擬宝珠) Kochi-ken (高知県) Shikoku Island (四国)© Rojiura-no-Giboshi 路地裏の ギボウシこと 後ノ上 憲一 Horticultural Progeny: H.

i var. polyneuron is available in

clones observed have similar macromorphology, i.e., tight vein spacing, straight but leaning scapes, white flowers with dark purple anthers (before dehiscing). H. kikutii

olyneuron is not listed as a pollen parent in the AHS Registry (URL: egistrar.org). It may have been used for hybridizing non-registered cultivars. ectly identified H. kikutii var. polyneuron specimens may have been used for izing in Asia, North Amhybrid

indicated in some records is subject to correction.

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JapanSportReportlectors

clivity polynetome mnumbe

of the and sports are listed, but many re unique and not available in

H. ‘Yosh スダレギボウシ

Hosta Library – le

ese Cultivars and s: s from Japanese col- and pictorial material

on Web sites and seen in hosta exhibitions indicate the pro-

of H. kikutii var. uron to undergo plas-

utations. Only a small r of the many sports

extant can be illustrated. Some more popular cultivars

a

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the trade and for this reason only the better know sports are featured. A number of these sports have been fount in the wild and are usually given names, which indicate the general location of discovery, as for example Yoshinogawa (= Yoshino River 吉野川), as seen © HL/R. Duback

H. ‘Yoshiスダレギボウシ

Hamanako Flower E

Court.: 浜名湖

nogawa’ ‘吉野川’

xposition - ぎぼうし

花博 擬宝珠

inogawa’ ‘吉野川’

af closeup

at the Hamanako Flower Exposition. H. kikutii var. polyneuron has produced many mutations in the wild and H. ‘Yoshinogawa’ is but one of many. Some sports have not been nam-ed but have been shown in Hosta exhibitions in Japan. As with various pheno-types of the species, the mutations also show nar-row- and wide-leaf forms. Many are reported to be unstable in their variega-tion patterns. Most origi-nate along mountain riv- ers and streams in Kochi-

ダレギボウシ (簾擬宝) hree examples of wild llected mutations ourt.: ©高知ギボウシ

好会事務局 ochi Hosta L. Assn.

op: 仁淀川▲ ダレギボウシ iyodogawa)

flowered

ken (高知県) and Ehime-en (愛媛県).k

H. kikutii var. olyneuron ►▼▲ p

スTcoC愛K

Tス(N(白花) white

Center: 新庄川►

スダレギボウシ(Shinjougawa) Bottom: 四万十川▼

スダレギボウシ(Shimantogawa)

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Fig. 1 Fig. 2

Also seen frequently are yellowprimarily found in Kochi-ken (高and other streams on Shikoku ギボウシ 黄中斑 (Kinakafu Suoccasion). As can be seen from the(Nishiki = 錦), variegated, spottvariegated in center (Nakafu 中斑yellow-margined sport on this pakikutii var. polyneuron phenotypeKifukurin Sudare Gibōshi. No uyellow sports, two stable, white mHideko Gowen, a native Japanes

medio-variegated sports (Fig 1 and 2). They are 知県) in the headwaters of Niyodogawa (仁淀川)

Island (四国). The are collectively called スダレ

dare Gibōshi) and plus the location of discovery on illustrations, most of these sports come streaked

ed or speckled (斑), margined (Fukurin = 覆輪) or ) or all yellow color. Also exhibited has been the ge (see photo below▼), which is a wide-leaf H.

(= スダレギボウシ 黄覆輪), which transliterates to nique cultivar name is known. Aside from streaky argined sports are in the trade. They originated with e residing in the U.S., who obtained them in Japan.

Several white-margined culti-vars are available, Schmid (1991) added the Japanese geographic source names the

m. A larger ギボウシ

the . ‘Shiro-

yama’ (Fig. 3). The Waka-yama-ken ( 和 歌 山 県 ) location has has not been verified, but the name is in use. H. ‘Shirofukurin Su-dare Shikoku’ (スダレギボ

ウシ 白覆輪 四国) equates to the Western H. ‘Shiro-fukurin Sudare Shikoku’ (Fig 4).

ダレギボウシ 黄覆輪 yellow-margined form of H. kikutii var. polyneuron

y collected in Kochi-ken (高知県); Agawa-gun (吾川郡); Niyodo-chō (仁淀川町)

distinguish theform called スダレ

白覆輪 和歌山 has cultivar name Hfukurin Sudare Waka-

A

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Reportedl

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Fig. 3

Fig. 4

スダレギボウシ 白覆輪

►

◄◄◄ H. kikutii var. polyneuron

(yellow sport) ダレギボウシ黄金鶴

H. ‘Golden Crane’ A yellow form of called

‘黄金鶴’ (= ‘Golden Crane) (in Japan called “tsuru”;

Grus japonensis, of the family Gruidae)

See pages ►►11 to 13

◄◄◄ H. kikutii var. polyneuron

ow sport with green margin) Court.: © Orimiki 写真共有サイト

Hi

(A yell

▼▼▼H. kikutii var. polyneuron

(A yellow sport with green margin) Court.: © 瓦ジスタ

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◄◄◄ H. kikutii var.

polyneuron (in situ)

gashiakaishi-yama (東赤石山) Ehime-ken (愛媛県 四国)

2010-06-19

ceae)

.E., 1988. Biosystematics of the Eastern Asian species of Hosta). University of Georgia, Athens. (Personal communications).

rdtman, G., 1966. Pollen morphology and plant taxonomy. Angiosperms. Hafner Publishing Company, New York.

genus Hosta (Liliaceae) in Japan. Acta Phytotaxonomica et Geobotanica, Vol. 27, 3/4:77-80 1976.

atus in Hosta (Asparagaceae). Acta Phytotaxonomica et Geobotanica, 59, 1: 31-36.

Botanical Nomenclature = ICBN, adopted by the International Botanical Congress, Nomenclature Section, Vienna Code 2005.

InteInternational Commission for the Nomenclature of Cultivated Plants of the

BS Division of Botany, 2004. nica,

ect. 3 Botany, Vol. 5:375, ic. 3, 4 p.

(Big Dictionary of Garden Craft),

yclopediaSeibundoshinkosha, Tokyo, Vol. 3:1105–1109.

Maekawa F. 1952 in Nakai, T. Iconographia Plantarum 4:496–497.

Murata, G., and H. Koyama. 1980. 襲速紀地域を中心とした日

特性について (Characteristics in the flora of he PMemoirs of the National Science Museum. Vol.13

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© W.George Schmid 2010: The text and illustrations are copy-

righted and are available for personal reference only. Other contributors retain their copyright of featured photographs as noted

in captions. The content may not be published inwithout the author’s written permission. Web

W. George Schmid HostaLibrary.org/species/.

の観察と栽培Yu, Y. 2002. Classific d cultivars

id); publish summary in Sauve, TSU Graduate School (with W.G. Schm

Nursery, Iome size and pollen viability as

nc.

iology, 3, pp. 176-

printed form quote reference:

2010-06-19 - 17 -

kutii var. polyneuron(in situ) スダレギボウシ

ing along steep rock ravine in Ehime-ken (愛媛県); Shikoku Island (四国) Note small seedlings below the main population further up

H. kiGrow

Court.: © Rojiura-no-Giboshi ▪ 路地裏のギボウシこと 後ノ上 憲一