Insects of Saudi Arabia Neuroptera Fam. Myrmeleonidae

Insects of Saudi Arabia

Neuroptera : Fam. Myrmeleonidae

H. Holzel

A b s t r a c t : 70 species of Myrmeleonidae are recorded from Saudi Arabia, 39 of them for the first time; 10 new species - Myrmecaelurus parvulus, M. (Nohoveus) saudiarabicus, Iranoleon arabicus, Solter wittmeri, Solter buettikeri, Solter pallidus, Gepus cunctatus, Cueta amseli, Cueta asirica and Creoleon desertus - are described.

Myrmeleonidae 'UL~ : ~ Y I a,:<; *

I :'u r+ a p J>Y 9~ yq ~ 8 y +J+I -I ip Myrmeleonidae 'UL~ ip w y. d3U b+s

Iranoleon a rab icus , M . (Nohoveus) saudiarabicus , Mymecael urus parvul us : & L c++ €'+I

Gepus cunctatus , Sol t e r p a l l i d u s , Sol t e r b u e t t i k e r i , Sol t e r wi t tmeri

. -3 6- .Creoleon desertus , Cueta a s i r i c a , Cueta amseli

First records of Ant-lions from Arabia were made about 140 years ago. KLUG (1834) described 6 species from "Arabia felix" without further particulars. In the following decades numerous publica- tions with descriptions of single species from Arabia were published, most of them by NAVLS (1916, 1926, 1934). KIMMINS (1943) studied a collection of Myrmeleonidae made in Saudi Arabia by Mr. H.St. J.Philby in 1930-1934 and described 7 new species. In a paper on the Myrmeleonidae of Anterior Asia (HOLZEL 1972) I treated all records of Myrmeleonidae from Saudi Arabia known to me (all in all 31 species) and described on this occasion 4 new species from this country.

The present paper is based on the large collections of Myrmeleonidae made in Saudi Arabia by Prof. W.Biittiker in 1975-1979 and Dr. W.Wittmer and Prof. W.Biittiker in 1976, Mr. A.Pittaway in 1977-1979 and Dr. H.G.Amse1 in 1979. I am gateful to Dr. W.Wittmer (Basle), Dr. H.G.Amse1 (Karlsruhe) and Dr. P.C.Barnard (London) for the opportunity of studying this material,

I also wish to express my thanks to Dr. F. Espafiol, Museo de Zoologia, Barcelona, Dr. 0. S. Flint, U. S. National Museum Smithsonian Institution, Washington (MW), Dr. W. Forster, Zoolog. Samm- lung des Bayerischen Staates, Miinchen (MM), Dr. B.Hauser, Museum dYHistoire naturelle, Genf (MG), Dr. D.E. Kimmins, British Museum Natural History, London (BM), Pater Palazon de Lattre, Colegio Salvador, Zaragoza (MZ), Mr. D.Simon, Zoological Institute, University of Tel Aviv, Dr. J.R.Steffan, Museum National dYHistoire naturelle, Paris (MP) and Dipl. Biol. H.Wendt, Zoolog. Museum der Humboldt-Universitat, Berlin (MB), who enabled me to study the necessary type material deposited in their museum collections.

Most helpful in interpreting the distribution of Arabian species was the knowledge of a collection of Myrmeleonidae from Sudan, on the opposite side of the Red Sea, made by Prof. Dr. R.Remane

FAUNA OF SAUDI ARABIA 4, 1982 245

(Marburg) in 1961-1962. As the results of this expedition never have been published - on the other hand the material is cited in this paper very often - it seems convenient to provide data of the two essential sampling sites: Sudan-Kassala = Kassala province, Red Sea-hills, Erkowit mountains 1100-1300 my R.Remane; Sudan-Hudeiba = Northern province, Hudeiba 380 my Wustenrand- Nilufer (Akazien-Dornbuschsteppe mit eingestreutem Kulturland) R.Remane. I am grateful to my friend Dr. P.Ohm (Zool. Museum der Universitat Kiel) who most generously enabled me to examine this important material which is deposited in his collection.

Evidences of other hitherto unpublished records cited in this paper are either deposited in the named museum collections or in collection Holzel. All holotypes of new described species are - if not- explicitly stated otherwise - deposited in the collections of Naturhistorisches Museum, Basle, paratypes - if available - also in collection Holzel. The terminology of the wing venation and of genital structures as used herein follows ASPOCK & al. (1980).

The photographs are the work of Mr. H . Elsasser (Institut fur Umweltwissenschaften und Natur- schutz der Usterr. Akademie der Wissenschaften, Graz). I am most grateful for his assistance.

Su bfam. Palparinae Genus Tomatarella Kimmins

Tomatarella Kimmins, 1952, Entomologist 85: 34 (Typus generis: Tomatarella markli Kimmins, 1952).

Tomatarella markli Kimmins Tomatarella markli Kimmins, 1952, Entomologist 85 : 34.

Saudi Arabia: Ghudaiyer (Kimmins 1952); 1 9, W.Duwassir, 5 km SW Khamis Mushayt, 25.VIII. 1976, W. Buttiker.

Distribution: So far Saudi Arabia only.

Genus Palpares Rambur Palpares Rambur, 1842, Hist. nat. Ins. NCvr.: 365 (Typus generis: Myrmeleon libelluloides Linnaeus, 1767).

Palpares dispar Navls Palpares dispar Navis, 1912, Annls Soc. scient. Brux. 36: 213; HOLZEL (1972).

Saudi Arabia: Riyadh (HOLZEL 1972); 3 99, Hail, 25.1X.1976; 1 9, Maraba, 3.IX.1978; 1 6, W. Huraymala, 12. VIII. 1976; 1 9 , Wadi Khumra, 16. IX. 1976, W. Buttiker.

Distribution : Saudi Arabia, Israel; Africa : Egypt.

Palpares cephalotes (Klug) Myrmeleon cephalotes Klug, 1834, Symb. phys. 4 : p1.35,Fig. 1. Palpares cephalotes (Klug): HOLZEL (1972).

Saudi Arabia: Riyadh (HOLZEL 1972); 1 d , 1 9 , Wadi Mizbil, 16.IX. 1976; 1 d, Bahara, 8.V. 1978; 3 99, W. Huraymala, 12. VIII. 1976; 1 9, Durma, 15. VIII. 1977 W. Buttiker.

Distribution: Saudi Arabia, Sinai, Iran, Afghanistan; Africa: Egypt, Sudan (Hudeiba), Ethiopia.

Palpares papilionoides (Klug) Myrmeleon papilionoides Klug, 1934, Symb. phys. 4: pl. 35, fig.2.

Saudi Arabia : 1 8, Adama (Asir), 22. IX. 1978, W. Buttiker. Distribution: Saudi Arabia; Africa: Ethiopia.

Palpares klugi Kolbe Myvmeleon papilionoides var. Klug, 1834, Symb. phys. 4: pl. 35, fig.3. Palpares klugi Kolbe, 1898 Stettin. ent. Ztg: 230; ESBEN-PETERSEN (1930).

Saudi Arabia: 1 9, Maraba, 30. IX. 1978 W. Buttiker. Distribution : Saudi Arabia; Africa : Sudan, Ethiopia.

Palpares walkeri McLachlan Palpaves walkeri McLachlan, 1894, Entomologist's mon. Mag. 5: 173; STITZ (1912). Palpares laticaudus Navis, 1915, Mems R. Acad. Cienc. Artes Barcelona 11 : 378.

Saudi Arabia: 1 8 , Adnan, 21. IX. 1978, W. Buttiker ; first record! Distribution : Saudi Arabia, S-Yemen ; Africa : Sudan.

Palpares angustus McLachlan Palpares angustus McLachlan, 1898, Trans. ent. Soc. Lond.: 152; STITZ (1912); KIMMINS (1960).

Saudi Arabia: 7 99, Maraba 30.IX. 1978; 1 9 , Adnan, 21.IX. 1978 W.Buttiker; 1 9 , Asir, 2350 m, 5 km S of Namas, 17. IV. 1979 H . G. Amsel; first record!

Distribution: Saudi Arabia, Jordan, Syria; Africa: Egypt, Somalia, Libya, Algeria.

Subfam. Myrmeleoninae Genus Acanthaclisis Rambur

Acanthaclisis Rambur, 1842, Hist nat. Ins. Nivr . : 378 (Typus generis : Myvmeleon occitanicum Villers, 1789).

Acanthaclisis mesopotamica Holzel Acanthaclisis mesopotamica Holzel, 1972 : 13.

Saudi Arabia: 1 8 , 1 Q, Hofuf, 10.X. 1977, W.Buttiker; 1 9, Hofuf, 31 .V. 1977, A. Pittaway ; first record !

Distribution : Saudi Arabia, Iraq.

Acanthaclisis formosa Holzel Acanthaclisis formosa Holzel, 1980, Israel J. Ent. 14: 30.

Saudi Arabia: 1 Q, Asir, 850 m, Wadi Tihama, 23. IV. 1979, H . G.Amse1 leg. ; 2 99, Wadi Shuqub, 1390 m, 21.X. 1979, W.Biittiker; first record!

Distribution: Saudi Arabia, Sinai, Palestine; Africa: Sudan (Kassala).

Genus Syngenes Kolbe Syngenes Kolbe, 1897, Deutsch Ost-Afrika 4: 16 (Typus generis: Acanthaclisis debilis Gerstacker, 1888).

Syngenes arabicus Kimmins Syngenes arabicus Kimmins, 1943 : 154.

Saudi Arabia: 1 9 (holotype) Taif (KIMMINS 1943); not in present material! Distribution: So far Saudi Arabia only.

Genus Centroclisis Navis Centroclisis Navis, 1909 Butll. Inst. catal. Hist. nat. 6 : 71 (Typus generis: Acanthaclisis cervina Gerstacker, 1863).

FAUNA O F SAUDI ARABIA 4, 1982

Centroclisis cervina (Gerstacker) Acanthaclisis cervina Gerstacker, 1863, Stettin. ent. Ztg. 24: 176. Centroclisis cervina (Gerstacker) : HOLZEL (1972).

Saudi Arabia: Riyadh (HOLZEL 1972); 1 d , 1 9, Afif, 8.X. 1975; 1 6 , 6 99 , Arai'da, Selouly's farm 9. X. 1975 Buttiker; 3 99, 5 km W Dammam 30. IX. 1977 A. Pittaway.

Distribution : Saudi Arabia, Sinai, Palestine ; Africa : Egypt, Sudan (Hudeiba).

Centroclisis distincta (Rambur) Acanthaclisis distincta Rambur, 1842, Hist. nat. Ins. NCvr.: 380.

Saudi Arabia: 1 6 , 1 9, Wadi Marah (Asir), 2000 m, 81 km S Biljurshi, 29.IV. 1979 H.G.Amse1; first record !

Distribution: Saudi Arabia; Africa: Egypt, Sudan, Mozambique.

Genus Myrmecaelurus Costa Myrmecaelurus Costa, 1855 Fauna Napol.: 10 (Typus generis: Myrmeleon trigrammus Pallas, 1781).

Myrmecaelurus (Myrmecaelurus) parvulus n. sp. Saudi Arabia: 1 d (holotype), 5 99 (paratypes), Wadi Nisah, 15.IX. 1977; 1 6 , 1 9, Wadi Awsat,

10.VIII. 1978; 1 9, Bahara, 31.VIII. 1977; 2 99 Dammam, 13.IX.1976, W. Buttiker; 1 d , 3 9 9 , 5 km W Dammam; 1 9, Dammam, 5.VIII. 1977,9.IX. 1977, A-Pittaway; 8 d d , 2 99,30 km N Medina, 500 my 27. IV. 1979 KAU/NHMB Exped.

Description of holotype: Length of fore wing 16 mm, of hind wing 14 mm, of body 17 mm. General colour yellow with obscure pale brown markings. Head with a small obscure interantennal spot and two small round spots on epicranium. Pronotum a little longer than broad with traces of three pale brown longitudinal lines, laterals extending only from the base to the median sulcus; meso- and metanotum with obscure pale brown markings which are arranged in three interrupted longitudinal stripes. Wings as in fig.35; venation pale with brown areas, membrane hyaline but nearly all cross veins fringed with pale brown; pterostigma distinct. Hind wing with 3 (left) and 2 (right) presectoral cross veins. Legs yellow, femora with pale, tibiae and tarsi with dark hairs and bristles; tibia1 spurs about as long as the basal tarsal segments. Abdomen clothed with pale silky hairs, with indistinct pale brown lines dorsally and ventrally and a few spots laterally; a pair of pleuritosquamae on segment 6 and 7; apex as in fig.1, gonarcus and parameres as in fig.2.

0 .5 rnrn

Figs 1-2: Myrmecaelurus parvulus n. sp. (holotype 8) : 1, apex of abdomen laterally. 2, gonarcus with parameres, laterally.

The paratypes are rather varying in size and coloration of body and wings. Length of fore wings (d) 18 mm, (9) 18-22 mm; sometimes the membrane may be considerably suffused with pale brown especially on the wing margins, in other examples the wings are nearly pale without brown shadings. The number of presectoral cross veins in hind wings varies from 2-3.

M.parvulus may easily be distinguished from all other known Myrmecaelurus species by its smaller size, the speckled membrane and the number of presectoral cross veins in hind wings.

Myrmecaelurus (Myrmecaelurus) laetus (Klug) Myrmeleon laetus Klug, 1834, Symb. phys. 4 : pl. 36, fig.4. Myrmecaelurus laetus (Klug) : K I M M I N ~ (1943, 1950) ; HOLZEL (1972).

Saudi Arabia: not in present material; Quatar: 1 9, 20 km N Doha, 12.V. 1979, A.Pittaway. Distribution: Saudi Arabia, Iran; Africa: Egypt.

Myrmecaelurus (Myrmecaelurus) acerbus (Walker) Myrmeleon acerbus Walker, 1853, Cat. Neur. Ins. 2 : 366. Myrmecaelurus acerbus (Walker) : HOLZEL (1972).

Saudi Arabia: Riyadh (HOLZEL 1972); numerous spec. from: Wadi Hanifa, 600 m, 25.IV. 1976; Wadi Shaib Luha, 27.V.1976, 30.1X.1976; Hieth, 40 km S Riyadh, 2.V.1976, 13.V.1977; Wadi Khumra, 14.V. 1976, 20.V. 1977, 12.V. and 24.VIII. 1978; Wadi Mizbil, 13.IV. 1977; Kushum a1 Buwaybiyat 27.IV. and 26.V.1978; Bahara, 8.V.1978; Ath Thamamah, 23.V.1978; A1 Khubra 29.V. 1978; Kushm Dibi, 7.IX. 1978, all W.Biittiker; Asir, 1100 my 30 km from Bisha, 25. IV. 1979; Asir, 1150 m Wadi Turabah, 145 km S Taif, 28. IV. 1979; Asir, 2000 my Wadi Marah, 81 km S Biljurshi, 29. IV. 1979, H . G. Amsel.

Distribution : Saudi Arabia, Palestine, Iraq, Turkey, Afghanistan, Pakistan; Africa : Egypt.

Myrmecaelurus (Myrmecaelurus) peterseni Kimmins Myrmecaelurus peterseni Kimmins, 1943 : 149.

Saudi Arabia : Taif (KIMMINS 1943) ; 1 d 1 9 Hieth, 40 km S Riyadh, 2. V. 1976 ; 2 99, Wadi Mizbil, 13.IV.1977; 1 d 1 9, vic. of Riyadh, 27.IV. 1978; 1 9, Salim, 20.1X.1978; 3 d d 1 9 , Jebel an Nir, 990 m, 2.X. 1979; 1 d 4 99, Wadi Tabalah, 19.X. 1979; 2 99, Wadi Shuqub, 1390 my 21.X. 1979, all W.Biittiker; 1 9, Wadi Khumra, 27.IV. 1979, W.Wittmer.

Distribution: So far Saudi Arabia only!

Myrmecaelurus (Myrmecaelurus) lobatus Navls Myrmecaelurus lobatus Navis, 1912, Broteria 10 : 89 : NAVLS (1926).

Saudi Arabia: Kinina, Merda, Hali, Sibbihat (NAVAS 1926); not in present material! I have seen the holotype from Agami, Egypt (MZ) and also one of the specimens which NAVAS

(1926) listed from Egypt (coll. Alfieri, MW). The species bears some superficial resemblance to M . (Nohoveus) zigan Aspock et Aspock et Holzel from East Europe but is easily to distinguish by the strong brown markings of the wings.

Distribution : Saudi Arabia, Palestine ; Africa : Egypt.

Subgenus Nohoveus Navls Nohoveus Navls, 1919, Insecta 8 : 169 (Typus subgeneris : Myrmeleon lepidus Klug, 1834).

FAUNA OF SAUDI ARABIA 4, 1982

Myrmecaelurus (Nohoveus) lepidus (Klug) Myrmeleon lepidus Klug, 1834, Symb. phys. 4: p1.36, fig.9. Nohoveus lepidus (Klug): HOLZEL (1972).

Saudi Arabia: Riyadh (HOLZEL 1972); 1 d , 1 9, Wadi Mizbil, 16.IX.1976, 13.1V.1977; 4 dd , 2 99, Salim, 20. IX. 1978, W. Biittiker.

Distribution : Saudi Arabia; Africa : Egypt, Algeria.

Myrmecaelurus (Nohoveus) philbyi Kimmins Myrmecaelurus philbyi Kimmins, 1943 : 151.

Saudi Arabia: Hafair, Samuda, Taif (KIMMINS 1943); not in present material! I have seen the type series in the collections of British Museum, London. The species very much

resembles M. lepidus (Klug) and most probably may not be more than a pale phenon of this species. Distribution: So far only the type series from Saudi Arabia known.

Myrmecaelurus (Nohoveus) palpalis (Klapilek) Myrmeleon palpalis Klapilek, 1914, Sber. Akad. Wiss. Wien 73: 723. Nohoveus palpalis (Klapilek) : HOLZEL (1972).

Saudi Arabia: Riyadh (HOLZEL 1972); 1 d Arai'da, Selouly's farm, 26.VI. 1975; 1 d , Jebel Banban, 9. VI. 1978 ; 1 d , 2 99, Bahara, 12. VI. 1978,29. VIII. 1979, all W. Buttiker ; 5 68 , 1 9,30 km N Medina, 27. IV. 1979, KAUINHMB Exped.

Distribution : Saudi Arabia, Iran; Africa : Libya, Algeria.

Myrmecaelurus (Nohoveus) saudiarabicus n. sp. Saudi Arabia: 1 d (holotype), 1 d , 3 9 9 (paratypes), Jeddah, 6.V. 1978; 7 d d , 7 99, Bahara, 14.VI.,

24.VIII., 29.IX. 1976,29.VIII. 1979; 1 9, Wadi Arai'da near Hair, 12.VI. 1975; 2 99, Arai'da, Selouly's farm, 9.X.1975; 1 9, Wadi Khumra, 10.1X.1976; 1 9, Unaizeh, 24.IX.1976, all W.Buttiker; 4 99, 16 km W Badr Hunayn, 18.V. 1979 KAUINHMB Exped.; Sudan: 1 S 1 9, Kassala prov., Port Sudan, 23.VI. 1962 R. Remane.

Description of holotype: Length of fore wing 26 mm, of hind wing 23 mm, of body 29 mm. General colour yellowish with dark brown markings. Head luteous with an obscure interantennal spot, with a narrow median stripe and four rounded spots on vertex. Antennae with basal segments brownish, flagellum with pale and brown annulations. Pronotum longer than broad with three narrow longitudinal brown stripes, laterals not reaching the anterior margin; meso- and metanotum also with three dark stripes, laterals interrupted. Wings as in fig. 36; membrane hyaline, posterior margin of both wings and many cross veins in anterior wings fringed with brown; pterostigma distinct. Venation pale with brown areas; hind wings less strongly marked, with 3 (left) and 4 (right) presectoral cross veins. Legs yellowish, femora with large brown areas, hairs and bristles black; tibia1 spurs slightly longer than the basal tarsal segments. Abdomen brown, on each side of tergites 2-5 a broad longitudinal yellow stripe; pale silky hairs, particularly long on the fourth tergite; a pair of pleuritosquamae on segments 6 and 7; apex as in fig.3; gonarcus and parameres as in fig.5.

The paratypes agree in all essential details with the holotype, the intensity of wing coloration being rather variable. Length of fore wings 25-28 mm. Apex of 9 abdomen see fig.4. The species bears some resemblance to Nophis flava HOLZEL, 1972, but may be distinguished by its narrower and more heavy speckled wings.

Figs 3-5 : Myrmecaelurus saudiarabicus n. sp. : 3, apex of abdomen, laterally (holotype 3). 4, apex of abdomen, laterally (paratype 9). 5, gonarcus with parameres, laterally.

Genus Nophis Navis Nophis Navbs, 1912, Mems R. Acad. Cienc. Artes Barcelona 10: 181 (Typus generis: Nophis teillardiNavbs, 1912).

Nophis teillardi Navis Nophis teillardi Navis, 1912, Mems R. Acad. Cienc. Artes Barcelona 10: 182; HOLZEL (1972).

Saudi Arabia: 1 8 , A1 Khubra, 30.V. 1978, W-Biittiker; 18, 2 99, 10 km passed A1 Fresh dir. Medina, 740 m, 20. IV. 1979; 1 8 , 1 9, 11 1 km before A1 Ula, 860 my 21. IV. 1979 all KAU/NHMB Exped. ; Quatar: 1 8 , 28 km S Doha, 28. V. 1979 A. Pittaway.

Distribution: Saudi Arabia, Palestine; Africa: Egypt, Algeria.

Nophis lutea Holzel Nophis lutea Hijlzel, 1972: 20.

Saudi Arabia: Riyadh (HOLZEL 1972); 1 9 Araida Selouly's farm, 26.VI. 1975; 1 d, 6 99, Wadi Shaib Luha, 27.V. 1976; 1 9, Wadi Mizbil, 16.IX. 1976; 2 99, Kushum a1 Buwaybiyat, 26.V. 1978; 2 99, Wadi Khumra, 24.VIII. 1978, 17.VIII.1979; 1 d , 1 9, Kushm Dibi, 7.IX.1978; 5 99, Riyadh, 19.1X.1978; 1 8 , 3 99, Salim, 20.1X.1978; 2 99, Adnan, 21.1X.1978; 1 9, Wadi Raydan, 22.VIII. 1979; 3 dd , Wadi Birk, 10.IX. 1979; 2 99, Wadi Durmah, 25. IX. 1979, all W.Biittiker; 4 dd , 1 9, 16 km W Badr Hunayn, 18.IV. 1979 KAU/NHMB Exped.; 1 9, Hilwah, 3.IX. 1978, W.Sanger.



Nophis flava Holzel Nophis flava Holzel, 1972 : 20.

Saudi Arabia: Riyadh (HOLZEL 1972); 2 99 , Wadi Hanifa, 30.VI11.1976; 1 9, Wadi Mizbil, 16.1X.1976; 1 9 , Hail, 25.IX.1976; 3 99 , Wadi Khumra, 4.VIII.1977, 24.VIII.1978; 3 d d , 3 99 , Wadi Nisah, 16.IX. 1977; 1 d , Hieth, 40 km S Riyadh, 13.V. 1977; 1 d, Wadi Awsat, 10.VIII. 1978 ; 1 9, Wadi Dawasir, 2.1X.1978; 1 9, Sharoura, 29.111.1979, all W.Biittiker; 1 9, 16 km W Badr Hunayn, 18.IV. 1979, KAU/NHMB Exped.

Distribution : Saudi Arabia, Sinai ; Africa : Sudan (Kassala).

Genus Iranoleon Holzel Iranoleon Holzel, 1968: 7 (Typus generis: Iranoleon vartianae HOLZEL, 1968).

Iranoleon darius Holzel Iranoleon darius Holzel, 1972 : 21.

Saudi Arabia: 1 9 Jeddah, 13.11.1979, W. Biittiker ; first record! Distribution: Saudi Arabia, Iran.

Figs 6-9 : Iranoleon arabicus n, sp. : 6, apex of abdomen, laterally (holotype 6). 7, apex of abdomen laterally (paratype 0 ) . 8, gonarcus with parameres, caudally. 9, same, laterally.

Iranoleon arabicus n, sp. Saudi Arabia: 1 d (holotype), 3 99 (paratypes), Wadi Mizbil, 13.IV. 1977,l d , 2 99, Hieth, 40 km

S Riyadh, 2. V. 1976, 13.V. 1977; 3 dd, 4 99, A1 Khardj, 16. IV. 1978 all W. Buttiker. Description of holotype: Length of fore wing 21 mm, of hind wing 19 mm, of body 21 mm.

General colour pale brown with dark patches. Head with brown interantennal mark extending below the antenna1 bases; vertex with a narrow median stripe and two interrupted transverse bands. Anten- nae with pale and brown annulations. Pronotum longer than broad with three narrow longitudinal brown stripes and two spots laterally on hind margin; meso- and metanotum also with three interrupted dark stripes. Wings as in fig.37; membrane hyaline, posterior margin of fore wing slightly suffused with brown; pterostigma distinct; venation pale with brown areas, hind wings generally paler; 4 presectoral cross veins in hind wings. Legs: femora at large parts brownish, tibiae with dark brown rings at apices and in basal half; tarsi pale with apices of segments brown; short pale hairs and black and pale spines; tibia1 spurs of anterior and median legs about as long as the basal tarsal segments. Abdomen dark brown with large luteous patches laterally, a pair of pleuritosquamae on segment, 5, 6 and 7, short dark hairs; apex as in fig.6, gonarcus and parameres as in figs 8, 9.

The paratypes agree in all essential details with the holotype; length of fore wings 20-23 mm. Genital segments of 9 see figs 7, 29.

This is the second known species of the genus Iranoleon with three pairs of pleuritosquamae on 8 abdomen, all other hitherto described species of this genus have pleuritosquamae only on segments 6 and 7.

I.arabicus resembles I, octavus from Palestine but is distinctly smaller (length of fore wings in I . octavus 24 mm). There are also differences in the shape of d and 9 genital structures.

Genus Lopezus NavPs Lopezus Navis, 1913, Novit. zool. 20: 449 (Typus generis: Myrmecaelurus fedtschenkoi McLachlan, 1875).

Lopezus fedtschenkoi arabicus Holzel Lopezus fedtschenkoi arabicus Holzel, 1972 : 24.

Saudi Arabia: Riyadh (HOLZEL 1972); 1 Q Hieth 40 km S Riyadh, 2.V.1976; 3 9, A1 Khardj, 16. IV. 1978 ; 1 9 Riyadh, 19. IX. 1979 all W. Buttiker ; 1 d , Dammam, 7. IV. 1977 A. Pittaway.


Genus Maracanda McLachlan Maracanda Mc Lachlan, 1875 Fedschenko's travels in Turkestan : 2 (Typus generis : Maracanda amoena McLach-

Ian, 1875).

Maracanda lineata NavPs Maracanda lineata Navis, 1913, Novit. 2001. 20: 455; HOLZEL (1972).

Saudi Arabia: 5 9, Dammam, 18.V.1976; 1 d , 1 9, Hieth, 40 km S Riyadh, 13.V.1977; 1 9, A1 Thamamah, 23.V. 1978, 1 9 , Riyadh, 19.IX. 1978, all W.Biittiker; 3 88 , 2 99, Dammam, 3. and 7.V. 1977 A-Pittaway; 8 99, 30 km N Medina 27.IV. 1979, KAU/NHMB Exped.

Distribution: Saudi Arabia, Palestine; Africa: Algeria.

Genus Gepus Navas Gepus Navis, 1912, Mems R. Acad. Cienc. Artes Barcelona 10: 170 (Typus generis: Gepus invisus Navis, 1912).


Gepus invisus Navis Gepus invisus Navis, 1912, Mems R. Acad. Cienc. Artes Barcelona 10: 180; HOLZEL (1972). ? Gepus curvatus Navis, 1914, Mems R. Acad. Cienc. Artes Barcelona 11 : 200; HOLZEL (1968, 1969, 1972).

Saudi Arabia: Riyadh (HOLZEL 1972); 4 dd, 9 99, Bahara, 24.VIII. 1976, 8.V. 1978; 1 9 , Hofuf, 21.VIII.1977 all W.Buttiker; 1 d , 2 99, Dammam, 5.VIII., 6.VIII. and 16.VIII.1977, A-Pittaway; Quatar: 1 d , 23 km N Shahainyeh 1.VI. 1979 A.Pittaway.

N ~ v i i s (1912) based his description of Gepus invisus upon two examples (1 d , 1 9 ) from Egypt. Later on he considered the two syntypes as belonging to two different species and described the 9 under the name Gepus curvatus (Navbs 1914). According to the description this species differs from G. invisus in the markings of the anterior wings : distal parts of RS, base of Cua and posterior Banksian line are strongly outlined in dark brown. Some authors who in the following studied Gepus species nevertheless considered G.curvatus as a synonym of G.invisus (MORTON 1921, ESBEN-PETERSEN 1930). A few years ago I also published records of Gepus species from Anterior Asia and North Africa (HOLZEL 1968, 1969, 1972). Though I was not able to detect any differences in the 8 genital structures I continued to use the name curvatus for specimens with brown lined fore wings.

Quite recently I had the opportunity to study rather large material of Gepus species both from Anterior Asia and North Africa. O n this occasion I found that at least three species appear to have two phena: fore wings with brown stripes ("curvatus-likeJ') or fore wings without brown stripes ("invisus- likeJJ). So arises the question which of these species has Navbs named curvatus? When I checked the Navis-collections in Barcelona and Zaragoza I found there only the d from Egypt which represents undoubtedly the holotype of Gepus invisus. The 9 which Navbs described under the name G. curvatus seems to be lost. It is therefore impossible to state which of the species with striped fore wings Navis named curvatus. So I propose to consider the name Gepus curvatus Navis, 1914 as a nomen dubium.

Distribution: Saudi Arabia, Palestine, Iran; Africa: Egypt, Sudan (Hudeiba).

Gepus cunctatus n. sp. Saudi Arabia: 1 d (holotype), 1 9 (paratype), Wadi Mizbil, 16.IX. 1976; 2 99, W Huraymala

12.VIII.1976; 1 9, Wadi Khumra, 4.VIII.1977; 1 9 , Hofuf, 15.X. 1977; 1 9, Kushm Dibi, 7.IX. 1978; 1 d 2 99, Wadi Raydan, 21.VIII.1979, all W.Buttiker; Israel: 2 99, Ein Gedi, 16.VIII.1957, 18.VI.1958 J.Wahrman leg.; Iran: 1 9 , Prov. Fars, 41 km S Fasa, 13.VII.1965, Neal.

Description of holotype: Length of fore wing 31 mm, of hind wing 28 mm, of body about 24 mm. Pale brown species with blackish markings. Head with an obscure mark between and a fine line below the antenna1 bases; vertex with two interrupted transversal brown bands. Palpi luteous, terminal segment of labial sligthly dilated and with a small brown spot on outer surface. Antennae with pale and brownish annulations. Pronotum about as long as broad, with five longitudinal brown stripes (partly interrupted), laterals extending only from the base to the median sulcus; meso- and metanotum with numerous blackish lines and spots. Wings as in fig.38; membrane hyaline, posterior margins of fore wings sligthly suffused with brown; pterostigma distinct, yellowish with three black veinlets on the proximal side; venation pale with dark brown areas; hind wings less strongly marked, with 4 presectoral cross veins. Legs yellowish with numerous small brown dots; tibiae of two anterior pairs with brown annulations at base, middle and apex, posterior pair at apex only; tarsi pale with apices of segments brown; pale and dark hairs and spines; coxae and femora of anterior legs with prominent long white bristles. Tibia1 spurs about as long as the basal tarsal segments. Abdomen with interrupted dorsal line and rather broad waved lateral lines with short streaks within; apex as in fig. 10; gonarcus and parameres as in figs 12, 13. Apex of 9 abdomen see fig. 11.

Figs 10-13 : Gepus cunctatus n. sp. : 10, apex of abdomen, laterally (holotype 8). 11, apex of abdomen, laterally (paratype 0 ) . 12, gonarcus with parameres, dorsally. 13, same, laterally.

The paratypes are rather varying in the markings of body and wings. 1 ? from Ein Gedi has striped fore wings (brown shadings along the distal part of Rs, the base of Cua and the posterior Banksian line). Length of fore wings 26-32 mm.

Gepus cunctatus n. sp. bears considerable resemblance to G. invisus NavSs and may be distinguished with certainty only with the help of examination of the 8 and 9 genital structures.

Genus Isoleon Esben-Petersen Isoleon Esben-Petersen, 1930, Sber. Akad. Wiss. Wien 102: 204 (Typus generis: Myrmeleon pumilio Klapilek,

1914).


Isoleon arabicus Holzel Isoleon arabicus Holzel, 1972 : 27.

Saudi Arabia: Riyadh (HOLZEL 1972); 1 6, 1 9, Araida Selouly's farm, 9. and 20.X. 1975 ; 1 6 , Wadi Mizbil, 16. IX. 1976, W. Buttiker; 1 6 , Hilwah, 3. IX. 1978, W. Sanger.

Distribution : Saudi Arabia, Palestine.

Genus Cueta Navis Cueta Navis, 1911, Insecta 1 : 242 (Typus generis : Cueta trilineata Navis, 1911).

Cueta lineosa (Rambur) Myrmeleon lineosus Rambur, 1842, Hist. nat. Ins. NCvr.: 389. Cueta lineosa (Rambur): HOLZEL (1968, 1972); ASPOCK, ASPOCK, HOLZEL (1980).

Saudi Arabia: Riyadh (HOLZEL 1972); 60 spec. from: Gizan, airport 24. IV. 1976; Wadi Hanifa, 25.1V.1976; Hieth, 40 km S Riyadh, 2.V.1976; Wadi Khumra, 14.V.1976, 12.V.1978; Wadi Ammariyah, 8. IV. 1977; Wadi Mizbil, 13. IV. 1977; Hofuf, 19. IV., 23.V., 10. X. 1977; Jeddah, 6.V.1978; Bahara, 12.VI.1978; Kushm Dibi, 7.IX.1979; Adnan, 21.1X.1978; Quwayiyah, 16.XI. 1978, all W.Buttiker; Wadi a1 Faraah, N Yumbo, 180 my 19. IV. 1979; 111 km before A1 Ula, 860 m, 21.1V.1979, KAU/NHMB Exped.; Wadi Maraba, 142 km N Gizan 350 m 13.1V.1979; Hakimah near Abu Arish, 35 km from Gizan, 140 my 15.1V.1979; Asir, 1150 m, Wadi Turabah, 145 km S Taif, 28.IV. 1979, H . G.Amse1.

This is a fine series which shows once again the broad band of variability within the populations of this species. It may be of interest to note that the ectoprocts of 6 from Eastern Arabia (Hofuf) and from the opposite side of the Gulf (Bandar Abbas) are distinctly longer and more slender.

Distribution : Saudi Arabia, Palestine, Lebanon, Iraq, Turkey, Cyprus, Iran, Afghanistan, Pakis- tan, Russia; Africa: Egypt, Sudan (Hudeiba), Djibouti; Europe: Greece, Italy, Albania (ASPOCK et al. 1980).

Cueta modesta Holzel Cueta modesta Holzel, 1972 : 30.

Saudi Arabia: 1 9, Bahara, 24.VIII. 1976; 1 6 , Kushm Dibi, 7.IX. 1978; 1 9 , Jeddah, 13.XI. 1979, W. Biittiker ; first record !

Distribution : Saudi Arabia, Iran.

Cueta striata Kimmins Cueta striata Kirnrnins, 1943: 153; HOLZEL (1972).

Saudi Arabia: Zilfi, S.Hedjaz (KIMMINS 1943); Riyadh (HOLZEL 1972); 1 9, Wadi Hanifa, 30.VIII.1976; 1 9, Wadi Salbukh, 29.VIII.1977; 1 9 , Salim, 20.IX. 1978 W.Buttiker; 1 8, Hilwah, 3.IX. 1978, W.Sanger.

Distribution : Saudi Arabia, Kuweit, Iran, Afghanistan.

Cueta klugi n. nom. Myrmeleon variegatus Klug, 1834, Symb. phys. 4: pl. 30, fig.4 - preoccupied by M.variegatum Olivier, 1811. Cueta variegata (Klug): HOLZEL (1969, 1972).

Saudi Arabia: Klug described the species from "Arabia felix"; not in present material. Distribution: Arabia; Africa: Senegal, Sudan (Hudeiba).


distally ; membrane hyaline with brown speckles (fig. 40). 7 and 9 presectoral cross-veins in anterior, 7 and 8 in posterior wings. Legs luteous with small brown dots, especially femora marked with brown; hind femora with long dark hairs above; tibia1 spurs shorter than the basal tarsal segments. Abdomen dark brown with interrupted pale longitudinal stripes laterally; hairiness dense. Apex as in fig. 16, gonarcus with parameres as in fig. 17.

The paratype agrees well with the holotype; length of fore wing 22 mm (see also C.asirica). I have much pleasure in dedicating this species to Dr. H. G. Amsel, Karlsruhe.

Figs 16-1 7 : Cueta amseli n. sp. (holotype d) : 16, apex of abdomen, laterally. 17, gonarcus with parameres, laterally.

Cueta pallens (Klug) Myrmeleon pallens Klug, 1834, Symb. phys. 4: pl. 36, fig. 6. Cueta chryseus (Navis) : KIMMINS (1943). Cueta pallens (Klug) : HOLZEL (1969, 1972).

Saudi Arabia: Taif (KIMMINS 1943); 1 9, Araida, Selouly's farm, X. 1975; 2 dd, 1 9, Wadi Shaib Luna, 27.V. and 30.IX. 1976; 2 dd , 4 99, Wadi Mizbil, 16.IX. 1976; 2 99, Wadi Salbukh, 9.V. 1977; 3 99, Wadi Khumra, 28.IX. 1977,24.VIII. 1978; 1 d , Bahara, 8.V. 1978; 1 d , Quwayiyah, 16.XI. 1978, all W-Biittiker; 1 d 5 99, Wadi Turabah (Asir), 1150 m, 145 km S Taif, 28. IV. 1979, H. G.Amse1.

Distribution: Saudi Arabia, Israel; Africa: Egypt, Sudan (Hudeiba), Tunisia.

Genus Gepella Holzel Gepella Holzel, 1968: 14 (Typus generis: Gepella modesta Holzel, 1968).

Gepella modesta Holzel Gepella modesta Holzel, 1968: 14

Saudi Arabia: 1 9, Kushum a1 Buwaybiyat, 26.V. 1978, W.Biittiker; Quatar: 1 ? , I 4 km W Doha, 13.V. 1979 A.Pittaway; first record!

Distribution: Saudi Arabia, Qatar, Iran, Afghanistan.

Genus Solter Navis Solter Navis, 1912, Broteria 10: 32 (Typus generis: Solter liber Navis, 1912).

Solter hardei Holzel Solter hardei Holzel, 1968 : 17.

Saudi Arabia: 1 9, Wadi Awsat, 1O.VIII. 1978 ; 1 6 1 6 , Wadi Raydan, 22.VIII. 1979, W.Buttiker ; 1 d 1 9, 30 km N Medina, 500 m, 27.IV. 1979, KAUINHMB Exped.; first record!

Distribution: Saudi Arabia, Iran (Baluchistan).

Solter propheticus Holzel Soher pvopheticus Holzel, 1980, Israel J. Ent. 14: 38.

Saudi Arabia: 3 66 , Wadi Huraymala, 12.VIII. 1976; 1 9, Bahara, 24.VIII. 1976; 2 66 1 9, Wadi Mizbil, 16.1X.1976; 1 9, Wadi Khumra, 4.VIII. 1977; 1 d, Wadi Salbukh, 9.VIII. 1977; 2 66 , A1 Thamamah, 23.V. 1978; 1 6 2 99, Salim, 20. IX. 1978, all W.Biittiker; first record!

Distribution : Saudi Arabia, Palestine ; Africa : Sudan (Hudeiba).

Solter wittmeri n. sp. Saudi Arabia: 1 d (holotype), 11 68 1 9 (paratypes) Wadi a1 Faraah, N Yanbo, 180 m, 19. IV. 1979,

KAU/NHMB Exped.; Sinai: 2 99, Phyran 29.VIII. 1967, G.Enaani. Description of holotype: Length of fore wing 20 mm (width 6 mm), of hind wing 19 mm, of body

about 18 mm. General coloration brown. Head luteous, above antennae with a broad brown mark, vertex almost totally brown. Antennae with luteous and brownish annulations. Pronotum fairly exactly as long as broad with traces of three brown longitudinal lines and small obscure spots laterally.

Figs 18-21 : Solter wittmeri n.sp.: 18, apex of abdomen, laterally (holotype d). 19, apex of abdomen, laterally (paratype 0 ) . 20, gonarcus with parameres caudally. 21, same, laterally.


Meso- and matanotum predominantly dark brown. Wings as in fig. 41 ; venation pale with a few brown spots; membrane hyaline, in the anterior wing with traces of pale brown speckles; pterostigma distinct. Legs luteous with pale spines. Femora, tibiae and tarsi brownish at apices, tibiae also with a brown spot at base and a brownish ring in basal half. Tibiae shorter than femora, tarsi shorter than tibiae; spurs of anterior and median legs as long as the tarsal segments 1 +2+3, in posterior legs as long as the segments 1+2. Abdomen brown with luteous patches, apex as in fig. 18; gonarcus and parameres as in figs 20, 21.

The paratypes agree in all essential details with the holotype, length of anterior wings 19-21 mm. Genital segments of ? : apex of sternite 7 and pregenital plate see figs 19, 31.

I have much pleasure in dedicating this species to Dr. W.Wittmer, Basle.

2 3 I 24 0 5 rnrn

Figs 22-24 : Solter buettikeri n. sp. (holotype 6) : 22, apex of abdomen, laterally. 23, gonarcus with parameres, caudally. 24, same, laterally.

Figs 25-28: Solterpallidus n. sp. : 25, apex of abdomen, laterally (holotype 8). 26, apex of abdomen, laterally (paratype 0 ) . 27, gonarcus with parameres, caudally. 28, same, laterally.

I , 0 .5 mm , '

2 9 30 3 1 Figs 29-31 : ? apex of sternite 7, ventrally : 29, Iranoleon arabicus n. sp. : 30, Solter pallidus n. sp. 31, Solter wittmeri n. sp.


Solter buettikeri n. sp. Saudi Arabia : 1 d (holotype), Abha-Gizan km 53 Wadi Ad Dilla 300 m, 21. IV. 1976, W. Buttiker ;

1 d , Asir, Wadi Tihama, 850 m, 23.IV. 1979, H . G.Amse1. Description of holotype: Length of anterior wing 31 mm (width 9 mm), of posterior wing 29 mm,

of body 26 mm. General coloration dark brown. Head luteous, above antennae with a broad brown mark, vertex with numerous irregularly situated brown spots. Antennae with luteous and brown annulations. Pronotum scarcely longer than broad, with traces of three narrow brown longitudinal lines and two large spots on each side. Meso- and metanotum predominantly dark brown with small pale spots. Wings as in fig. 42 ; venation brown with pale areas ; membrane hyaline, in the anterior wing with a few brown speckles at Cua; pterostigma distinct, brown basally, yellowish distally. Legs luteous with pale and a few black (in hind legs) spines, dark spots at apices of femora, tibiae and tarsi; femora (in anterior legs) with a large spot in basal half; tibiae with brown rings at base and in basal half; spurs of anterior and median legs as long as the tarsal segments 1 +2 + 3, in posterior legs as long as segments 1 +2. Abdomen luteous with large brown patches dorsally ; apex as in fig. 22 ; gonarcus and parameres as in figs 23, 24.

The paratpye agrees in all details with the holotype, length of anterior wing 32 mm. I have much pleasure in dedicating this species to Prof. W.Buttiker, Riyadh.

Solter pallidus n. sp. Saudi Arabia: 1 d (holotype), Wadi Khumra, 24.VIII. 1978, 1 ? (paratype), Wadi Shaib Luha,

30.IX. 1976, W. Buttiker. Description of holotype : Length of fore wing 28 mm, of hind wing 24 mm, of body about 25 mm.

General coloration pale brown. Head with a brown mark above antennae, vertex with irregularly situated brownish spots. Antennae pale brown with brownish annulations. Pronotum rather as long as broad, with three narrow interrupted longitudinal lines, with short brown hairs and pale bristles laterally. Meso- and metanotum predominantly pale with small brownish spots. Wings as in fig. 43; membrane hyaline without spots and shadings; venation almost totally pale, only in fore wings with small brown dots and lines, pterostigma distinctly brown. Legs pale brown with pale and a few black (in hind legs) spines; spurs of anterior and median legs slightly longer than the tarsal segments 1 +2, in posterior legs longer than the basal tarsal segment. Abdomen pale brown with large brown patches dorsally; apex as in fig.25; gonarcus and parameres as in figs 27, 28.

The ? paratype agrees well with the holotype; length of anterior wing 29 mm; genital structures: apex of sternite 7 and pregenital plate as in figs 26, 30.

The Solter species of Saudi Arabia may be distinguished with the help of the following key:

1 Anterior wings distinctly longer than 25 mm . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 - Anterior wings shorter than 25 mm . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 2 Posterior legs : spurs distinctly longer than the two basal tarsal segments

S.propheticus Holzel - Posterior legs: spurs shorter or as long as the two basal tarsal segments . . . . . . . . . . . . . . . . 3 3 Body and wings marked with dark brown spots and patches S. buettikeri n. sp. - Body and wings pale brown S.pallidus n. sp. 4 Anterior legs : spurs as long as the two basal tarsal segments S. hardei Holzel - Anterior legs : spurs as long as the three basal tarsal segments S. wittmeri n. sp.

Genus Myrmeleon Linnaeus Myrmeleon Linnaeus, 1767, Syst. Nat. 12: 913 (Typus generis: Myrmeleon formicarius Linnaeus, 1767).

Subgenus Morter Navis Mems R. Acad. Cienc. Artes Barcelona 11 : 466

Morter Navis, 1915, (Typus generis: M~rrneleon h~al inus Olivier, 1811).

Myrmeleon (Morter) hyalinus Olivier Myrmeleon hyalinus Olivier, 181 1, Encycl. meth. 8 : 126. Myrmeleon (Morter) hyalinus Oliver: HOLZEL 1972; ASPOCK, ASPOCK, HOLZEL (1980).

Saudi Arabia: 1 9, Wadi Durmah, 29.1V.1976; 1 9, Wadi Khumra, 14.V.1976; 1 d, Bahara, 20.XII. 1976; 1 9, Wadi A1 Ammari~ah, XII. 1977; 1 9, Maraba, 30. IX. 1978 ; 9, Jeddah, 13.11.1979, all W.Buttiker; 2 99, 16 km W Badr Hunayn, 18.IV. 1979, KAU/NHMB Exped.; 1 9, Asir, 1150 my Wadi Turabah, 145 km S Taif, 28.IV. 1979, H.G.Amse1; 1 9, Dammam, 5.XI.1977; 1 9, Khobar, 11. XI. 1977, A. Pittaway.

Distribution: Saudi Arabia, Palestine, Lebanon, Iraq, Turkey, Cyprus, Iran, Afghanistan, Russia; Africa: Egypt, Algeria, Morocco ; Canary Islands ; Europe : Spain, Italy, Greece (ASPOCK et al. 1980).

Myrmeleon (Morter) fasciatus (Navis) Nesoleon fasciatus Navis, 1912, Annls Soc. scient. Brux. 36: 238. My~meleon (Morter) fasciatus (Navis): HOLZEL (1972); ASPOCK, ASPOCK, HOLZEL (1980).

Saudi Arabia: 3 dd , Wadi a1 Ammari~ah, 18.IV. 1977; 1 9, Ath Thamamah, 27.X. 1978,W.But- tiker; 1 9, 30 km N Medina, 500 my 27.1V.1979 KAUINHMB Exped.; 1 9, Wadi Marah (Asir), 2000 my 81 km S Biljurshi, 29.IV. 1979, H . G.Amse1.

Distribution: Saudi Arabia, Palestine, Sinai; Africa: Egypt, Algeria; Europe: Greece (Rhodes) (Aspock et al. 1980).

Genus Mesonemurus Navis Mesonemurus Navis, 1919, Novit. 2001. 26.: 283 (Typus generis: Mesonemurus harterti Navis).

Mesonemurus harterti Navis Mesonemurus harterti Navis, 1919, Novit. zool. 26: 284; HOLZEL (1972).

Saudi Arabia: 1 9, Wadi A1 Ammariyah, 18. IV. 1977, W. Buttiker; first record! Distribution: Saudi Arabia, Palestine, Syria, Iraq, Iran, Afghanistan, Pakistan; Africa: Egypt,

Algeria, Tunisia.

Genus Geyria Esben-Petersen Geyria Esben-Petersen, 1920, Arch. Naturgesch. 84: 146 (Typus generis: Geyria saharica Esben-Petersen, 1920).

Geyria saharica Esben-Petersen Geyria saharica Esben-Petersen, 1920, Arch. Naturgesch. 84: 147.

Saudi Arabia: 1 8, Bahara, 12.VI. 1978, W.Buttiker; first record! I have compared this specimen with the holotype (1 d , Aim Taiba, Algeria, coll. MB). The species

bears some resemblance to G. belutschistana Holzel, from Afghanistan and Iran but may be distinguished by its narrower wings and the shorter d ectoprocts.

Distribution: Saudi Arabia, Palestine; Africa: Sudan (Kassala), Algeria, Tunisia.


Geyria lepidula (Navis) Macronemurus lepidulus Navis, 1912, 8. Congr. 2001. Graz: 748 Geyria lepidula (Navis): HOLZEL (1972).

Saudi Arabia: Riyadh (HOLZEL 1972); numerous spec. from: Araida, Selouly's farm, 30.1.1975; Afif, 8.X. 1975; Bahara, 24.VIII. 1976, 31.VIII. 1977; Wadi Mizbil, 16. IX. 1976; Hail 25.IX. 1976; Wadi Salbukh, 29.VIII. 1977; Wadi Nisah, 16.IX. 1977; Wadi Awsat, 10.VIII. 1978; Kushm Dibi, 7. IX. 1978 ; Wadi Raydan, 22.VIII. 1979; Todiah, 9. IX. 1979; Wadi Birk, 10. IX. 1979; Wadi Durmah, 25.IX. 1979; Wadi Khumra, 28. IX. 1979; Jebel an Nir, 990 my 2.X. 1979; Wadi Tabalah, 19.X. 1979; Wadi A1 Kharj, 22.X. 1979, all W.Biittiker.

Distribution: Saudi Arabia, Sinai, Palestine, Iran; Africa: Egypt, Sudan (Hudeiba).

Genus Macronemurus Costa Macronemurus Costa, 1855, Fauna Napol.: 8 (Typus generis: Myrmeleon appendiculatus Latreille, 1807).

Macronemurus delicatulus Morton Macronemurus delicatulus Morton, 1926, Trans. ent. Soc. Lond. 1925: 409; HOLZEL (1972). Saudi Arabia: 1 9, Hieth, 40 km S Riyadh, 13.V. 1977; 1 9, Kushm Dibi, 20.IV. 1978; 3 66 3 99,

Kushum a1 Buwaybiyat, 27. IV. and 26.V. 1978; 1 9, Wadi Khumra, 12.V. 1978, all W. Buttiker; 1 9, 23 km N Shahaniyeh, l.VI.1979; Quatar: 1 9, 14 km W Doha, 13.V. 1979 A.Pittaway; first record!

Distribution: Saudi Arabia, Qatar, Palestine; Africa: Algeria.

Genus Quinemurus Kimmins Quinemurus Kimmins, 1943: 148 (Typus generis: Quinemurus cinereus Kimmins, 1943).

Quinemurus cinereus Kimmins Quinemurus cinereus Kimmins, 1943: 148; HOLZEL (1972).

Saudi Arabia: S.Hedjaz, Qui'iya (KIMMINS 1943); Riyadh (HOLZEL 1972); 1 6 1 9 , Arai'da, Selouly's farm, 20.X. 1975; 1 9, Hail, 25.IX. 1976; 2 86 1 9, Salim, 20.IX. 1978, all W.Buttiker; 2 99 , Hafar, 4.X. 1977; 2 99, Dammam, 3.XI. and 5.XI. 1977, A.Pittaway.

Distribution : Saudi Arabia, Iran.

Genus Ganguilus Navbs Ganguilus Navis, 1912, Broteria 10: 72 (Typus generis: Ganguilus pallescens Navis, 1912). Cordeses Navis, 1914, Memorie Accad. pont. Nuovi Lincei 32: 94 (Typus generis: Cordeses oblitusNavis, 1914) -

new synonymy!

Ganguilus pallescens Navis Ganguilus pallescens Navis, 1912, Broteria 10: 72; HOLZEL (1972). Cordeses oblitus Navis, 1914, Memorie Accad. pont. Nuovi Lincei 32: 95, new synonymy!

Saudi Arabia: 3 99, Hail, 25.IX. 1976; 9 99, Salim, 20.IX. 1978; 1 9, Adnan, 21.IX. 1978, all W.Buttiker; 1 9, 5 km W Dammam, 25.IX. 1977; 1 9 , Hafar-al-Batin, 5.X. 1977, A. Pittaway ; first record !

I have seen the holotype of G.pallescens (1 9 from Nigeria, Zungeno, coll. BM) and of Cordeses oblitus (1 9 Obock, coll. MP); both belong to the same species.

Distribution: Saudi Arabia, Palestine, Iran; Africa: Egypt, Sudan (Hudeiba), Nigeria, Somalia.

Genus Distoleon Banks Distoleon Banks, 1910, Ann. ent. Soc. Am. 3 : 42 (Typus generis: Distoleon verticalis Banks, 1910).

Distoleon laticollis (Navls) Formicaleo laticollis Navis, 1913, Ent. obozr. 13 : 278. Formicaleo alienus Navis, 1913, Mitt. miinch. ent. Ges. 4: 12, new synonymy! Distoleon laticollis (Navis) : HOLZEL (1972).

Saudi Arabia: 1 9, Jeddah-Taif, 1200-1800 my 1.V. 1979, KAUINHMB Exped. ; 1 9, Wadi Marah, 2000 m, (Asir), 81 km S Biljurshi, 22.IV. 1979, H.G.Amse1; first record!

I have examined the holotype of Formicaleo laticollis (1 9 "Syria", coll. MG) and of Formicaleo alienus (1 9 Beirut, Lebanon, coll. MM); both are undoubtedly the same species.

Distribution : Saudi Arabia, Palestine, Lebanon, Syria, Turkey ; Africa : Sudan (Kassala), Ethiopia.

Genus Pseudoformicaleo Van der Weele Pseudoformicaleo Van der Weele, 1908, Notes Leyden Mus. 31 : 25 (Typus generis: Myrmeleon gracilis Klug,

1834).

Pseudoformicaleo gracilis (Klug) Myrmeleon gracilis Klug, 1834, Symb. phys. 4 : pl. 35, fig. 5. Pseudoformicaleo gracilis (Klug): HOLZEL (1972).

Saudi Arabia: 1 8, Dammam, 16.VIII. 1976, W.Buttiker ; 1 9, Jeddah-Taif, 1200-1800 my 1 .V. 1979 KAUINHMB Exped. ; first record !

Distribution: Saudi Arabia, Palestine, Syria, Iran; Africa: Algeria.

Genus Neuroleon Navis Neuroleon Navis, 1909, Mem. I. Congr. Nat. Esp., Zaragoza 1 : 148 (Typus generis: Myrmeleon arenarius Navis,

1904).

Neuroleon leptaleus (Navls) Ganussa leptalea Navis, 1912, Insecta 2: 222. Neuroleon leptaleus (Navis): HOLZEL (1972).

Saudi Arabia: Riyadh (HOLZEL 1972); 37 spec. from: Wadi Durmah, 9.V. 1976; Wadi Khumra, 10. IX. 1976, 20. V. 1977, 27. IV. 1979; Wadi Mizbil, 16. IX. 1976, 13. IV. 1977; Wadi A1 Ammariyah, 18.IV. 1977; Hieth, 40 km S Riyadh, 13.V. 1977; Wadi Salbukh, 9.1X.1977; A1 Khubra, 15.X. 1978; Rumah, 7.XI. 1979, all W.Buttiker; 30 km N Medina, 500 my 27.IV. 1979, KAUINHMB Exped.; 5 km W Dammam, 25. IX. 1977, A. Pittway.

Distribution : Saudi Arabia, Palestine, Iraq, Iran; Africa : Algeria, Tunisia.

Neuroleon tenellus (Klug) Myrmeleon tenellus Klug, 1834, Symb. phys. 4: pl.35, fig.7. Neuroleon tenellus (Klug): HOLZEL (1972); ASPOCK, ASPOCK, HOLZEL (1980).

Saudi Arabia: Riyadh (HOLZEL 1972); 2 99, Hofuf, 23.V. and 9.X. 1977, W.Biittiker. Distribution: Saudi Arabia, Palestine, Lebanon, Iraq, Turkey, Iran, Afghanistan; Africa: Egypt,

Sudan (Hudeiba), Tunisia; Europe: Greece (Crete, Naxos) (ASPOCK ET AL. 1980).

Neuroleon taifensis Kimmins Neuroleon taifensis Kimmins, 1943 : 145.


Saudi Arabia: Taif (KIMMINS 1943); 2 88 1 9 , Wadi Khumra, 16.1X.1976, 28.1X.1977, 24.VIII. 1978 ; 6 99, Wadi Shaib Luha, 30. IX. 1976; 1 9, Quwayiyah, 16. IX. 1978, all W. Buttiker.

Distribution : Saudi Arabia, Palestine.

Neuroleon longipennis (Esben-Petersen) Oligoleon longipennis Esben-Petersen, 1930, Sber. Akad. Wiss. Wien 102: 202. Neuroleon longipennis (Esben-Petersen): HOLZEL (1972).

Saudi Arabia: Jeddah (coll. BM); 1 ? , I 6 km W Badr Hunayn, 18. V. 1979, KAU/NHBM Exped. Distribution : Saudi Arabia; Africa : Sudan.

Neuroleon pardalice (Banks) nov. comb. Nemoleon pardalice Banks, 1911, Ann. ent. Soc. Am. 4: 21.

Saudi Arabia: 1 9, Adama (Asir), 22.1X.1978, W.Buttiker; 1 9, Wadi Tihama (Asir), 850 m, 23.IV. 1979, H . G.Amse1; first record!

Distribution: Saudi Arabia; Africa: Sudan (Kassala), Ethiopia.

Neuroleon parvus Kimmins Neuroleon parvus Kimmins, 1943 : 146.

Saudi Arabia: Mastura (KIMMINS 1943) ; numerous spec. from : Bahara, 24.VIII., 29. IX., 26.X. 1976, 26. IV. 1977, 12.VI. 1978, 30. I., 29,VIII. 1979; Jeddah, 7.V. 1978, all W. Buttiker; 16 km W Badr. Hunayn, 18. IV. 1979, KAUINHMB Exped.

Distribution : Saudi Arabia only !

Neuroleon erato Holzel Neuroleon erato Holzel, 1972 : 53.

Saudi Arabia: 1 9, Wadi Shaib Luha, 27.V.1976; 1 9, Wadi Mizbil, 10.1X.1976; 1 9, Wadi Raydan, 22.VIII. 1979, all W.Buttiker; 1 8 , 30 km N Medina, 500 m, 27. IV. 1979, KAU/NHMB Exped.; first record!

Distribution: Saudi Arabia, Palestine, Iran, Afghanistan, Pakistan.

Neuroleon lugubris (Navls) nov. comb. Gymnoleon lugubris Navis, 1926, Bull. Soc, ent. Egypte 19: 36.

Saudi Arabia: 2 99, Adnan, 21.IX.1978; 1 9, Adama (Asir), 22.1X.1978; 1 8 5 99, Wadi Turabah, Camp 2, 1580 m, 7.4.X.1979; 1 9, Maraba, 16.X.1979; 3 99, Wadi Shuqub, 1390 m, 21. X. 1979, all W. Buttiker; 4 99, Wadi Tihama (Asir), 850 m, 23.IV. 1978 ; 1 8 2 99, Wadi Maraba (Asir), 350 m, 142 km N Gizan, 13.IV. 1979; 4 88 3 99, Wadi Turabah (Asir), 145 km S Taif, 1150 m, 28. IV. 1979, all H . G. Amsel.

The absence of tibial spurs caused Navls to place this species in the genus Gymnoleon Banks, 191 1. As he did not notice that the reduction and loss of tibial spurs apparently has occured in many evolutionary lines of the family (in different tribes) he established for all "spurless" genera the tribe Gymnocemini (NAVLS 1926).

The syntypes ( ? 9 9 from Kinina and Merda in "South Arabia") seem to be lost, at least they are not deposited in the Alfieri collection (coll. MW, personal communication Dr. 0. S. Flint.) The species can be recognised with the help of the description and the figure of thorax, but above all by the absence of tibial spurs. So I hope to be right when I consider the present specimens to belong to this species which accordingly is to be placed in the genus Neuroleon Navis.

Distribution : Saudi Arabia, Palestine ; Africa : Sudan (Kassala).

G e n u s Creoleon Ti l lyard

Creoleon Tillyard, 1918, Proc. Linn. Soc. N. S. W. 43 : 436 (Typus generis: Myrmeleon lugdunensis Villers, 1789).

Creoleon griseus (Klug)

Myrmeleon griseus Klug, 1834, Symb. phys. 4: p1.31, fig. 8. Creoleon griseus (Klug): HOLZEL (1972).

Saudi A r a b i a : 1 9, A1 K h a r d j , 19. IX. 1975, W . B u t t i k e r ; f i rs t record! D i s t r i b u t i o n : Saudi Arab ia , Palest ine, I r a n , N - I n d i a ; Afr ica : E g y p t , Sudan ( H u d e i b a ) .

Creoleon parallelus (Klap i lek) nov. comb. Myrmeleon irroratus Klug, 1834, Symb. phys. 4 : pl. 35, fig. 6, preoccupied by Myrmeleon irroratus Olivier, 1811. Creagris parallelus Klapilek, 191 1, Ent. 2. 25: 76. Creagris surcoufi Navis, 1912, Insecta 2: 218, new synonymy! Creoleon klugi Navis, 1926, Bull. Soc. ent. Egypte 19: 50, nov. nom. for M.iworatus Klug, new synonymy! Creoleon iworatus (Klug) : HOLZEL (1972).

Saudi Arab ia : R i y a d h (HOLZEL 1972); 2 99, W a d i K h u m r a , 14.V. 1976 a n d 24 .VII I .1978; 1 9 Bure ida , 28.V. 1978, W. But t iker .

F A U N A OF SAUDI ARABIA 4, 1982 267

I have examined the holotype of C. surcoufi Navls (1 9 Laghouat, coll. MZ) and compared it with the syntypes of Myrmeleon irroratus Klug from Arabia (coll. MB). They belong to the same species which Klapilek described from Ethiopia under the name parallelus.

Distribution: Saudi Arabia, Palestine, Iran; Africa: Egypt, Sudan (Hudeiba, Kassala), Ethiopia, Algeria, Tunisia.

Creoleon desertus n. sp. Saudi Arabia: 1 d (holotype), 3 99 (paratypes), Wadi A1 Ammariyah, 18.IV. 1977; 1 9, Wadi

Hanifa, 600 m, 25.IV. 1976; 2 99, Salim, 20. IX. 1978; 1 9, A1 Hunayy, 22.X. 1978, all W.Biittiker; 1 9, Wadi Khumra, 27.IV. 1979, W.Wittmer; 2 99, Sinai, Wadi Firan, 27.IX. 1977, D. Simon.

Description of holotype: Length of fore wing 33 mm, of hind wing 31 mm, of body 32 mm. General colour pale brown. Head with a brownish mark behind antennae, vertex with large brown spots. Antennae pale brown. Pronotum scarcely longer than broad with two narrow median stripes in caudal half and indistinct brownish shadings laterally; short pale hairs and long pale bristles laterally; meso- and metanotum with interrupted brown stripes laterally and brown spots on prescutum and mesoscutellum. Wings as in fig. 44 ; membrane hyaline without shadings ; venation totally pale ; pterostigma indistinctly brownish. Legs pale brown with black (and a few pale) hairs and bristles; spurs of anterior and median legs as long as the tarsal segments 1 +2, in posterior legs sligthly longer than the basal tarsal segments. Abdomen pale with a median dark line dorsally and indistinct dark areas laterally; apex as in fig.32, gonarcus and parameres as in figs 33, 34.

The paratypes agree in all essential details with the holotype, some specimens show an obscure interantennal spot; length of fore wings 31-36 mm. C. desertus n.sp. resembles C. remanei Holzel, 1972, from Iraq, but differs in the length of d abdomen and of tibial spurs. In C.remanei the d abdomen is distinctly longer than the wings and the tibial spurs (of anterior legs) are as long as the four basal tarsal segments together.

Creoleon antennatus (Navis) nov. comb. Creagris antennata Navis, 1914, Mems R. Acad. Cienc. Artes Barcelona 10: 20 Creoleon indigus Navis, 1926, Bull. Soc. ent. Egypte 19: 52, new synonymy!

Saudi Arabia: 2 d d 1 9 , Wadi Mizbil, 16.IX. 1976 and 13.IV. 1977; 1 d , Wadi Shaib Luha, 27.V.1976; 1 d, Wadi Salbukh, 15.1V.1977; 1 9, Kushm Dibi, 7.1X.1978; 1 9, Quwayiyah,

Figs 35-44 : Wings o f : 35, Myrmecaelurusparvulus n. sp. (paratype 0). 36, Myrmecaelurus saudiarabicus n. sp. (paratype 0 ) . 37, Iranoleon arabicus n.sp. (holotype 8). 38, Gepus cunctatus n.sp. (paratype 0 ) . 39, Cueta asirica n.sp. (paratype 6). 40, Cueta amseli n.sp. (paratype d41, Solter wittmeri n. sp. (paratype d). 42, Solter buettikerin.sp. (paratype 8). 43, Solter pallidus n. sp. (paratype 0). 44, Creoleon desertus n. sp. (holotype 8).

FAUNA OF SAUDI ARABIA 4,1982 269

16.XI. 1978, all W.Buttiker; 1 S, Wadi a1 Faraah N Yambo, 180 my 19.IV. 1979; 1 9,111 km before A1 Ula, 860 my 21. IV. 1979; 1 9, 155 km before Tayma road to Tabuk, 1150 my 22.IV. 1979; 1 ? , I 7 km before Khaybar, Wadi Ghamra, 660 my 25.IV. 1979, KAU/NHMB Exped.; 1 9 , Hilwah, 3.IX.1978, W. Siinger; 1 9, Asir, 1150 my Wadi Turabah, 145 km S Taif, 28. IV. 1979, H . G. Amsel; first record!

I have examined the holotypes of Creagris antennata (1 S Egypt, Kossei'r, coll. MG) and Creoleon

indigus (1 9 Egypt, Wadi Digla, coll. MZ), both belong to the same species. Distribution : Saudi Arabia, Palestine, Sinai, Iran; Africa : Egypt, Sudan.

Creoleon elegans Holzel Creoleon elegans Holzel, 1968: 27; HOLZEL 1972.

Saudi Arabia: Riyadh (HOLZEL 1972); 2 99, Arai'da Selouly's farm, 20.X. 1975; 1 9, Wadi Khumra, 14.V.1976; 1 9 , Wadi Shaib Luha, 27.V.1976; 2 d S 2 99, Bahara, 24.VIII.1976 and 30.1.1979; 1 9, Wadi Mizbil, 16.IX. 1976; 2 SS 1 9, Wadi Ammariyah, 18.IV. 1977; 3 99, Hieth, 40 km S Riyadh, 13.V. 1977; 1 9, A1 Hunayy, 22.X. 1978, all W.Biittiker.

Distribution : Saudi Arabia, Syria, Iraq, Iran, Pakistan.

Creoleon persicus Holzel Creoleon persicus Holzel, 1972 : 62.

Saudi Arabia : 1 S 5 99, Adama (Asir), 22. IX. 1978; 1 9, Ash Sharayi, 24. IX. 1978, W. Buttiker ; 1 9, 155 km before Tayma, road to Tabuk, 1150 my 22.IV.1979, KAU/NHMB Exped.; 1 S 1 9, Asir, Wadi Turabah, 1150 my 145 km S Taif, 28.IV. 1979, H . G. Amsel; first record!

Distribution: Saudi Arabia, Palestine, Iran, Afghanistan.

Genus Noaleon Holzel Noaleon Holzel, 1972 : 64 (Typus generis : Neuroleon limbatellus Navis, 1913).

Noaleon limbatellus (Navis) Neuroleon limbatellus Navis, 1913, Novit. 2001. 20: 451. Noaleon limbatellus (Navis) : HOLZEL ( 1 972).

Saudi Arabia: 1 8 , Afif, 8.X. 1975; 1 9, Bahara, 8.V. 1978, W.Biittiker; 2 SS, 111 km before A1 Ula, 860 my 21.IV. 1979, KAU/NHMB Exped.; first record!

Distribution : Saudi Arabia, Palestine, Iran; Africa: Sudan (Kassala), Algeria.

CONCLUSION

As far as can be judged from the hitherto examined Myrmeleonidae of Saudi Arabia the fauna of this country consists mainly of elements of the Palearctic Region. Within the 70 recorded species we find only 8 which most probably represent elements of the Ethiopian Region: Palpares papilionoides,

l? klugi, l? walkeri, Centroclisis distincta, Cueta klugi, Neuroleon pardalice, N. lugubris and N. lon-

gipennis. N o elements of the Oriental Region are known at present. If we try a preliminary zoogeographical analysis of the Palearctic species we find 27 species with

Central Eremian distribution (14 species only in Saudi Arabia - 6 of them only in the western mountain region, 13 species from Saudi Arabia and other countries of the Arabian Peninsula - 8 of them also in Northeast-Africa), 14 species with Western Eremian distribution (8 Saharo-Arabian, 6 Saharo-Iranian) and 16 species with Eastern Eremian distribution. 5 species - Cueta lineosa, Myrme-

leon hyalinus, Mesonemurus harterti, Ganguilus pallescens and Neuroleon tenellus represent apparently Paneremian elements, but this classification should be treated with caution: a further examination of more material from other regions will most probably show that these species (or at least some of them) are represented in Saudi Arabia by more or less well distinguishable subspecies.

REFERENCES

ASPOCK, A,, ASPOCK, U., HOLZEL, H . (unter Mitarbeit von H.RAUSCH), 1980. Die Neuropteren Europas. 2 Bde. Goecke u. Evers, Krefeld.

ESBEN-PETERSEN, P., 1930. Wissenschaftliche Ergebnisse der mit Unterstutzung der Akademie der Wissenschaften in Wien aus der Erbschaft Treitl von F. Werner unternommenen zoologischen Expedition nach dem Anglo-Agypti- schen Sudan (Kordofan) 1914. XVZZ. Neuropteren. Denkschr. Akad. Wiss. Wien 102: 201-206.

HOLZEL, H. , 1968. Zur Kenntnis der Myrmeleoniden des Iran (Planipennia, Myrmeleonidae). Stuttg. Beitr. Naturk. 181: 1-32.

HOLZEL, H., 1969. Beitrag zur Systematik der Myrmeleoniden (Neuroptera-Planipennia, Myrmeleonidae). Annln naturh. Mus. Wien 73: 275-320.

HOLZEL, H. , 1972. Die Neuropteren Vorderasiens I V . Myrmeleonidae. Beitr. naturk. Forsch. SiidwDtl. 1 : 3-103. HOLZEL, H., 1980. Notes on Ant-lions of Israel and nearby countries, together with descriptions of 11 new species

Planipennia, Myrmeleonidae). Israel J . Ent. 14: 29-46. KIMMINS, D.E., 1943. N e w species of Arabian Myrmeleonidae. Ann. Mag. Nat. Hist. (11)lO: 145-156. KIMMINS, D.E., 1950. Results of the Armstrong college expedition to Siwa Oasis (Libyan Desert), 1935. Under the

leadership of Prof. J. Omer-Cooper. Odonata and Neuroptera. Bull. Soc. Fouad I. Ent. 34: 151-157. KIMMINS, D.E., 1961. The Odonata and Neuroptera of the island of Socotra. Ann. Mag. Nat. Hist. (13)3: 385-392. KLUG, J . C. F., 1834. Symbolae physicae, sei icones et descriptiones insectorum, quae ex itinere per African borealem et

Asiam F. G. Hemprich et C . H . Ehrenberg studio novae aut illustrate redierunt. Bd. 4. Berlin. MORTON, K. J., 1921. Neuroptera, Mecoptera and Odonata from Mesopotamia and Persia. Entomologist's mon. Mag.

7: 213-222. N A V ~ S , L. 1912. Znsectos Neuropteros nuevos opoco conocidos. Mems. R. Acad. Cienc. Artes Barcelona 10: 135-202. NAVAS, L., 1914. Neuropteros nuevos o poco conocidos I I I . Mems R. Acad. Cienc. Artes Barcelona 11: 193-205. NAVAS, L., 1915. Neuropteros nuevos o poco conocidos IV . Mems R. Acad. Cienc. Artes Barcelona 11: 373-400. N A V ~ S , L., 1926. NivroptPres dYEgypte et de Palestine ZZ. Bull. Soc. ent. Egypte 19: 26-62. NAVAS, L., 1934. Insectos del Museo de Hamburgo. Mems R. Acad. Cienc. Artes Barcelona 23: 499-506. RAMBUR, P., 1842. Niuroptires in: Histoire naturelle des insectes. 529 pp., Paris. STITZ, H . , 1912. Palpares aus der Sammlung des Berliner Museums. Mitt. 2001. Mus. Berl. 6: 105-116.

Author's address : Dr. Herbert Holzel, Annenheim 160, A-9520 Sattendorf

Bibliography of the Neuropterida Bibliography of the Neuropterida Reference number (r#): 2988 Reference Citation: Hölzel, H. 1982 [1982.??.??]. Insects of Saudi Arabia. Neuroptera: Fam. Myrmeleonidae. Fauna of Saudi Arabia 4:244-270. Copyrights: Any/all applicable copyrights reside with, and are reserved by, the publisher(s), the author(s) and/or other entities as allowed by law. No copyrights belong to the Bibliography of the Neuropterida. Work made available through the Bibliography of the Neuropterida with permission(s) obtained, or with copyrights believed to be expired. Notes: This work made available through the Bibliography of the Neuropterida through the kind permission of Syngenta. File: File produced for the Bibliography of the Neuropterida (BotN) component of the Global Lacewing Digital Library (GLDL) Project, 2006.

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Insects of Saudi Arabia Neuroptera Fam. Myrmeleonidae