INTERNATIONAL LICHENOLOGICAL NEWSLETTER … · The International Lichenological Newsletter is the official publication of IAL. ... (SPM), Universidad del ... Kondratyuk’s presentation

Official publication of theInternational Association for Lichenology

Editor:A. SUIJA

University of Tartu, Lai street 36-40, Tartu, EE51005, [email protected], phone (+372) 7376 177

Editorial Board:

P. SCHOLZ (Schkeuditz) , M.R.D. SEAWARD (Bradford),ISSN: 0731 2830

The opinions expressed in the Newsletter are not necessarily those held by the InternationalAssociation for Lichenology

INTERNATIONAL

LICHENOLOGICAL

NEWSLETTER Vol. 49, no. 1, July 2016

International Association for LichenologyThe International Association for Lichenology (IAL) promotes the study and conservation oflichens. It organizes symposia, field trips, and distributes a biannual newsletter. There is alistserver that enables on-line discussion of topics of interest. Webpages devoted to lichenologyare also maintained by members of the Association. People wishing to renew their membershipor become members of IAL are requested to send their subscription (one payment of 40 USD for2012-2016) to either Treasurers.The International Lichenological Newsletter is the official publication of IAL. It is issuedtwice a year (July and December) in English. The Newsletter is also available on the Internet. TheNewsletter is divided into fourmain sections:1)Associationnews:official informationconcern-ing the Association, such as minutes of Council meetings, proposals of Constitutional changes,newmembers, changesof addresses, etc. 2)News: informationabout lichenologists, institutionalprojects, herbaria, requests of collaboration, announcements of meetings, book reviews, etc. 3)Reports: reports of past activities, short lectures, obituaries, short historical novelties, etc. 4)Reviews: presentation of recent progress and other topics of interest in lichenology with optionaldiscussion. When the material exceeds the available space, the Editor will prepare a summary,on prior agreement with the contributors.Any information intended for publication should reach the Editor on or before June 15 andNovember 15 for inclusion in the July and December issues, respectively.

IALaffairs aredirectedbyanExecutiveCouncil electedduring the lastGeneralMeeting.Councilmembers elected at the IAL7 Symposium (Bangkok, Thailand, 2012) are listed below, and willserve until 2016.

IAL Council 2012-2016President: Helge Thorsten Lumbsch, The Field Museum of Natural History, Department of

Botany, 1400 S. Lake Shore Drive, Chicago, IL 60605-2496, USA. E-mail:[email protected]

Vice President: Mats Wedin, Swedish Museum of Natural History, Department of CryptogamicBotany, P.O. Box 500 07, 104 05 Stockholm, Sweden. E-mail: [email protected]

Secretary: Sergio Pérez-Ortega, Real Jardín Botánico, CSIC, Madrid, Spain. E-mail:[email protected]

Treasurer: Volker Otte, Senckenberg Museum für Naturkunde Görlitz, PF 300 154, 02806Görlitz, Germany. E-mail: [email protected]

Assistant Treasurer: Christian Printzen, Senckenberg Forschungsinstitut und NaturmuseumFrankfurt, Senckenberganlage 25, D-60325 Frankfurt am Main, Germany. E-mail:[email protected]

Editor: Ave Suija, Institute of Ecology and Earth Sciences, University of Tartu, Lai street 36-40,EE-51005, Tartu, Estonia. E-mail: [email protected]

Members-at-Large: Heidi Döring. – Jolanta Miadlikowska, Department of Biology, DukeUniversity, Box 90338, Durham, NC 27708, USA. – Adriano Spielmann, UniversidadeFederal de Mato Grosso do Sul (UFMS), Centro de Ciências Biológicas e da Saúde,Departamento de Biologia, Laboratório de Botânica, Cidade Universitária, s/n, CaixaPostal 549, CEP 79070-900, Campo Grande, MS. Brazil. – Marko Hyvärinen, FinnishMuseum of Natural History, PL 7 (Unioninkatu 44), 122 Helsingin Yliopisto, Helsinki,Finland.

INTERNATIONAL LICHENOLOGICAL NEWSLETTER 49 (1) 1

ASSOCIATION NEWS

Bid for IAL9 – 2020

We would like to propose Brazil as the host country for the 9th IAL Symposium in 2020. Thevenue, the Bonito Convention Center, in Bonito, Mato Grosso do Sul, in the Center-Westregion of Brazil (http://www.bonito-brazil.com/), has previously hosted the 49th AnnualMeeting of the Association for Tropical Biology and Conservation (http://www.eventus.com.br/atbc2012/welcome.html) with over 1200 participants; furthermore, thecity of Bonito has the infrastructure to host large international events with about 70 lodgingoptions.

Currently, we have a large group of leading Brazilian lichenologists from different parts of thecountry that as a team would be involved with the organization of the IAL meeting, including,apart from us, Emerson L. Gumboski (Univille), Eugênia Pereira (UFPE), Iane Cunha Dias(UEMA), Marcelo P. Marcelli (IBt), Michel N. Benatti (IBt), Patrícia Jungbluth (UFSM),Sionara Eliasaro (UFPR), and Suzana Azevedo Martins (FZB-RS).

We have also planned at least four different field trips to areas with resident lichenologists andappropriate infrastructure to host field workshops focusing on unique Brazilian ecosystems:Reserva Ducke (Amazon Forest); Serra Gaúcha (Atlantic Forest); Porto Murtinho andsurroundings (Brazilian Chaco); and the Catimbau National Park (Caatinga).

The host city, Bonito, is one of the most popular destinations in Brazil. Although little knownabroad, its name translates to “beautiful”, a reference to an area important for ecotourism dueto its natural beauties.Bonito has manycaves,waterfalls and streamswith crystal-clearwaters,and one of the world's largest sinkhole is nearby, hosting one of the largest populations ofscarlet macaws.

Brazil is one of the world's leading countries in science and technology and has historicallybeen important for its numerous botanical expeditions into the Amazon, the Pantanal and thesoutheastern Atlantic Forest. Due to the work of Müller, Krempelhuber, Vainio and Malme,many species have historically been described from Brazil. In recent decades, lichenologistsfrom inside and outside the country have been extremely active, making Brazil one of thecountries with the best known and most diverse lichen biota. Additionally, it would be the firstIAL meeting in South America and the Southern Hemisphere.

We are excited to announce our candidacy and will work hard to offer the participants of the9th IAL Symposium in Brazil a great experience and look forward to seeing our colleagues inHelsinki, Finland.

Thank you,

Marcela Cáceres (President, UFS)

Adriano Spielmann (Vice-President, UFMS)

Luciana Canêz (Secretary, UFMS)

Manuela Dal Forno (Treasurer, Smithsonian Institution)


NEWS

Lichenology courses at the Eagle Hill Institute

Steuben, Maine – 2016

Photo: Craig Snapp

Eagle Hill field courses are of special interest because they focus on the natural history of oneofNorthAmerica’smost spectacular andpristinenatural areas, the coast of easternMaine fromAcadia National Park to Petit Manan National Wildlife Refuge and beyond. Our summer fieldcourses are taught by experts in their respective fields. Course participants include beginningto advanced amateurs, graduate and undergraduate students, teachers, professional fieldbiologists, university professors, and personnel from federal and state agencies and numerousenvironmental organizations.

For general program information, http://www.eaglehill.us/programs/nhs/natural-history-seminars.shtml. For our complete course listing, go to http://eaglehill.us/seminars. Titles arehot-linked to course descriptions and instructor biographies. Go to http://www.eaglehill.us/programs/general/application-info.shtml. Institute homepage: http://www.eaglehill.us. Formore information, contact Marilyn Mayer: [email protected] or 207-546-2821 x1

Lichens and Lichen Ecology July 3–9, 2016

Instructors: David Richardson and Mark Seaward

This seminar is designed to help teachers, students, amateurs and those working in state orfederal positions become more knowledgeable about lichens. It is suitable for beginners andthosewhohave some familiaritywith lichens.With small student numbers and two instructors,we happily accommodate a range of abilities. Lichens are abundant and diverse along the coastof Maine, so participants can pursue topics of interest and also to develop identification skills.This seminar emphasizes both fieldwork and laboratory studies. We focus on identification ofspecimens using books, keys and chemical spot tests. We provide instruction for how to dothese tests and how to cut sections of lichens and their fruiting structures in order to examinethe spores. Lichens from open, forested, and seashore habitats will be studied, with anemphasis on the macrolichens, although some crustose species will also receive attention.Lectures and slide presentations will cover topics such as the structure, reproduction, and


ecology of lichens, as well as their use by man and their value for pollution monitoring.Thosecompleting this seminar can go on to advanced lichen seminars, such as CrustoseLichens of Coastal Maine.

Dr. David Richardson ([email protected]), Professor and Dean Emeritus at SaintMary’s University in Halifax, Nova Scotia, is a specialist in the effects of air pollution onlichens and has published many research papers and the books, The Vanishing Lichens andPollution Monitoring with Lichens. His studies have taken him to England, Ireland, Canada,and Australia.

Dr. Mark Seaward ([email protected]), Professor of Environmental Biology atBradford University, England, is a lichen ecologist. He has written/contributed to eight booksand edited the well-known Lichen Ecology published by Academic Press, as well as some 400other publications. He has studied lichens in Eastern and Western Europe, the Middle East,Indian Ocean islands, and Hong Kong.

Crustose Lichen Identification: July 10–16, 2016

Instructor: Irwin Brodo

Although the lichens of Maine’s coastal rocks and forest are a conspicuous component of thevegetation, it is usually the larger lichens, the foliose and fruticose species that attract one’sattention. This seminar will concentrate on the less conspicuous, but equally diverse andimportant, crustose lichens. Collections will be made from bark, wood, rocks and soil, and theywill then be identified in the laboratory. The newly expanded keys to crustose lichens fromLichens of North America, will be used, as well as other modern keys from the world literature.Techniques for sectioning, staining, and interpreting the tissues of crustose lichen fruitingbodies will be introduced, with special attention being devoted to staining various ascus typeswith iodine. Thin layer chromatography will be introduced and used to demonstrate how toanalyze the chemistry of some crustose lichens, especially sterile species, as the interest ofparticipants and time permits. Techniques for testing crustose lichens with para-phenylenediamine, hypochlorite solution (bleach), potassium hydroxide, nitric acid, andiodine will be discussed and used regularly for identifications. (Special $10 lab fee).

Dr. Irwin M. Brodo ([email protected]) was the research lichenologist at the CanadianMuseum of Nature in Ottawa for 35 years and remains a Research Associate there. He is theauthor of over 90 research papers on lichens, most of them on the taxonomy of North Americanspecies, but also including studies of ecology, air pollution, and general lichen biology.Together with photographer/naturalists Sylvia and Stephen Sharnoff, he has written anencyclopedic reference book entitled Lichens of North America illustrated with over 920 colorphotographs. The book was published in October 2001 by Yale University Press and is in itsfifth printing. He has fully revised and expanded the keys from that book published by YaleUniversity Press in 2015.


21st Symposium of Cryptogamic Botany

21–24 June 2017

We are delighted to announce the 21th edition of the Symposium of Cryptogamic Botany thatwill take place 21–24 of June 2017. This edition is organized by University Rey Juan Carlos,and will be hosted at Pavia building, old headquarters that nowadays is part of the Campus ofthis University in Aranjuez.Venue: Aranjuez, situated in the Madrid Region, lies42 km south of Madrid and 44 from Toledo, and iseasily reached by bus, train and car.

The Royal Site and Villa of Aranjuez nestles in thevalley at the confluence of the Tagus (locally Tajo)and Jarama rivers, in a privileged natural settlingwith vegetable gardens. An old quarter which is aHistoric-Artistic Site, royal palaces and gardens onthe banks of the Tagus forms the layout of Aranjuez.declared a World Heritage Site by UNESCO in 2001.

The current layout dates back to the XVIcentury, during the reign of Felipe II, whoawarded this town the title of Royal Site. Inthe following centuries (17th and 18th)Aranjuez became a centre of the court withpalace architecture and gardens designfollowing the aesthetic taste of theEnlightenment.


Symposium Center: Old Headquarters Building Pavia. It was built at the time of FernandoVII (1752), and subsequently used with military purposes. It is currently one of the historicalbuildings of Aranjuez, and part of the Campus of University Rey Juan Carlos (URJC).

The motto of the Symposium is “The Hidden Biodiversity” which highlights the efforts ofresearchers on Lichens, Ferns, Algae, Fungi and Bryophytes to reveal and make public therichness of these groups, their ecology and their conservation value.

Local Organizing Committee, members of the Area ofBiodiversity and Conservation at URJC:

Isabel Martínez Moreno

María Prieto Álvaro

Ana M Millanes Romero

Rosa M Viejo GarcíaBrezo Martínez Díaz-Caneja

Gregorio Aragón Rubio

Luis García QuintanillaM Carmen Molina Cobos

Further information:

More information about the Symposium will be soonavailable in the web page http://www.criptogami-a2017.com. For questions and suggestions, please sendan email to: [email protected].

Languages: Spanish, Portuguese, English.

Please, mark the dates 21–24 June 2017 in yourcalendar! We are looking forward to seeing you in theSymposium.

11th International Mycological Congress

16–21 July 2018, San Juan, Puerto Rico

The Mycological Society of America (MSA), theInternational Mycological Association, the Latin AmericanMycological Association (ALM), the Puerto RicanMycological Society (SPM), Universidad del Turabo andMeet Puerto Rico are proud to sponsor the 11th InternationalMycological Congress in San Juan, Puerto Rico.

San Juan, the capital city, possesses a state of the art


convention center, excellent hotels, excellent cultural activities and an amazing culinaryexperience. The Luis Muñoz Marin International Airport is a main hub for different airlinesreceiving direct flights from major airports in the USA, Europe, Central and South America.

New Cryptogamic Herbarium (ICH) in Iran

I am pleased to provide the link of the new lichen herbarium (http://sweetgum.nybg.org/science/ih/herbarium_details.php?irn=245024) founded in May 2016 at the Iranian ResearchOrganization for Science and Technology (IROST).

The Iranian Cryptogamic Herbarium currently contains mostly lichenized and lichenicolousfungi, aswell aswood-inhabitingmacrofungi.This collectionhasanestimatednumberof2000specimens mainly from Iran, and some duplicates from around the world, with an emphasis onCaspian Hyrcanian mixed forests, Oak forests of the Zagros Mountains, arid and semiaridsteppes. It is a fundamental resource for biological systematics andclassification, diversity andconservation, biotechnological applications, biological monitoring, biodeterioration andbioremediation studies, climate change, ecology, and pharmaceutical bioprospecting. Thecollection is a resource for academics, students, government, private organizations, and thepublic.

The new herbarium is one of the fundamental infrastructures of the Iranian Lichen Floraproject, which will start as a Multi-Authored International Research Project in the near future.Youaremostwelcome tovisit ICH, to request loans, exchange lichenmaterial, initiatebilateralcooperation, and to receive gift samples which will be safely deposited in ICH.

Mohammad Sohrabi, Tehran

REPORTS

Second Young Lichenologists’ Workshop

12–15 November 2015, Budapest, Hungary

In view of the success of the First Young Lichenologists’ Workshop in 2009, we decided toattend the second meeting in Hungary. This event was organized especially for Ph.D. studentsand young scientists with deep interests in lichenology, who could present their currentresearch works in front of an international audience and thus improve their discussion abilitiesand language skills. Since it was free of charge, the event was available for a wider audienceof interested students. The meeting brought together 17 participants from six Europeancountries (Croatia, Finland, Hungary, Poland, Slovakia, Ukraine). The main topics of theworkshopwere focusedon lichenbiodiversityandchemistry, but lichenological investigationse.g. taxonomy, morphology, ecology, ecophysiology and climate change, were welcome.

On the first day, the organizers welcomed us in the Danube Research Institute, MTA Centrefor Ecological Research of the Hungarian Academy of Sciences in Budapest. Péter Ódor,Director of the Institute of Ecology and Botany, Centre for Ecological Research of the MTA,opened the workshop with a presentation on the dynamics of epiphytic species composition inHungarian mixed forests. At the evening welcome reception we could freely talk with otherparticipants and enjoyed delicious home-made specialities prepared by the organizers. Thesecond day was devoted to presentations of invited speakers S. Ya. Kondratyuk and R. Engel,


as well as student participants. S. Kondratyuk’s presentation was about novel lichenicolousfungi described by him from Ukraine, Israel and South Korea, and R. Engel presented workon the detection of active secondary compounds from herbs and lichens by HPLC. Edit Farkas’presentation covered the chemistry of lichens and standard HPTLC methods used in her work– these three presentations provided a very useful summary to all of us. In total, 12 oralcontributions and three posters were presented. The ideal atmosphere generated duringdiscussions enabled one to pose many questions and encouraged us to share, develop and guideour ideas.

During the next days, the programme was enriched by an excursion to Mt Odvas-hegy in theBuda Mts, which are the part of the Transdanubian Mountain Range. We observed the lichendiversity on opened xerothermic slopes with predominantly grassland vegetation andconglomerate outcrops, where we recorded various species, despite the windy weather, suchas Agonimia opuntiella, Cladonia convoluta, Fulgensia fulgens, Lecanora argopholis,Leptogium schraderi, Psora decipiens, Squamarina cartilaginea and Toninia sedifolia. A raresaxicolous species Rusavskia papillifera (= Xanthoria p.) discovered about 100 years ago wasfound on the calcareous rocks of the SE slopes. After the excursion we checked somespecimens under microscopes, learnt about some lichen taxa, and discussed several questions.

We also visited the Institute of Ecology and Botany, MTA and the National Botanical Gardenin Vácrátót, which contains c. 11 600 plant taxa, resulting in the richest scientific plantcollection of Hungary; we also checked materials in the lichen herbarium (VBI).

Theaimsof theSecondYoungLichenologists’Workshopwere fulfilled andwe returnedhomewith new knowledge that will help us to increase our research expertise. For us, it was pleasurebeing a part of this unforgettable event and therefore we would like to thank the organizers Edit

Group photo (from left) Katalin Veres, Edit Farkas, Nóra Varga, Zsanett Laufer, Rita Engel,Sergej Kondratyuk, Mónika Sinigla, Gábor Matus, Zuzana Fačkovcová, Zsolt Csintalan,Natalia Matura, László Lőkös, Maja Maslać, Aino Hämäläinen. Photo: Nora Varga


In the field – Mt Odvas-hegy. Photo: Lazlo Lőkös.

Lichen identification after the excursion. Photo: Nora Varga


Farkas and Nóra Varga of the Institute of Ecology and Botany, MTA Centre for EcologicalResearch of the Hungarian Academy of Sciences. We hope that the idea of this workshop willcontinue.

Zuzana Fačkovcová and Mónika Sinigla

Institute of Botany, Slovak Academy of Sciences, Dúbravská cesta 14, 845 23 Bratislava,Slovakia; [email protected]

Bakony Museum of the Hungarian Natural History Museum, H-8420, Zirc, Rákóczi tér 3–5., Hungary; [email protected]

Lichens under the microscope. Photo: Jarek Jõepera

“The bold and the beautiful” exhibition, Tallinn

Despite the work done by schools, many people do not know the difference between lichensand mosses. To pour oil on the fire, the words ‘lichen’ and ‘moss’ are very similar in Estonian,namely samblik and sammal respectively and hence people frequently assume that these aresynonymous names for one organism. An exhibition of lichens and mosses in the EstonianMuseum of Natural History introduces these two groups to a wider audience and shows howbold and beautiful they really are. People often do not recognize the existence of lichens andmosses, and remain invisible because they are usually small. To change this, we bought lichensand mosses to our exhibition hall, where visitors can look and explore them very closely usingavarietyofways, suchas touch, smell, viewed throughmagnifyingglasses and light anddigitalmicroscopes, and use of hands-on interactive models. Lichens and mosses are brought fromthe background to the foreground, turned from extras to superstars (as often demonstrated inthe wild). Two themes – lichens and mosses – are divided into two different exhibition rooms,


Journey to the lichen diversity. Photo: Jarek Jõepera

their physical separation emphasising that these are two fundamentally different groups. Onesignpost on the floor leads visitors to the fungal kingdom, and the other to the plant kingdom.

The bold and the beautiful. Starred by: lichen and moss is the largest and most comprehensiveexhibition of lichens made in Estonia. In the fungal kingdom, visitors can learn lichen ‘basics’,and dig deeper to explore their appearance, hardiness, significance, etc. Boards with texts,drawings, photographs and different species placed on walls and exhibition displays a varietyof topics, helping to open up lichens world to visitors. Texts, drawings, etc. have been designedto aid both people who are just getting acquainted with lichens as well as those who wish toexpand their knowledge of them.

In the exhibition we showcase the commonest and most conspicuous lichens from differenthabitats and varieties of substrates in Estonia. Genuine tree trunks, rocks, etc. have beenintroduced into the exhibit room, so that visitors could touch and have a closer look at lichenson different surfaces. In addition to Estonian lichens, outstanding species from around theworld are also presented. Altogether about 80 species are displayed, most of which werecollected for theexhibition,but somespecimensareon loan fromthe lichenological collectionsof the University of Tartu and the Estonian Museum of Natural History.

The part of the exhibition devoted to lichens consists of ten topics.

1. Lichen = fungus + alga; 2. Lichen as a fungal greenhouse; 3. Lichens in the distant past

In the exciting journey of discovery, visitors can peek into lichens via an interactive model andexamine under the microscope a X 100 enlarged cross section of lichen thallus. Visitors canexamine the rhizines with a magnifying glass and read about their true function. The followingquestions will find an answer: is the relationship mutualistic or parasitic, did lichens alreadyexist before the dinosaurs, and how old is the oldest confirmed lichen fossil?


4. Variegated folks; 5. Where is its home ground?

Visitors can learn from text boards that lichens colonize all region of the world, but aredominant in about 8% of the land surface, notably in extreme polar areas, deserts and highmountains. However, they can also be found growing in the realm of water. Lichens aredisplayed to show a variety of lichen substrates starting with trees and ending with a soldier’sboot. A hands-on device introduces the extraordinary diversity of shapes and colours exhibitedby lichens, and enables the visitor to find lichens with different growth forms on trees by justpressing a button – to confirm the accuracy of its search, or simply learn rapidly.

6. Reproduction – together or alone

A digital microscope helps visitors to take a closer look at fungal fruiting bodies and vegetativereproduction of lichens, and to explore their diversity. Through the microscope, drawings andtexts visitors can learn about the two different ways of reproduction and their importance indifferent environmental conditions.

7. Hardy but sensitive

An interactive solution has been created to illustrate a lichen’s resistance to extremetemperatures, radiation and moisture fluctuations on the one hand, and their sensitivity to airpollution on the other hand. Via text boards how and why lichens are used to detect air qualityis explained, visitors can also acquaint themselves with several indicator species.

8. The 1000 reasons for uniqueness – lichen substances; 9. Lichens at the service of humans

The exhibition brings light to the well-hidden secret of lichens – lichen substances of which

Special UV-light device for observing fluorescing lichen substances. Photo: Jarek Jõepera


there are more than 1000, mostly unique to lichens, some fluorescing in ultraviolet radiation– visitors can switch on and off a specially made UV “searchlight” device. Guests can also readabout the role and importance of lichen substances to lichens. The human use of lichens, fromhaving a meal to making mummies, is displayed on rotating text boards.

10. Jobs in the ecosystem

Lichens play a number of important roles in the ecosystem, from soil formation to offering ahabitat and shelter to small invertebrates. To show lichens as soil makers, rocks with lichensgrowing on them are demonstrated in the exhibition. In addition, visitors can find a chaffinch’snest constructed mainly of lichens on a branch – and look for a disguised butterfly.

The exhibition, enhanced by educational programmes, lectures and workshops, supported bythe Environmental Investment Centre in Estonia, is open in the Estonian Museum of NaturalHistory in Tallinn’s old town (Lai 29a) until 8 January 2017. All texts are in Estonian, Englishand Russian.

Marja-Liisa Kämärä, Tallinn

OBITUARY

Lars-Erik Emil Muhr

25 October 1936 – 24 May 2015

On May 24th 2015 Lars-Erik Muhr died, aged 78. For many years he was deeply engaged inlichenology. Like his predecessor and friend Sigurd Sundell, he mainly concentrated on thelichen flora of his home province Värmland, but he also collected elsewhere in Scandinavia.He was one of very few lichenologists who set up a TLC laboratory at home and his knowledgein lichen chemistry was valuable in his study of sorediate crusts, often in collaboration with


Lars-Erik Muhr in 1983. Photo: Roland Moberg

Tor Tønsberg. He was also keen on picking up news from abroad, particularly from Britishlichenologists.

Lars-Erik published a number of floristic and taxonomic papers, alone or in collaboration withothers, in Svensk Botanisk Tidskrift, Graphis Scripta, The Lichenologist and Nordic Journalof Botany. New species co-authored by him included Buellia violaceofusca, Japewiasubaurifera, Micarea marginata, Pertusaria flavocorallina and Schaereria corticola. He alsodiscoveredStereocaulonurceolatum,whichwas first describedasMuhriaurceolata in honourof him. The main part of his collections are housed at UPS, which 2544 are currently registeredin the UPS database (available at the home page of The Museum of Evolution).

During the last 20 years Lars-Erik pursued other interests, partly due to the cancer he managedto successfully fight for a long time. Two of these were entomology and the compilation ofcrosswords. His large collection of hoverflies (Syrphidae) will soon be incorporated into thezoological part of The Museum of Evolution, and his crosswords are still manifested by smallamounts of money now and then sent to his wife from different newspapers and magazines.

Our acquaintance with Lars-Erik was mainly in the 1980s and 1990s, when we met at severalexcursions arranged privately or by NLF and SLF. We profited greatly from his deepknowledge and sharp eyes, which made us discover details we would not have seen withouthis assistance, and we rejoiced in his sympathetic personality and his sense of humour. We willforever remember him as a good friend and an extraordinary lichenologist.

Roland Moberg and Anders Nordin, Uppsala

[Published in Graphis Scripta 28-1, 2016]

PERSONALIA

A new lichenologist at the Canadian Museum of Nature(CANL).

R. Troy McMullin has accepted the position of Research Lichenologist at the CanadianMuseum of Nature in Ottawa. He bega his tenure on April 3, 2016. This is the position vacatedby Irwin (Ernie) Brodo in November 2000 when he retired, although Ernie has continued todo lichen research at the Museum ever since.

Troy was born in Montreal, Quebec, but received much of his later education in Ontario,earning several degrees and diplomas in Environmental Studies. His interests turned to lichensin Nova Scotia where he earned his Master’s Degree at Dalhousie University (Halifax)studying lichens inoldgrowth forests.Hecontinuedhis lichenological studies at theUniversityof Guelph in Ontario dealing mainly with woodland caribou habitat in Ontario focusing on thegenus Bryoria. Since earning his Ph.D. at Guelph, he has held several postdoctoral fellowshipsthere, especially at the Centre for Biodiversity Genomics.

Troy’s interest in lichen floristics has taken him far and wide in North America, especially inCanada, and he has amassed a sizable lichen collection and deep knowledge of lichens of all


kinds, which will be a particular value in his new job. He has written or contributed to severalguidebooks on lichens, the latest being Common lichens of the Northeastern North America(New York Botanical Garden Press, 2014), collaborating with Nova Scotia lichenologist,Frances Anderson. In recent years, Troy has made many important additions to the Ontariolichen flora and has made the stubble lichens (calicioid lichens) his special field of interest. (Herecently described the new species, Chaenotheca balsamconensis and has another in press.)Troy recently was appointed to the Subcommittee on Lichens and Bryophytes of theCommitteeon theStatusEndangeredWildlife inCanada (COSEWIC)keeping trackof raritiesin the country and writing assessment reports.

Troy’s acceptance of the position begins a new era for lichenology at the Canadian Museumof Nature and promises to bring new life to Canadian lichenology in general.

Irwin Brodo, Ottawa

Boros Ádám Award in Hungary 2016

Ádám Boros (1900–1973) was a famous Hungarian botanist with a special interest in bryologyfrom an early age. He spent his life investigating the flora and vegetation of the CarpathianBasin, particularly cryptogams, and many taxa were named after him.

The Boros Ádám Award was established in 1998, on the 25th anniversary of his death, byAlbert Tóth, Attila V. Molnár and Róbert Vidéki to honour the achievements of botanistscontributing outstanding work on the flora and vegetation in the Carpathian Basin. The award


is given every second year during a ceremony at the International Conference Advances inresearch on the flora and vegetation of the Carpato-Pannonian region. The aim of the awardis to inspire and give moral support to botanists for their future endeavours and hopefullyinspiring them to become eminent lichenologists. During the 11th Conference (Budapest, 12–14 February, 2016) two Hungarian lichenologists, Edit Farkas (Institute of Ecology andBotany, MTA Centre for Ecological Research) and László Lőkös (Botanical Department,Hungarian Natural History Museum) were honoured by this Award. In his speech, Prof. AttilaBorhidi announced that these lichenologists had made outstanding and pioneer workthemselves, as well as organizing the mapping of the lichen flora in Hungary, discovering alarge number of lichen-forming fungi for the country as floristic novelties, introducing newchemical methods for identifying lichen compounds in Hungary, and involving and teachingmany young students and scientists in their collaborations.

Congratulations to Edit and Laszló and my best wishes to continue their work for a long timeto come.

Nóra Varga, [email protected]

Institute of Ecology and Botany, MTA Centre for Ecological Research, Vácrátót, Hungary

BOOK REVIEWS

ĀBOLIŅA , A., PITERĀNS, A. & BAMBE, B. (2015): Lichens and Bryophytes in Latvia.Checklist. (Latvijas Ķērpji un Sūnas. Taksonu saraksts) – Salaspils:Latvijas Valstsmežzinātnes institūts ‘Silava’, DU AA ‘Saule’, 213 Pages, Hardcover. ISBN978-9984-14-735-2. Price: unknown.

Latvian lichens remain the least studied of all the three BalticStates and during the last two decades comparatively very fewpapers dealing with lichen flora of this country have beenpublished. Among them, there was the first lichen checklist in2001 compiled by the author of the present checklist whichincludes 573 species and, for the first time, lichenicolous fungi(albeit listed separately from the lichens). The checklistfollows the same format as the previous one, namely: name ofspecies, synonyms, substratum data and frequency in Latvia.It can be an unrewarding job to compile checklists ofunderstudied but easily accessible areas since such lists ofspecies often become obsolete very soon after published.Sadly this is what happened with the latest checklist of Latvianlichens. Several recent papers, published or in press, add c. 100new lichenized and lichenicolous fungi species for the country

(i.e. over one-sixth of this checklist). Still, the book can be of some use for these interested inlichen distribution in the eastern Baltic region, but should be regarded with certain reservationsdue to numerous misprints, misuse of taxon names, lack of modern revision of some species


or species complexes, or a critical evaluation of the presence in the country of species that areknown only from historical literature data.

Jurga Motiejūnaitė, Vilnius

BRODO,I.M. (2016):Keys toLichensofNorthAmerica:RevisedandExpanded. –CanadianMuseum of Nature. 427 Pages, 43 Figures (13 color, 30 b/w). ISBN 978-0-30019573-6. Price:$29.95 8 1/2 x 11

Fifteen years ago, natural history enthusiasts andlichenologists alike were astounded by the exquisite photosand excellent keys and descriptions in Brodo, Sharnoff andSharnoff’s instant classic, the Lichens of North America.Brodo’s new Keys to Lichens of North America: Revised andExpanded (2016) complements the original work by almostdoubling the number of species and providing nomenclaturalupdates. The revised and expanded keys include 2028 speciesin 382 genera, including 106 new genera. The format,paperback and spiral bound, makes it easy to use for lichenidentification in a classroom or workshop. The keys arecomprehensive yet straightforward so that both professionalsin the field and other lichen enthusiasts will find it a valuableaddition to their libraries.

As in Lichens of North America, the keys are presented and organized in way that makes themaccessible. The book begins with an identification key to major groups followed by keys to thegenera in each of these major groups. The identification keys to species are organizedalphabetically by genus, which makes it easy to find the genus you are looking for. The keysinclude leads on geography, habitat type, lichen color, external and internal structures,responses to spot tests and to illumination with UV light. Often when describing spot testresults, Brodo includes in parentheses the name of the secondary compound(s) responsible forthe response, as in, “K+ red (salazinic acid)”, which is helpful for students developingunderstanding of the chemical diversity of lichens. Brodo has been accepting comments on hisLichens of North America Key for over a decade, and has improved many of the couplets forclarity.

The book also includes many valuable sections for people just learning lichenology, many ofwhich were also in Lichens of North America, with some new ones included. The diagrams ofSusan Laurie-Bourque include those describing thallus growth forms, vegetative propaguletypes, the types of fruiting bodies including cross-sections, types of asci and types of spores;these diagrams are very helpful for clarifying the language in the keys. Another valuablediagram for beginners depicts where on a foliose lichen one should measure the lobe size. Newfigures include illustrations of the 5 algal and 5 cyanobacterial photobionts, the difference inlichen appearance between wet and dry specimens, the variation in types of isidia, thebranching and measurement of Usnea, and spores of Rinodina.


The literature cited section directs readers to more detailed treatments of the taxa included; thefact that 76% of the sources were published since 1998, when Lichens of North America wasfirst prepared, underlines how important it was that Brodo complied this revised and expandedkey, which is a superb resource for lichenologists of North America and beyond.

Natalie Howe, Edison, New Jersey, United States

DAVEY, S. with DAVEY, A. (2015): The Lichens of Jersey. – Société Jersiaise Jersey. xi +195 Pages, Paperback. Available from The Société Jersiaise, http://shop.societe-jersiaise.orgor through http://www.tiliapublishinguk.co.uk, ISBN 978-0-901897-58-9. Price £15 +postage.

The Lichens of Jersey is a significant, comprehensive andattractive contribution to our understanding of the lichens ofthis, the largest of the Channel Islands. The authors, Simonand Amanda Davey, provide us with accounts of its richdiversity, 560 recorded species, of which over one half aregenerously illustrated by the authors’ colour photographs.

Resulting from their enthusiasm and frequent visits to theisland, many jointly with Peter James, the authors have takenthe opportunity of drawing on background and contextualinformation, and lichen collections, seldom made available insuch a publication. This makes the book uniquely useful andof interest both to lichenologists seeking more than an up-to-date list of the lichen species of Jersey and also to a wideraudience.

Thebook is skilfullyorganised intoninemainchapterswithinwhich species accounts (Chapter6) occupy half of the volume. It is the enhancement of supporting and background chapterswhich provides the special strength of the book: Chapter 1 (Introduction), Chapter 2 (Historyof Lichenology in Jersey), Chapter 3 (The Difficulties of Species Names over Time), Chapter4 (Topography and Geology), Chapter 5 (Habitats), Chapter 7 (Jersey Lichen Herbaria). Thesechapters, together with a Glossary (Chapter 8) and Bibliography (Chapter 9) and sectionsdevoted to Dedication, Acknowledgements, Foreword, and Indices (both general and to lichenspecies) make up the rest of the book.

Chapter 4 (four pages) alerts the reader to the diversity of habitats in Jersey, rich in coastalgranites (see map on page 23), and to its affinity with the nearby French coast, as also is thecase for other Channel Islands. The map shows the positions of the 48 sites visitedsuperimposed on the underlying geology. Habitat descriptions in Chapter 5 (36 pages) includephotographs and mention the influences of the geology of the island on the lichens. Selectedcorticolous, saxicolous (which includes churchyards, castles and forts) and terricolous(grassland) lichen communities in these habitats are discussed, accompanied by photographsand with lists of noteworthy lichens of described sites. Conservation status is given forcorticolous communities as, for example, in St Catherine’s Woods (see page 27) and mentionis made in species accounts (Chapter 6) of nationally scarce Jersey lichens.


Charles Larbalestier, the eminent 19th century lichenologist, is the main protagonist in thehistory of lichenology of the island (Chapters 2, 3 and 7). He was born and lived in Jersey andstudied at Cambridge; the account given in this book includes his interaction with otherprominent lichenologists of the day (eg William Nylander), reference to his collecting inJersey, other Channel Islands and in Ireland and makes a valuable and enlightening read.Larbalestier was part of an active network of Victorian lichen collectors who made boundexsiccati, exchanged and sold them and sent exsiccati to museums worldwide.

The book remains a summative account, the value of which should not be eclipsed byweaknesses that can be improved in any future edition. Inevitably, a few minor inconsistenciesand typographical errors occur. In Chapter 6 photographs can be difficult to match with therelevant species accounts as the caption should include the species name, as well as (or insteadof) the site. It would also be helpful to include magnification or scale for species photographed.Some images reproduced are disappointing, as for very dark lichens (e.g. Verrucarianigrescens, page157)which canbenotoriouslydifficult, requiring adjustment in exposure andextra care in printing, to reveal detail. Most serious is the absence of keys to genera (and tospecies where applicable) or reference to diagnostic characters which reduces the usefulnessof the book for identification purposes. An additional section or chapter could be included,rather than presume reliance on another publication to provide keys, as found in The LichenFlora of Great Britain and Ireland edited by Smith et al. or in Lichens an Illustrated Guide tothe British and Irish Species by Dobson (the omission of which in the Bibliography isregrettable).

The Lichens of Jersey is recommended to those interested in the lichens of this and otherChannel Islands, both from contemporary and historical perspectives. This study should alsoappeal to the natural historian and ecologist, attracted by its broader scope. It is important tobear in mind that the lichen flora of Jersey, as so much of our coastline, is rapidly changing withresultant losses (reduced diversity) and impact on lichen communities. At the same time, moreremote areas, such as inaccessible headlands and small islands, may provide reservoirs of relicspecies lost from elsewhere, a situation requiring monitoring. The information in this book, inaddition toprovidingacurrent list of Jersey lichens, contributes toourknowledgeof ecologicalchange.

Ann Allen, Bideford, Devon

DIETRICH,M.&DANNER,E. (2014):Flechten.FaszinierendeVielfalt inderBergwelt umEngelberg. Auf den Spuren von Pater Fintan Greter (1899–1984). (NaturforschendeGesellschaft Obwalden und Nidwalden NAGON Bd. 5) – Grafenort: NAGON, 239 pages,Hardback. ISBN 3-9521401-5-5. Price: CHF 40.

This nicely produced book, despite its title Lichens. Fascinating diversity in the mountainousworld around Engelberg. On the tracks of Father Fintan Greter (1899–1984), is mostly anintroduction to lichens in general and therefore of much broader interest then its title maysuggest. The book is the result of the revision of the lichen collections of Fintan Greter, aBenedictine monk from the monastery of Engelberg working for much of his life as a highschool teacher of biology and other natural sciences. These collections are now housed in the


Natural History Museum of Lucerne in Switzerland (Natur-Museum Luzern) and have been determined or revised by thepresent authors. Fintan Greter only published his thesis onmosses during his lifetime, but had a strong interest in lichensas well, collecting more than 3000 specimens (˃ 500 species)from the area of Engelberg in central Switzerland. He was alsoa founding member of the Swiss Bryological andLichenological Society (today Bryolich) and the organizer ofits second meeting in Engelberg.

On 20 pages the tradition of natural history research andcollections in the monasteries of Engelberg and Einsiedeln areexplained. Short biographical notes of 10 botanists among theBenedictines of these two monasteries during the last 200years are presented, followed by a short biography of Fintan

Greter. 18 pages are devoted to an introduction of the area studied by Greter: the upperEngelberg Valley south of Lucerne reaching altitudes of over 3000 m. The next 28 pages area general introduction to lichens, their diversity, morphology and ecology. A further 40 pagescover the occurrences of lichens in the local habitats, and finally 96 species are portrayed withgood colour photographs and descriptions. A glossary, a list of illustrated species and, mostimportant from a scientific point of view, a complete list of all the c. 500 species collected byGreter together with notes on their occurrence in the Cantons of Obwalden, Nidwalden and Uriare provided.

Sadly the photographs lack scales which are especially important for beginners and non-lichenologists for whom the book is mainly intended. Nevertheless the book can be warmlyrecommended for everybody who require an introduction to lichens and lichenology of theAlps in general.

Peter Scholz, Schkeuditz

GAGARINA, L.V. (2015): Gyalectoid lichens (families Gyalectaceae Stizenb. andCoenogoniaceae (Fr.) Stizenb. in extratropical Eurasia [In Russian] – Nestor-Historia, Sankt-Petersburg. 240 pages ISBN 978-5-4469-0662-8. Price: unknown.

This comprehensive monograph arose from the author’s dissertation on extratropicalgyalectoid lichens. It starts with an in-depth overview on different past and present conceptsof these lichens, followed by a detailed description of their morpho-anatomy, including SEMphotographs showing the different types of ascomata and microstructures of the ascospore’ssurface. One interesting aspect is the inclusion of original b/w drawings describing thedevelopment of ascospores and the formation of septa in gyalectoid genera. Less coverage isprovided on chemical characterization, ecological aspects and geographical distributions. Thefinal chapter summarizes the current knowledge of molecular systematics in gyalectoidlichens. In addition to standard methods commonly used to identify lichen secondarymetabolites (i.e. spot tests, TLC, HPTLC), the author has used a metabolomic method tocharacterize the whole set of metabolites. From five species representing three genera


JAKLITSCH, W., BARAL, H.-O., LÜCKING, R. & LUMBSCH, H.T. (2016):Ascomycota. – In: Frey, W. (ed.): Syllabus of Plant Families. Adolf Engler’s Syllabus derPflanzenfamilien 13th edition, Part 1/2. – Stuttgart: Gebr. Borntraeger, 322 pages, Hardback.ISBN 978-3-443-01089-8. Price: 119 EUR.

(Coenogonium, Gyalecta and Pachyphiale), 573 substanceswere detected, 17% of which were identifiable as polyols,carbohydrates, amino acids, etc. Based on a statistical analysisof the range of substances it would appear that each genus hasits own unique set of metabolites.

The major part of the book contains detailed descriptions andkeys for 62 species of Belonia, Cryptotechia, Gyalecta,Pachyphiale, Ramonia and Coenogonium, three of which,Gyalecta hokkaidica Gagarina, Pachyphiale filamentosa G.Thor & Gagarina and Ramonia elongata Gagarina, aredescribed, and 34 species are lectotypified. One species –Gyalecta levuschensis Vain. is synonymized with Biatorasphaeroides (Dicks.) Hornem.The book is richly illustrated by photographs and b/w drawings, some of which are originaldrawings of main characters (asci, ascospores, paraphyses). Three appendices are given, twoof which are tables summarizing the main characters of extra-tropical gyalectoid lichens andtheir substrate preferences, and the third comprises distribution maps.

The Editor

Some lichenologists may be surprised by a work whichincludes a volume on lichenized and non-lichenizedAscomycota in its 13th edition that they may not have heardmuch about before. However, so-called cryptogams have beenincluded in this work for more than 100 years due to the famousbotanist,AdolfEngler,Directorof theBerlinBotanicalGardenand Museum, who wrote a single volume himself whichincluded all fungi (with lichens) on only 21 pages. It started in1892 with a slightly different title and soon became a standardtext with new editions every few years, the 6th editionappearing with minor alterations in 1909. The 7th edition in1912 saw the inclusion of illustrations and therefore anextension of the fungal chapter to nearly 40 pages. Engler wassupported by E. Gilg hereafter until the 11th edition in 1936whichwas issuedbyLudwigDiels as a singlevolume.Only the12th edition, edited by H. Melchior & E. Werdermann in two volumes in 1954 and 1964,contained a special chapter (15 pages) on lichens by F. Mattick.


After a break of about 60 years a completely new edition of several volumes written in Englishby well-known international specialists and edited by W. Frey has appeared. From the formereditions in German it follows the general concept, title and tradition of including the fungi aswell as blue-green algae, myxomycetes etc.

The short introduction of 12 pages is followed by a synopsis of classification of the phylumAscomycota on 14 pages. This continues with the systematic arrangement of taxa – the maincorpus of the book. Here all accepted taxonomic entities to family level are described usingmorphological as well as molecular data. Taxonomic uncertainties are discussed and extendedlists of references are separately given for every class. Accepted genera in every family arelisted with an estimation of the number of species in brackets. This part is illustrated by 7 platesof line drawing taken from the 12th edition and 17 plates of c. 150 new colour photographs ofhigh quality. The orders Thelocarpales Lücking & Lumbsch and Vezdeales Lumbsch &Lücking are proposed as taxonomic novelties.

The text of the introduction and the systematic arrangement is written by Thorsten Lumbschand Robert Lücking with contributions by the other two authors. The lichenized Ascomycotaare treated by Lumbsch and Lücking, whereas the non-lichenized Ascomycota are partlywritten by Walter Jaklitsch of Vienna or Hans-Otto Baral of Tübingen.

This is a standard reference work mandatory for every honest botanical library. Forlichenologists the parts including lichen photo-symbionts may also be of interest.Photoautotrophic eukaryotic Algae have been published as volume 2/1 in 2015, blue-greenAlgaeare included involume1/1published in2012, andBasidiomycotaare still inpreparation.


RANDLANE,T., SAAG,A.,MARTIN,L.,TIMDAL,E.&NIMIS,P.L. (2011):Epiphyticmacrolichens of Estonia. – University of Tartu, Tartu. 326 Pages, Paperback. ISBN978-9949-19-652-4. Price: 12–19 EUR.

The book provides an identification key, species descriptions,glossary, distribution maps, phototographs and drawings for 120epiphytic macrolichens found in Estonia. This is the first three-language (Estonian, English, Russian) richly illustratedlichenological book not only in the Baltics, but on a wider scale toencourage interested people to study epiphytic macrolichens.

The book starts with the identification key (with lichencharacteristics in three languages) supplemented with pictures anddrawings, making it easy to use. Although a page reference at theend of each species identification to its description is not given, thelatter are fortunately presented in alphabetical order to aid locatingthem. The species descriptions provide key information onmorphology, chemistry, distribution and ecology (in threelanguages), each species description is illustrated by a high quality


macro-photograph,where typical lichen thallus structures are accentuated;drawingsare addedfor some descriptions to elaborate particular characters. Each species is complemented by a

VUST, M. (2015): Inventaire des lichens du canton de Genève. – Boissiera 69. – Genève:Editions des Conservatoire et Jardin botaníques, 144 pages, Paperback. ISBN978-2-8277-0085-1. Price: 75–100 CHF.http://www.ville-ge.ch/cjb/publications/publications_pdf/liste_publications.pdf.

map to show its distribution in Estonia. A useful glossary in three languages defines the mostimportant terms used in the book.

Those interested in this excellent book could be both students as well as experiencedlichenologists interested in temperate region epiphytic macrolichens. The book has in user-friendly format, and is easy use in the laboratory and to take in a backpack for fieldwork. Theidentification key is also available in a digital form via the internet and smart-phoneapplications. The price of the paper printed version is very attractive.

Anna Mežaka, Rēzekne

The inventory of the lichens in the Canton of Geneva(Switzerland) is part of a major project of lichen research inthis area. Somewhat surprisingly the main list of acceptedspecies is not included in this paper, but has been publishedpreviously in Herzogia (vol. 28 (1): 153–184) by M. Vust, P.Clerc, C. Habashi & J.-C. Mermilliod in English. The presentFrench publication with an English abstract includes a generalintroduction to the project, the studied area and the history oflichen inventories in Geneva. Despite the recent detailedinvestigations there are two major sources of historicalinformation on lichens in the area, the first on the notes andcollections of Jean Müller (Müller Argoviensis), whopublished in 1862 a list of lichens in the vicinity of Geneva, butcontinued to make notes on their distribution over more than30 years until the end of his life (1896), and the second on the


more than 1200 unpublished collections of Jacques Rome (1831–1888) made during 1874–1887 and now part of the lichen herbarium of Geneva (G). The general results of the inventoryare presented and discussed. The red list of the lichens of the Canton is also discussed, but notincluded; however, it can be downloaded from ttp://www.ville-ge.ch/cjb/publications/publications_pdf/LR_lichen.pdf.

A major part (50 pages) of the present publication is an overview of lichen communities “quitelikely to exist in the Canton”, which includes 52 associations in 14 classes. Most of them areillustrated by photographs and their characteristics are discussed but no vegetationassessments have been given.

The publication concludes with two important appendices of the species listed by Jean Müllerand of the unpublished collections of Jaques Rome, complete with localities, dates andaccession numbers at G.

All those who have been involved in the inventory can be congratulated on the many detailedresults assembled for a rather smallCanton, especially themain investigator,MathiasVust, butI would have preferred to see the results in a single volume instead of several publications.


New theses

Margrét Auður Sigurbjörnsdóttir (University of Iceland) defended her Ph.D. thesis Thelichen-associated microbiome: taxonomy and functional roles of lichen-associated bacteriaon March 7th, 2016

Aino Hämälainen (University of Eastern Finland) defended her Ph. D. thesis Retentionforestry and intensified biomass harvest: epiphytic lichen assemblages under opposingecological effects in pine-dominated boreal forests on June 10th, 2016.

New members

Guillermo Amo de Paz, e-mail: [email protected], C\ La Alegría de la Huerta, 22 esc1ª 6ºA, 28041 Madrid, Spain

PilarHurtadoAragüés, e-mail:[email protected],CalleAntonioLopez67,Portale 13e, E-28019 Madrid, Spain

Elisa Banchi, e-mail: [email protected], [email protected], Life SciencesDepartment, University of Trieste, via Giorgieri 10, 34127 Trieste, Italy

Kristine Bogomazova, e-mail: [email protected], Royal Botanic GardenEdinburgh, 20A Inverleith Row, Edinburgh EH3 5LR, Scotland, UK

Prof. Joel Boustie, e-mail: [email protected], Université de Rennes 1 – Facultédes Sciences pharmaceutiques et biologiques, Laboratoire de Pharmacognosie et deMycologie 2, Av. du Professeur Léon Bernard - 35043 Rennes Cedex, France

Dr. Marylene Chollet-Krugler, Université de Rennes 1 – Faculté des Sciencespharmaceutiques et biologiques, Laboratoire de Pharmacognosie et de Mycologie 2, Av. duProfesseur Léon Bernard - 35043 Rennes Cedex, France

Jessica Coyle, e-mail: [email protected], 4308 Judah Apt #4, San Francisco, CA 94122,USA

Romain Darnajoux, e-mail: [email protected], 1480 Rue laterriere app 113Sherbrooke, Quebec J1K3A9, Canada

Evgeniy Davydov, e-mail: [email protected], Russia

Sally Eaton, Royal Botanical Garden Edinburgh, Edinburgh, Scotland, UK

David Díaz Escandón, e-mail: [email protected], Cra 79 #13b. - 159 apto 902-5, Cali,Valle, 760033, Colombia


Mgr. Zuzana Fačkovcová, Institute of Botany, Slovak Academy of Sciences, Dúbravskácesta 9, 845 23 Bratislava, Slovakia

Luis Fernando, e-mail: [email protected], [email protected], CocaHerbario Universidad de Caldas, Caldas, Manizales, Colombia

Lorenzo Fortuna, e-mail: [email protected], Via Licio Giorgieri 10, 34127Trieste, Italy

Julia Gerasimova, e-mail: [email protected], Komarov Botanical Institute, Laboratoryof Lichenology and Bryology, Popova street 2, 197376 St. Petersburg, Russia

Ricardo Miranda Gonzalez, e-mail: [email protected], 6105 NW Happy ValleyDr., Corvallis, OR-97330, USA

Georg Hillmann, e-mail: [email protected], Gnejsvaegen 8, SE-74731 Alunda, Sweden

Natalie Howe, e-mail: [email protected], 66 Fox Rd, 9B Edison, NJ 08817, USA

Beata Kazimiera Guzow-Krzeminska, University of Gdansk, Poland

Elisa Lagostina, Germany

Dr. Francoise Le Deveheat, Université de Rennes 1 – Faculté des Sciences pharmaceutiqueset biologiques, Laboratoire de Pharmacognosie et de Mycologie 2, Av. du Professeur LéonBernard - 35043 Rennes Cedex, France

James Lendemer, e-mail: [email protected], Institute of Systematic Botany, The NewYork Botanical Garden, Bronx, NY 10458, USA

Grzegorz Leśniański, e-mail: [email protected], ul. Wojska Polskiego 19a, 32-500 Chrzanów,Poland

Hanna Lindgren, e-mail: [email protected], 6151 N Winthrop Ave Apt 404,Chicago, IL 60660, USA

Tetiana Lutsak, e-mail: [email protected], Athener Str. 8, 60327 Frankfurt am Main,Germany

Jiří Malíček, e-mail: [email protected], Department of Botany, Charles University inPrague, Benátská 2, 128 01 Praha 2, Czech Republic

Joel Mercado, e-mail: [email protected], 5466 S. Harper 2A, Chicago, IL-60615, USA

Gaurav K. Mishra, e-mail: [email protected], CSIR – National Botanical ResearchInstitute, Rana Pratap, Marg Lucknow-226001, India

Alice Montagner, e-mail: [email protected], Via Licio Giorgieri 10, 34127Trieste, Italy

Carlos Pardo, e-mail: [email protected], 311 S LASALLE ST Apt. 34 L,Durham, NC 27705, USA

Dinah Parker, e-mail: [email protected], 77 Pond Point Avenue, Milford, CT 06460,USA


Eva Peña, e-mail: [email protected], Plz San Juan de Covas 16 8º A, 28925Alcorcon, Madrid, Spain

Vasun Poengsungnoen, e-mail: [email protected], Lichen Research Unit,Ramkhamhaeng University, Huamark, Bang Kapi, Bangkok, 10240, Thailand

Siri Rui, e-mail: [email protected], Tormods vei 19b, 1184 Oslo, Norway

Klara Scharnagl, e-mail: [email protected], 520 N Clemens Ave, Lansing,MI-48912, USA

Angela Figas Segura, e-mail: [email protected], Avda. Cataluña n64, 4F 12200 Onda,Castellon, Spain

Antoine Simon, e-mail: [email protected], Quai Marcellis 9, boîte 10, 4020 Liège,Belgium

Jana Steinová, e-mail: [email protected], Faculty of Science, Charles University inPrague, Albertov 6, Praha 2, 12843, Czech Republic

Victoria Tarasova, e-mail: [email protected], Varlamova 39-103, Petrozavodsk, 185005,Russia

Jonathan Terry, e-mail: [email protected], 2 Rhodes St, Newtown, Wellington,Wellington 6021, New Zealand

Kajonsak Vongshewarat, e-mail: [email protected], Biology Department,Faculty of Science, Ramkhamhaeng University, Bangkapi, Bangkok, 10240, Thailand

Monika Wagner, e-mail: [email protected], Fasaneriestraße 6/C6, A-5020Salzburg, Austria

Yolanda Wiersma, e-mail: [email protected], 37 Roche St., StJohns NewfoundlandA1B1L6, Canada

Karina Wilk, e-mail: [email protected], Institute of Botany, Polish Academy of Sciences,Lubicz 46, 31-512 Krakow, Poland

Tyler Wright, e-mail: [email protected], 8504 16th St. Apt 614, Silver Spring,MD 20910, USA

Zakieh Zakeri, e-mail: [email protected], Senckenberg Museum of NaturalHistory Görlitz, Am Museum 1, 02826 Görlitz, Germany


IAL Advisory Committee

Laurens Sparrius – Dutch Bryological and Lichenological Society – [email protected]

Arne Thell – Nordic Lichen Society – [email protected]

Chris Ellis – British Lichen Society – [email protected]

Paolo Giordani – Italian Lichen Society – [email protected]

Nobuo Hamada – Lichen Society of Japan – [email protected]

Susan Will-Wolf – American Bryological and Lichenological Society – [email protected]

Ana Rosa Burgaz – Spanish Lichenological Society – [email protected]

Volker John – Bryologisch-Lichenologische Arbeitsgemeinschaft für Mitteleuropa –

[email protected]

Mikhail Zhurbenko – Russia – [email protected]

Susana Calvelo – South America – [email protected]

Gintaras Kantvilas – Australasia – [email protected]

Paul Kirika – Africa – [email protected]

Khwanruan Papong – Asia – [email protected]

Paulina Bawingan – South East Asia – [email protected]

Auditor:

Ulf Arup, Botanical Museum, Lund University, Sölvegatan 37, 223 62 Lund, Sweden. E-mail: [email protected]

Vice Auditor:

Starri Heiðmarsson, Icelandic Institute of Natural History, Akureyri Division, Borgir vidNordurslod, IS-600 Akureyri, Iceland. E-mail: [email protected]

The cover-page illustration

Ramalina intermedia by Lucy C. Taylor, first published in American Arctic Lichens Vol 1.Macrolichens by J.W. Thomson

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