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ANNALS OF GEOPHYSICS, VOL. 48, N. 3, June 2005

Key words ionospheric tomography – total elec-tron content – ionospheric propagation – ionospher-ic modeling

1. Introduction

The development of tomographic imaginghas been one of the great scientific achieve-ments of the final decades of the twentieth cen-tury. While the mathematical basis of themethod was formulated early in the century, theadvances in computing technology in the 1960sgave practical realization to the technique as animaging tool. The method is hailed primarilyfor transforming medical diagnostics, but it hasalso lead to significant progress in many otherfields, including geophysics. The underlyingprinciples behind the method and descriptionsof some of the early applications can be foundin the special issue of Proceeding of the IEEE,71 (3), 1983. Further material of a fundamental

nature can also be found in the book by Kakand Slaney (1988).

A number of authors have reviewed the ap-plication of the method to radio tomographicimaging of the ionized atmosphere. Early de-velopments in the field are documented in spe-cial issues of the journals International Journalof Imaging Systems and Technology, 5 (2),1994 and Annales Geophysicae, 13 (12), 1996,while in further reviews Leitinger (1996, 1999)discussed the theoretical basis and the limita-tions of the ionospheric application. More re-cently, ionospheric tomography, includingcomprehensive descriptions of a number of the-oretical formulations, has been the subject of abook by Kunitsyn and Tereshchenko (2003). Inaddition, Pryse (2003) has reviewed experi-mental results from the application of radio to-mography to ionospheric imaging, concentrat-ing in the main on use of the method in investi-gations directed towards understanding of theunderlying geophysics. The present paper aimsto focus on aspects of the use of tomographicimaging as an aid to practical radio science ap-plications. Material relating to purely geophys-ical studies of the ionized atmosphere, whichwas presented in the oral version of the paper,can be found in the earlier review by Pryse(2003) and will not be repeated here.

Ionospheric tomography and its applications in radio science

and geophysical investigations

Leonard KersleyRadio and Space Physics Group, Department of Physics, University of Wales, Aberystwyth, U.K.

AbstractThe paper reviews recent progress in the development of radio tomographic imaging of the ionized atmosphere,particularly in relation to the use of the technique to assist in the mitigation of propagation effects on practicalradio systems.

Mailing address: Prof. Leonard Kersley, Radio andSpace Physics Group, Department of Physics, Universityof Wales, Aberystwyth, Dyfed SY23 3BZ, U.K.; e-mail:[email protected]

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The experimental results discussed here in-volve measurements of the line integral of theelectron density made along ray paths to groundstations from satellites in low Earth polar orbits.The Total Electron Content (TEC) is determinedusing the phase coherent signals from satellitesin the Navy Ionospheric Monitoring System(NIMS), monitored at a chain of receiving loca-tions aligned essentially in longitude, but sepa-rated in latitude. Inversion of the data set in a re-construction algorithm yields a pixelised imagein two dimensions of the spatial distribution ofthe electron density throughout the region of theray-path intersections. While many groupsworldwide have been involved in the develop-ment of radio tomographic imaging and thetechnique has now been used in many situa-tions, the results presented here are concerned inthe main with investigations of the potentialusefulness of the method to applications relatedto practical radio systems. The experimental ob-servations have been made for the most part atlocations instrumented by the UWA group inU.K. and Scandinavia. Maps with the current

stations operated by the group are shown in fig.1, though it should be noted that some of thework discussed here relates to results from ear-lier measurements, including those made at fivelocations in U.K. spanning the latitudinal rangebetween about 50°N and 61°N and campaignsinvolving additional sites in mainland Sweden.

The development work carried out in recentyears has demonstrated that radio tomographyhas now matured into a powerful experimentaltechnique capable of providing images of large-scale spatial structures in the ionosphere from alimited number of ground stations. The spatialnature of the observations is complementary tothose from many other types of experimentalmeasurements of the ionosphere. The limitationsto the method were thoroughly investigated andwell understood from early simulation studies.These result primarily from the limited-anglegeometry of the satellite-to-ground observationsso that there is incomplete information in theraw measurements about the vertical profile.However, even at an early stage of the develop-ment, algorithms were formulated capable of re-

Fig. 1. Maps showing locations of satellite receiving station chains in U.K. and Scandinavia currently in useby UWA group.

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Ionospheric tomography and its applications in radio science and geophysical investigations

producing ionospheric layers at different heightsthroughout the image (Fremouw et al., 1993). Inaddition, the temporal coverage is dependent onthe number and orbits of the available satelliteswith appropriate beacon transmitters.

2. Tomography and ionosondes

The ionosonde remains the basic experi-mental tool used to monitor the state of the ion-ized atmosphere for radio systems applications.

Fig. 2. Estimates of peak electron density (NmF2)obtained from the ionosondes at Chilton (left upper)and Lannion (left lower) as a function of time, togeth-er with (right) tomographic images from satellitepasses at three times throughout the night. The F de-notes spread-F conditions. From Kersley et al. (1997),reproduced by permission of the American Geophysi-cal Union.

Significant advances have been made in recentyears in the development of the instrument, notjust in the electronic hardware and digital pro-cessing of the sounding signals, but also in in-version software that can yield electron densityversus height profiles below the layer peak innear-real time. However, ionosondes can onlyprovide measurements for specific locations atdiscrete times. The spatial imaging capabilityof the radio tomography technique forms a nat-ural complement to the ionosonde for routinemonitoring, with synergies of mutual benefit to

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both methods. The addition of informationabout the horizontal structure of the ionospherefrom tomographic imaging can be used to placethe point measurements from the sounder intheir wider spatial context, while inclusion ofinput from an ionosonde in the reconstructionalgorithm can be used to mitigate the limitedcapability of the tomographic technique to de-termine heights.

The complementary nature of the two ex-perimental techniques is demonstrated in fig. 2.The upper and lower panels to the left show re-spectively measurements of the electron densi-ty at the layer peak (NmF2) obtained from theionosondes at Chilton (51.5°N) from Lannion(48.8°N), as a function of time during a night ofvery severe geomagnetic disturbance. The in-terpretation of these observations can be eluci-dated from the tomographic images obtainedfrom satellite passes at three times throughoutthe night (Kersley et al., 1997). The spatial im-ages show that the spread-F conditions are re-lated to irregularities at the poleward edge ofthe trough, while it is a boundary blob that is re-sponsible for the increased ionisation seen laterin the night at Slough, during these exceptionconditions where the trough minimum wasfound over Northern France.

Measurements by ionosondes have alsobeen used to improve the representation of thevertical profile in tomographic reconstructions.Discussion of the role of ionosonde input in re-construction algorithms, used as complementa-ry information to the TEC observations, can befound in Kersley et al. (1993). The paper wasalso the first of several that include verificationof the resultant image of electron density usingindependent results from the EISCAT incoherentscatter radar.

3. Tomography and empirical or parameterised models

Radio tomography is an experimental tech-nique that can be used to provide independentmeasurements with which to test the function-ality and reliability of the empirical and para-meterized models that are in use for the mitiga-tion of ionospheric effects on radio systems.

Fig. 3. An actual tomographic image of electrondensities (upper panel), with corresponding outputsfrom three ionospheric models: (in descending orderof panels) IRI-95, PIM and COST 238. From Dabasand Kersley (2003), reproduced by permission of theAmerican Geophysical Union.

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The region of the main trough, a dynamic fea-ture whose location and structure is highly vari-able and which is found over Northern Europeon many nights, presents particular problemsfor such models. Most models do not even at-tempt to incorporate the feature, while for oth-ers, placing the trough in the wrong place orwith inappropriate gradients may represent apoorer solution for a particular radio applica-tion than complete omission of the feature. To-mographic imaging of large-scale spatial struc-tures like the main trough provides a powerfultool to aid the verification of such models bycomparison with actual observations.

The panels of fig. 3, taken from Dabas andKersley (2003), show an example of such com-parison between a tomographic image and threedifferent ionospheric models, the InternationalReference Ionosphere (IRI-95) (Bilitza et al.,1993), the Parameterised Ionospheric Model(PIM) (Daniell et al., 1995) and the Europeanregional model developed during the COST238 Action (Bradley, 1999). The experimentalresults from observation at a tomographic chainof stations in U.K. show an ionosphere with aclear trough and in addition a mid-latitudenighttime enhancement to the south. The outputfrom the IRI-95 model for the same geophysi-cal conditions fails to replicate either feature,

while that from PIM does show a trough, butwith no localized enhancement to the south.The results from the COST 238 model do ap-pear to show a trough, though extreme cautionmust be used in the interpretation because thisparticular model was designed specifically tocover the European region below 55°N latitude,with default matching to the IRI model abovethat latitude, so that the feature shown is likelyto be an artifact of this process.

4. Tomography and the mapping of ionospheric parameters

Maps of ionospheric parameters, on a re-gional or global basis, provide another tool thatis used to correct for propagation effects inpractical radio systems. Dabas and Kersley(2003) also investigated the potential use of to-mographic images in mapping. They demon-strated that improvements could be achieved tothe reliability of maps of the F2-layer criticalfrequency and peak electron density over Eu-rope by incorporating information from experi-mental tomography into the mapping process.Figure 4 shows a comparison of a map ofNmF2, predicted by the IRI-95 model for par-ticular conditions, with one in which an actual

Fig. 4. Contour maps of NmF2 over Europe obtained from IRI-95 alone (left) and using input from an actualtomographic image plus the IRI-95 model (right). From Dabas and Kersley (2003), reproduced by permissionof the American Geophysical Union.

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tomographic image obtained from observationsmade in U.K. has been used to provide the lati-tudinal gradient. In essence, the electron densi-ties at the layer peak, determined from the im-age, were mapped in longitude, with zonal gra-

dients from the model being used to create a re-vised map that was more realistic of the actualconditions. The locations of the spatial struc-tures that were not present in the model outputalone can be identified from the map modifiedby the use of tomographic input.

Verification of the improvement afforded bythe use of tomographic observations in the map-ping procedure was obtained by comparing themapped values with actual measurements of foF2from the network of ionosondes throughout Eu-rope. Figure 5 shows the results of such compar-ison for the PIM model. It can be seen that inclu-sion of tomographic input with the PIM modelimproved the correlation coefficient between themapped NmF2 and actual ionosonde measure-ments to 0.92, while a smaller coefficient of 0.83was found when the PIM model was used alone.

5. Tomography and oblique ionograms

The possible role of radio tomography as anaid to oblique ionospheric sounding has been in-vestigated by Heaton et al. (2001). A network ofIRIS oblique sounders was established in U.K.

Fig. 5. Correlation of NmF2 from tomography plusPIM maps (upper) and PIM alone maps (lower) withcorresponding values from ionosondes. From Dabasand Kersley (2003), reproduced by permission of theAmerican Geophysical Union

Fig. 6. An oblique ionogram and bottomside pro-file of electron density from IRIS observations inU.K., together with a complete profile at the mid-point obtained from a corresponding tomographicimage. From Heaton et al. (2001), reproduced bypermission of the American Geophysical Union.

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during an experimental campaign, covering theregion spanned by a chain of tomographic re-ceivers. The results demonstrated that the maxi-mum density at the mid-point of the oblique pathestimated by inversion of the oblique ionogramwas in reasonable agreement with the density atthe corresponding height and latitude in the to-mographic image. The example shown in fig. 6confirms this finding, though discrepancies canbe seen at E-layer heights. It was concluded fromthe study that images of path conditions from thetomographic technique could have a role to playin assessing the applicability of the assumptionof spherical symmetry in the reduction ofoblique incidence ionograms.

6. Tomography and HF ray tracing

The observations from the IRIS oblique in-cidence sounder network in U.K. were also usedin a study of the potential of radio tomographyin aiding the ray tracing of HF propagationpaths. The results are discussed in detail inRogers et al. (2001). The study investigated theeffects of various types of input information thatcan be used to constrain the vertical electrondensity structure in the tomographic reconstruc-

tions. It was found that the use of a fine heightresolution (5 km) and incorporation of inputfrom one vertical ionosonde in the reconstruc-tion process made significant improvements tothe overall reliability of the tomographic image.As expected, E-layer propagation was better de-fined using a climatological model than by thetomographic method. It can be seen from fig. 7that the use of tomographic images reduced theRMS error in the determination of the F2-layerMaximum Useable Frequency (MUF), yieldingsignificantly smaller errors than those obtainedfrom three independent ionospheric models(FAIM, PIM and IRI-95).

7. Tomography and HF direction finding

Warrington et al. (2002) presented observa-tions from an HF direction-finding experiment,on a link between Uppsala in Sweden andLeicester in U.K., where the propagation pathwas aligned approximately with the orientationof the main ionospheric trough. An example isshown here that demonstrates significant devia-tions from the great-circle path. The resultsfrom the days 326-327, reproduced here in fig.8, show anomalous out-of-great-circle propaga-

Fig. 7. Histograms of RMS errors in the estimation of the maximum useable frequency showing that use of to-mographic images gives smaller errors for F2-layer MUF than the FAIM, PIM and IRI 95 models. From Rogerset al. (2001), reproduced by permission of the American Geophysical Union.

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tion during the night, with consequent in-creased times of flight and doppler frequencyshifts. The azimuth of the received signalsshows a sudden transition from the great-circle

direction, followed by a smooth rotation and re-versal over a period of several hours before re-turning to the original direction. In a subse-quent study, images of electron density from

Fig. 8. Results from an HF direction-finding experiment between Uppsala to Leicester during a three-day pe-riod in November 2001. Note in particular the anomalous azimuthal directions of the received signals during thenight of 326-327. From Warrington et al. (2002), reproduced by permission of the authors.

Fig. 9. A sequence of tomographic images from four satellite passes throughout the night of 326-327, moni-tored at the stations in U.K., which show a deep trough with a strong ionisation enhancement or boundary blobpolewards of the minimum that is acting as a reflector for the out-of-great-circle HF propagation found on theUppsala to Leicester path shown in fig. 8.

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the U.K. tomography chain, from a successionof satellite passes throughout the night, re-vealed a trough with a steep poleward wall thatadvanced equatorwards during the early part ofthe night before receding again northwards (fig.9). The anomalous propagation found in the re-sults from the HF experiment can be explainedin terms of reflection by the enhanced densitiesto be seen in the tomographic images polewardsof the trough minimum.

8. Tomography and vertical TEC

Kersley et al. (2002) investigated the limita-tions to the accuracy of experimental measure-ments of total electron content imposed by thenecessary assumptions required to convert the

slant observations to equivalent vertical meas-urements. The assumed height of the thin-shellionospheric layer was found to be a critical fac-tor. However, it was demonstrated that someadvantage could be gained by assessing the ver-tical TEC by integration through a tomographicimage as opposed to using the equivalent verti-cal estimate of the parameter for an assumedthin-layer height. In the example shown in fig.10 it can be seen that to the north of the troughthe tomographic image reveals that the peakheight of the actual ionosphere was only some200 km. The lower panel shows that the result-ant equivalent vertical total electron contentsestimated from the slant measurements at theindividual stations for an assumed ionosphericheight of 350 km are all much less than the ver-tical TEC (shown by the dotted line) in that re-

Fig. 10. Tomographic image from observations in U.K. (upper panel), with plots of equivalent vertical totalelectron content from the individual station observations (solid lines) and the vertical total electron content com-puted through the tomographic image (dotted line) shown in the lower panel. From Kersley et al. (2002).

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gion obtained from the tomographic imageabove. In a similar way, an assumed thin-shellionosphere at too great a height leads to overes-timation of the equivalent vertical TEC. It wasargued that the vertical TEC obtained from thetomographic image was a more reliable esti-mate than the equivalent vertical TECs deter-mined from the slant measurements.

9. Tomography and trough parameters

Vertical TEC estimates from tomographicimages of the kind described above are beingused in a project to characterize the main troughin terms of a set of defined parameters. The pa-rameters have been chosen to identify both thelocation of the trough and the local structure ofthe ionosphere in the vicinity of the feature. Inaddition to the latitude of the minimum, the pa-rameters include the gradients of both the equa-torward and poleward walls and evidence for alocal maximum that would represent a bound-ary blob poleward of the minimum. A databaseof the parameters has been compiled, using theobservations from the tomographic chain of sta-tions in U.K., covering more than one year. Theobjectives of the study are two fold. First, theaim is to use the parameters in the extensivedatabase to try to further understanding of thebehaviour of the trough under a wide variety ofgeophysical conditions. In addition, characteri-sation of the feature in this defined way isplanned to provide parameters that can be cal-culated directly from models so that the actualobservations can be used in a direct test of thereliability of the model in the trough region.

10. Tomography and the SUCTIP model

While the work outlined above involved thetesting of models that had been developedspecifically for radio systems applications, to-mographic images have also been used to as-sess the ability of a physical model of the cou-pled ionosphere/thermosphere system to repli-cate conditions in the high-latitude ionosphere.Experimental observations from a chain ofsatellite stations in the Scandinavian sector

were used to provide ‘ground truth’ informationfor comparison with output from the SheffieldUniversity Coupled Thermosphere IonospherePlasmasphere (SUCTIP) model. Figure 11, takenfrom the paper by Idenden et al. (1998) report-ing the results of the study, shows an exampleof a tomographic image containing a daysidetrough in the high-latitude ionosphere that canbe compared with the output from the SUCTIP

model for the appropriate conditions, which isreproduced in the panel below.

A subsequent study by Balthazor et al. (2002)compared the locations of the trough minimum,determined experimentally from tomographicimages made under quiet geomagnetic condi-tions, with those predicted by the model (fig. 12).

The results of the comparison shown in fig.12 demonstrate that the model tends consistent-

Fig. 11. Tomographic image from experimental ob-servations in Scandinavia with corresponding outputfrom the SUPIM model. From Idenden et al. (1998),reproduced by permission of Annales Geophysicae.

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ly to place the trough at slightly lower latitudesthan those identified experimentally from thetomographic observations.

11.Tomography and space weather

Many of the processes that can have adverseeffects on the performance of radio systems areof particular significance during times when dis-turbed space weather impacts on the terrestrialenvironment. Radio tomographic imaging ofionospheric structures found at high latitudes hasbeen shown to provide insight into space-weath-er processes. The convergence of the geomag-netic field close to the Earth results in flux tubeswith very different plasma populations that arecharacteristic of widely separate regions of spacebeing brought into close spatial proximity in thedayside cusp ionosphere. In consequence, struc-tures in the ionospheric plasma represent signa-tures of physical processes that are operating farout in space. Tomographic imaging of such spa-tial features has demonstrated that they containfootprints of important mechanisms in solar-ter-

restrial coupling. Several papers have addressedsuch issues, including two that illustrate the sig-natures in the dayside cusp ionosphere of the re-connection processes that link the InterplanetaryMagnetic Field (IMF), carried by the solar wind,to the geomagnetic field.

The location of the magnetic reconnectionand the resultant behaviour and flow of theionospheric plasma is dependent, in particular,on the orientation of the north/south- or z-com-ponent of the IMF. When Bz is directed south-wards the IMF is oriented anti-parallel to the ge-omagnetic field at the magnetopause at low lati-tudes, so that the reconnection occurs close to theequatorial plane. The resultant tension exerted bythe solar wind on the geomagnetic field drivesthe ionospheric plasma in the familiar two-cellpattern at high latitudes, with anti-sunward flowacross the polar cap, as is illustrated in fig. 13.

Walker et al. (1998), using observations fromthe chain of tomographic stations in the vicinityof Svalbard, investigated the ionospheric foot-prints of low-latitude reconnection. An image, re-produced in fig. 14, shows characteristic featuresthat can be identified with signatures of the re-connection process. The open/closed field lineboundary is marked by a steep meridional gradi-ent in the F2-layer ionospheric electron density.The increase in the peak height of the layer to thenorth was associated with the dispersed spectrumof the particles precipitating from the magne-topause reconnection site, with the softer energies

Fig. 12. Polar plot in the magnetic reference framecomparing average locations of the trough minimumunder quiet geomagnetic conditions obtained fromtomographic images (+) with estimates by the SUCTIP

model (∗). From Balthazor et al. (2002).

Fig. 13. Magnetic reconnection close to the equato-rial plane and the consequent convection of plasmain the high-latitude ionosphere, driven in an anti-sun-ward flow across the polar cap. From Lockwood(1998), reproduced by permission of the publishers.

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taking longer to impact the ionosphere as the fluxtube is convected polewards. The tomographicimage also revealed ionospheric signatures of thefield-aligned current system driven by the recon-nection. The E-layer enhancement just south ofthe boundary was linked to harder electron pre-cipitation from the central plasma sheet associat-ed with an upward current, while the narrow bandof increased densities extending to high altitudesimmediately polewards of the reconnection loca-tion marked a downward field-aligned current.

When the IMF is directed northwards theanti-parallel condition necessary for reconnec-tion with the geomagnetic field is now satisfiedat high latitudes in the magnetospheric lobe.The resultant lobe reconnection and the conse-quent convection of plasma in the polar iono-sphere are illustrated in fig. 15. Pryse et al.(1999) identified a tomographic image that con-tained the spatial footprints in the high-latitudedayside ionosphere of lobe reconnection, whichis reproduced here as fig. 16. There is a markedcontrast between this signature of reconnectionin the magnetospheric lobe and that shown ear-lier for the low-latitude situation. Figure 15 il-lustrates that for lobe reconnection the initialmotion of the flux tubes, driven by the tensionof the IMF solar wind, is equatorwards with aconsequent reversed dispersion of the spectrumof the precipitation as a function of latitude. Theresultant signature of this process can be seen inthe tomographic image, where the increasingpeak height of plasma in the flux tubes convect-ing from the north towards the adiaroic bound-ary reflects the softening of the precipitationdriven by the reconnection process. The adi-aroic boundary, across which there is no trans-fer of flux, is marked by a steep gradient in thedensity. Plasma from the south is driven in theanti-sunward direction up to the boundary and

Fig. 14. Tomographic image showing signatures in the high-latitude ionosphere of reconnection at the equato-rial magnetopause. From Walker et al. (1998), reproduced in modified form by permission of the American Geo-physical Union.

Fig. 15. Magnetic reconnection in the magnetos-pheric lobe and the consequent convection of plasmain the high-latitude ionosphere, which polewards ofthe adiaroic boundary is initially in a sunward direc-tion. From Lockwood (1998), reproduced by permis-sion of the publishers.

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then carried zonally by viscous processes. Itshould be noted that the trajectory of the satel-lite, which was to the east of the chain of ob-serving stations, gives rise to a geometrical dis-tortion resulting in the apparent reverse slopewith altitude of the field-aligned boundary.

Tomographic images from observations athigh latitudes have also been used to investigateother signatures of the impact of space-weatherprocesses in the ionospheric plasma. For exam-ple, Moen et al. (1998) related structures in elec-tron density seen in tomographic images to opti-cal aurorae, with the precipitation responsible forgreen-line emissions being linked to density en-hancements in the E-layer, while the softer parti-cles given rise to red-line aurora produced dis-crete features in the plasma at F-layer heights.

12. Conclusions

The results reviewed here demonstrate thepotential role of radio tomographic imaging ofthe ionized atmosphere in both applied radio sci-ence, directed towards the mitigation of propa-gation effects on practical radio systems, and in

studies of the geophysics of the underlying phys-ical processes. The experimental technique isable to provide a unique capability to monitorlarge-scale spatial structures in the electron den-sity over a wide region from observations madeat a limited number of ground stations. The ad-vantages of method, given by wide coverageeven in inaccessible and environmentally hostileregions, coupled with the low cost of a passivesystem, far out way the known limitations. Thelatter result primarily from the lack of horizontalray paths in satellite-to-ground propagation,though they can be mitigated by use of appropri-ate reconstruction algorithms and with addition-al input from ionosondes or other independentsources. The recent studies outlined here haveshown the potential of radio tomography infields as diverse as the validation of both empir-ical and physical models of the ionosphere,oblique sounding, HF ray tracing, direction find-ing and the mapping of ionospheric parameters.The versatility of tomographic imaging in ap-plied radio science adds to the potential of themethod as a new experimental technique that canbe used in geophysical studies to identify signa-tures of space-weather processes in the ionized

Fig. 16. Tomographic image showing signatures in the high-latitude ionosphere of reconnection at the magne-tospheric lobe. From Pryse et al. (1999), reproduced by permission of the American Geophysical Union.

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atmosphere at high latitudes. A fuller review ofthe wider use of tomographic imaging in globalaspects of geophysics can be found in Pryse(2003).

Acknowledgements

The development of radio tomography bythe Radio and Space Physics Group at UWAhas been supported financially at various stagesby the U.K. Particle Physics and AstronomyResearch Council, the RadiocommunicationsAgency, DERA and EOARD. The importantcontributions of many individuals, both withinthe group and as external scientific collabora-tors are acknowledged with grateful thanks, asis the assistance of many people and organisa-tions for the practical support provided for theexperimental activities.

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Ionospheric tomography and its applications in …...537 Ionospheric tomography and its applications in radio science and geophysical investigations producing ionospheric layers at