17
Cretaceous Research (1999) 20, 447–463 Article No. cres.1999.0160, available online at http://www.idealibrary.com on Palynology and sequence stratigraphy of the Cretaceous of eastern Venezuela *J. Helenes and D. Somoza *CICESE, Departamento de Geologia, Carretera Tijuana-Ensenada, Ensenada, Baja California 22860, Me ´xico ²Servicios Geolo ´gicos Especializados, Exploracio ´n y Produccio ´n-PDVSA, Puerto La Cruz, Edo. Anzoa ´tegui, Venezuela Revised manuscript accepted 7 April 1999 From Jurassic to Tertiary times, northeastern Venezuela represented the continental shelf of a tectonically passive margin. A dominantly marine Cretaceous sedimentary section, within an overthrusted allochthonous block and with a maximum thickness of 3658 m, is found in the subsurface in this area. Palynomorph recovery from these strata, together with planktonic foraminifera and calcareous nannofossils, indicate an age-range from at least Aptian to Maastrichtian. The palynological assemblages observed exhibit changes with time and environments, which are here related to the sedimentary evolution. Integrated analyses of palynology, lithology, and well logs are the basis for the reconstruction of the sedimentary evolution in the area. The oldest marine sediments in this section correspond to the uppermost part of the Lower Zuni Megacycle and contain a maximum flooding surface correlative with that dated at 111 Ma. The rest of the sequence represents a major transgressive-regressive cycle (Upper Zuni) and contains seven sequences correlative with global eustatic cycles. The Turonian transgression dated at 91.5 Ma is the most important marine transgression in the entire section and is recognizable in all sections studied, while the Maastrichtian-Paleocene boundary dated at 68 Ma is not always clearly recognizable. ? 1999 Academic Press K W: Cretaceous; Aptian-Maastrichtian; dinoflagellates; biostratigraphy; paleoenvironments; sequence stratigraphy; passive margin; eastern Venezuela. 1. Introduction 1.1. Objectives This paper reports on the stratigraphy and paly- nological content, particularly dinoflagellates, of Cretaceous strata drilled in an allochthonous thrust block from the Eastern Basin of Venezuela (Figure 1). The objectives of the paper are to expand our knowl- edge on the stratigraphic distribution of Cretaceous dinoflagellate assemblages in tropical, shallow mar- ine environments, and to present a chronostrati- graphic model for the evolution of the Cretaceous in Venezuela’s Eastern Basin. The model proposed is based on a sequence stratigraphic analysis and corre- lation with global eustatic cycles (Haq et al., 1988) and serves to correlate the strata studied with other Cretaceous sections in the region. The data on paly- nomorphs will help to refine the known biostrati- graphic ranges of the species included and to improve our paleoenvironmental interpretations of Tethyan palynomorph assemblages. 1.2. General setting Global eustatic cycles for the Mesozoic and Cenozoic were first proposed by Vail et al. (1977). These cycles were then correlated with published biostratigraphic zonations and calibrated with a paleomagnetic scale and radiomagnetic ages by Haq et al. (1988). In the stratigraphic record of a continental margin, the cycles are recognized by transgressions and regressions, which can be linked to the rise and fall of the global (or local) sea level. The amount of sediments reaching a basin and the space available to accommodate them are controlled by changes in the volume of the world’s oceans, changes in the amount of water filling them and continental uplift or subsidence (Vail et al., 1991). This uplift or subsidence can be caused by tectonic events or by lithostatic reactions to the loading and unloading of the continental margins. Tectonic processes can obscure the relation of transgressive-regressive sedimentary successions to global eustatic cycles. A tectonically passive, con- tinental margin such as the Cretaceous of eastern 0195–6671/99/040447+17 $30.00/0 ? 1999 Academic Press

Palynology and sequence stratigraphy of the Cretaceous of ... (1).pdfVenezuela’s Eastern Basin. The palynological results are based mainly on the analysis of samples from the following

  • Upload
    others

  • View
    10

  • Download
    0

Embed Size (px)

Citation preview

Page 1: Palynology and sequence stratigraphy of the Cretaceous of ... (1).pdfVenezuela’s Eastern Basin. The palynological results are based mainly on the analysis of samples from the following

Cretaceous Research (1999) 20, 447–463Article No. cres.1999.0160, available online at http://www.idealibrary.com on

Palynology and sequence stratigraphy of theCretaceous of eastern Venezuela

*J. Helenes and †D. Somoza

*CICESE, Departamento de Geologia, Carretera Tijuana-Ensenada, Ensenada, Baja California 22860, Mexico†Servicios Geologicos Especializados, Exploracion y Produccion-PDVSA, Puerto La Cruz, Edo. Anzoategui,Venezuela

Revised manuscript accepted 7 April 1999

From Jurassic to Tertiary times, northeastern Venezuela represented the continental shelf of a tectonically passive margin. Adominantly marine Cretaceous sedimentary section, within an overthrusted allochthonous block and with a maximumthickness of 3658 m, is found in the subsurface in this area. Palynomorph recovery from these strata, together with planktonicforaminifera and calcareous nannofossils, indicate an age-range from at least Aptian to Maastrichtian. The palynologicalassemblages observed exhibit changes with time and environments, which are here related to the sedimentary evolution.Integrated analyses of palynology, lithology, and well logs are the basis for the reconstruction of the sedimentary evolution inthe area. The oldest marine sediments in this section correspond to the uppermost part of the Lower Zuni Megacycle andcontain a maximum flooding surface correlative with that dated at 111 Ma. The rest of the sequence represents a majortransgressive-regressive cycle (Upper Zuni) and contains seven sequences correlative with global eustatic cycles. TheTuronian transgression dated at 91.5 Ma is the most important marine transgression in the entire section and is recognizablein all sections studied, while the Maastrichtian-Paleocene boundary dated at 68 Ma is not always clearly recognizable.

? 1999 Academic Press

K W: Cretaceous; Aptian-Maastrichtian; dinoflagellates; biostratigraphy; paleoenvironments; sequencestratigraphy; passive margin; eastern Venezuela.

1. Introduction

1.1. Objectives

This paper reports on the stratigraphy and paly-nological content, particularly dinoflagellates, ofCretaceous strata drilled in an allochthonous thrustblock from the Eastern Basin of Venezuela (Figure 1).The objectives of the paper are to expand our knowl-edge on the stratigraphic distribution of Cretaceousdinoflagellate assemblages in tropical, shallow mar-ine environments, and to present a chronostrati-graphic model for the evolution of the Cretaceous inVenezuela’s Eastern Basin. The model proposed isbased on a sequence stratigraphic analysis and corre-lation with global eustatic cycles (Haq et al., 1988)and serves to correlate the strata studied with otherCretaceous sections in the region. The data on paly-nomorphs will help to refine the known biostrati-graphic ranges of the species included and to improveour paleoenvironmental interpretations of Tethyanpalynomorph assemblages.

0195–6671/99/040447+17 $30.00/0

1.2. General setting

Global eustatic cycles for the Mesozoic and Cenozoicwere first proposed by Vail et al. (1977). These cycleswere then correlated with published biostratigraphiczonations and calibrated with a paleomagnetic scaleand radiomagnetic ages by Haq et al. (1988). In thestratigraphic record of a continental margin, the cyclesare recognized by transgressions and regressions,which can be linked to the rise and fall of the global(or local) sea level.

The amount of sediments reaching a basin and thespace available to accommodate them are controlledby changes in the volume of the world’s oceans,changes in the amount of water filling them andcontinental uplift or subsidence (Vail et al., 1991).This uplift or subsidence can be caused by tectonicevents or by lithostatic reactions to the loading andunloading of the continental margins.

Tectonic processes can obscure the relation oftransgressive-regressive sedimentary successions toglobal eustatic cycles. A tectonically passive, con-tinental margin such as the Cretaceous of eastern

? 1999 Academic Press

Page 2: Palynology and sequence stratigraphy of the Cretaceous of ... (1).pdfVenezuela’s Eastern Basin. The palynological results are based mainly on the analysis of samples from the following

448 J. Helenes and D. Somoza

Venezuela, could be a good place to try and separateeustatic from tectonic cycles. In addition, shallowmarine environments represented in continental mar-gin settings, usually exhibit lithologic and micro-paleontological changes that are obvious. Therefore inpassive margin, marine-dominated sedimentary suc-cessions, transgressive-regressive cycles are normallyrecognizable and could be related to eustatic cycles.

Cretaceous strata in eastern Venezuela have beenextensively studied from outcrops in the Serranıa del

Interior and areas to the north (Gonzalez de Juana etal., 1980; Morris et al., 1990; Bellizzia & Dengo,1990; Erikson, 1994; Stephan et al., 1994). Depo-sitional environments represented in these strata rangefrom continental and nearshore in the south and west,to deep-water marine paleoenvironments in the northand northeast.

The known stratigraphic record from the sub-surface in eastern Venezuela indicates that mostlyshallow marine sediments were deposited during the

Figure 1. Main tectonic and geologic elements in Venezuela and location of wells studied.

Page 3: Palynology and sequence stratigraphy of the Cretaceous of ... (1).pdfVenezuela’s Eastern Basin. The palynological results are based mainly on the analysis of samples from the following

Palynology and sequence stratigraphy of the Cretaceous of eastern Venezuela 449

Cretaceous Period (Gonzalez de Juana et al., 1980).During most of this time, the northern margin ofSouth America was passive, particularly in the areanow occupied by eastern Venezuela. Therefore theCretaceous sequences in eastern Venezuela provide anexcellent opportunity to compare paleoenvironmentalchanges in the sedimentary record to global eustaticchanges, without important tectonic components. Ananalysis of the micropaleontological record of thesesequences, as the one undertaken in this study, shouldprovide evidence to relate all these changes.

1.3. Tectonic setting

Prior to the Jurassic, the Caribbean did not exist andthe North and South American continental plateswere united. During the Jurassic, plate separationoccurred and by Campanian-Maastrichtian times theproto-Caribbean plate appeared, extending from thewest (Pindell & Barrett, 1990; Pindell & Erikson,1993). Today the northern edge of the SouthAmerican continent is extensively deformed by right-lateral, strike-slip faulting, and an oblique subductionarea is present to the north (Molnar & Sykes, 1969;Burke et al., 1984; Pindell & Barrett, 1990).

As a result of this evolution, the Caribbean region istectonically complex and consists of a mixed oceanicand continental North American plate; a mostlyoceanic Caribbean plate; the mixed oceanic andcontinental South American plate; and the oceanicPacific and Nazca plates.

In Venezuela (Figure 1), the main geologicalelements (Gonzalez de Juana et al., 1980) are thestable Guyana Shield to the south, the Miocene oryounger Andes mountains to the northwest, and theMiocene or younger right-lateral, strike-slip faultalong the Andes and to the east, known as theBocono-Moron-El Pilar Fault System (Schubert,1984), labeled as Bocono F. Z. and El Pilar F. Z. onFigure 1.

The Eastern Basin of Venezuela (Figure 1) islocated north of the Guyana Shield and south ofthe Serranıa del Interior, which was formed bylate Tertiary compression of the Caribbean and theSouth American plates. In the subsurface, a complexstructural regime is dominated by the late MiocenePirital Thrust Fault which dips to the north. In thispaper we present data on the sedimentary sequencesdrilled in an allochthonous block overlying the PiritalFault, and located east of the Urica Fault (see inset onFigure 1).

1.4. Stratigraphy

The oil-producing Tertiary strata of the Eastern Basinof Venezuela have been drilled and studied for a long

time (Gonzalez de Juana et al., 1980). However, thebasin also contains Cretaceous strata with an age-range from Barremian to Maastrichtian (Paredeset al., 1994). These Cretaceous rocks representalmost continuous marine sedimentation from atleast Aptian to Maastrichtian times. PresumedBarremian strata represent mainly continental toparalic paleoenvironments.

In the allochthonous block, the Cretaceoussequence is unconformably overlain by Tertiary strataand overlies thrusted Miocene strata (Figure 2). TheCretaceous units drilled have been correlated withstratigraphic units cropping out in the Serranıa delInterior (Figure 2), and the same stratigraphicnomenclature is applied to the subsurface sections. Ingeneral, the stratigraphic units in the subsurfacerepresent shallower facies than those present in theSerranıa del Interior; but deeper than those present tothe south (Gonzalez de Juana, et al. 1980).

Cretaceous Units. The oldest drilled unit is theBarranquın Formation, which is composed mainly ofcoarse to medium grained clastics. Conformably over-lying this unit is the calcareous El Cantil Formation,which includes a basal shaly member known as theMiembro Garcıa. Conformably overlying the ElCantil limestones rest the black, thin-bedded,calcareous shales of the Querecual Formation. Con-formably overlying the black shales, the San AntonioFormation consists of alternating sandstones andshales. The uppermost unit of the Cretaceoussequence is the San Juan Formation, which consists ofthick-bedded, medium to coarse grained sands.

The San Juan Formation is conformably overlainby alternating siltstones and dark gray shalesassigned to the Vidono Formation (Paleocene),and/or by glauconitic sandstones and siltstones of theCaratas Formation (Eocene). Unconformably over-lying these units, are the coarse clastics of theMorichito Formation (Pliocene) in the western partof the study area, and the medium to coarse clasticsof the Merecure Group (Oligocene to Miocene) inthe eastern part.

2. Material and methods

2.1. Data

The material used includes core and cuttings samplesfrom four exploratory wells drilled by CORPOVEN (aVenezuelan oil company, now part of PDVSA) inVenezuela’s Eastern Basin. The palynological resultsare based mainly on the analysis of samples from thefollowing wells.

Page 4: Palynology and sequence stratigraphy of the Cretaceous of ... (1).pdfVenezuela’s Eastern Basin. The palynological results are based mainly on the analysis of samples from the following

450 J. Helenes and D. Somoza

Well Cretaceousdrilledm (ft)

Cuttingssamplesanalyzed

Coresamplesanalyzed

PIC-1E 3,600 (11,630) 134PIC-3E 3,674 (11,870) 19PIC-3E 2,400 (7,803) 84SBC-3 900 (2,800) 43 6

Lithological interpretations are based on the gammaray well logs and the cuttings samples. Microfossilcontent was used to construct a biostratigraphic andpaleoenvironmental framework for the sequencesanalyzed. Age assignments of the sections studied arebased mainly on the stratigraphic ranges of the paly-nomorphs contained in cuttings and core samples.When assigning ages based on fossils contained incuttings samples, mainly uppermost appearances oftaxa were considered.

2.2. Sample preparation

Samples for palynological analyses were prepared fol-lowing the normal treatment with HCl, HF andZnBr2 (Wood et al., 1996). The residue was thensieved through 100 ìm and 20 ìm mesh sieves, and

strew mounts were prepared with the residue retainedon the 20 ìm. One slide per sample was analyzedunder the microscope, and numerical abundances ofmarine and terrestrial palynomorphs were recorded.

2.3. Palynological Marine Index

Since the recovery from all samples was low tomoderate, a precise statistical analysis was not under-taken. Instead, to help in the interpretation of thedepositional environments, a Palynological MarineIndex (PMI) was calculated for all samples analyzed.This PMI was calculated using the formula:PMI=(Rm/Rt+1) 100, where Rm=Richness ofmarine palynomorphs (dinoflagellates, acritarchs andchitinous internal molds of foraminifera), counted asthe number of taxa per sample; and Rt=Richness ofterrestrial palynomorphs (pollen and spores), alsocounted as the number of taxa per sample.

Null values of PMI indicate samples without marinepalynomorphs, and are interpreted to represent terres-trial or freshwater environments. Low values of PMIare interpreted as indicative of brackish water influ-ence, and higher values are interpreted as indicative ofmarine conditions of deposition. When compared to a

Figure 2. Cretaceous stratigraphy of the Serranıa del Interior, as applied to the allochthonous block.

Page 5: Palynology and sequence stratigraphy of the Cretaceous of ... (1).pdfVenezuela’s Eastern Basin. The palynological results are based mainly on the analysis of samples from the following

Palynology and sequence stratigraphy of the Cretaceous of eastern Venezuela 451

paleobathymetric curve based on benthonic foraminif-era, the higher values of the PMI generally correspondto neritic environments, while lower PMI valuescorrespond to either paralic or bathyal environments.

Palynological analyses of Recent sediments from theOrinoco Delta (Muller, 1959) indicate that dinoflag-ellate cyst recovery is restricted to brackish and marineenvironments, being more abundant in fully marineenvironments. On the other hand, the abundance ofspores and pollen in marine environments generally

decreases offshore (Hoffmeister, 1954; Muller, 1959;Heusser, 1978).

The lower PMI values in bathyal environmentsreflect the tendency of cyst-forming dinoflagellates toinhabit shelf or coastal areas (Dale, 1996) whereseasonal changes, such as temperature, that triggerexcystment are more pronounced (Evitt, 1985).

The PMI is particularly useful when comparingadjacent samples. Significant differences suggestchanges in depositional environment represented in

Figure 3. Gamma ray log, palynological statistics, sequences, and formations recognized in PIC-1E.

Page 6: Palynology and sequence stratigraphy of the Cretaceous of ... (1).pdfVenezuela’s Eastern Basin. The palynological results are based mainly on the analysis of samples from the following

452 J. Helenes and D. Somoza

Page 7: Palynology and sequence stratigraphy of the Cretaceous of ... (1).pdfVenezuela’s Eastern Basin. The palynological results are based mainly on the analysis of samples from the following

Palynology and sequence stratigraphy of the Cretaceous of eastern Venezuela 453

the section and help in the recognition of sequenceboundaries and maximum flooding surfaces.Sequence boundaries will normally be found in inter-vals with upward decreasing values of PMI; whilemaximum flooding surfaces will be near the maximumvalues of PMI in each sequence.

2.4. Other analyses

Foraminifera and calcareous nannofossils have beenstudied (Paredes et al., 1994) and the age assignmentsbased on these calcareous fossils are used here inconjunction with those determined on the basis ofpalynology.

The gamma ray and conductivity logs wereanalyzed to determine grain size gradation. Resultsfrom these analyses, together with the lithologyobserved in core and cuttings samples, and the PMIvalues obtained from the palynological analyses, allowinterpretation of the depositional systems and thestratigraphic sequences represented. Finally, theintegration of the stratigraphic sequences recognizedand the palynological ages assigned, are the bases for acorrelation with the chronostratigraphic chart of Haqet al. (1988).

2.5. Previous palynological studies

Most of the information on the stratigraphic distribu-tion of Cretaceous dinoflagellates is based on samplesfrom either mid- to high latitudes in the northernhemisphere (Helenes, 1984; Williams & Bujak, 1985;Aurisano, 1989; Powell, 1992; Williams et al., 1993),or from the southern hemisphere (Cookson &

Eisenack, 1960a, b, 1962; Helby et al., 1987; Stover &Helby, 1987).

The palynological content of Cretaceous stratafrom Venezuela and the Tethyan region in general,particularly the terrestrial palynomorphs, have beenextensively studied (Germeraad et al., 1968; Mulleret al., 1987; Colmenares & Teran, 1993). Fossildinoflagellates from these areas are not so well known.However, some of the publications on Cretaceousdinoflagellates related to this region are: Lentin &Williams (1980); Malloy (1972); Jain & Millepied(1973); Pares-Regali et al. (1974); Davey (1978);Below (1984); Fasola & Paredes-de-Ramos (1991);Paredes et al. (1994); Helenes et al. (1991, 1994,1998).

3. Paleoenvironments and sequences

3.1. General observations

The paleoenvironmental changes in the section drilledare interpreted from the gamma ray logs and thepalynomorph assemblages. PMI values are used tosummarize the palynological data, providing ameans of discriminating marine from non-marineenvironments.

PIC-1E (Figures 3, 4) had the best palynomorphrecovery and highest abundances of marine paly-nomorphs, and therefore provides the best record ofpaleoenvironmental changes in the basin. Compara-tively, PIC-5E (Figure 5) contains fewer marinepalynomorphs and PIC-3E (Figure 6) and SBC-3(Figure 7) have less complete Cretaceous sections.

The values of PMI from PIC-1E and PIC-5E(Figures 4, 5) show relatively poor recovery of

Figure 4. Main fossils, paleontological ages, and sequences recognized in PIC-1E (on facing page). Key to symbols andabbreviations (above) also applies to Figures 5–7: Low.=lowermost.

Page 8: Palynology and sequence stratigraphy of the Cretaceous of ... (1).pdfVenezuela’s Eastern Basin. The palynological results are based mainly on the analysis of samples from the following

454 J. Helenes and D. Somoza

Figure 5. Main fossils, paleontological ages, and sequences recognized in PIC-5E. For key to symbols and abbreviations, seeFigure 4.

Page 9: Palynology and sequence stratigraphy of the Cretaceous of ... (1).pdfVenezuela’s Eastern Basin. The palynological results are based mainly on the analysis of samples from the following

Palynology and sequence stratigraphy of the Cretaceous of eastern Venezuela 455

marine palynomorphs in the bottom half and aconsiderable increase in the upper part of thesection. These characteristics of the palynomorphassemblages and the interpretation of the gammaray logs are used to define seven Cretaceoussequences, labeled from B to H, in PIC-1E. Based ontheir lithologic and palynological characteristics,some of the sequences are also recognized in theother wells.

3.2. Tertiary sequences

Sequences A, I, and J are Tertiary in age, so they arenot described in detail in this paper. Their generalcharacteristics are the following.

Sequence A. Corresponds to the Miocene sedimentaryrocks underlying the thrust fault and representsSupercycle TB-2 of Haq et al. (1988).

Figure 6. Main fossils, paleontological ages, and sequences recognized in PIC-3E. For key to symbols and abbreviations, seeFigure 4.

Page 10: Palynology and sequence stratigraphy of the Cretaceous of ... (1).pdfVenezuela’s Eastern Basin. The palynological results are based mainly on the analysis of samples from the following

456 J. Helenes and D. Somoza

Sequence I. Present only in short intervals of PIC-3Eand SBC-3, it is probably incompletely represented.Characterized by low values of the PMI, it containsthe dinoflagellate species Muratodinium fimbriatum,Nematosphaeropsis reticulense, Phelodinium magnificumand the terrestrial palynomorphs Proxapertites cursusand Psilatricolporites crassus. In SBC-3, this sequencecontains late Paleocene planktonic foraminifera.Sequence I is Paleocene to early Eocene in age andrepresents Supercycle TA-2 of Haq et al. (1988).

Sequence J. This sequence is also Miocene in age, butis found overlying the unconformity and, as SequenceA, also represents Supercycle TB-2 (Haq et al., 1988).

3.3. Cretaceous sequences

The seven Cretaceous sequences (B–H) interpretedwith their associated dinoflagellate assemblages are inascending order from oldest to youngest:

Sequence B. Recognized in PIC-1E and PIC-5E only,this sequence is characterized by low PMI and diver-sities, particularly in the lower part. The paleoenviron-ments represented are fluvial in the lower part and

deltaic to shallow marine in the upper part. The lowerpart corresponds to the Barranquın Formation, whilethe upper represents the basal part of El Cantil.

The sequence contains the dinoflagellate speciesApteodinium spinosum, Cepadinium ventriosum, Cribro-peridinium conjunctum, Trichodinium castaneum andXenascus plotei. Together with the spores Auritulin-asporites deltaformis, Callialasporites trilobatus, Car-diangulina crassiparietalis, Exesipollenties tumulus,Impardecispora (Trilobosporites) trioreticulosa, andMatonisporites SCI 56 of Jardine et Magloire (1965),this assemblage indicates a Barremian-early Aptianage.

Sequence C. Recognized in PIC-1E and PIC-5E only,this sequence also shows low values of PMI. Thepaleoenvironments represented are marine, probablyinner to middle neritic in the lower part and middleneritic in the upper part. The latter includes the basalMiembro Garcia of the El Cantil Formation(Gonzalez de Juana, et al., 1980; Falcon, 1988;Giovannina et al., 1994).

The sequence yields the dinoflagellate speciesApteodinium spinosum, Oligosphaeridium albertense, O.pulcherrimum, Pseudoceratium eisenackii, P. interiorense,

Figure 7. Main fossils, paleontological ages, and sequences recognized in SBC-3. For key to symbols and abbreviations, seeFigure 4.

Page 11: Palynology and sequence stratigraphy of the Cretaceous of ... (1).pdfVenezuela’s Eastern Basin. The palynological results are based mainly on the analysis of samples from the following

Palynology and sequence stratigraphy of the Cretaceous of eastern Venezuela 457

P. polymorphum, P. securigerum, Subtilisphaera terrula,and the terrestrial palynomorphs Aequitriradites spinu-losus, Araucariacites australis, Auritulinasporites delta-formis, Crybelosporites pannuceus, and Impardecispora(Trilobosporites) trioreticulosa. The age assignation isAptian-early Albian.

Sequence D. Recognized in PIC-1E, PIC-5E andPIC-3E, this sequence is characterized by a slightincrease in the values of the PMI. The paleoenviron-ments represented are marine, mostly middle neritic,with some transgressions reaching outer neriticdepths. The upper part of the sequence containscalcareous nannofossils in PIC-1E.

The sequence contains the dinoflagellate speciesCepadinium ventriosum, Chichaouadinium arabicum,Cribroperidinium diaphane, Pseudoceratium interiorense,Subtilisphaera senegalensis, S. zawia and the lowermostoccurrences of Dinopterygium cladoides, and thespores Auritulinasporites deltaformis and Elaterosporitesprotensus.

Sequence E. Recognized in PIC-1E, PIC-5E, andPIC-3E, the PMI values in this sequence are low inthe lower part and increase towards the top. Thepaleoenvironments represented are marine, mainlyinner neritic in the lower part and outer neritic tobathyal in the upper part. The Querecual Formation iswithin the upper part of this sequence. The lithologi-cal characteristics of this unit (described above), andthe content of planktonic foraminifera and calcareousnannofossils indicate that it was deposited in upperbathyal, anoxic environments, reflecting the deepestenvironments and the greatest extent of flooding inthe entire section.

The upper part of sequence E contains the lower-most occurrence of the dinoflagellate genus Dino-gymnium sp. In addition, the dinoflagellate speciesCerbia tabulata, Cribroperidinium cooksoniae and thelowermost occurrences of Palaeohystrichophora infuso-rioides are encountered. Sequence E also yields theterrestrial palynomorphs Afropollis jardinus, Corollinasp., Elaterosporites klaszi, E. verrucosus, Ephedripites sp.,and Sofrepites legouxae. The assigned age is lateAlbian-Turonian.

Sequence F. Recognized in PIC-1E, PIC-5E and PIC-3E, this sequence is characterized by a gradualincrease in PMI values in the upper part. Thepaleoenvironments represented are marine, outer tomiddle neritic. The sequence corresponds to the lowerpart of the San Antonio Formation. It yields thedinoflagellate species Canningia senonica, Cyclo-nephelium vannophorum, Kallosphaeridium helbyi,

Subtilisphaera pirnaensis and Trichodinium castaneum.Terrestrial palynomorphs are scarce to absent. Theage assignation is Turonian-early Campanian.

Sequence G. Recognized in all four wells studied, thissequence is characterized by the highest PMI values,particularly in the upper part. The paleoenvironmentsrepresented are marine, mainly middle neritic, withoccasional transgressions to outer neritic. Thesequence corresponds to the upper part of the SanAntonio Formation, which contains more sand thanthe lower part.

The sequence contains the dinoflagellate speciesAldorfia deflandrei, Andalusiella spp., Cerodinium spp.,Cometodinium whitei, Dinogymnium heterocostatum,Isabelidinium spp., Kallosphaeridium ringnesiorum,Odontochitina operculata, Palaeohystrichophora infuso-rioides, Palynodinium grallator, Senegalinium bicavatum,Spinidinium echinoideum, and Xenascus ceratiodes.Terrestrial palynomorphs are very scarce. The ageassignation is early Campanian-Maastrichtian.

Sequence H. Recognized in all four wells studied, thissequence is characterized by a sharp decrease in thePMI values as compared to the underlying sequence.It corresponds to the sandy San Juan Formation. Thepaleoenvironments represented are shallow marine todeltaic. The sequence is partially Paleocene in age,representing Supercycle TA-1.0 of Haq et al. (1988).

It contains the highest occurrence of the dinoflagel-late genus Dinogymnium sp. and the dinoflagellatespecies Andalusiella polymorpha, Circulodinium distinc-tum, Palaeocystodinium australinum, and Senegaliniumobscurum. Also present are the terrestrial paly-nomorphs Ariadnaesporites spp., Buttinia andreevi,Echitriporites trianguliformis, Foveotriletes margaritae, F.perforatus, and Proteacidites sigalii. The age assignationis late Maastrichtian-early Paleocene.

4. Ages and correlation with global record

4.1. Chronostratigraphy

Based on dinoflagellates, planktonic foraminifera, andcalcareous nannofossil analyses, it is apparent thatsediments of Barremian-Maastrichtian age (Paredeset al., 1994) are represented in the Eastern Basinof Venezuela. In general, the majority of the agesassigned are based on the stratigraphic ranges of themarine palynomorphs reported here (Figure 8).

The combination of the paleontological ages withthe sedimentary sequences defined previously, allowscorrelation with the second Order Cycles of theMesozoic-Cenozoic Cycle Chart of global eustatic

Page 12: Palynology and sequence stratigraphy of the Cretaceous of ... (1).pdfVenezuela’s Eastern Basin. The palynological results are based mainly on the analysis of samples from the following

458 J. Helenes and D. Somoza

events (Haq et al., 1988). Correlation with this chart isthe basis for the assignment of numerical ages to thesequence boundaries defined in the sections studied(Figure 9). The Cycle Chart is related to the globalcycle chart of sea-level changes proposed by Vail et al.(1977), which was constructed with results fromseismic stratigraphy studies of various regionsincluding eastern Venezuela (Vail et al., 1977, fig. 4).

The Cretaceous section from Venezuela’s EasternBasin represents marine sedimentation in a passivemargin setting, from approximately 112 Ma to about

65 Ma. Considering only the age assignments for thesequence boundaries recognized, there are 64 Marepresented in approximately 4000 m of sediments.

4.2. Correlation of sequences to supercycles

The sequences described above can be correlated tothe supercycles proposed by Haq et al. (1988). Asnoted above, Sequence A corresponds to the Miocenesedimentary rocks underlying the thrust fault andrepresents Supercycle TB-2. Sequence J is also

Figure 8. Stratigraphic ranges of the main marine palynomorph taxa found in the Eastern Basin of Venezuela.

Page 13: Palynology and sequence stratigraphy of the Cretaceous of ... (1).pdfVenezuela’s Eastern Basin. The palynological results are based mainly on the analysis of samples from the following

Palynology and sequence stratigraphy of the Cretaceous of eastern Venezuela 459

Miocene, but overlying the unconformity and alsorepresents Supercycle TB-2. Sequence I is earlyEocene and represents Supercycle TA-2, whilesequence H is partially Paleocene in age and repre-sents Supercycle TA-1. The rest of the sequences areCretaceous, and in ascending stratigraphic order are:

Sequence B. Barremian-early Aptian, corresponds tothe upper part of Supercycle LZB-3 (Haq et al., 1988)dating from 117.5 to 112.2 Ma.

Sequence C. Aptian-early Albian, corresponds toSupercycle LZB-4 (Haq et al., 1988) dating from 112to 107.5 Ma. In addition to the palynomorphsreported above, the upper half of sequence C containsAptian calcareous nannofossils in PIC-1E.

Sequence D. Albian, corresponds to Supercycle UZA-1(Haq et al., 1988) dating from 107.5 to 98 Ma. Theupper half of this sequence contains late Albiancalcareous nannofossils in PIC-1E.

Sequence E. Late Albian-Turonian, corresponds toSupercycle UZA-2 (Haq et al., 1988) dating from 98

to 90 Ma. This sequence in PIC-1E contains Albian-Cenomanian calcareous nannofossils in the lowerpart, and Cenomanian-Santonian planktonic foramin-ifera in the middle.

Sequence F. Turonian-early Campanian, correspondsto Supercycle UZA-3 (Haq et al., 1988) ranging from90 to 80 Ma. In PIC-1E, this sequence containscalcareous nannofossils indicating age-ranges fromTuronian to Santonian. Barremian-Coniacian plank-tonic foraminifera are present in the middle part of thesequence in PIC-5E.

Sequence G. Early Campanian to late Maastrichtian,corresponds to Supercycle UZA-4 (Haq et al., 1988)dating from 80 to 68 Ma.

Sequence H. Late Maastrichtian-early Paleocene, cor-responds to the lower part of Supercycle TA-1 (Haqet al., 1988) ranging from 68 to 58.5 Ma. The locationof the Cretaceous-Tertiary boundary cannot be deter-mined paleontologically because of the poor fossilrecovery from the sandy part of the San Juan For-mation. It is likely to be unconformable. In SBC-3 the

Figure 9. Cretaceous sequences and chronostratigraphy in the Eastern Basin of Venezeula. Supercycles from Haq et al.(1988)). A to J are the stratigraphic sequences recognized in this study. Lithologic symbols are the same as those inFigure 2.

Page 14: Palynology and sequence stratigraphy of the Cretaceous of ... (1).pdfVenezuela’s Eastern Basin. The palynological results are based mainly on the analysis of samples from the following

460 J. Helenes and D. Somoza

interval from 1018 to 815 m contains a mixture ofPaleocene and Maastrichtian taxa. Core samples at815 m and 817 m contain scarce specimens of theCretaceous dinoflagellate Circulodinium distinctum andthe spore Bahiaporites reticularis which we consider tobe reworked, based on their poor state of preservationand the combined presence of several Paleocenespecies, such as (Paleocene to middle Eocene)Nematosphaeropsis reticulensis and Cordosphaeridiumexilimurum (Paleocene to early Eocene).

5. Regional distribution

Based on our chronostratigraphic framework, theanalysis of the sedimentary evolution of the areastudied allows us to recognize the following regionaldistribution of some of the most importantstratigraphic units and sedimentological events.

5.1. Barremian to early Aptian

The Barremian to early Aptian non-marine to margin-ally marine sediments of the Barranquin Formationare widespread in eastern Venezuela (Erikson, 1994).Apparently these strata represent the initial stages ofan extensional regime in the area.

In western Venezuela, the Rio Negro and lower partof the Apon Formations (Lugo, 1994; Parnaud et al.,1995) represent paralic to shallow marine environ-ments. In Colombia, the Paja Formation representscompletely marine conditions related to the develop-ment of the Pacific margin of South America (Cooperet al., 1995).

5.2. Aptian transgression

The first important flooding surface observed in thesesections corresponds to the lower part of the El CantilFormation, and represents the Aptian transgressiondated at 111 Ma. In our area of study this transgres-sion attained middle neritic depths as indicated pre-viously. In the Serranıa del Interior, this transgressionis represented by the Miembro Garcıa of the El CantilFormation (Falcon, 1988), and represents outerneritic to upper bathyal environments as indicated bythe presence of common planktonic foraminifera andcalcareous nannoplankton.

In western Venezuela (Lugo, 1994; Parnaud et al.,1995) this event correlates with the Apon Formation,and in Colombia it is represented by the Caballos andTablazo Formations (Cooper et al., 1995).

5.3. Albian transgression

The Albian transgression indicated in the upper partof the El Cantil Formation is correlatable with thatdated at 98.5 Ma, which corresponds to the upperpart of Supercycle UZA-1 (Haq et al., 1988). Thisevent is closely related to the first marine transgressionobserved in the Barinas-Apure Basin of westernVenezuela by Lugo (1994), Helenes et al. (1994,1998), and Parnaud et al. (1995). In the MaracaiboBasin area of western Venezuela (Lugo, 1994;Parnaud et al., 1995) and in northwestern and centralColombia (Cooper et al., 1995), this transgressiveevent is related to the early phases of deposition of theLa Luna Formation.

5.4. Turonian transgression

The black shales of the Querecual Formation ineastern Venezuela represent deposition in bathyalenvironments (Gonzalez de Juana et al., 1980;Villamil, 1996). These strata commonly containplanktonic foraminifera and calcareous nannofossils,and are the product of one of the most importanttransgressions observed in the sections studied. Theycorrespond to the Turonian transgression (91.5 Ma)found in the upper part of Supercycle UZA-2 (Haqet al., 1988).

In western Venezuela, these strata can be correlatedto the La Morita Member of the Navay Formation ofthe Barinas-Apure Basin (Lugo, 1994; Helenes et al.,1998), and parts of the La Luna Formation in theMaracaibo Basin (Lugo, 1994; Parnaud et al., 1995)and in central and northern Colombia (Martinez &Hernandez, 1992; Cooper et al., 1995).

In Central America, there are few reports ofCretaceous sediments correlatable with these events.In Costa Rica (Sprechmann, 1984, fig. 16), mid-Cretaceous rocks in the west are related to the ophio-lites of the Nicoya Complex while in the east they havenot been reported. In Honduras (Finch, 1981), themid- to upper Cretaceous Los Angeles Group isrepresented mostly by red beds. However, theyinclude the Cenomanian Jaitique Formation, which iscomposed of thick-bedded limestones with shallowwater benthonic foraminifera in the lower part (theJaitique Member), and by thin-bedded, dark, shalylimestones with planktonic foraminifera in the upperpart (Guare Member). The Guare Member mayrepresent the Turonian transgression.

5.5. Santonian to Maastrichtian transgressions

In general, from Coniacian to Maastrichtian the areawas completely within the marine realm. However,

Page 15: Palynology and sequence stratigraphy of the Cretaceous of ... (1).pdfVenezuela’s Eastern Basin. The palynological results are based mainly on the analysis of samples from the following

Palynology and sequence stratigraphy of the Cretaceous of eastern Venezuela 461

the San Antonio Formation contains two main flood-ing surfaces; the Santonian one correlates with thecondensed section dated at 86 Ma, and that of theearly Maastrichtian correlates with the condensedsection dated at 73.5 Ma. Similar conditions alsoprevailed in western Venezuela during deposition ofthe Burguita Formation in the Barinas-Apure Basin(Lugo, 1994; Helenes et al., 1998), the upper partof the La Luna and the Colon Formations in theMaracaibo Basin (Lugo, 1994; Parnaud et al., 1995),and in much of central and northern Colombia(Martınez & Hernandez, 1992; Cooper et al., 1995).In the Llanos Basin of eastern Colombia, only one ofthese transgressions is represented by fine grainedclastics in the Guadalupe Formation (Cooper et al.,1995).

5.6. Maastrichtian-Paleocene fall in sea-level

The coarse to medium sandstones of the San JuanFormation represent deposition after a sharp fall in sealevel, corresponding to the 68 Ma sequence boundary.In western Venezuela and Colombia, this event isobscured by the tectonic accretion of the WesternCordillera in Colombia (Cooper et al., 1995).

6. Conclusions

In the relatively shallow marine environments ofdeposition commonly indicated in the sectionsstudied, dinoflagellates provide better biostratigraphicresolution than either planktonic foraminifera or cal-careous nannofossils. A Palynomorph Marine Index isuseful in identifying flooding surfaces and sequenceboundaries within these deposits.

Cretaceous strata in the subsurface of easternVenezuela have a maximum thickness of 3658 mand encompass ages ranging from Barremian toMaastrichtian. Seven stratigraphic sequences arerecognized and correlated with Supercycles LZB-3 toTA-1 of Haq et al. (1988).

The first important flooding surface correspondsto the lower part of the El Cantil Formation, andrepresents the Aptian transgression correlated to thecondensed section dated at 111 Ma. The Albiantransgression found in the upper part of the El CantilFormation is correlatable to the condensed sectiondated at 98.5 Ma. After this transgression, and formost of the Cenomanian, deposition in shallowmarine environments, with an input of large amountsof terrestrial palynomorphs, is indicated.

The most important maximum flooding surfaceobserved corresponds to the Turonian transgression(91.5 Ma) of the Querecual Formation, and can be

correlated with sections in western Venezuela andColombia.

The Santonian flooding surface in the lower part ofthe San Antonio Formation is correlated with thecondensed section dated at 86 Ma, and the earlyMaastrichtian flooding surface in the upper part of theSan Antonio Formation with the condensed sectiondated at 73.5 Ma. The San Juan Formation representsdeposition after a sharp fall in sea level, correspondingto the 68 Ma sequence boundary.

Acknowledgements

Thanks are extended to the Exploration Manage-ment of CORPOVEN (now Servicios GeologicosEspecializados de Exploracion y Produccion-PDVSA)for permission to publish this paper, and to our friendsin the laboratory, for all their help. The studywas conducted at the Laboratorio Geologico fromCORPOVEN in Puerto La Cruz, Venezuela.

References

Aurisano, R. W. 1989. Upper Cretaceous dinoflagellate bio-stratigraphy of the subsurface Atlantic Coastal Plain of NewJersey and Delaware, U.S.A. Palynology 13, 143–180.

Bellizzia, A. & Dengo, G. 1990. The Caribbean mountainsystem, northern South America; a summary. In The Caribbeanregion (eds Dengo, G. & Case, J. E.), pp. 167–175 (The Geologyof North America, vol. H; Geological Society of America,Boulder).

Below, R. 1984. Aptian to Cenomanian dinoflagellate cystsfrom the Mazagan Plateau, northwest Africa (Sites 545 and547, Deep Sea Drilling Project Leg 79) Initial Reports of the DeepSea Drilling Project 79 (eds Hinz, K., Winterer, E. L. et al.),621–649.

Burke, K., Cooper, C., Dewey, J. F., Mann, J. P. & Pindell, J. 1984.Caribbean tectonics and relative plate motions. In The Caribbean-South American plate boundary and regional tectonics (eds Bonini,W. E., Hargraves, R. B. & Shagam, R.), Geological Society ofAmerica, Memoir 62, 31–64.

Colmenares, O. & Teran, L. 1993. A biostratigraphic study ofPaleogene sequences in southwestern Venezuela. Palynology 17,67–90.

Cookson, I. C. & Eisenack, A. 1960a. Microplankton fromAustralian Cretaceous sediments. Micropaleontology 6, 1–18.

Cookson, I. C. & Eisenack, A. 1960b. Upper Mesozoic micro-plankton from Australia and New Guinea. Palaeontology 2,243–261.

Cookson, I. C. & Eisenack, A. 1962. Additional microplanktonfrom Australian Cretaceous sediments. Micropaleontology 8, 485–507.

Cooper, M. A., Addison, F. T., Alvarez, R., Coral, M., Graham,R. H., Hayward, A. B., Howe, S., Martinez, J., Naar, J., Penas,R., Pulham, A. J. & Taborda, A. 1995. Basin development andtectonic history of the Llanos Basin, Eastern Cordillera, andMiddle Magdalena Valley, Colombia. American Association ofPetroleum Geologists, Bulletin 79, 1421–1443.

Dale, B. 1996. Dinoflagellate cyst ecology: modeling and geologicalimplications. In Palynology: principles and applications, Volume 3(eds Jansonius, J. & McGregor, D. C.), pp. 1249–1275 (AmericanAssociation of Stratigraphic Palynologists Foundation, Dallas).

Davey, R. J. 1978. Marine Cretaceous palynology of site 361,DSDP Leg 40, off southwestern Africa. Initial reports of the Deep

Page 16: Palynology and sequence stratigraphy of the Cretaceous of ... (1).pdfVenezuela’s Eastern Basin. The palynological results are based mainly on the analysis of samples from the following

462 J. Helenes and D. Somoza

Sea Drilling Project 40 (eds Bolli, H. M., Ryan, W. B. F. et al.),883–913.

Erikson, J. P. 1994. A Lower Cretaceous shelf and delta in theEastern Venezuela Basin. Memorias del V Simposio Bolivariano—Exploracion Petrolera en cuencas Subandinas, pp. 174–189(Sociedad Venezolana de Geologicos, Caracas).

Evitt, W. R. 1985. Sporopollenin dinoflagellate cysts—their morphologyand interpretation, 333 pp. (American Association of StratigraphicPalynologists Foundation, Dallas).

Falcon, R. A. 1988. Estratigrafia de las formaciones Barranquin yGarcia en el flanco sur de la Serrania del Interior Oriental deVenezuela. Unpublished thesis, Universidad Central deVenezuela, 186 pp.

Fasola, A. & Paredes-de-Ramos, I. 1991. Late Cretaceous palyno-logical assemblages from El Furrial area wells. Revista Tecnica deINTEVEP 11 (1), 3–13.

Finch, R. C. 1981. Mesozoic stratigraphy of central Honduras.American Association of Petroleum Geologists, Bulletin 65,1320–1333.

Giovannina, R., Falcon, R., Salazar, M. & Gonzales, G. 1994.Analisis secuencial de las formaciones Barranquin y El Cantil enel flanco sur de la Serrania del Interior Oriental de Venezuela.Memorias del V Simposio Bolivariano-Exploracion Petrolera en cuen-cas Subandinas, pp. 20–22 (Sociedad Venezolana de Geologos,Caracas).

Germeraad, J. H., Hopping, C. A. & Muller, J. 1968. Palynology ofTertiary sediments from tropical areas. Review of Palaeobotanyand Palynology 6, 189–348.

Gonzalez de Juana, C., Iturralde de Arozena, J. M. & PicardCadillat, X. 1980. Geologıa de Venezuela y de sus cuencaspetrolıferas, Tomos 1 y II, 1030 pp (Ediciones FONINVES,Caracas).

Haq, B. U., Hardenbol, J. & Vail, P. R. 1988. Mesozoic andCenozoic chronostratigraphy and eustatic cycles. In Sea-levelchanges: an integrated approach (eds Wilgus, C. K., Posamentier,H., Ross, C. A. & Kendall, G. St. C.), SEPM (Society forSedimentary Geology) Special Paper 42, 71–108.

Helby, R., Morgan, R. & Partridge, A. D. 1987. A palynologicalzonation of the Australian Mesozoic. In Studies in AustralianMesozoic palynology (ed. Jell, P. A.), Association of AustralasianPalaeontologists, Memoir 4, 1–85.

Helenes, J. 1984. Dinoflagellates from Cretaceous to Early Tertiaryrocks of the Sebastian Vizcaıno Basin, Baja California, Mexico. InGeology of the Baja California Peninsula (ed. Frizzell, V. A. Jr),Proceedings of the Pacific Section of the SEPM 39, 80–106.

Helenes, J., Somoza, D. & Vasquez, J. 1991. Upper Cretaceousdinoflagellates, paleoenvironments and stratigraphic sequences ofEastern Venezuela. Abstracts, 24th Annual Meeting, AmericanAssociation of Stratigraphic Palynologists, San Diego.

Helenes, J., de-Guerra, C. & Vasquez, J. 1998. Palynology andchronostratigraphy of the Upper Cretaceous in the subsurface ofthe Barinas area, western Venezuela. American Association ofPetroleum Geologists, Bulletin 82, 1308–1328.

Helenes-Escamilla, J., de-Guerra, C. & Vasquez, J. 1994.Estratigrafia por secuencias del Cretacico Superior en el subsuelodel area de Barinas. Memorias del V Simposio Bolivariano—Exploracion Petrolera en cuencas Subandinas, pp. 29–39 (SociedadVenezolana de Geologicos, Caracas).

Heusser, L. E. 1978. Spores and pollen in the marine realm.In Introduction to marine micropaleontology (eds Haq, B. V. &Boersma, A.), pp. 327–339 (Elsevier, New York).

Hoffmeister, W. S. 1954. Microfossil prospecting for petroleum.U.S. Patent: 2,686,108.

Jain, K. P. & Millepied, P. 1973. Cretaceous microplankton fromSenegal Basin, NW Africa, 1. Some new genera, species andcombinations of dinoflagellates. Palaeobotanist 20, 22–32.

Lentin, J. K. & Williams, G. L. 1980. Dinoflagellate provincial-ism with emphasis on Campanian peridiniaceans. AmericanAssociation of Stratigraphic Palynologists, Contributions Series 7,1–47.

Lugo, J. 1994. Arco de Merida: control tectonico sobre lasedimentacion desde el Mesozoico Tardıo hasta el CenozoicoTemprano en el Occidente Venezolano. Memorias del V SimposioBolivariano—Exploracion Petrolera en cuencas Subandinas,pp. 291–310 (Sociedad Venezolana de Geologicos, Caracas).

Malloy, R. E. 1972. An Upper Cretaceous dinoflagellate cystlineage from Gabon, West Africa. Geoscience and Man 4, 57–65.

Martınez, R. J. I. & Hernandez, R. 1992. Evolution and drowningof the Late Cretaceous Venezuelan carbonate platform. Journal ofSouth American Earth Sciences 5, 197–210.

Molnar, P. & Sykes, L. R. 1969. Tectonics of the Caribbean andMiddle America regions from focal mechanisms and seismicity.Geological Society of America, Bulletin 80, 1639–1684.

Morris, A. E. L., Meyerhoff, A. A., Taner, I., Bueno-Salazar, R. &Young, G. A. 1990. Energy resources of the Caribbean region. InThe Caribbean region (eds Dengo, G. & Case, J. E.), pp. 483–510(The Geology of North America, vol. H; Geological Society ofAmerica, Boulder).

Muller, J. 1959. Palynology of Recent Orinoco Delta and shelfsediments. Micropaleontology 5, 1–32.

Muller, J., Di Giacomo, E. & Van Erve, A. W. 1987. A palynologi-cal zonation for the Cretaceous, Tertiary and Quaternary ofnorthern South America. American Association of StratigraphicPalynologists, Contributions Series 19, 7–76.

Pares-Regali, M. S., Uesugui, N. & Santos, A. D. S. 1974.Palinologia dos sedimentos meso-cenozoicos do Brasil (II).Boletim Tecnico da Petrobras 17, 263–301.

Paredes, I., Guzman, J. I., Helenes, J., Carrillo, M., De Cabrera, S.,Crux, J., Castro-Mora, M. & Luna, F. 1994. Fluctuaciones delnivel del mar entre el Barremiense y el Paleogeno, CuencaOriental de Venezuela. Memorias del V Simposio Bolivariano—Exploracion Petrolera en cuencas Subandinas, pp. 234–236 (Socie-dad Venezolano de Geologicos, Caracas).

Parnaud, F., Gou, Y., Pascual, J. C., Capello, M. A., Truskowski,I. & Passalacqua, H. 1995. Stratigraphic synthesis of westernVenezuela. In Petroleum basins of South America (eds Tankard,A. J. & Suarez, S. R.), American Association of Petroleum Geologists,Memoir 62, 681–698.

Pindell, J. L. 1985. Alleghenian reconstruction and the subsequentevolution of the Gulf of Mexico, Bahamas, and Proto-CaribbeanSea. Tectonics 4, 1–39.

Pindell, J. L. & Barrett, S. F. 1990. Geological evolution of theCaribbean region; a plate-tectonic perspective. In The Caribbeanregion (eds Dengo, G. & Case, J. E.), pp. 405–432 (The Geologyof North America, vol. H; Geological Society of America,Boulder).

Pindell, J. L. & Erikson, J. P. 1993. The Mesozoic passive margin ofnorthern South America. In Cretaceous tectonics in the Andes (ed.Vogel, A.), pp. 1–30 (International Monograph Series, EarthEvolution Sciences, Vieweg Publishing, Wiesbaden, FRG).

Powell, A. J. (ed.) 1992. A stratigraphic index of dinoflagellate cysts,290 pp. (Chapman & Hall, London).

Schubert, C. 1984. Basin formation along the Bocono-Moron-ElPilar Fault System, Venezuela. Journal of Geophysical Research 89,5711–5718.

Sprechmann, P. (ed.) 1994. Manual de geologıa de Costa Rica.320 pp. (Editorial de la Universidad de Costa Rica, San Jose).

Stephan, J. F., Beck, C., Chaplet, M., Sauvage, J. F., Vivas, V.,Bellizzia, A., Saint-Marc, P., Bulot, L., Arnaud, H., Arnaud-Vanneau, A., Ivaldi, J. P., Chorowicz, J., Muller, C., Masse, J. P.,Iatzoura, A., Dubar, M. & Macsotay, O. 1994. Historia sedimen-taria y tectonica de la parte Oriental de la Serranıa del InteriorOriental (Venezuela). Memorias del V Simposio Bolivariano—Exploracion Petrolera en cuencas Subandinas, pp. 267–268(Sociedad Venezolana de Geologicos, Caracas).

Stover, L. E. & Helby, R. 1987. Some Australian Mesozoicmicroplankton index species. In Studies in Australian Mesozoicpalynology (ed. Jell, P. A.), Association of AustralasianPalaeontologists, Memoir 4, 101–134.

Page 17: Palynology and sequence stratigraphy of the Cretaceous of ... (1).pdfVenezuela’s Eastern Basin. The palynological results are based mainly on the analysis of samples from the following

Palynology and sequence stratigraphy of the Cretaceous of eastern Venezuela 463

Vail, P. R., Mitchum, R. M. & Thompson, S. III. 1977. Seismicstratigraphy and global changes of sea level, part 4. Global cyclesof relative changes of sea level. In Seismic stratigraphy—applicationto hydrocarbon exploration (ed. Payton, C. E.), AmericanAssociation of Petroleum Geologists, Memoir 26, 83–98.

Vail, P. R., Audemard, F., Bowman, S. A., Eisner, P. N. &Perez-Cruz, C. 1991. The stratigraphic signatures of tectonics,eustacy and sedimentology—an overview. In Cycles and events instratigraphy (eds Einsele, G., Ricken, W. & Seilacher, A.)pp. 617–659 (Springer-Verlag, Berlin).

Villamil, T. 1996. Paleobiology of two new species of the bivalveAnomia from Colombia and Venezuela and the importance of thegenus in recognition of the base of the Turonian. CretaceousResearch 17, 607–632.

Williams, G. L. & Bujak, J. P. 1985. Mesozoic and Cenozoicdinoflagellates. In Plankton stratigraphy (eds Bolli, H. M.,Saunders, J. B. & Perch-Nielsen, K.), pp. 847–964 (CambridgeUniversity Press, Cambridge).

Williams, G. L., Stover, L. E. & Kidson, E. J. 1993. Morphologyand stratigraphic ranges of selected Mesozoic-Cenozoic dino-flagellate taxa in the northern hemisphere. Geological Survey ofCanada, Paper 92-10, 1–137.

Wood, G. D., Gabriel, A. M. & Lawson, J. C. 1996. Palynologicaltechniques—processing and microscopy. In Palynology: principlesand applications, volume 1 (eds Jansonius, J. & McGregor, D. C.),pp. 29–50 (American Association of Stratigraphic PalynologistsFoundation, Dallas).