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Plant Internal Structure and Growth
BAMBOO is said to be the fastest
growing plant. It can grow 3 to 4
feet in height in one day. The General
Sherman Sequoia is the largest living
organism on earth. It is 275 feet tall
and weighs 1,385 tons. How these and
other plants grow is discussed in this
unit.
Objectives:
� 1. Describe plant cells and
tissues.
2. Explain the processes of plant growth.
Key Terms:
�
E-unit: Plant Internal Structure and Growth
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apical meristem
bud primordia
cell
cell differentiation
cell division
cell elongation
cell wall
cellulose
chlorophyll
chloroplasts
chromatin
chromosomes
collenchyma cells
companion cells
cork cambium
cork cells
cytoplasm
dermal tissues
endoplasmic
reticulum (ER)
fiber
genes
golgi complex
ground tissue
lateral meristem
leaf primordia
lignin
meristems
mitochondria
nucleolus
nucleus
organelles
parenchyma cells
periderm
phloem
pith
plasma
membrane
plastids
primary plant
growth
ribosomes
rough
endoplasmic
reticulum
sclereid
sclerenchyma
cells
secondary plant
growth
sieve tube
member
smooth
endoplasmic
reticulum
tissues
tracheid cells
vacuole
vascular
cambium
vascular tissues
vessel elements
xylem
Plant Cells and Tissues
The cell is the most basic unit of life. It is the smallest unit that can carry out functions of
life. Interconnected groups of cells that perform similar functions are tissues.
COMPONENTS OF A PLANT CELL
A plant cell is made up of a cell wall, a plasma membrane, cytoplasm, and a variety of
organelles.
Cell Wall
The cell wall is made of multiple layers of cellulose. Cellulose is a polysaccharide, or
complex sugar molecule. The layers of cellulose offer great strength. Cellulose is the primary
ingredient of wood. Once a cell has stopped growing, the cell wall thickens and becomes rigid.
Plasma Membrane
The feature that really defines a cell is the plasma membrane. A plasma membrane, or
cell membrane, is a thin sheet that holds the contents of the cell together and serves as a pro-
tective barrier from the surrounding environment. The fluid mosaic model of the plasma
membrane consists of a lipid bi-layer in which proteins are imbedded. While the plasma mem-
brane restricts the movement of some materials, it allows, and even initiates, the movement of
materials into and out of the cell.
Cytoplasm
Cytoplasm is the living semi-fluid material inside the cell membrane. Cytoplasm is the
home to a number of specialized structures called organelles.
Organelles
Organelles are small structures inside the cell that carry out the physiological processes of
the organism. The organelles within a cell have thin membranes used to compartmentalize or
isolate different conditions from other organelles. Within each organelle is a different
intracellular pH, different enzyme systems, and other differences. This enables the cell to carry
out different metabolic activities at the same time.
Plastids are organelles that produce and store food. Chloroplasts are a type of plastid
that contains green pigments called chlorophyll. Chlorophyll traps light energy for photosyn-
thesis. Chloroplasts are found in cells exposed to light. They are abundant in leaves and absent
from root cells. Chromoplasts are plastids that give flowers and fruits their color.
The endoplasmic reticulum (ER) is an internal network of membranes extending
throughout the cytoplasm. The endoplasmic reticulum contains many types of enzymes that
catalyze different types of chemical reactions. There are two distinct forms of endoplasmic
E-unit: Plant Internal Structure and Growth
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reticulum, the smooth and the rough endoplasmic reticulum. The smooth endoplasmic
reticulum is the site for the production of lipids (fats) and hormones. It is also a site that
breaks down toxic chemicals. The rough endoplasmic reticulum produces the proteins for
the cell secretion and cell membrane.
Proteins are processed, sorted, or modified in the golgi complex. These processes result
in the complex molecules needed for plant growth. Many proteins manufactured in the ER
pass through the golgi complex.
E-unit: Plant Internal Structure and Growth
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Vacuole
Chloroplast
Smoothendoplasmic
reticulum
Roughendoplasmic
reticulum
Cell wall
Golgiapparatus
Mitochondrion
Nucleus
Nuclearmembrane
FIGURE 1. Detailed plant cell.
Mitochondria convert sugars
into energy through cellular res-
piration. It is with mitochondria
that the plant is able to convert
the food it has made and stored
back into a form of energy.
The nucleus is the command
center of the cell. It holds the
genetic information for a cell’s
activities within a nuclear mem-
brane. The nucleus is surrounded
by a nuclear membrane. Flowing
through the nuclear membrane
are materials needed for protein
synthesis and fluids carrying chemical messages between the cytoplasm and the nucleus. Chro-
mosomes contained in the nucleus hold the genetic blueprint of the organism and direct all
functions of the cell, including protein synthesis. Chromosomes are made of chromatin.
Chromatin is a complex of deoxyribonucleic acid (DNA) molecules (35%), RNA (5%), and
protein (60%). Segments of chromosomes, or genes, are units of hereditary data that dictate
the activity and structure of a cell. Each chromosome contains thousands of genes. A nucleo-
lus is a specialized structure in a nucleus that is the site of ribosome synthesis.
Ribosomes are beadlike structures found on the endoplasmic reticulum. The ribosomes
are the major site for the assembly of amino acids into proteins.
The vacuole is a large sac bound by a membrane. It may occupy up to 90 percent of the
cell. It serves to transport and store water, foods, salts, minerals, pigments, proteins, and
wastes.
TYPES OF PLANT CELLS
Genetic information and environmental conditions determine the types of cells that a plant
produces. With the appropriate signals, various types of cells are produced.
Parenchyma Cells
Parenchyma cells are found throughout a plant and typically lack a secondary cell wall
associated with rigid strength for support. The palisade layer and spongy mesophyll cells in
leaves are parenchyma cells. Modified parenchyma cells make up a large portion of the pith
(center part) of a stem. There, the parenchyma cells serve as storage facilities for starches, oils,
water, and salts. The pith is very evident in corn and sugar cane.
Collenchyma Cells
Collenchyma cells are located under the epidermis of the stem, along leaf veins, and at
corners of angular stems. They have thick, yet flexible, cell walls that provide structural sup-
port for the plant.
E-unit: Plant Internal Structure and Growth
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Nucleolus
Nuclearpores
Chromatin
Nuclearenvelope
FIGURE 2. The nucleus.
Sclerenchyma Cells
Sclerenchyma cells are found throughout a plant. Their role is one of support. They
have a hard, thick secondary cell wall composed, in part, of a material called lignin. Lignin is a
molecule that provides strength and rigidity to the cell. A long type of sclerenchyma cell is
called fiber. Fiber cells are abundant in wood and bark. A form of sclerenchyma cell that is
short and cubical is the sclereid. It is found in the shell of nuts and the stones of fruit, such as
peaches.
Tracheid Cells
Tracheid cells are one of four types of cells found in the xylem. The others are vessel ele-
ments, parenchyma, and fibers. Tracheids are long and tapering in shape. They are dead cells
and hollow. Their role is to conduct water and minerals throughout the plant. Water passes
from one cell to another through thin places in the cell wall, called pits. Tracheids also contrib-
ute to the structural support of the plant.
Vessel Elements
The vessel elements are found in the xylem, where they conduct water and minerals
and provide structural support for the plant. Vessel element cells are hollow. At the end of the
cell walls there are perforations or holes that allow the free flow of water from one cell to the
next. Vessel cells also have pits along the sides of the cells that permit the lateral movement of
water.
Sieve Tube Members
The sieve tube member is a major
element of the phloem. Other cells that
make up phloem tissue are companion
cells, parenchyma cells, and fibers. Sieve
tube members are stacked end to end to
form sieve tubes. There are holes at the
end of each cell, called sieve plates,
through which sugars are moved. Sieve
tube members are living cells at maturity,
but many organelles, including the
nucleus, vacuole, and ribosomes, vanish
as the cell matures.
Companion Cells
Companion cells are associated
with sieve tube members. They have
many connections with adjacent sieve
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Pores of sieve plate
Sieve plate
Sieve tube member
Companion cell
FIGURE 3. Major elements of the phloem.
tube members. Companion cells do not conduct sugar throughout the plant. They do, how-
ever, play an important role in transporting sugars from photosynthetic parenchyma cells to
the sieve tube members.
Cork Cells
Cork cells are produced to the outside of the cork cambium and result in the outer cover-
ing of secondary growth. Cork cells have waterproof characteristics and fire retardant attrib-
utes.
TYPES OF TISSUES
Vascular plants have three types of mature tissues. They are the dermal (protective) tissues,
the vascular tissues, and the ground (fundamental) tissues.
Dermal Tissues
Dermal tissues consist of
the epidermis (external covering
of the leaves), stems, and roots.
Dermal tissues offer a barrier to
infectious organisms and other
invaders.
The epidermis is made of
closely packed cells that secrete a
waxy cuticle to reduce water loss.
An epidermis is associated with
plants that exhibit primary
growth. The epidermis is a single
layer of cells. Specialized epider-
mal cells, called guard cells, control the opening and closing of the stomata, through which the
diffusion of carbon dioxide, oxygen, and water vapor is allowed.
Dermal tissue on secondary woody growth is a few, to many, cells thick and forms the
familiar bark of trees.
Vascular Tissues
Vascular tissues are the conductive vessels of the plant, and they transport water, miner-
als, foods, and hormones. Specialized vascular tissues include the xylem, which conducts
water and minerals upward from the roots, and the phloem, which transports food.
Ground Tissue
Ground tissue includes all the parts of the plant other than dermal or vascular tissues.
The bulk of a plant consists of ground tissue made up of parenchyma, collenchyma, and
sclerenchyma cells.
E-unit: Plant Internal Structure and Growth
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Water fromstem
Waterto stem
Cuticle
Xylem
Phloem
Spongymaterial
Epidermis
StomataGuard cell
Air space
FIGURE 4. A cross section of a leaf.
Plant Growth
The growth of a plant takes place in specialized areas of the plant called the meristems.
The meristem tissue consists of unspecialized cells capable of mitosis. There are two types of
growth resulting from cell division in meristem regions. They are termed primary and second-
ary growth.
CELL DIVISION, CELL ELONGATION, AND
CELL DIFFERENTIATION
Growth of higher plants takes place through three essential processes. These are cell divi-
sion, cell elongation, and cell differentiation.
Cell Division
Cell division is simply the increase in the number of cells. This is accomplished through
mitosis.
Cell Elongation
Cell elongation is the enlargement of the individual cells. The elongation of individual
plant cells results in growth of the entire tissue or organ. Both auxins and gibberellins are
responsible for cell elongation.
Cell Differentiation
Cell differentiation allows
cells to take on specific functions.
Cell differentiation is important
to multi-cellular plants, whereas
single-cell plants have no need to
have different types of cells. The
cells of the higher plants are pro-
grammed to become specialized
and make up the various plant
organs. Some cells specialize as
root cells that absorb water and
nutrients. Some conduct water
and minerals throughout the
plant. Others become leaf cells
containing chloroplasts and func-
tion to produce food for the plant.
E-unit: Plant Internal Structure and Growth
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Root hairs
Xylem
Phloem
Apical meristem
AREA OF CELL
DIFFERENTIATION
AREA OF CELL
ELONGATION
AREA OF CELL
DIVISION
Root cap
Developing
secondary
root
FIGURE 5. Cell division, elongation, and differentiation.
PRIMARY PLANT GROWTH
Primary plant growth occurs in areas called apical meristems. Apical meristem tissue
is found at the tips of roots and at the end of stems. Growth at the apical meristem increases
the length of the plant. Primary growth occurs in both herbaceous and woody plants.
The meristem of a root is located right behind the root cap. The root cap is a layer of cells
that protect the meristem as the root grows through the soil. Just behind the root cap is the
area of cell division. Further back is an area of cell elongation. As the cells in the root mature,
they differentiate to perform specific functions. In a root system, the cells might become root
hairs, xylem, phloem, and so on.
The meristem of the shoot has areas of cell division, cell elongation, and cell maturation.
The structure of the stem apical meristem differs from the root apical meristem in that it has
embryonic leaves known as leaf primordia and embryonic buds called bud primordia. As
the cells in these immature structures divide, elongate, and mature, they become the leaves and
stems of the plant.
SECONDARY PLANT GROWTH
Secondary plant growth takes place only
in woody plants and occurs in the lateral
meristem. Lateral meristem tissue is found on
the sides of roots and stems. It involves cell divi-
sion in layers ringing the stem. The result is an
increase in the width of the stem or trunk of the
plant. Two lateral meristems involved in sec-
ondary growth are the vascular cambium and
cork cambium.
Vascular cambium is a layer of
meristematic tissue found between the wood
and the bark. It produces secondary xylem to the
inside of the vascular cambium and secondary
phloem to the outside. Cell division occurs only
when the plant is actively growing, primarily
during the spring and summer.
The cork cambium is located in the outer bark region, and the cells produced there form the
periderm, or the outer bark. Cork cambium produces cork cells that replace the epidermis present
during primary growth. The periderm ranges from several cells in thickness to many cells.
Summary:
� The cell is the most basic unit of life. Plant cells are made up of cell walls, mem-branes, cytoplasm, and organelles, including plastids, endoplasmic reticulum, golgicomplex, mitochondria, nucleus, ribosomes, and vacuole.
E-unit: Plant Internal Structure and Growth
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Periderms with their cork cambia
Inner bark (secondary phloem)Outer bark
Wood (secondary xylem)
Surface of vascular cambium
Bark
FIGURE 6. Secondary growth occurs in the vascular
cambium of a tree.
Types of cells include parenchyma cells, collenchyma cells, sclerenchyma cells,tracheid cells, vessel elements, sieve tube members, companion cells, and cork cells.
Interconnected groups of cells that perform similar functions are tissues. Threetypes of mature tissues found in vascular plants are the dermal (protective) tissues,the vascular tissues, and the ground (fundamental) tissues.
The growth of a plant takes place in specialized areas of the plant called themeristems. Growth of higher plants takes place through cell division, cell elonga-tion, and cell differentiation.
Primary plant growth occurs in areas called apical meristems. Secondary plant growthtakes place only in the lateral meristem of woody plants. Two lateral meristemsinvolved in secondary growth are the vascular cambium and cork cambium.
Checking Your Knowledge:
� 1. What are the components of a plant cell?
2. What are the types of plant cells?
3. What are three types of mature tissues found in vascular plants?
4. Where does plant growth occur?
5. How do primary plant growth and secondary plant growth differ?
Expanding Your Knowledge:
� Ask your teacher or a biology teacher if they have prepared slides showing differenttypes of cells and meristematic tissues. View these under a microscope. Obtain across section of a woody trunk. Sand it smooth and determine the lateral meristemand cell types.
Web Links:
� Plant Cell Types
http://www.biologie.uni-hamburg.de/b-online/library/plant_biology/celltypes.html
Plant Cells and Tissues
http://arnica.csustan.edu/boty1050/Tissues/tissues.htm
Secondary Growth
http://plantphys.info/Plant_Biology/secondary.html
Primary and Secondary Plant Growth
http://www.cas.muohio.edu/~meicenrd/mudescd/Dendrology/SCDRYGTH/1&2gwrtp2.html
Agricultural Career Profiles
http://www.mycaert.com/career-profiles
E-unit: Plant Internal Structure and Growth
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