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Ch. 29 Warm-Up
1. Describe the process of how H2O gets into the plant and up to the leaves.
2. Compare and contrast apoplastic flow to symplastic flow.
3. Explain the mass flow of materials in the phloem (source to sink).
Ch. 29 Warm-Up
1. What is transpiration?
2. What are mycorrhizae?
3. What is the function of the Casparian strip?
Chapter 29Resource Acquisition, Nutrition, and Transport in
Vascular Plants
Plant Nutrition
Adaptations for Acquiring Resources
Shoots = capture light efficiently
Roots = acquire water and minerals
Nutritional Requirements
Essential element: required for plant to
complete life cycle and produce another
generation
Macronutrients (large amounts): CHNOPS +
K, Ca, Mg
◦Nitrogen = most important!
Micronutrients (small amounts): Fe, Mn, Zn,
Cu, etc.
Mutualistic Relationships:
1. Rhizobium bacteria supply nitrogen at roots
(fix atmospheric N2 to usable N)
• Plant supplies sugar & amino acids
2. Mycorrhizae (plant + fungus)
Role of soil bacteria in the nitrogen
nutrition of plants
Unusual nutritional adaptations:
epiphytes, parasitic plants, canivorous
plants
Epiphyte: grow on
surface of another
plant, absorb H2O from
rain through leaves
Parasitic Plants: not photosynthetic; absorb
sugar and minerals from living hosts
Carnivorous Plants: photosynthetic, but obtain
some nitrogen and minerals by digesting small
animals; found in nitrogen-poor soil
Transport Processes
Vascular Tissues: conduct molecules
Xylem Phloem
Nonliving functional Living functional
Xylem sap = H2O &
minerals
Phloem sap = sucrose,
minerals, amino acids,
hormones
Source to sink
(sugar made) to (sugar
consumed/stored)
Transport pathways in plant tissues:
Apoplast = materials travel between cells
Symplast = materials cross cell membrane, move
through cytosol & plasmodesmata
TRANSPORT OF WATER
Selectively permeable membrane: control
movement of substances in/out of cells
Osmosis: passive transport of water
Proton pump: active transport; uses E to
pump H+ out of cell proton gradient
Cotransport: couple H+ diffusion with
sucrose transport
Aquaporin: channel protein which
controls H2O uptake/loss
Review:
Solute transport
across plant cell
plasma membranes
Osmosis
**Water potential (ψ): H2O moves from high ψ
low ψ potential, solute conc. & pressure
◦ Water potential equation: ψ = ψS + ψP
◦ Solute potential (ψS) – osmotic potential
◦ Pressure potential (ψP) – physical pressure on
solution
◦ Pure water: ψS = 0 Mpa
◦ Ψ is always negative!
◦ Turgor pressure = force on cell wall
Bulk flow: move H2O in plant from regions of high
pressure low pressure
** Review AP Bio Investigation 4 (Osmosis & Diffusion)
Turgid: firm (healthy plant)
Flaccid: limp (wilting)
Plasmolysis: cell membrane shrinks & pulls
away from cell wall due to H2O loss; kills
most plant cells
Turgid Plant Cell Plasmolysis
Plant roots absorb essential elements
from soil
Root hairs: increase surface area of
absorption at root tips
Mycorrhizae: symbiotic relationship
between fungus + roots
◦ Increase H2O/mineral absorption
The white mycelium of the fungus ensheathes these roots of a pine tree.
Transport of H2O and minerals into xylem:
Root epidermis cortex [Casparian Strip]
vascular cylinder xylem tissue shoot system
Casparian strip controls entry into
vascular cylinder
How does material move vertically (against gravity)?
Transpiration: loss of H2O via
evaporation from leaves into air
Cohesion-tension hypothesis:
◦ Transpiration provides pull
◦ Cohesion of H2O transmits
pull from roots shoots
◦ Adhesion of H2O to xylem
walls (fight gravity)
◦ Negative pressure of xylem
sap (tension)
Stomata regulate rate of transpiration Stomata – pores in epidermis of leaves/stems, allow gas
exchange (CO2 in, O2 out) and transpiration (H2O out)
Guard cells – open/close stoma by changing shape
◦ Take up K+ lowers ψ take up H2O pore opens
◦ Lose K+ lose H2O cells less bowed pore closes
Cells stimulated to open by: light, loss of CO2 in
leaf, circadian rhythms
Stomata closure: drought, high temperature,
wind
BIOFLIX: WATER TRANSPORT
IN PLANTS
TRANSPORT OF SUGAR
Sugar Transport
Translocation: transport of sugars into phloem
by pressure flow
Source Sink
◦ Source = produce sugar (leaf photosynthesis)
◦ Sink = consume/store sugar (fruit, roots)
Via sieve-tube elements
Active transport of sucrose
Bulk flow by
positive pressure
flow in a sieve
tube
(active transport)
Material movement via
symplast is dynamic
Plasmodesmata allows movement of RNA &
proteins between cells
Phloem can carry rapid, long-distance electrical
signaling
◦ Nerve-like function
◦ Swift communication
◦ Changes in gene expression, respiration,
photosynthesis